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<document id="342D4591B6005BBDC64B1A43DD7CE072" ID-CLB-Dataset="36842" ID-DOI="10.11646/zootaxa.3808.1.1" ID-GBIF-Dataset="154a67eb-e80e-451e-8afd-497de87d068a" ID-ISSN="1175-5326" ID-Zenodo-Dep="286171" ID-ZooBank="E4C0E93F-DBAB-4788-A8F7-C03E2C6F0306" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460429779140" checkinUser="plazi" docAuthor="Boumans, Louis &amp; Murányi, Dávid" docDate="2014" docId="B023F04EFFCEFF80D2ADFEC3FA273EBD" docLanguage="en" docName="zt03808p091.pdf" docOrigin="Zootaxa 3808 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Zwicknia komica Muranyi &amp; Boumans" docType="treatment" docVersion="8" lastPageNumber="14" masterDocId="4C1A8836FFC4FF8DD23AFF88FFBB3F38" masterDocTitle="Two new species of Zwicknia Murányi, with molecular data on the phylogenetic position of the genus (Plecoptera, Capniidae)" masterLastPageNumber="91" masterPageNumber="1" pageNumber="11" updateTime="1698331207130" updateUser="ExternalLinkService">
<mods:mods id="C0D8F94D3AEF4527212C217E7D7DDE80" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CD789FCB45375A1269393C34C366FB6A">
<mods:title id="7E3F52B792C3F574D1A24769914FD168">Two new species of Zwicknia Murányi, with molecular data on the phylogenetic position of the genus (Plecoptera, Capniidae)</mods:title>
</mods:titleInfo>
<mods:name id="F3D6D5CAE3CC973958EC000E553D9CBC" type="personal">
<mods:role id="92D24B0846910B7D00A273201A793F27">
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<mods:namePart id="4FE80EDA7235B464B7EDC1A305F2ED95">Boumans, Louis</mods:namePart>
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<mods:namePart id="8FF376CDF2C9180BD9C808BC07CECD10">Murányi, Dávid</mods:namePart>
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<mods:title id="45AEF18AE842909BCF3F2E0BB1343DF6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DFFDD0AAB47C66A318DAB65D4822D22D">
<mods:date id="08EB8E32487B9529710625B8BAAE7B85">2014</mods:date>
<mods:detail id="544E4ADA30AB09409978FE45C8356053" type="volume">
<mods:number id="0F253159F18836B38B7AF172165A71BF">3808</mods:number>
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<mods:number id="DF1A5C1825E8E3A6E8CB2CC1B669292E">1</mods:number>
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<mods:classification id="1D62D80C779B95D8031BE4B50B37F313">journal article</mods:classification>
<mods:identifier id="B8690A76606566E3BEB26EC8A38CC4E0" type="CLB-Dataset">36842</mods:identifier>
<mods:identifier id="C2CB0B6D7DDE8D4805C72EDA44FD3699" type="DOI">10.11646/zootaxa.3808.1.1</mods:identifier>
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<mods:identifier id="903C4A3272F7C671AD9E2EDE194AEE27" type="Zenodo-Dep">286171</mods:identifier>
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<treatment id="B023F04EFFCEFF80D2ADFEC3FA273EBD" ID-DOI="http://doi.org/10.5281/zenodo.5618748" ID-GBIF-Taxon="127668163" ID-Zenodo-Dep="5618748" LSID="urn:lsid:plazi:treatment:B023F04EFFCEFF80D2ADFEC3FA273EBD" httpUri="http://treatment.plazi.org/id/B023F04EFFCEFF80D2ADFEC3FA273EBD" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11">
<subSubSection id="709012D3FFCEFF87D2ADFEC3FE863DDC" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="38354158FFCEFF87D2ADFEC3FDC23E5E" blockId="10.[151,633,331,391]" box="[151,633,331,358]" pageId="10" pageNumber="11">
<heading id="637DF634FFCEFF87D2ADFEC3FDC23E5E" bold="true" box="[151,633,331,358]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<taxonomicName id="FF8A3ADBFFCEFF87D2ADFEC3FDC23E5E" ID-CoL="7GGKS" authority="Muranyi &amp; Boumans" authorityName="Muranyi &amp; Boumans" box="[151,633,331,358]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="komica">
<emphasis id="0AFE9D4AFFCEFF87D2ADFEC3FDC23E5E" bold="true" box="[151,633,331,358]" pageId="10" pageNumber="11">
<emphasis id="0AFE9D4AFFCEFF87D2ADFEC3FEDD3E5D" bold="true" box="[151,358,331,357]" italics="true" pageId="10" pageNumber="11">Zwicknia komica</emphasis>
Murányi &amp; Boumans
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="38354158FFCEFF87D2ADFEE6FE093EBF" blockId="10.[151,633,331,391]" box="[151,434,366,391]" pageId="10" pageNumber="11">
(
<figureCitation id="A0B15DDDFFCEFF87D2A5FEE6FEA33EBF" box="[159,280,366,391]" captionStart="FIGURES 9 – 13" captionStartId="11.[151,264,1892,1915]" captionTargetBox="[189,1402,207,1861]" captionTargetId="figure@11.[151,1436,193,1871]" captionTargetPageId="11" captionText="FIGURES 9 – 13. Male terminalia of Zwicknia komica Murányi &amp; Boumans, sp. n.; paratype, Russia, Komi Republic, Tyla-yu stream. 9: dorso-caudal view; 10: dorsal view; 11: lateral view; 12: ventral view; 13: caudal view—scale 0.5 mm." httpUri="https://zenodo.org/record/286175/files/figure.png" pageId="10" pageNumber="11">Figs. 9–13</figureCitation>
,
<figureCitation id="A0B15DDDFFCEFF87D31FFEE6FE113EBF" box="[293,426,366,391]" captionStart="FIGURES 14 – 21" captionStartId="12.[151,264,1638,1661]" captionTargetBox="[312,1266,182,1638]" captionTargetId="figure@12.[293,1293,147,1648]" captionTargetPageId="12" captionText="FIGURES 14 – 21. Male terminalia of Zwicknia westermanni Boumans &amp; Murányi, sp. n. and Z. komica Murányi &amp; Boumans, sp. n. — 14, 16, 18, 20: main epiproct sclerite (Ep-scl), lateral view; 15, 17, 19, 21: hump of tergite 9, caudal view— 14 – 17: Z. westermanni paratypes, Germany, Lower Saxony, Hohenförsiek; 18 – 19: Z. westermanni paratypes France, Burgundy, Curgy; 20 – 21: Z. komica paratype, Russia, Komi Republic, Tyla-yu stream—scale 0.5 mm." httpUri="https://zenodo.org/record/286176/files/figure.png" pageId="10" pageNumber="11">Figs. 20–21</figureCitation>
)
</paragraph>
<paragraph id="38354158FFCEFF87D2ADFE3FFE683ED3" blockId="10.[151,1436,438,491]" pageId="10" pageNumber="11">
<taxonomicName id="FF8A3ADBFFCEFF87D2ADFE3FFE523EF4" authority="Newman, 1838" authorityName="Newman" authorityYear="1838" box="[151,489,438,460]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="0AFE9D4AFFCEFF87D2ADFE3FFE883EF4" box="[151,307,438,460]" italics="true" pageId="10" pageNumber="11">Capnia bifrons</emphasis>
(Newman, 1838)
</taxonomicName>
—
<bibRefCitation id="5C1B3CA9FFCEFF87D03EFE3EFD153EF4" author="Loskutova" box="[516,686,438,460]" pageId="10" pageNumber="11" refString="Loskutova, O. (2003) Fauna of stoneflies (Plecoptera) of the European North-East Russia and its change under the influence of anthropogenic factors. In: Gaino, E. (Ed.), Research Update on Ephemeroptera &amp; Plecoptera. Universita di Perugia, Perugia, pp. 357 - 362." type="book chapter" year="2003">Loskutova 2003</bibRefCitation>
(faunistic data, at least some of the northwestern Russian populations belong to the new species)
</paragraph>
<paragraph id="38354158FFCEFF87D2ADFD9FFE863DDC" blockId="10.[151,1437,535,2037]" pageId="10" pageNumber="11">
<emphasis id="0AFE9D4AFFCEFF87D2ADFD9FFE783D08" bold="true" box="[151,451,535,560]" pageId="10" pageNumber="11">Morphological diagnosis.</emphasis>
Process of male tergite 9 high, perpendicularly elevated, 3× as wide as epiproct sclerite (
<figureCitation id="A0B15DDDFFCEFF87D2A5FDB4FE963D6C" box="[159,301,572,597]" captionStart="FIGURES 9 – 13" captionStartId="11.[151,264,1892,1915]" captionTargetBox="[189,1402,207,1861]" captionTargetId="figure@11.[151,1436,193,1871]" captionTargetPageId="11" captionText="FIGURES 9 – 13. Male terminalia of Zwicknia komica Murányi &amp; Boumans, sp. n.; paratype, Russia, Komi Republic, Tyla-yu stream. 9: dorso-caudal view; 10: dorsal view; 11: lateral view; 12: ventral view; 13: caudal view—scale 0.5 mm." httpUri="https://zenodo.org/record/286175/files/figure.png" pageId="10" pageNumber="11">Figs. 10, 13</figureCitation>
), rectangular, the membranous portion sides parallel in caudal view (
<figureCitation id="A0B15DDDFFCEFF87D66EFDB4FB0E3D6C" box="[1108,1205,572,597]" captionStart="FIGURES 9 – 13" captionStartId="11.[151,264,1892,1915]" captionTargetBox="[189,1402,207,1861]" captionTargetId="figure@11.[151,1436,193,1871]" captionTargetPageId="11" captionText="FIGURES 9 – 13. Male terminalia of Zwicknia komica Murányi &amp; Boumans, sp. n.; paratype, Russia, Komi Republic, Tyla-yu stream. 9: dorso-caudal view; 10: dorsal view; 11: lateral view; 12: ventral view; 13: caudal view—scale 0.5 mm." httpUri="https://zenodo.org/record/286175/files/figure.png" pageId="10" pageNumber="11">Figs. 13</figureCitation>
,
<figureCitation id="A0B15DDDFFCEFF87D6FEFDB4FB583D6C" box="[1220,1251,572,596]" captionStart="FIGURES 14 – 21" captionStartId="12.[151,264,1638,1661]" captionTargetBox="[312,1266,182,1638]" captionTargetId="figure@12.[293,1293,147,1648]" captionTargetPageId="12" captionText="FIGURES 14 – 21. Male terminalia of Zwicknia westermanni Boumans &amp; Murányi, sp. n. and Z. komica Murányi &amp; Boumans, sp. n. — 14, 16, 18, 20: main epiproct sclerite (Ep-scl), lateral view; 15, 17, 19, 21: hump of tergite 9, caudal view— 14 – 17: Z. westermanni paratypes, Germany, Lower Saxony, Hohenförsiek; 18 – 19: Z. westermanni paratypes France, Burgundy, Curgy; 20 – 21: Z. komica paratype, Russia, Komi Republic, Tyla-yu stream—scale 0.5 mm." httpUri="https://zenodo.org/record/286176/files/figure.png" pageId="10" pageNumber="11">21</figureCitation>
). Main epiproct sclerite (Ep-scl): narrow and acute in dorsal view (
<figureCitation id="A0B15DDDFFCEFF87D0DDFDD7FC863D40" box="[743,829,607,632]" captionStart="FIGURES 9 – 13" captionStartId="11.[151,264,1892,1915]" captionTargetBox="[189,1402,207,1861]" captionTargetId="figure@11.[151,1436,193,1871]" captionTargetPageId="11" captionText="FIGURES 9 – 13. Male terminalia of Zwicknia komica Murányi &amp; Boumans, sp. n.; paratype, Russia, Komi Republic, Tyla-yu stream. 9: dorso-caudal view; 10: dorsal view; 11: lateral view; 12: ventral view; 13: caudal view—scale 0.5 mm." httpUri="https://zenodo.org/record/286175/files/figure.png" pageId="10" pageNumber="11">Fig. 10</figureCitation>
), tip upcurved in lateral view (
<figureCitation id="A0B15DDDFFCEFF87D692FDD7FAB33D40" box="[1192,1288,607,632]" captionStart="FIGURES 9 – 13" captionStartId="11.[151,264,1892,1915]" captionTargetBox="[189,1402,207,1861]" captionTargetId="figure@11.[151,1436,193,1871]" captionTargetPageId="11" captionText="FIGURES 9 – 13. Male terminalia of Zwicknia komica Murányi &amp; Boumans, sp. n.; paratype, Russia, Komi Republic, Tyla-yu stream. 9: dorso-caudal view; 10: dorsal view; 11: lateral view; 12: ventral view; 13: caudal view—scale 0.5 mm." httpUri="https://zenodo.org/record/286175/files/figure.png" pageId="10" pageNumber="11">Figs. 11</figureCitation>
,
<figureCitation id="A0B15DDDFFCEFF87D72DFDE8FA8C3D40" box="[1303,1335,608,632]" captionStart="FIGURES 14 – 21" captionStartId="12.[151,264,1638,1661]" captionTargetBox="[312,1266,182,1638]" captionTargetId="figure@12.[293,1293,147,1648]" captionTargetPageId="12" captionText="FIGURES 14 – 21. Male terminalia of Zwicknia westermanni Boumans &amp; Murányi, sp. n. and Z. komica Murányi &amp; Boumans, sp. n. — 14, 16, 18, 20: main epiproct sclerite (Ep-scl), lateral view; 15, 17, 19, 21: hump of tergite 9, caudal view— 14 – 17: Z. westermanni paratypes, Germany, Lower Saxony, Hohenförsiek; 18 – 19: Z. westermanni paratypes France, Burgundy, Curgy; 20 – 21: Z. komica paratype, Russia, Komi Republic, Tyla-yu stream—scale 0.5 mm." httpUri="https://zenodo.org/record/286176/files/figure.png" pageId="10" pageNumber="11">20</figureCitation>
); ventral membranous section ends far before the base in lateral view, apical spines stout, distributed on membranous portion and a few also on the sclerite (
<figureCitation id="A0B15DDDFFCEFF87D075FD2FFD1E3DF8" box="[591,677,679,704]" captionStart="FIGURES 14 – 21" captionStartId="12.[151,264,1638,1661]" captionTargetBox="[312,1266,182,1638]" captionTargetId="figure@12.[293,1293,147,1648]" captionTargetPageId="12" captionText="FIGURES 14 – 21. Male terminalia of Zwicknia westermanni Boumans &amp; Murányi, sp. n. and Z. komica Murányi &amp; Boumans, sp. n. — 14, 16, 18, 20: main epiproct sclerite (Ep-scl), lateral view; 15, 17, 19, 21: hump of tergite 9, caudal view— 14 – 17: Z. westermanni paratypes, Germany, Lower Saxony, Hohenförsiek; 18 – 19: Z. westermanni paratypes France, Burgundy, Curgy; 20 – 21: Z. komica paratype, Russia, Komi Republic, Tyla-yu stream—scale 0.5 mm." httpUri="https://zenodo.org/record/286176/files/figure.png" pageId="10" pageNumber="11">Fig. 20</figureCitation>
). Male micropterous, ventral vesicle half as wide as the subgenital plate (
<figureCitation id="A0B15DDDFFCEFF87D2E7FD44FE8B3DDC" box="[221,304,716,741]" captionStart="FIGURES 9 – 13" captionStartId="11.[151,264,1892,1915]" captionTargetBox="[189,1402,207,1861]" captionTargetId="figure@11.[151,1436,193,1871]" captionTargetPageId="11" captionText="FIGURES 9 – 13. Male terminalia of Zwicknia komica Murányi &amp; Boumans, sp. n.; paratype, Russia, Komi Republic, Tyla-yu stream. 9: dorso-caudal view; 10: dorsal view; 11: lateral view; 12: ventral view; 13: caudal view—scale 0.5 mm." httpUri="https://zenodo.org/record/286175/files/figure.png" pageId="10" pageNumber="11">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="709012D3FFCEFF87D2FDFD67FB723CA0" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="38354158FFCEFF87D2FDFD67FB5F3C15" blockId="10.[151,1437,535,2037]" pageId="10" pageNumber="11">
<emphasis id="0AFE9D4AFFCEFF87D2FDFD67FECE3C30" bold="true" box="[199,373,751,776]" pageId="10" pageNumber="11">
<typeStatus id="E731FFFAFFCEFF87D2FDFD67FEB93C30" box="[199,258,751,776]" pageId="10" pageNumber="11">Type</typeStatus>
material.
</emphasis>
<typeStatus id="E731FFFAFFCEFF87D346FD67FE5F3C30" box="[380,484,751,776]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="409D01C8FFCEFF87D00AFD78FD233C30" box="[560,664,752,776]" name="Russia" pageId="10" pageNumber="11">RUSSIA</collectingCountry>
: Komi Republic, Vyl’gort, Tyla-yu stream, N 6134.699' E 5039.635',
<quantity id="FF72ECBDFFCEFF87D2ADFC9CFF693C14" box="[151,210,788,813]" metricMagnitude="1" metricUnit="m" metricValue="9.5" pageId="10" pageNumber="11" unit="m" value="95.0">95 m</quantity>
,
<date id="4C346798FFCEFF87D2E7FC9CFEE53C15" box="[221,350,788,813]" pageId="10" pageNumber="11" value="2012-04-25">25.04.2012</date>
, leg. O. Loskutova (
<collectionCode id="5E9BD99DFFCEFF87D07BFC9CFD273C14" box="[577,668,788,812]" httpUri="http://grbio.org/cool/2knt-7f1r" name="Natural History Museum, University of Oslo" pageId="10" pageNumber="11">NHMO</collectionCode>
: 10073065; used for molecular studies as No.1653).
</paragraph>
<paragraph id="38354158FFCEFF87D2FDFCB0FB723CA0" blockId="10.[151,1437,535,2037]" pageId="10" pageNumber="11">
<typeStatus id="E731FFFAFFCEFF87D2FDFCB0FE8C3C68" box="[199,311,824,848]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="2E8C8AD1FFCEFF87D37CFCBFFE1D3C68" box="[326,422,823,848]" pageId="10" pageNumber="11" type="male">5 males</specimenCount>
:
<collectingCountry id="409D01C8FFCEFF87D38EFCB0FDA73C68" box="[436,540,824,848]" name="Russia" pageId="10" pageNumber="11">RUSSIA</collectingCountry>
: Komi Republic, Vyl’gort, Tyla-yu Stream, N 6134.699' E 5039.635',
<quantity id="FF72ECBDFFCEFF87D762FCBFFA2C3C77" box="[1368,1431,823,848]" metricMagnitude="1" metricUnit="m" metricValue="9.5" pageId="10" pageNumber="11" unit="m" value="95.0">95 m</quantity>
,
<date id="4C346798FFCEFF87D2ADFCD4FEA33C4D" box="[151,280,860,885]" pageId="10" pageNumber="11" value="2012-04-25">25.04.2012</date>
, leg. O. Loskutova:
<specimenCount id="2E8C8AD1FFCEFF87D033FCD4FD973C4C" box="[521,556,860,884]" pageId="10" pageNumber="11" type="generic">2♂</specimenCount>
(
<collectionCode id="5E9BD99DFFCEFF87D005FCD4FD203C4C" LSID="urn:lsid:biocol.org:col:34991" box="[575,667,860,884]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="11">HNHM</collectionCode>
: PLP4281; used for drawings
<figureCitation id="A0B15DDDFFCEFF87D1C4FCD4FBC23C4C" box="[1022,1145,860,885]" captionStart="FIGURES 9 – 13" captionStartId="11.[151,264,1892,1915]" captionTargetBox="[189,1402,207,1861]" captionTargetId="figure@11.[151,1436,193,1871]" captionTargetPageId="11" captionText="FIGURES 9 – 13. Male terminalia of Zwicknia komica Murányi &amp; Boumans, sp. n.; paratype, Russia, Komi Republic, Tyla-yu stream. 9: dorso-caudal view; 10: dorsal view; 11: lateral view; 12: ventral view; 13: caudal view—scale 0.5 mm." httpUri="https://zenodo.org/record/286175/files/figure.png" pageId="10" pageNumber="11">Figs. 9–13</figureCitation>
,
<figureCitation id="A0B15DDDFFCEFF87D6B3FCD4FAAD3C4C" box="[1161,1302,860,885]" captionStart="FIGURES 14 – 21" captionStartId="12.[151,264,1638,1661]" captionTargetBox="[312,1266,182,1638]" captionTargetId="figure@12.[293,1293,147,1648]" captionTargetPageId="12" captionText="FIGURES 14 – 21. Male terminalia of Zwicknia westermanni Boumans &amp; Murányi, sp. n. and Z. komica Murányi &amp; Boumans, sp. n. — 14, 16, 18, 20: main epiproct sclerite (Ep-scl), lateral view; 15, 17, 19, 21: hump of tergite 9, caudal view— 14 – 17: Z. westermanni paratypes, Germany, Lower Saxony, Hohenförsiek; 18 – 19: Z. westermanni paratypes France, Burgundy, Curgy; 20 – 21: Z. komica paratype, Russia, Komi Republic, Tyla-yu stream—scale 0.5 mm." httpUri="https://zenodo.org/record/286176/files/figure.png" pageId="10" pageNumber="11">Figs. 20–21</figureCitation>
); Idem:
<specimenCount id="2E8C8AD1FFCEFF87D743FCD4FA273C4C" box="[1401,1436,860,884]" pageId="10" pageNumber="11" type="generic">3♂</specimenCount>
(
<collectionCode id="5E9BD99DFFCEFF87D2A5FC08FF403CA0" box="[159,251,896,920]" httpUri="http://grbio.org/cool/2knt-7f1r" name="Natural History Museum, University of Oslo" pageId="10" pageNumber="11">NHMO</collectionCode>
: 10073143, 10073149, 10073227; used for molecular studies as No.1654, 1655, 1660);
</paragraph>
</subSubSection>
<subSubSection id="709012D3FFCEFF87D2FDFC2CFE743988" pageId="10" pageNumber="11" type="description">
<paragraph id="38354158FFCEFF87D2FDFC2CFE9E3B1F" blockId="10.[151,1437,535,2037]" pageId="10" pageNumber="11">
<emphasis id="0AFE9D4AFFCEFF87D2FDFC2CFEEA3C85" bold="true" box="[199,337,932,957]" pageId="10" pageNumber="11">Description</emphasis>
: Head, thorax, appendages and basal segments of the abdomen as is typical for the genus. Males micropterous; females macropterous (personal communication O. Loskutova, specimens were not available for study). Body length:
<typeStatus id="E731FFFAFFCEFF87D3BEFC64FE5D3B3D" box="[388,486,1004,1029]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
<quantity id="FF72ECBDFFCEFF87D3D4FC64FDFD3B3C" box="[494,582,1004,1028]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="10" pageNumber="11" unit="mm" value="7.3">7.3 mm</quantity>
, male
<typeStatus id="E731FFFAFFCEFF87D0AAFC65FD403B3D" box="[656,763,1005,1029]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
5.5–6.0, forewing length:
<typeStatus id="E731FFFAFFCEFF87D61CFC64FB333B3D" box="[1062,1160,1004,1029]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
<quantity id="FF72ECBDFFCEFF87D6B5FC64FB5C3B3C" box="[1167,1255,1004,1028]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="11" unit="mm" value="1.2">1.2 mm</quantity>
, male
<typeStatus id="E731FFFAFFCEFF87D70BFC65FA273B3D" box="[1329,1436,1005,1029]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
<quantity id="FF72ECBDFFCEFF87D2ADFB98FE9A3B1F" box="[151,289,1040,1064]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="10" pageNumber="11" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.1–1.2 mm</quantity>
.
</paragraph>
<paragraph id="38354158FFCEFF87D2FDFBBCFC9E3A8D" blockId="10.[151,1437,535,2037]" pageId="10" pageNumber="11">
<emphasis id="0AFE9D4AFFCEFF87D2FDFBBCFE323B75" bold="true" box="[199,393,1076,1101]" pageId="10" pageNumber="11">Male terminalia</emphasis>
(
<figureCitation id="A0B15DDDFFCEFF87D3A2FBBCFDA93B74" box="[408,530,1076,1101]" captionStart="FIGURES 9 – 13" captionStartId="11.[151,264,1892,1915]" captionTargetBox="[189,1402,207,1861]" captionTargetId="figure@11.[151,1436,193,1871]" captionTargetPageId="11" captionText="FIGURES 9 – 13. Male terminalia of Zwicknia komica Murányi &amp; Boumans, sp. n.; paratype, Russia, Komi Republic, Tyla-yu stream. 9: dorso-caudal view; 10: dorsal view; 11: lateral view; 12: ventral view; 13: caudal view—scale 0.5 mm." httpUri="https://zenodo.org/record/286175/files/figure.png" pageId="10" pageNumber="11">Figs. 9–13</figureCitation>
,
<figureCitation id="A0B15DDDFFCEFF87D025FBBCFD123B74" box="[543,681,1076,1101]" captionStart="FIGURES 14 – 21" captionStartId="12.[151,264,1638,1661]" captionTargetBox="[312,1266,182,1638]" captionTargetId="figure@12.[293,1293,147,1648]" captionTargetPageId="12" captionText="FIGURES 14 – 21. Male terminalia of Zwicknia westermanni Boumans &amp; Murányi, sp. n. and Z. komica Murányi &amp; Boumans, sp. n. — 14, 16, 18, 20: main epiproct sclerite (Ep-scl), lateral view; 15, 17, 19, 21: hump of tergite 9, caudal view— 14 – 17: Z. westermanni paratypes, Germany, Lower Saxony, Hohenförsiek; 18 – 19: Z. westermanni paratypes France, Burgundy, Curgy; 20 – 21: Z. komica paratype, Russia, Komi Republic, Tyla-yu stream—scale 0.5 mm." httpUri="https://zenodo.org/record/286176/files/figure.png" pageId="10" pageNumber="11">Figs. 20–21</figureCitation>
): Process of tergite 9 high, perpendicularly raised, its apex is three times as wide as the medial section of the main epiproct sclerite (Ep-scl); its shape is rectangular, the apex bearing two hump-like apices which are visible only in dorsal view; sides slightly indented in caudal view, the membranous portion parallel. Tergite 10, basal and laterobasal epiproct sclerites as is typical for the genus. Main epiproct sclerite narrow and pointed in dorsal view, medially not swollen; its medial width is half the basal width; tip upcurved and moderately acute in lateral view, divided section long. Ventral membranous part between the left and right divisions of the main epiproct sclerite ends far before the base in lateral view; apical spines stout, distributed on membranous portion, few spines occur on the sclerite. Inner sclerite (I-scl) as is typical for the genus, eversible crest (Ec) long but contorted on the studied specimens. Sternite 9 slightly projecting medially, vesicle rather large, half as wide as the subgenital plate. Subgenital plate rounded with triangular shape, tip rounded. Paraprocts, fusion plate, retractoral plate and cerci as is typical for the genus.
</paragraph>
<paragraph id="38354158FFCEFF87D2FDFA37FDCE39B5" blockId="10.[151,1437,535,2037]" pageId="10" pageNumber="11">
<emphasis id="0AFE9D4AFFCEFF87D2FDFA37FE5D3AE0" bold="true" box="[199,486,1471,1496]" pageId="10" pageNumber="11">Morphological affinities</emphasis>
: Males of the species can be easily distinguished from congeners on the basis of the very large, rectangular process of tergite 9. The epiproct resembles that of
<taxonomicName id="FF8A3ADBFFCEFF87D62BFA6DFBCA3AC4" box="[1041,1137,1509,1532]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="acuta">
<emphasis id="0AFE9D4AFFCEFF87D62BFA6DFBCA3AC4" box="[1041,1137,1509,1532]" italics="true" pageId="10" pageNumber="11">Z. acuta</emphasis>
</taxonomicName>
, but differs in having an upcurved, moderately acute tip, not straight and acute. The rather large ventral vesicle is probably also distinctive, but given that only one population was examined, the value of this feature, variable in some other species, is questionable. Female specimens were not available for study, but are most probably indistinguishable from congeners, like other females in the genus.
</paragraph>
<paragraph id="38354158FFCEFF87D2FDF91FFE743988" blockId="10.[151,1437,535,2037]" box="[199,463,1687,1712]" pageId="10" pageNumber="11">
<emphasis id="0AFE9D4AFFCEFF87D2FDF91FFEE93988" bold="true" box="[199,338,1687,1712]" pageId="10" pageNumber="11">Drumming.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="709012D3FFCEFF87D2FDF934FDF738B1" pageId="10" pageNumber="11" type="distribution">
<paragraph id="38354158FFCEFF87D2FDF934FDF738B1" blockId="10.[151,1437,535,2037]" pageId="10" pageNumber="11">
<emphasis id="0AFE9D4AFFCEFF87D2FDF934FE4E39ED" bold="true" box="[199,501,1724,1749]" pageId="10" pageNumber="11">Distribution and ecology.</emphasis>
We describe this species on the basis of six males from a single population in the Komi Republic,
<collectingCountry id="409D01C8FFCEFF87D36BF957FE2739C0" box="[337,412,1759,1784]" name="Russia" pageId="10" pageNumber="11">Russia</collectingCountry>
(
<figureCitation id="A0B15DDDFFCEFF87D396F957FDBA39C0" box="[428,513,1759,1784]" captionStart="FIGURE 22" captionStartId="13.[151,250,1126,1148]" captionTargetBox="[312,1274,451,1090]" captionTargetId="figure@13.[312,1275,451,1091]" captionTargetPageId="13" captionText="FIGURE 22. Known distribution of Zwicknia westermanni Boumans &amp; Murányi, sp. n. (full circles, type locality indicated with a central dot) and Z. komica Murányi &amp; Boumans, sp. n. (open circle) — known area of the genus Zwicknia Murányi, 2014 indicated in red, grey areas are above 1000 m sea level." httpUri="https://zenodo.org/record/286177/files/figure.png" pageId="10" pageNumber="11">Fig. 22</figureCitation>
). The
<typeStatus id="E731FFFAFFCEFF87D07DF969FDCC39C1" box="[583,631,1761,1785]" pageId="10" pageNumber="11">type</typeStatus>
locality is a lowland forest stream, tributary of the Sysola River, Dvina drainage basin. Further populations of
<taxonomicName id="FF8A3ADBFFCEFF87D05AF88CFD7D3824" box="[608,710,1796,1820]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="0AFE9D4AFFCEFF87D05AF88CFD7D3824" box="[608,710,1796,1820]" italics="true" pageId="10" pageNumber="11">Zwicknia</emphasis>
</taxonomicName>
from European Northeast
<collectingCountry id="409D01C8FFCEFF87D639F88CFBF53825" box="[1027,1102,1796,1821]" name="Russia" pageId="10" pageNumber="11">Russia</collectingCountry>
(
<bibRefCitation id="5C1B3CA9FFCEFF87D65BF88CFA9D3824" author="Loskutova" box="[1121,1318,1796,1821]" pageId="10" pageNumber="11" refString="Loskutova, O. (2003) Fauna of stoneflies (Plecoptera) of the European North-East Russia and its change under the influence of anthropogenic factors. In: Gaino, E. (Ed.), Research Update on Ephemeroptera &amp; Plecoptera. Universita di Perugia, Perugia, pp. 357 - 362." type="book chapter" year="2003">Loskutova, 2003</bibRefCitation>
) probably belong to this species, but the actual distribution of this species in the East European Plain or Russian Plain is unknown.
<taxonomicName id="FF8A3ADBFFCEFF87D32DF8C4FE64385C" box="[279,479,1868,1892]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="komica">
<emphasis id="0AFE9D4AFFCEFF87D32DF8C4FE64385C" box="[279,479,1868,1892]" italics="true" pageId="10" pageNumber="11">Zwicknia komica</emphasis>
</taxonomicName>
and
<taxonomicName id="FF8A3ADBFFCEFF87D013F8C5FD25385C" box="[553,670,1868,1892]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="0AFE9D4AFFCEFF87D013F8C5FD25385C" box="[553,670,1868,1892]" italics="true" pageId="10" pageNumber="11">Z. bifrons</emphasis>
</taxonomicName>
from
<collectingCountry id="409D01C8FFCEFF87D0CCF8C5FCEB385D" box="[758,848,1869,1893]" name="Norway" pageId="10" pageNumber="11">Norway</collectingCountry>
(
<bibRefCitation id="5C1B3CA9FFCEFF87D152F8C4FBF7385C" author="Lillehammer" box="[872,1100,1868,1893]" pageId="10" pageNumber="11" refString="Lillehammer, A. (1988) Stoneflies (Plecoptera) of Fennoscandia and Denmark. Brill, Leiden, 165 pp." type="book" year="1988">Lillehammer, 1988</bibRefCitation>
;
<bibRefCitation id="5C1B3CA9FFCEFF87D658F8C5FAA7385C" author="Boumans" box="[1122,1308,1868,1892]" pageId="10" pageNumber="11" refString="Boumans, L. (2011) Databasing the stoneflies (Plecoptera) at the Natural History Museum in Oslo reveals new Norwegian province records. Norwegian Journal of Entomology, 58, 170 - 179." type="journal article" year="2011">Boumans, 2011</bibRefCitation>
) have the northernmost distribution of the genus.
</paragraph>
</subSubSection>
<subSubSection id="709012D3FFCEFF80D2FDF81CFA273EBD" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11" type="etymology">
<paragraph id="38354158FFCEFF87D2FDF81CFD3838CD" blockId="10.[151,1437,535,2037]" pageId="10" pageNumber="11">
<emphasis id="0AFE9D4AFFCEFF87D2FDF81CFEF53895" bold="true" box="[199,334,1940,1965]" pageId="10" pageNumber="11">Etymology.</emphasis>
The species is dedicated to the Komi Republic where the
<typeStatus id="E731FFFAFFCEFF87D1E3F81DFBB23895" box="[985,1033,1941,1965]" pageId="10" pageNumber="11">type</typeStatus>
specimens were found. The species epithet ends in the Latin adjectival suffix –
<emphasis id="0AFE9D4AFFCEFF87D0B4F831FD0038E8" box="[654,699,1977,2000]" italics="true" pageId="10" pageNumber="11">icus</emphasis>
“pertaining to”. Agreeing with the genus name in grammatical gender, the adjective has the feminine form.
</paragraph>
<caption id="6CF511D0FFCFFF86D2ADF8ECFAAF38A1" httpUri="https://zenodo.org/record/286175/files/figure.png" pageId="11" pageNumber="12" targetBox="[189,1402,207,1861]" targetPageId="11">
<paragraph id="38354158FFCFFF86D2ADF8ECFAAF38A1" blockId="11.[151,1435,1892,1945]" pageId="11" pageNumber="12">
<emphasis id="0AFE9D4AFFCFFF86D2ADF8ECFEFC3842" bold="true" box="[151,327,1892,1915]" pageId="11" pageNumber="12">FIGURES 9–13.</emphasis>
Male terminalia of
<taxonomicName id="FF8A3ADBFFCFFF86D02BF8ECFC253842" authority="Muranyi &amp; Boumans" authorityName="Muranyi &amp; Boumans" box="[529,926,1892,1914]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="komica" status="sp. nov.">
<emphasis id="0AFE9D4AFFCFFF86D02BF8ECFD053842" box="[529,702,1892,1914]" italics="true" pageId="11" pageNumber="12">Zwicknia komica</emphasis>
Murányi &amp; Boumans
</taxonomicName>
,
<taxonomicNameLabel id="11CD2031FFCFFF86D193F8EDFC583842" box="[937,995,1893,1914]" pageId="11" pageNumber="12" rank="species">
<emphasis id="0AFE9D4AFFCFFF86D193F8EDFC583842" bold="true" box="[937,995,1893,1914]" pageId="11" pageNumber="12">sp. n.</emphasis>
</taxonomicNameLabel>
; paratype, Russia, Komi Republic, Tyla-yu stream. 9: dorso-caudal view; 10: dorsal view; 11: lateral view; 12: ventral view; 13: caudal view—scale 0.5 mm.
</paragraph>
</caption>
<caption id="6CF511D0FFC8FF81D2ADF9EEFC5339E1" httpUri="https://zenodo.org/record/286176/files/figure.png" pageId="12" pageNumber="13" targetBox="[312,1266,182,1638]" targetPageId="12">
<paragraph id="38354158FFC8FF81D2ADF9EEFC5339E1" blockId="12.[151,1437,1638,1753]" pageId="12" pageNumber="13">
<emphasis id="0AFE9D4AFFC8FF81D2ADF9EEFEEF3944" bold="true" box="[151,340,1638,1661]" pageId="12" pageNumber="13">FIGURES 14–21.</emphasis>
Male terminalia of
<taxonomicName id="FF8A3ADBFFC8FF81D026F9EFFC5A3945" authority="Boumans &amp; Muranyi" authorityName="Boumans &amp; Muranyi" box="[540,993,1639,1661]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="westermanni" status="sp. nov.">
<emphasis id="0AFE9D4AFFC8FF81D026F9EFFCBA3945" box="[540,769,1639,1661]" italics="true" pageId="12" pageNumber="13">Zwicknia westermanni</emphasis>
Boumans &amp; Murányi
</taxonomicName>
,
<taxonomicNameLabel id="11CD2031FFC8FF81D1D0F9EFFB983944" box="[1002,1059,1639,1660]" pageId="12" pageNumber="13" rank="species">
<emphasis id="0AFE9D4AFFC8FF81D1D0F9EFFB983944" bold="true" box="[1002,1059,1639,1660]" pageId="12" pageNumber="13">sp. n.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="FF8A3ADBFFC8FF81D66EF9E0FA233945" authority="Muranyi &amp; Boumans" authorityName="Muranyi &amp; Boumans" box="[1108,1432,1639,1661]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="komica" status="sp. nov.">
<emphasis id="0AFE9D4AFFC8FF81D66EF9E0FB033945" box="[1108,1208,1639,1661]" italics="true" pageId="12" pageNumber="13">Z. komica</emphasis>
Murányi &amp; Boumans
</taxonomicName>
,
<emphasis id="0AFE9D4AFFC8FF81D2ADF90EFF6B39A3" bold="true" box="[151,208,1670,1691]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="11CD2031FFC8FF81D2ADF90EFF6B39A3" box="[151,208,1670,1691]" pageId="12" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
—14, 16, 18, 20: main epiproct sclerite (Ep-scl), lateral view; 15, 17, 19, 21: hump of tergite 9, caudal view—14–17:
<taxonomicName id="FF8A3ADBFFC8FF81D7BDF90FFEA33983" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="0AFE9D4AFFC8FF81D7BDF90FFEA33983" italics="true" pageId="12" pageNumber="13">Z. westermanni</emphasis>
</taxonomicName>
paratypes, Germany, Lower Saxony, Hohenförsiek; 18–19:
<taxonomicName id="FF8A3ADBFFC8FF81D1B9F92EFB9A3983" box="[899,1057,1702,1723]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="0AFE9D4AFFC8FF81D1B9F92EFB9A3983" box="[899,1057,1702,1723]" italics="true" pageId="12" pageNumber="13">Z. westermanni</emphasis>
</taxonomicName>
paratypes France, Burgundy, Curgy; 20–21:
<taxonomicName id="FF8A3ADBFFC8FF81D2DEF94CFEFC39E1" box="[228,327,1731,1753]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="komica">
<emphasis id="0AFE9D4AFFC8FF81D2DEF94CFEFC39E1" box="[228,327,1731,1753]" italics="true" pageId="12" pageNumber="13">Z. komica</emphasis>
</taxonomicName>
paratype, Russia, Komi Republic, Tyla-yu stream—scale 0.5 mm.
</paragraph>
</caption>
<paragraph id="38354158FFC8FF81D2ADF897FB4A3800" blockId="12.[151,1265,1823,1848]" box="[151,1265,1823,1848]" pageId="12" pageNumber="13">
Contribution to the morphological key of adult
<taxonomicName id="FF8A3ADBFFC8FF81D090F897FCAA380F" box="[682,785,1823,1847]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="0AFE9D4AFFC8FF81D090F897FCAA380F" box="[682,785,1823,1847]" italics="true" pageId="12" pageNumber="13">Zwicknia</emphasis>
</taxonomicName>
males, published in
<bibRefCitation id="5C1B3CA9FFC8FF81D1C2F897FB563800" box="[1016,1261,1823,1848]" pageId="12" pageNumber="13" refString="Muranyi, D., Gamboa, M. &amp; Orci, K. M. (2014) Zwicknia gen. n. for the Capnia bifrons species group, with descriptions of three new species based on morphology, drumming signals and molecular genetics, and a synopsis of the West Palaearctic and Nearctic genera of Capniidae (Plecoptera). Zootaxa, in press." type="book">
Murányi
<emphasis id="0AFE9D4AFFC8FF81D665F8A8FB2C380F" box="[1119,1175,1823,1847]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
(2014)
</bibRefCitation>
:
</paragraph>
<paragraph id="38354158FFC8FF81D2ADF8E0FCF63847" blockId="12.[151,845,1895,1920]" box="[151,845,1895,1920]" pageId="12" pageNumber="13">
<taxonomicName id="FF8A3ADBFFC8FF81D2ADF8E0FEBE3847" box="[151,261,1895,1919]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="komica">
<emphasis id="0AFE9D4AFFC8FF81D2ADF8E0FEBE3847" box="[151,261,1895,1919]" italics="true" pageId="12" pageNumber="13">Z. komica</emphasis>
</taxonomicName>
keys out to couplet 3, which should be modified as:
</paragraph>
<paragraph id="38354158FFC8FF80D2ADF820FA2C3FF4" blockId="12.[151,1435,1960,2011]" lastBlockId="13.[151,1436,151,269]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
3a Process of tergite 9 low and caudally projecting (
<bibRefCitation id="5C1B3CA9FFC8FF81D08CF820FCC13886" author="Muranyi" box="[694,890,1960,1982]" pageId="12" pageNumber="13" refString="Muranyi, D., Gamboa, M. &amp; Orci, K. M. (2014) Zwicknia gen. n. for the Capnia bifrons species group, with descriptions of three new species based on morphology, drumming signals and molecular genetics, and a synopsis of the West Palaearctic and Nearctic genera of Capniidae (Plecoptera). Zootaxa, in press." type="book" year="2014">
Murányi
<emphasis id="0AFE9D4AFFC8FF81D12AF821FCFA3886" box="[784,833,1960,1982]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2014
</bibRefCitation>
: Figs. 92, 157–159)..................
<taxonomicName id="FF8A3ADBFFC8FF81D724F821FA203886" box="[1310,1435,1960,1982]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="rupprechti">
<emphasis id="0AFE9D4AFFC8FF81D724F821FA203886" box="[1310,1435,1960,1982]" italics="true" pageId="12" pageNumber="13">Z. rupprechti</emphasis>
</taxonomicName>
- Process of tergite 9 high and perpendicularly raised........................................................... 3 3 Process of tergite 9 three times as wide as main epiproct sclerite (
<figureCitation id="A0B15DDDFFC9FF80D16EFF1FFC643F95" box="[852,991,151,173]" captionStart="FIGURES 9 – 13" captionStartId="11.[151,264,1892,1915]" captionTargetBox="[189,1402,207,1861]" captionTargetId="figure@11.[151,1436,193,1871]" captionTargetPageId="11" captionText="FIGURES 9 – 13. Male terminalia of Zwicknia komica Murányi &amp; Boumans, sp. n.; paratype, Russia, Komi Republic, Tyla-yu stream. 9: dorso-caudal view; 10: dorsal view; 11: lateral view; 12: ventral view; 13: caudal view—scale 0.5 mm." httpUri="https://zenodo.org/record/286175/files/figure.png" pageId="13" pageNumber="14">Figs. 9–10, 13</figureCitation>
,
<figureCitation id="A0B15DDDFFC9FF80D1D0FF1FFBB83F95" box="[1002,1027,151,173]" captionStart="FIGURES 14 – 21" captionStartId="12.[151,264,1638,1661]" captionTargetBox="[312,1266,182,1638]" captionTargetId="figure@12.[293,1293,147,1648]" captionTargetPageId="12" captionText="FIGURES 14 – 21. Male terminalia of Zwicknia westermanni Boumans &amp; Murányi, sp. n. and Z. komica Murányi &amp; Boumans, sp. n. — 14, 16, 18, 20: main epiproct sclerite (Ep-scl), lateral view; 15, 17, 19, 21: hump of tergite 9, caudal view— 14 – 17: Z. westermanni paratypes, Germany, Lower Saxony, Hohenförsiek; 18 – 19: Z. westermanni paratypes France, Burgundy, Curgy; 20 – 21: Z. komica paratype, Russia, Komi Republic, Tyla-yu stream—scale 0.5 mm." httpUri="https://zenodo.org/record/286176/files/figure.png" pageId="13" pageNumber="14">21</figureCitation>
).........................
<taxonomicName id="FF8A3ADBFFC9FF80D707FF10FA203F95" box="[1341,1435,151,173]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="komica">
<emphasis id="0AFE9D4AFFC9FF80D707FF10FA203F95" box="[1341,1435,151,173]" italics="true" pageId="13" pageNumber="14">Z. komica</emphasis>
</taxonomicName>
- Process of tergite 9 less than 1.5× as wide as main epiproct sclerite (
<bibRefCitation id="5C1B3CA9FFC9FF80D14AFF3FFB8F3FF5" author="Muranyi" box="[880,1076,183,205]" pageId="13" pageNumber="14" refString="Muranyi, D., Gamboa, M. &amp; Orci, K. M. (2014) Zwicknia gen. n. for the Capnia bifrons species group, with descriptions of three new species based on morphology, drumming signals and molecular genetics, and a synopsis of the West Palaearctic and Nearctic genera of Capniidae (Plecoptera). Zootaxa, in press." type="book" year="2014">
Murányi
<emphasis id="0AFE9D4AFFC9FF80D1F0FF30FC403FF5" box="[970,1019,183,205]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2014
</bibRefCitation>
: Figs. 82, 85, 120, 152–156).......
</paragraph>
<paragraph id="38354158FFC9FF80D2D9FF5EFC303E34" blockId="13.[151,1436,151,269]" pageId="13" pageNumber="14">
.............................................................................................
<taxonomicName id="FF8A3ADBFFC9FF80D705FF5EFA273FD3" box="[1343,1436,213,235]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="0AFE9D4AFFC9FF80D705FF5EFA273FD3" box="[1343,1436,213,235]" italics="true" pageId="13" pageNumber="14">Z. bifrons</emphasis>
</taxonomicName>
<taxonomicName id="FF8A3ADBFFC9FF80D2ADFF7DFEF83E34" box="[151,323,245,268]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="0AFE9D4AFFC9FF80D2ADFF7DFEF83E34" box="[151,323,245,268]" italics="true" pageId="13" pageNumber="14">Z. westermanni</emphasis>
</taxonomicName>
keys out to couplet 4, which should be modified as:
</paragraph>
<paragraph id="38354158FFC9FF80D2ADFEBCFA273EBD" blockId="13.[151,1436,308,389]" pageId="13" pageNumber="14">
4a Brachypterous (
<figureCitation id="A0B15DDDFFC9FF80D340FEBCFE0F3E72" box="[378,436,308,330]" captionStart="FIGURE 6" captionStartId="8.[151,250,1071,1093]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@8.[151,1436,193,1049]" captionTargetPageId="8" captionText="FIGURE 6. Habitus of Zwicknia westermanni Boumans &amp; Murányi, sp. n., mating pair, France, Burgundy, Saône-et-Loire department, Curgy. Photo A. Ruffoni." httpUri="https://zenodo.org/record/286172/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
); ventral vesicle half as wide as the subgenital plate (Fig. 4).......................
<taxonomicName id="FF8A3ADBFFC9FF80D73FFEBDFA273E72" box="[1285,1436,309,330]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="0AFE9D4AFFC9FF80D73FFEBDFA273E72" box="[1285,1436,309,330]" italics="true" pageId="13" pageNumber="14">Z. westermanni</emphasis>
</taxonomicName>
- Micropterous or macropterous (
<bibRefCitation id="5C1B3CA9FFC9FF80D02BFED9FD6F3E5F" author="Muranyi" box="[529,724,337,359]" pageId="13" pageNumber="14" refString="Muranyi, D., Gamboa, M. &amp; Orci, K. M. (2014) Zwicknia gen. n. for the Capnia bifrons species group, with descriptions of three new species based on morphology, drumming signals and molecular genetics, and a synopsis of the West Palaearctic and Nearctic genera of Capniidae (Plecoptera). Zootaxa, in press." type="book" year="2014">
Murányi
<emphasis id="0AFE9D4AFFC9FF80D051FEDAFD273E5F" box="[619,668,337,359]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2014
</bibRefCitation>
: Figs. 57, 59, 61); ventral vesicle narrower than half the width of subgenital plate (
<bibRefCitation id="5C1B3CA9FFC9FF80D300FEE7FE463EBD" author="Muranyi" box="[314,509,367,389]" pageId="13" pageNumber="14" refString="Muranyi, D., Gamboa, M. &amp; Orci, K. M. (2014) Zwicknia gen. n. for the Capnia bifrons species group, with descriptions of three new species based on morphology, drumming signals and molecular genetics, and a synopsis of the West Palaearctic and Nearctic genera of Capniidae (Plecoptera). Zootaxa, in press." type="book" year="2014">
Murányi
<emphasis id="0AFE9D4AFFC9FF80D3AEFEF8FE7E3EBD" box="[404,453,367,389]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2014
</bibRefCitation>
: Figs. 79, 95, 99, 103).......................................................... 4
</paragraph>
</subSubSection>
</treatment>
</document>
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