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<document id="EF2482FF09C392AC66F0C779101F9832" ID-CLB-Dataset="39303" ID-DOI="10.11646/zootaxa.4057.1.9" ID-GBIF-Dataset="a3d9451b-2e0c-40e6-bafb-2d8a958fd8de" ID-ISSN="1175-5326" ID-Zenodo-Dep="233231" ID-ZooBank="7AC91C1D-5EAB-40A6-B177-3A154AF2F9EA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461354776227" checkinUser="plazi" docAuthor="Jacobs, Carmen T., Scholtz, Clarke H. &amp; Strümpher, Werner P." docDate="2015" docId="B01287C7FFB28811FF283C2DC93AE0EE" docLanguage="en" docName="zt04057p142.pdf" docOrigin="Zootaxa 4057 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Colophon stokoei Barnard 1929" docType="treatment" docVersion="9" lastPageNumber="137" masterDocId="4C2BFFBFFFB38813FFBF397ACD4FE223" masterDocTitle="Taxonomy of Colophon Gray (Coleoptera: Lucanidae): new species and a status change" masterLastPageNumber="142" masterPageNumber="135" pageNumber="136" updateTime="1698661467203" updateUser="ExternalLinkService">
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<mods:title id="E0773F8EDA7357B0876CDEE75EA83035">Taxonomy of Colophon Gray (Coleoptera: Lucanidae): new species and a status change</mods:title>
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<mods:namePart id="767944C77DCDB8B9C175508D4EBCD151">Jacobs, Carmen T.</mods:namePart>
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<treatment id="B01287C7FFB28811FF283C2DC93AE0EE" ID-DOI="http://doi.org/10.5281/zenodo.6093229" ID-GBIF-Taxon="119647053" ID-Zenodo-Dep="6093229" LSID="urn:lsid:plazi:treatment:B01287C7FFB28811FF283C2DC93AE0EE" httpUri="http://treatment.plazi.org/id/B01287C7FFB28811FF283C2DC93AE0EE" lastPageId="2" lastPageNumber="137" pageId="1" pageNumber="136">
<subSubSection id="70A1655AFFB28812FF283C2DCCC8E7B0" pageId="1" pageNumber="136" type="nomenclature">
<paragraph id="380436D1FFB28812FF283C2DCF69E751" blockId="1.[151,550,1367,1427]" box="[151,550,1367,1394]" pageId="1" pageNumber="136">
<heading id="634C81BDFFB28812FF283C2DCF69E751" bold="true" box="[151,550,1367,1394]" fontSize="11" level="1" pageId="1" pageNumber="136" reason="1">
<taxonomicName id="FFBB4D52FFB28812FF283C2DCF69E751" ID-CoL="X846" authority="Barnard, 1929" authorityName="Barnard" authorityYear="1929" box="[151,550,1367,1394]" class="Insecta" family="Lucanidae" genus="Colophon" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="stokoei">
<emphasis id="0ACFEAC3FFB28812FF283C2DCF69E751" bold="true" box="[151,550,1367,1394]" pageId="1" pageNumber="136">
<emphasis id="0ACFEAC3FFB28812FF283C2DCC25E752" bold="true" box="[151,362,1367,1393]" italics="true" pageId="1" pageNumber="136">Colophon stokoei</emphasis>
<bibRefCitation id="5C2A4B20FFB28812FECE3C22CF69E751" author="Barnard" box="[369,550,1368,1394]" pageId="1" pageNumber="136" refString="Barnard, K. H. (1929) A study of the genus Colophon Gray (Coleoptera: Lucanidae). Transactions of the Royal Society of South Africa, 18, 163 - 182. http: // dx. doi. org / 10.1080 / 00359192909518797" type="journal article" year="1929">Barnard, 1929</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="380436D1FFB28812FF283C00CCC8E7B0" blockId="1.[151,550,1367,1427]" box="[151,391,1402,1427]" pageId="1" pageNumber="136">
(
<figureCitation id="A0802A54FFB28812FF203C00CDA7E7B0" box="[159,232,1402,1427]" captionStart="FIGURE 1" captionStartId="2.[151,250,1641,1663]" captionTargetBox="[244,1340,779,1610]" captionTargetId="figure@2.[240,1347,772,1617]" captionTargetPageId="2" captionText="FIGURE 1. Adult habitus of Colophon stokoei Barnard: (a) male dorsal view; (b) male ventral view; (c) female dorsal view; (d) male protibia; (e) aedeagus dorsal view; (f) aedeagus lateral view; (g) ventral plate of genital mass; (h) dorsal plate of genital mass." httpUri="https://zenodo.org/record/233232/files/figure.png" pageId="1" pageNumber="136">Figs. 1</figureCitation>
ah;
<tableCitation id="7539036AFFB28812FE9B3C00CC3AE7B0" box="[292,373,1402,1427]" captionStart="TABLE 1" captionStartId="5.[151,239,152,175]" captionText="TABLE 1. Diagnostic features of the head, mentum, and mandibles to aid in the identification of Colophon deschodti, C. stokoei, C. struempheri, and C. switalae." pageId="1" pageNumber="136">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="70A1655AFFB28812FF283CB8CE0EE7FB" box="[151,833,1474,1496]" pageId="1" pageNumber="136" type="reference_group">
<paragraph id="380436D1FFB28812FF283CB8CE0EE7FB" blockId="1.[151,833,1474,1496]" box="[151,833,1474,1496]" pageId="1" pageNumber="136">
<treatmentCitationGroup id="18AB11FFFFB28812FF283CB8CE0EE7FB" box="[151,833,1474,1496]" pageId="1" pageNumber="136">
<taxonomicName id="FFBB4D52FFB28812FF283CB8CF59E7FB" ID-CoL="X846" authority="Barnard, 1929: 168" authorityName="Barnard" authorityPageNumber="168" authorityYear="1929" box="[151,534,1474,1496]" class="Insecta" family="Lucanidae" genus="Colophon" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="stokoei">
<emphasis id="0ACFEAC3FFB28812FF283CB8CC06E7FB" box="[151,329,1474,1496]" italics="true" pageId="1" pageNumber="136">Colophon stokoei</emphasis>
<treatmentCitation id="B91A10C0FFB28812FEF03CB8CF59E7FB" author="Barnard" box="[335,534,1474,1496]" page="168" pageId="1" pageNumber="136" year="1929">
<bibRefCitation id="5C2A4B20FFB28812FEF03CB8CCACE7FB" author="Barnard" box="[335,483,1474,1496]" pageId="1" pageNumber="136" refString="Barnard, K. H. (1929) A study of the genus Colophon Gray (Coleoptera: Lucanidae). Transactions of the Royal Society of South Africa, 18, 163 - 182. http: // dx. doi. org / 10.1080 / 00359192909518797" type="journal article" year="1929">Barnard, 1929</bibRefCitation>
: 168
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="B91A10C0FFB28812FD9E3CB8CE72E7FB" author="Endrody-Younga" box="[545,829,1474,1496]" page="393" pageId="1" pageNumber="136" year="1988">
<bibRefCitation id="5C2A4B20FFB28812FD9E3CB8CE45E7FB" author="Endrody-Younga" box="[545,778,1474,1496]" pageId="1" pageNumber="136" refString="Endrody-Younga, S. (1988) Evidence for the low-altitude origin of the Cape Mountain biome derived from the systematic revision of the genus Colophon Gray (Coleoptera, Lucanidae). Annals of the South African Museum, 96, 359 - 424." type="journal article" year="1988">Endrődy-Younga 1988</bibRefCitation>
: 393
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="70A1655AFFB28812FF283F7FCF4EE445" pageId="1" pageNumber="136" type="materials_examined">
<paragraph id="380436D1FFB28812FF283F7FCF4EE445" blockId="1.[151,1437,1541,2034]" pageId="1" pageNumber="136">
<emphasis id="0ACFEAC3FFB28812FF283F7FCCF7E43D" bold="true" box="[151,440,1541,1566]" pageId="1" pageNumber="136">
<typeStatus id="E7008873FFB28812FF283F7FCD9DE43D" box="[151,210,1541,1566]" pageId="1" pageNumber="136">Type</typeStatus>
material examined
</emphasis>
.
<typeStatus id="E7008873FFB28812FE783F7FCF10E43D" box="[455,607,1541,1566]" pageId="1" pageNumber="136" type="paralectotype">Paralectotype</typeStatus>
(
<specimenCount id="2EBDFD58FFB28812FDCE3F7CCFDAE43D" box="[625,661,1542,1566]" pageId="1" pageNumber="136" type="generic">1♂</specimenCount>
): Hott.-Holl. [Hottentots
<collectingCountry id="40AC7641FFB28812FC0C3F7FC95FE43D" box="[947,1040,1541,1566]" name="Netherlands" pageId="1" pageNumber="136">Holland</collectingCountry>
] Mts., 4 000ft. [4 0005 000] ft. [= 1 2201
<quantity id="FF439B34FFB28812FF463F53CC09E462" box="[249,326,1577,1602]" metricMagnitude="2" metricUnit="m" metricValue="5.24" pageId="1" pageNumber="136" unit="m" value="524.0">524 m</quantity>
], Caledon C.C. [Cape Colony], Barnard, [January] 1916 (
<collectionCode id="5EAAAE14FFB28812FC5F3F50C963E461" box="[992,1068,1578,1602]" pageId="1" pageNumber="136">ISAM</collectionCode>
). Additional material examined:
<specimenCount id="2EBDFD58FFB28812FF283F37CD84E445" box="[151,203,1613,1638]" pageId="1" pageNumber="136" type="generic">62♂</specimenCount>
, 3♀ (
<collectionCode id="5EAAAE14FFB28812FEB73F37CC1BE445" box="[264,340,1613,1638]" pageId="1" pageNumber="136">ISAM</collectionCode>
);
<specimenCount id="2EBDFD58FFB28812FED83F37CCD6E445" box="[359,409,1613,1638]" pageId="1" pageNumber="136" type="generic">10♂</specimenCount>
(
<collectionCode id="5EAAAE14FFB28812FE183F37CCBAE445" box="[423,501,1613,1638]" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="1" pageNumber="136">UPSA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="70A1655AFFB28812FF783F0BCC4CE539" pageId="1" pageNumber="136" type="diagnosis">
<paragraph id="380436D1FFB28812FF783F0BCC4CE539" blockId="1.[151,1437,1541,2034]" pageId="1" pageNumber="136">
<emphasis id="0ACFEAC3FFB28812FF783F0BCC0EE4A9" bold="true" box="[199,321,1649,1674]" pageId="1" pageNumber="136">Diagnosis.</emphasis>
<taxonomicName id="FFBB4D52FFB28812FEF63F0BCF41E4AA" box="[329,526,1649,1673]" class="Insecta" family="Lucanidae" genus="Colophon" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="stokoei">
<emphasis id="0ACFEAC3FFB28812FEF63F0BCF41E4AA" box="[329,526,1649,1673]" italics="true" pageId="1" pageNumber="136">Colophon stokoei</emphasis>
</taxonomicName>
is well represented in collections and is relatively easy to identify despite being superficially similar to
<taxonomicName id="FFBB4D52FFB28812FE1F3FECCF62E48E" box="[416,557,1685,1709]" class="Insecta" family="Lucanidae" genus="Colophon" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="deschodti" status="sp. nov.">
<emphasis id="0ACFEAC3FFB28812FE1F3FECCCFCE48E" box="[416,435,1686,1709]" italics="true" pageId="1" pageNumber="136">C</emphasis>
.
<emphasis id="0ACFEAC3FFB28812FE7D3FEFCF62E48E" box="[450,557,1685,1709]" italics="true" pageId="1" pageNumber="136">deschodti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="11FC57B8FFB28812FD8A3FEFCF8EE48D" box="[565,705,1685,1710]" pageId="1" pageNumber="136" rank="species">
<emphasis id="0ACFEAC3FFB28812FD8A3FEFCF8EE48D" bold="true" box="[565,705,1685,1710]" pageId="1" pageNumber="136">new species</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="FFBB4D52FFB28812FD6F3FECCE00E48E" box="[720,847,1685,1709]" class="Insecta" family="Lucanidae" genus="Colophon" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="switalae" status="sp. nov.">
<emphasis id="0ACFEAC3FFB28812FD6F3FECCE00E48E" box="[720,847,1685,1709]" italics="true" pageId="1" pageNumber="136">C. switalae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="11FC57B8FFB28812FCE73FEFCEACE48D" box="[856,995,1685,1710]" pageId="1" pageNumber="136" rank="species">
<emphasis id="0ACFEAC3FFB28812FCE73FEFCEACE48D" bold="true" box="[856,995,1685,1710]" pageId="1" pageNumber="136">new species</emphasis>
</taxonomicNameLabel>
, and
<taxonomicName id="FFBB4D52FFB28812FB9C3FECC983E48E" box="[1059,1228,1685,1709]" class="Insecta" family="Lucanidae" genus="Colophon" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="struempheri" status="sp. nov.">
<emphasis id="0ACFEAC3FFB28812FB9C3FECC983E48E" box="[1059,1228,1685,1709]" italics="true" pageId="1" pageNumber="136">C. struempheri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="11FC57B8FFB28812FB6A3FEFC82FE48D" box="[1237,1376,1685,1710]" pageId="1" pageNumber="136" rank="species">
<emphasis id="0ACFEAC3FFB28812FB6A3FEFC82FE48D" bold="true" box="[1237,1376,1685,1710]" pageId="1" pageNumber="136">new species</emphasis>
</taxonomicNameLabel>
. It is best characterised by a distinct tooth situated on the inner basal margin of the sickle-shaped mandible, which is small or obsolete in the male specimens of the other three species. The shape of the mentum is also diagnostic (
<tableCitation id="7539036AFFB28812FF213E7BCDBDE539" box="[158,242,1793,1818]" captionStart="TABLE 1" captionStartId="5.[151,239,152,175]" captionText="TABLE 1. Diagnostic features of the head, mentum, and mandibles to aid in the identification of Colophon deschodti, C. stokoei, C. struempheri, and C. switalae." pageId="1" pageNumber="136">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="70A1655AFFB28811FF783E5FCE73E08B" lastPageId="2" lastPageNumber="137" pageId="1" pageNumber="136" type="description">
<paragraph id="380436D1FFB28811FF783E5FCE73E08B" blockId="1.[151,1437,1541,2034]" lastBlockId="2.[151,1436,151,717]" lastPageId="2" lastPageNumber="137" pageId="1" pageNumber="136">
<emphasis id="0ACFEAC3FFB28812FF783E5FCC3BE51D" bold="true" box="[199,372,1829,1854]" pageId="1" pageNumber="136">Redescription.</emphasis>
<emphasis id="0ACFEAC3FFB28812FEC23E5CCCE5E51E" box="[381,426,1830,1853]" italics="true" pageId="1" pageNumber="136">Size</emphasis>
: Male:
<quantity id="FF439B34FFB28812FDBA3E5CCFCEE51E" box="[517,641,1830,1854]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.7" metricValueMin="2.1" pageId="1" pageNumber="136" unit="mm" value="24.0" valueMax="27.0" valueMin="21.0">2127 mm</quantity>
, pronotal width:
<quantity id="FF439B34FFB28812FCFA3E5CCEFDE51E" box="[837,946,1830,1854]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="1" pageNumber="136" unit="mm" value="10.5" valueMax="12.0" valueMin="9.0">912 mm</quantity>
(
<emphasis id="0ACFEAC3FFB28812FC7C3E5CCE9EE51E" box="[963,977,1830,1853]" italics="true" pageId="1" pageNumber="136">n</emphasis>
= 53). Female:
<quantity id="FF439B34FFB28812FB343E5CC848E51E" box="[1163,1287,1830,1854]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="1" pageNumber="136" unit="mm" value="20.0" valueMax="22.0" valueMin="18.0">1822 mm</quantity>
, width:
<quantity id="FF439B34FFB28812FADA3E5CCD8CE542" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="1" pageNumber="136" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">911 mm</quantity>
(
<emphasis id="0ACFEAC3FFB28812FF6D3E30CDAFE542" box="[210,224,1866,1889]" italics="true" pageId="1" pageNumber="136">n</emphasis>
= 3).
<emphasis id="0ACFEAC3FFB28812FE9D3E33CC14E542" box="[290,347,1865,1889]" italics="true" pageId="1" pageNumber="136">Body</emphasis>
: Convex, dull black (
<figureCitation id="A0802A54FFB28812FDF43E33CFCBE541" box="[587,644,1865,1890]" captionStart="FIGURE 1" captionStartId="2.[151,250,1641,1663]" captionTargetBox="[244,1340,779,1610]" captionTargetId="figure@2.[240,1347,772,1617]" captionTargetPageId="2" captionText="FIGURE 1. Adult habitus of Colophon stokoei Barnard: (a) male dorsal view; (b) male ventral view; (c) female dorsal view; (d) male protibia; (e) aedeagus dorsal view; (f) aedeagus lateral view; (g) ventral plate of genital mass; (h) dorsal plate of genital mass." httpUri="https://zenodo.org/record/233232/files/figure.png" pageId="1" pageNumber="136">Fig 1</figureCitation>
a, b).
<emphasis id="0ACFEAC3FFB28812FD7A3E33CE4EE542" box="[709,769,1865,1889]" italics="true" pageId="1" pageNumber="136">Head</emphasis>
: Transversely oblong, anterior lateral angle almost pointed; lateral margins straight, upper surface with a compressed tubercle on either side at the base of the mandible and an oblique ridge over the eye; internal subocular crests rounded but distinct, with tubercle at the anterior inner end of each crest; frons surface pitted, evenly depressed between tubercles and arcuately slanting to clypeus; clypeus meets triangular labrum on the same level (
<tableCitation id="7539036AFFB28812FD393EA3CF94E5D1" box="[646,731,2009,2034]" captionStart="TABLE 1" captionStartId="5.[151,239,152,175]" captionText="TABLE 1. Diagnostic features of the head, mentum, and mandibles to aid in the identification of Colophon deschodti, C. stokoei, C. struempheri, and C. switalae." pageId="1" pageNumber="136">Table 1</tableCitation>
).
<emphasis id="0ACFEAC3FFB28812FD4B3EA0CE11E5D2" box="[756,862,2010,2033]" italics="true" pageId="1" pageNumber="136">Antennae</emphasis>
: Elbowed, antennomeres of club cannot be folded together.
<emphasis id="0ACFEAC3FFB18811FEBF39E2CC39E293" box="[256,374,152,176]" italics="true" pageId="2" pageNumber="137">Mandibles</emphasis>
: Short and simple, each nearly semicircular; dorsal surface of apical portion strongly furcate; inner basal margin of lower surface with a distinct tooth, which is prominently visible in dorsal as well as ventral view; yellow setae between mandibles (
<tableCitation id="7539036AFFB18811FDEE39A5CFEAE2DB" box="[593,677,223,248]" captionStart="TABLE 1" captionStartId="5.[151,239,152,175]" captionText="TABLE 1. Diagnostic features of the head, mentum, and mandibles to aid in the identification of Colophon deschodti, C. stokoei, C. struempheri, and C. switalae." pageId="2" pageNumber="137">Table 1</tableCitation>
).
<emphasis id="0ACFEAC3FFB18811FD03399BCE57E2DB" box="[700,792,225,248]" italics="true" pageId="2" pageNumber="137">Mentum</emphasis>
: Thick and projecting prominently in lateral view; evenly elevating from base in lateral view, and forming a vertical line or surface anteriorly; transversely quadrangular in ventral view, evenly elevating from base in lateral view, and forming a vertical line or surface anteriorly; anterior surface large and deeply excavated; anterior margin transverse (
<tableCitation id="7539036AFFB18811FC2D3836CEA3E347" box="[914,1004,332,357]" captionStart="TABLE 1" captionStartId="5.[151,239,152,175]" captionText="TABLE 1. Diagnostic features of the head, mentum, and mandibles to aid in the identification of Colophon deschodti, C. stokoei, C. struempheri, and C. switalae." pageId="2" pageNumber="137">Table 1</tableCitation>
).
<emphasis id="0ACFEAC3FFB18811FBB53837C936E347" box="[1034,1145,333,356]" italics="true" pageId="2" pageNumber="137">Pronotum</emphasis>
: Lateral margins evenly arcuate, hardly and only slightly emarginate near posterior angle, in general appearance moderately enlarged.
<emphasis id="0ACFEAC3FFB18811FAE0380BC8DBE3AB" box="[1375,1428,369,392]" italics="true" pageId="2" pageNumber="137">Legs</emphasis>
: Protibia strongly curved and dilated towards the apex, ventral longitudinal crest distinct, but not sharply raised basally; apicoventral process long and narrow; apex externally bidentate, proximal to which is a semicircular excision, dorsal surface convex with a blunt median keel, ventral surface also keeled, distally concave, with a short forwardly directed tooth near insertion of tarsus (
<figureCitation id="A0802A54FFB18811FD753885CE43E03B" box="[714,780,511,536]" captionStart="FIGURE 1" captionStartId="2.[151,250,1641,1663]" captionTargetBox="[244,1340,779,1610]" captionTargetId="figure@2.[240,1347,772,1617]" captionTargetPageId="2" captionText="FIGURE 1. Adult habitus of Colophon stokoei Barnard: (a) male dorsal view; (b) male ventral view; (c) female dorsal view; (d) male protibia; (e) aedeagus dorsal view; (f) aedeagus lateral view; (g) ventral plate of genital mass; (h) dorsal plate of genital mass." httpUri="https://zenodo.org/record/233232/files/figure.png" pageId="2" pageNumber="137">Fig. 1</figureCitation>
d).
<emphasis id="0ACFEAC3FFB18811FC893B7ACE94E03B" box="[822,987,512,536]" italics="true" pageId="2" pageNumber="137">Male genitalia</emphasis>
: Aedeagus symmetrical. Dorsal line of left paramere in lateral view emarginated in apical third, and forms an angle where it turns into the finely convex basal two-thirds (
<figureCitation id="A0802A54FFB18811FEE43B3DCCD5E043" box="[347,410,583,608]" captionStart="FIGURE 1" captionStartId="2.[151,250,1641,1663]" captionTargetBox="[244,1340,779,1610]" captionTargetId="figure@2.[240,1347,772,1617]" captionTargetPageId="2" captionText="FIGURE 1. Adult habitus of Colophon stokoei Barnard: (a) male dorsal view; (b) male ventral view; (c) female dorsal view; (d) male protibia; (e) aedeagus dorsal view; (f) aedeagus lateral view; (g) ventral plate of genital mass; (h) dorsal plate of genital mass." httpUri="https://zenodo.org/record/233232/files/figure.png" pageId="2" pageNumber="137">Fig. 1</figureCitation>
eh).
<emphasis id="0ACFEAC3FFB18811FE613B32CF7DE043" box="[478,562,584,608]" italics="true" pageId="2" pageNumber="137">Female</emphasis>
: Body convex, dull black; pronotum slightly narrower but similar in shape to that of the males; elytra similar to that of males; punctation on head, pronotum, and elytra much more distinct than in males; head and mandibles conspicuously small (
<figureCitation id="A0802A54FFB18811FD633BF5CE52E08B" box="[732,797,655,680]" captionStart="FIGURE 1" captionStartId="2.[151,250,1641,1663]" captionTargetBox="[244,1340,779,1610]" captionTargetId="figure@2.[240,1347,772,1617]" captionTargetPageId="2" captionText="FIGURE 1. Adult habitus of Colophon stokoei Barnard: (a) male dorsal view; (b) male ventral view; (c) female dorsal view; (d) male protibia; (e) aedeagus dorsal view; (f) aedeagus lateral view; (g) ventral plate of genital mass; (h) dorsal plate of genital mass." httpUri="https://zenodo.org/record/233232/files/figure.png" pageId="2" pageNumber="137">Fig. 1</figureCitation>
c).
</paragraph>
</subSubSection>
<subSubSection id="70A1655AFFB18811FF783BCEC93AE0EE" box="[199,1141,692,717]" pageId="2" pageNumber="137" type="distribution">
<paragraph id="380436D1FFB18811FF783BCEC93AE0EE" blockId="2.[151,1436,151,717]" box="[199,1141,692,717]" pageId="2" pageNumber="137">
<emphasis id="0ACFEAC3FFB18811FF783BCECC16E0EE" bold="true" box="[199,345,692,717]" pageId="2" pageNumber="137">Distribution</emphasis>
. Hottentots
<collectingCountry id="40AC7641FFB18811FE5C3BCECF72E0EE" box="[483,573,692,717]" name="Netherlands" pageId="2" pageNumber="137">Holland</collectingCountry>
Mountains, Western Cape Province,
<collectingCountry id="40AC7641FFB18811FC613BCEC93EE0EE" box="[990,1137,692,717]" name="South Africa" pageId="2" pageNumber="137">South Africa</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>