425 lines
63 KiB
XML
425 lines
63 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3792148" ID-GBIF-Dataset="a7e349dc-b498-4233-ade4-a73f3271e501" ID-GBIF-Taxon="163747532" ID-ISSN="1313–2970" ID-Zenodo-Dep="3792148" ID-ZooBank="702BCAE3-5B0A-49EA-8603-D60826241F2C" approvalRequired="3" approvalRequired_for_textStreams="3" checkinTime="1587954994003" checkinUser="plazi" docAuthor="Mesibov, Robert" docDate="2009" docId="9B0D9E62FF9CCC31FF5E5E9D854FB194" docLanguage="en" docName="ZK_article_1996.pdf" docOrigin="ZooKeys 12 (12)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@52c7884c" docStyleName="zookeys.2008.journal_article" docTitle="Agathodesmus steeli Silvestri 1910" docType="treatment" docVersion="6" lastPageId="11" lastPageNumber="98" masterDocId="6734E61AFF99CC3AFFD25A1D811BB31E" masterDocTitle="Revision of Agathodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae)" masterLastPageNumber="110" masterPageNumber="87" pageId="5" pageNumber="92" updateTime="1643568681921" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of Agathodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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||
<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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||
</mods:role>
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<mods:namePart>Mesibov, Robert</mods:namePart>
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<mods:affiliation>Queen Victoria Museum and Art Gallery, Launceston, Australia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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||
<mods:titleInfo>
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||
<mods:title>ZooKeys</mods:title>
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||
</mods:titleInfo>
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||
<mods:part>
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||
<mods:date>2009</mods:date>
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||
<mods:detail type="pubDate">
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||
<mods:number>2009-06-18</mods:number>
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||
</mods:detail>
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||
<mods:detail type="volume">
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||
<mods:number>12</mods:number>
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||
</mods:detail>
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||
<mods:detail type="issue">
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||
<mods:number>12</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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||
<mods:start>87</mods:start>
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||
<mods:end>110</mods:end>
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||
</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.12.206</mods:identifier>
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<mods:identifier type="GBIF-Dataset">a7e349dc-b498-4233-ade4-a73f3271e501</mods:identifier>
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||
<mods:identifier type="ISSN">1313–2970</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">576462</mods:identifier>
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||
<mods:identifier type="ZooBank">702BCAE3-5B0A-49EA-8603-D60826241F2C</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792148" ID-GBIF-Taxon="163747532" ID-Zenodo-Dep="3792148" LSID="urn:lsid:plazi:treatment:9B0D9E62FF9CCC31FF5E5E9D854FB194" httpUri="http://treatment.plazi.org/id/9B0D9E62FF9CCC31FF5E5E9D854FB194" lastPageId="11" lastPageNumber="98" pageId="5" pageNumber="92">
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<subSubSection box="[140,552,1152,1179]" pageId="5" pageNumber="92" type="nomenclature">
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<paragraph blockId="5.[140,586,1152,1214]" box="[140,552,1152,1179]" pageId="5" pageNumber="92">
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<heading box="[140,552,1152,1179]" fontSize="11" level="2" pageId="5" pageNumber="92" reason="3">
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<taxonomicName authority="Silvestri, 1910" authorityName="Silvestri" authorityYear="1910" box="[140,552,1152,1179]" class="Diplopoda" family="Haplodesmidae" genus="Agathodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="steeli">
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<emphasis bold="true" box="[140,552,1152,1179]" pageId="5" pageNumber="92">
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<emphasis bold="true" box="[140,374,1152,1179]" italics="true" pageId="5" pageNumber="92">Agathodesmus steeli</emphasis>
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<bibRefCitation author="Silvestri" box="[381,552,1152,1178]" firstAuthor="Silvestri" pageId="5" pageNumber="92" pagination="357 - 364" refId="ref8865" refString="Silvestri F (1910) Descrizioni preliminari di nuovi generi di Diplopodi. I. Polydesmoidea. Zoologischer Anzeiger 35: 357 - 364." type="journal article" year="1910">Silvestri, 1910</bibRefCitation>
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[140,586,1187,1214]" pageId="5" pageNumber="92" type="description">
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<paragraph blockId="5.[140,586,1152,1214]" box="[140,586,1187,1214]" pageId="5" pageNumber="92">
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<figureCitation box="[140,207,1187,1214]" captionStart="Figure 1" captionStartId="1.[140,212,1622,1644]" captionTargetBox="[140,1108,1260,1603]" captionTargetId="figure@1.[140,1108,1260,1603]" captionTargetPageId="1" captionText="Figure 1. Holotype slide, photographed 1 May 2009. The label reads ‘Agathodesmus / Steeli Silv. / ♀ Typus! / Australia / Avoca. N.S.W.’" figureDoi="http://doi.org/10.5281/zenodo.3813062" httpUri="https://zenodo.org/record/3813062/files/figure.png" pageId="5" pageNumber="92">Figs 1</figureCitation>
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,
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<figureCitation box="[221,234,1188,1214]" captionStart="Figure 3" captionStartId="6.[140,212,1590,1612]" captionTargetBox="[140,1108,529,1570]" captionTargetId="figure@6.[140,1108,529,1570]" captionTargetPageId="6" captionText="Figure 3. Agathodesmus steeli Silvestri, 1910. A-C Right lateral views of anterior end; A female holotype, B female (cleared specimen), C male. D Male, ventral view of head. B-D ex AM KS107964. (A), (B) to same unknown scale; scale bars in (C), (D) = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3767246" httpUri="https://zenodo.org/record/3767246/files/figure.png" pageId="5" pageNumber="92">3</figureCitation>
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A-D,
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<figureCitation box="[301,315,1188,1214]" captionStart="Figure 4" captionStartId="7.[140,212,1179,1201]" captionTargetBox="[140,1108,165,1159]" captionTargetId="figure@7.[140,1109,165,1159]" captionTargetPageId="7" captionText="Figure 4. Agathodesmus steeli Silvestri, 1910, ex AM KS107964. A Female (left) and male (right); red colour is imaging artifact. B Male, left lateral view of posterior end; o = ozopore area. C Male, ventral view of telson. D Enlargement of C showing spinnerets. Scale bars: (A) = 2.5 mm, (B) = 0.25 mm, (C) = 0.1 mm, (D) = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3767248" httpUri="https://zenodo.org/record/3767248/files/figure.png" pageId="5" pageNumber="92">4</figureCitation>
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A-D,
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<figureCitation box="[381,394,1187,1214]" captionStart="Figure 5" captionStartId="9.[140,212,1558,1580]" captionTargetBox="[140,1108,165,1538]" captionTargetId="figure@9.[140,1109,165,1539]" captionTargetPageId="9" captionText="Figure 5. Agathodesmus steeli Silvestri, 1910, ex AM KS107964. A Male midbody sternite (anterior towards top). B Anterior view of extended left gonopod. Agathodesmus johnsi sp. n., ex AM KS94156. C Left ventrolateral view of male midbody ring (anterior towards left); no spiracles are evident above leg bases. Scale bars: (A) = 0.5 mm; (B), (C) = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767250" httpUri="https://zenodo.org/record/3767250/files/figure.png" pageId="5" pageNumber="92">5</figureCitation>
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A,
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<figureCitation box="[429,443,1187,1214]" captionStart="Figure 5" captionStartId="9.[140,212,1558,1580]" captionTargetBox="[140,1108,165,1538]" captionTargetId="figure@9.[140,1109,165,1539]" captionTargetPageId="9" captionText="Figure 5. Agathodesmus steeli Silvestri, 1910, ex AM KS107964. A Male midbody sternite (anterior towards top). B Anterior view of extended left gonopod. Agathodesmus johnsi sp. n., ex AM KS94156. C Left ventrolateral view of male midbody ring (anterior towards left); no spiracles are evident above leg bases. Scale bars: (A) = 0.5 mm; (B), (C) = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767250" httpUri="https://zenodo.org/record/3767250/files/figure.png" pageId="5" pageNumber="92">5</figureCitation>
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B,
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<figureCitation box="[476,489,1187,1213]" captionStart="Figure 6" captionStartId="10.[140,212,1558,1580]" captionTargetBox="[140,1108,423,1538]" captionTargetId="figure@10.[140,1109,422,1538]" captionTargetPageId="10" captionText="Figure 6. Agathodesmus steeli Silvestri, 1910, female ex AM KS107964. A Tubercle on midbody metatergite. B Bisegmented seta adjoining tubercle on midbody metatergite. C Dorsal view of midbody prozonite; limbus on next anterior ring at top, edge of metazonite at bottom; pl = primary limbus element, sl = secondary limbus element. Scale bars: (A) = 0.025 mm, (B) = 0.02 mm, (C) = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3767252" httpUri="https://zenodo.org/record/3767252/files/figure.png" pageId="5" pageNumber="92">6</figureCitation>
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A-C,
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<figureCitation box="[553,566,1188,1214]" captionStart="Figure 7" captionStartId="12.[140,212,1583,1605]" captionTargetBox="[140,1108,165,1563]" captionTargetId="figure@12.[140,1108,165,1564]" captionTargetPageId="12" captionText="Figure 7. Ventral views of retracted male gonopods. A Agathodesmus steeli Silvestri, 1910, ex AM KS107964. B Agathodesmus johnsi sp. n., ex AM KS94156. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767254" httpUri="https://zenodo.org/record/3767254/files/figure.png" pageId="5" pageNumber="92">7</figureCitation>
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A
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="5" pageNumber="92" type="reference_group">
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||
<paragraph blockId="5.[140,1108,1258,1320]" pageId="5" pageNumber="92">
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<treatmentCitationGroup pageId="5" pageNumber="92">
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<taxonomicName authorityName="Silvestri" authorityYear="1910" box="[140,357,1258,1284]" class="Diplopoda" family="Haplodesmidae" genus="Agathodesmus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="steeli">
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<emphasis box="[140,357,1258,1284]" italics="true" pageId="5" pageNumber="92">Agathodesmus steeli</emphasis>
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||
</taxonomicName>
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||
.
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<treatmentCitation box="[378,595,1258,1284]" pageId="5" pageNumber="92">
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||
<bibRefCitation author="Silvestri" box="[378,595,1258,1284]" firstAuthor="Silvestri" pageId="5" pageNumber="92" pagination="357 - 364" refId="ref8865" refString="Silvestri F (1910) Descrizioni preliminari di nuovi generi di Diplopodi. I. Polydesmoidea. Zoologischer Anzeiger 35: 357 - 364." type="journal article" year="1910">Silvestri 1910:362</bibRefCitation>
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</treatmentCitation>
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||
.
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||
<treatmentCitation box="[613,820,1258,1284]" pageId="5" pageNumber="92">
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||
<bibRefCitation author="Attems" box="[613,820,1258,1284]" firstAuthor="Attems" pageId="5" pageNumber="92" pagination="1 - 398" refId="ref7777" refString="Attems C (1914) Die indoaustralischen Myriopoden. Archiv fur Naturgeschichte (A) 80 (4): 1 - 398." type="journal article" year="1914">Attems 1914:283</bibRefCitation>
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</treatmentCitation>
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||
,
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||
<treatmentCitation box="[839,953,1258,1284]" pageId="5" pageNumber="92">
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||
<bibRefCitation author="Attems" box="[839,953,1258,1284]" firstAuthor="Attems" pageId="5" pageNumber="92" refId="ref7801" refString="Attems C (1940) Das Tierreich. 70. Polydesmoidea. III. Fam. Polydesmidae, Vanhoeffeniidae, Cryptodesmidae, Oniscodesmidae, Sphaerotrichopidae, Peridontodesmidae, Rhachidesmidae, Macellolophidae, Pandirodesmidae. Walter de Gruyter and Co, Berlin, 577 pp." type="book" year="1940">1940:488</bibRefCitation>
|
||
</treatmentCitation>
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||
.
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||
<treatmentCitation pageId="5" pageNumber="92">
|
||
<bibRefCitation author="Chamberlin" firstAuthor="Chamberlin" pageId="5" pageNumber="92" pagination="1 - 269" refId="ref8017" refString="Chamberlin RV (1920) Th e Myriopoda of the Australian region. Bulletin of the Museum of Comparative Zoology 64 (1): 1 - 269." type="journal article" year="1920">Chamberlin 1920:137</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
<treatmentCitation box="[310,496,1293,1319]" pageId="5" pageNumber="92">
|
||
<bibRefCitation author="Jeekel" box="[310,496,1293,1319]" firstAuthor="Jeekel" pageId="5" pageNumber="92" pagination="1 - 412" refId="ref8359" refString="Jeekel CAW (1971) Nomenclator generum et familiarum Diplopodorum: a list of the genus and family-group names in the Class Diplopoda from the 10 th edition of Linnaeus, 1758, to the end of 1957. Monografieen van de Nederlandse Entomologische Vereniging 5: i-xii, 1 - 412." type="journal article" year="1971">Jeekel 1971:310</bibRefCitation>
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||
</treatmentCitation>
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||
,
|
||
<treatmentCitation box="[507,606,1293,1320]" pageId="5" pageNumber="92">
|
||
<bibRefCitation author="Jeekel" box="[507,606,1293,1320]" firstAuthor="Jeekel" pageId="5" pageNumber="92" pagination="50 - 55" refId="ref8552" refString="Jeekel CAW (1985) Millipedes from Australia, 9: A new polydesmoid millipede from Queensland (Diplopoda, Polydesmida: Dalodesmidae). Entomologische Berichten (Amsterdam) 45: 50 - 55." type="journal article" year="1985">1985:50</bibRefCitation>
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||
</treatmentCitation>
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||
.
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||
<treatmentCitation box="[617,887,1293,1320]" pageId="5" pageNumber="92">
|
||
<bibRefCitation author="Golovatch" box="[617,887,1293,1320]" etAl="et al." firstAuthor="Golovatch" pageId="5" pageNumber="92" pagination="1 - 53" refId="ref8177" refString="Golovatch SI, Geoffroy J-J, Mauries J-P, Vanden Spiegel D (2009) Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys 7: 1 - 53." type="journal article" year="2009">Golovatch et al. 2009:3</bibRefCitation>
|
||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="5" pageNumber="92" type="materials_examined">
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||
<paragraph blockId="5.[140,1108,1363,1637]" pageId="5" pageNumber="92">
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||
<materialsCitation ID-GBIF-Occurrence="2608811664" collectionCode="NSW, MCSN" country="Australia" county="In" location="Female" municipality="Thomas Steel" pageId="5" pageNumber="92" specimenCount="1" typeStatus="holotype">
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||
<emphasis bold="true" box="[140,260,1363,1389]" pageId="5" pageNumber="92">
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||
<emphasis box="[140,256,1363,1389]" inLineHeading="true" pageId="5" pageNumber="92" reason="1">
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||
<typeStatus box="[140,256,1363,1389]" pageId="5" pageNumber="92">Holotype</typeStatus>
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||
</emphasis>
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||
.
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</emphasis>
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<location LSID="urn:lsid:plazi:treatment:9B0D9E62FF9CCC31FF5E5E9D854FB194:167B79AFFF9CCC3FFEDF5F4E807BB673" box="[269,352,1363,1389]" country="Australia" county="In" municipality="Thomas Steel" name="Female" pageId="5" pageNumber="92">Female</location>
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, permanently mounted on microscope slide in two pieces, with a break between rings 9 and 10 (
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<figureCitation box="[485,554,1399,1425]" captionStart="Figure 1" captionStartId="1.[140,212,1622,1644]" captionTargetBox="[140,1108,1260,1603]" captionTargetId="figure@1.[140,1108,1260,1603]" captionTargetPageId="1" captionText="Figure 1. Holotype slide, photographed 1 May 2009. The label reads ‘Agathodesmus / Steeli Silv. / ♀ Typus! / Australia / Avoca. N.S.W.’" figureDoi="http://doi.org/10.5281/zenodo.3813062" httpUri="https://zenodo.org/record/3813062/files/figure.png" pageId="5" pageNumber="92">Figs 1</figureCitation>
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||
, 3A).
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||
<location LSID="urn:lsid:plazi:treatment:9B0D9E62FF9CCC31FF5E5E9D854FB194:167B79AFFF9CCC3FFDA25F6A83ACB68F" box="[624,695,1399,1425]" country="Australia" county="In" municipality="Thomas Steel" name="Avoca" pageId="5" pageNumber="92">Avoca</location>
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||
,
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||
<collectionCode box="[706,771,1399,1425]" pageId="5" pageNumber="92">NSW</collectionCode>
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||
,
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||
<collectingCountry box="[782,885,1399,1425]" name="Australia" pageId="5" pageNumber="92">Australia</collectingCountry>
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||
,
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||
<collectingMunicipality box="[896,1049,1399,1425]" pageId="5" pageNumber="92">Thomas Steel</collectingMunicipality>
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||
, date not known.
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||
<collectingCounty box="[279,304,1434,1460]" pageId="5" pageNumber="92">In</collectingCounty>
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||
<collectionCode box="[311,399,1434,1460]" country="Italy" httpUri="http://grbio.org/cool/353t-cj9k" name="Museo Civico di Storia Naturale, Verona" pageId="5" pageNumber="92">MCSN</collectionCode>
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||
.
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph blockId="5.[140,1108,1363,1637]" box="[185,513,1469,1496]" pageId="5" pageNumber="92">
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||
<emphasis bold="true" box="[185,308,1469,1495]" pageId="5" pageNumber="92">
|
||
<emphasis box="[185,303,1469,1495]" inLineHeading="true" pageId="5" pageNumber="92" reason="1">
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||
<typeStatus box="[185,303,1469,1495]" pageId="5" pageNumber="92">Paratypes</typeStatus>
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||
</emphasis>
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||
.
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||
</emphasis>
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||
None designated.
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||
</paragraph>
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||
<paragraph blockId="5.[140,1108,1363,1637]" pageId="5" pageNumber="92">
|
||
<emphasis bold="true" box="[185,493,1504,1530]" pageId="5" pageNumber="92">
|
||
<emphasis box="[185,257,1504,1530]" inLineHeading="true" pageId="5" pageNumber="92" reason="1">Other</emphasis>
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||
material examined.
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||
</emphasis>
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||
<specimenCount box="[500,589,1504,1531]" pageId="5" pageNumber="92" type="male">7 males</specimenCount>
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,
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<specimenCount box="[601,710,1504,1530]" pageId="5" pageNumber="92" type="female">4 females</specimenCount>
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, 1 stadium VI female, 2 stadium V females, 1 stadium IV female, Knights Hill,
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||
<collectionCode box="[651,716,1540,1566]" pageId="5" pageNumber="92">NSW</collectionCode>
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||
,
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||
<geoCoordinate box="[729,859,1540,1566]" degrees="34" direction="south" minutes="37" orientation="latitude" pageId="5" pageNumber="92" precision="15" seconds="07" value="-34.61861">34°37’07”S</geoCoordinate>
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||
<geoCoordinate box="[868,1015,1539,1566]" degrees="150" direction="east" minutes="42" orientation="longitude" pageId="5" pageNumber="92" precision="15" seconds="38" value="150.71056">150°42’38”E</geoCoordinate>
|
||
±
|
||
<quantity box="[1040,1103,1539,1566]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="92" unit="m" value="25.0">25 m</quantity>
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||
,
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<quantity box="[140,218,1575,1601]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="5" pageNumber="92" unit="m" value="720.0">720 m</quantity>
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||
,
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||
<date box="[230,388,1575,1601]" pageId="5" pageNumber="92" value="2009-05-14">14 May 2009</date>
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||
, R. Mesibov and T. Moule, wet eucalypt forest,
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||
<collectionCode box="[932,977,1575,1601]" country="Australia" name="Australian Museum" pageId="5" pageNumber="92" type="Museum">AM</collectionCode>
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||
<accessionNumber box="[985,1108,1575,1601]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/KS107964" pageId="5" pageNumber="92">KS107964</accessionNumber>
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||
(three males, two females dissected).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="6" pageNumber="93" type="diagnosis">
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||
<paragraph blockId="6.[140,1108,166,475]" pageId="6" pageNumber="93">
|
||
<emphasis bold="true" box="[185,312,166,192]" pageId="6" pageNumber="93">
|
||
<emphasis box="[185,307,166,192]" inLineHeading="true" pageId="6" pageNumber="93" reason="1">Diagnosis</emphasis>
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||
.
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||
</emphasis>
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||
Head + 19 rings; gonopod telopodite with distal portion directed basally and slightly laterally near origin and with broad lateral branch apically expanded and divided into three anterobasally curving lobes.
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection lastPageId="10" lastPageNumber="97" pageId="6" pageNumber="93" type="description">
|
||
<paragraph blockId="6.[140,1108,166,475]" pageId="6" pageNumber="93">
|
||
<emphasis bold="true" box="[185,335,272,298]" pageId="6" pageNumber="93">
|
||
<emphasis box="[185,330,272,298]" inLineHeading="true" pageId="6" pageNumber="93" reason="1">Description</emphasis>
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||
.
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||
</emphasis>
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||
The original description (
|
||
<bibRefCitation author="Silvestri" box="[629,788,272,298]" firstAuthor="Silvestri" pageId="6" pageNumber="93" pagination="357 - 364" refId="ref8865" refString="Silvestri F (1910) Descrizioni preliminari di nuovi generi di Diplopodi. I. Polydesmoidea. Zoologischer Anzeiger 35: 357 - 364." type="journal article" year="1910">Silvestri 1910</bibRefCitation>
|
||
) is quoted in the Appendix. What follows is based on my examination of both the
|
||
<typeStatus box="[763,862,307,333]" pageId="6" pageNumber="93">holotype</typeStatus>
|
||
and the Knights Hill material.
|
||
</paragraph>
|
||
<paragraph blockId="6.[140,1108,166,475]" lastBlockId="8.[140,1108,167,1672]" lastPageId="8" lastPageNumber="95" pageId="6" pageNumber="93">
|
||
Adult with head + 19 rings (
|
||
<figureCitation box="[512,602,378,404]" captionStart="Figure 4" captionStartId="7.[140,212,1179,1201]" captionTargetBox="[140,1108,165,1159]" captionTargetId="figure@7.[140,1109,165,1159]" captionTargetPageId="7" captionText="Figure 4. Agathodesmus steeli Silvestri, 1910, ex AM KS107964. A Female (left) and male (right); red colour is imaging artifact. B Male, left lateral view of posterior end; o = ozopore area. C Male, ventral view of telson. D Enlargement of C showing spinnerets. Scale bars: (A) = 2.5 mm, (B) = 0.25 mm, (C) = 0.1 mm, (D) = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3767248" httpUri="https://zenodo.org/record/3767248/files/figure.png" pageId="6" pageNumber="93">Fig. 4A</figureCitation>
|
||
). Live and freshly preserved adults pale with faint reddish pigmentation dorsally; in some individuals, pigment concentrated in transverse band at rear of metazonite. Male/female approximate dimensions: length 4.5/5.5 mm, maximum width with paranota 0.5/0.6 mm, midbody vertical diameter 0.4/0.4 mm. Head (
|
||
<figureCitation box="[368,511,1399,1425]" captionStart="Figure 3" captionStartId="6.[140,212,1590,1612]" captionTargetBox="[140,1108,529,1570]" captionTargetId="figure@6.[140,1108,529,1570]" captionTargetPageId="6" captionText="Figure 3. Agathodesmus steeli Silvestri, 1910. A-C Right lateral views of anterior end; A female holotype, B female (cleared specimen), C male. D Male, ventral view of head. B-D ex AM KS107964. (A), (B) to same unknown scale; scale bars in (C), (D) = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3767246" httpUri="https://zenodo.org/record/3767246/files/figure.png" pageId="7" pageNumber="94">Figs 3C, 3D</figureCitation>
|
||
) about as wide as collum, overall body width almost uniform, tapering only slightly posteriorly from collum. Head facing downwards (
|
||
<figureCitation captionStart="Figure 3" captionStartId="6.[140,212,1590,1612]" captionTargetBox="[140,1108,529,1570]" captionTargetId="figure@6.[140,1108,529,1570]" captionTargetPageId="6" captionText="Figure 3. Agathodesmus steeli Silvestri, 1910. A-C Right lateral views of anterior end; A female holotype, B female (cleared specimen), C male. D Male, ventral view of head. B-D ex AM KS107964. (A), (B) to same unknown scale; scale bars in (C), (D) = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3767246" httpUri="https://zenodo.org/record/3767246/files/figure.png" pageId="7" pageNumber="94">Fig. 3C</figureCitation>
|
||
), with clypeus, frons and ventral part of vertex almost parallel to substrate and only slightly convex. Antenna (
|
||
<figureCitation box="[438,528,1504,1530]" captionStart="Figure 3" captionStartId="6.[140,212,1590,1612]" captionTargetBox="[140,1108,529,1570]" captionTargetId="figure@6.[140,1108,529,1570]" captionTargetPageId="6" captionText="Figure 3. Agathodesmus steeli Silvestri, 1910. A-C Right lateral views of anterior end; A female holotype, B female (cleared specimen), C male. D Male, ventral view of head. B-D ex AM KS107964. (A), (B) to same unknown scale; scale bars in (C), (D) = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3767246" httpUri="https://zenodo.org/record/3767246/files/figure.png" pageId="7" pageNumber="94">Fig. 3D</figureCitation>
|
||
) short, stout, clavate, held close to head, antennomeres 2 and 3 lying in broad, shallow excavation on head; antennomere 6 widest and longest; antennomeres 2-5 about equal in length, decreasing slightly in diameter from 5 to 2. Collum with slightly convex anterior margin and broadly convex posterior margin; corners rounded (
|
||
<figureCitation box="[448,542,1645,1671]" captionStart="Figure 3" captionStartId="6.[140,212,1590,1612]" captionTargetBox="[140,1108,529,1570]" captionTargetId="figure@6.[140,1108,529,1570]" captionTargetPageId="6" captionText="Figure 3. Agathodesmus steeli Silvestri, 1910. A-C Right lateral views of anterior end; A female holotype, B female (cleared specimen), C male. D Male, ventral view of head. B-D ex AM KS107964. (A), (B) to same unknown scale; scale bars in (C), (D) = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3767246" httpUri="https://zenodo.org/record/3767246/files/figure.png" pageId="7" pageNumber="94">Fig. 3C</figureCitation>
|
||
). Ring 2 tergite largest, extending basally, later- ally and anteriorly well below collum corner (
|
||
<figureCitation box="[657,846,167,193]" captionStart="Figure 3" captionStartId="6.[140,212,1590,1612]" captionTargetBox="[140,1108,529,1570]" captionTargetId="figure@6.[140,1108,529,1570]" captionTargetPageId="6" captionText="Figure 3. Agathodesmus steeli Silvestri, 1910. A-C Right lateral views of anterior end; A female holotype, B female (cleared specimen), C male. D Male, ventral view of head. B-D ex AM KS107964. (A), (B) to same unknown scale; scale bars in (C), (D) = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3767246" httpUri="https://zenodo.org/record/3767246/files/figure.png" pageId="8" pageNumber="95">Figs 3A, 3B, 3C</figureCitation>
|
||
). Ring 2 and 3 tergites edged with 5-6 and 4 large tubercles, respectively (
|
||
<figureCitation box="[721,912,202,228]" captionStart="Figure 3" captionStartId="6.[140,212,1590,1612]" captionTargetBox="[140,1108,529,1570]" captionTargetId="figure@6.[140,1108,529,1570]" captionTargetPageId="6" captionText="Figure 3. Agathodesmus steeli Silvestri, 1910. A-C Right lateral views of anterior end; A female holotype, B female (cleared specimen), C male. D Male, ventral view of head. B-D ex AM KS107964. (A), (B) to same unknown scale; scale bars in (C), (D) = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3767246" httpUri="https://zenodo.org/record/3767246/files/figure.png" pageId="8" pageNumber="95">Figs 3A, 3B, 3C</figureCitation>
|
||
); posterior rings, including apodous ring 18, with row of 4 large tubercles just above leg bases forming all or part of lateral extension of metatergite, the anteriormost tubercle smaller than the posterior 3. Prozonites sharply demarcated from metazonites (
|
||
<figureCitation box="[839,931,307,334]" captionStart="Figure 3" captionStartId="6.[140,212,1590,1612]" captionTargetBox="[140,1108,529,1570]" captionTargetId="figure@6.[140,1108,529,1570]" captionTargetPageId="6" captionText="Figure 3. Agathodesmus steeli Silvestri, 1910. A-C Right lateral views of anterior end; A female holotype, B female (cleared specimen), C male. D Male, ventral view of head. B-D ex AM KS107964. (A), (B) to same unknown scale; scale bars in (C), (D) = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3767246" httpUri="https://zenodo.org/record/3767246/files/figure.png" pageId="8" pageNumber="95">Figs 3C</figureCitation>
|
||
,
|
||
<figureCitation box="[943,978,308,334]" captionStart="Figure 4" captionStartId="7.[140,212,1179,1201]" captionTargetBox="[140,1108,165,1159]" captionTargetId="figure@7.[140,1109,165,1159]" captionTargetPageId="7" captionText="Figure 4. Agathodesmus steeli Silvestri, 1910, ex AM KS107964. A Female (left) and male (right); red colour is imaging artifact. B Male, left lateral view of posterior end; o = ozopore area. C Male, ventral view of telson. D Enlargement of C showing spinnerets. Scale bars: (A) = 2.5 mm, (B) = 0.25 mm, (C) = 0.1 mm, (D) = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3767248" httpUri="https://zenodo.org/record/3767248/files/figure.png" pageId="8" pageNumber="95">4B</figureCitation>
|
||
,
|
||
<figureCitation box="[991,1029,307,334]" captionStart="Figure 6" captionStartId="10.[140,212,1558,1580]" captionTargetBox="[140,1108,423,1538]" captionTargetId="figure@10.[140,1109,422,1538]" captionTargetPageId="10" captionText="Figure 6. Agathodesmus steeli Silvestri, 1910, female ex AM KS107964. A Tubercle on midbody metatergite. B Bisegmented seta adjoining tubercle on midbody metatergite. C Dorsal view of midbody prozonite; limbus on next anterior ring at top, edge of metazonite at bottom; pl = primary limbus element, sl = secondary limbus element. Scale bars: (A) = 0.025 mm, (B) = 0.02 mm, (C) = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3767252" httpUri="https://zenodo.org/record/3767252/files/figure.png" pageId="8" pageNumber="95">6C</figureCitation>
|
||
). Ozopore (
|
||
<figureCitation box="[205,289,343,369]" captionStart="Figure 4" captionStartId="7.[140,212,1179,1201]" captionTargetBox="[140,1108,165,1159]" captionTargetId="figure@7.[140,1109,165,1159]" captionTargetPageId="7" captionText="Figure 4. Agathodesmus steeli Silvestri, 1910, ex AM KS107964. A Female (left) and male (right); red colour is imaging artifact. B Male, left lateral view of posterior end; o = ozopore area. C Male, ventral view of telson. D Enlargement of C showing spinnerets. Scale bars: (A) = 2.5 mm, (B) = 0.25 mm, (C) = 0.1 mm, (D) = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3767248" httpUri="https://zenodo.org/record/3767248/files/figure.png" pageId="8" pageNumber="95">Fig. 4B</figureCitation>
|
||
) very small, not raised, in small, non-tuberculated area just above middle of group of 4 larger tubercles forming lateral metatergal extension; pore formula 5, 7, 9, 10, 12, 13, 15-18. Sternites on diplosegments (
|
||
<figureCitation box="[751,840,413,440]" captionStart="Figure 5" captionStartId="9.[140,212,1558,1580]" captionTargetBox="[140,1108,165,1538]" captionTargetId="figure@9.[140,1109,165,1539]" captionTargetPageId="9" captionText="Figure 5. Agathodesmus steeli Silvestri, 1910, ex AM KS107964. A Male midbody sternite (anterior towards top). B Anterior view of extended left gonopod. Agathodesmus johnsi sp. n., ex AM KS94156. C Left ventrolateral view of male midbody ring (anterior towards left); no spiracles are evident above leg bases. Scale bars: (A) = 0.5 mm; (B), (C) = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767250" httpUri="https://zenodo.org/record/3767250/files/figure.png" pageId="8" pageNumber="95">Fig. 5A</figureCitation>
|
||
) longer than wide, not setose, with distinct longitudinal and transverse impressions. Legs short, stout; relative podomere lengths tarsus>(prefemur, femur)>(postfemur, tibia); claw large, about twothirds tarsus length. Spiracles not evident. Telson facing downwards (
|
||
<figureCitation box="[947,1037,519,545]" captionStart="Figure 4" captionStartId="7.[140,212,1179,1201]" captionTargetBox="[140,1108,165,1159]" captionTargetId="figure@7.[140,1109,165,1159]" captionTargetPageId="7" captionText="Figure 4. Agathodesmus steeli Silvestri, 1910, ex AM KS107964. A Female (left) and male (right); red colour is imaging artifact. B Male, left lateral view of posterior end; o = ozopore area. C Male, ventral view of telson. D Enlargement of C showing spinnerets. Scale bars: (A) = 2.5 mm, (B) = 0.25 mm, (C) = 0.1 mm, (D) = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3767248" httpUri="https://zenodo.org/record/3767248/files/figure.png" pageId="8" pageNumber="95">Fig. 4B</figureCitation>
|
||
), anal valves parallel to substrate and almost flat. Hypoproct trapezoidal (
|
||
<figureCitation box="[885,973,554,580]" captionStart="Figure 4" captionStartId="7.[140,212,1179,1201]" captionTargetBox="[140,1108,165,1159]" captionTargetId="figure@7.[140,1109,165,1159]" captionTargetPageId="7" captionText="Figure 4. Agathodesmus steeli Silvestri, 1910, ex AM KS107964. A Female (left) and male (right); red colour is imaging artifact. B Male, left lateral view of posterior end; o = ozopore area. C Male, ventral view of telson. D Enlargement of C showing spinnerets. Scale bars: (A) = 2.5 mm, (B) = 0.25 mm, (C) = 0.1 mm, (D) = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3767248" httpUri="https://zenodo.org/record/3767248/files/figure.png" pageId="8" pageNumber="95">Fig. 4C</figureCitation>
|
||
); spinnerets in square array (
|
||
<figureCitation box="[316,458,589,615]" captionStart="Figure 4" captionStartId="7.[140,212,1179,1201]" captionTargetBox="[140,1108,165,1159]" captionTargetId="figure@7.[140,1109,165,1159]" captionTargetPageId="7" captionText="Figure 4. Agathodesmus steeli Silvestri, 1910, ex AM KS107964. A Female (left) and male (right); red colour is imaging artifact. B Male, left lateral view of posterior end; o = ozopore area. C Male, ventral view of telson. D Enlargement of C showing spinnerets. Scale bars: (A) = 2.5 mm, (B) = 0.25 mm, (C) = 0.1 mm, (D) = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3767248" httpUri="https://zenodo.org/record/3767248/files/figure.png" pageId="8" pageNumber="95">Figs 4C, 4D</figureCitation>
|
||
); spinneret setae with single, low sheath, each seta in deep, walled depression.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767246" ID-Zenodo-Dep="3767246" httpUri="https://zenodo.org/record/3767246/files/figure.png" pageId="6" pageNumber="93" startId="6.[140,212,1590,1612]" targetBox="[140,1108,529,1570]" targetPageId="6">
|
||
<paragraph blockId="6.[140,1108,1590,1677]" pageId="6" pageNumber="93">
|
||
<emphasis bold="true" box="[140,237,1590,1612]" pageId="6" pageNumber="93">Figure 3.</emphasis>
|
||
<taxonomicName authority="Silvestri, 1910" authorityName="Silvestri" authorityYear="1910" box="[242,552,1590,1612]" class="Diplopoda" family="Haplodesmidae" genus="Agathodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="steeli">
|
||
<emphasis box="[242,412,1590,1611]" italics="true" pageId="6" pageNumber="93">Agathodesmus steeli</emphasis>
|
||
<bibRefCitation author="Silvestri" box="[418,552,1590,1612]" firstAuthor="Silvestri" pageId="6" pageNumber="93" pagination="357 - 364" refId="ref8865" refString="Silvestri F (1910) Descrizioni preliminari di nuovi generi di Diplopodi. I. Polydesmoidea. Zoologischer Anzeiger 35: 357 - 364." type="journal article" year="1910">Silvestri, 1910</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis bold="true" box="[560,606,1590,1612]" pageId="6" pageNumber="93">A -C</emphasis>
|
||
Right lateral views of anterior end;
|
||
<emphasis bold="true" box="[931,950,1590,1612]" pageId="6" pageNumber="93">A</emphasis>
|
||
female holotype,
|
||
<emphasis bold="true" box="[140,157,1622,1644]" pageId="6" pageNumber="93">B</emphasis>
|
||
female (cleared specimen),
|
||
<emphasis bold="true" box="[414,433,1622,1644]" pageId="6" pageNumber="93">C</emphasis>
|
||
male.
|
||
<emphasis bold="true" box="[495,514,1622,1644]" pageId="6" pageNumber="93">D</emphasis>
|
||
Male, ventral view of head.
|
||
<emphasis bold="true" box="[778,822,1622,1644]" pageId="6" pageNumber="93">B -D</emphasis>
|
||
ex AM KS107964. (A), (B) to same unknown scale; scale bars in (C), (D) = 0.25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767248" ID-Zenodo-Dep="3767248" httpUri="https://zenodo.org/record/3767248/files/figure.png" pageId="7" pageNumber="94" startId="7.[140,212,1179,1201]" targetBox="[140,1108,165,1159]" targetPageId="7">
|
||
<paragraph blockId="7.[140,1107,1179,1297]" pageId="7" pageNumber="94">
|
||
<emphasis bold="true" box="[140,237,1179,1201]" pageId="7" pageNumber="94">Figure 4.</emphasis>
|
||
<taxonomicName authority="Silvestri, 1910" authorityName="Silvestri" authorityYear="1910" box="[245,562,1179,1201]" class="Diplopoda" family="Haplodesmidae" genus="Agathodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="94" phylum="Arthropoda" rank="species" species="steeli">
|
||
<emphasis box="[245,418,1179,1200]" italics="true" pageId="7" pageNumber="94">Agathodesmus steeli</emphasis>
|
||
<bibRefCitation author="Silvestri" box="[426,562,1179,1201]" firstAuthor="Silvestri" pageId="7" pageNumber="94" pagination="357 - 364" refId="ref8865" refString="Silvestri F (1910) Descrizioni preliminari di nuovi generi di Diplopodi. I. Polydesmoidea. Zoologischer Anzeiger 35: 357 - 364." type="journal article" year="1910">Silvestri, 1910</bibRefCitation>
|
||
</taxonomicName>
|
||
, ex AM KS107964.
|
||
<emphasis bold="true" box="[759,778,1179,1201]" pageId="7" pageNumber="94">A</emphasis>
|
||
Female (left) and male (right); red colour is imaging artifact.
|
||
<emphasis bold="true" box="[389,406,1211,1233]" pageId="7" pageNumber="94">B</emphasis>
|
||
Male, left lateral view of posterior end; o = ozopore area.
|
||
<emphasis bold="true" box="[958,977,1211,1233]" pageId="7" pageNumber="94">C</emphasis>
|
||
Male, ventral view of telson.
|
||
<emphasis bold="true" box="[279,298,1243,1265]" pageId="7" pageNumber="94">D</emphasis>
|
||
Enlargement of
|
||
<emphasis bold="true" box="[453,472,1243,1265]" pageId="7" pageNumber="94">C</emphasis>
|
||
showing spinnerets. Scale bars: (A) = 2.5 mm, (B) = 0.25 mm, (C) = 0.1 mm, (D) = 0.02 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="8.[140,1108,167,1672]" pageId="8" pageNumber="95">
|
||
Integument richly and densely sculptured (
|
||
<figureCitation box="[674,769,659,686]" captionStart="Figure 3" captionStartId="6.[140,212,1590,1612]" captionTargetBox="[140,1108,529,1570]" captionTargetId="figure@6.[140,1108,529,1570]" captionTargetPageId="6" captionText="Figure 3. Agathodesmus steeli Silvestri, 1910. A-C Right lateral views of anterior end; A female holotype, B female (cleared specimen), C male. D Male, ventral view of head. B-D ex AM KS107964. (A), (B) to same unknown scale; scale bars in (C), (D) = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3767246" httpUri="https://zenodo.org/record/3767246/files/figure.png" pageId="8" pageNumber="95">Figs 3D</figureCitation>
|
||
,
|
||
<figureCitation box="[782,819,660,686]" captionStart="Figure 4" captionStartId="7.[140,212,1179,1201]" captionTargetBox="[140,1108,165,1159]" captionTargetId="figure@7.[140,1109,165,1159]" captionTargetPageId="7" captionText="Figure 4. Agathodesmus steeli Silvestri, 1910, ex AM KS107964. A Female (left) and male (right); red colour is imaging artifact. B Male, left lateral view of posterior end; o = ozopore area. C Male, ventral view of telson. D Enlargement of C showing spinnerets. Scale bars: (A) = 2.5 mm, (B) = 0.25 mm, (C) = 0.1 mm, (D) = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3767248" httpUri="https://zenodo.org/record/3767248/files/figure.png" pageId="8" pageNumber="95">4C</figureCitation>
|
||
,
|
||
<figureCitation box="[833,918,659,686]" captionStart="Figure 5" captionStartId="9.[140,212,1558,1580]" captionTargetBox="[140,1108,165,1538]" captionTargetId="figure@9.[140,1109,165,1539]" captionTargetPageId="9" captionText="Figure 5. Agathodesmus steeli Silvestri, 1910, ex AM KS107964. A Male midbody sternite (anterior towards top). B Anterior view of extended left gonopod. Agathodesmus johnsi sp. n., ex AM KS94156. C Left ventrolateral view of male midbody ring (anterior towards left); no spiracles are evident above leg bases. Scale bars: (A) = 0.5 mm; (B), (C) = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767250" httpUri="https://zenodo.org/record/3767250/files/figure.png" pageId="8" pageNumber="95">5A, 5C</figureCitation>
|
||
,
|
||
<figureCitation box="[932,945,659,685]" captionStart="Figure 6" captionStartId="10.[140,212,1558,1580]" captionTargetBox="[140,1108,423,1538]" captionTargetId="figure@10.[140,1109,422,1538]" captionTargetPageId="10" captionText="Figure 6. Agathodesmus steeli Silvestri, 1910, female ex AM KS107964. A Tubercle on midbody metatergite. B Bisegmented seta adjoining tubercle on midbody metatergite. C Dorsal view of midbody prozonite; limbus on next anterior ring at top, edge of metazonite at bottom; pl = primary limbus element, sl = secondary limbus element. Scale bars: (A) = 0.025 mm, (B) = 0.02 mm, (C) = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3767252" httpUri="https://zenodo.org/record/3767252/files/figure.png" pageId="8" pageNumber="95">6</figureCitation>
|
||
A-C, 7A, 7B). Most of body covered with cuticle raised in cellular mesh of narrow folds, often with minute bumps (adorned with even smaller bumps) at or near fold junctions. Integument raised further as tubercles (
|
||
<figureCitation box="[522,610,765,792]" captionStart="Figure 6" captionStartId="10.[140,212,1558,1580]" captionTargetBox="[140,1108,423,1538]" captionTargetId="figure@10.[140,1109,422,1538]" captionTargetPageId="10" captionText="Figure 6. Agathodesmus steeli Silvestri, 1910, female ex AM KS107964. A Tubercle on midbody metatergite. B Bisegmented seta adjoining tubercle on midbody metatergite. C Dorsal view of midbody prozonite; limbus on next anterior ring at top, edge of metazonite at bottom; pl = primary limbus element, sl = secondary limbus element. Scale bars: (A) = 0.025 mm, (B) = 0.02 mm, (C) = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3767252" httpUri="https://zenodo.org/record/3767252/files/figure.png" pageId="8" pageNumber="95">Fig. 6A</figureCitation>
|
||
) of varying sizes on head, collum, tergites, metatergites and telson, the largest tubercles forming paranotum-like extensions on posterior rings; tubercles and some other parts of integument with minute, finger-like projections (
|
||
<figureCitation box="[278,372,871,897]" captionStart="Figure 4" captionStartId="7.[140,212,1179,1201]" captionTargetBox="[140,1108,165,1159]" captionTargetId="figure@7.[140,1109,165,1159]" captionTargetPageId="7" captionText="Figure 4. Agathodesmus steeli Silvestri, 1910, ex AM KS107964. A Female (left) and male (right); red colour is imaging artifact. B Male, left lateral view of posterior end; o = ozopore area. C Male, ventral view of telson. D Enlargement of C showing spinnerets. Scale bars: (A) = 2.5 mm, (B) = 0.25 mm, (C) = 0.1 mm, (D) = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3767248" httpUri="https://zenodo.org/record/3767248/files/figure.png" pageId="8" pageNumber="95">Figs 4D</figureCitation>
|
||
,
|
||
<figureCitation box="[382,464,871,898]" captionStart="Figure 6" captionStartId="10.[140,212,1558,1580]" captionTargetBox="[140,1108,423,1538]" captionTargetId="figure@10.[140,1109,422,1538]" captionTargetPageId="10" captionText="Figure 6. Agathodesmus steeli Silvestri, 1910, female ex AM KS107964. A Tubercle on midbody metatergite. B Bisegmented seta adjoining tubercle on midbody metatergite. C Dorsal view of midbody prozonite; limbus on next anterior ring at top, edge of metazonite at bottom; pl = primary limbus element, sl = secondary limbus element. Scale bars: (A) = 0.025 mm, (B) = 0.02 mm, (C) = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3767252" httpUri="https://zenodo.org/record/3767252/files/figure.png" pageId="8" pageNumber="95">6A, 6B</figureCitation>
|
||
), often arising along ‘mesh-cell’ boundaries. Cell boundaries at rear of metazonite extended as lappets, forming secondary limbus above primary limbus of uniform, triangular elements (
|
||
<figureCitation box="[608,700,941,968]" captionStart="Figure 6" captionStartId="10.[140,212,1558,1580]" captionTargetBox="[140,1108,423,1538]" captionTargetId="figure@10.[140,1109,422,1538]" captionTargetPageId="10" captionText="Figure 6. Agathodesmus steeli Silvestri, 1910, female ex AM KS107964. A Tubercle on midbody metatergite. B Bisegmented seta adjoining tubercle on midbody metatergite. C Dorsal view of midbody prozonite; limbus on next anterior ring at top, edge of metazonite at bottom; pl = primary limbus element, sl = secondary limbus element. Scale bars: (A) = 0.025 mm, (B) = 0.02 mm, (C) = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3767252" httpUri="https://zenodo.org/record/3767252/files/figure.png" pageId="8" pageNumber="95">Fig. 6C</figureCitation>
|
||
). Setae of normal
|
||
<typeStatus box="[916,964,942,968]" pageId="8" pageNumber="95">type</typeStatus>
|
||
on legs and some other surfaces; a bisegmented seta with flattened, expanded tip (
|
||
<figureCitation box="[921,1005,976,1003]" captionStart="Figure 6" captionStartId="10.[140,212,1558,1580]" captionTargetBox="[140,1108,423,1538]" captionTargetId="figure@10.[140,1109,422,1538]" captionTargetPageId="10" captionText="Figure 6. Agathodesmus steeli Silvestri, 1910, female ex AM KS107964. A Tubercle on midbody metatergite. B Bisegmented seta adjoining tubercle on midbody metatergite. C Dorsal view of midbody prozonite; limbus on next anterior ring at top, edge of metazonite at bottom; pl = primary limbus element, sl = secondary limbus element. Scale bars: (A) = 0.025 mm, (B) = 0.02 mm, (C) = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.3767252" httpUri="https://zenodo.org/record/3767252/files/figure.png" pageId="8" pageNumber="95">Fig. 6B</figureCitation>
|
||
) on each ‘paranotum’ tubercle and in association with some dorsal tubercles.
|
||
</paragraph>
|
||
<paragraph blockId="8.[140,1108,167,1672]" lastBlockId="10.[140,1108,167,369]" lastPageId="10" lastPageNumber="97" pageId="8" pageNumber="95">
|
||
Male with gonopore opening at tip of cylindrical projection about 1/3 the length of leg 2 coxa, arising distomedially on the coxa. First legs somewhat swollen (
|
||
<figureCitation box="[1005,1093,1082,1108]" captionStart="Figure 3" captionStartId="6.[140,212,1590,1612]" captionTargetBox="[140,1108,529,1570]" captionTargetId="figure@6.[140,1108,529,1570]" captionTargetPageId="6" captionText="Figure 3. Agathodesmus steeli Silvestri, 1910. A-C Right lateral views of anterior end; A female holotype, B female (cleared specimen), C male. D Male, ventral view of head. B-D ex AM KS107964. (A), (B) to same unknown scale; scale bars in (C), (D) = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3767246" httpUri="https://zenodo.org/record/3767246/files/figure.png" pageId="8" pageNumber="95">Fig. 3C</figureCitation>
|
||
), no other anterior legs enlarged; neither sphaerotrichomes nor brush setae on any legs. Leg 7 bases well separated; leg 6 bases slightly separated, with a pair of short, rounded projections between coxae. Gonopod aperture oval (
|
||
<figureCitation box="[725,813,1187,1214]" captionStart="Figure 7" captionStartId="12.[140,212,1583,1605]" captionTargetBox="[140,1108,165,1563]" captionTargetId="figure@12.[140,1108,165,1564]" captionTargetPageId="12" captionText="Figure 7. Ventral views of retracted male gonopods. A Agathodesmus steeli Silvestri, 1910, ex AM KS107964. B Agathodesmus johnsi sp. n., ex AM KS94156. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767254" httpUri="https://zenodo.org/record/3767254/files/figure.png" pageId="8" pageNumber="95">Fig. 7A</figureCitation>
|
||
), rim a little raised laterally. Gonocoxae (
|
||
<figureCitation box="[313,398,1223,1249]" captionStart="Figure 7" captionStartId="12.[140,212,1583,1605]" captionTargetBox="[140,1108,165,1563]" captionTargetId="figure@12.[140,1108,165,1564]" captionTargetPageId="12" captionText="Figure 7. Ventral views of retracted male gonopods. A Agathodesmus steeli Silvestri, 1910, ex AM KS107964. B Agathodesmus johnsi sp. n., ex AM KS94156. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767254" httpUri="https://zenodo.org/record/3767254/files/figure.png" pageId="8" pageNumber="95">Fig. 7A</figureCitation>
|
||
) occupying full width of aperture; tapering a little distally; with mesh-like integumental sculpture and without setae; firmly joined medially near distal end. Telopodite (
|
||
<figureCitation box="[332,418,1293,1319]" captionStart="Figure 7" captionStartId="12.[140,212,1583,1605]" captionTargetBox="[140,1108,165,1563]" captionTargetId="figure@12.[140,1108,165,1564]" captionTargetPageId="12" captionText="Figure 7. Ventral views of retracted male gonopods. A Agathodesmus steeli Silvestri, 1910, ex AM KS107964. B Agathodesmus johnsi sp. n., ex AM KS94156. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767254" httpUri="https://zenodo.org/record/3767254/files/figure.png" pageId="8" pageNumber="95">Fig. 7A</figureCitation>
|
||
) short, compact, when retracted reaching leg 7 base; broadly joined to gonocoxa (
|
||
<figureCitation box="[373,463,1328,1355]" captionStart="Figure 5" captionStartId="9.[140,212,1558,1580]" captionTargetBox="[140,1108,165,1538]" captionTargetId="figure@9.[140,1109,165,1539]" captionTargetPageId="9" captionText="Figure 5. Agathodesmus steeli Silvestri, 1910, ex AM KS107964. A Male midbody sternite (anterior towards top). B Anterior view of extended left gonopod. Agathodesmus johnsi sp. n., ex AM KS94156. C Left ventrolateral view of male midbody ring (anterior towards left); no spiracles are evident above leg bases. Scale bars: (A) = 0.5 mm; (B), (C) = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767250" httpUri="https://zenodo.org/record/3767250/files/figure.png" pageId="8" pageNumber="95">Figs 5B</figureCitation>
|
||
,
|
||
<figureCitation box="[476,513,1328,1354]" captionStart="Figure 7" captionStartId="12.[140,212,1583,1605]" captionTargetBox="[140,1108,165,1563]" captionTargetId="figure@12.[140,1108,165,1564]" captionTargetPageId="12" captionText="Figure 7. Ventral views of retracted male gonopods. A Agathodesmus steeli Silvestri, 1910, ex AM KS107964. B Agathodesmus johnsi sp. n., ex AM KS94156. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767254" httpUri="https://zenodo.org/record/3767254/files/figure.png" pageId="8" pageNumber="95">7A</figureCitation>
|
||
); no trace of cannula or prostatic groove; no integumental sculpturing; divided into more or less cylindrical basal portion and flattened distal portion. Basal portion of telopodite with blunt, basally directed projection arising posteromedial to junction with gonocoxa; portion terminating in flat, rounded tab bending posteriorly; with a few short setae on basal half of posterior surface of portion and three large setae in a row on lateral edge of posterior surface of terminal tab. Distal portion of telopodite a large, flattened structure arising on posterior surface of basal portion of telopodite just below terminal tab; curving basally and slightly laterally; divided near base into narrow medial branch, flattened apically with minute, spine-like protrusions on posterior and medial surfaces, and much larger lateral branch, the latter much expanded, curving anterobasally and divided by two deep notches into three broad, rounded lobes (
|
||
<figureCitation box="[395,483,202,229]" captionStart="Figure 5" captionStartId="9.[140,212,1558,1580]" captionTargetBox="[140,1108,165,1538]" captionTargetId="figure@9.[140,1109,165,1539]" captionTargetPageId="9" captionText="Figure 5. Agathodesmus steeli Silvestri, 1910, ex AM KS107964. A Male midbody sternite (anterior towards top). B Anterior view of extended left gonopod. Agathodesmus johnsi sp. n., ex AM KS94156. C Left ventrolateral view of male midbody ring (anterior towards left); no spiracles are evident above leg bases. Scale bars: (A) = 0.5 mm; (B), (C) = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767250" httpUri="https://zenodo.org/record/3767250/files/figure.png" pageId="10" pageNumber="97">Figs 5B</figureCitation>
|
||
,
|
||
<figureCitation box="[496,533,202,228]" captionStart="Figure 7" captionStartId="12.[140,212,1583,1605]" captionTargetBox="[140,1108,165,1563]" captionTargetId="figure@12.[140,1108,165,1564]" captionTargetPageId="12" captionText="Figure 7. Ventral views of retracted male gonopods. A Agathodesmus steeli Silvestri, 1910, ex AM KS107964. B Agathodesmus johnsi sp. n., ex AM KS94156. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767254" httpUri="https://zenodo.org/record/3767254/files/figure.png" pageId="10" pageNumber="97">7A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767250" ID-Zenodo-Dep="3767250" httpUri="https://zenodo.org/record/3767250/files/figure.png" pageId="9" pageNumber="96" startId="9.[140,212,1558,1580]" targetBox="[140,1108,165,1538]" targetPageId="9">
|
||
<paragraph blockId="9.[140,1109,1558,1677]" pageId="9" pageNumber="96">
|
||
<emphasis bold="true" box="[140,241,1558,1580]" pageId="9" pageNumber="96">Figure 5.</emphasis>
|
||
<taxonomicName authority="Silvestri, 1910" authorityName="Silvestri" authorityYear="1910" box="[250,571,1558,1580]" class="Diplopoda" family="Haplodesmidae" genus="Agathodesmus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="96" phylum="Arthropoda" rank="species" species="steeli">
|
||
<emphasis box="[250,424,1558,1579]" italics="true" pageId="9" pageNumber="96">Agathodesmus steeli</emphasis>
|
||
<bibRefCitation author="Silvestri" box="[434,571,1558,1580]" firstAuthor="Silvestri" pageId="9" pageNumber="96" pagination="357 - 364" refId="ref8865" refString="Silvestri F (1910) Descrizioni preliminari di nuovi generi di Diplopodi. I. Polydesmoidea. Zoologischer Anzeiger 35: 357 - 364." type="journal article" year="1910">Silvestri, 1910</bibRefCitation>
|
||
</taxonomicName>
|
||
, ex AM KS107964.
|
||
<emphasis bold="true" box="[774,793,1558,1580]" pageId="9" pageNumber="96">A</emphasis>
|
||
Male midbody sternite (anterior towards top).
|
||
<emphasis bold="true" box="[272,289,1590,1612]" pageId="9" pageNumber="96">B</emphasis>
|
||
Anterior view of extended left gonopod.
|
||
<taxonomicName authority="Mesibov, 2009" authorityName="Mesibov" authorityYear="2009" box="[684,867,1590,1611]" class="Diplopoda" family="Haplodesmidae" genus="Agathodesmus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="96" phylum="Arthropoda" rank="species" species="johnsi" status="sp. n.">
|
||
<emphasis box="[684,867,1590,1611]" italics="true" pageId="9" pageNumber="96">Agathodesmus johnsi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[875,929,1591,1612]" pageId="9" pageNumber="96" rank="species">sp. n.</taxonomicNameLabel>
|
||
, ex AM KS94156.
|
||
<emphasis bold="true" box="[140,159,1622,1644]" pageId="9" pageNumber="96">C</emphasis>
|
||
Left ventrolateral view of male midbody ring (anterior towards left); no spiracles are evident above leg bases. Scale bars: (A) = 0.5 mm; (B), (C) = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="10.[140,1108,167,369]" pageId="10" pageNumber="97">
|
||
Female longer and more robust than male (
|
||
<figureCitation box="[699,790,237,263]" captionStart="Figure 4" captionStartId="7.[140,212,1179,1201]" captionTargetBox="[140,1108,165,1159]" captionTargetId="figure@7.[140,1109,165,1159]" captionTargetPageId="7" captionText="Figure 4. Agathodesmus steeli Silvestri, 1910, ex AM KS107964. A Female (left) and male (right); red colour is imaging artifact. B Male, left lateral view of posterior end; o = ozopore area. C Male, ventral view of telson. D Enlargement of C showing spinnerets. Scale bars: (A) = 2.5 mm, (B) = 0.25 mm, (C) = 0.1 mm, (D) = 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.3767248" httpUri="https://zenodo.org/record/3767248/files/figure.png" pageId="10" pageNumber="97">Fig. 4A</figureCitation>
|
||
); epigynum inconspicuous, posterior margin barely raised; cyphopods not examined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="11" lastPageNumber="98" pageId="10" pageNumber="97" type="distribution">
|
||
<paragraph blockId="10.[140,1108,167,369]" lastBlockId="11.[140,1108,167,651]" lastPageId="11" lastPageNumber="98" pageId="10" pageNumber="97">
|
||
<emphasis bold="true" box="[185,490,307,333]" pageId="10" pageNumber="97">
|
||
<emphasis box="[185,336,307,333]" inLineHeading="true" pageId="10" pageNumber="97" reason="1">Distribution</emphasis>
|
||
and habitat.
|
||
</emphasis>
|
||
Known so far from eucalypt forest (historically in the case of Avoca) at two localities ca.
|
||
<quantity box="[554,632,343,369]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="97" unit="km" value="20.0">20 km</quantity>
|
||
apart in southeastern
|
||
<collectingRegion box="[899,1108,343,369]" country="Australia" name="New South Wales" pageId="10" pageNumber="97">New South Wales</collectingRegion>
|
||
(
|
||
<figureCitation box="[147,291,167,193]" captionStart="Figure 2" captionStartId="2.[140,212,1526,1548]" captionTargetBox="[140,1108,306,1507]" captionTargetId="figure@2.[141,1107,306,1505]" captionTargetPageId="2" captionText="Figure 2. Agathodesmus localities. A Australia with rectangle indicating extent of map B; arrow points to type locality of A. bucculentus. B New South Wales localities (squares). 1 = Mt Aggie, A. johnsi sp. n.; 2 = Avoca, 3 = Knights Hill, A. steeli Silvestri, 1910; rectangle around 2 and 3 indicates extent of map C. C Southern highlands places (open circles), approximate sites for ANIC berlesates (triangles), 2009 search sites on sandstone and shale (crosses) and basalt (squares). Mercator projections." figureDoi="http://doi.org/10.5281/zenodo.3767244" httpUri="https://zenodo.org/record/3767244/files/figure.png" pageId="11" pageNumber="98">Figs 2B, 2C</figureCitation>
|
||
). Both sites are above
|
||
<quantity box="[554,630,167,193]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="98" unit="m" value="700.0">700 m</quantity>
|
||
with annual rainfall probably>
|
||
<quantity box="[1001,1103,167,193]" metricMagnitude="-1" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="98" unit="mm" value="900.0">900 mm</quantity>
|
||
, in a temperate climate with cool winters. At the Knights Hill site, the 15
|
||
<taxonomicName authorityName="Silvestri" authorityYear="1910" box="[1020,1108,202,228]" class="Diplopoda" family="Haplodesmidae" genus="Agathodesmus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="98" phylum="Arthropoda" rank="species" species="steeli">
|
||
<emphasis box="[1020,1108,202,228]" italics="true" pageId="11" pageNumber="98">A. steeli</emphasis>
|
||
</taxonomicName>
|
||
specimens were found in narrow spaces in part of a large, moist, well-rotted log, either a
|
||
<taxonomicName box="[160,275,272,298]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="98" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[160,275,272,298]" italics="true" pageId="11" pageNumber="98">Eucalyptus</emphasis>
|
||
</taxonomicName>
|
||
species or
|
||
<taxonomicName box="[400,614,272,299]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="11" pageNumber="98" phylum="Tracheophyta" rank="species" species="melanoxylon">
|
||
<emphasis box="[400,614,272,299]" italics="true" pageId="11" pageNumber="98">Acacia melanoxylon</emphasis>
|
||
</taxonomicName>
|
||
. Also in that part of the log were Siphonophorida, Symphyla,
|
||
<taxonomicName box="[369,463,308,334]" class="Chilopoda" family="Cryptopidae" genus="Cryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="11" pageNumber="98" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[369,463,308,334]" italics="true" pageId="11" pageNumber="98">Cryptops</emphasis>
|
||
</taxonomicName>
|
||
sp. centipedes, fly and beetle larvae and terrestrial isopod crustaceans.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3767252" ID-Zenodo-Dep="3767252" httpUri="https://zenodo.org/record/3767252/files/figure.png" pageId="10" pageNumber="97" startId="10.[140,212,1558,1580]" targetBox="[140,1108,423,1538]" targetPageId="10">
|
||
<paragraph blockId="10.[140,1108,1558,1677]" pageId="10" pageNumber="97">
|
||
<emphasis bold="true" box="[140,244,1558,1580]" pageId="10" pageNumber="97">Figure 6.</emphasis>
|
||
<taxonomicName authority="Silvestri, 1910" authorityName="Silvestri" authorityYear="1910" box="[255,584,1558,1580]" class="Diplopoda" family="Haplodesmidae" genus="Agathodesmus" kingdom="Animalia" order="Polydesmida" pageId="10" pageNumber="97" phylum="Arthropoda" rank="species" species="steeli">
|
||
<emphasis box="[255,432,1558,1579]" italics="true" pageId="10" pageNumber="97">Agathodesmus steeli</emphasis>
|
||
<bibRefCitation author="Silvestri" box="[444,584,1558,1580]" firstAuthor="Silvestri" pageId="10" pageNumber="97" pagination="357 - 364" refId="ref8865" refString="Silvestri F (1910) Descrizioni preliminari di nuovi generi di Diplopodi. I. Polydesmoidea. Zoologischer Anzeiger 35: 357 - 364." type="journal article" year="1910">Silvestri, 1910</bibRefCitation>
|
||
</taxonomicName>
|
||
, female ex AM KS107964.
|
||
<emphasis bold="true" box="[868,887,1558,1580]" pageId="10" pageNumber="97">A</emphasis>
|
||
Tubercle on midbody metatergite.
|
||
<emphasis bold="true" box="[256,273,1590,1612]" pageId="10" pageNumber="97">B</emphasis>
|
||
Bisegmented seta adjoining tubercle on midbody metatergite.
|
||
<emphasis bold="true" box="[857,876,1590,1612]" pageId="10" pageNumber="97">C</emphasis>
|
||
Dorsal view of midbody prozonite; limbus on next anterior ring at top, edge of metazonite at bottom; pl = primary limbus element, sl = secondary limbus element. Scale bars: (A) = 0.025 mm, (B) = 0.02 mm, (C) = 0.05 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="11" pageNumber="98" type="discussion">
|
||
<paragraph blockId="11.[140,1108,167,651]" pageId="11" pageNumber="98">
|
||
<emphasis bold="true" box="[185,298,378,404]" pageId="11" pageNumber="98">
|
||
<emphasis box="[185,293,378,404]" inLineHeading="true" pageId="11" pageNumber="98" reason="1">Remarks</emphasis>
|
||
.
|
||
</emphasis>
|
||
Live
|
||
<taxonomicName authorityName="Silvestri" authorityYear="1910" box="[362,447,378,404]" class="Diplopoda" family="Haplodesmidae" genus="Agathodesmus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="98" phylum="Arthropoda" rank="species" species="steeli">
|
||
<emphasis box="[362,447,378,404]" italics="true" pageId="11" pageNumber="98">A. steeli</emphasis>
|
||
</taxonomicName>
|
||
are very slow-moving and do not curl up, even when disturbed. Unlike adults of the morphologically, ecologically and behaviourally similar species of
|
||
<taxonomicName authority="Silvestri, 1910" authorityName="Silvestri" authorityYear="1910" box="[255,582,448,474]" class="Diplopoda" family="Dalodesmidae" genus="Asphalidesmus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="98" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[255,411,448,474]" italics="true" pageId="11" pageNumber="98">Asphalidesmus</emphasis>
|
||
<bibRefCitation author="Silvestri" box="[419,582,448,474]" firstAuthor="Silvestri" pageId="11" pageNumber="98" pagination="357 - 364" refId="ref8865" refString="Silvestri F (1910) Descrizioni preliminari di nuovi generi di Diplopodi. I. Polydesmoidea. Zoologischer Anzeiger 35: 357 - 364." type="journal article" year="1910">Silvestri, 1910</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Mesibov" box="[597,765,448,474]" firstAuthor="Mesibov" pageId="11" pageNumber="98" pagination="531 - 540" refId="ref8692" refString="Mesibov R (2002) Redescriptions of Asphalidesmus leae Silvestri, 1910 and A. parvus (Chamberlin, 1920) comb. nov. from Tasmania, Australia (Diplopoda: Polydesmida: Haplodesmidae). Memoirs of Museum Victoria 59 (2): 531 - 540." type="journal article" year="2002">Mesibov 2002</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Mesibov" box="[777,840,448,474]" firstAuthor="Mesibov" pageId="11" pageNumber="98" pagination="55 - 74" refId="ref8771" refString="Mesibov R (2009) A new millipede genus and a new species of Asphalidesmus Silvestri, 1910 (Diplopoda: Polydesmida: Dalodesmidea) from southern Tasmania, Australia. ZooKeys 7: 55 - 74." type="journal article" year="2009">2009</bibRefCitation>
|
||
),
|
||
<taxonomicName authorityName="Silvestri" authorityYear="1910" box="[860,945,448,475]" class="Diplopoda" family="Haplodesmidae" genus="Agathodesmus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="98" phylum="Arthropoda" rank="species" species="steeli">
|
||
<emphasis box="[860,945,448,475]" italics="true" pageId="11" pageNumber="98">A. steeli</emphasis>
|
||
</taxonomicName>
|
||
adults are not heavily encrusted with soil particles.
|
||
</paragraph>
|
||
<paragraph blockId="11.[140,1108,167,651]" pageId="11" pageNumber="98">
|
||
The apparent absence of well-defined spiracles in
|
||
<taxonomicName authorityName="Silvestri" authorityYear="1910" box="[748,833,519,545]" class="Diplopoda" family="Haplodesmidae" genus="Agathodesmus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="98" phylum="Arthropoda" rank="species" species="steeli">
|
||
<emphasis box="[748,833,519,545]" italics="true" pageId="11" pageNumber="98">A. steeli</emphasis>
|
||
</taxonomicName>
|
||
is remarkable. I have so far been unable to detect spiracles either with light microscopy (cleared specimens) or scanning electron microscopy (see also
|
||
<figureCitation box="[622,712,589,616]" captionStart="Figure 5" captionStartId="9.[140,212,1558,1580]" captionTargetBox="[140,1108,165,1538]" captionTargetId="figure@9.[140,1109,165,1539]" captionTargetPageId="9" captionText="Figure 5. Agathodesmus steeli Silvestri, 1910, ex AM KS107964. A Male midbody sternite (anterior towards top). B Anterior view of extended left gonopod. Agathodesmus johnsi sp. n., ex AM KS94156. C Left ventrolateral view of male midbody ring (anterior towards left); no spiracles are evident above leg bases. Scale bars: (A) = 0.5 mm; (B), (C) = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767250" httpUri="https://zenodo.org/record/3767250/files/figure.png" pageId="11" pageNumber="98">Fig. 5C</figureCitation>
|
||
). A histological study is needed to determine whether the tracheal system is also modified from the norm in
|
||
<taxonomicName authorityName="Pocock" authorityYear="1887" box="[958,1104,624,650]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="98" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |