265 lines
32 KiB
XML
265 lines
32 KiB
XML
<document id="8B63C3D7D0AEC609A25C2186594BD795" ID-CLB-Dataset="22413" ID-DOI="10.11646/zootaxa.4768.3.2" ID-GBIF-Dataset="43c18a9c-5545-4f3d-b285-956094e43d43" ID-ISSN="1175-5326" ID-Zenodo-Dep="3784036" ID-ZooBank="A005B93A-2070-46F5-A06E-6F6268A207EB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588576355177" checkinUser="plazi" docAuthor="Hu, Fang-Shuo" docDate="2020" docId="997687A2FFC6FF91FACC30CD509FA052" docLanguage="en" docName="zootaxa.4768.3.2.pdf" docOrigin="Zootaxa 4768 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ontholestes oculatus" docType="treatment" docVersion="6" lastPageNumber="346" masterDocId="654FFFDAFFCCFF9DFA5B34445127A409" masterDocTitle="New distributional records of Staphylinina in Taiwan, including a new species of Miobdelus Sharp (Coleoptera: Staphylinidae: Staphylininae: Staphylinini)" masterLastPageNumber="360" masterPageNumber="334" pageNumber="344" updateTime="1698838343837" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="437060786EDEECF178F742E64AF4F9B7">New distributional records of Staphylinina in Taiwan, including a new species of Miobdelus Sharp (Coleoptera: Staphylinidae: Staphylininae: Staphylinini)</mods:title>
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<mods:name id="801A2670D588CAB98591B4293E4FF5A4" type="personal">
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<mods:roleTerm id="A8B7AAC856FD23289B33EDFF5D95E6AC">Author</mods:roleTerm>
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<mods:namePart id="9A077D720CE1393779D69B6E4DB741A9">Hu, Fang-Shuo</mods:namePart>
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<mods:title id="65D0F9FD2AB95773F3DC07884FFFB920">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="054B6BFC2F93393BC67FCDE28006EEB5">
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<mods:date id="EA06210BA81DD61AB44619B5A76D24A3">2020</mods:date>
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<mods:number id="467F74B5DDF8AF43743D0473639F24D2">2020-05-04</mods:number>
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<mods:number id="A69900078EB1E18C809662E49D3CB084">4768</mods:number>
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<mods:number id="5F5DA5EEA65BEA1E24B7EDA882605008">3</mods:number>
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<mods:classification id="88E9DA45C9F0ED313CC44E9142D082A3">journal article</mods:classification>
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<mods:identifier id="049574A2FA25BA3511E0652A3E6629B7" type="DOI">10.11646/zootaxa.4768.3.2</mods:identifier>
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<treatment id="997687A2FFC6FF91FACC30CD509FA052" ID-DOI="http://doi.org/10.5281/zenodo.3794832" ID-GBIF-Taxon="163885981" ID-Zenodo-Dep="3794832" LSID="urn:lsid:plazi:treatment:997687A2FFC6FF91FACC30CD509FA052" httpUri="http://treatment.plazi.org/id/997687A2FFC6FF91FACC30CD509FA052" lastPageId="12" lastPageNumber="346" pageId="10" pageNumber="344">
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<subSubSection id="59C5653FFFC6FF97FACC30CD5363A0AA" box="[151,580,1161,1188]" pageId="10" pageNumber="344" type="nomenclature">
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<paragraph id="116036B4FFC6FF97FACC30CD5363A0AA" blockId="10.[151,580,1161,1224]" box="[151,580,1161,1188]" pageId="10" pageNumber="344">
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<heading id="4A2881D8FFC6FF97FACC30CD5363A0AA" bold="true" box="[151,580,1161,1188]" fontSize="11" level="1" pageId="10" pageNumber="344" reason="1">
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<taxonomicName id="D6DF4D37FFC6FF97FACC30CD5363A0AA" ID-CoL="49MB4" authority="(Sharp, 1874)" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[151,580,1161,1188]" class="Insecta" family="Staphylinidae" genus="Ontholestes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="344" phylum="Arthropoda" rank="species" species="oculatus">
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||
<emphasis id="23ABEAA6FFC6FF97FACC30CD5363A0AA" bold="true" box="[151,580,1161,1188]" pageId="10" pageNumber="344">
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<emphasis id="23ABEAA6FFC6FF97FACC30CD50B4A0AD" bold="true" box="[151,403,1161,1188]" italics="true" pageId="10" pageNumber="344">Ontholestes oculatus</emphasis>
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(Sharp, 1874)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="59C5653FFFC6FF97FACC30E950A5A0C1" box="[151,386,1197,1224]" pageId="10" pageNumber="344" type="description">
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<paragraph id="116036B4FFC6FF97FACC30E950A5A0C1" blockId="10.[151,580,1161,1224]" box="[151,386,1197,1224]" pageId="10" pageNumber="344">
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||
(
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<figureCitation id="89E42A31FFC6FF97FAC430E95000A0C1" box="[159,295,1197,1224]" captionStart="FIGURES 14–19" captionStartId="16.[151,264,1911,1936]" captionTargetBox="[161,1430,370,1883]" captionTargetId="figure@16.[151,1435,345,1886]" captionTargetPageId="16" captionText="FIGURES 14–19. 14–15. Ontholestes oculatus: male and female. 16. Platydracus juang: male. 17. Thoracostrongylus formosanus: male. 18–19. Thoracostrongylus miyakei: male, lectotype and attached labels; male. Scale bar: 3 mm" figureDoi="http://doi.org/10.5281/zenodo.3784042" httpUri="https://zenodo.org/record/3784042/files/figure.png" pageId="10" pageNumber="344">Figs. 14–15</figureCitation>
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,
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<figureCitation id="89E42A31FFC6FF97FB6830E9505DA0C1" box="[307,378,1197,1224]" captionStart="FIGURES 35–40" captionStartId="20.[151,264,1962,1987]" captionTargetBox="[196,1392,181,1937]" captionTargetId="figure@20.[196,1392,181,1937]" captionTargetPageId="20" captionText="FIGURES 35–40. Ontholestes oculatus 35. Aedeagus in ventral view; 36. Aedeagus in lateral view; 37. Sternite 9, male; 38. Sternite 8, male; 39. Tergite 10, male; 40. Tergite 10, female. Scale bar: 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.3784050" httpUri="https://zenodo.org/record/3784050/files/figure.png" pageId="10" pageNumber="344">35–40</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="59C5653FFFC6FF97FACC30B15209A175" pageId="10" pageNumber="344" type="materials_examined">
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<paragraph id="116036B4FFC6FF97FACC30B15209A175" blockId="10.[151,1437,1268,2016]" pageId="10" pageNumber="344">
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<emphasis id="23ABEAA6FFC6FF97FACC30B150A5A119" bold="true" box="[151,386,1269,1296]" pageId="10" pageNumber="344">Material examined.</emphasis>
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<materialsCitation id="A1B73CE9FFC6FF97FBD030B1532BA13A" ID-GBIF-Occurrence="2609799059" collectingDate="2018-04-09" collectorName="Jinsha To. & B. H. Ho" country="Taiwan" location="Tianpu Reservoir" pageId="10" pageNumber="344" specimenCount="1" specimenCount-male="1" stateProvince="Kinmen">
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<emphasis id="23ABEAA6FFC6FF97FBD030B150DBA119" bold="true" box="[395,508,1269,1296]" pageId="10" pageNumber="344">
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<collectingCountry id="69C87624FFC6FF97FBD030B150DBA119" box="[395,508,1269,1296]" name="Taiwan" pageId="10" pageNumber="344">TAIWAN</collectingCountry>
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</emphasis>
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:
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<collectingRegion id="D31BF856FFC6FF97F85730B153E1A106" box="[524,710,1269,1295]" country="Taiwan" name="Kinmen" pageId="10" pageNumber="344">Kinmen County</collectingRegion>
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:
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<specimenCount id="07D9FD3DFFC6FF97F88930B15205A106" box="[722,802,1269,1295]" pageId="10" pageNumber="344" type="male">1 male</specimenCount>
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,
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<location id="1400606FFFC6FF97F97530B152D5A106" LSID="urn:lsid:plazi:treatment:997687A2FFC6FF91FACC30CD509FA052:1400606FFFC6FF97F97530B152D5A106" box="[814,1010,1269,1295]" country="Taiwan" name="Tianpu Reservoir" pageId="10" pageNumber="344" stateProvince="Kinmen">Tianpu Reservoir</location>
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(
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<emphasis id="23ABEAA6FFC6FF97FE5F30B055A5A102" bold="true" box="[1028,1154,1268,1291]" pageId="10" pageNumber="344">aảửw)</emphasis>
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,
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<collectorName id="BC2A5362FFC6FF97FEC930B15420A119" box="[1170,1287,1269,1296]" pageId="10" pageNumber="344">Jinsha To.</collectorName>
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,
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<date id="65611074FFC6FF97FF4F30B154BFA106" box="[1300,1432,1269,1295]" pageId="10" pageNumber="344" value="2018-04-09">
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<collectingDate id="7525E99CFFC6FF97FF4F30B154BFA106" box="[1300,1432,1269,1295]" pageId="10" pageNumber="344" value="2018-04-09">09-IV-2018</collectingDate>
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</date>
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, leg.
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<collectorName id="BC2A5362FFC6FF97FA9C315E500DA13D" box="[199,298,1306,1332]" pageId="10" pageNumber="344">B. H. Ho</collectorName>
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(cow dung) (FSHc)
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</materialsCitation>
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;
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<materialsCitation id="A1B73CE9FFC6FF97F84C315D54BFA13A" ID-GBIF-Occurrence="2609799084" box="[535,1432,1305,1332]" collectingDate="2019-04-01" collectorName="Y. H. Ho" country="Taiwan" location="Tianpu Reservoir" pageId="10" pageNumber="344" specimenCount="2" specimenCount-female="2" stateProvince="Kinmen County">
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<specimenCount id="07D9FD3DFFC6FF97F84C315D53A2A13A" box="[535,645,1305,1331]" pageId="10" pageNumber="344" type="female">2 females</specimenCount>
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, same locality as above,
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<date id="65611074FFC6FF97F9CC315D553BA13A" box="[919,1052,1305,1331]" pageId="10" pageNumber="344" value="2019-04-01">
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||
<collectingDate id="7525E99CFFC6FF97F9CC315D553BA13A" box="[919,1052,1305,1331]" pageId="10" pageNumber="344" value="2019-04-01">01-IV-2019</collectingDate>
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</date>
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, leg.
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<collectorName id="BC2A5362FFC6FF97FE0F315E5591A13D" box="[1108,1206,1306,1332]" pageId="10" pageNumber="344">Y. H. Ho</collectorName>
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(cow dung) (FSHc)
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</materialsCitation>
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;
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<materialsCitation id="A1B73CE9FFC6FF97FACC31795229A15E" ID-GBIF-Occurrence="2609799062" box="[151,782,1341,1368]" collectingDate="2019-04-03" collectorName="Y. H. Ho" country="Taiwan" location="Tianpu Reservoir" pageId="10" pageNumber="344" specimenCount="1" specimenCount-female="1" stateProvince="Kinmen County">
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||
<specimenCount id="07D9FD3DFFC6FF97FACC317951DCA15E" box="[151,251,1341,1367]" pageId="10" pageNumber="344" type="female">1 female</specimenCount>
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, same data as above except
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<date id="65611074FFC6FF97F86B31795395A15E" box="[560,690,1341,1367]" pageId="10" pageNumber="344" value="2019-04-03">
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||
<collectingDate id="7525E99CFFC6FF97F86B31795395A15E" box="[560,690,1341,1367]" pageId="10" pageNumber="344" value="2019-04-03">03-IV-2019</collectingDate>
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</date>
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(FSHc)
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||
</materialsCitation>
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;
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<materialsCitation id="A1B73CE9FFC6FF97F9423179520DA175" ID-GBIF-Occurrence="2609799130" collectingDate="2019-04-30" collectionCode="NMNS, TARI" collectorName="Lieyu To. & C. H. Ma" country="Taiwan" latitude="24.4469" location="Lieyu" longLatPrecision="7" longitude="118.258" pageId="10" pageNumber="344" specimenCount="3" specimenCount-male="3" stateProvince="Kinmen County">
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<specimenCount id="07D9FD3DFFC6FF97F94231795256A15E" box="[793,881,1341,1367]" pageId="10" pageNumber="344" type="male">3 males</specimenCount>
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,
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<location id="1400606FFFC6FF97F9273179529AA15E" LSID="urn:lsid:plazi:treatment:997687A2FFC6FF91FACC30CD509FA052:1400606FFFC6FF97F9273179529AA15E" box="[892,957,1341,1367]" country="Taiwan" latitude="24.4469" longLatPrecision="7" longitude="118.258" name="Lieyu" pageId="10" pageNumber="344" stateProvince="Kinmen County">Lieyu</location>
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(
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<emphasis id="23ABEAA6FFC6FF97F99631785534A15A" bold="true" box="[973,1043,1340,1363]" pageId="10" pageNumber="344">ȗë)</emphasis>
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,
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<collectorName id="BC2A5362FFC6FF97FE7B317955A8A151" box="[1056,1167,1341,1368]" pageId="10" pageNumber="344">Lieyu To.</collectorName>
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,
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<geoCoordinate id="74EB5073FFC6FF97FEC2317955D0A15E" box="[1177,1271,1341,1367]" degrees="24.4469" direction="north" orientation="latitude" pageId="10" pageNumber="344" precision="5" value="24.4469">24.4469</geoCoordinate>
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,
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<geoCoordinate id="74EB5073FFC6FF97FF593179544AA151" box="[1282,1389,1341,1368]" degrees="118.2580" direction="east" orientation="longitude" pageId="10" pageNumber="344" precision="5" value="118.258">118.2580</geoCoordinate>
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,
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<date id="65611074FFC6FF97FF2C317951D1A172" pageId="10" pageNumber="344" value="2019-04-30">
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<collectingDate id="7525E99CFFC6FF97FF2C317951D1A172" pageId="10" pageNumber="344" value="2019-04-30">30- IV-2019</collectingDate>
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</date>
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, leg.
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<collectorName id="BC2A5362FFC6FF97FB69312550BDA175" box="[306,410,1377,1404]" pageId="10" pageNumber="344">C. H. Ma</collectorName>
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(cow dung) (FSHc,
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<collectionCode id="77CEAE71FFC6FF97F824312553F4A172" box="[639,723,1377,1403]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="10" pageNumber="344">NMNS</collectionCode>
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,
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<collectionCode id="77CEAE71FFC6FF97F88531265205A175" box="[734,802,1378,1404]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="10" pageNumber="344">TARI</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="59C5653FFFC6FF91FA9C31C250ABA71E" lastPageId="12" lastPageNumber="346" pageId="10" pageNumber="344" type="description">
|
||
<paragraph id="116036B4FFC6FF97FA9C31C252ECA1CA" blockId="10.[151,1437,1268,2016]" pageId="10" pageNumber="344">
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<emphasis id="23ABEAA6FFC6FF97FA9C31C25059A1A9" bold="true" box="[199,382,1414,1440]" pageId="10" pageNumber="344">Measurements.</emphasis>
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BL: 11.86 (11.31–12.75); HL: 1.75 (1.56–1.83); HW: 2.85 (2.52–3.15); EyL: 1.29 (1.12–1.45); TL: 0.28 (0.25–0.30); PL: 2.35 (2.05–2.50); ElL: 2.98 (2.74–3.15) (mm).
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</paragraph>
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<paragraph id="116036B4FFC6FF97FA9C3189524FA30E" blockId="10.[151,1437,1268,2016]" pageId="10" pageNumber="344">
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<emphasis id="23ABEAA6FFC6FF97FA9C31895059A1E1" bold="true" box="[199,382,1485,1512]" pageId="10" pageNumber="344">Re-description.</emphasis>
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Medium sized species. Habitus (
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||
<figureCitation id="89E42A31FFC6FF97F8B131895257A1E1" box="[746,880,1485,1512]" captionStart="FIGURES 14–19" captionStartId="16.[151,264,1911,1936]" captionTargetBox="[161,1430,370,1883]" captionTargetId="figure@16.[151,1435,345,1886]" captionTargetPageId="16" captionText="FIGURES 14–19. 14–15. Ontholestes oculatus: male and female. 16. Platydracus juang: male. 17. Thoracostrongylus formosanus: male. 18–19. Thoracostrongylus miyakei: male, lectotype and attached labels; male. Scale bar: 3 mm" figureDoi="http://doi.org/10.5281/zenodo.3784042" httpUri="https://zenodo.org/record/3784042/files/figure.png" pageId="10" pageNumber="344">Figs. 14–15</figureCitation>
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) of both sexes with black head, pronotum and elytra covered with brownish yellow and silver setae; scutellum black, with brownish yellow setae on anterior margin; meso- and metathorax covered with gray setae on margin in ventral view; tergites 3–5 covered with brownish yellow setae, but without pair of dark tomentose spots at middle, some individuals also with mixed few silver setae on tergites; anterior margin of tergites 6–8 and each sternite covered with gray setae on anterior part, rather distinct on margin. Colouration with distinct sexual dimorphism. Antennal segments 1–4 brownish yellow in male but darker in female, antennal segments 5–11 dark brown in both sexes; in male, anterior part of labrum brownish yellow, basal part of labrum black, in female entire labrum black; legs brownish yellow, with black spots on femur in male, legs dark brown and femur with slightly larger black spots in female.
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</paragraph>
|
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<paragraph id="116036B4FFC6FF97FA9C335553FFA3D6" blockId="10.[151,1437,1268,2016]" pageId="10" pageNumber="344">Head obtusely quadrangular, not dilated posteriad, with rounded posterior angles, markedly wider than long (ratio: 1.63), eyes very large but convex, tempora markedly shorter than length of eyes in dorsal view (ratio: 0.22); dorsal surface of head densely covered with fine punctation.Antennae moderately long, segment 1 (scape) markedly longer than other segments; segment 2 (pedicel) shortest; segment 3 slightly shorter than segment 1, but markedly longer than other segments; segments 4–10 wider than long, gradually but not obviously becoming shorter; last segment longer than wide, asymmetrically emarginate.</paragraph>
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<caption id="45A0663CFFC7FF96FACC33025237A3AF" ID-DOI="http://doi.org/10.5281/zenodo.3784040" ID-Zenodo-Dep="3784040" httpUri="https://zenodo.org/record/3784040/files/figure.png" pageId="11" pageNumber="345" startId="11.[151,264,1862,1887]" targetBox="[151,1412,185,1837]" targetPageId="11">
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||
<paragraph id="116036B4FFC7FF96FACC33025237A3AF" blockId="11.[151,1436,1862,1959]" pageId="11" pageNumber="345">
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||
<emphasis id="23ABEAA6FFC7FF96FACC33025060A356" bold="true" box="[151,327,1862,1887]" pageId="11" pageNumber="345">FIGURES 8–13.</emphasis>
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8.
|
||
<taxonomicName id="D6DF4D37FFC7FF96FB3E33035307A356" authority="Hu, 2020" authorityName="Hu" authorityYear="2020" box="[357,544,1863,1887]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="345" phylum="Arthropoda" rank="species" species="humilis" status="sp. nov.">
|
||
<emphasis id="23ABEAA6FFC7FF96FB3E33035307A356" box="[357,544,1863,1887]" italics="true" pageId="11" pageNumber="345">Miobdelus humilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="23ABEAA6FFC7FF96F87E33035352A356" bold="true" box="[549,629,1863,1887]" pageId="11" pageNumber="345">
|
||
<taxonomicNameLabel id="389857DDFFC7FF96F87E33035352A356" box="[549,629,1863,1887]" pageId="11" pageNumber="345" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: male, paratype. 9.
|
||
<taxonomicName id="D6DF4D37FFC7FF96F96033035586A356" authority="taiwanensis" authorityName="taiwanensis Smetana" authorityYear="2001" box="[827,1185,1863,1887]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="345" phylum="Arthropoda" rank="subSpecies" species="taiwanensis" subSpecies="taiwanensis">
|
||
<emphasis id="23ABEAA6FFC7FF96F96033035586A356" box="[827,1185,1863,1887]" italics="true" pageId="11" pageNumber="345">Miobdelus taiwanensis taiwanensis</emphasis>
|
||
</taxonomicName>
|
||
: male. 10.
|
||
<taxonomicName id="D6DF4D37FFC7FF96FF54330351C3A38A" class="Insecta" family="Lucanidae" genus="Nelmanwasus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="345" phylum="Arthropoda" rank="species" species="ornatus">
|
||
<emphasis id="23ABEAA6FFC7FF96FF54330351C3A38A" italics="true" pageId="11" pageNumber="345">Nelmanwasus ornatus</emphasis>
|
||
</taxonomicName>
|
||
: male. 11–12.
|
||
<taxonomicName id="D6DF4D37FFC7FF96FBD8332F5345A38A" authorityName="Hu" authorityYear="2020" baseAuthorityName="Bernhauer" baseAuthorityYear="1943" box="[387,610,1899,1923]" class="Insecta" family="Staphylinidae" genus="Sphaerobulbus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="345" phylum="Arthropoda" rank="species" species="miwai" status="comb. nov.">
|
||
<emphasis id="23ABEAA6FFC7FF96FBD8332F5345A38A" box="[387,610,1899,1923]" italics="true" pageId="11" pageNumber="345">Sphaerobulbus miwai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="23ABEAA6FFC7FF96F830332F53C6A38A" bold="true" box="[619,737,1899,1923]" pageId="11" pageNumber="345">
|
||
<taxonomicNameLabel id="389857DDFFC7FF96F830332F53C6A38A" box="[619,737,1899,1923]" pageId="11" pageNumber="345" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: male, lectotype of
|
||
<taxonomicName id="D6DF4D37FFC7FF96F9EC332F5436A38A" box="[951,1297,1899,1923]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="345" phylum="Arthropoda" rank="species" species="miwai" subGenus="Xanthocypus">
|
||
<emphasis id="23ABEAA6FFC7FF96F9EC332F550AA38A" box="[951,1069,1899,1923]" italics="true" pageId="11" pageNumber="345">Staphylinus</emphasis>
|
||
(
|
||
<emphasis id="23ABEAA6FFC7FF96FE64332F55E5A38A" box="[1087,1218,1899,1923]" italics="true" pageId="11" pageNumber="345">Xanthocypus</emphasis>
|
||
)
|
||
<emphasis id="23ABEAA6FFC7FF96FE8833285436A38A" box="[1235,1297,1900,1923]" italics="true" pageId="11" pageNumber="345">miwai</emphasis>
|
||
</taxonomicName>
|
||
and attached labels; male. 13.
|
||
<taxonomicName id="D6DF4D37FFC7FF96FB1F33CB532BA3AE" authorityName="Shibata" authorityYear="1979" box="[324,524,1935,1959]" class="Insecta" family="Staphylinidae" genus="Naddia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="345" phylum="Arthropoda" rank="species" species="taiwanensis">
|
||
<emphasis id="23ABEAA6FFC7FF96FB1F33CB532BA3AE" box="[324,524,1935,1959]" italics="true" pageId="11" pageNumber="345">Naddia taiwanensis</emphasis>
|
||
</taxonomicName>
|
||
: female. Scale bar: 3 mm
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="116036B4FFC0FF91FA9C34DD50B4A582" blockId="12.[151,1437,153,1115]" pageId="12" pageNumber="346">Thorax. Pronotum about as long as wide, slightly dilated in anterior part; anterior margin with acute angle on both sides; with widely rounded posterior margin; disc of pronotum with narrow impunctate midline, not distinct in anterior part but markedly expanded in front of posterior margin; punctation similar to that of head. Scutellum very large, triangular with dense setose punctures. First four segments of front tarsus markedly dilated, sub-bilobed, covered dense setae ventrally, without distinct difference between sexes. Elytra about as long as wide; slightly longer than pronotum (ratio: 1.27); anterior margin of humeral area markedly dilated; slightly dilated posteriad. Hind wings well developed.</paragraph>
|
||
<paragraph id="116036B4FFC0FF91FA9C35D15438A5DA" blockId="12.[151,1437,153,1115]" pageId="12" pageNumber="346">Abdomen. Tergite 7 (fifth visible) with apical palisade fringe; tergite 2 (anterior to first fully visible tergite) with fine punctation; general punctation of tergites moderately fine and dense, spread evenly on each tergite.</paragraph>
|
||
<paragraph id="116036B4FFC0FF91FA9C359954A5A6C6" blockId="12.[151,1437,153,1115]" pageId="12" pageNumber="346">
|
||
Male. Sternite 8 (
|
||
<figureCitation id="89E42A31FFC0FF91FBD5359950C6A5FE" box="[398,481,477,503]" captionStart="FIGURES 35–40" captionStartId="20.[151,264,1962,1987]" captionTargetBox="[196,1392,181,1937]" captionTargetId="figure@20.[196,1392,181,1937]" captionTargetPageId="20" captionText="FIGURES 35–40. Ontholestes oculatus 35. Aedeagus in ventral view; 36. Aedeagus in lateral view; 37. Sternite 9, male; 38. Sternite 8, male; 39. Tergite 10, male; 40. Tergite 10, female. Scale bar: 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.3784050" httpUri="https://zenodo.org/record/3784050/files/figure.png" pageId="12" pageNumber="346">Fig. 38</figureCitation>
|
||
) with obvious, deep, and obtuse medioapical emargination; sternite 9 (
|
||
<figureCitation id="89E42A31FFC0FF91FF5C3599547DA5FE" box="[1287,1370,477,503]" captionStart="FIGURES 35–40" captionStartId="20.[151,264,1962,1987]" captionTargetBox="[196,1392,181,1937]" captionTargetId="figure@20.[196,1392,181,1937]" captionTargetPageId="20" captionText="FIGURES 35–40. Ontholestes oculatus 35. Aedeagus in ventral view; 36. Aedeagus in lateral view; 37. Sternite 9, male; 38. Sternite 8, male; 39. Tergite 10, male; 40. Tergite 10, female. Scale bar: 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.3784050" httpUri="https://zenodo.org/record/3784050/files/figure.png" pageId="12" pageNumber="346">Fig. 37</figureCitation>
|
||
) with slightly broad basal portion, apical portion with deep and narrow medioapical emargination. Tergite 10 (
|
||
<figureCitation id="89E42A31FFC0FF91FF1A364554B3A612" box="[1345,1428,513,539]" captionStart="FIGURES 35–40" captionStartId="20.[151,264,1962,1987]" captionTargetBox="[196,1392,181,1937]" captionTargetId="figure@20.[196,1392,181,1937]" captionTargetPageId="20" captionText="FIGURES 35–40. Ontholestes oculatus 35. Aedeagus in ventral view; 36. Aedeagus in lateral view; 37. Sternite 9, male; 38. Sternite 8, male; 39. Tergite 10, male; 40. Tergite 10, female. Scale bar: 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.3784050" httpUri="https://zenodo.org/record/3784050/files/figure.png" pageId="12" pageNumber="346">Fig. 39</figureCitation>
|
||
) subtrapezoidal, with a shallow arcuate apical margin; numerous pale setae on posterior margin. Aedeagus (
|
||
<figureCitation id="89E42A31FFC0FF91FF3F366151F9A66A" captionStart="FIGURES 35–40" captionStartId="20.[151,264,1962,1987]" captionTargetBox="[196,1392,181,1937]" captionTargetId="figure@20.[196,1392,181,1937]" captionTargetPageId="20" captionText="FIGURES 35–40. Ontholestes oculatus 35. Aedeagus in ventral view; 36. Aedeagus in lateral view; 37. Sternite 9, male; 38. Sternite 8, male; 39. Tergite 10, male; 40. Tergite 10, female. Scale bar: 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.3784050" httpUri="https://zenodo.org/record/3784050/files/figure.png" pageId="12" pageNumber="346">Figs. 35–36</figureCitation>
|
||
) with both median lobe and paramere symmetrical; median lobe in ventral view sub-parallel for most of its length, slightly constricted at the middle, dilated and obtuse apically in ventral view; in lateral view, apical portion hook-like. Paramere slender, moderately long, apex narrowly rounded, with three to four pairs of long setae apically; base of paramere with two parallel, sclerotized keels, upper one with shallow emargination medioapically.
|
||
</paragraph>
|
||
<paragraph id="116036B4FFC0FF91FA9C369D50ABA71E" blockId="12.[151,1437,153,1115]" pageId="12" pageNumber="346">
|
||
Female genital segment. Tergite 10 (
|
||
<figureCitation id="89E42A31FFC0FF91F82B369D53E2A6FA" box="[624,709,729,755]" captionStart="FIGURES 35–40" captionStartId="20.[151,264,1962,1987]" captionTargetBox="[196,1392,181,1937]" captionTargetId="figure@20.[196,1392,181,1937]" captionTargetPageId="20" captionText="FIGURES 35–40. Ontholestes oculatus 35. Aedeagus in ventral view; 36. Aedeagus in lateral view; 37. Sternite 9, male; 38. Sternite 8, male; 39. Tergite 10, male; 40. Tergite 10, female. Scale bar: 0.5 mm" figureDoi="http://doi.org/10.5281/zenodo.3784050" httpUri="https://zenodo.org/record/3784050/files/figure.png" pageId="12" pageNumber="346">Fig. 40</figureCitation>
|
||
) subtrapezoidal, similar to male tergite 10, apex more broadly rounded than in male.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59C5653FFFC0FF91FA9C37645278A7AE" pageId="12" pageNumber="346" type="diagnosis">
|
||
<paragraph id="116036B4FFC0FF91FA9C37645598A756" blockId="12.[151,1437,153,1115]" pageId="12" pageNumber="346">
|
||
<emphasis id="23ABEAA6FFC0FF91FA9C37645065A732" bold="true" box="[199,322,800,827]" pageId="12" pageNumber="346">Diagnosis.</emphasis>
|
||
<taxonomicName id="D6DF4D37FFC0FF91FB1137655311A732" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[330,566,801,827]" class="Insecta" family="Staphylinidae" genus="Ontholestes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="346" phylum="Arthropoda" rank="species" species="oculatus">
|
||
<emphasis id="23ABEAA6FFC0FF91FB1137655311A732" box="[330,566,801,827]" italics="true" pageId="12" pageNumber="346">Ontholestes oculatus</emphasis>
|
||
</taxonomicName>
|
||
is currently the only species of the genus in
|
||
<collectingCountry id="69C87624FFC0FF91FE67376555B5A732" box="[1084,1170,801,827]" name="Taiwan" pageId="12" pageNumber="346">Taiwan</collectingCountry>
|
||
. It can be easily distinguished from other
|
||
<taxonomicName id="D6DF4D37FFC0FF91FB2837015326A756" authorityName="Latreille" authorityYear="1802" box="[371,513,837,863]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="346" phylum="Arthropoda" rank="subTribe" subTribe="Staphylinina">Staphylinina</taxonomicName>
|
||
by the generic character: pronotum with acute anterior angles.
|
||
</paragraph>
|
||
<paragraph id="116036B4FFC0FF91FA9C372C5278A7AE" blockId="12.[151,1437,153,1115]" pageId="12" pageNumber="346">
|
||
<emphasis id="23ABEAA6FFC0FF91FA9C372C506CA78A" bold="true" box="[199,331,872,899]" pageId="12" pageNumber="346">Bionomics.</emphasis>
|
||
Live specimens of
|
||
<taxonomicName id="D6DF4D37FFC0FF91F87E372D5381A78A" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[549,678,873,899]" class="Insecta" family="Staphylinidae" genus="Ontholestes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="346" phylum="Arthropoda" rank="species" species="oculatus">
|
||
<emphasis id="23ABEAA6FFC0FF91F87E372D531EA78A" box="[549,569,873,899]" italics="true" pageId="12" pageNumber="346">O</emphasis>
|
||
.
|
||
<emphasis id="23ABEAA6FFC0FF91F81C372D5381A78A" box="[583,678,873,899]" italics="true" pageId="12" pageNumber="346">oculatus</emphasis>
|
||
</taxonomicName>
|
||
were observed to be active on the surface of fresh cow dung during the entire day (Cheng-Han Ma personal communication, 2019).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59C5653FFFC0FF91FA9C37F450AAA7F9" pageId="12" pageNumber="346" type="distribution">
|
||
<paragraph id="116036B4FFC0FF91FA9C37F450AAA7F9" blockId="12.[151,1437,153,1115]" pageId="12" pageNumber="346">
|
||
<emphasis id="23ABEAA6FFC0FF91FA9C37F45046A7C2" bold="true" box="[199,353,944,971]" pageId="12" pageNumber="346">Distribution.</emphasis>
|
||
In the lowland regions of
|
||
<collectingCountry id="69C87624FFC0FF91F8CA37F553F3A7C2" box="[657,724,945,971]" name="China" pageId="12" pageNumber="346">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="D31BF856FFC0FF91F8BF37F5526EA7C2" box="[740,841,945,971]" country="China" name="Guangxi" pageId="12" pageNumber="346">Guangxi</collectingRegion>
|
||
,
|
||
<collectingCountry id="69C87624FFC0FF91F90E37F552F9A7C2" box="[853,990,945,971]" name="Hong Kong" pageId="12" pageNumber="346">Hong Kong</collectingCountry>
|
||
),
|
||
<collectingCountry id="69C87624FFC0FF91F9A837F55514A7C2" box="[1011,1075,945,971]" name="Japan" pageId="12" pageNumber="346">Japan</collectingCountry>
|
||
(Kyushu) and
|
||
<collectingCountry id="69C87624FFC0FF91FE8537F55417A7C2" box="[1246,1328,945,971]" name="Taiwan" pageId="12" pageNumber="346">Taiwan</collectingCountry>
|
||
(
|
||
<collectingRegion id="D31BF856FFC0FF91FF1B37F554BBA7C2" box="[1344,1436,945,971]" country="Taiwan" name="Kinmen" pageId="12" pageNumber="346">Kinmen</collectingRegion>
|
||
islands) (new record).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59C5653FFFC0FF91FA9C37BD509FA052" pageId="12" pageNumber="346" type="discussion">
|
||
<paragraph id="116036B4FFC0FF91FA9C37BD509FA052" blockId="12.[151,1437,153,1115]" pageId="12" pageNumber="346">
|
||
<emphasis id="23ABEAA6FFC0FF91FA9C37BD501CA01A" bold="true" box="[199,315,1017,1043]" pageId="12" pageNumber="346">Remarks.</emphasis>
|
||
<bibRefCitation id="754E4B45FFC0FF91FB1A37BD530AA01A" author="Yang, Z. & Zhou, H. - Z." box="[321,557,1017,1043]" pageId="12" pageNumber="346" pagination="1 - 20" refId="ref18582" refString="Yang, Z. & Zhou, H. - Z. (2012) Review of Chinese species of the genus Ontholestes Ganglbauer, 1895 (Coleoptera: Staphylinidae: Staphylininae) with description of two species new to science. Zootaxa, 3191 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 3191.1.1" type="journal article" year="2012">Yang & Zhou (2012)</bibRefCitation>
|
||
provided illustrations of
|
||
<taxonomicName id="D6DF4D37FFC0FF91F91137BD52ECA01A" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[842,971,1017,1043]" class="Insecta" family="Staphylinidae" genus="Ontholestes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="346" phylum="Arthropoda" rank="species" species="oculatus">
|
||
<emphasis id="23ABEAA6FFC0FF91F91137BD5279A01A" box="[842,862,1017,1043]" italics="true" pageId="12" pageNumber="346">O</emphasis>
|
||
.
|
||
<emphasis id="23ABEAA6FFC0FF91F93737BD52ECA01A" box="[876,971,1017,1043]" italics="true" pageId="12" pageNumber="346">oculatus</emphasis>
|
||
</taxonomicName>
|
||
but they ignored the shape of the base of the paramere, which made identification confusing. The illustration of
|
||
<bibRefCitation id="754E4B45FFC0FF91F9E03059558DA03E" author="Yang, Z. & Zhou, H. - Z." box="[955,1194,1053,1079]" pageId="12" pageNumber="346" pagination="1 - 20" refId="ref18582" refString="Yang, Z. & Zhou, H. - Z. (2012) Review of Chinese species of the genus Ontholestes Ganglbauer, 1895 (Coleoptera: Staphylinidae: Staphylininae) with description of two species new to science. Zootaxa, 3191 (1), 1 - 20. https: // doi. org / 10.11646 / zootaxa. 3191.1.1" type="journal article" year="2012">Yang & Zhou (2012)</bibRefCitation>
|
||
should be ignored to prevent misidentification.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |