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<document id="D3804EAA0F9629A9916CD474212788CE" ID-CLB-Dataset="24728" ID-DOI="10.11646/zootaxa.4702.1.8" ID-GBIF-Dataset="f10d83aa-41cc-4a2b-8b37-dfabe65d5fd6" ID-ISSN="1175-5326" ID-Zenodo-Dep="3562843" ID-ZooBank="F5D0A1FE-C0BC-4132-A1EF-C671113B3577" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575479123297" checkinUser="plazi" docAuthor="Ho, Hsuan-Ching" docDate="2019" docId="992D879EE60CFFCCFF7EB1A4FD04FD6B" docLanguage="en" docName="zootaxa.4702.1.8.pdf" docOrigin="Zootaxa 4702 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterophycis spatium Ho 2019, sp. nov." docType="treatment" docUuid="A3E35168-14BF-455C-AA78-AE626E91CC5E" docUuidSource="ZooBank" docVersion="7" lastPageNumber="40" masterDocId="6514FFE6E60DFFCAFFE9B725FFC8FFC0" masterDocTitle="A new genus and species of the cod fish family Moridae (Order Gadiiformes) from southwestern Taiwan" masterLastPageNumber="40" masterPageNumber="32" pageNumber="32" updateTime="1702001654194" updateUser="plazi">
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<mods:title id="6553EC47FC91BEE3D5198DE4E556AF23">A new genus and species of the cod fish family Moridae (Order Gadiiformes) from southwestern Taiwan</mods:title>
</mods:titleInfo>
<mods:name id="CBD36709E05E5EE263B77780753A98A2" type="personal">
<mods:role id="2FEA048A39FF29EAECF97D3955355EC6">
<mods:roleTerm id="FEC33CE7B11ED7FA15C91906D884C191">Author</mods:roleTerm>
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<mods:namePart id="57A06CBF908C0F14BE74EED5A8C38959">Ho, Hsuan-Ching</mods:namePart>
</mods:name>
<mods:typeOfResource id="67D8D9A8A2796D09A4421A17655E838B">text</mods:typeOfResource>
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<mods:title id="BB032B5006775B32F7ECE6CC786630FB">Zootaxa</mods:title>
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<mods:part id="81B916BB14617F6DB4F0A7430C06E5BC">
<mods:date id="18E46CA5FC04AE14985BCFA9A6B3209F">2019</mods:date>
<mods:detail id="AE74DE6A101CA92A3EEC160336338ABA" type="pubDate">
<mods:number id="F1E4CCE20C24947F16F831E6EADACE48">2019-12-04</mods:number>
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<mods:detail id="DEA465FD09D4E618CDA3DD34B5A3F75B" type="volume">
<mods:number id="A8A544E43C0D1C3A491EFFB5FC48CD5C">4702</mods:number>
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<mods:number id="C773FF229CD76A316D8676FA245AA8C1">1</mods:number>
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<mods:classification id="3E6199B20F8FDAF2CA5909EE3FB60ED1">journal article</mods:classification>
<mods:identifier id="FC42EB43C8D5F98FA52A465B15767386" type="CLB-Dataset">24728</mods:identifier>
<mods:identifier id="BCDA26A9C49D349757959B5E048681D4" type="DOI">10.11646/zootaxa.4702.1.8</mods:identifier>
<mods:identifier id="99953EB852515ABE08B2C269563F4DB3" type="GBIF-Dataset">f10d83aa-41cc-4a2b-8b37-dfabe65d5fd6</mods:identifier>
<mods:identifier id="449D405B1253AF656E5F3F087434B66D" type="ISSN">1175-5326</mods:identifier>
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<treatment id="992D879EE60CFFCCFF7EB1A4FD04FD6B" ID-GBIF-Taxon="161786749" LSID="urn:lsid:zoobank.org:act:A3E35168-14BF-455C-AA78-AE626E91CC5E" httpUri="http://treatment.plazi.org/id/992D879EE60CFFCCFF7EB1A4FD04FD6B" lastPageId="6" pageId="1">
<subSubSection id="599E6503E60CFFCBFF7EB1A4FDCCF900" pageId="1" type="nomenclature">
<paragraph id="113B3688E60CFFCBFF7EB1A4FE39F95C" blockId="1.[151,687,1665,1799]" box="[151,497,1665,1692]" pageId="1">
<heading id="4A7381E4E60CFFCBFF7EB1A4FE39F95C" bold="true" box="[151,497,1665,1692]" fontSize="11" level="1" pageId="1" reason="1">
<emphasis id="23F0EA9AE60CFFCBFF7EB1A4FE39F95C" bold="true" box="[151,497,1665,1692]" pageId="1">
<taxonomicName id="D6844D0BE60CFFCBFF7EB1A4FE44F95B" authority="Ho, 2019" authorityName="Ho" authorityYear="2019" box="[151,396,1665,1692]" class="Actinopterygii" family="Moridae" genus="Pterophycis" kingdom="Animalia" order="Gadiformes" pageId="1" phylum="Chordata" rank="species" species="spatium" status="sp. nov.">
<emphasis id="23F0EA9AE60CFFCBFF7EB1A4FE44F95B" bold="true" box="[151,396,1665,1692]" italics="true" pageId="1">Pterophycis spatium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="38C357E1E60CFFCBFE7DB1A7FE39F95C" box="[404,497,1666,1692]" pageId="1" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="113B3688E60CFFCBFF7EB183FDCCF900" blockId="1.[151,687,1665,1799]" box="[151,516,1701,1728]" pageId="1">New English name: Bigfin mora</paragraph>
</subSubSection>
<subSubSection id="599E6503E60CFFCBFF7EB1EFFE01F8C7" pageId="1" type="description">
<paragraph id="113B3688E60CFFCBFF7EB1EFFD67F924" blockId="1.[151,687,1665,1799]" box="[151,687,1736,1764]" pageId="1">
New Chinese name:
<emphasis id="23F0EA9AE60CFFCBFE90B1EDFE21F91F" bold="true" box="[377,489,1736,1759]" pageId="1">ƛñëḋ</emphasis>
(Da-chi-zhi-shue)
</paragraph>
<paragraph id="113B3688E60CFFCBFF7EB1C8FE01F8C7" blockId="1.[151,687,1665,1799]" box="[151,457,1773,1799]" pageId="1">
<figureCitation id="89BF2A0DE60CFFCBFF7EB1C8FF32F8C7" box="[151,250,1773,1799]" captionStart="FIGURE 1" captionStartId="2.[151,250,1910,1935]" captionTargetBox="[322,1264,205,1872]" captionTargetId="figure@2.[322,1264,181,1885]" captionTargetPageId="2" captionText="FIGURE 1. Ventral view of abdominal region comparing with the pelvic fins, light organ (left arrow), anus (middle arrow) and genital papilla (right arrow). A. Pterophycis gen. nov., NMMB-P22891. B. Holotype of Physiculus dalwigkii, type species of Physiculus, MNHN 1996-1380, 190 mm SL. C. Physiculus chigodarana, NMMB-P16401, 118 mm SL." figureDoi="http://doi.org/10.5281/zenodo.3562845" httpUri="https://zenodo.org/record/3562845/files/figure.png" pageId="1">Figs. 1A</figureCitation>
,
<figureCitation id="89BF2A0DE60CFFCBFEEFB1C8FEE2F8C7" box="[262,298,1773,1799]" captionStart="FIGURE 2" captionStartId="3.[151,250,1671,1696]" captionTargetBox="[227,1361,181,1645]" captionTargetId="figure@3.[227,1361,181,1645]" captionTargetPageId="3" captionText="FIGURE 2. Dorsolateral view of head showing the anterior extension of swimbladder (arrowed). A. Pterophycis gen. nov., NMMB-P22891. B. Physiculus dalwigkii, type species of Physiculus, MNHN A.4502, 197 mm SL." figureDoi="http://doi.org/10.5281/zenodo.3562847" httpUri="https://zenodo.org/record/3562847/files/figure.png" pageId="1">2A</figureCitation>
,
<figureCitation id="89BF2A0DE60CFFCBFEDFB1C8FE8AF8C7" box="[310,322,1773,1799]" captionStart="FIGURE 3" captionStartId="4.[151,250,1944,1969]" captionTargetBox="[214,1373,181,1917]" captionTargetId="figure@4.[214,1373,181,1918]" captionTargetPageId="4" captionText="FIGURE 3. Pterophycis spatium sp. nov., holotype, NMMB-P22891, 243 mm SL. A. Lateral view. B. Ventral view of head. C. Lateral view of head." figureDoi="http://doi.org/10.5281/zenodo.3562849" httpUri="https://zenodo.org/record/3562849/files/figure.png" pageId="1">3</figureCitation>
AC, 4AB
</paragraph>
</subSubSection>
<subSubSection id="599E6503E60CFFCBFF7EB013FA93F858" pageId="1" type="materials_examined">
<paragraph id="113B3688E60CFFCBFF7EB013FC1BF8B4" blockId="1.[151,1436,1845,1980]" pageId="1">
<materialsCitation id="A1EC3CD5E60CFFCBFF7EB013FC1BF8B4" ID-GBIF-Occurrence="2556252195" collectingDate="2015-05-04" collectionCode="NMMB-P" country="Taiwan" location="Strait" pageId="1" specimenCode="NMMB-P22891" specimenCount="1" stateProvince="Kaohsiung" typeStatus="holotype">
<emphasis id="23F0EA9AE60CFFCBFF7EB013FEC3F890" bold="true" box="[151,267,1846,1872]" pageId="1">
<typeStatus id="CE3F882AE60CFFCBFF7EB013FECEF890" box="[151,262,1846,1872]" pageId="1" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="41229EF3E60CFFCBFEFCB010FE04F88F" box="[277,460,1845,1871]" collectionCode="NMMB-P" pageId="1">NMMB-P22891</specimenCode>
(
<quantity id="D67C9B6DE60CFFCBFE37B010FDF7F890" box="[478,575,1845,1872]" metricMagnitude="-1" metricUnit="m" metricValue="2.43" pageId="1" unit="mm" value="243.0">243 mm</quantity>
SL), Ke-tzu-liao,
<collectingRegion id="D340F86AE60CFFCBFCFAB010FC47F88F" box="[787,911,1845,1871]" country="Taiwan" name="Kaohsiung" pageId="1">Kaohsiung</collectingRegion>
, southwestern
<collectingCountry id="69937618E60CFFCBFBD3B010FB47F88F" box="[1082,1167,1845,1871]" name="Taiwan" pageId="1">Taiwan</collectingCountry>
, western
<collectingCountry id="69937618E60CFFCBFB14B010FA87F88F" box="[1277,1359,1845,1871]" name="Taiwan" pageId="1">Taiwan</collectingCountry>
<location id="145B6053E60CFFCBFAB1B010FA50F88F" LSID="urn:lsid:plazi:treatment:992D879EE60CFFCCFF7EB1A4FD04FD6B:145B6053E60CFFCBFAB1B010FA50F88F" box="[1368,1432,1845,1871]" country="Taiwan" name="Strait" pageId="1" stateProvince="Kaohsiung">Strait</location>
, purchased from fish market, collected by local bottom trawl,
<date id="653A1048E60CFFCBFCADB07CFC18F8B4" box="[836,976,1881,1908]" pageId="1" value="2015-05-04">
<collectingDate id="757EE9A0E60CFFCBFCADB07CFC18F8B4" box="[836,976,1881,1908]" pageId="1" value="2015-05-04">4 May 2015</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="113B3688E60CFFCBFF2EB05BFA93F858" blockId="1.[151,1436,1845,1980]" box="[199,1371,1917,1944]" pageId="1">
<materialsCitation id="A1EC3CD5E60CFFCBFF2EB05BFA9FF858" ID-GBIF-Occurrence="2556252194" box="[199,1367,1917,1944]" collectingDate="1985-12-01" collectionCode="ASIZP" country="Taiwan" location="Chong-chou" pageId="1" specimenCode="ASIZP 58039" specimenCount="1" stateProvince="Kaohsiung" typeStatus="paratype">
<emphasis id="23F0EA9AE60CFFCBFF2EB05BFEF3F858" bold="true" box="[199,315,1918,1944]" pageId="1">
<typeStatus id="CE3F882AE60CFFCBFF2EB05BFEFEF858" box="[199,310,1918,1944]" pageId="1" type="paratype">Paratype</typeStatus>
.
</emphasis>
<specimenCode id="41229EF3E60CFFCBFEA9B058FE12F858" box="[320,474,1917,1944]" collectionCode="ASIZP" country="Taiwan" httpUri="http://grbio.org/cool/zgdx-i3kf" name="Academia Sinica Institute of Zoology, Ichthyology Collection" pageId="1">ASIZP 58039</specimenCode>
(1, 210+ mm SL),
<location id="145B6053E60CFFCBFD5BB058FCF7F857" LSID="urn:lsid:plazi:treatment:992D879EE60CFFCCFF7EB1A4FD04FD6B:145B6053E60CFFCBFD5BB058FCF7F857" box="[690,831,1917,1943]" country="Taiwan" name="Chong-chou" pageId="1" stateProvince="Kaohsiung">Chong-chou</location>
,
<collectingRegion id="D340F86AE60CFFCBFCA3B058FC0EF857" box="[842,966,1917,1943]" country="Taiwan" name="Kaohsiung" pageId="1">Kaohsiung</collectingRegion>
, southwestern
<collectingCountry id="69937618E60CFFCBFB85B058FB09F857" box="[1132,1217,1917,1943]" name="Taiwan" pageId="1">Taiwan</collectingCountry>
,
<date id="653A1048E60CFFCBFB25B058FA9FF858" box="[1228,1367,1917,1944]" pageId="1" value="1985-12-01">
<collectingDate id="757EE9A0E60CFFCBFB25B058FA9FF858" box="[1228,1367,1917,1944]" pageId="1" value="1985-12-01">1 Dec. 1985</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="599E6503E60CFFCBFF2EB084FD84F87C" box="[199,588,1953,1980]" pageId="1" type="diagnosis">
<paragraph id="113B3688E60CFFCBFF2EB084FD84F87C" blockId="1.[151,1436,1845,1980]" box="[199,588,1953,1980]" pageId="1">
<emphasis id="23F0EA9AE60CFFCBFF2EB084FE8AF87C" bold="true" box="[199,322,1953,1980]" pageId="1">Diagnosis.</emphasis>
As given for the genus.
</paragraph>
</subSubSection>
<caption id="45FB6600E60FFFC8FF7EB053FB7CF817" ID-DOI="http://doi.org/10.5281/zenodo.3562845" ID-Zenodo-Dep="3562845" httpUri="https://zenodo.org/record/3562845/files/figure.png" pageId="2" startId="2.[151,250,1910,1935]" targetBox="[322,1264,205,1872]" targetPageId="2">
<paragraph id="113B3688E60FFFC8FF7EB053FB7CF817" blockId="2.[151,1437,1910,2007]" pageId="2">
<emphasis id="23F0EA9AE60FFFC8FF7EB053FEDBF84F" bold="true" box="[151,275,1910,1935]" pageId="2">FIGURE 1.</emphasis>
Ventral view of abdominal region comparing with the pelvic fins, light organ (left arrow), anus (middle arrow) and genital papilla (right arrow). A.
<taxonomicName id="D6844D0BE60FFFC8FE09B0BEFD90F873" authority="Ho, 2019" authorityName="Ho" authorityYear="2019" box="[480,600,1947,1971]" class="Actinopterygii" family="Moridae" genus="Pterophycis" kingdom="Animalia" order="Gadiformes" pageId="2" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="23F0EA9AE60FFFC8FE09B0BEFD90F873" box="[480,600,1947,1971]" italics="true" pageId="2">Pterophycis</emphasis>
</taxonomicName>
<emphasis id="23F0EA9AE60FFFC8FDB6B0BEFD77F873" bold="true" box="[607,703,1947,1971]" pageId="2">
<taxonomicNameLabel id="38C357E1E60FFFC8FDB6B0BEFD77F873" box="[607,703,1947,1971]" pageId="2" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, NMMB-P22891. B. Holotype of
<taxonomicName id="D6844D0BE60FFFC8FBCBB0BEFB3DF873" authorityName="Kaup" authorityYear="1858" box="[1058,1269,1947,1971]" class="Actinopterygii" family="Moridae" genus="Physiculus" kingdom="Animalia" order="Gadiformes" pageId="2" phylum="Chordata" rank="species" species="dalwigkii">
<emphasis id="23F0EA9AE60FFFC8FBCBB0BEFB3DF873" box="[1058,1269,1947,1971]" italics="true" pageId="2">Physiculus dalwigkii</emphasis>
</taxonomicName>
, type species of
<taxonomicName id="D6844D0BE60FFFC8FF7EB09AFECDF817" authorityName="Kaup" authorityYear="1858" box="[151,261,1983,2007]" class="Actinopterygii" family="Moridae" genus="Physiculus" kingdom="Animalia" order="Gadiformes" pageId="2" phylum="Chordata" rank="genus">
<emphasis id="23F0EA9AE60FFFC8FF7EB09AFECDF817" box="[151,261,1983,2007]" italics="true" pageId="2">Physiculus</emphasis>
</taxonomicName>
, MNHN 1996-1380, 190 mm SL. C.
<taxonomicName id="D6844D0BE60FFFC8FD6BB09AFCBFF817" authorityName="Paulin" authorityYear="1989" box="[642,887,1983,2007]" class="Actinopterygii" family="Moridae" genus="Physiculus" kingdom="Animalia" order="Gadiformes" pageId="2" phylum="Chordata" rank="species" species="chigodarana">
<emphasis id="23F0EA9AE60FFFC8FD6BB09AFCBFF817" box="[642,887,1983,2007]" italics="true" pageId="2">Physiculus chigodarana</emphasis>
</taxonomicName>
, NMMB-P16401, 118 mm SL.
</paragraph>
</caption>
<caption id="45FB6600E60EFFC9FF7EB1A2FB4FF904" ID-DOI="http://doi.org/10.5281/zenodo.3562847" ID-Zenodo-Dep="3562847" httpUri="https://zenodo.org/record/3562847/files/figure.png" pageId="3" startId="3.[151,250,1671,1696]" targetBox="[227,1361,181,1645]" targetPageId="3">
<paragraph id="113B3688E60EFFC9FF7EB1A2FB4FF904" blockId="3.[151,1436,1671,1733]" pageId="3">
<emphasis id="23F0EA9AE60EFFC9FF7EB1A2FEDDF960" bold="true" box="[151,277,1671,1696]" pageId="3">FIGURE 2.</emphasis>
Dorsolateral view of head showing the anterior extension of swimbladder (arrowed). A.
<taxonomicName id="D6844D0BE60EFFC9FB5DB1ADFAE4F960" authority="Ho, 2019" authorityName="Ho" authorityYear="2019" box="[1204,1324,1672,1696]" class="Actinopterygii" family="Moridae" genus="Pterophycis" kingdom="Animalia" order="Gadiformes" pageId="3" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="23F0EA9AE60EFFC9FB5DB1ADFAE4F960" box="[1204,1324,1672,1696]" italics="true" pageId="3">Pterophycis</emphasis>
</taxonomicName>
<emphasis id="23F0EA9AE60EFFC9FADCB1ADFA5DF960" bold="true" box="[1333,1429,1672,1696]" pageId="3">
<taxonomicNameLabel id="38C357E1E60EFFC9FADCB1ADFA5DF960" box="[1333,1429,1672,1696]" pageId="3" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, NMMB-P22891. B.
<taxonomicName id="D6844D0BE60EFFC9FE81B189FDF2F904" authorityName="Kaup" authorityYear="1858" box="[360,570,1708,1732]" class="Actinopterygii" family="Moridae" genus="Physiculus" kingdom="Animalia" order="Gadiformes" pageId="3" phylum="Chordata" rank="species" species="dalwigkii">
<emphasis id="23F0EA9AE60EFFC9FE81B189FDF2F904" box="[360,570,1708,1732]" italics="true" pageId="3">Physiculus dalwigkii</emphasis>
</taxonomicName>
, type species of
<taxonomicName id="D6844D0BE60EFFC9FD0AB189FC99F904" authorityName="Kaup" authorityYear="1858" box="[739,849,1708,1732]" class="Actinopterygii" family="Moridae" genus="Physiculus" kingdom="Animalia" order="Gadiformes" pageId="3" phylum="Chordata" rank="genus">
<emphasis id="23F0EA9AE60EFFC9FD0AB189FC99F904" box="[739,849,1708,1732]" italics="true" pageId="3">Physiculus</emphasis>
</taxonomicName>
, MNHN A.4502, 197 mm SL.
</paragraph>
</caption>
<subSubSection id="599E6503E60EFFCCFF2EB033FC13FDDB" lastPageId="6" pageId="3" type="description">
<paragraph id="113B3688E60EFFC9FF2EB033FBDBF894" blockId="3.[151,1436,1814,1985]" pageId="3">
<emphasis id="23F0EA9AE60EFFC9FF2EB033FE91F8F1" bold="true" box="[199,345,1814,1841]" pageId="3">Description.</emphasis>
The following data are given for the
<typeStatus id="CE3F882AE60EFFC9FCE9B033FCADF8F0" box="[768,869,1814,1840]" pageId="3" type="holotype">holotype</typeStatus>
, followed by the value of
<typeStatus id="CE3F882AE60EFFC9FB64B032FB25F8F1" box="[1165,1261,1815,1841]" pageId="3" type="paratype">paratype</typeStatus>
in parentheses, when available. Characters given in the diagnosis of the genus are not repeated.
</paragraph>
<paragraph id="113B3688E60EFFC9FF2EB07BFC57F801" blockId="3.[151,1436,1814,1985]" pageId="3">First dorsal-fin rays 8 (8), second dorsal-fin rays 67 (47+); pectoral-fin rays 27 (29); anal-fin rays 71 (47+); pelvic-fin rays 7 (7/5); scales in longitudinal series ca. 125 (ca. 105+); scale rows between first dorsal-fin base and lateral line 12 (11); gill rakers 3+8=11 (3+4=7); vertebrae 15+40=55.</paragraph>
<caption id="45FB6600E609FFCEFF7EB0BDFEA7F815" ID-DOI="http://doi.org/10.5281/zenodo.3562849" ID-Zenodo-Dep="3562849" httpUri="https://zenodo.org/record/3562849/files/figure.png" pageId="4" startId="4.[151,250,1944,1969]" targetBox="[214,1373,181,1917]" targetPageId="4">
<paragraph id="113B3688E609FFCEFF7EB0BDFEA7F815" blockId="4.[151,1437,1944,2005]" pageId="4">
<emphasis id="23F0EA9AE609FFCEFF7EB0BDFEDAF871" bold="true" box="[151,274,1944,1969]" pageId="4">FIGURE 3.</emphasis>
<taxonomicName id="D6844D0BE609FFCEFEF0B0BCFE2FF870" authority="Ho, 2019" authorityName="Ho" authorityYear="2019" box="[281,487,1945,1969]" class="Actinopterygii" family="Moridae" genus="Pterophycis" kingdom="Animalia" order="Gadiformes" pageId="4" phylum="Chordata" rank="species" species="spatium" status="sp. nov.">
<emphasis id="23F0EA9AE609FFCEFEF0B0BCFE2FF870" box="[281,487,1945,1969]" italics="true" pageId="4">Pterophycis spatium</emphasis>
</taxonomicName>
<emphasis id="23F0EA9AE609FFCEFE04B0BCFDF5F871" bold="true" box="[493,573,1945,1969]" pageId="4">
<taxonomicNameLabel id="38C357E1E609FFCEFE04B0BCFDF5F871" box="[493,573,1945,1969]" pageId="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, NMMB-P22891, 243 mm SL. A. Lateral view. B. Ventral view of head. C. Lateral view of head.
</paragraph>
</caption>
<caption id="45FB6600E608FFCFFF7EB3E7FE88FB3F" ID-DOI="http://doi.org/10.5281/zenodo.3562851" ID-Zenodo-Dep="3562851" httpUri="https://zenodo.org/record/3562851/files/figure.png" pageId="5" startId="5.[151,250,1218,1243]" targetBox="[175,1435,202,1129]" targetPageId="5">
<paragraph id="113B3688E608FFCFFF7EB3E7FE88FB3F" blockId="5.[151,1436,1218,1279]" pageId="5">
<emphasis id="23F0EA9AE608FFCFFF7EB3E7FEDBFB1B" bold="true" box="[151,275,1218,1243]" pageId="5">FIGURE 4.</emphasis>
Paratype of
<taxonomicName id="D6844D0BE608FFCFFE7DB3E6FDAAFB1B" authority="Ho, 2019" authorityName="Ho" authorityYear="2019" box="[404,610,1219,1243]" class="Actinopterygii" family="Moridae" genus="Pterophycis" kingdom="Animalia" order="Gadiformes" pageId="5" phylum="Chordata" rank="species" species="spatium" status="sp. nov.">
<emphasis id="23F0EA9AE608FFCFFE7DB3E6FDAAFB1B" box="[404,610,1219,1243]" italics="true" pageId="5">Pterophycis spatium</emphasis>
</taxonomicName>
<emphasis id="23F0EA9AE608FFCFFD80B3E6FD72FB1B" bold="true" box="[617,698,1219,1243]" pageId="5">
<taxonomicNameLabel id="38C357E1E608FFCFFD80B3E6FD72FB1B" box="[617,698,1219,1243]" pageId="5" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ASIZP 58039, 210+ mm SL. A. lateral view (reversed). B. Ventral view of anterior body.
</paragraph>
</caption>
<paragraph id="113B3688E608FFCFFF2EB208FE05FA18" blockId="5.[151,1437,1325,2035]" pageId="5">Head length 4.0 times in SL; predorsal length 3.3; prepectoral length 3.7; prepelvic length 4.2; preanal length 2.7; dorsal-fin base 1.5; anal-fin base 1.5; pectoral-fin length 6.0; pelvic-fin length 4.1. Snout 3.3 (3.1) times in head length; eye diameter 4.0 (3.9); interorbital width 4.5 (4.3); upper-jaw length 2.2 (2.1); post-orbital length 1.9 (2.0); barbel 6.6 (6.5); longest first dorsal-fin ray 2.8 (2.8); longest second dorsal-fin ray 2.8; caudal-peduncle length 6.5; caudal-peduncle height 8.5.</paragraph>
<paragraph id="113B3688E608FFCFFF2EB2C4FF37F96F" blockId="5.[151,1437,1325,2035]" pageId="5">Body moderately elongate, robust anteriorly, gradually compressed and narrowing to a narrow caudal peduncle and caudal fin. Head large and broadly rounded, slightly depressed anteriorly; snout wide with a round anterior outline; eye large, its diameter slightly wider than interorbital space. Mouth large and terminal; maxilla extends to a vertical near the posterior margin of eye; upper jaw slightly overhanging the lower jaw. Gill slit wide. Teeth villiform, in wide band on both jaws; vomerine and palatine toothless. Chin barbel present, its length shorter than eye diameter.</paragraph>
<paragraph id="113B3688E608FFCFFF2EB19FFEACF847" blockId="5.[151,1437,1325,2035]" pageId="5">Two dorsal fins. Origin of first dorsal fin clearly behind a vertical of insertion of pectoral fin; origin of anal fin slightly behind a vertical of that of second dorsal fin; pectoral fin with broad base, rounded posteriorly, with lower rays gradually shorter; pelvic fin thoracic, extremely large, inserting slightly anterior to pectoral fin, when appressed, it tip extends to base of 15th (21st) ray of anal fin, all rays elongate, subequal in length; caudal fin small, well separated from dorsal and anal fins, rounded and symmetrical posteriorly. Caudal peduncle short, its length less than eye diameter.</paragraph>
<paragraph id="113B3688E608FFCFFF2EB0B4FD49F833" blockId="5.[151,1437,1325,2035]" pageId="5">Scales cycloid, small, oval-shaped; scale patch extends to anterior nostril and below middle of eye; naked area on snout large, not reaching to middle of eye. Gular region and branchiostegal membrane naked. Scales in front of pelvic-fin base gradually smaller anteriorly.</paragraph>
<paragraph id="113B3688E60BFFCCFF2EB7BCFC93FEA7" blockId="6.[151,1436,153,683]" pageId="6">
Luminous organ large, distance between its anterior and posterior margins 12.7% (22.0%) of distance between interventral line and origin of anal fin (InV-af); its locality closer to interventral line than anus (at middle point of interventral line and anus in
<typeStatus id="CE3F882AE60BFFCCFE35B7C4FD89FF3B" box="[476,577,225,251]" pageId="6" type="paratype">paratype</typeStatus>
), distance between interventral line and anterior margin of light organ (InV- LO) 21.7% (29.8%) of InV-af; anus large and rounded, circled by black naked skin, situating slightly closer to interventral line than origin of anal fin (at middle point of interventral line and origin of anal fin in
<typeStatus id="CE3F882AE60BFFCCFB31B60CFAF6FE83" box="[1240,1342,297,323]" pageId="6" type="paratype">paratype</typeStatus>
); genital papilla separated posteriorly from anus by several small scales.
</paragraph>
<paragraph id="113B3688E60BFFCCFF2EB655FC13FDDB" blockId="6.[151,1436,153,683]" pageId="6">
<emphasis id="23F0EA9AE60BFFCCFF2EB655FE87FE4B" bold="true" box="[199,335,368,395]" pageId="6">Coloration.</emphasis>
When fresh body reddish brown and trunk blue-tan; branchiostegal membrane black; first dorsal fin and pectoral- and pelvic- fin bases darker; second dorsal and anal fins with indistinct black margins. The preserved
<typeStatus id="CE3F882AE60BFFCCFF01B69CFE82FE13" box="[232,330,441,467]" pageId="6" type="holotype">holotype</typeStatus>
is darker than its fresh condition overall. The long-term preserved
<typeStatus id="CE3F882AE60BFFCCFBABB69CFB6AFE13" box="[1090,1186,441,467]" pageId="6" type="paratype">paratype</typeStatus>
lost most of its coloration; head light brown; gular region, branchiostegal membranes, naked area of snout and outer margins of dorsal and anal fins deep brownish, with all other portions of body creamy white.
</paragraph>
</subSubSection>
<subSubSection id="599E6503E60BFFCCFF2EB501FD44FDA3" pageId="6" type="distribution">
<paragraph id="113B3688E60BFFCCFF2EB501FD44FDA3" blockId="6.[151,1436,153,683]" pageId="6">
<emphasis id="23F0EA9AE60BFFCCFF2EB501FEA9FDFF" bold="true" box="[199,353,548,575]" pageId="6">Distribution.</emphasis>
Only known from the shallow waters (less than
<quantity id="D67C9B6DE60BFFCCFC61B500FC19FDFF" box="[904,977,549,575]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="6" unit="m" value="100.0">100 m</quantity>
) in southwestern
<collectingCountry id="69937618E60BFFCCFB73B500FB24FDFF" box="[1178,1260,549,575]" name="Taiwan" pageId="6">Taiwan</collectingCountry>
off
<collectingRegion id="D340F86AE60BFFCCFAF5B500FA50FDFF" box="[1308,1432,549,575]" country="Taiwan" name="Kaohsiung" pageId="6">Kaohsiung</collectingRegion>
, western
<collectingCountry id="69937618E60BFFCCFF1CB56CFE8FFDA3" box="[245,327,585,611]" name="Taiwan" pageId="6">Taiwan</collectingCountry>
Strait, southwestern
<collectingCountry id="69937618E60BFFCCFDDAB56CFD41FDA3" box="[563,649,585,611]" name="Taiwan" pageId="6">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="599E6503E60BFFCCFF2EB548FD04FD6B" pageId="6" type="etymology">
<paragraph id="113B3688E60BFFCCFF2EB548FD04FD6B" blockId="6.[151,1436,153,683]" pageId="6">
<emphasis id="23F0EA9AE60BFFCCFF2EB548FE86FD47" bold="true" box="[199,334,621,647]" pageId="6">Etymology.</emphasis>
The specific name
<taxonomicName id="D6844D0BE60BFFCCFDC1B54BFD49FD47" authorityName="Ho" authorityYear="2019" box="[552,641,622,647]" class="Actinopterygii" family="Moridae" genus="Pterophycis" kingdom="Animalia" order="Gadiformes" pageId="6" phylum="Chordata" rank="species" species="spatium">
<emphasis id="23F0EA9AE60BFFCCFDC1B54BFD49FD47" box="[552,641,622,647]" italics="true" pageId="6">spatium</emphasis>
</taxonomicName>
, means distance in Latin, is referred to the clear gape between the anus and genital papilla, which is unique for the family.
</paragraph>
</subSubSection>
</treatment>
</document>