516 lines
50 KiB
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516 lines
50 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/alpento.3.37308" ID-GBIF-Dataset="adf22e94-ae9e-41f1-82d0-1487cdac4674" ID-GBIF-Taxon="159811133" ID-Pensoft-Pub="2535-0889-3-169" ID-Pensoft-UUID="3E9C8DB48EB65E35B3879BD2D61FE75F" LSID="urn:lsid:zoobank.org:act:39B9B98B-9B70-4A56-9361-D87B38716289" ModsDocID="2535-0889-3-169" checkinTime="1571756603840" checkinUser="pensoft" docAuthor="Balke, Michael, Ospina-Torres, Rodulfo, Megna, Yoandri S., Laython, Marco & Hendrich, Lars" docDate="2019" docId="8C3ED96BC7A55FF6BD5E4933BD972BB6" docLanguage="en" docName="AlpEntomol 3: 169-174" docOrigin="Alpine Entomology 3" docSource="http://dx.doi.org/10.3897/alpento.3.37308" docTitle="Rhantus bogotensis Balke, Ospina-Torres, Megna, Laython & Hendrich, 2019, sp. nov." docType="treatment" docUuid="39B9B98B-9B70-4A56-9361-D87B38716289" docUuidSource="ZooBank" docVersion="4" id="3E9C8DB48EB65E35B3879BD2D61FE75F" lastPageId="2" lastPageNumber="171" masterDocId="3E9C8DB48EB65E35B3879BD2D61FE75F" masterDocTitle="A new species of Rhantus diving beetles from the wetlands of the City of Bogota and surroundings (Coleoptera, Dytiscidae, Colymbetinae)" masterLastPageNumber="174" masterPageNumber="169" pageId="0" pageNumber="169" updateTime="1643508593820" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new species of Rhantus diving beetles from the wetlands of the City of Bogota and surroundings (Coleoptera, Dytiscidae, Colymbetinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balke, Michael</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ospina-Torres, Rodulfo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Megna, Yoandri S.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Laython, Marco</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hendrich, Lars</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Alpine Entomology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>169</mods:start>
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<mods:end>174</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/alpento.3.37308</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/alpento.3.37308</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2535-0889-3-169</mods:identifier>
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<mods:identifier type="Pensoft-UUID">3E9C8DB48EB65E35B3879BD2D61FE75F</mods:identifier>
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<mods:identifier type="ZooBank">5B852E87-BCC2-4902-988B-3CE4EFD88EC0</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159811133" LSID="urn:lsid:zoobank.org:act:39B9B98B-9B70-4A56-9361-D87B38716289" httpUri="http://treatment.plazi.org/id/8C3ED96BC7A55FF6BD5E4933BD972BB6" lastPageId="2" lastPageNumber="171" pageId="0" pageNumber="169">
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<subSubSection pageId="0" pageNumber="169" type="nomenclature">
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<paragraph pageId="0" pageNumber="169">
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<taxonomicName LSID="8C3ED96B-C7A5-5FF6-BD5E-4933BD972BB6" class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus bogotensis" order="Coleoptera" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="bogotensis">Rhantus bogotensis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="169">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Rhantus andinus (A), R. bogotensis sp. nov. (B), R. vicinus (C), R. franzi (D)." figureDoi="10.3897/alpento.3.37308.figure1" httpUri="https://binary.pensoft.net/fig/349401" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure1">Figs 1B</figureCitation>
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,
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Rhantus spp.: Pronotal margin of R. bogotensis sp. nov. female (A), male (B); R. vicinus female (C); R. bogotensis: surface sculpture on head, frons (D), middle part of pronotum with base and disc (E) elytron, basal area (F)." figureDoi="10.3897/alpento.3.37308.figure2" httpUri="https://binary.pensoft.net/fig/349402" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure2">2A, B, D, E, F</figureCitation>
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,
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Rhantus spp., males: Rhantus bogotensis sp. nov. parameres (A), median lobe of aedeagus in lateral view (B), same in ventral view (C), mesotarsus (D), mesotarsomere V and claws (E), foretarsus (F), protarsomere V and claws (G); R. vicinus: protarsomere V and claws (H); R. andinus: protarsomere V and claws (I); R. crypticus: protarsomere V and claws (J); R. franzi median lobe of aedeagus in lateral view (K), protarsomere III-V and claws (L), mesotarsus (M), mesotarsomere V and anterior claw, posterior claw missing (N). " a " denotes anterior claw." figureDoi="10.3897/alpento.3.37308.figure3" httpUri="https://binary.pensoft.net/fig/349403" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure3">
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3
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<normalizedToken originalValue="A–G">A-G</normalizedToken>
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</figureCitation>
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,
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitat of Rhantus bogotensis sp. nov. in the Altiplano of Bogota, Colombia. Florida wetland, deeper water bodies where we did encounter only very few specimens (A), shallow ponds, most likely ephemeral (B, C)." figureDoi="10.3897/alpento.3.37308.figure4" httpUri="https://binary.pensoft.net/fig/349404" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure4">4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="169" type="type locality">
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<paragraph pageId="0" pageNumber="169">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="169">Colombia, Bogota city, Juan Amarillo / Tibabuyes wetland.</paragraph>
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<paragraph pageId="0" pageNumber="169">
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<emphasis bold="true" pageId="0" pageNumber="169">Holotype.</emphasis>
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Male (ICN-UNAL): "Colombia,
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<normalizedToken originalValue="Bogotá">Bogota</normalizedToken>
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D.C.,
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<normalizedToken originalValue="Bogotá">Bogota</normalizedToken>
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, Humedal Tibabuyes, 2,500m, 25.ix.2003,
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<geoCoordinate degrees="4.7187" direction="north" orientation="latitude" precision="5" value="4.7187">4.7187</geoCoordinate>
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,
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<geoCoordinate degrees="74.0976" direction="west" orientation="longitude" precision="5" value="-74.0976">-74.0976</geoCoordinate>
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; dry mounted out of an Ethanol vial labelled "
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<taxonomicName lsidName="" order="Coleoptera" pageId="0" pageNumber="169" rank="order">Coleoptera</taxonomicName>
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<taxonomicName family="Dytiscidae" lsidName="" pageId="0" pageNumber="169" rank="family">Dytiscidae</taxonomicName>
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356,
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus" order="Coleoptera" pageId="0" pageNumber="169" phylum="Arthropoda" rank="genus">Rhantus</taxonomicName>
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, det. M. Laython 2016"; "HOLOTYPE
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus bogotensis" order="Coleoptera" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="bogotensis">
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<emphasis italics="true" pageId="0" pageNumber="169">Rhantus bogotensis</emphasis>
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</taxonomicName>
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sp. nov. Balke, Ospina-Torres, Megna, Laython & Hendrich, 2019" [red printed label].
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</paragraph>
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<paragraph pageId="0" pageNumber="169">
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<emphasis bold="true" pageId="0" pageNumber="169">240 Paratypes.</emphasis>
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2 exx, same data as holotype (ICN-ICN-UNAL); 1 ex., "Colombia: Bogota, La Florida, 2,436m, 19.iv.2017,
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<geoCoordinate degrees="4.728" direction="north" orientation="latitude" precision="55" value="4.728">4.728</geoCoordinate>
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,
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<geoCoordinate degrees="74.142" direction="west" orientation="longitude" precision="55" value="-74.142">-74.142</geoCoordinate>
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, Y.S. Megna & N. Stiven (12)", (MB 7823) (ZSM); 118 exx, "Colombia, Cundinamarca, Humedal La Florida, 2,400m, 19.xi.2018,
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<geoCoordinate degrees="4.729" direction="north" orientation="latitude" precision="55" value="4.729">4.729</geoCoordinate>
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<geoCoordinate degrees="74.143" direction="west" orientation="longitude" precision="55" value="-74.143">-74.143</geoCoordinate>
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, Ospina, Balke & Megna (COL_MB_2018_08)" (ICN-UNAL, ZSM); 119 exx, "Colombia, Cundinamarca, Humedal La Florida, 2,400m, 22.xi.2018,
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<geoCoordinate degrees="4.729" direction="north" orientation="latitude" precision="55" value="4.729">4.729</geoCoordinate>
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<geoCoordinate degrees="74.143" direction="west" orientation="longitude" precision="55" value="-74.143">-74.143</geoCoordinate>
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, Ospina, Balke & Megna (COL_MB_2018_12)" (ICN-UNAL, ZSM). All specimens bear our red printed paratype labels.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="169" type="description">
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<paragraph pageId="0" pageNumber="169">Description.</paragraph>
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<paragraph pageId="0" pageNumber="169">
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Holotype. A medium sized
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus" order="Coleoptera" pageId="0" pageNumber="169" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="169">Rhantus</emphasis>
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</taxonomicName>
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species, total length of holotype 11.3 mm, length without head 10.2 mm, greatest width 5.9 mm.
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</paragraph>
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<paragraph pageId="0" pageNumber="169">
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<emphasis bold="true" pageId="0" pageNumber="169">Colour</emphasis>
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: Head black with contrasting orange marks as in
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Rhantus andinus (A), R. bogotensis sp. nov. (B), R. vicinus (C), R. franzi (D)." figureDoi="10.3897/alpento.3.37308.figure1" httpUri="https://binary.pensoft.net/fig/349401" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure1">Fig. 1B</figureCitation>
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. Pronotum yellow orange, base as well discal patch somewhat black. Elytron orange with black irrorations. Body appendages, prothorax, prosternal process, epipleuron, coxae, postcoxal process and hind margins of abdominal sternites orange to dark ferruginous, rest of venter black.
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</paragraph>
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<paragraph pageId="0" pageNumber="169">
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<emphasis bold="true" pageId="0" pageNumber="169">Surface sculpture</emphasis>
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: Head with irregular meshes and dense, distinct punctation, no microreticulation (MR) visible (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Rhantus spp.: Pronotal margin of R. bogotensis sp. nov. female (A), male (B); R. vicinus female (C); R. bogotensis: surface sculpture on head, frons (D), middle part of pronotum with base and disc (E) elytron, basal area (F)." figureDoi="10.3897/alpento.3.37308.figure2" httpUri="https://binary.pensoft.net/fig/349402" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure2">Fig. 2D</figureCitation>
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); clypeus with double punctation only. Pronotum with irregular meshes and dense, distinct double punctation, disc with double punctation only; no MR visible on pronotum (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Rhantus spp.: Pronotal margin of R. bogotensis sp. nov. female (A), male (B); R. vicinus female (C); R. bogotensis: surface sculpture on head, frons (D), middle part of pronotum with base and disc (E) elytron, basal area (F)." figureDoi="10.3897/alpento.3.37308.figure2" httpUri="https://binary.pensoft.net/fig/349402" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure2">Fig. 2E</figureCitation>
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). Elytron with distinct, irregular polygonal meshes and fine punctation; with distinct though sometimes faint MR within the meshes (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Rhantus spp.: Pronotal margin of R. bogotensis sp. nov. female (A), male (B); R. vicinus female (C); R. bogotensis: surface sculpture on head, frons (D), middle part of pronotum with base and disc (E) elytron, basal area (F)." figureDoi="10.3897/alpento.3.37308.figure2" httpUri="https://binary.pensoft.net/fig/349402" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure2">Fig. 2F</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="169">
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<emphasis bold="true" pageId="0" pageNumber="169">Structures</emphasis>
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: Pronotum with broad and conspicuous lateral bead which does however not reach the anterior angle (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Rhantus spp.: Pronotal margin of R. bogotensis sp. nov. female (A), male (B); R. vicinus female (C); R. bogotensis: surface sculpture on head, frons (D), middle part of pronotum with base and disc (E) elytron, basal area (F)." figureDoi="10.3897/alpento.3.37308.figure2" httpUri="https://binary.pensoft.net/fig/349402" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure2">Fig. 2B</figureCitation>
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). Lateral wings of metaventrite triangular; hind wings fully developed. Last ventrite rounded apically.
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</paragraph>
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<paragraph pageId="0" pageNumber="169">
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<emphasis bold="true" pageId="0" pageNumber="169">Tarsal characters</emphasis>
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: Protarsal claws (
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Rhantus spp., males: Rhantus bogotensis sp. nov. parameres (A), median lobe of aedeagus in lateral view (B), same in ventral view (C), mesotarsus (D), mesotarsomere V and claws (E), foretarsus (F), protarsomere V and claws (G); R. vicinus: protarsomere V and claws (H); R. andinus: protarsomere V and claws (I); R. crypticus: protarsomere V and claws (J); R. franzi median lobe of aedeagus in lateral view (K), protarsomere III-V and claws (L), mesotarsus (M), mesotarsomere V and anterior claw, posterior claw missing (N). " a " denotes anterior claw." figureDoi="10.3897/alpento.3.37308.figure3" httpUri="https://binary.pensoft.net/fig/349403" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure3">Fig. 3F, G</figureCitation>
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) with anterior and posterior claw of almost equal length, posterior claw slightly longer and about the same length as tarsomere V. Claws slender and with no particular modification (e.g. not dentate). Mesotarsal claws sinuate, with posterior claw shorter and more strongly curved than anterior claw, shorter than fifth protarsomere (
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Rhantus spp., males: Rhantus bogotensis sp. nov. parameres (A), median lobe of aedeagus in lateral view (B), same in ventral view (C), mesotarsus (D), mesotarsomere V and claws (E), foretarsus (F), protarsomere V and claws (G); R. vicinus: protarsomere V and claws (H); R. andinus: protarsomere V and claws (I); R. crypticus: protarsomere V and claws (J); R. franzi median lobe of aedeagus in lateral view (K), protarsomere III-V and claws (L), mesotarsus (M), mesotarsomere V and anterior claw, posterior claw missing (N). " a " denotes anterior claw." figureDoi="10.3897/alpento.3.37308.figure3" httpUri="https://binary.pensoft.net/fig/349403" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure3">Fig. 3D, E</figureCitation>
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). Pro- and mesotarsomeres 1-3 not dilated laterally (
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Rhantus spp., males: Rhantus bogotensis sp. nov. parameres (A), median lobe of aedeagus in lateral view (B), same in ventral view (C), mesotarsus (D), mesotarsomere V and claws (E), foretarsus (F), protarsomere V and claws (G); R. vicinus: protarsomere V and claws (H); R. andinus: protarsomere V and claws (I); R. crypticus: protarsomere V and claws (J); R. franzi median lobe of aedeagus in lateral view (K), protarsomere III-V and claws (L), mesotarsus (M), mesotarsomere V and anterior claw, posterior claw missing (N). " a " denotes anterior claw." figureDoi="10.3897/alpento.3.37308.figure3" httpUri="https://binary.pensoft.net/fig/349403" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure3">Fig. 3D, F</figureCitation>
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), with four transverse rows of stalked suction discs ventrally. Number of discs per row (tarsomere on which row occurs given in parentheses): 5(I)/6(I)/6(II)/5(III).
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</paragraph>
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<paragraph pageId="0" pageNumber="169">
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<emphasis bold="true" pageId="0" pageNumber="169">Genital structure</emphasis>
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: Median lobe of aedeagus as in
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Rhantus spp., males: Rhantus bogotensis sp. nov. parameres (A), median lobe of aedeagus in lateral view (B), same in ventral view (C), mesotarsus (D), mesotarsomere V and claws (E), foretarsus (F), protarsomere V and claws (G); R. vicinus: protarsomere V and claws (H); R. andinus: protarsomere V and claws (I); R. crypticus: protarsomere V and claws (J); R. franzi median lobe of aedeagus in lateral view (K), protarsomere III-V and claws (L), mesotarsus (M), mesotarsomere V and anterior claw, posterior claw missing (N). " a " denotes anterior claw." figureDoi="10.3897/alpento.3.37308.figure3" httpUri="https://binary.pensoft.net/fig/349403" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure3">Fig. 3B, C</figureCitation>
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, in lateral view appearing of longish curvature; parameres with a fringe of yellow hairs which are apically acute, not trumpet shaped (
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Rhantus spp., males: Rhantus bogotensis sp. nov. parameres (A), median lobe of aedeagus in lateral view (B), same in ventral view (C), mesotarsus (D), mesotarsomere V and claws (E), foretarsus (F), protarsomere V and claws (G); R. vicinus: protarsomere V and claws (H); R. andinus: protarsomere V and claws (I); R. crypticus: protarsomere V and claws (J); R. franzi median lobe of aedeagus in lateral view (K), protarsomere III-V and claws (L), mesotarsus (M), mesotarsomere V and anterior claw, posterior claw missing (N). " a " denotes anterior claw." figureDoi="10.3897/alpento.3.37308.figure3" httpUri="https://binary.pensoft.net/fig/349403" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure3">Fig. 3A</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="169">
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<emphasis bold="true" pageId="0" pageNumber="169">Female</emphasis>
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: Color and surface sculpture as in male. Tarsomere without stalked suction discs ventrally. The anterior margin of the anterior corner of pronotum (
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Rhantus spp.: Pronotal margin of R. bogotensis sp. nov. female (A), male (B); R. vicinus female (C); R. bogotensis: surface sculpture on head, frons (D), middle part of pronotum with base and disc (E) elytron, basal area (F)." figureDoi="10.3897/alpento.3.37308.figure2" httpUri="https://binary.pensoft.net/fig/349402" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure2">Fig. 2A</figureCitation>
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, white asterisk) thicker than in male specimens (
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Rhantus spp.: Pronotal margin of R. bogotensis sp. nov. female (A), male (B); R. vicinus female (C); R. bogotensis: surface sculpture on head, frons (D), middle part of pronotum with base and disc (E) elytron, basal area (F)." figureDoi="10.3897/alpento.3.37308.figure2" httpUri="https://binary.pensoft.net/fig/349402" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure2">Fig. 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="169" type="variation">
|
||
<paragraph pageId="0" pageNumber="169">Variation.</paragraph>
|
||
<paragraph pageId="0" pageNumber="169">
|
||
Body size varies from 11.0-12.1 mm. The discal dark spot on pronotum can be obsolete, or slightly more extended than in
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Rhantus andinus (A), R. bogotensis sp. nov. (B), R. vicinus (C), R. franzi (D)." figureDoi="10.3897/alpento.3.37308.figure1" httpUri="https://binary.pensoft.net/fig/349401" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure1">Fig. 1B</figureCitation>
|
||
, the same is true for the basal dark spot. Both can be approaching but do not connect.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="169" type="etymology">
|
||
<paragraph pageId="0" pageNumber="169">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="169">Named after the type locality.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="2" lastPageNumber="171" pageId="0" pageNumber="169" type="comparisons">
|
||
<paragraph pageId="0" pageNumber="169">Comparative notes.</paragraph>
|
||
<paragraph pageId="0" pageNumber="169">
|
||
A species well delineated from the other Colombian
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus" order="Coleoptera" pageId="0" pageNumber="169" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="169">Rhantus</emphasis>
|
||
</taxonomicName>
|
||
of similar size by the following set of features:
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="169">
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus franzi" order="Coleoptera" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="169">Rhantus franzi</emphasis>
|
||
</taxonomicName>
|
||
, described from the Cauca Valley and Ecuador (holotype male studied here) is similar in size but with pronotal coloration more similar to
|
||
<taxonomicName genus="R." lsidName="R. andinus" pageId="0" pageNumber="169" rank="species" species="andinus">
|
||
<emphasis italics="true" pageId="0" pageNumber="169">R. andinus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Rhantus andinus (A), R. bogotensis sp. nov. (B), R. vicinus (C), R. franzi (D)." figureDoi="10.3897/alpento.3.37308.figure1" httpUri="https://binary.pensoft.net/fig/349401" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure1">Fig. 1A, D</figureCitation>
|
||
). The protarsal claws are similar to
|
||
<taxonomicName genus="R." lsidName="R. bogotensis" pageId="0" pageNumber="169" rank="species" species="bogotensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="169">R. bogotensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Rhantus spp., males: Rhantus bogotensis sp. nov. parameres (A), median lobe of aedeagus in lateral view (B), same in ventral view (C), mesotarsus (D), mesotarsomere V and claws (E), foretarsus (F), protarsomere V and claws (G); R. vicinus: protarsomere V and claws (H); R. andinus: protarsomere V and claws (I); R. crypticus: protarsomere V and claws (J); R. franzi median lobe of aedeagus in lateral view (K), protarsomere III-V and claws (L), mesotarsus (M), mesotarsomere V and anterior claw, posterior claw missing (N). " a " denotes anterior claw." figureDoi="10.3897/alpento.3.37308.figure3" httpUri="https://binary.pensoft.net/fig/349403" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure3">Fig. 3L</figureCitation>
|
||
), but the anterior mesotarsal claw is simply curved in
|
||
<taxonomicName genus="R." lsidName="R. franzi" pageId="0" pageNumber="169" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="169">R. franzi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Rhantus spp., males: Rhantus bogotensis sp. nov. parameres (A), median lobe of aedeagus in lateral view (B), same in ventral view (C), mesotarsus (D), mesotarsomere V and claws (E), foretarsus (F), protarsomere V and claws (G); R. vicinus: protarsomere V and claws (H); R. andinus: protarsomere V and claws (I); R. crypticus: protarsomere V and claws (J); R. franzi median lobe of aedeagus in lateral view (K), protarsomere III-V and claws (L), mesotarsus (M), mesotarsomere V and anterior claw, posterior claw missing (N). " a " denotes anterior claw." figureDoi="10.3897/alpento.3.37308.figure3" httpUri="https://binary.pensoft.net/fig/349403" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure3">Fig. 3M, N</figureCitation>
|
||
) and not more broadly triangular as in
|
||
<taxonomicName genus="R." lsidName="R. bogotensis" pageId="0" pageNumber="169" rank="species" species="bogotensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="169">R. bogotensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Rhantus spp., males: Rhantus bogotensis sp. nov. parameres (A), median lobe of aedeagus in lateral view (B), same in ventral view (C), mesotarsus (D), mesotarsomere V and claws (E), foretarsus (F), protarsomere V and claws (G); R. vicinus: protarsomere V and claws (H); R. andinus: protarsomere V and claws (I); R. crypticus: protarsomere V and claws (J); R. franzi median lobe of aedeagus in lateral view (K), protarsomere III-V and claws (L), mesotarsus (M), mesotarsomere V and anterior claw, posterior claw missing (N). " a " denotes anterior claw." figureDoi="10.3897/alpento.3.37308.figure3" httpUri="https://binary.pensoft.net/fig/349403" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure3">Fig. 3D, E</figureCitation>
|
||
). The posterior mesotarsal claw is missing in the holotype of
|
||
<taxonomicName genus="R." lsidName="R. franzi" pageId="0" pageNumber="169" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="169">R. franzi</emphasis>
|
||
</taxonomicName>
|
||
studied here (and tarsus missing on other side of body). The median lobe of aedeagus only slightly more elongate in
|
||
<taxonomicName genus="R." lsidName="R. franzi" pageId="0" pageNumber="169" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="0" pageNumber="169">R. franzi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Rhantus spp., males: Rhantus bogotensis sp. nov. parameres (A), median lobe of aedeagus in lateral view (B), same in ventral view (C), mesotarsus (D), mesotarsomere V and claws (E), foretarsus (F), protarsomere V and claws (G); R. vicinus: protarsomere V and claws (H); R. andinus: protarsomere V and claws (I); R. crypticus: protarsomere V and claws (J); R. franzi median lobe of aedeagus in lateral view (K), protarsomere III-V and claws (L), mesotarsus (M), mesotarsomere V and anterior claw, posterior claw missing (N). " a " denotes anterior claw." figureDoi="10.3897/alpento.3.37308.figure3" httpUri="https://binary.pensoft.net/fig/349403" pageId="0" pageNumber="169" tableDoi="10.3897/alpento.3.37308.figure3">Fig. 3K</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="1" pageNumber="170">
|
||
<pageBreakToken pageId="1" pageNumber="170" start="start">Males</pageBreakToken>
|
||
of
|
||
<taxonomicName genus="R." lsidName="R. vicinus" pageId="1" pageNumber="170" rank="species" species="vicinus">
|
||
<emphasis italics="true" pageId="1" pageNumber="170">R. vicinus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName genus="R." lsidName="R. crypticus" pageId="1" pageNumber="170" rank="species" species="crypticus">
|
||
<emphasis italics="true" pageId="1" pageNumber="170">R. crypticus</emphasis>
|
||
</taxonomicName>
|
||
have the pro- and mesotarsomeres 1-3 distinctly dilated laterally, see
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Rhantus spp., males: Rhantus bogotensis sp. nov. parameres (A), median lobe of aedeagus in lateral view (B), same in ventral view (C), mesotarsus (D), mesotarsomere V and claws (E), foretarsus (F), protarsomere V and claws (G); R. vicinus: protarsomere V and claws (H); R. andinus: protarsomere V and claws (I); R. crypticus: protarsomere V and claws (J); R. franzi median lobe of aedeagus in lateral view (K), protarsomere III-V and claws (L), mesotarsus (M), mesotarsomere V and anterior claw, posterior claw missing (N). " a " denotes anterior claw." figureDoi="10.3897/alpento.3.37308.figure3" httpUri="https://binary.pensoft.net/fig/349403" pageId="1" pageNumber="170" tableDoi="10.3897/alpento.3.37308.figure3">Fig. 3J</figureCitation>
|
||
(not so in
|
||
<taxonomicName genus="R." lsidName="R. bogotensis" pageId="1" pageNumber="170" rank="species" species="bogotensis">
|
||
<emphasis italics="true" pageId="1" pageNumber="170">R. bogotensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Rhantus spp., males: Rhantus bogotensis sp. nov. parameres (A), median lobe of aedeagus in lateral view (B), same in ventral view (C), mesotarsus (D), mesotarsomere V and claws (E), foretarsus (F), protarsomere V and claws (G); R. vicinus: protarsomere V and claws (H); R. andinus: protarsomere V and claws (I); R. crypticus: protarsomere V and claws (J); R. franzi median lobe of aedeagus in lateral view (K), protarsomere III-V and claws (L), mesotarsus (M), mesotarsomere V and anterior claw, posterior claw missing (N). " a " denotes anterior claw." figureDoi="10.3897/alpento.3.37308.figure3" httpUri="https://binary.pensoft.net/fig/349403" pageId="1" pageNumber="170" tableDoi="10.3897/alpento.3.37308.figure3">Fig. 3F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph lastPageId="2" lastPageNumber="171" pageId="1" pageNumber="170">
|
||
Males of
|
||
<taxonomicName genus="R." lsidName="R. vicinus" pageId="1" pageNumber="170" rank="species" species="vicinus">
|
||
<emphasis italics="true" pageId="1" pageNumber="170">R. vicinus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName genus="R." lsidName="R. crypticus" pageId="1" pageNumber="170" rank="species" species="crypticus">
|
||
<emphasis italics="true" pageId="1" pageNumber="170">R. crypticus</emphasis>
|
||
</taxonomicName>
|
||
have the protarsal claws of unequal length and shape (
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Rhantus spp., males: Rhantus bogotensis sp. nov. parameres (A), median lobe of aedeagus in lateral view (B), same in ventral view (C), mesotarsus (D), mesotarsomere V and claws (E), foretarsus (F), protarsomere V and claws (G); R. vicinus: protarsomere V and claws (H); R. andinus: protarsomere V and claws (I); R. crypticus: protarsomere V and claws (J); R. franzi median lobe of aedeagus in lateral view (K), protarsomere III-V and claws (L), mesotarsus (M), mesotarsomere V and anterior claw, posterior claw missing (N). " a " denotes anterior claw." figureDoi="10.3897/alpento.3.37308.figure3" httpUri="https://binary.pensoft.net/fig/349403" pageId="1" pageNumber="170" tableDoi="10.3897/alpento.3.37308.figure3">Fig. 3H, J</figureCitation>
|
||
), anterior one much shorter and broader than posterior, with dent like shape in
|
||
<taxonomicName genus="R." lsidName="R. crypticus" pageId="1" pageNumber="170" rank="species" species="crypticus">
|
||
<emphasis italics="true" pageId="1" pageNumber="170">R. crypticus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Rhantus spp., males: Rhantus bogotensis sp. nov. parameres (A), median lobe of aedeagus in lateral view (B), same in ventral view (C), mesotarsus (D), mesotarsomere V and claws (E), foretarsus (F), protarsomere V and claws (G); R. vicinus: protarsomere V and claws (H); R. andinus: protarsomere V and claws (I); R. crypticus: protarsomere V and claws (J); R. franzi median lobe of aedeagus in lateral view (K), protarsomere III-V and claws (L), mesotarsus (M), mesotarsomere V and anterior claw, posterior claw missing (N). " a " denotes anterior claw." figureDoi="10.3897/alpento.3.37308.figure3" httpUri="https://binary.pensoft.net/fig/349403" pageId="1" pageNumber="170" tableDoi="10.3897/alpento.3.37308.figure3">Fig. 3J</figureCitation>
|
||
) (subequal in
|
||
<taxonomicName genus="R." lsidName="R. bogotensis" pageId="1" pageNumber="170" rank="species" species="bogotensis">
|
||
<emphasis italics="true" pageId="1" pageNumber="170">R. bogotensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Rhantus spp., males: Rhantus bogotensis sp. nov. parameres (A), median lobe of aedeagus in lateral view (B), same in ventral view (C), mesotarsus (D), mesotarsomere V and claws (E), foretarsus (F), protarsomere V and claws (G); R. vicinus: protarsomere V and claws (H); R. andinus: protarsomere V and claws (I); R. crypticus: protarsomere V and claws (J); R. franzi median lobe of aedeagus in lateral view (K), protarsomere III-V and claws (L), mesotarsus (M), mesotarsomere V and anterior claw, posterior claw missing (N). " a " denotes anterior claw." figureDoi="10.3897/alpento.3.37308.figure3" httpUri="https://binary.pensoft.net/fig/349403" pageId="1" pageNumber="170" tableDoi="10.3897/alpento.3.37308.figure3">Fig. 3F, G</figureCitation>
|
||
). The protarsal and mesotarsal claws of
|
||
<taxonomicName genus="R." lsidName="R. andinus" pageId="1" pageNumber="170" rank="species" species="andinus">
|
||
<emphasis italics="true" pageId="1" pageNumber="170">R. andinus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Rhantus spp., males: Rhantus bogotensis sp. nov. parameres (A), median lobe of aedeagus in lateral view (B), same in ventral view (C), mesotarsus (D), mesotarsomere V and claws (E), foretarsus (F), protarsomere V and claws (G); R. vicinus: protarsomere V and claws (H); R. andinus: protarsomere V and claws (I); R. crypticus: protarsomere V and claws (J); R. franzi median lobe of aedeagus in lateral view (K), protarsomere III-V and claws (L), mesotarsus (M), mesotarsomere V and anterior claw, posterior claw missing (N). " a " denotes anterior claw." figureDoi="10.3897/alpento.3.37308.figure3" httpUri="https://binary.pensoft.net/fig/349403" pageId="1" pageNumber="170" tableDoi="10.3897/alpento.3.37308.figure3">Fig. 3I</figureCitation>
|
||
) are similar to
|
||
<taxonomicName genus="R." lsidName="R. bogotensis" pageId="2" pageNumber="171" rank="species" species="bogotensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">
|
||
<pageBreakToken pageId="2" pageNumber="171" start="start">R</pageBreakToken>
|
||
. bogotensis
|
||
</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., but the beetles are easily separable by body size and coloration alone (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Rhantus andinus (A), R. bogotensis sp. nov. (B), R. vicinus (C), R. franzi (D)." figureDoi="10.3897/alpento.3.37308.figure1" httpUri="https://binary.pensoft.net/fig/349401" pageId="2" pageNumber="171" tableDoi="10.3897/alpento.3.37308.figure1">Fig. 1A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="2" pageNumber="171">
|
||
Females of
|
||
<taxonomicName genus="R." lsidName="R. vicinus" pageId="2" pageNumber="171" rank="species" species="vicinus">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">R. vicinus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName genus="R." lsidName="R. crypticus" pageId="2" pageNumber="171" rank="species" species="crypticus">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">R. crypticus</emphasis>
|
||
</taxonomicName>
|
||
have the anterior angle of the pronotum distinctly extended (
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Rhantus spp.: Pronotal margin of R. bogotensis sp. nov. female (A), male (B); R. vicinus female (C); R. bogotensis: surface sculpture on head, frons (D), middle part of pronotum with base and disc (E) elytron, basal area (F)." figureDoi="10.3897/alpento.3.37308.figure2" httpUri="https://binary.pensoft.net/fig/349402" pageId="2" pageNumber="171" tableDoi="10.3897/alpento.3.37308.figure2">Fig. 2C</figureCitation>
|
||
, arrow). This is not the case in
|
||
<taxonomicName genus="R." lsidName="R. bogotensis" pageId="2" pageNumber="171" rank="species" species="bogotensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">R. bogotensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Rhantus spp.: Pronotal margin of R. bogotensis sp. nov. female (A), male (B); R. vicinus female (C); R. bogotensis: surface sculpture on head, frons (D), middle part of pronotum with base and disc (E) elytron, basal area (F)." figureDoi="10.3897/alpento.3.37308.figure2" httpUri="https://binary.pensoft.net/fig/349402" pageId="2" pageNumber="171" tableDoi="10.3897/alpento.3.37308.figure2">Fig. 2A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="2" pageNumber="171">
|
||
Specimens of
|
||
<taxonomicName genus="R." lsidName="R. vicinus" pageId="2" pageNumber="171" rank="species" species="vicinus">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">R. vicinus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Rhantus andinus (A), R. bogotensis sp. nov. (B), R. vicinus (C), R. franzi (D)." figureDoi="10.3897/alpento.3.37308.figure1" httpUri="https://binary.pensoft.net/fig/349401" pageId="2" pageNumber="171" tableDoi="10.3897/alpento.3.37308.figure1">Fig. 1C</figureCitation>
|
||
) and
|
||
<taxonomicName genus="R." lsidName="R. crypticus" pageId="2" pageNumber="171" rank="species" species="crypticus">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">R. crypticus</emphasis>
|
||
</taxonomicName>
|
||
are generally of darker appearance than
|
||
<taxonomicName genus="R." lsidName="R. bogotensis" pageId="2" pageNumber="171" rank="species" species="bogotensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">R. bogotensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Rhantus andinus (A), R. bogotensis sp. nov. (B), R. vicinus (C), R. franzi (D)." figureDoi="10.3897/alpento.3.37308.figure1" httpUri="https://binary.pensoft.net/fig/349401" pageId="2" pageNumber="171" tableDoi="10.3897/alpento.3.37308.figure1">Fig. 1B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="171" type="distribution">
|
||
<paragraph pageId="2" pageNumber="171">Distribution.</paragraph>
|
||
<paragraph pageId="2" pageNumber="171">Altiplano of the Bogota region.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="171" type="habitat">
|
||
<paragraph pageId="2" pageNumber="171">Habitat.</paragraph>
|
||
<paragraph pageId="2" pageNumber="171">
|
||
Collected from richly vegetated ponds in full sun or slightly shaded, in abundance from flooded mats of grasses (
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitat of Rhantus bogotensis sp. nov. in the Altiplano of Bogota, Colombia. Florida wetland, deeper water bodies where we did encounter only very few specimens (A), shallow ponds, most likely ephemeral (B, C)." figureDoi="10.3897/alpento.3.37308.figure4" httpUri="https://binary.pensoft.net/fig/349404" pageId="2" pageNumber="171" tableDoi="10.3897/alpento.3.37308.figure4">Fig. 4B, C</figureCitation>
|
||
). Bottom of these habitats was usually black, foul smelling mud. In the La Florida wetland, the beetles seemed to avoid larger, deeper water bodies, possibly to avoid predation by fish (
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitat of Rhantus bogotensis sp. nov. in the Altiplano of Bogota, Colombia. Florida wetland, deeper water bodies where we did encounter only very few specimens (A), shallow ponds, most likely ephemeral (B, C)." figureDoi="10.3897/alpento.3.37308.figure4" httpUri="https://binary.pensoft.net/fig/349404" pageId="2" pageNumber="171" tableDoi="10.3897/alpento.3.37308.figure4">Fig. 4A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="171" type="notes">
|
||
<paragraph pageId="2" pageNumber="171">Notes.</paragraph>
|
||
<paragraph pageId="2" pageNumber="171">
|
||
Collected with aquatic nets as well as bottle traps. In association with the following other
|
||
<taxonomicName family="Dytiscidae" lsidName="" pageId="2" pageNumber="171" rank="family">Dytiscidae</taxonomicName>
|
||
:
|
||
<taxonomicName genus="R." lsidName="R. andinus" pageId="2" pageNumber="171" rank="species" species="andinus">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">R. andinus</emphasis>
|
||
</taxonomicName>
|
||
(abundant),
|
||
<taxonomicName genus="R." lsidName="R. vicinus" pageId="2" pageNumber="171" rank="species" species="vicinus">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">R. vicinus</emphasis>
|
||
</taxonomicName>
|
||
(only few, this species is abundant on higher elevations as well as in the
|
||
<normalizedToken originalValue="Páramos">Paramos</normalizedToken>
|
||
above the altiplano of Bogota), two species of
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">Copelatus</emphasis>
|
||
</taxonomicName>
|
||
Erichson, 1832 and
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Liodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liodessus bogotensis" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="bogotensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">Liodessus bogotensis</emphasis>
|
||
</taxonomicName>
|
||
Guignot, 1953, as well as different
|
||
<taxonomicName family="Hydrophilidae" lsidName="" pageId="2" pageNumber="171" rank="family">Hydrophilidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption doi="10.3897/alpento.3.37308.figure1" httpUri="https://binary.pensoft.net/fig/349401" pageId="2" pageNumber="171" start="Figure 1" startId="F1">
|
||
<paragraph pageId="2" pageNumber="171">
|
||
<emphasis bold="true" pageId="2" pageNumber="171">Figure 1.</emphasis>
|
||
Dorsal habitus of
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus andinus" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="andinus">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">Rhantus andinus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis bold="true" pageId="2" pageNumber="171">A</emphasis>
|
||
),
|
||
<taxonomicName genus="R." lsidName="R. bogotensis" pageId="2" pageNumber="171" rank="species" species="bogotensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">R. bogotensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">B</emphasis>
|
||
),
|
||
<taxonomicName genus="R." lsidName="R. vicinus" pageId="2" pageNumber="171" rank="species" species="vicinus">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">R. vicinus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis bold="true" pageId="2" pageNumber="171">C</emphasis>
|
||
),
|
||
<taxonomicName genus="R." lsidName="R. franzi" pageId="2" pageNumber="171" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">R. franzi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis bold="true" pageId="2" pageNumber="171">D</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/alpento.3.37308.figure2" httpUri="https://binary.pensoft.net/fig/349402" pageId="2" pageNumber="171" start="Figure 2" startId="F2">
|
||
<paragraph pageId="2" pageNumber="171">
|
||
<emphasis bold="true" pageId="2" pageNumber="171">Figure 2.</emphasis>
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">Rhantus</emphasis>
|
||
</taxonomicName>
|
||
spp.: Pronotal margin of
|
||
<taxonomicName genus="R." lsidName="R. bogotensis" pageId="2" pageNumber="171" rank="species" species="bogotensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">R. bogotensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. female (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">A</emphasis>
|
||
), male (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">B</emphasis>
|
||
);
|
||
<taxonomicName genus="R." lsidName="R. vicinus" pageId="2" pageNumber="171" rank="species" species="vicinus">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">R. vicinus</emphasis>
|
||
</taxonomicName>
|
||
female (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">C</emphasis>
|
||
);
|
||
<taxonomicName genus="R." lsidName="R. bogotensis" pageId="2" pageNumber="171" rank="species" species="bogotensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">R. bogotensis</emphasis>
|
||
</taxonomicName>
|
||
: surface sculpture on head, frons (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">D</emphasis>
|
||
), middle part of pronotum with base and disc (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">E</emphasis>
|
||
) elytron, basal area (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">F</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/alpento.3.37308.figure3" httpUri="https://binary.pensoft.net/fig/349403" pageId="2" pageNumber="171" start="Figure 3" startId="F3">
|
||
<paragraph pageId="2" pageNumber="171">
|
||
<emphasis bold="true" pageId="2" pageNumber="171">Figure 3.</emphasis>
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">Rhantus</emphasis>
|
||
</taxonomicName>
|
||
spp., males:
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus bogotensis" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="bogotensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">Rhantus bogotensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. parameres (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">A</emphasis>
|
||
), median lobe of aedeagus in lateral view (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">B</emphasis>
|
||
), same in ventral view (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">C</emphasis>
|
||
), mesotarsus (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">D</emphasis>
|
||
), mesotarsomere V and claws (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">E</emphasis>
|
||
), foretarsus (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">F</emphasis>
|
||
), protarsomere V and claws (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">G</emphasis>
|
||
);
|
||
<taxonomicName genus="R." lsidName="R. vicinus" pageId="2" pageNumber="171" rank="species" species="vicinus">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">R. vicinus</emphasis>
|
||
</taxonomicName>
|
||
: protarsomere V and claws (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">H</emphasis>
|
||
);
|
||
<taxonomicName genus="R." lsidName="R. andinus" pageId="2" pageNumber="171" rank="species" species="andinus">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">R. andinus</emphasis>
|
||
</taxonomicName>
|
||
: protarsomere V and claws (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">I</emphasis>
|
||
);
|
||
<taxonomicName genus="R." lsidName="R. crypticus" pageId="2" pageNumber="171" rank="species" species="crypticus">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">R. crypticus</emphasis>
|
||
</taxonomicName>
|
||
: protarsomere V and claws (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">J</emphasis>
|
||
);
|
||
<taxonomicName genus="R." lsidName="R. franzi" pageId="2" pageNumber="171" rank="species" species="franzi">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">R. franzi</emphasis>
|
||
</taxonomicName>
|
||
median lobe of aedeagus in lateral view (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">K</emphasis>
|
||
), protarsomere
|
||
<normalizedToken originalValue="III–V">III-V</normalizedToken>
|
||
and claws (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">L</emphasis>
|
||
), mesotarsus (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">M</emphasis>
|
||
), mesotarsomere V and anterior claw, posterior claw missing (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">N</emphasis>
|
||
).
|
||
<normalizedToken originalValue="“a”">"a"</normalizedToken>
|
||
denotes anterior claw.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/alpento.3.37308.figure4" httpUri="https://binary.pensoft.net/fig/349404" pageId="2" pageNumber="171" start="Figure 4" startId="F4">
|
||
<paragraph pageId="2" pageNumber="171">
|
||
<emphasis bold="true" pageId="2" pageNumber="171">Figure 4.</emphasis>
|
||
Habitat of
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Rhantus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhantus bogotensis" order="Coleoptera" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="bogotensis">
|
||
<emphasis italics="true" pageId="2" pageNumber="171">Rhantus bogotensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in the Altiplano of Bogota, Colombia. Florida wetland, deeper water bodies where we did encounter only very few specimens (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">A</emphasis>
|
||
), shallow ponds, most likely ephemeral (
|
||
<emphasis bold="true" pageId="2" pageNumber="171">B, C</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |