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<document id="DD7501D14E99C47ACC0B516B0C9227ED" ID-CLB-Dataset="4488" ID-DOI="10.11646/phytotaxa.480.2.2" ID-GBIF-Dataset="702b3b1f-bd77-4ce5-a439-32b6d953cbac" ID-ISSN="1179-3163" ID-Zenodo-Dep="5415987" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630358620745" checkinUser="marcus" docAuthor="Ranjbar, Massoud, Askari, Ali &amp; Mohammadi, Somayeh" docDate="2021" docId="836F1A24FFCFFFA4FF0BFF68FDF4F947" docLanguage="en" docName="phytotaxa.480.2.2.pdf" docOrigin="Phytotaxa 480 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.480.2.2" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Centaurea kuhdashtensis Ranjbar &amp; Mohamadi 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="132" masterDocId="7F56625CFFC8FFA3FF83FFF6FF97FFE7" masterDocTitle="Two new species of Centaurea sect. Microlophus (Asteraceae, Cardueae) from Iran" masterLastPageNumber="138" masterPageNumber="125" pageNumber="132" updateTime="1699252135494" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="119A8522F47D033D381EB10C7BF1237F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FC3134D8CDC85DF794C2DC346287396B">
<mods:title id="64985C7530A6625F5CF4C72EFE81A33A">Two new species of Centaurea sect. Microlophus (Asteraceae, Cardueae) from Iran</mods:title>
</mods:titleInfo>
<mods:name id="5CAB7E6985A3D5E991092BE605D68599" type="personal">
<mods:role id="A6D4091BB14C9DDC9E25854C387C405C">
<mods:roleTerm id="73686B2694E10F174B44457931703DB9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0E2933B066ED1C7F2BFF400265498960">Ranjbar, Massoud</mods:namePart>
<mods:nameIdentifier id="C7A7CC7D8A277C4D9EF5D5CB8C0F6031" type="ORCID">0000-0003-0939-3878</mods:nameIdentifier>
<mods:affiliation id="5C8DB4790991F79C77E51CCBF83D78AA">Department of Biology, Herbarium Division, Bu-Ali Sina University, P. O. Box 65175 / 4111, Hamedan, Iran. &amp; ranjbar @ basu. ac. ir; https: // orcid. org / 0000 - 0003 - 0939 - 3878</mods:affiliation>
<mods:nameIdentifier id="9B1462FE28D85AE597FA5373E933FB55" type="email">ranjbar@basu.ac.ir</mods:nameIdentifier>
</mods:name>
<mods:name id="0D2425058845E954E0D16CC1BB6F01AA" type="personal">
<mods:role id="FBD8570033D9C8202125672CF55DF953">
<mods:roleTerm id="3780571F9560924CD4486DEA939578BA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AD446D22A990287ECFBD794AA3CCE942">Askari, Ali</mods:namePart>
<mods:nameIdentifier id="FA03AE170887DBD844A6401A31E3EAAD" type="ORCID">0000-0002-4043-6306</mods:nameIdentifier>
<mods:affiliation id="D82393964A3245FBCCDDD14FC6BE7A71">Department of Biology, Herbarium Division, Bu-Ali Sina University, P. O. Box 65175 / 4111, Hamedan, Iran. &amp; aliiiiaskari 34 @ gmail. com; https: // orcid. org / 0000 - 0002 - 4043 - 6306</mods:affiliation>
<mods:nameIdentifier id="7C5B975A0F7C7B9A95D61A713CEDB917" type="email">aliiiiaskari34@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="B3B1A54C814CCA501DBFF4847B369EE1" type="personal">
<mods:role id="DB349F6366A9DD9961690E5FB9941042">
<mods:roleTerm id="98367FE60B96FA1ECBE830B6514689FB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8DFDDFFA1274F9C1B313E54ABCEB0BDA">Mohammadi, Somayeh</mods:namePart>
<mods:nameIdentifier id="2C40C987F205D60BAE327FFD59DB0075" type="ORCID">0000-0001-8925-5824</mods:nameIdentifier>
<mods:affiliation id="17DAC4D35255D9E69E90E5BFB02A9F52">Department of Biology, Herbarium Division, Bu-Ali Sina University, P. O. Box 65175 / 4111, Hamedan, Iran. &amp; s. mohamadi 6868 @ gmail. com; https: // orcid. org / 0000 - 0001 - 8925 - 5824</mods:affiliation>
<mods:nameIdentifier id="9D8B0374B32606DDD3C432978311AE6B" type="email">s.mohamadi6868@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="14B9636DC6B0D5304FE28D38AFA196F1">text</mods:typeOfResource>
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<mods:titleInfo id="F38BE35C0274C5FBF7038B7FCDB146A1">
<mods:title id="E16C467F4F93D00A75BDBF541937FE52">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="EE9CB3D60180BF352FE88C9342C4BB66">
<mods:date id="EA38790D23BF0C9D9AECFAA8325ABF44">2021</mods:date>
<mods:detail id="13178B993CC60625C7FC9E37D34F8494" type="pubDate">
<mods:number id="8FF27DD76AEC1B5D281469F9B2C09D5A">2021-01-20</mods:number>
</mods:detail>
<mods:detail id="68A9702A553755A901EC3E00F4D769AE" type="volume">
<mods:number id="F6D624308517A03F723B630F353CAC5C">480</mods:number>
</mods:detail>
<mods:detail id="8A4741618E70AFEFB8E95D0A7AF5A200" type="issue">
<mods:number id="4C0AABB90377ADDD881C24B2448E865A">2</mods:number>
</mods:detail>
<mods:extent id="411E864A92324397043CE62FD56462ED" unit="page">
<mods:start id="AC011CC7D1BC4A31264ED712594D9934">125</mods:start>
<mods:end id="F2A178633591294F3C4D93E02810D9EF">138</mods:end>
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<mods:location id="9C75D5A9557CE64D555FBA486E5A2256">
<mods:url id="F2B98A1C38F4FEE68209228EC628E747">http://dx.doi.org/10.11646/phytotaxa.480.2.2</mods:url>
</mods:location>
<mods:classification id="4B1429BA9CF96E09F2C72FE4D4D6CDA1">journal article</mods:classification>
<mods:identifier id="0B26E5C03BD1A4CB29A62CA1D93B2E6D" type="CLB-Dataset">4488</mods:identifier>
<mods:identifier id="01C82F4BF61CD496A5C38C98309B2C92" type="DOI">10.11646/phytotaxa.480.2.2</mods:identifier>
<mods:identifier id="C8F4A1A35D31A306A9D33D0C3D4A5125" type="GBIF-Dataset">702b3b1f-bd77-4ce5-a439-32b6d953cbac</mods:identifier>
<mods:identifier id="81A3196086BA431A0114DEEDC63C7A97" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="611BA6A841F5FAA5F049E56EDAF36A53" type="Zenodo-Dep">5415987</mods:identifier>
</mods:mods>
<treatment id="836F1A24FFCFFFA4FF0BFF68FDF4F947" ID-DOI="http://doi.org/10.5281/zenodo.5916074" ID-GBIF-Taxon="185389039" ID-Zenodo-Dep="5916074" LSID="urn:lsid:plazi:treatment:836F1A24FFCFFFA4FF0BFF68FDF4F947" httpUri="http://treatment.plazi.org/id/836F1A24FFCFFFA4FF0BFF68FDF4F947" lastPageNumber="132" pageId="7" pageNumber="132">
<subSubSection id="43DCF8B9FFCFFFA4FF0BFF68FC35FF5F" box="[136,930,158,185]" pageId="7" pageNumber="132" type="nomenclature">
<paragraph id="0B79AB32FFCFFFA4FF0BFF68FC35FF5F" blockId="7.[136,930,158,185]" box="[136,930,158,185]" pageId="7" pageNumber="132">
<heading id="50311C5EFFCFFFA4FF0BFF68FC35FF5F" box="[136,930,158,185]" fontSize="11" level="2" pageId="7" pageNumber="132" reason="4">
<emphasis id="39B27720FFCFFFA4FF0BFF68FF0AFF5F" bold="true" box="[136,157,158,184]" pageId="7" pageNumber="132">2.</emphasis>
<taxonomicName id="CCC6D0B1FFCFFFA4FF27FF68FD51FF5F" ID-CoL="8478Z" authority="Ranjbar &amp; Mohamadi" authorityName="Ranjbar &amp; Mohamadi" authorityYear="2021" box="[164,710,158,185]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="132" phylum="Tracheophyta" rank="species" species="kuhdashtensis" status="sp. nov.">
<emphasis id="39B27720FFCFFFA4FF27FF68FE5EFF5E" bold="true" box="[164,457,158,185]" italics="true" pageId="7" pageNumber="132">Centaurea kuhdashtensis</emphasis>
Ranjbar &amp; Mohamadi
</taxonomicName>
,
<emphasis id="39B27720FFCFFFA4FD57FF69FCBDFF5E" bold="true" box="[724,810,159,185]" italics="true" pageId="7" pageNumber="132">
<taxonomicNameLabel id="2281CA5BFFCFFFA4FD57FF69FCBDFF5E" box="[724,810,159,185]" pageId="7" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<figureCitation id="93FDB7B7FFCFFFA4FCB1FF68FC0AFF5F" box="[818,925,158,184]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStartId-0="10.[136,229,2013,2035]" captionStartId-1="11.[136,229,2013,2035]" captionStartId-2="12.[136,229,2013,2035]" captionTargetBox-0="[151,1436,190,1989]" captionTargetBox-1="[151,1436,190,1989]" captionTargetBox-2="[151,1436,190,1989]" captionTargetId-0="figure-27@10.[151,1436,190,1989]" captionTargetId-1="figure-18@11.[151,1436,190,1989]" captionTargetId-2="figure-27@12.[151,1436,190,1989]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURE 7. Centaurea kuhdashtensis (M. Ranjbar 33645, BASU). Scale = 5 cm." captionText-1="FIGURE 8. Centaurea kuhdashtensis (M. Ranjbar 33645, BASU). A1, close up of capitula, B1–B3, outer phyllaries, C1–C3, median phyllaries, D1–D3, inner phyllaries, E, leaf stomata, F and G, leaf indumentum. Scale A = 1 cm." captionText-2="FIGURE 9. Measured morphological characters in Centaurea kuhdashtensis (M. Ranjbar 33645, BASU). A, middle stem leaf, B, central floret, C, involucre, D, involucral bract. For character explanations see Table 1." figureDoi-0="http://doi.org/10.5281/zenodo.5416003" figureDoi-1="http://doi.org/10.5281/zenodo.5416007" figureDoi-2="http://doi.org/10.5281/zenodo.5416009" httpUri-0="https://zenodo.org/record/5416003/files/figure.png" httpUri-1="https://zenodo.org/record/5416007/files/figure.png" httpUri-2="https://zenodo.org/record/5416009/files/figure.png" pageId="7" pageNumber="132">Figs. 7–9</figureCitation>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="43DCF8B9FFCFFFA4FF0BFF10FB19FE3E" pageId="7" pageNumber="132" type="diagnosis">
<paragraph id="0B79AB32FFCFFFA4FF0BFF10FB19FE3E" blockId="7.[136,1452,230,1697]" pageId="7" pageNumber="132">
<emphasis id="39B27720FFCFFFA4FF0BFF10FF6CFEE7" bold="true" box="[136,251,230,256]" pageId="7" pageNumber="132">Diagnosis</emphasis>
:—
<taxonomicName id="CCC6D0B1FFCFFFA4FE9CFF11FDABFEE7" authorityName="Ranjbar &amp; Mohamadi" authorityYear="2021" box="[287,572,230,257]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="132" phylum="Tracheophyta" rank="species" species="kuhdashtensis">
<emphasis id="39B27720FFCFFFA4FE9CFF11FDABFEE7" box="[287,572,230,257]" italics="true" pageId="7" pageNumber="132">Centaurea kuhdashtensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="CCC6D0B1FFCFFFA4FD5CFF11FCFFFEE7" box="[735,872,230,257]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="132" phylum="Tracheophyta" rank="species" species="koeieana">
<emphasis id="39B27720FFCFFFA4FD5CFF11FCFFFEE7" box="[735,872,230,257]" italics="true" pageId="7" pageNumber="132">C. koeieana</emphasis>
</taxonomicName>
but differs by hispid (vs. arachnoid) hairs, upper cauline leaves decurrent for
<quantity id="CC3E06D7FFCFFFA4FE4EFEFCFDADFEC2" box="[461,570,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="7" pageNumber="132" unit="mm" value="6.5" valueMax="10.0" valueMin="3.0">3–10 mm</quantity>
(vs.
<quantity id="CC3E06D7FFCFFFA4FDF7FEFCFD40FEC2" box="[628,727,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="7" pageNumber="132" unit="mm" value="4.5" valueMax="7.0" valueMin="2.0">2–7 mm</quantity>
), lower cauline leaves 35–38 ×
<quantity id="CC3E06D7FFCFFFA4FBC6FEFCFB5DFEC2" box="[1093,1226,266,293]" metricMagnitude="-1" metricUnit="m" metricValue="1.075" metricValueMax="1.15" metricValueMin="1.0" pageId="7" pageNumber="132" unit="cm" value="10.75" valueMax="11.5" valueMin="10.0">10–11.5 cm</quantity>
(vs. 15–17 ×
<quantity id="CC3E06D7FFCFFFA4FAEFFEFCFF39FEAE" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="7" pageNumber="132" unit="cm" value="2.75" valueMax="3.5" valueMin="2.0">2–3.5 cm</quantity>
), median [lower] cauline leaves pinnatipartite- pinnatilobate or pinnatifid (vs. lanceolate or lanceolate-oblong), margin entire-dentate (vs. irregularly dentate or entire), segments lower cauline leaves 15–17 pairs (vs. no segment), basal cauline leaves 44–46 ×
<quantity id="CC3E06D7FFCFFFA4FE59FE80FDDCFE76" box="[474,587,374,401]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="7" pageNumber="132" unit="cm" value="14.0" valueMax="15.0" valueMin="13.0">13–15 cm</quantity>
(vs. 15.5–17 ×
<quantity id="CC3E06D7FFCFFFA4FC81FE80FCCCFE76" box="[770,859,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="7" pageNumber="132" unit="cm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 cm</quantity>
), basal cauline leaves pinnatipartite-pinnatilobate or pinnatifid (vs. oblong lyrate or pinnatilobate), basal cauline leaves 14–15 pairs (vs. 1–2), margin basal cauline leaves irregular dentate (vs. dentate or lobate), involucres 10–16 ×
<quantity id="CC3E06D7FFCFFFA4FCAAFE48FC11FE3E" box="[809,902,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="7" pageNumber="132" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
(vs. 15–18 ×
<quantity id="CC3E06D7FFCFFFA4FBA2FE48FB15FE3E" box="[1057,1154,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="7" pageNumber="132" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="43DCF8B9FFCFFFA4FF3EFE14FC9FFB17" pageId="7" pageNumber="132" type="materials_examined">
<paragraph id="0B79AB32FFCFFFA4FF3EFE14FC31FDC7" blockId="7.[136,1452,230,1697]" pageId="7" pageNumber="132">
<emphasis id="39B27720FFCFFFA4FF3EFE14FF6FFE1B" bold="true" box="[189,248,482,508]" pageId="7" pageNumber="132">Type</emphasis>
:—
<materialsCitation id="BBAEA16FFFCFFFA4FE9FFE14FD81FDC7" ID-GBIF-Occurrence="3340970302" collectingDate="2013-05-01" collectionCode="M" country="Iran" county="Bolouran" elevation="1504" latitude="33.55611" location="Baba Zeyd" longLatPrecision="20" longitude="47.300278" municipality="Baba Zeyd" pageId="7" pageNumber="132" specimenCount="1" stateProvince="Lorestan">
<collectingCountry id="73D1EBA2FFCFFFA4FE9FFE14FEF7FE1B" box="[284,352,482,508]" name="Iran" pageId="7" pageNumber="132">IRAN</collectingCountry>
.
<collectingRegion id="C90265D0FFCFFFA4FEECFE15FE43FE1A" box="[367,468,483,509]" country="Iran" name="Lorestan" pageId="7" pageNumber="132">Lorestan</collectingRegion>
,
<collectingCounty id="E218D3BEFFCFFFA4FE63FE14FDDFFE1B" box="[480,584,482,508]" pageId="7" pageNumber="132">Bolouran</collectingCounty>
to
<collectingMunicipality id="EB1D3148FFCFFFA4FDEEFE14FD7CFE1B" box="[621,747,482,508]" pageId="7" pageNumber="132">Baba Zeyd</collectingMunicipality>
,
<quantity id="CC3E06D7FFCFFFA4FD74FE14FCA8FE1B" box="[759,831,482,509]" metricMagnitude="4" metricUnit="m" metricValue="6.5" pageId="7" pageNumber="132" unit="km" value="65.0">65 km</quantity>
to
<location id="0E19FDE9FFCFFFA4FCE7FE14FC76FE1B" LSID="urn:lsid:plazi:treatment:836F1A24FFCFFFA4FF0BFF68FDF4F947:0E19FDE9FFCFFFA4FCE7FE14FC76FE1B" box="[868,993,482,508]" country="Iran" county="Bolouran" latitude="33.55611" longLatPrecision="20" longitude="47.300278" municipality="Baba Zeyd" name="Baba Zeyd" pageId="7" pageNumber="132" stateProvince="Lorestan">Baba Zeyd</location>
,
<quantity id="CC3E06D7FFCFFFA4FC6DFE14FBA2FE1B" box="[1006,1077,482,509]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="7" pageNumber="132" unit="km" value="35.0">35 km</quantity>
to
<location id="0E19FDE9FFCFFFA4FBD8FE14FB5EFE1B" LSID="urn:lsid:plazi:treatment:836F1A24FFCFFFA4FF0BFF68FDF4F947:0E19FDE9FFCFFFA4FBD8FE14FB5EFE1B" box="[1115,1225,482,508]" country="Iran" county="Bolouran" latitude="33.55611" longLatPrecision="20" longitude="47.300278" municipality="Baba Zeyd" name="Kuhdasht" pageId="7" pageNumber="132" stateProvince="Lorestan">Kuhdasht</location>
,
<geoCoordinate id="6EF2CDF5FFCFFFA4FB56FE14FAEFFE1A" box="[1237,1400,482,509]" degrees="33" direction="north" minutes="33" orientation="latitude" pageId="7" pageNumber="132" precision="15" seconds="22" value="33.55611">33° 33’ 22’’ N</geoCoordinate>
,
<geoCoordinate id="6EF2CDF5FFCFFFA4FA07FE14FF60FDC6" degrees="47" direction="east" minutes="18" orientation="longitude" pageId="7" pageNumber="132" precision="15" seconds="01" value="47.300278">47° 18’ 01’’ E</geoCoordinate>
,
<quantity id="CC3E06D7FFCFFFA4FE80FDF0FECDFDC6" box="[259,346,518,545]" metricMagnitude="3" metricUnit="m" metricValue="1.504" pageId="7" pageNumber="132" unit="m" value="1504.0">
<elevation id="80EB4C01FFCFFFA4FE80FDF0FECDFDC6" box="[259,346,518,545]" metricMagnitude="3" metricUnit="m" metricValue="1.504" pageId="7" pageNumber="132" unit="m" value="1504.0">1504 m</elevation>
</quantity>
,
<date id="7F788DF2FFCFFFA4FEE5FDF0FE67FDC7" box="[358,496,518,545]" pageId="7" pageNumber="132" value="2013-05-01">
<collectingDate id="6F3C741AFFCFFFA4FEE5FDF0FE67FDC7" box="[358,496,518,545]" pageId="7" pageNumber="132" value="2013-05-01">1 May 2013</collectingDate>
</date>
,
<collectionCode id="6DD733F7FFCFFFA4FE78FDF1FD81FDC7" box="[507,534,519,544]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="7" pageNumber="132" type="Herbarium">
<emphasis id="39B27720FFCFFFA4FE78FDF1FD81FDC7" box="[507,534,519,544]" italics="true" pageId="7" pageNumber="132">M</emphasis>
</collectionCode>
</materialsCitation>
<emphasis id="39B27720FFCFFFA4FD95FDF1FD8DFDC7" box="[534,538,519,544]" italics="true" pageId="7" pageNumber="132">.</emphasis>
<materialsCitation id="BBAEA16FFFCFFFA4FDA3FDF0FC35FDC7" ID-GBIF-Occurrence="3340970304" box="[544,930,518,545]" collectionCode="BASU" country="Iran" location="Ranjbar" pageId="7" pageNumber="132" specimenCount="1" stateProvince="Lorestan" typeStatus="holotype">
<emphasis id="39B27720FFCFFFA4FDA3FDF0FDEBFDC7" box="[544,636,518,544]" italics="true" pageId="7" pageNumber="132">
<location id="0E19FDE9FFCFFFA4FDA3FDF0FDEBFDC7" LSID="urn:lsid:plazi:treatment:836F1A24FFCFFFA4FF0BFF68FDF4F947:0E19FDE9FFCFFFA4FDA3FDF0FDEBFDC7" box="[544,636,518,544]" country="Iran" name="Ranjbar" pageId="7" pageNumber="132" stateProvince="Lorestan">Ranjbar</location>
</emphasis>
33645 (
<typeStatus id="D47D1590FFCFFFA4FD5BFDF0FCD7FDC7" box="[728,832,518,544]" pageId="7" pageNumber="132" type="holotype">holotype</typeStatus>
:
<collectionCode id="6DD733F7FFCFFFA4FCC9FDF0FC0DFDC7" box="[842,922,518,544]" country="Iran" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15764" lsid="urn:lsid:biocol.org:col:15764" name="Bu-Ali Sina University" pageId="7" pageNumber="132" type="Herbarium">BASU</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="0B79AB32FFCFFFA4FF3EFDDCFC9FFB17" blockId="7.[136,1452,230,1697]" pageId="7" pageNumber="132">
Perennial plants with thickened rootstock, usually completely green up to
<quantity id="CC3E06D7FFCFFFA4FB8CFDDCFBC1FDA2" box="[1039,1110,554,581]" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="132" unit="cm" value="80.0">80 cm</quantity>
tall, in the lower part loosely covered with hispid hairs. Stem erect, often branched above, sometimes branched in the median to upper parts striate striped, many branched, usually striped,
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in diam. at base, cylindrical. Sometimes ascending, branched often at the base, or branched in median part, thinly white striate, rigid densely leafy throughout. Leaves and stem loosely hispid, venation actinodromous at basal and lower stem leaves. Basal cauline leaves pinnatipartite, pinnatilobate or pinnatifid, 44–46 ×
<quantity id="CC3E06D7FFCFFFA4FF7BFD28FEE6FD1E" box="[248,369,734,761]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="7" pageNumber="132" unit="cm" value="14.0" valueMax="15.0" valueMin="13.0">13–15 cm</quantity>
, terminal segments larger, lanceolate-linear, lateral segments 14–15 pairs, lanceolate-linear, segments sometimes lobed near the base, acute at apex, margin irregular dentate. Lower cauline leaves pinnatipartite to pinnately divided, odd pinnate in outline, 35–38 ×
<quantity id="CC3E06D7FFCFFFA4FD5EFCD0FCF4FCA6" box="[733,867,806,833]" metricMagnitude="-1" metricUnit="m" metricValue="1.075" metricValueMax="1.15" metricValueMin="1.0" pageId="7" pageNumber="132" unit="cm" value="10.75" valueMax="11.5" valueMin="10.0">10–11.5 cm</quantity>
, terminal segments larger, oblong-lanceolate, lateral segments 15–17 pairs, oblong-lanceolate, segments sometimes lobed near base, acute at apex, margin dentate. Median [lower] cauline leaves pinnatipartite, pinnatilobate or pinnatifid, odd pinnate in outline, 10–13 ×
<quantity id="CC3E06D7FFCFFFA4FB34FC98FA9BFC6E" box="[1207,1292,878,905]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="7" pageNumber="132" unit="cm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 cm</quantity>
, margin entire dentate, decurrent for
<quantity id="CC3E06D7FFCFFFA4FE07FC64FD96FC4A" box="[388,513,914,941]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="7" pageNumber="132" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 mm</quantity>
, margin dentate, venation reticulate. Median [upper] cauline leaves sessile, simple oblong-lanceolate, 9–10.5 ×
<quantity id="CC3E06D7FFCFFFA4FE46FC40FDD3FC36" box="[453,580,950,977]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="7" pageNumber="132" unit="cm" value="0.85" valueMax="0.9" valueMin="0.8">0.8–0.9 cm</quantity>
, acute to acuminate at apex, decurrent for
<quantity id="CC3E06D7FFCFFFA4FB97FC40FB19FC36" box="[1044,1166,950,977]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="7" pageNumber="132" unit="mm" value="45.0" valueMax="50.0" valueMin="40.0">40–50 mm</quantity>
, perfoliate, margin entire. Upper cauline leaves sessile, simple, oblong to oblong-lanceolate in outline, 1–3 ×
<quantity id="CC3E06D7FFCFFFA4FBB5FC2CFB20FC12" box="[1078,1207,986,1013]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="7" pageNumber="132" unit="cm" value="0.4" valueMax="0.5" valueMin="0.3">0.3–0.5 cm</quantity>
, shortly decurrent for
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, acuminate at apex, venation reticulate, margin entire. Involucres conical contracted towards top, diamond, 10–16 ×
<quantity id="CC3E06D7FFCFFFA4FF68FBD4FEDCFBDA" box="[235,331,1058,1085]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="7" pageNumber="132" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
. Phyllaries multiseriate, imbricate, coriaceous, yellowish or green, hairy. Outer phyllaries ovate,
<quantity id="CC3E06D7FFCFFFA4FA02FBD4FF23FB86" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="7" pageNumber="132" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
long. Median phyllaries ovate to oblong
<quantity id="CC3E06D7FFCFFFA4FDFFFBB0FD4FFB86" box="[636,728,1094,1121]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="7" pageNumber="132" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
long, spine
<quantity id="CC3E06D7FFCFFFA4FCDDFBB0FC08FB86" box="[862,927,1094,1121]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="132" unit="mm" value="1.0">1 mm</quantity>
long. Inner phyllaries oblong,
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long. Appendages forming a minute deciduous tip, not decurrent, outer phyllaries deciduous, inner phyllaries persistent, always small, not concealing the basal part of phyllaries. Flowers yellowish-green; central flowers hermaphrodite, up to
<quantity id="CC3E06D7FFCFFFA4FF25FB44FF60FB2A" box="[166,247,1202,1229]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="132" unit="mm" value="14.0">14 mm</quantity>
long; peripheral sterile, not radiant, limb lobes linear. Achene oblong,
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long. Pappus multiseriate, scabrous, the outer
<quantity id="CC3E06D7FFCFFFA4FEE1FB20FE28FB16" box="[354,447,1238,1265]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="7" pageNumber="132" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3–6 mm</quantity>
long, the inner much shorter.
</paragraph>
</subSubSection>
<subSubSection id="43DCF8B9FFCFFFA4FF3EFB0CFC8EFADE" pageId="7" pageNumber="132" type="etymology">
<paragraph id="0B79AB32FFCFFFA4FF3EFB0CFACBFAF3" blockId="7.[136,1452,230,1697]" box="[189,1372,1274,1301]" pageId="7" pageNumber="132">
<emphasis id="39B27720FFCFFFA4FF3EFB0CFED0FAF3" bold="true" box="[189,327,1274,1300]" pageId="7" pageNumber="132">Etymology:</emphasis>
—The specific epithet refers to Kuhdasht city, near which the
<typeStatus id="D47D1590FFCFFFA4FC7CFB0DFBB8FAF2" box="[1023,1071,1275,1301]" pageId="7" pageNumber="132">type</typeStatus>
specimens were collected.
</paragraph>
<paragraph id="0B79AB32FFCFFFA4FF3EFAE8FC8EFADE" blockId="7.[136,1452,230,1697]" box="[189,793,1310,1337]" pageId="7" pageNumber="132">
<emphasis id="39B27720FFCFFFA4FF3EFAE8FED4FADF" bold="true" box="[189,323,1310,1336]" pageId="7" pageNumber="132">Phenology:</emphasis>
—Flowering March (second half) to May.
</paragraph>
</subSubSection>
<subSubSection id="43DCF8B9FFCFFFA4FF3EFAB4FDF4F947" pageId="7" pageNumber="132" type="distribution">
<paragraph id="0B79AB32FFCFFFA4FF3EFAB4FE65FA42" blockId="7.[136,1452,230,1697]" pageId="7" pageNumber="132">
<emphasis id="39B27720FFCFFFA4FF3EFAB4FE7EFABB" bold="true" box="[189,489,1346,1372]" pageId="7" pageNumber="132">Distribution and habitat:</emphasis>
—
<taxonomicName id="CCC6D0B1FFCFFFA4FD86FAB5FC89FABB" authorityName="Ranjbar &amp; Mohamadi" authorityYear="2021" box="[517,798,1346,1373]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="132" phylum="Tracheophyta" rank="species" species="kuhdashtensis">
<emphasis id="39B27720FFCFFFA4FD86FAB5FC89FABB" box="[517,798,1346,1373]" italics="true" pageId="7" pageNumber="132">Centaurea kuhdashtensis</emphasis>
</taxonomicName>
occurs in Kuhdasht area, a mountainous region in western
<collectingRegion id="C90265D0FFCFFFA4FF0BFA91FEC0FA67" box="[136,343,1382,1409]" country="Iran" name="Lorestan" pageId="7" pageNumber="132">Lorestan province</collectingRegion>
,
<collectingCountry id="73D1EBA2FFCFFFA4FEE1FA91FE04FA66" box="[354,403,1383,1409]" name="Iran" pageId="7" pageNumber="132">Iran</collectingCountry>
. Phytogeographically it is an Irano-Turanian element and grows in dry clay, stony slopes and screes, at
<quantity id="CC3E06D7FFCFFFA4FF75FA7CFE06FA42" box="[246,401,1418,1445]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="7" pageNumber="132" unit="m" value="1500.0" valueMax="1600.0" valueMin="1400.0">1400–1600 m</quantity>
(
<figureCitation id="93FDB7B7FFCFFFA4FE23FA7CFE73FA42" box="[416,484,1418,1445]" captionStart="FIGURE 6" captionStartId="9.[136,229,1675,1697]" captionTargetBox="[174,1387,752,1625]" captionTargetId="figure-114@9.[151,1436,742,1651]" captionTargetPageId="9" captionText="FIGURE 6. Distribution of Centaurea in Iran. A, C. dezfulica, B, C. pabotii, C, C. kuhdashtensis and D, C. koeieana." figureDoi="http://doi.org/10.5281/zenodo.5416001" httpUri="https://zenodo.org/record/5416001/files/figure.png" pageId="7" pageNumber="132">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="0B79AB32FFCFFFA4FF3EFA58FC77F9D3" blockId="7.[136,1452,230,1697]" pageId="7" pageNumber="132">
<emphasis id="39B27720FFCFFFA4FF3EFA58FE7BFA2F" bold="true" box="[189,492,1454,1480]" pageId="7" pageNumber="132">IUCN Red List category:</emphasis>
—
<taxonomicName id="CCC6D0B1FFCFFFA4FD8BFA59FCB4FA2F" authorityName="Ranjbar &amp; Mohamadi" authorityYear="2021" box="[520,803,1454,1481]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="132" phylum="Tracheophyta" rank="species" species="kuhdashtensis">
<emphasis id="39B27720FFCFFFA4FD8BFA59FCB4FA2F" box="[520,803,1454,1481]" italics="true" pageId="7" pageNumber="132">Centaurea kuhdashtensis</emphasis>
</taxonomicName>
is known from a few small populations, consisting of no more than five plants, observed in the mountains between Bolouran and Baba Zeyd village (M. Ranjbar, pers. obs.). According to the IUCN Red List criteria (
<bibRefCitation id="6F57D6C3FFCFFFA4FDDAFA00FD75F9F7" author="IUCN &amp; Species Survival Commission, IUCN" box="[601,738,1526,1552]" pageId="7" pageNumber="132" refId="ref4249" refString="IUCN (2001) IUCN Red List Categories and Criteria. Version 3. 1. Species Survival Commission, IUCN, Gland and Cambridge, 33 pp." type="book" year="2001">IUCN 2001</bibRefCitation>
), this species could be classified as Critically Endangered (CR). This category includes taxa in which fewer than 50 mature individuals exist.
</paragraph>
<paragraph id="0B79AB32FFCFFFA4FF0BF9C9FDF4F947" blockId="7.[136,1452,230,1697]" pageId="7" pageNumber="132">
In recent years, several morphological and molecular studies have been done on this section (
<bibRefCitation id="6F57D6C3FFCFFFA4FB0CF9C8FA37F9BE" author="Garcia-Jacas, N. &amp; Uysal, T. &amp; Romashchenko, K. &amp; Suarez Santiago, V. N. &amp; Ertudrul, K. &amp; Susanna, A." box="[1167,1440,1598,1625]" pageId="7" pageNumber="132" pagination="741 - 753" refId="ref4181" refString="Garcia-Jacas, N., Uysal, T., Romashchenko, K., Suarez Santiago, V. N., Ertudrul, K. &amp; Susanna, A. (2006) Centaurea revisited: A molecular survey of the Jacea Group. Annals of Botany 98: 741 - 753." type="journal article" year="2006">
Garcia-Jacas
<emphasis id="39B27720FFCFFFA4FAA5F9C9FACAF9BF" box="[1318,1373,1598,1624]" italics="true" pageId="7" pageNumber="132">et al.</emphasis>
2006
</bibRefCitation>
). The evaluated morphological characters were most suitable for separating
<taxonomicName id="CCC6D0B1FFCFFFA4FC44F995FB3FF99B" authorityName="Ranjbar &amp; Mohamadi" authorityYear="2021" box="[967,1192,1634,1661]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="132" phylum="Tracheophyta" rank="species" species="dezfulica">
<emphasis id="39B27720FFCFFFA4FC44F995FB3FF99B" box="[967,1192,1634,1661]" italics="true" pageId="7" pageNumber="132">Centaurea dezfulica</emphasis>
</taxonomicName>
from
<taxonomicName id="CCC6D0B1FFCFFFA4FB6FF995FACBF99B" authorityName="Wagenitz" authorityYear="1980" box="[1260,1372,1634,1661]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="132" phylum="Tracheophyta" rank="species" species="pabotii">
<emphasis id="39B27720FFCFFFA4FB6FF995FACBF99B" box="[1260,1372,1634,1661]" italics="true" pageId="7" pageNumber="132">C. pabotii</emphasis>
</taxonomicName>
and
<taxonomicName id="CCC6D0B1FFCFFFA4FA11F995FEB2F947" authorityName="Ranjbar &amp; Mohamadi" authorityYear="2021" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="132" phylum="Tracheophyta" rank="species" species="kuhdashtensis">
<emphasis id="39B27720FFCFFFA4FA11F995FEB2F947" italics="true" pageId="7" pageNumber="132">C. kuhdashtensis</emphasis>
</taxonomicName>
from
<taxonomicName id="CCC6D0B1FFCFFFA4FEE9F971FE78F947" box="[362,495,1670,1697]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="132" phylum="Tracheophyta" rank="species" species="koeieana">
<emphasis id="39B27720FFCFFFA4FEE9F971FE78F947" box="[362,495,1670,1697]" italics="true" pageId="7" pageNumber="132">C. koeieana</emphasis>
</taxonomicName>
(
<tableCitation id="46449E89FFCFFFA4FE7DF970FDC6F947" box="[510,593,1670,1696]" captionStart="TABLE 1" captionStartId="7.[136,224,1744,1768]" captionText="TABLE 1. Diagnostic morphological characters of Centaurea dezfulica, C. koeieana, C. kuhdashtensis and C. pabotii." pageId="7" pageNumber="132">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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