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<document id="B07F2BEDD611CDF01CB8E9D1C1726DCC" ID-DOI="10.11646/zootaxa.3893.4.3" ID-GBIF-Dataset="26a81425-6fa8-4f17-8518-9500e9b18cae" ID-ISSN="1175-5326" ID-Zenodo-Dep="250414" ID-ZooBank="CF82AC1C-372C-495A-8053-2BAA0370354C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460996181066" checkinUser="plazi" docAuthor="Schwendinger, Peter J. &amp; Hongpadharakiree, Komson" docDate="2014" docId="83648792FFEB1D1BFF4843B9FB608AE7" docLanguage="en" docName="zt03893p550.pdf" docOrigin="Zootaxa 3893 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Prothemenops irineae, sp. n." docType="treatment" docVersion="9" lastPageNumber="537" masterDocId="7F5DFFEAFFE91D13FFDF407FFFA28B5C" masterDocTitle="Three new Prothemenops species (Araneae: Idiopidae) from central Thailand" masterLastPageNumber="550" masterPageNumber="530" pageNumber="532" updateTime="1698562868504" updateUser="plazi">
<mods:mods id="F37A3DE5EBE0F6E23CEE2C3339E8C12B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="904924A2F0AEFD2AE7ADF804F7793057">
<mods:title id="47A6B5925EB50B2CA8506D53EC5534C1">Three new Prothemenops species (Araneae: Idiopidae) from central Thailand</mods:title>
</mods:titleInfo>
<mods:name id="D8CD45082A5C71BCE9C7B2E14DE6AFAB" type="personal">
<mods:role id="3554F64E813C8F1FF869599F7AF49DE8">
<mods:roleTerm id="F7C959207CC46C551ED3544C8F076449">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E57AE5B11FFAEB9582E2C498B80E0CDD">Schwendinger, Peter J.</mods:namePart>
</mods:name>
<mods:name id="27112A99AD000F3C3F1205262CB1FD4A" type="personal">
<mods:role id="EEF7CA4A7C7AD49AB3C6064C52E6A6D4">
<mods:roleTerm id="7795B7FA05E55A4B3349498E3F9B70E9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9EF57E15B5818E16FA8D8605399DA73D">Hongpadharakiree, Komson</mods:namePart>
</mods:name>
<mods:typeOfResource id="E858BAA781676A7BDFFA464E530984CF">text</mods:typeOfResource>
<mods:relatedItem id="57BA6854ECDBE06F40CE9EA97DEA8D5C" type="host">
<mods:titleInfo id="6E62FCCB4E0DE5E87CDB7062B9D54729">
<mods:title id="8AF2DFCCB7BBC638BFB08DB419A90F1A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="129C4029DD7EBD63ED7777AE69C2317E">
<mods:date id="1466D664A6AE1BE049085DDF303BAA1C">2014</mods:date>
<mods:detail id="8AA8B6CCE91DAC81D00D7DCA9A25EE48" type="volume">
<mods:number id="FA30EF30F223DA64592FFE5FA9B8EC24">3893</mods:number>
</mods:detail>
<mods:detail id="F1AD4EA219B82130379EEA01A2EAFA52" type="issue">
<mods:number id="D75C5FD39888A6068686F7D644FCCBAD">4</mods:number>
</mods:detail>
<mods:extent id="860E50F63F5373D5462A6B6AD8E09ADD" unit="page">
<mods:start id="75FF3E507AF8BA0F23BA6FDD7D82A9D3">530</mods:start>
<mods:end id="912016CC45FE98E41C2EFA296464E186">550</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5DC31C3B45807A8329AC6C37E214D67A">journal article</mods:classification>
<mods:identifier id="DDCFD32654C5B503D247E8D24F5CB6EE" type="DOI">10.11646/zootaxa.3893.4.3</mods:identifier>
<mods:identifier id="5BE7104C2FF734F2BE1C86A8B950AFBA" type="GBIF-Dataset">26a81425-6fa8-4f17-8518-9500e9b18cae</mods:identifier>
<mods:identifier id="6334B42FC3B13CB4721856F24782252E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="49B08F5BE468D933EBFC826D1104B591" type="Zenodo-Dep">250414</mods:identifier>
<mods:identifier id="0827DC3ED5DC8878235495BBD20B5B02" type="ZooBank">CF82AC1C-372C-495A-8053-2BAA0370354C</mods:identifier>
</mods:mods>
<treatment id="83648792FFEB1D1BFF4843B9FB608AE7" ID-DOI="http://doi.org/10.5281/zenodo.6123477" ID-GBIF-Taxon="119597359" ID-Zenodo-Dep="6123477" LSID="urn:lsid:plazi:treatment:83648792FFEB1D1BFF4843B9FB608AE7" httpUri="http://treatment.plazi.org/id/83648792FFEB1D1BFF4843B9FB608AE7" lastPageId="8" lastPageNumber="537" pageId="2" pageNumber="532">
<subSubSection id="43D7650FFFEB1D11FF4843B9FE4888BC" box="[151,490,966,992]" pageId="2" pageNumber="532" type="nomenclature">
<paragraph id="0B723684FFEB1D11FF4843B9FE4888BC" blockId="2.[151,490,966,1026]" box="[151,490,966,992]" pageId="2" pageNumber="532">
<heading id="503A81E8FFEB1D11FF4843B9FE4888BC" bold="true" box="[151,490,966,992]" fontSize="11" level="1" pageId="2" pageNumber="532" reason="1">
<emphasis id="39B9EA96FFEB1D11FF4843B9FE4888BC" bold="true" box="[151,490,966,992]" pageId="2" pageNumber="532">
<taxonomicName id="CCCD4D07FFEB1D11FF4843B9FE0388BC" ID-CoL="4N44N" authority="Schwendinger &amp; Hongpadharakiree 2014" box="[151,417,966,992]" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="532" phylum="Arthropoda" rank="species" species="irineae" status="sp. nov.">
<emphasis id="39B9EA96FFEB1D11FF4843B9FE0388BC" bold="true" box="[151,417,966,992]" italics="true" pageId="2" pageNumber="532">Prothemenops irineae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="228A57EDFFEB1D11FE7743B9FE4888BC" box="[424,490,966,992]" pageId="2" pageNumber="532" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="43D7650FFFEB1D11FF484396FE3F8F5E" box="[151,413,1001,1026]" pageId="2" pageNumber="532" type="description">
<paragraph id="0B723684FFEB1D11FF484396FE3F8F5E" blockId="2.[151,490,966,1026]" box="[151,413,1001,1026]" pageId="2" pageNumber="532">
<figureCitation id="93F62A01FFEB1D11FF484396FEBC8F5E" box="[151,286,1001,1026]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1357,1378]" captionStartId-1="5.[151,250,1871,1893]" captionStartId-2="6.[151,250,1215,1237]" captionStartId-3="7.[151,250,1341,1363]" captionTargetBox-0="[159,1423,165,1319]" captionTargetBox-1="[152,1431,343,1842]" captionTargetBox-2="[165,1433,183,1184]" captionTargetBox-3="[159,1432,223,1282]" captionTargetId-0="figure@3.[151,1436,155,1331]" captionTargetId-1="figure@5.[149,1437,338,1847]" captionTargetId-2="figure@6.[160,1442,175,1194]" captionTargetId-3="figure@7.[151,1436,195,1320]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 1. Prothemenops irineae sp. n., SEM micrographs of female (not a paratype). A, dorsolateral view of metatarsus and tarsus of right leg I showing distribution of club-shaped trichobothria (no clavate trichobothria present). B, same in dorsodistal view. C, detail of club-shaped trichobothrium. D, scopular setae on metatarsus I. E, tarsal organ (with adhering dust particles) of right tarsus I. F, tarsal organ of right tarsus II." captionText-1="FIGURE 2. Prothemenops irineae sp. n., male holotype (A, C, E – L) and female allotype (B, D). A – B, habitus, dorsal view. C – D, carapace, lateral view. E, eye group, dorsal view. F, distal part of right palp, retrodorsal view. G, same, proventral view. H, same, ventral view. I, tibia and metatarsus of right leg I, ventral view. J, tibia and metatarsus of right leg II, ventral view. K, labium and proximal part of left palpal coxa showing bristles with slightly swollen bases. L, recurved fovea with indentation in posterior margin." captionText-2="FIGURE 3. Prothemenops irineae sp. n., A – B, habitus of two males (B, male in defence posture). C – D, habitus of two females." captionText-3="FIGURE 4. Prothemenops irineae sp. n. A, right palpal organ of male holotype, retrolateral and slightly distal view. B, same, ventral and slightly distal view. C, left receptacle of exuvia of paratype, ental view (anterior is to the right, posterior to the left). D – G, vulvae of four female paratypes, dorsal view (D = allotype). Abbreviations: a = genital atrium, ae = apex of embolus, b = base of receptacle, be = base of embolus, dh = distal hematodocha, h = head of receptacle, pa = para-embolic apophysis, s = stalk of receptacle, st = subtegulum, t = tegulum, ue = uterus externus. Scale lines 0.1 mm (C), 0.5 mm (A – B, D – G)." httpUri-0="https://zenodo.org/record/250415/files/figure.png" httpUri-1="https://zenodo.org/record/250416/files/figure.png" httpUri-2="https://zenodo.org/record/250417/files/figure.png" httpUri-3="https://zenodo.org/record/250418/files/figure.png" pageId="2" pageNumber="532">Figures 1–4</figureCitation>
,
<figureCitation id="93F62A01FFEB1D11FEF54395FEE78F5E" box="[298,325,1002,1026]" captionStart="FIGURE 11" captionStartId="17.[151,250,1461,1483]" captionTargetBox="[211,1378,186,1441]" captionTargetId="figure@17.[211,1379,183,1450]" captionTargetPageId="17" captionText="FIGURE 11. Burrows of Prothemenops khirikhan sp. n. (A – B), P. irineae sp. n. (C, F) and P. phanthurat sp. n. (D – E). A, intact straight burrow just below the soil surface (1 a = open front door, 1 b = closed hind door). B, three such burrows close to each other (1 a – 3 a front door, 1 b – 3 b hind door of each burrow); upper side of longest burrow (shown in A) removed to show its shallow course. C, typical Y-shaped burrow still intact, with its two doors open and distinctly apart from each other. D, Yshaped burrow (in captivity) dug up to show its below-ground shape. E, Y-shaped burrow running into limestone crack, with both doors close to each other. F, straight, deep, blind-ending large burrow with its single door closed." httpUri="https://zenodo.org/record/250425/files/figure.png" pageId="2" pageNumber="532">11</figureCitation>
C, F, 12
</paragraph>
</subSubSection>
<subSubSection id="43D7650FFFEB1D11FF48444EFD858ED2" pageId="2" pageNumber="532" type="materials_examined">
<paragraph id="0B723684FFEB1D11FF48444EFEDC8E1A" blockId="2.[151,1437,1073,2034]" pageId="2" pageNumber="532">
<emphasis id="39B9EA96FFEB1D11FF48444EFEE48F16" bold="true" box="[151,326,1073,1098]" pageId="2" pageNumber="532">
<typeStatus id="D4768826FFEB1D11FF48444EFF708F16" box="[151,210,1073,1098]" pageId="2" pageNumber="532">Type</typeStatus>
material.
</emphasis>
<collectingCountry id="73DA7614FFEB1D11FE91444DFE438F16" box="[334,481,1074,1098]" name="Thailand" pageId="2" pageNumber="532">THAILAND</collectingCountry>
,
<collectingMunicipality id="EB16ACFEFFEB1D11FE32444EFCEE8F16" box="[493,844,1073,1098]" pageId="2" pageNumber="532">Prachuap Khiri Khan Province</collectingMunicipality>
,
<collectingRegion id="C909F866FFEB1D11FC87444EFBAC8F16" box="[856,1038,1073,1098]" pageId="2" pageNumber="532">Kuiburi District</collectingRegion>
,
<location id="0E12605FFFEB1D11FBC4444EFA3A8F16" LSID="urn:lsid:plazi:treatment:83648792FFEB1D1BFF4843B9FB608AE7:0E12605FFFEB1D11FBC4444EFA3A8F16" box="[1051,1432,1073,1098]" country="Thailand" municipality="Prachuap Khiri Khan Province" name="Khao Sam Roi Yot National Park" pageId="2" pageNumber="532" stateProvince="Kuiburi District">Khao Sam Roi Yot National Park</location>
, site 1, 5–
<quantity id="CC359B61FFEB1D11FEDC442AFEE08F31" box="[259,322,1109,1134]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="532" unit="m" value="80.0">
<elevation id="80E0D1B7FFEB1D11FEDC442AFEE08F31" box="[259,322,1109,1134]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="532" unit="m" value="80.0">80 m</elevation>
</quantity>
, secondary forest on limestone:
<materialsCitation id="BBA53CD9FFEB1D11FD61442AFBFF8F32" ID-GBIF-Occurrence="3707969309" box="[702,1117,1109,1134]" collectionCode="MHNG" country="Thailand" elevation="80" location="Khao Sam Roi Yot National Park" municipality="Prachuap Khiri Khan Province" pageId="2" pageNumber="532" specimenCount="1" specimenCount-male="1" stateProvince="Kuiburi District" typeStatus="holotype">
<specimenCount id="1DCBFD0DFFEB1D11FD61442AFD568F32" box="[702,756,1109,1134]" pageId="2" pageNumber="532" type="male">male</specimenCount>
<typeStatus id="D4768826FFEB1D11FD20442AFCC38F32" box="[767,865,1109,1134]" pageId="2" pageNumber="532" type="holotype">holotype</typeStatus>
(matured
<date id="7F731044FFEB1D11FC04442AFBF28F32" box="[987,1104,1109,1134]" pageId="2" pageNumber="532" value="2012-09-05">5.IX.2012</date>
),
</materialsCitation>
<materialsCitation id="BBA53CD9FFEB1D11FBB7442AFD6E8FCE" ID-GBIF-Occurrence="3707969311" country="Thailand" elevation="80" location="Khao Sam Roi Yot National Park" municipality="Prachuap Khiri Khan Province" pageId="2" pageNumber="532" specimenCount="5" specimenCount-male="5" stateProvince="Kuiburi District" typeStatus="paratype">
<specimenCount id="1DCBFD0DFFEB1D11FBB7442AFB148F32" box="[1128,1206,1109,1134]" pageId="2" pageNumber="532" type="male">5 male</specimenCount>
<typeStatus id="D4768826FFEB1D11FB1E4429FA8E8F32" box="[1217,1324,1110,1134]" pageId="2" pageNumber="532" type="paratype">paratypes</typeStatus>
(matured
<date id="7F731044FFEB1D11FF484405FEBA8FCE" box="[151,280,1146,1170]" pageId="2" pageNumber="532" value="2011-11-29">29.XI.2011</date>
, 7.VIII.,
<date id="7F731044FFEB1D11FE5F4405FE7F8FCE" box="[384,477,1146,1170]" pageId="2" pageNumber="532" value="2012-08">12.VIII.</date>
,
<date id="7F731044FFEB1D11FE364405FDE48FCE" box="[489,582,1146,1170]" pageId="2" pageNumber="532" value="2012-08">14.VIII.</date>
,
<date id="7F731044FFEB1D11FD8C4405FD1C8FCE" box="[595,702,1146,1170]" pageId="2" pageNumber="532" value="2012-10-01">1.X.2012</date>
),
</materialsCitation>
<materialsCitation id="BBA53CD9FFEB1D11FD0C4406FD438F86" ID-GBIF-Occurrence="3707969303" collectedFrom="secondary forest on limestone" collectingDate="2011-07-03" collectionCode="MHNG" collectorName="P. J. Schwendinger &amp; K. Hongpadharakiree &amp; D. Manathamkamon" elevation="0" location="Khao Sam Roi Yot National Park" pageId="2" pageNumber="532" specimenCount="6" specimenCount-female="6" typeStatus="paratype">
<specimenCount id="1DCBFD0DFFEB1D11FD0C4406FC948FCE" box="[723,822,1145,1170]" pageId="2" pageNumber="532" type="female">6 female</specimenCount>
<typeStatus id="D4768826FFEB1D11FCE14405FC088FCE" box="[830,938,1146,1170]" pageId="2" pageNumber="532" type="paratype">paratypes</typeStatus>
(one of them the allotype);
<date id="7F731044FFEB1D11FB364405FAC48FCE" box="[1257,1382,1146,1170]" pageId="2" pageNumber="532" value="2011-07-03">
<collectingDate id="6F37E9ACFFEB1D11FB364405FAC48FCE" box="[1257,1382,1146,1170]" pageId="2" pageNumber="532" value="2011-07-03">3.VII.2011</collectingDate>
</date>
; leg.
<collectorName id="A6385352FFEB1D11FF4844E1FEC58FEA" box="[151,359,1181,1206]" pageId="2" pageNumber="532">P.J. Schwendinger</collectorName>
,
<collectorName id="A6385352FFEB1D11FEAB44E1FDC48FEA" box="[372,614,1181,1206]" pageId="2" pageNumber="532">K. Hongpadharakiree</collectorName>
&amp;
<collectorName id="A6385352FFEB1D11FD5244E1FCDB8FEA" box="[653,889,1181,1206]" pageId="2" pageNumber="532">D. Manathamkamon</collectorName>
; sample TH-11/04.
<location id="0E12605FFFEB1D11FB8444E2FF6F8F86" LSID="urn:lsid:plazi:treatment:83648792FFEB1D1BFF4843B9FB608AE7:0E12605FFFEB1D11FB8444E2FF6F8F86" name="Khao Sam Roi Yot National Park" pageId="2" pageNumber="532">Khao Sam Roi Yot National Park</location>
, site 2, 50 m,
<collectedFrom id="B8D17579FFEB1D11FEA244BEFD7F8F86" box="[381,733,1217,1242]" pageId="2" pageNumber="532">secondary forest on limestone</collectedFrom>
:
</materialsCitation>
<materialsCitation id="BBA53CD9FFEB1D11FD3444BDFB0F8F86" ID-GBIF-Occurrence="3707969320" box="[747,1197,1217,1242]" country="Thailand" elevation="80" location="Khao Sam Roi Yot National Park" municipality="Prachuap Khiri Khan Province" pageId="2" pageNumber="532" specimenCount="1" specimenCount-male="1" stateProvince="Kuiburi District" typeStatus="paratype">
<specimenCount id="1DCBFD0DFFEB1D11FD3444BDFC988F86" box="[747,826,1217,1242]" pageId="2" pageNumber="532" type="male">1 male</specimenCount>
<typeStatus id="D4768826FFEB1D11FC9944BDFC048F86" box="[838,934,1218,1242]" pageId="2" pageNumber="532" type="paratype">paratype</typeStatus>
(matured
<date id="7F731044FFEB1D11FBFE44BDFB008F86" box="[1057,1186,1218,1242]" pageId="2" pageNumber="532" value="2014-04-21">21.IV.2014</date>
),
</materialsCitation>
<materialsCitation id="BBA53CD9FFEB1D11FB6644BDFCCC8FA2" ID-GBIF-Occurrence="3707969306" collectingDate="2012-12-21" collectorName="K. Hongpadharakiree &amp; D. Manathamkamon" country="Thailand" elevation="80" location="Khao Sam Roi Yot National Park" municipality="Prachuap Khiri Khan Province" pageId="2" pageNumber="532" specimenCount="2" specimenCount-female="2" stateProvince="Kuiburi District" typeStatus="paratype">
<specimenCount id="1DCBFD0DFFEB1D11FB6644BDFABC8F86" box="[1209,1310,1217,1242]" pageId="2" pageNumber="532" type="female">2 female</specimenCount>
<typeStatus id="D4768826FFEB1D11FAF644BDFA3A8F86" box="[1321,1432,1218,1242]" pageId="2" pageNumber="532" type="paratype">paratypes</typeStatus>
;
<date id="7F731044FFEB1D11FF484499FE818FA2" box="[151,291,1254,1278]" pageId="2" pageNumber="532" value="2012-12-21">
<collectingDate id="6F37E9ACFFEB1D11FF484499FE818FA2" box="[151,291,1254,1278]" pageId="2" pageNumber="532" value="2012-12-21">21.XII.2012</collectingDate>
</date>
; leg.
<collectorName id="A6385352FFEB1D11FEBC4499FDF48FA2" box="[355,598,1253,1278]" pageId="2" pageNumber="532">K. Hongpadharakiree</collectorName>
&amp;
<collectorName id="A6385352FFEB1D11FDA14499FCC88FA2" box="[638,874,1253,1278]" pageId="2" pageNumber="532">D. Manathamkamon</collectorName>
.
</materialsCitation>
<materialsCitation id="BBA53CD9FFEB1D11FCA84499FA268FA2" ID-GBIF-Occurrence="3707969301" box="[887,1412,1253,1278]" pageId="2" pageNumber="532" typeStatus="paratype">
<specimenCount id="1DCBFD0DFFEB1D11FCA84499FC668FA2" box="[887,964,1253,1278]" pageId="2" pageNumber="532" type="male">1 male</specimenCount>
<typeStatus id="D4768826FFEB1D11FC124499FB8F8FA2" box="[973,1069,1254,1278]" pageId="2" pageNumber="532" type="paratype">paratype</typeStatus>
(THNHM-I-2014-00001) and
</materialsCitation>
<materialsCitation id="BBA53CD9FFEB1D11FA514499FD238E7E" ID-GBIF-Occurrence="3707969322" pageId="2" pageNumber="532" typeStatus="paratype">
<specimenCount id="1DCBFD0DFFEB1D11FA514499FF418E7E" pageId="2" pageNumber="532" type="female">1 female</specimenCount>
<typeStatus id="D4768826FFEB1D11FF2B4575FEF68E7E" box="[244,340,1290,1314]" pageId="2" pageNumber="532" type="paratype">paratype</typeStatus>
(THNHM-I-2014-00002)
</materialsCitation>
deposited in
<collectionCode id="6DDCAE41FFEB1D11FCEC4575FC3F8E7D" box="[819,925,1290,1313]" pageId="2" pageNumber="532">THNHM</collectionCode>
, all other specimens (without registration numbers) in
<collectionCode id="6DDCAE41FFEB1D11FEFB4551FED88E1A" box="[292,378,1326,1350]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="532">MHNG</collectionCode>
.
</paragraph>
<paragraph id="0B723684FFEB1D11FF18452EFD858ED2" blockId="2.[151,1437,1073,2034]" pageId="2" pageNumber="532">
<emphasis id="39B9EA96FFEB1D11FF18452EFE5D8E36" bold="true" box="[199,511,1361,1386]" pageId="2" pageNumber="532">Other material examined.</emphasis>
Same data as for
<typeStatus id="D4768826FFEB1D11FD10452EFC978E36" box="[719,821,1361,1386]" pageId="2" pageNumber="532" type="holotype">holotype</typeStatus>
;
<materialsCitation id="BBA53CD9FFEB1D11FC9E452DFD858ED2" ID-GBIF-Occurrence="3707969314" collectionCode="MHNG" country="Thailand" elevation="80" location="Khao Sam Roi Yot National Park" municipality="Prachuap Khiri Khan Province" pageId="2" pageNumber="532" specimenCount="5" specimenCount-female="2" specimenCount-juvenile="2" stateProvince="Kuiburi District">
<specimenCount id="1DCBFD0DFFEB1D11FC9E452DFC108E36" box="[833,946,1361,1386]" pageId="2" pageNumber="532" type="female">2 females</specimenCount>
,
<specimenCount id="1DCBFD0DFFEB1D11FC60452DFB398E36" box="[959,1179,1361,1386]" pageId="2" pageNumber="532">1 penultimate male</specimenCount>
,
<specimenCount id="1DCBFD0DFFEB1D11FB76452DFA868E36" box="[1193,1316,1361,1386]" pageId="2" pageNumber="532" type="juvenile">2 juveniles</specimenCount>
(all partly decomposed; deposited in
<collectionCode id="6DDCAE41FFEB1D11FE1E4509FDBE8ED2" box="[449,540,1398,1422]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="2" pageNumber="532">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="43D7650FFFEB1D11FF1845E6FD378E8A" pageId="2" pageNumber="532" type="etymology">
<paragraph id="0B723684FFEB1D11FF1845E6FD378E8A" blockId="2.[151,1437,1073,2034]" pageId="2" pageNumber="532">
<emphasis id="39B9EA96FFEB1D11FF1845E6FEEC8EEE" bold="true" box="[199,334,1433,1458]" pageId="2" pageNumber="532">Etymology.</emphasis>
This species is named after Darika Manathamkamon, nicknamed “Irine”, the wife of the second author, who first discovered this new species.
</paragraph>
</subSubSection>
<subSubSection id="43D7650FFFEB1D11FF18459EFEC48CAE" pageId="2" pageNumber="532" type="diagnosis">
<paragraph id="0B723684FFEB1D11FF18459EFEC48CAE" blockId="2.[151,1437,1073,2034]" pageId="2" pageNumber="532">
<emphasis id="39B9EA96FFEB1D11FF18459EFEE38EA6" bold="true" box="[199,321,1505,1530]" pageId="2" pageNumber="532">Diagnosis.</emphasis>
Distinguished from
<taxonomicName id="CCCD4D07FFEB1D11FDF6459DFD108EA5" box="[553,690,1506,1529]" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="532" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="39B9EA96FFEB1D11FDF6459DFD108EA5" box="[553,690,1506,1529]" italics="true" pageId="2" pageNumber="532">P. siamensis</emphasis>
</taxonomicName>
by males with a more slender palpal tibia carrying a much shorter retroventral apophysis lacking spicules; palpal organ with a basally narrower embolus equipped with an indistinct para-embolic apophysis; tibia of leg I more slender, without coupling spurs and megaspines; patella III with fewer (5–8 instead of 9–10) prodorsal spines; tarsus III aspinose (6/7 spines in
<typeStatus id="D4768826FFEB1D11FC184632FB8B8D3A" box="[967,1065,1613,1638]" pageId="2" pageNumber="532" type="holotype">holotype</typeStatus>
of
<emphasis id="39B9EA96FFEB1D11FB904631FB718D39" box="[1103,1235,1614,1637]" italics="true" pageId="2" pageNumber="532">P. s i a m e ns i s</emphasis>
). Females can be distinguished from those of
<taxonomicName id="CCCD4D07FFEB1D11FE0D460DFDF98DD5" box="[466,603,1650,1673]" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="532" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="39B9EA96FFEB1D11FE0D460DFDF98DD5" box="[466,603,1650,1673]" italics="true" pageId="2" pageNumber="532">P. siamensis</emphasis>
</taxonomicName>
by vulva with smaller, more globular receptacular bases, and receptacular stalks and heads bent dorsad instead of outward (ectad); tibia I and metatarsus III with weak scopula (absent in both
<typeStatus id="D4768826FFEB1D11FF4846C5FEA08D8E" box="[151,258,1722,1746]" pageId="2" pageNumber="532" type="paratype">paratypes</typeStatus>
of
<emphasis id="39B9EA96FFEB1D11FEEE46C5FE188D8D" box="[305,442,1722,1745]" italics="true" pageId="2" pageNumber="532">P. s i a m e ns i s</emphasis>
); lunate prolateral depression of same colour as rest of palpal coxa (lighter in both
<typeStatus id="D4768826FFEB1D11FF4846A1FEA08DAA" box="[151,258,1758,1782]" pageId="2" pageNumber="532" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="CCCD4D07FFEB1D11FEF146A1FE198DA9" box="[302,443,1758,1781]" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="532" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="39B9EA96FFEB1D11FEF146A1FE198DA9" box="[302,443,1758,1781]" italics="true" pageId="2" pageNumber="532">P. siamensis</emphasis>
</taxonomicName>
). Distinguished from
<taxonomicName id="CCCD4D07FFEB1D11FD6346A1FCEA8DA9" box="[700,840,1757,1781]" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="532" phylum="Arthropoda" rank="species" species="khirikhan" status="sp. nov.">
<emphasis id="39B9EA96FFEB1D11FD6346A1FCEA8DA9" box="[700,840,1757,1781]" italics="true" pageId="2" pageNumber="532">P. khirikhan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="228A57EDFFEB1D11FC8D46A1FC338DA9" box="[850,913,1758,1781]" pageId="2" pageNumber="532" rank="species">sp. n.</taxonomicNameLabel>
by palp of males possessing a shorter, more distally situated tibial apophysis, and a smaller para-embolic apophysis; females with a scopula on tibia I and with slightly larger receptacular bases. Distinguished from
<taxonomicName id="CCCD4D07FFEB1D11FD2D4759FC2B8C61" box="[754,905,1829,1853]" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="532" phylum="Arthropoda" rank="species" species="phanthurat" status="sp. nov.">
<emphasis id="39B9EA96FFEB1D11FD2D4759FC2B8C61" box="[754,905,1829,1853]" italics="true" pageId="2" pageNumber="532">P. phanthurat</emphasis>
</taxonomicName>
<taxonomicNameLabel id="228A57EDFFEB1D11FC4E4759FC6E8C61" box="[913,972,1830,1853]" pageId="2" pageNumber="532" rank="species">sp. n.</taxonomicNameLabel>
by larger body size; males with narrower apophysis lacking spicules on palpal tibia, with angular para-embolic apophysis on retrolateral side of embolus (crescent-shaped and situated on posterior side in
<taxonomicName id="CCCD4D07FFEB1D11FD164711FCC28CD9" box="[713,864,1901,1925]" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="532" phylum="Arthropoda" rank="species" species="phanthurat" status="sp. nov.">
<emphasis id="39B9EA96FFEB1D11FD164711FCC28CD9" box="[713,864,1901,1925]" italics="true" pageId="2" pageNumber="532">P. phanthurat</emphasis>
</taxonomicName>
<taxonomicNameLabel id="228A57EDFFEB1D11FCB64711FC048CDA" box="[873,934,1902,1926]" pageId="2" pageNumber="532" rank="species">sp. n.</taxonomicNameLabel>
), without sigmoid, elongate, subapical spine on prolateral side of tibia I; females with lunate prolateral depression of same colour as rest of palpal coxa, and with distinct scopula on tarsus III (rudimentary in
<taxonomicName id="CCCD4D07FFEB1D11FD2647C9FC348C91" box="[761,918,1973,1997]" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="532" phylum="Arthropoda" rank="species" species="phanthurat" status="sp. nov.">
<emphasis id="39B9EA96FFEB1D11FD2647C9FC348C91" box="[761,918,1973,1997]" italics="true" pageId="2" pageNumber="532">P. phanthurat</emphasis>
</taxonomicName>
<taxonomicNameLabel id="228A57EDFFEB1D11FC7B47C9FC478C92" box="[932,997,1974,1998]" pageId="2" pageNumber="532" rank="species">sp. n.</taxonomicNameLabel>
) and on metatarsus III (absent in
<taxonomicName id="CCCD4D07FFEB1D11FA5847C9FEB68CAD" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="532" phylum="Arthropoda" rank="species" species="phanthurat" status="sp. nov.">
<emphasis id="39B9EA96FFEB1D11FA5847C9FEB68CAD" italics="true" pageId="2" pageNumber="532">P. phanthurat</emphasis>
</taxonomicName>
<taxonomicNameLabel id="228A57EDFFEB1D11FEC547A5FEFA8CAE" box="[282,344,2010,2034]" pageId="2" pageNumber="532" rank="species">sp. n.</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
<caption id="5FB2660CFFEA1D10FF484532FDDD8E94" httpUri="https://zenodo.org/record/250415/files/figure.png" pageId="3" pageNumber="533" targetBox="[159,1423,165,1319]" targetPageId="3">
<paragraph id="0B723684FFEA1D10FF484532FDDD8E94" blockId="3.[151,1436,1357,1480]" pageId="3" pageNumber="533">
<emphasis id="39B9EA96FFEA1D10FF484532FEB18E3E" bold="true" box="[151,275,1357,1378]" pageId="3" pageNumber="533">FIGURE 1.</emphasis>
<taxonomicName id="CCCD4D07FFEA1D10FEC64532FE5B8E3F" box="[281,505,1357,1379]" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="533" phylum="Arthropoda" rank="species" species="irineae" status="sp. nov.">
<emphasis id="39B9EA96FFEA1D10FEC64532FE5B8E3F" box="[281,505,1357,1379]" italics="true" pageId="3" pageNumber="533">Prothemenops irineae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="228A57EDFFEA1D10FE204531FD948E3F" box="[511,566,1358,1379]" pageId="3" pageNumber="533" rank="species">sp. n.</taxonomicNameLabel>
, SEM micrographs of female (not a paratype). A, dorsolateral view of metatarsus and tarsus of right leg I showing distribution of club-shaped trichobothria (no clavate trichobothria present). B, same in dorsodistal view. C, detail of club-shaped trichobothrium. D, scopular setae on metatarsus I. E, tarsal organ (with adhering dust particles) of right tarsus I. F, tarsal organ of right tarsus II.
</paragraph>
</caption>
<subSubSection id="43D7650FFFEA1D15FF18458EFE7A8CE0" lastPageId="6" lastPageNumber="536" pageId="3" pageNumber="533" type="description">
<paragraph id="0B723684FFEA1D10FF18458EFDE08DA0" blockId="3.[151,1437,1521,2031]" pageId="3" pageNumber="533">
<emphasis id="39B9EA96FFEA1D10FF18458EFEFB8D56" bold="true" box="[199,345,1521,1546]" pageId="3" pageNumber="533">Description.</emphasis>
MALE (
<typeStatus id="D4768826FFEA1D10FE1C458EFD898D56" box="[451,555,1521,1546]" pageId="3" pageNumber="533" type="holotype">holotype</typeStatus>
). Colour in alcohol (
<figureCitation id="93F62A01FFEA1D10FCC2458EFCC68D55" box="[797,868,1521,1546]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[152,1431,343,1842]" captionTargetId="figure@5.[149,1437,338,1847]" captionTargetPageId="5" captionText="FIGURE 2. Prothemenops irineae sp. n., male holotype (A, C, E – L) and female allotype (B, D). A – B, habitus, dorsal view. C – D, carapace, lateral view. E, eye group, dorsal view. F, distal part of right palp, retrodorsal view. G, same, proventral view. H, same, ventral view. I, tibia and metatarsus of right leg I, ventral view. J, tibia and metatarsus of right leg II, ventral view. K, labium and proximal part of left palpal coxa showing bristles with slightly swollen bases. L, recurved fovea with indentation in posterior margin." httpUri="https://zenodo.org/record/250416/files/figure.png" pageId="3" pageNumber="533">Fig. 2</figureCitation>
A; for colouration of live males, see
<figureCitation id="93F62A01FFEA1D10FACD458EFAF78D55" box="[1298,1365,1521,1546]" captionStart="FIGURE 3" captionStartId="6.[151,250,1215,1237]" captionTargetBox="[165,1433,183,1184]" captionTargetId="figure@6.[160,1442,175,1194]" captionTargetPageId="6" captionText="FIGURE 3. Prothemenops irineae sp. n., A – B, habitus of two males (B, male in defence posture). C – D, habitus of two females." httpUri="https://zenodo.org/record/250417/files/figure.png" pageId="3" pageNumber="533">Fig. 3</figureCitation>
A–B): carapace, dorsal sides of leg femora and of palpal femora, and distal part of chelicerae reddish brown; coxae and trochanters plus proximal portion of chelicerae and distal half of cymbium brown; patellae to tarsi light brown; all tarsi with a cream-coloured median zone (= pseudosegmentation). Anterior portion of dorsal side of opisthosoma grey-brown, with an indistinct light central area; posterior portion of opisthosoma with six dark transversal bands interconnected along mid-line, their lateral ends tapering and dissolving into dark spots. Booklung plates light orange-brown; rest of opisthosoma (including spinnerets) light yellow-brown. Ventral side of whole animal generally much paler than dorsal side.
</paragraph>
<paragraph id="0B723684FFEA1D17FF184779FE1E8A99" blockId="3.[151,1437,1521,2031]" lastBlockId="4.[151,1437,151,2025]" lastPageId="4" lastPageNumber="534" pageId="3" pageNumber="533">
Total length 15.0. Carapace 5.8 long, 4.8 wide, hirsute, covered with fine white adpressed hairs, with small dark bristles on both sides of eye mound, and with strong dark bristles on eye mound, on lateral and posterior margin of carapace, on coxal elevations and on area behind fovea. Pars cephalica low (
<figureCitation id="93F62A01FFEA1D10FB574734FB6C8C38" box="[1160,1230,1867,1892]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[152,1431,343,1842]" captionTargetId="figure@5.[149,1437,338,1847]" captionTargetPageId="5" captionText="FIGURE 2. Prothemenops irineae sp. n., male holotype (A, C, E – L) and female allotype (B, D). A – B, habitus, dorsal view. C – D, carapace, lateral view. E, eye group, dorsal view. F, distal part of right palp, retrodorsal view. G, same, proventral view. H, same, ventral view. I, tibia and metatarsus of right leg I, ventral view. J, tibia and metatarsus of right leg II, ventral view. K, labium and proximal part of left palpal coxa showing bristles with slightly swollen bases. L, recurved fovea with indentation in posterior margin." httpUri="https://zenodo.org/record/250416/files/figure.png" pageId="3" pageNumber="533">Fig. 2</figureCitation>
C). Fovea slightly recurved (median portion straight, both ends recurved), narrow, occupying 10% of carapace width at that point, with a distinct, narrow indentation in its posterior margin (
<figureCitation id="93F62A01FFEA1D10FCE247EEFC208CF5" box="[829,898,1937,1962]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[152,1431,343,1842]" captionTargetId="figure@5.[149,1437,338,1847]" captionTargetPageId="5" captionText="FIGURE 2. Prothemenops irineae sp. n., male holotype (A, C, E – L) and female allotype (B, D). A – B, habitus, dorsal view. C – D, carapace, lateral view. E, eye group, dorsal view. F, distal part of right palp, retrodorsal view. G, same, proventral view. H, same, ventral view. I, tibia and metatarsus of right leg I, ventral view. J, tibia and metatarsus of right leg II, ventral view. K, labium and proximal part of left palpal coxa showing bristles with slightly swollen bases. L, recurved fovea with indentation in posterior margin." httpUri="https://zenodo.org/record/250416/files/figure.png" pageId="3" pageNumber="533">Fig. 2</figureCitation>
A, L). Eye group (
<figureCitation id="93F62A01FFEA1D10FB8447EEFB028CF5" box="[1115,1184,1937,1962]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[152,1431,343,1842]" captionTargetId="figure@5.[149,1437,338,1847]" captionTargetPageId="5" captionText="FIGURE 2. Prothemenops irineae sp. n., male holotype (A, C, E – L) and female allotype (B, D). A – B, habitus, dorsal view. C – D, carapace, lateral view. E, eye group, dorsal view. F, distal part of right palp, retrodorsal view. G, same, proventral view. H, same, ventral view. I, tibia and metatarsus of right leg I, ventral view. J, tibia and metatarsus of right leg II, ventral view. K, labium and proximal part of left palpal coxa showing bristles with slightly swollen bases. L, recurved fovea with indentation in posterior margin." httpUri="https://zenodo.org/record/250416/files/figure.png" pageId="3" pageNumber="533">Fig. 2</figureCitation>
A, E) 0.83 long, front width 0.68, back width 0.93, occupying slightly less than one-third of “head” width, raised on low mound carrying an extra-long sigmoid bristle between AME and ALE (
<figureCitation id="93F62A01FFEA1D10FCD747A9FCEE8CB3" box="[776,844,2006,2031]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[152,1431,343,1842]" captionTargetId="figure@5.[149,1437,338,1847]" captionTargetPageId="5" captionText="FIGURE 2. Prothemenops irineae sp. n., male holotype (A, C, E – L) and female allotype (B, D). A – B, habitus, dorsal view. C – D, carapace, lateral view. E, eye group, dorsal view. F, distal part of right palp, retrodorsal view. G, same, proventral view. H, same, ventral view. I, tibia and metatarsus of right leg I, ventral view. J, tibia and metatarsus of right leg II, ventral view. K, labium and proximal part of left palpal coxa showing bristles with slightly swollen bases. L, recurved fovea with indentation in posterior margin." httpUri="https://zenodo.org/record/250416/files/figure.png" pageId="3" pageNumber="533">Fig. 2</figureCitation>
C, E). MOQ 0.47 long, front width 0.61, back width 0.60. Eye diameters: AME 0.31, ALE 0.29, PME 0.24, PLE 0.30. Sternum 3.1 long, 2.5 wide; all sigilla small, anterior two pairs very close to margin, posterior pair slightly more distant from it. Labio-sternal suture very short but not interrupted in middle (reduced to thin transversal stripe and essentially interrupted in middle in
<typeStatus id="D4768826FFED1D17FAF940A1FAC38BAA" box="[1318,1377,222,246]" pageId="4" pageNumber="534">types</typeStatus>
of
<taxonomicName id="CCCD4D07FFED1D17FA5840A1FEA68A4B" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="534" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="39B9EA96FFED1D17FA5840A1FEA68A4B" italics="true" pageId="4" pageNumber="534">P. siamensis</emphasis>
</taxonomicName>
, see
<bibRefCitation id="6F5C4B75FFED1D17FE9C4080FDC68A44" author="Schwendinger, P. J." box="[323,612,255,280]" firstAuthor="Schwendinger" journalOrPublisher="Bulletin of the British arachnological Society" page="233" pageId="4" pageNumber="534" pagination="233 - 240" part="8" refId="ref13579" refString="Schwendinger, P. J. (1991) Two new trap-door spiders from Thailand (Araneae, Mygalomorphae, Idiopidae). Bulletin of the British arachnological Society, 8 (8), 233 - 240." title="Two new trap-door spiders from Thailand (Araneae, Mygalomorphae, Idiopidae)" type="journal article" year="1991">Schwendinger 1991: 233</bibRefCitation>
, fig. 5). Labium 0.4 long, 0.9 wide, without cuspules or spicules, with relatively long bristles in two groups on anterior margin (
<figureCitation id="93F62A01FFED1D17FC92415DFC348A67" box="[845,918,290,315]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[152,1431,343,1842]" captionTargetId="figure@5.[149,1437,338,1847]" captionTargetPageId="5" captionText="FIGURE 2. Prothemenops irineae sp. n., male holotype (A, C, E – L) and female allotype (B, D). A – B, habitus, dorsal view. C – D, carapace, lateral view. E, eye group, dorsal view. F, distal part of right palp, retrodorsal view. G, same, proventral view. H, same, ventral view. I, tibia and metatarsus of right leg I, ventral view. J, tibia and metatarsus of right leg II, ventral view. K, labium and proximal part of left palpal coxa showing bristles with slightly swollen bases. L, recurved fovea with indentation in posterior margin." httpUri="https://zenodo.org/record/250416/files/figure.png" pageId="4" pageNumber="534">Fig. 2</figureCitation>
K). Palpal coxae 1.9 long, 1.0 wide; lunate prolateral zone of ventral side slightly depressed and of same colour as rest of article; some prolateral-proximal bristles slightly swollen at base (
<figureCitation id="93F62A01FFED1D17FDD54118FDEC8ADC" box="[522,590,359,384]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[152,1431,343,1842]" captionTargetId="figure@5.[149,1437,338,1847]" captionTargetPageId="5" captionText="FIGURE 2. Prothemenops irineae sp. n., male holotype (A, C, E – L) and female allotype (B, D). A – B, habitus, dorsal view. C – D, carapace, lateral view. E, eye group, dorsal view. F, distal part of right palp, retrodorsal view. G, same, proventral view. H, same, ventral view. I, tibia and metatarsus of right leg I, ventral view. J, tibia and metatarsus of right leg II, ventral view. K, labium and proximal part of left palpal coxa showing bristles with slightly swollen bases. L, recurved fovea with indentation in posterior margin." httpUri="https://zenodo.org/record/250416/files/figure.png" pageId="4" pageNumber="534">Fig. 2</figureCitation>
K). Chelicerae with sessile rastellum composed of six stout blunt spines in two rows (four spines in anterior row, two in posterior); cheliceral groove with 7/7 teeth on promargin plus 3/5 tiny denticles in a median row.
</paragraph>
<paragraph id="0B723684FFED1D17FF1841AFFB988881" blockId="4.[151,1437,151,2025]" pageId="4" pageNumber="534">
Legs 3214; lengths: leg I 17.7 (4.8 + 3.0 + 3.9 + 3.9 + 2.1), leg II 16.1 (4.5 + 2.5 + 3.5 + 3.6 + 2.0), leg III 14.8 (4.0 + 2.0 + 2.8 + 3.8 + 2.2), leg IV 21.1 (5.3 + 2.7 + 5.1 + 5.4 + 2.6). Three relatively weak, light, glabrous longitudinal stripes (two dorsal and one retrolateral) on all femora of palps and legs; the retrolateral one on palps and legs I–II shorter than on legs III–IV. Two such dorsal stripes (wider but less distinct than on femora) also on all patellae and on tibia of palp. Tibia I and metatarsus I without modifications (
<figureCitation id="93F62A01FFED1D17FC254221FB9D892B" box="[1018,1087,606,631]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[152,1431,343,1842]" captionTargetId="figure@5.[149,1437,338,1847]" captionTargetPageId="5" captionText="FIGURE 2. Prothemenops irineae sp. n., male holotype (A, C, E – L) and female allotype (B, D). A – B, habitus, dorsal view. C – D, carapace, lateral view. E, eye group, dorsal view. F, distal part of right palp, retrodorsal view. G, same, proventral view. H, same, ventral view. I, tibia and metatarsus of right leg I, ventral view. J, tibia and metatarsus of right leg II, ventral view. K, labium and proximal part of left palpal coxa showing bristles with slightly swollen bases. L, recurved fovea with indentation in posterior margin." httpUri="https://zenodo.org/record/250416/files/figure.png" pageId="4" pageNumber="534">
Fig.
<date id="7F731044FFED1D17FBF04220FB9D892B" box="[1071,1087,607,631]" pageId="4" pageNumber="534">2</date>
</figureCitation>
I); metatarsi I–II each with one strong proventral spine and two weak retroventral spines (stiff bristles) (
<figureCitation id="93F62A01FFED1D17FC1F42FDFBA689C7" box="[960,1028,642,667]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[152,1431,343,1842]" captionTargetId="figure@5.[149,1437,338,1847]" captionTargetPageId="5" captionText="FIGURE 2. Prothemenops irineae sp. n., male holotype (A, C, E – L) and female allotype (B, D). A – B, habitus, dorsal view. C – D, carapace, lateral view. E, eye group, dorsal view. F, distal part of right palp, retrodorsal view. G, same, proventral view. H, same, ventral view. I, tibia and metatarsus of right leg I, ventral view. J, tibia and metatarsus of right leg II, ventral view. K, labium and proximal part of left palpal coxa showing bristles with slightly swollen bases. L, recurved fovea with indentation in posterior margin." httpUri="https://zenodo.org/record/250416/files/figure.png" pageId="4" pageNumber="534">
Fig.
<date id="7F731044FFED1D17FC2A42FCFBA689C7" box="[1013,1028,643,667]" pageId="4" pageNumber="534">2</date>
</figureCitation>
I–J). Patella III with row of six short prodorsal spines on right side and with row of five such spines on left side; all tibiae and metatarsi of legs with ventral spines, those on posterior legs more numerous and extending to pro- and retrodorsal sides; all leg tarsi aspinose (but see “Variation”). Filiform trichobothria on tibiae to tarsi of legs; club-shaped trichobothria (
<figureCitation id="93F62A01FFED1D17FAE04292FADD885A" box="[1343,1407,749,774]" captionStart="FIGURE 1" captionStartId="3.[151,250,1357,1378]" captionTargetBox="[159,1423,165,1319]" captionTargetId="figure@3.[151,1436,155,1331]" captionTargetPageId="3" captionText="FIGURE 1. Prothemenops irineae sp. n., SEM micrographs of female (not a paratype). A, dorsolateral view of metatarsus and tarsus of right leg I showing distribution of club-shaped trichobothria (no clavate trichobothria present). B, same in dorsodistal view. C, detail of club-shaped trichobothrium. D, scopular setae on metatarsus I. E, tarsal organ (with adhering dust particles) of right tarsus I. F, tarsal organ of right tarsus II." httpUri="https://zenodo.org/record/250415/files/figure.png" pageId="4" pageNumber="534">Fig. 1</figureCitation>
C, showing female) in distal two-thirds of metatarsi only (
<figureCitation id="93F62A01FFED1D17FCDE436EFCE28876" box="[769,832,785,810]" captionStart="FIGURE 1" captionStartId="3.[151,250,1357,1378]" captionTargetBox="[159,1423,165,1319]" captionTargetId="figure@3.[151,1436,155,1331]" captionTargetPageId="3" captionText="FIGURE 1. Prothemenops irineae sp. n., SEM micrographs of female (not a paratype). A, dorsolateral view of metatarsus and tarsus of right leg I showing distribution of club-shaped trichobothria (no clavate trichobothria present). B, same in dorsodistal view. C, detail of club-shaped trichobothrium. D, scopular setae on metatarsus I. E, tarsal organ (with adhering dust particles) of right tarsus I. F, tarsal organ of right tarsus II." httpUri="https://zenodo.org/record/250415/files/figure.png" pageId="4" pageNumber="534">Fig. 1</figureCitation>
A–B, showing female). Scopula thin on entire ventral side of tarsus I and II, very thin on entire ventral side of tarsus III, very thin and only in distal two-thirds of tarsus IV; thin distally and very thin proximally on ventral side of metatarsi I–II (on II slightly denser than on I;
<figureCitation id="93F62A01FFED1D17FAF64326FACB882D" box="[1321,1385,857,882]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[152,1431,343,1842]" captionTargetId="figure@5.[149,1437,338,1847]" captionTargetPageId="5" captionText="FIGURE 2. Prothemenops irineae sp. n., male holotype (A, C, E – L) and female allotype (B, D). A – B, habitus, dorsal view. C – D, carapace, lateral view. E, eye group, dorsal view. F, distal part of right palp, retrodorsal view. G, same, proventral view. H, same, ventral view. I, tibia and metatarsus of right leg I, ventral view. J, tibia and metatarsus of right leg II, ventral view. K, labium and proximal part of left palpal coxa showing bristles with slightly swollen bases. L, recurved fovea with indentation in posterior margin." httpUri="https://zenodo.org/record/250416/files/figure.png" pageId="4" pageNumber="534">
Fig.
<date id="7F731044FFED1D17FA844326FACB882D" box="[1371,1385,857,881]" pageId="4" pageNumber="534">2</date>
</figureCitation>
I–J), very thin and only in distal fourth on metatarsus III, absent from metatarsus IV. Scopular setae hairbrush-shaped (
<figureCitation id="93F62A01FFED1D17FF4043DFFF4088E5" box="[159,226,928,953]" captionStart="FIGURE 1" captionStartId="3.[151,250,1357,1378]" captionTargetBox="[159,1423,165,1319]" captionTargetId="figure@3.[151,1436,155,1331]" captionTargetPageId="3" captionText="FIGURE 1. Prothemenops irineae sp. n., SEM micrographs of female (not a paratype). A, dorsolateral view of metatarsus and tarsus of right leg I showing distribution of club-shaped trichobothria (no clavate trichobothria present). B, same in dorsodistal view. C, detail of club-shaped trichobothrium. D, scopular setae on metatarsus I. E, tarsal organ (with adhering dust particles) of right tarsus I. F, tarsal organ of right tarsus II." httpUri="https://zenodo.org/record/250415/files/figure.png" pageId="4" pageNumber="534">Fig. 1</figureCitation>
D, showing female). Tarsal organ low, cowpat-shaped (
<figureCitation id="93F62A01FFED1D17FCB343DFFC0F88E5" box="[876,941,928,953]" captionStart="FIGURE 1" captionStartId="3.[151,250,1357,1378]" captionTargetBox="[159,1423,165,1319]" captionTargetId="figure@3.[151,1436,155,1331]" captionTargetPageId="3" captionText="FIGURE 1. Prothemenops irineae sp. n., SEM micrographs of female (not a paratype). A, dorsolateral view of metatarsus and tarsus of right leg I showing distribution of club-shaped trichobothria (no clavate trichobothria present). B, same in dorsodistal view. C, detail of club-shaped trichobothrium. D, scopular setae on metatarsus I. E, tarsal organ (with adhering dust particles) of right tarsus I. F, tarsal organ of right tarsus II." httpUri="https://zenodo.org/record/250415/files/figure.png" pageId="4" pageNumber="534">Fig. 1</figureCitation>
E–F, showing female). Paired tarsal claws armed with row of 5–7 teeth (second tooth from base largest); unpaired claws bare.
</paragraph>
<paragraph id="0B723684FFED1D17FF184398FDCB8FA6" blockId="4.[151,1437,151,2025]" pageId="4" pageNumber="534">
Palp 7.2 long (2.5 + 1.8 + 1.9 + 1.0), slender. Relatively slender tibia with short retroventral apophysis situated some distance from distal margin and carrying only bristles (no spines or spicules;
<figureCitation id="93F62A01FFED1D17FB834474FB028F78" box="[1116,1184,1035,1060]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[152,1431,343,1842]" captionTargetId="figure@5.[149,1437,338,1847]" captionTargetPageId="5" captionText="FIGURE 2. Prothemenops irineae sp. n., male holotype (A, C, E – L) and female allotype (B, D). A – B, habitus, dorsal view. C – D, carapace, lateral view. E, eye group, dorsal view. F, distal part of right palp, retrodorsal view. G, same, proventral view. H, same, ventral view. I, tibia and metatarsus of right leg I, ventral view. J, tibia and metatarsus of right leg II, ventral view. K, labium and proximal part of left palpal coxa showing bristles with slightly swollen bases. L, recurved fovea with indentation in posterior margin." httpUri="https://zenodo.org/record/250416/files/figure.png" pageId="4" pageNumber="534">Fig. 2</figureCitation>
F–H). Short cymbium with rounded lobe prodistally and rather angular, indistinct lobe retrodistally, carrying weak spines (stiff bristles) on distal margin plus club-shaped trichobothria on dorsal surface. Palpal organ with oval tegulum, its distal haematodocha vertically divided by narrow pigmented stripe and not reaching base of embolus (
<figureCitation id="93F62A01FFED1D17FB024409FA848FD3" box="[1245,1318,1142,1167]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[152,1431,343,1842]" captionTargetId="figure@5.[149,1437,338,1847]" captionTargetPageId="5" captionText="FIGURE 2. Prothemenops irineae sp. n., male holotype (A, C, E – L) and female allotype (B, D). A – B, habitus, dorsal view. C – D, carapace, lateral view. E, eye group, dorsal view. F, distal part of right palp, retrodorsal view. G, same, proventral view. H, same, ventral view. I, tibia and metatarsus of right leg I, ventral view. J, tibia and metatarsus of right leg II, ventral view. K, labium and proximal part of left palpal coxa showing bristles with slightly swollen bases. L, recurved fovea with indentation in posterior margin." httpUri="https://zenodo.org/record/250416/files/figure.png" pageId="4" pageNumber="534">Figs 2</figureCitation>
H, 4A–B); embolus long, slender and bent away from axis of palp; base of embolus occupying more than half of total embolus length, ending in an indistinct, short triangular para-embolic apophysis retrolaterally; apex of embolus thin, flexible, flagelliform (
<figureCitation id="93F62A01FFED1D17FE50449DFE748FA6" box="[399,470,1250,1275]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[152,1431,343,1842]" captionTargetId="figure@5.[149,1437,338,1847]" captionTargetPageId="5" captionText="FIGURE 2. Prothemenops irineae sp. n., male holotype (A, C, E – L) and female allotype (B, D). A – B, habitus, dorsal view. C – D, carapace, lateral view. E, eye group, dorsal view. F, distal part of right palp, retrodorsal view. G, same, proventral view. H, same, ventral view. I, tibia and metatarsus of right leg I, ventral view. J, tibia and metatarsus of right leg II, ventral view. K, labium and proximal part of left palpal coxa showing bristles with slightly swollen bases. L, recurved fovea with indentation in posterior margin." httpUri="https://zenodo.org/record/250416/files/figure.png" pageId="4" pageNumber="534">Figs 2</figureCitation>
F–H, 4A–B).
</paragraph>
<paragraph id="0B723684FFED1D17FF18457AFDFD8E3A" blockId="4.[151,1437,151,2025]" pageId="4" pageNumber="534">Opisthosoma covered with adpressed white hairs (only on dorsal side) and erect dark bristles (strongest in anterodorsal region), 6.8 long, 4.3 wide. PMS 0.5 long, PLS 2.4 long: apical article 0.4 (slightly longer than wide), median article 0.7, basal article 1.3 long.</paragraph>
<paragraph id="0B723684FFED1D17FF18450DFB748EF1" blockId="4.[151,1437,151,2025]" pageId="4" pageNumber="534">
FEMALE (allotype).
<collectingCountry id="73DA7614FFED1D17FE64450DFE788ED6" box="[443,474,1394,1418]" name="American Samoa" pageId="4" pageNumber="534">As</collectingCountry>
in male, except for: Colour in alcohol generally more greyish; chelicerae and dorsal pattern of opisthosoma distinctly darker (
<figureCitation id="93F62A01FFED1D17FDBA45EBFD0B8EF1" box="[613,681,1428,1453]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[152,1431,343,1842]" captionTargetId="figure@5.[149,1437,338,1847]" captionTargetPageId="5" captionText="FIGURE 2. Prothemenops irineae sp. n., male holotype (A, C, E – L) and female allotype (B, D). A – B, habitus, dorsal view. C – D, carapace, lateral view. E, eye group, dorsal view. F, distal part of right palp, retrodorsal view. G, same, proventral view. H, same, ventral view. I, tibia and metatarsus of right leg I, ventral view. J, tibia and metatarsus of right leg II, ventral view. K, labium and proximal part of left palpal coxa showing bristles with slightly swollen bases. L, recurved fovea with indentation in posterior margin." httpUri="https://zenodo.org/record/250416/files/figure.png" pageId="4" pageNumber="534">Fig. 2</figureCitation>
B; for colouration of live females see
<figureCitation id="93F62A01FFED1D17FB9045EBFB2D8EF1" box="[1103,1167,1428,1453]" captionStart="FIGURE 3" captionStartId="6.[151,250,1215,1237]" captionTargetBox="[165,1433,183,1184]" captionTargetId="figure@6.[160,1442,175,1194]" captionTargetPageId="6" captionText="FIGURE 3. Prothemenops irineae sp. n., A – B, habitus of two males (B, male in defence posture). C – D, habitus of two females." httpUri="https://zenodo.org/record/250417/files/figure.png" pageId="4" pageNumber="534">Fig. 3</figureCitation>
C–D).
</paragraph>
<paragraph id="0B723684FFED1D16FF1845C7FB2B8A63" blockId="4.[151,1437,151,2025]" lastBlockId="5.[151,1436,151,319]" lastPageId="5" lastPageNumber="535" pageId="4" pageNumber="534">
Total length 21.5. Carapace 7.6 long, 6.1 wide, bristles on its lateral and posterior margins thinner than in male; pars cephalica moderately raised (
<figureCitation id="93F62A01FFED1D17FDCC45A3FDF48EA9" box="[531,598,1500,1525]" captionStart="FIGURE 2" captionStartId="5.[151,250,1871,1893]" captionTargetBox="[152,1431,343,1842]" captionTargetId="figure@5.[149,1437,338,1847]" captionTargetPageId="5" captionText="FIGURE 2. Prothemenops irineae sp. n., male holotype (A, C, E – L) and female allotype (B, D). A – B, habitus, dorsal view. C – D, carapace, lateral view. E, eye group, dorsal view. F, distal part of right palp, retrodorsal view. G, same, proventral view. H, same, ventral view. I, tibia and metatarsus of right leg I, ventral view. J, tibia and metatarsus of right leg II, ventral view. K, labium and proximal part of left palpal coxa showing bristles with slightly swollen bases. L, recurved fovea with indentation in posterior margin." httpUri="https://zenodo.org/record/250416/files/figure.png" pageId="4" pageNumber="534">Fig. 2</figureCitation>
D), clearly higher than in male. Fovea occupying 15% of carapace width at that point; median portion of anterior margin slightly procurved, both ends recurved; posterior margin with wider indentation than in male. Eye group 1.10 long, front width 0.83, back width 1.38, occupying slightly less than onethird of “head” width. MOQ 0.59 long, front width 0.75, back width 0.74. Eye diameters: AME 0.36, ALE 0.31, PME 0.15, PLE 0.37. Sternum 4.4 long, 3.3 wide, its sigilla slightly more distinct than in male. Labium 0.7 long, 1.3 wide. Palpal coxae 2.8 long, 1.5 wide, carrying 43/46 thick cuspules proximally-prolaterally; lunate prolateral zone of ventral side slightly depressed and mostly of same colour as rest of article except for narrow light band along prolateral margin next to long, curved, reddish marginal hairs. Chelicerae more robust than in male; rastellum composed of about 20 short strong spines in 6–7 rows; cheliceral furrows with 7/7 teeth on promargin and 6/7 denticles (larger than in male) in a median row. Legs stouter than in male; lengths: leg I 15.4 (4.7 + 3.6 + 3.2 + 2.5 + 1.4), leg II 14.0 (4.4 + 3.1 + 2.7 + 2.4 + 1.4), leg III 12.8 (3.9 + 2.6 + 2.2 + 2.7 + 1.4), leg IV 19.8 (5.4 + 3.6 + 4.6 + 4.5 + 1.7). Patella III with 9/11 prodorsal spines, dorsal bristles stronger than in male; prolateral side of femur IV distally with dense group of much stronger bristles than in male. Scopula dense on entire ventral side of tarsi and metatarsi I–II (
<figureCitation id="93F62A01FFED1D17FE1F47D2FE5C8C99" box="[448,510,1965,1990]" captionStart="FIGURE 1" captionStartId="3.[151,250,1357,1378]" captionTargetBox="[159,1423,165,1319]" captionTargetId="figure@3.[151,1436,155,1331]" captionTargetPageId="3" captionText="FIGURE 1. Prothemenops irineae sp. n., SEM micrographs of female (not a paratype). A, dorsolateral view of metatarsus and tarsus of right leg I showing distribution of club-shaped trichobothria (no clavate trichobothria present). B, same in dorsodistal view. C, detail of club-shaped trichobothrium. D, scopular setae on metatarsus I. E, tarsal organ (with adhering dust particles) of right tarsus I. F, tarsal organ of right tarsus II." httpUri="https://zenodo.org/record/250415/files/figure.png" pageId="4" pageNumber="534">Fig. 1</figureCitation>
A–B), thin on entire ventral side of tarsus III, in distal half of ventral and prolateral sides of metatarsus III and distally on prolateral side of tibia I, absent on leg IV; scopula of palp dense on entire ventral side of tarsus and proventrally to prolaterally in distal half of tibia. All tarsi of palps and legs integral (not pseudosegmented as in males) and armed with spines: two widely spaced basal ones and 3/4 closely spaced distal ones on palpal tarsus; three distoventral ones on tarsi I–II; a prodorsal one plus 7–10 ventral ones on tarsus III; 11–12 prolateral, ventral and retroventral ones on tarsus IV; spines on other leg articles mostly as in male. Paired tarsal claws with row of 4–5 teeth, unpaired claws bare; palpal claw with row of 3/4 teeth.
</paragraph>
<caption id="5FB2660CFFEC1D16FF484730FEE48CBC" httpUri="https://zenodo.org/record/250416/files/figure.png" pageId="5" pageNumber="535" targetBox="[152,1431,343,1842]" targetPageId="5">
<paragraph id="0B723684FFEC1D16FF484730FEE48CBC" blockId="5.[151,1437,1871,2016]" pageId="5" pageNumber="535">
<emphasis id="39B9EA96FFEC1D16FF484730FEB68C38" bold="true" box="[151,276,1871,1893]" pageId="5" pageNumber="535">FIGURE 2.</emphasis>
<taxonomicName id="CCCD4D07FFEC1D16FEC34730FE5F8C3A" box="[284,509,1871,1894]" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="535" phylum="Arthropoda" rank="species" species="irineae" status="sp. nov.">
<emphasis id="39B9EA96FFEC1D16FEC34730FE5F8C3A" box="[284,509,1871,1894]" italics="true" pageId="5" pageNumber="535">Prothemenops irineae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="228A57EDFFEC1D16FDDA472FFD9C8C39" box="[517,574,1872,1893]" pageId="5" pageNumber="535" rank="species">sp. n.</taxonomicNameLabel>
, male holotype (A, C, E–L) and female allotype (B, D). A–B, habitus, dorsal view. C–D, carapace, lateral view. E, eye group, dorsal view. F, distal part of right palp, retrodorsal view. G, same, proventral view. H, same, ventral view. I, tibia and metatarsus of right leg I, ventral view. J, tibia and metatarsus of right leg II, ventral view. K, labium and proximal part of left palpal coxa showing bristles with slightly swollen bases. L, recurved fovea with indentation in posterior margin.
</paragraph>
</caption>
<caption id="5FB2660CFFEF1D15FF4844C0FF4F8FA8" httpUri="https://zenodo.org/record/250417/files/figure.png" pageId="6" pageNumber="536" targetBox="[165,1433,183,1184]" targetPageId="6">
<paragraph id="0B723684FFEF1D15FF4844C0FF4F8FA8" blockId="6.[151,1436,1215,1268]" pageId="6" pageNumber="536">
<emphasis id="39B9EA96FFEF1D15FF4844C0FEB48F88" bold="true" box="[151,278,1215,1237]" pageId="6" pageNumber="536">FIGURE 3.</emphasis>
<taxonomicName id="CCCD4D07FFEF1D15FEFE44C0FDA68F8A" box="[289,516,1215,1238]" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="irineae" status="sp. nov.">
<emphasis id="39B9EA96FFEF1D15FEFE44C0FDA68F8A" box="[289,516,1215,1238]" italics="true" pageId="6" pageNumber="536">Prothemenops irineae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="228A57EDFFEF1D15FDD144BFFDE88F89" box="[526,586,1216,1237]" pageId="6" pageNumber="536" rank="species">sp. n.</taxonomicNameLabel>
, A–B, habitus of two males (B, male in defence posture). C–D, habitus of two females.
</paragraph>
</caption>
<paragraph id="0B723684FFEF1D15FF184564FEE58E04" blockId="6.[151,1437,1307,1980]" pageId="6" pageNumber="536">Opisthosoma 10.0 long, 6.9 wide. PMS 0.6 long; PLS 3.0 long: apical article 0.5, median article 0.9, basal article 1.6 long.</paragraph>
<paragraph id="0B723684FFEF1D15FF18451CFACD8D50" blockId="6.[151,1437,1307,1980]" pageId="6" pageNumber="536">
Vulva (of
<typeStatus id="D4768826FFEF1D15FEE5451BFE0B8E20" box="[314,425,1380,1404]" pageId="6" pageNumber="536" type="paratype">paratypes</typeStatus>
;
<figureCitation id="93F62A01FFEF1D15FE6A451CFE5A8E20" box="[437,504,1379,1404]" captionStart="FIGURE 4" captionStartId="7.[151,250,1341,1363]" captionTargetBox="[159,1432,223,1282]" captionTargetId="figure@7.[151,1436,195,1320]" captionTargetPageId="7" captionText="FIGURE 4. Prothemenops irineae sp. n. A, right palpal organ of male holotype, retrolateral and slightly distal view. B, same, ventral and slightly distal view. C, left receptacle of exuvia of paratype, ental view (anterior is to the right, posterior to the left). D – G, vulvae of four female paratypes, dorsal view (D = allotype). Abbreviations: a = genital atrium, ae = apex of embolus, b = base of receptacle, be = base of embolus, dh = distal hematodocha, h = head of receptacle, pa = para-embolic apophysis, s = stalk of receptacle, st = subtegulum, t = tegulum, ue = uterus externus. Scale lines 0.1 mm (C), 0.5 mm (A – B, D – G)." httpUri="https://zenodo.org/record/250418/files/figure.png" pageId="6" pageNumber="536">Fig. 4</figureCitation>
C–G) with wide short atrium running into two widely separated sperm receptacles composed of thick-walled, strongly pigmented, ovoid basal portion (called “sclerotized lateral pouches” in description of
<taxonomicName id="CCCD4D07FFEF1D15FE9945D3FE778E9F" box="[326,469,1452,1475]" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="39B9EA96FFEF1D15FE9945D3FE778E9F" box="[326,469,1452,1475]" italics="true" pageId="6" pageNumber="536">P. siamensis</emphasis>
</taxonomicName>
, see
<bibRefCitation id="6F5C4B75FFEF1D15FDC445D4FCE68E98" author="Schwendinger, P. J." box="[539,836,1451,1476]" firstAuthor="Schwendinger" journalOrPublisher="Bulletin of the British arachnological Society" page="235" pageId="6" pageNumber="536" pagination="233 - 240" part="8" refId="ref13579" refString="Schwendinger, P. J. (1991) Two new trap-door spiders from Thailand (Araneae, Mygalomorphae, Idiopidae). Bulletin of the British arachnological Society, 8 (8), 233 - 240." title="Two new trap-door spiders from Thailand (Araneae, Mygalomorphae, Idiopidae)" type="journal article" year="1991">Schwendinger 1991: 235</bibRefCitation>
, fig. 12), and slightly less pigmented, tubuliform, convoluted stalk terminating in slightly enlarged, rounded head. The latter as large as or smaller than base, and distally distinctly less sclerotized and pigmented than stalk. Receptacular stalk and head bent dorsad (
<figureCitation id="93F62A01FFEF1D15FAD6458CFAEF8D50" box="[1289,1357,1523,1548]" captionStart="FIGURE 4" captionStartId="7.[151,250,1341,1363]" captionTargetBox="[159,1432,223,1282]" captionTargetId="figure@7.[151,1436,195,1320]" captionTargetPageId="7" captionText="FIGURE 4. Prothemenops irineae sp. n. A, right palpal organ of male holotype, retrolateral and slightly distal view. B, same, ventral and slightly distal view. C, left receptacle of exuvia of paratype, ental view (anterior is to the right, posterior to the left). D – G, vulvae of four female paratypes, dorsal view (D = allotype). Abbreviations: a = genital atrium, ae = apex of embolus, b = base of receptacle, be = base of embolus, dh = distal hematodocha, h = head of receptacle, pa = para-embolic apophysis, s = stalk of receptacle, st = subtegulum, t = tegulum, ue = uterus externus. Scale lines 0.1 mm (C), 0.5 mm (A – B, D – G)." httpUri="https://zenodo.org/record/250418/files/figure.png" pageId="6" pageNumber="536">Fig. 4</figureCitation>
C).
</paragraph>
<paragraph id="0B723684FFEF1D15FF184668FE7A8CE0" blockId="6.[151,1437,1307,1980]" pageId="6" pageNumber="536">
<emphasis id="39B9EA96FFEF1D15FF184668FE998D6C" bold="true" box="[199,315,1559,1584]" pageId="6" pageNumber="536">Variation.</emphasis>
Carapace lengths of seven males (and the six largest females in parentheses) range 5.1–7.0 (6.8–8.3), carapace widths 4.3–5.7 (5.3–6.3). In the
<typeStatus id="D4768826FFEF1D15FD034644FC9C8D08" box="[732,830,1595,1620]" pageId="6" pageNumber="536" type="holotype">holotype</typeStatus>
and allotype the AME is slightly larger than the ALE, but in most other conspecific specimens examined the AME is as large as or slightly smaller than the ALE. The posterior margin of the fovea has a more or less distinct median indentation in all males and four females, whereas in two females this is not visible (presumably hidden under the overlapping anterior margin of the fovea). Patella III of males has 5–8 prodorsal spines in an irregular row, that of females 6–12. Tarsus IV is aspinose in four males (including the
<typeStatus id="D4768826FFEF1D15FEE54690FE038C54" box="[314,417,1775,1800]" pageId="6" pageNumber="536" type="holotype">holotype</typeStatus>
); it has 0/2, 1/1 and 3/9 spines in the other three males. Tarsi I–III are aspinose in all males. The male from the second locality has a slightly smaller tibial apophysis on its palp and a less distinct para-embolic apophysis than the males from the first locality. All eight female
<typeStatus id="D4768826FFEF1D15FC5E4747FC4D8C0C" box="[897,1007,1848,1872]" pageId="6" pageNumber="536" type="paratype">paratypes</typeStatus>
, two other females which were killed and overgrown by the pathogenic fungus
<taxonomicName id="CCCD4D07FFEF1D15FDB04723FCF78C28" box="[623,853,1884,1908]" class="Arachnida" family="Idiopidae" genus="Nomourea" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="atypicola">
<emphasis id="39B9EA96FFEF1D15FDB04723FCF78C28" box="[623,853,1884,1908]" italics="true" pageId="6" pageNumber="536">Nomourea atypicola</emphasis>
</taxonomicName>
, and a subadult male have a thin scopula on tibia I (very indistinct in subadult male, distinct in all others) and on metatarsus III. Variation in the shape of the vulva of four females, see
<figureCitation id="93F62A01FFEF1D15FE8347DCFE3C8CE0" box="[348,414,1955,1980]" captionStart="FIGURE 4" captionStartId="7.[151,250,1341,1363]" captionTargetBox="[159,1432,223,1282]" captionTargetId="figure@7.[151,1436,195,1320]" captionTargetPageId="7" captionText="FIGURE 4. Prothemenops irineae sp. n. A, right palpal organ of male holotype, retrolateral and slightly distal view. B, same, ventral and slightly distal view. C, left receptacle of exuvia of paratype, ental view (anterior is to the right, posterior to the left). D – G, vulvae of four female paratypes, dorsal view (D = allotype). Abbreviations: a = genital atrium, ae = apex of embolus, b = base of receptacle, be = base of embolus, dh = distal hematodocha, h = head of receptacle, pa = para-embolic apophysis, s = stalk of receptacle, st = subtegulum, t = tegulum, ue = uterus externus. Scale lines 0.1 mm (C), 0.5 mm (A – B, D – G)." httpUri="https://zenodo.org/record/250418/files/figure.png" pageId="6" pageNumber="536">Fig. 4</figureCitation>
D–G.
</paragraph>
</subSubSection>
<caption id="5FB2660CFFEE1D14FF484542FA968E92" httpUri="https://zenodo.org/record/250418/files/figure.png" pageId="7" pageNumber="537" targetBox="[159,1432,223,1282]" targetPageId="7">
<paragraph id="0B723684FFEE1D14FF484542FA968E92" blockId="7.[151,1437,1341,1486]" pageId="7" pageNumber="537">
<emphasis id="39B9EA96FFEE1D14FF484542FEB18E0F" bold="true" box="[151,275,1341,1363]" pageId="7" pageNumber="537">FIGURE 4.</emphasis>
<taxonomicName id="CCCD4D07FFEE1D14FEC54542FE588E08" box="[282,506,1341,1364]" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="537" phylum="Arthropoda" rank="species" species="irineae" status="sp. nov.">
<emphasis id="39B9EA96FFEE1D14FEC54542FE588E08" box="[282,506,1341,1364]" italics="true" pageId="7" pageNumber="537">Prothemenops irineae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="228A57EDFFEE1D14FDDE4541FD9A8E0F" box="[513,568,1342,1363]" pageId="7" pageNumber="537" rank="species">sp. n.</taxonomicNameLabel>
A, right palpal organ of male holotype, retrolateral and slightly distal view. B, same, ventral and slightly distal view. C, left receptacle of exuvia of paratype, ental view (anterior is to the right, posterior to the left). D–G, vulvae of four female paratypes, dorsal view (D = allotype). Abbreviations: a = genital atrium, ae = apex of embolus, b = base of receptacle, be = base of embolus, dh = distal hematodocha, h = head of receptacle, pa = para-embolic apophysis, s = stalk of receptacle, st = subtegulum, t = tegulum, ue = uterus externus. Scale lines 0.1 mm (C), 0.5 mm (A–B, D–G).
</paragraph>
</caption>
<subSubSection id="43D7650FFFEE1D14FF18466BFD638D0C" pageId="7" pageNumber="537" type="discussion">
<paragraph id="0B723684FFEE1D14FF18466BFD638D0C" blockId="7.[151,1436,1556,2013]" pageId="7" pageNumber="537">
<emphasis id="39B9EA96FFEE1D14FF18466BFED28D71" bold="true" box="[199,368,1556,1581]" pageId="7" pageNumber="537">Relationships.</emphasis>
Strong morphological similarities in copulatory organs of both sexes show that this species is probably the closest relative of
<taxonomicName id="CCCD4D07FFEE1D14FE284646FDDC8D0C" box="[503,638,1592,1616]" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="537" phylum="Arthropoda" rank="species" species="khirikhan" status="sp. nov.">
<emphasis id="39B9EA96FFEE1D14FE284646FDDC8D0C" box="[503,638,1592,1616]" italics="true" pageId="7" pageNumber="537">P. khirikhan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="228A57EDFFEE1D14FD5A4646FD638D0C" box="[645,705,1593,1616]" pageId="7" pageNumber="537" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="43D7650FFFEE1D14FF184623FBD58DBD" pageId="7" pageNumber="537" type="distribution">
<paragraph id="0B723684FFEE1D14FF184623FBD58DBD" blockId="7.[151,1436,1556,2013]" pageId="7" pageNumber="537">
<emphasis id="39B9EA96FFEE1D14FF184623FEC28D29" bold="true" box="[199,352,1628,1653]" pageId="7" pageNumber="537">Distribution.</emphasis>
<taxonomicName id="CCCD4D07FFEE1D14FEB54623FDC18D28" box="[362,611,1628,1652]" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="537" phylum="Arthropoda" rank="species" species="irineae" status="sp. nov.">
<emphasis id="39B9EA96FFEE1D14FEB54623FDC18D28" box="[362,611,1628,1652]" italics="true" pageId="7" pageNumber="537">Prothemenops irineae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="228A57EDFFEE1D14FDB24622FD098D28" box="[621,683,1629,1652]" pageId="7" pageNumber="537" rank="species">sp. n.</taxonomicNameLabel>
was found at three sites (within a distance of
<quantity id="CC359B61FFEE1D14FB194622FAA68D29" box="[1222,1284,1628,1653]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="537" unit="km" value="3.0">3 km</quantity>
) in the small isolated limestone massif that comprises the Khao Sam Roi Yot National Park (
<figureCitation id="93F62A01FFEE1D14FB8A46FFFB0C8DC5" box="[1109,1198,1664,1689]" captionStart="FIGURE 12" captionStartId="18.[151,250,1064,1086]" captionTargetBox="[341,1244,196,1033]" captionTargetId="figure@18.[335,1251,193,1039]" captionTargetPageId="18" captionText="FIGURE 12. Map of Southeast Asia (without southernmost Thailand and without peninsular Malaysia) showing localities of the three species treated in here (squares with numbers) and the currently known geographic range of the genus Prothemenops (grey area). 1, P. khirikhan sp. n.; 2, P. irineae sp. n.; 3, P. phanthurat sp. n." httpUri="https://zenodo.org/record/250426/files/figure.png" pageId="7" pageNumber="537">Fig. 12</figureCitation>
, locality 2). Mature specimens are available from only two of these sites, but we believe that all
<taxonomicName id="CCCD4D07FFEE1D14FBE946DBFB758DE0" box="[1078,1239,1700,1724]" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="537" phylum="Arthropoda" rank="genus">
<emphasis id="39B9EA96FFEE1D14FBE946DBFB758DE0" box="[1078,1239,1700,1724]" italics="true" pageId="7" pageNumber="537">Prothemenops</emphasis>
</taxonomicName>
in that park are conspecific. The new species appears to be locally abundant but restricted to this massif.
</paragraph>
</subSubSection>
<subSubSection id="43D7650FFFEE1D1BFF184693FB608AE7" lastPageId="8" lastPageNumber="538" pageId="7" pageNumber="537" type="biology_ecology">
<paragraph id="0B723684FFEE1D14FF184693FB4F8CE5" blockId="7.[151,1436,1556,2013]" pageId="7" pageNumber="537">
<emphasis id="39B9EA96FFEE1D14FF184693FE858C59" bold="true" box="[199,295,1772,1797]" pageId="7" pageNumber="537">Biology.</emphasis>
The spiders were found on the steep slopes of limestone hills covered with a low, thorny secondary vegetation interspersed with introduced
<taxonomicName id="CCCD4D07FFEE1D14FDB1476EFC318C75" box="[622,915,1808,1833]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="7" pageNumber="537" phylum="Tracheophyta" rank="genus">
<emphasis id="39B9EA96FFEE1D14FDB1476EFD688C74" box="[622,714,1809,1832]" italics="true" pageId="7" pageNumber="537">Opuntia</emphasis>
(Euphorbiaceae)
</taxonomicName>
. They lived in burrows which were usually unbranched and closed by a single thin trapdoor (
<figureCitation id="93F62A01FFEE1D14FD1B474BFCB18C11" box="[708,787,1844,1869]" captionStart="FIGURE 11" captionStartId="17.[151,250,1461,1483]" captionTargetBox="[211,1378,186,1441]" captionTargetId="figure@17.[211,1379,183,1450]" captionTargetPageId="17" captionText="FIGURE 11. Burrows of Prothemenops khirikhan sp. n. (A – B), P. irineae sp. n. (C, F) and P. phanthurat sp. n. (D – E). A, intact straight burrow just below the soil surface (1 a = open front door, 1 b = closed hind door). B, three such burrows close to each other (1 a – 3 a front door, 1 b – 3 b hind door of each burrow); upper side of longest burrow (shown in A) removed to show its shallow course. C, typical Y-shaped burrow still intact, with its two doors open and distinctly apart from each other. D, Yshaped burrow (in captivity) dug up to show its below-ground shape. E, Y-shaped burrow running into limestone crack, with both doors close to each other. F, straight, deep, blind-ending large burrow with its single door closed." httpUri="https://zenodo.org/record/250425/files/figure.png" pageId="7" pageNumber="537">Fig. 11</figureCitation>
F) in large spiders, Y-shaped and closed by two trapdoors (
<figureCitation id="93F62A01FFEE1D14FF404727FF4C8C2D" box="[159,238,1880,1905]" captionStart="FIGURE 11" captionStartId="17.[151,250,1461,1483]" captionTargetBox="[211,1378,186,1441]" captionTargetId="figure@17.[211,1379,183,1450]" captionTargetPageId="17" captionText="FIGURE 11. Burrows of Prothemenops khirikhan sp. n. (A – B), P. irineae sp. n. (C, F) and P. phanthurat sp. n. (D – E). A, intact straight burrow just below the soil surface (1 a = open front door, 1 b = closed hind door). B, three such burrows close to each other (1 a – 3 a front door, 1 b – 3 b hind door of each burrow); upper side of longest burrow (shown in A) removed to show its shallow course. C, typical Y-shaped burrow still intact, with its two doors open and distinctly apart from each other. D, Yshaped burrow (in captivity) dug up to show its below-ground shape. E, Y-shaped burrow running into limestone crack, with both doors close to each other. F, straight, deep, blind-ending large burrow with its single door closed." httpUri="https://zenodo.org/record/250425/files/figure.png" pageId="7" pageNumber="537">Fig. 11</figureCitation>
C) in small and medium-sized spiders. The largest burrow (of a female) was about
<quantity id="CC359B61FFEE1D14FB404727FB478C2C" box="[1183,1253,1880,1905]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="537" unit="cm" value="15.0">15 cm</quantity>
long, and had a
<quantity id="CC359B61FFEE1D14FF484703FF538CC8" box="[151,241,1916,1941]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="7" pageNumber="537" unit="cm" value="1.75">1.75 cm</quantity>
long and
<quantity id="CC359B61FFEE1D14FEBE4702FE0F8CC8" box="[353,429,1917,1941]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="537" unit="cm" value="2.2">2.2 cm</quantity>
wide trapdoor. Adult males (all raised in captivity) had 1.00–
<quantity id="CC359B61FFEE1D14FBBC4703FB1E8CC8" box="[1123,1212,1916,1941]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="7" pageNumber="537" unit="cm" value="1.25">1.25 cm</quantity>
long and
<quantity id="CC359B61FFEE1D14FAF34703FF1B8CE4" metricMagnitude="-2" metricUnit="m" metricValue="1.275" metricValueMax="1.4" metricValueMin="1.15" pageId="7" pageNumber="537" unit="cm" value="1.275" valueMax="1.4" valueMin="1.15">1.15–1.40 cm</quantity>
wide trapdoors; two of these males had Y-shaped burrows with two trapdoors of different sizes.
</paragraph>
<paragraph id="0B723684FFEE1D1BFF1847BBFB608AE7" blockId="7.[151,1436,1556,2013]" lastBlockId="8.[151,1437,151,443]" lastPageId="8" lastPageNumber="538" pageId="7" pageNumber="537">
The first male matured at the end of
<date id="7F731044FFEE1D14FD8247BBFCAC8C81" box="[605,782,1988,2013]" pageId="7" pageNumber="537" value="2011-12">December 2011</date>
after less than six months in captivity; five others matured in August to October of the following year; the last one matured in late
<date id="7F731044FFE11D1BFC1540E8FBEA8BEC" box="[970,1096,151,176]" pageId="8" pageNumber="538" value="2014-04">April 2014</date>
, presumably an untimely date caused by conditions during 16 months of captivity. When disturbed, adult males and large females display an aggressive defence behaviour similar to that of theraphosid spiders: they raise their palps, first pair of legs and the prosoma (
<figureCitation id="93F62A01FFE11D1BFED44083FEF08A48" box="[267,338,252,277]" captionStart="FIGURE 3" captionStartId="6.[151,250,1215,1237]" captionTargetBox="[165,1433,183,1184]" captionTargetId="figure@6.[160,1442,175,1194]" captionTargetPageId="6" captionText="FIGURE 3. Prothemenops irineae sp. n., A – B, habitus of two males (B, male in defence posture). C – D, habitus of two females." httpUri="https://zenodo.org/record/250417/files/figure.png" pageId="8" pageNumber="538">Fig. 3</figureCitation>
B), and jump (only for a short distance) towards or even onto the object that threatens them. Occasionally, the spiders then land on their back. Moults of two large females were recorded in mid-September 2012 and in late
<date id="7F731044FFE11D1BFEBD4141FE468A0B" box="[354,484,318,343]" pageId="8" pageNumber="538" value="2013-04">April 2013</date>
.
<collectingCountry id="73DA7614FFE11D1BFE2D4140FDB38A0B" box="[498,529,319,343]" name="American Samoa" pageId="8" pageNumber="538">As</collectingCountry>
the spiders moult inside their burrows and then usually deposit their heavily fragmented exuviae at the bottom of the burrow, moults are rarely observed. In captivity two females were killed in their burrows by
<taxonomicName id="CCCD4D07FFE11D1BFE8A41FDFCA58AC5" authority="(Yasuda) Samson" authorityName="(Yasuda) Samson" box="[341,775,385,410]" class="Sordariomycetes" family="Clavicipitaceae" genus="Nomuraea" kingdom="Fungi" order="Hypocreales" pageId="8" pageNumber="538" phylum="Ascomycota" rank="species" species="atypicola">
<emphasis id="39B9EA96FFE11D1BFE8A41FDFD988AC5" box="[341,570,385,409]" italics="true" pageId="8" pageNumber="538">Nomuraea atypicola</emphasis>
(Yasuda) Samson
</taxonomicName>
, a common pathogenic fungus of spiders (
<bibRefCitation id="6F5C4B75FFE11D1BFB0241FEFF728AE7" author="Greenstone, M. H. &amp; Ignoffo, C. M. &amp; Samson, R. A." etAl="et al." firstAuthor="Greenstone" journalOrPublisher="The Journal of Arachnology" pageId="8" pageNumber="538" pagination="266 - 268" part="15" refId="ref13399" refString="Greenstone, M. H., Ignoffo, C. M. &amp; Samson, R. A. (1987) Susceptibility of spider species to the fungus Nomuraea atypicola. The Journal of Arachnology, 15, 266 - 268." title="Susceptibility of spider species to the fungus Nomuraea atypicola" type="journal article" year="1987">
Greenstone
<emphasis id="39B9EA96FFE11D1BFABB41FDFA3E8AC5" box="[1380,1436,385,409]" italics="true" pageId="8" pageNumber="538">et al.</emphasis>
1987
</bibRefCitation>
) which has also been observed in other
<taxonomicName id="CCCD4D07FFE11D1BFD4F41DCFC938AE7" box="[656,817,419,443]" class="Arachnida" family="Idiopidae" genus="Prothemenops" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="538" phylum="Arthropoda" rank="genus">
<emphasis id="39B9EA96FFE11D1BFD4F41DCFC938AE7" box="[656,817,419,443]" italics="true" pageId="8" pageNumber="538">Prothemenops</emphasis>
</taxonomicName>
species (
<bibRefCitation id="6F5C4B75FFE11D1BFC4841DDFB178AE7" author="Schwendinger, P. J." box="[919,1205,418,443]" firstAuthor="Schwendinger" journalOrPublisher="Revue suisse de Zoologie" page="579" pageId="8" pageNumber="538" pagination="577 - 584" part="2" refId="ref13618" refString="Schwendinger, P. J. (1996) The fauna of orthognathous spiders (Araneae: Mesothelae, Mygalomorphae) in Thailand. Revue suisse de Zoologie, 2 (Special Volume), 577 - 584." title="The fauna of orthognathous spiders (Araneae: Mesothelae, Mygalomorphae) in Thailand" type="journal article" year="1996">Schwendinger 1996: 579</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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