240 lines
19 KiB
XML
240 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.878.37403" ID-GBIF-Dataset="83f095d1-8441-49cd-be4b-71f0ddfa311c" ID-PMC="PMC6791902" ID-Pensoft-Pub="1313-2970-878-73" ID-Pensoft-UUID="C93F9CBB14D25FEAB2ACCBC3E6282D28" ID-PubMed="31632177" ID-ZooBank="192214DE1D38467BA577ECD16EC5EAB5" ModsDocID="1313-2970-878-73" checkinTime="1570497196116" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo & Balke, Michael" docDate="2019" docId="807019FED654537594C76E6B28173514" docLanguage="en" docName="ZooKeys 878: 73-143" docOrigin="ZooKeys 878" docSource="http://dx.doi.org/10.3897/zookeys.878.37403" docTitle="Exocelina erteldi Balke 1998" docType="treatment" docVersion="6" id="C93F9CBB14D25FEAB2ACCBC3E6282D28" lastPageNumber="73" masterDocId="C93F9CBB14D25FEAB2ACCBC3E6282D28" masterDocTitle="Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="143" masterPageNumber="73" pageNumber="73" updateTime="1668167854570" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shaverdo, Helena</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Surbakti, Suriani</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Warikar, Evie L.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sagata, Katayo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balke, Michael</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>878</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>73</mods:start>
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<mods:end>143</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.878.37403</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.878.37403</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-878-73</mods:identifier>
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<mods:identifier type="ZooBank">192214DE1D38467BA577ECD16EC5EAB5</mods:identifier>
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<mods:identifier type="Pensoft-UUID">C93F9CBB14D25FEAB2ACCBC3E6282D28</mods:identifier>
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||
</mods:mods>
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<treatment ID-GBIF-Taxon="159654833" LSID="urn:lsid:plazi:treatment:807019FED654537594C76E6B28173514" httpUri="http://treatment.plazi.org/id/807019FED654537594C76E6B28173514" lastPageNumber="73" pageId="0" pageNumber="73">
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<subSubSection pageId="0" pageNumber="73" type="nomenclature">
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<paragraph pageId="0" pageNumber="73">
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7.
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<taxonomicName LSID="807019FE-D654-5375-94C7-6E6B28173514" authority="Balke, 1998" authorityName="Balke" authorityYear="1998" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina erteldi" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="erteldi">Exocelina erteldi (Balke, 1998)</taxonomicName>
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<figureCitation captionStart="Figures 12–17" captionStartId="F8" captionText="Figures 12 - 17. Habitus and colouration 12 Exocelina oiwa sp. nov. 13 E. akameku sp. nov. 14 E. erteldi (Balke, 1998) 15 E. oksibilensis sp. nov. 16 E. bacchusi (Balke, 1998) 17 E. bacchusi herzogensis sp. nov." figureDoi="10.3897/zookeys.878.37403.figures12-17" httpUri="https://binary.pensoft.net/fig/344976" pageId="0" pageNumber="73" tableDoi="10.3897/zookeys.878.37403.figures12-17">Figs 14</figureCitation>
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,
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<figureCitation captionStart="Figures 20, 21" captionStartId="F10" captionText="Figures 20, 21. 20 Exocelina erteldi (Balke, 1998) 21 E. oksibilensis sp. nov. A median lobe in ventral view B median lobe in lateral view C paramere in external view D male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.878.37403.figures20-21" httpUri="https://binary.pensoft.net/fig/344978" pageId="0" pageNumber="73" tableDoi="10.3897/zookeys.878.37403.figures20-21">20</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="73" type="reference_group">
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<paragraph pageId="0" pageNumber="73">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus (Papuadytes) erteldi" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="erteldi" subGenus="Papuadytes">Copelatus (Papuadytes) erteldi</taxonomicName>
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Balke, 1998: 330;
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<bibRefCitation pageId="0" pageNumber="73" refId="B9">Nilsson 2001</bibRefCitation>
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: 76 (catalogue).
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</paragraph>
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<paragraph pageId="0" pageNumber="73">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Papuadytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papuadytes erteldi" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="erteldi">Papuadytes erteldi</taxonomicName>
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(Balke, 1998):
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<bibRefCitation author="Nilsson, AN" journalOrPublisher="Koleopterologische Rundschau" pageId="0" pageNumber="73" pagination="55 - 74" refId="B11" refString="Nilsson, AN, Fery, H, 2006. World Catalogue of Dytiscidae - corrections and additions, 3 (Coleoptera: Dytiscidae). Koleopterologische Rundschau 76: 55 - 74" title="World Catalogue of Dytiscidae - corrections and additions, 3 (Coleoptera: Dytiscidae)." volume="76" year="2006">Nilsson and Fery 2006</bibRefCitation>
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: 56 (comb. nov.).
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</paragraph>
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<paragraph pageId="0" pageNumber="73">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina erteldi" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="erteldi">Exocelina erteldi</taxonomicName>
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(Balke, 1998):
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<bibRefCitation author="Nilsson, AN" journalOrPublisher="Latissimus" pageId="0" pageNumber="73" pagination="33 - 34" refId="B10" refString="Nilsson, AN, 2007. Exocelina Broun, 1886, is the valid name of Papuadytes Balke, 1998. Latissimus 23: 33 - 34" title="Exocelina Broun, 1886, is the valid name of Papuadytes Balke, 1998." volume="23" year="2007">Nilsson 2007</bibRefCitation>
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: 33 (comb. nov.).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="73" type="type locality">
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<paragraph pageId="0" pageNumber="73">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="73">
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Indonesia: Papua Province: Pegunungan Bintang Regency, Borme, ca.
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<geoCoordinate degrees="04" direction="south" minutes="24" orientation="latitude" precision="925" value="-4.4">04°24'S</geoCoordinate>
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,
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<geoCoordinate degrees="140" direction="east" minutes="25" orientation="longitude" precision="925" value="140.41667">140°25'E</geoCoordinate>
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, 1200 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="73" type="type material">
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<paragraph pageId="0" pageNumber="73">Type material studied.</paragraph>
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<paragraph pageId="0" pageNumber="73">
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<emphasis italics="true" pageId="0" pageNumber="73">Holotype</emphasis>
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: male "IRIAN JAYA Zentralmassive
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<geoCoordinate degrees="140" direction="east" minutes="25" orientation="longitude" precision="925" value="140.41667">140°25'E</geoCoordinate>
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<geoCoordinate degrees="04" direction="south" minutes="24" orientation="latitude" precision="925" value="-4.4">04°24'S</geoCoordinate>
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", "14./17.8.1992 Borme, 1900m leg. Balke (11)", "
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
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<taxonomicName lsidName="erteldi" pageId="0" pageNumber="73" rank="species" species="erteldi">erteldi</taxonomicName>
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Balke des. 1997" [red],
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<normalizedToken originalValue="“HOLOTYPUS”">"HOLOTYPUS"</normalizedToken>
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[red] (NHMW).
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<emphasis italics="true" pageId="0" pageNumber="73">Paratypes</emphasis>
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: 1 female with the same label as the holotype and additionally with a red label "Paratypus
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
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<taxonomicName lsidName="erteldi" pageId="0" pageNumber="73" rank="species" species="erteldi">erteldi</taxonomicName>
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Balke des. 1997" (NHMW). 54 males, 35 females "IRIAN JAYA Zentralmassive
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<geoCoordinate degrees="140" direction="east" minutes="25" orientation="longitude" precision="925" value="140.41667">140°25'E</geoCoordinate>
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<geoCoordinate degrees="04" direction="south" minutes="24" orientation="latitude" precision="925" value="-4.4">04°24'S</geoCoordinate>
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", "Borme, 1800m 16.8.1992 leg. Balke (12, 12 A)", one of the males with two additional labels "M.Balke 3273" [green] and "M.Balke 6404 DNA" [green text], another male with an additional green label "M.Balke 3273" (CGW, NHMW). Note: in the original description (
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<bibRefCitation pageId="0" pageNumber="73" refId="B2">Balke 1998</bibRefCitation>
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), number of specimens of the locality (12, 12 A) is erroneously given as "43 males, 46 females". 1 female "12./18.8.1992 Borme, 100m leg. Balke (7)", "Paratypus
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
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<taxonomicName lsidName="erteldi" pageId="0" pageNumber="73" rank="species" species="erteldi">erteldi</taxonomicName>
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Balke des. 1997", this paratyte does not belong to species of
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<taxonomicName lsidName="E. erteldi" pageId="0" pageNumber="73" rank="species" species="erteldi">
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<emphasis italics="true" pageId="0" pageNumber="73">E. erteldi</emphasis>
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</taxonomicName>
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but to
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<taxonomicName lsidName="E. bifida" pageId="0" pageNumber="73" rank="species" species="bifida">
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<emphasis italics="true" pageId="0" pageNumber="73">E. bifida</emphasis>
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</taxonomicName>
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.250.3715" author="Shaverdo, HV" journalOrPublisher="ZooKeys" pageId="0" pageNumber="73" pagination="1 - 76" refId="B13" refString="Shaverdo, HV, Surbakti, S, Hendrich, L, Balke, M, 2012. Introduction of the Exocelinaekari-group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 250: 1 - 76" title="Introduction of the Exocelinaekari-group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)." url="https://doi.org/10.3897/zookeys.250.3715" volume="250" year="2012">Shaverdo et al. 2012</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="73" type="materials_examined">
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<paragraph pageId="0" pageNumber="73">Additional material.</paragraph>
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<paragraph pageId="0" pageNumber="73">
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1 male "IRIAN JAYA Zentralmassive
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<geoCoordinate degrees="140" direction="east" minutes="25" orientation="longitude" precision="925" value="140.41667">140°25'E</geoCoordinate>
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<geoCoordinate degrees="04" direction="south" minutes="24" orientation="latitude" precision="925" value="-4.4">04°24'S</geoCoordinate>
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", "Borme, 1800m 16.8.1992 leg. Balke (12, 12 A)", "Paratypus
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="73" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
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<taxonomicName lsidName="fume" pageId="0" pageNumber="73" rank="species" species="fume">fume</taxonomicName>
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Balke des. 1997" [red] (NHMW).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="73" type="diagnosis">
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<paragraph pageId="0" pageNumber="73">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="73">
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For complete description, see
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<bibRefCitation pageId="0" pageNumber="73" refId="B2">Balke (1998</bibRefCitation>
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: 330). Beetle small (TL-H 3.45-3.75 mm), oblong-oval; brown to piceous, usually with paler pronotal sides; dorsally more or less shiny, with fine but conspicuous punctation and weakly impressed microreticulation; pronotum with distinct lateral bead (
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<figureCitation captionStart="Figures 12–17" captionStartId="F8" captionText="Figures 12 - 17. Habitus and colouration 12 Exocelina oiwa sp. nov. 13 E. akameku sp. nov. 14 E. erteldi (Balke, 1998) 15 E. oksibilensis sp. nov. 16 E. bacchusi (Balke, 1998) 17 E. bacchusi herzogensis sp. nov." figureDoi="10.3897/zookeys.878.37403.figures12-17" httpUri="https://binary.pensoft.net/fig/344976" pageId="0" pageNumber="73" tableDoi="10.3897/zookeys.878.37403.figures12-17">Fig. 14</figureCitation>
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); male protarsomere 4 with anterolateral seta thin, weakly curved, smaller than more laterally situated large seta; male protarsomere 5 ventrally with anterior band of ca. 70 and posterior row of 6 relatively long setae (
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<figureCitation captionStart="Figures 20, 21" captionStartId="F10" captionText="Figures 20, 21. 20 Exocelina erteldi (Balke, 1998) 21 E. oksibilensis sp. nov. A median lobe in ventral view B median lobe in lateral view C paramere in external view D male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.878.37403.figures20-21" httpUri="https://binary.pensoft.net/fig/344978" pageId="0" pageNumber="73" tableDoi="10.3897/zookeys.878.37403.figures20-21">Fig. 20D</figureCitation>
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); median lobe in lateral view evenly attenuated to elongate, thin apex, which slightly pointed in ventral view; paramere slightly concave on dorsal side and with distinct, long, dense, uniform setae, situated along dorsal margin (
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<figureCitation captionStart="Figures 20, 21" captionStartId="F10" captionText="Figures 20, 21. 20 Exocelina erteldi (Balke, 1998) 21 E. oksibilensis sp. nov. A median lobe in ventral view B median lobe in lateral view C paramere in external view D male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.878.37403.figures20-21" httpUri="https://binary.pensoft.net/fig/344978" pageId="0" pageNumber="73" tableDoi="10.3897/zookeys.878.37403.figures20-21">
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Fig. 20
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="73" type="affinities">
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<paragraph pageId="0" pageNumber="73">Affinities.</paragraph>
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<paragraph pageId="0" pageNumber="73">
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The species can be distinguished from the species co-occurring in the same area (
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<taxonomicName lsidName="E. ascendens" pageId="0" pageNumber="73" rank="species" species="ascendens">
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<emphasis italics="true" pageId="0" pageNumber="73">E. ascendens</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="E. aipomek" pageId="0" pageNumber="73" rank="species" species="aipomek">
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<emphasis italics="true" pageId="0" pageNumber="73">E. aipomek</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="E. takime" pageId="0" pageNumber="73" rank="species" species="takime">
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<emphasis italics="true" pageId="0" pageNumber="73">E. takime</emphasis>
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</taxonomicName>
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, the
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<taxonomicName lsidName="E. ekari" pageId="0" pageNumber="73" rank="species" species="ekari">
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<emphasis italics="true" pageId="0" pageNumber="73">E. ekari</emphasis>
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</taxonomicName>
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group:
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<taxonomicName lsidName="E. eme" pageId="0" pageNumber="73" rank="species" species="eme">
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<emphasis italics="true" pageId="0" pageNumber="73">E. eme</emphasis>
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</taxonomicName>
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Shaverdo and
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<taxonomicName lsidName="E. bifida" pageId="0" pageNumber="73" rank="species" species="bifida">
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<emphasis italics="true" pageId="0" pageNumber="73">E. bifida</emphasis>
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</taxonomicName>
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, the
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<taxonomicName lsidName="E. danae" pageId="0" pageNumber="73" rank="species" species="danae">
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<emphasis italics="true" pageId="0" pageNumber="73">E. danae</emphasis>
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</taxonomicName>
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group:
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<taxonomicName lsidName="E. damantiensis" pageId="0" pageNumber="73" rank="species" species="damantiensis">
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<emphasis italics="true" pageId="0" pageNumber="73">E. damantiensis</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="E. danae" pageId="0" pageNumber="73" rank="species" species="danae">
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<emphasis italics="true" pageId="0" pageNumber="73">E. danae</emphasis>
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</taxonomicName>
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, the
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<taxonomicName lsidName="E. okbapensis" pageId="0" pageNumber="73" rank="species" species="okbapensis">
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<emphasis italics="true" pageId="0" pageNumber="73">E. okbapensis</emphasis>
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</taxonomicName>
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group:
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<taxonomicName lsidName="E. ketembang" pageId="0" pageNumber="73" rank="species" species="ketembang">
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<emphasis italics="true" pageId="0" pageNumber="73">E. ketembang</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="E. talaki" pageId="0" pageNumber="73" rank="species" species="talaki">
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<emphasis italics="true" pageId="0" pageNumber="73">E. talaki</emphasis>
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</taxonomicName>
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, and
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<taxonomicName lsidName="E. okbapensis" pageId="0" pageNumber="73" rank="species" species="okbapensis">
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<emphasis italics="true" pageId="0" pageNumber="73">E. okbapensis</emphasis>
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</taxonomicName>
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, and all species of the
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<taxonomicName lsidName="E. aipo" pageId="0" pageNumber="73" rank="species" species="aipo">
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<emphasis italics="true" pageId="0" pageNumber="73">E. aipo</emphasis>
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</taxonomicName>
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group) by body size and colouration, presence of pronotal bead, fine but conspicuous dorsal punctation, and shape and setation of its median lobe, paramere, and male protarsomere 4. For the affinities within the group, see the
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<normalizedToken originalValue="“Key”">"Key"</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="73" type="distribution">
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<paragraph pageId="0" pageNumber="73">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="73">
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Indonesia: Papua Province: Pegunungan Bintang Regency, Borme (
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<figureCitation captionStart="Figure 25" captionStartId="F13" captionText="Figure 25. Map of the eastern part of New Guinea showing distribution of the species of the E. bacchusi group." figureDoi="10.3897/zookeys.878.37403.figure25" httpUri="https://binary.pensoft.net/fig/344981" pageId="0" pageNumber="73" tableDoi="10.3897/zookeys.878.37403.figure25">Fig. 25</figureCitation>
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). The species is known only from the type material.
|
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |