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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.888.34494" ID-GBIF-Dataset="da6676c3-83a5-4231-85d7-0a8411a98a7f" ID-PMC="PMC6861337" ID-Pensoft-Pub="1313-2970-888-1" ID-Pensoft-UUID="F8810409F4DA5A43BF94417F5D40DECE" ID-PubMed="31754317" ID-ZooBank="B995C05697EC41A49012B58F9D3AFDC1" ModsDocID="1313-2970-888-1" checkinTime="1573555956936" checkinUser="pensoft" docAuthor="Xu, Xin, Ono, Hirotsugu, Kuntner, Matjaz, Liu, Fengxiang &amp; Li, Daiqin" docDate="2019" docId="7F077D2BA4FC5DCC95C6D09E666966EF" docLanguage="en" docName="ZooKeys 888: 1-50" docOrigin="ZooKeys 888" docSource="http://dx.doi.org/10.3897/zookeys.888.34494" docTitle="Heptathela uken Xu, Ono, Kuntner, Liu &amp; Li, 2019, sp. nov." docType="treatment" docUuid="C538969C-60CB-4184-B388-E8958807AF33" docUuidSource="ZooBank" docVersion="5" id="F8810409F4DA5A43BF94417F5D40DECE" lastPageNumber="1" masterDocId="F8810409F4DA5A43BF94417F5D40DECE" masterDocTitle="A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands" masterLastPageNumber="50" masterPageNumber="1" pageNumber="1" updateTime="1668168046841" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A taxonomic monograph of the liphistiid spider genus Heptathela, endemic to Japanese islands</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Xu, Xin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ono, Hirotsugu</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kuntner, Matjaz</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu, Fengxiang</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Daiqin</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>888</mods:number>
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<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>50</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.888.34494</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.888.34494</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-888-1</mods:identifier>
<mods:identifier type="ZooBank">B995C05697EC41A49012B58F9D3AFDC1</mods:identifier>
<mods:identifier type="Pensoft-UUID">F8810409F4DA5A43BF94417F5D40DECE</mods:identifier>
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<treatment ID-GBIF-Taxon="160342577" LSID="urn:lsid:zoobank.org:act:C538969C-60CB-4184-B388-E8958807AF33" httpUri="http://treatment.plazi.org/id/7F077D2BA4FC5DCC95C6D09E666966EF" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="7F077D2B-A4FC-5DCC-95C6-D09E666966EF" class="Arachnida" family="Liphistiidae" genus="Heptathela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptathela uken" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="uken">Heptathela uken</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male and female genital anatomy of Heptathela uken sp. nov. A-G 3297 (holotype, short for XUX- 2013 - 297) H, K, J 3301 I, L, M 3309 A palp prolateral view B palp ventral view C palp retrolateral view D-G palp distal view H, I vulva dorsal view K, L vulva ventral view J, M vulva distal view; 3297, 3301: Uken-son, Amamioshima; 3309: Yamato-son, Amamioshima. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.888.34494.figure13" httpUri="https://binary.pensoft.net/fig/354504" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.888.34494.figure13">Fig. 13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: JAPAN · ♂; Kagoshima-ken, Amami, Uken-son, Oshima-gun, Road No. 85, Redsoil Park;
<geoCoordinate degrees="28.24" direction="north" orientation="latitude" precision="555" value="28.24">28.24N</geoCoordinate>
,
<geoCoordinate degrees="129.34" direction="east" orientation="longitude" precision="555" value="129.34">129.34E</geoCoordinate>
; alt. 260 m; 15 September 2013; D. Li and B. Wu leg.; XUX-2013-297.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: JAPAN · 2 ♂♂, 2 ♀♀; same data as for holotype; XUX-2013-298, 301, 302, 304 · 3 ♂♂, 5 ♀♀; Kagoshima-ken, Amami, Yamato-son, Amami Forest Park;
<geoCoordinate degrees="28.31" direction="north" orientation="latitude" precision="555" value="28.31">28.31N</geoCoordinate>
,
<geoCoordinate degrees="129.33" direction="east" orientation="longitude" precision="555" value="129.33">129.33E</geoCoordinate>
; alt. 300 m; 17 September 2013; D. Li and B. Wu leg.; XUX-2013-305 to 314.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Males of
<taxonomicName lsidName="H. uken" pageId="0" pageNumber="1" rank="species" species="uken">
<emphasis italics="true" pageId="0" pageNumber="1">H. uken</emphasis>
</taxonomicName>
sp. nov. can be distinguished from those of
<taxonomicName lsidName="H. kanenoi" pageId="0" pageNumber="1" rank="species" species="kanenoi">
<emphasis italics="true" pageId="0" pageNumber="1">H. kanenoi</emphasis>
</taxonomicName>
by the spiniform conductor apex (
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male and female genital anatomy of Heptathela uken sp. nov. A-G 3297 (holotype, short for XUX- 2013 - 297) H, K, J 3301 I, L, M 3309 A palp prolateral view B palp ventral view C palp retrolateral view D-G palp distal view H, I vulva dorsal view K, L vulva ventral view J, M vulva distal view; 3297, 3301: Uken-son, Amamioshima; 3309: Yamato-son, Amamioshima. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.888.34494.figure13" httpUri="https://binary.pensoft.net/fig/354504" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.888.34494.figure13">Fig. 13A, B, D, E</figureCitation>
), from those of
<taxonomicName lsidName="H. amamiensis" pageId="0" pageNumber="1" rank="species" species="amamiensis">
<emphasis italics="true" pageId="0" pageNumber="1">H. amamiensis</emphasis>
</taxonomicName>
by the dorsal extension of tegular terminal apophysis without dentation (
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male and female genital anatomy of Heptathela uken sp. nov. A-G 3297 (holotype, short for XUX- 2013 - 297) H, K, J 3301 I, L, M 3309 A palp prolateral view B palp ventral view C palp retrolateral view D-G palp distal view H, I vulva dorsal view K, L vulva ventral view J, M vulva distal view; 3297, 3301: Uken-son, Amamioshima; 3309: Yamato-son, Amamioshima. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.888.34494.figure13" httpUri="https://binary.pensoft.net/fig/354504" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.888.34494.figure13">Fig. 13F, G</figureCitation>
). Females of
<taxonomicName lsidName="H. uken" pageId="0" pageNumber="1" rank="species" species="uken">
<emphasis italics="true" pageId="0" pageNumber="1">H. uken</emphasis>
</taxonomicName>
sp. nov. cannot be diagnosed from those of the other Amami group
<taxonomicName class="Arachnida" family="Liphistiidae" genus="Heptathela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptathela" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Heptathela</emphasis>
</taxonomicName>
species morphologically (
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male and female genital anatomy of Heptathela uken sp. nov. A-G 3297 (holotype, short for XUX- 2013 - 297) H, K, J 3301 I, L, M 3309 A palp prolateral view B palp ventral view C palp retrolateral view D-G palp distal view H, I vulva dorsal view K, L vulva ventral view J, M vulva distal view; 3297, 3301: Uken-son, Amamioshima; 3309: Yamato-son, Amamioshima. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.888.34494.figure13" httpUri="https://binary.pensoft.net/fig/354504" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.888.34494.figure13">
Fig. 13
<normalizedToken originalValue="HM">H-M</normalizedToken>
</figureCitation>
), only by the following unique nucleotide substitutions in the standard DNA barcode alignment: T (161), T (191), G (227), C (236), T (287), T (297), A (299), C (389), T (395), G (413), C (416), G (503), C (509), C (510), T (527), T (558), G (560), G (569), C (578), T (584), C (596), T (614), G (629), G (635), C (650), C (665).
</paragraph>
<caption doi="10.3897/zookeys.888.34494.figure13" httpUri="https://binary.pensoft.net/fig/354504" pageId="0" pageNumber="1" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 13.</emphasis>
Male and female genital anatomy of
<taxonomicName class="Arachnida" family="Liphistiidae" genus="Heptathela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heptathela uken" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="uken">
<emphasis italics="true" pageId="0" pageNumber="1">Heptathela uken</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="AG">A-G</normalizedToken>
</emphasis>
3297 (holotype, short for XUX-2013-297)
<emphasis bold="true" pageId="0" pageNumber="1">H, K, J</emphasis>
3301
<emphasis bold="true" pageId="0" pageNumber="1">I, L, M</emphasis>
3309
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
palp prolateral view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
palp ventral view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
palp retrolateral view
<emphasis bold="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="DG">D-G</normalizedToken>
</emphasis>
palp distal view
<emphasis bold="true" pageId="0" pageNumber="1">H, I</emphasis>
vulva dorsal view
<emphasis bold="true" pageId="0" pageNumber="1">K, L</emphasis>
vulva ventral view
<emphasis bold="true" pageId="0" pageNumber="1">J, M</emphasis>
vulva distal view; 3297, 3301: Uken-son, Amamioshima; 3309: Yamato-son, Amamioshima. Scale bar: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Male</emphasis>
(Holotype). Carapace brown; opisthosoma light brown, with dark brown tergites; cheliceral groove with eight denticles; seven spinnerets. Measurements: BL 11.60, CL 6.10, CW 5.60, OL 6.00, OW 4.30; ALE&gt; PLE&gt; PME&gt; AME; leg I 17.47 (4.80 + 2.28 + 3.67+ 4.42 + 2.30), leg II 17.95 (4.65 + 2.30 + 3.60 + 4.90 + 2.50), leg III 18.80 (4.45 + 2.40 + 3.55 + 5.20 + 3.20), leg IV 23.60 (5.60 + 2.45 + 4.45 + 7.20 + 3.90).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Palp</emphasis>
.
</emphasis>
Prolateral side of paracymbium unpigmented and unsclerotised, numerous setae and spines at the tip of paracymbium (
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male and female genital anatomy of Heptathela uken sp. nov. A-G 3297 (holotype, short for XUX- 2013 - 297) H, K, J 3301 I, L, M 3309 A palp prolateral view B palp ventral view C palp retrolateral view D-G palp distal view H, I vulva dorsal view K, L vulva ventral view J, M vulva distal view; 3297, 3301: Uken-son, Amamioshima; 3309: Yamato-son, Amamioshima. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.888.34494.figure13" httpUri="https://binary.pensoft.net/fig/354504" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.888.34494.figure13">
Fig. 13
<normalizedToken originalValue="AC">A-C</normalizedToken>
</figureCitation>
). Contrategulum with serrated margin (
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male and female genital anatomy of Heptathela uken sp. nov. A-G 3297 (holotype, short for XUX- 2013 - 297) H, K, J 3301 I, L, M 3309 A palp prolateral view B palp ventral view C palp retrolateral view D-G palp distal view H, I vulva dorsal view K, L vulva ventral view J, M vulva distal view; 3297, 3301: Uken-son, Amamioshima; 3309: Yamato-son, Amamioshima. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.888.34494.figure13" httpUri="https://binary.pensoft.net/fig/354504" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.888.34494.figure13">Fig. 13A, B, D, F</figureCitation>
). Tegulum with smooth dorsal extension of terminal apophysis (
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male and female genital anatomy of Heptathela uken sp. nov. A-G 3297 (holotype, short for XUX- 2013 - 297) H, K, J 3301 I, L, M 3309 A palp prolateral view B palp ventral view C palp retrolateral view D-G palp distal view H, I vulva dorsal view K, L vulva ventral view J, M vulva distal view; 3297, 3301: Uken-son, Amamioshima; 3309: Yamato-son, Amamioshima. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.888.34494.figure13" httpUri="https://binary.pensoft.net/fig/354504" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.888.34494.figure13">Fig. 13F, G</figureCitation>
). Conductor wide, sclerotised and rugose, with several folds and a spiniform apex (
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male and female genital anatomy of Heptathela uken sp. nov. A-G 3297 (holotype, short for XUX- 2013 - 297) H, K, J 3301 I, L, M 3309 A palp prolateral view B palp ventral view C palp retrolateral view D-G palp distal view H, I vulva dorsal view K, L vulva ventral view J, M vulva distal view; 3297, 3301: Uken-son, Amamioshima; 3309: Yamato-son, Amamioshima. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.888.34494.figure13" httpUri="https://binary.pensoft.net/fig/354504" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.888.34494.figure13">Fig. 13A, B, D, F</figureCitation>
). Embolus sclerotised, wide with a flat opening, and wide saddle-shaped in the prolateral view (
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male and female genital anatomy of Heptathela uken sp. nov. A-G 3297 (holotype, short for XUX- 2013 - 297) H, K, J 3301 I, L, M 3309 A palp prolateral view B palp ventral view C palp retrolateral view D-G palp distal view H, I vulva dorsal view K, L vulva ventral view J, M vulva distal view; 3297, 3301: Uken-son, Amamioshima; 3309: Yamato-son, Amamioshima. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.888.34494.figure13" httpUri="https://binary.pensoft.net/fig/354504" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.888.34494.figure13">Fig. 13A, B, D, E</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Females</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 8). Carapace and opisthosoma colour as in male; cheliceral groove with 13-15 pronounced denticles; tergites similar to male; 7 or 8 spinnerets. Measurements: BL 11.80-16.00, CL 5.90-8.26, CW 5.20-7.20, OL 6.00-8.10, OW 4.60-6.00; ALE&gt; PLE&gt; PME&gt; AME; palp 10.04 (3.05 + 1.89 + 2.30 + 2.80), leg I 12.05 (3.90 + 2.10 + 2.25 + 2.40 + 1.40), leg II 11.30 (3.55 + 2.05 + 1.80+ 2.45 + 1.45), leg III 12.42 (3.60 + 2.05 + 2.10 + 3.00 + 1.67), leg IV 18.33 (5.15 + 2.40 + 3.30 + 4.78 + 2.70).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Female genitalia</emphasis>
.
</emphasis>
A pair of indistinct depressions on the ventro-lateral part of genital atrium (
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male and female genital anatomy of Heptathela uken sp. nov. A-G 3297 (holotype, short for XUX- 2013 - 297) H, K, J 3301 I, L, M 3309 A palp prolateral view B palp ventral view C palp retrolateral view D-G palp distal view H, I vulva dorsal view K, L vulva ventral view J, M vulva distal view; 3297, 3301: Uken-son, Amamioshima; 3309: Yamato-son, Amamioshima. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.888.34494.figure13" httpUri="https://binary.pensoft.net/fig/354504" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.888.34494.figure13">Fig. 13K</figureCitation>
). Two pairs of receptacular clusters along the anterior margin of bursa copulatrix, the inners almost globose, with short genital stalks in distal view, the laterals tuberculate, without genital stalks (
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Male and female genital anatomy of Heptathela uken sp. nov. A-G 3297 (holotype, short for XUX- 2013 - 297) H, K, J 3301 I, L, M 3309 A palp prolateral view B palp ventral view C palp retrolateral view D-G palp distal view H, I vulva dorsal view K, L vulva ventral view J, M vulva distal view; 3297, 3301: Uken-son, Amamioshima; 3309: Yamato-son, Amamioshima. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.888.34494.figure13" httpUri="https://binary.pensoft.net/fig/354504" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.888.34494.figure13">
Fig. 13
<normalizedToken originalValue="HM">H-M</normalizedToken>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">The species epithet, a noun in apposition, refers to the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
The species is known from the Japanese island Amamioshima (
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General somatic morphology of Heptathela kimurai (Kishida, 1920) A dorsal view B ventral view and map showing the sampling localities of Heptathela specimens from Kyushu to central Ryukyus (C)." figureDoi="10.3897/zookeys.888.34494.figure1" httpUri="https://binary.pensoft.net/fig/354492" pageId="0" pageNumber="1" tableDoi="10.3897/zookeys.888.34494.figure1">Fig. 1C</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>