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<document id="F3ACEB8C6C3C42D1A7B3D0AC83FDFF82" ID-CLB-Dataset="30789" ID-DOI="10.5852/ejt.2018.402" ID-GBIF-Dataset="8f1510f6-1b79-467f-9e8d-f193ac82cc4b" ID-Zenodo-Dep="1169559" ID-ZooBank="A3CE68AA-C5C6-40B7-B57C-6EF6D949149B" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1518136191129" checkinUser="plazi" docAuthor="Ruiz-C, Raquel I., Román-Valencia, César, Taphorn, Donald C., Buckup, Paulo A. &amp; Ortega, Hernán" docDate="2018" docId="2C0787E7FFD8D423FDDFFDA8FD46FA9E" docLanguage="en" docName="ejt-402_Ruiz-C_Roman-Valencia.pdf.imf" docOrigin="European Journal of Taxonomy 402" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Astyanax moorii Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega 2018, comb. nov." docType="treatment" docVersion="17" lastPageNumber="32" masterDocId="D03EFF9FFFC5D43CFFDCFFD0FE0FFFBE" masterDocTitle="Revision of the Astyanax orthodus species-group (Teleostei: Characidae) with descriptions of three new species" masterLastPageNumber="45" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1547298528796" originalUpdateUser="yanell" pageNumber="30" updateTime="1698520653973" updateUser="ExternalLinkService">
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<mods:title id="65FDAF3627C8E5E184F651332DABC9B1">Revision of the Astyanax orthodus species-group (Teleostei: Characidae) with descriptions of three new species</mods:title>
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<mods:namePart id="428B0ED14314217109F772671E3B6B8B">Ruiz-C, Raquel I.</mods:namePart>
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<mods:namePart id="680DD0DCDEB0C91C0FFC0E0076DC13D6">Román-Valencia, César</mods:namePart>
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<mods:namePart id="1EDE4B5F42F445FAB30D24E4EED00456">Taphorn, Donald C.</mods:namePart>
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<mods:namePart id="230388061098840A9C670117A9FA4101">Buckup, Paulo A.</mods:namePart>
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<mods:namePart id="6E4A1402CFF109E80544E1C241BB30CF">Ortega, Hernán</mods:namePart>
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<mods:title id="5F7D68E6A526D17CE6A5D03142CA0F8F">European Journal of Taxonomy</mods:title>
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<mods:date id="2A37F736E0211F77208250BA1EFF3BE5">2018</mods:date>
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<mods:number id="BE856E92A97261ACD61D1DD5A00EB0B5">2018-02-08</mods:number>
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<mods:number id="8FE48B3A1C632381AA5C39FC2775F702">402</mods:number>
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<treatment id="2C0787E7FFD8D423FDDFFDA8FD46FA9E" ID-DOI="http://doi.org/10.5281/zenodo.5622624" ID-GBIF-Taxon="140562868" ID-Zenodo-Dep="5622624" LSID="urn:lsid:plazi:treatment:2C0787E7FFD8D423FDDFFDA8FD46FA9E" httpUri="http://treatment.plazi.org/id/2C0787E7FFD8D423FDDFFDA8FD46FA9E" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30">
<subSubSection id="ECB4657AFFD8D421FDDFFDA8FA3EFD2D" box="[515,1073,632,659]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="A41136F1FFD8D421FDDFFDA8FA3EFD2D" blockId="29.[515,1073,632,695]" box="[515,1073,632,659]" pageId="29" pageNumber="30">
<heading id="FF59819DFFD8D421FDDFFDA8FA3EFD2D" box="[515,1073,632,659]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="30" reason="2">
<taxonomicName id="63AE4D72FFD8D421FDDFFDA8FDA7FD2C" authority="(Boulenger, 1892)" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" baseAuthorityName="Boulenger" baseAuthorityYear="1892" box="[515,936,632,658]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="moorii" status="comb. nov.">
<emphasis id="96DAEAE3FFD8D421FDDFFDA8FCC4FD2C" bold="true" box="[515,715,632,658]" italics="true" pageId="29" pageNumber="30">Astyanax moorii</emphasis>
(
<bibRefCitation id="C03F4B00FFD8D421FD06FDA8FD90FD2C" author="Boulenger G. A." box="[730,927,632,658]" journalOrPublisher="Annals and Magazine of Natural History (Series 6)" pageId="29" pageNumber="30" pagination="9 - 12" part="10 (55)" refId="ref19010" refString="Boulenger G. A. 1892. On some new or little-known fishes obtained by Dr. J. W. Evans and Mr. Spencer Moore during their recent expedition to the Province of Matto Grosso, Brazil. Annals and Magazine of Natural History (Series 6) 10 (55) (art. 2): 9 - 12." title="On some new or little-known fishes obtained by Dr. J. W. Evans and Mr. Spencer Moore during their recent expedition to the Province of Matto Grosso, Brazil." type="journal article" year="1892">Boulenger, 1892</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFD8D421FC73FDA8FA3EFD2D" box="[943,1073,632,659]" pageId="29" pageNumber="30">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ECB4657AFFD8D421FD71FD4DFD88FD09" box="[685,903,669,695]" pageId="29" pageNumber="30" type="description">
<paragraph id="A41136F1FFD8D421FD71FD4DFD88FD09" blockId="29.[515,1073,632,695]" box="[685,903,669,695]" pageId="29" pageNumber="30">
<figureCitation id="3C952A74FFD8D421FD71FD4DFCF7FD09" box="[685,760,669,695]" captionStart="Fig. 4" captionStartId="8.[189,232,2000,2026]" captionTargetBox="[189,1392,764,1921]" captionTargetId="figure@8.[151,1417,755,1932]" captionTargetPageId="8" captionText="Fig. 4.Distribution of speciesof the Astyanaxorthodus species-group(symbol includestypelocalities)." httpUri="https://zenodo.org/record/1169567/files/figure.png" pageId="29" pageNumber="30">Figs 4</figureCitation>
,
<figureCitation id="3C952A74FFD8D421FCD8FD4DFD2CFD09" box="[772,803,669,695]" captionStart="Fig. 13" captionStartId="30.[189,232,1929,1955]" captionTargetBox="[189,1398,1393,1892]" captionTargetId="figure@30.[189,1398,1393,1892]" captionTargetPageId="30" captionText="Fig. 13. Astyanax moorii (Boulenger, 1892), 64.7 mm SL, Creek near the Manso Hydroelectric Plant, Cuiabá River drainage, Fazenda Nova region, Mato Grosso State, Brazil (MNRJ 29284). Scale bar = 1 cm." httpUri="https://zenodo.org/record/1169585/files/figure.png" pageId="29" pageNumber="30">13</figureCitation>
,
<tableCitation id="E92C034AFFD8D421FCECFD4DFD88FD09" box="[816,903,669,695]" captionStart="Table 2" captionStartId="17.[145,214,609,635]" captionTargetPageId="17" captionText="Table 2. Morphometric data for some species of the Astyanax orthodus species-group." httpUri="http://table.plazi.org/id/F0D16679FFD4D42DFF4DFDB1FA88FDC5" pageId="29" pageNumber="30" tableUuid="F0D16679FFD4D42DFF4DFDB1FA88FDC5">Table 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="ECB4657AFFD8D421FF61FD31FD58FC37" pageId="29" pageNumber="30" type="reference_group">
<paragraph id="A41136F1FFD8D421FF61FD31FC87FCA1" blockId="29.[189,1398,736,800]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="84BE11DFFFD8D421FF61FD31FC87FCA1" pageId="29" pageNumber="30">
<taxonomicName id="63AE4D72FFD8D421FF61FD31FCDBFD45" authority="Boulenger, 1892: 11" authorityName="Boulenger" authorityPageNumber="11" authorityYear="1892" box="[189,724,736,763]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="moorii">
<emphasis id="96DAEAE3FFD8D421FF61FD31FFDAFD44" box="[189,469,737,762]" italics="true" pageId="29" pageNumber="30">Tetragonopterus moorii</emphasis>
<treatmentCitation id="250F10E0FFD8D421FE03FD30FCDBFD45" author="" box="[479,724,736,763]" page="11" pageId="29" pageNumber="30">
<bibRefCitation id="C03F4B00FFD8D421FE03FD30FCABFD45" author="Boulenger G. A." box="[479,676,736,763]" journalOrPublisher="Annals and Magazine of Natural History (Series 6)" pageId="29" pageNumber="30" pagination="9 - 12" part="10 (55)" refId="ref19010" refString="Boulenger G. A. 1892. On some new or little-known fishes obtained by Dr. J. W. Evans and Mr. Spencer Moore during their recent expedition to the Province of Matto Grosso, Brazil. Annals and Magazine of Natural History (Series 6) 10 (55) (art. 2): 9 - 12." title="On some new or little-known fishes obtained by Dr. J. W. Evans and Mr. Spencer Moore during their recent expedition to the Province of Matto Grosso, Brazil." type="journal article" year="1892">Boulenger, 1892</bibRefCitation>
: 11
</treatmentCitation>
</taxonomicName>
, pl. 2 fig. 1. Original description,
<typeStatus id="7B158853FFD8D421FBADFD31FAABFD45" box="[1137,1188,737,763]" pageId="29" pageNumber="30">type</typeStatus>
locality:
<location id="A171602AFFD8D421FACAFD30FF4CFCA1" LSID="urn:lsid:plazi:treatment:2C0787E7FFD8D423FDDFFDA8FD46FA9E:A171602AFFD8D421FACAFD30FF4CFCA1" country="Brazil" name="Chapala Plateau" pageId="29" pageNumber="30" stateProvince="Mato Grosso">Chapala Plateau</location>
,
<collectingRegion id="666AF813FFD8D421FE93FCD6FFE7FC9E" box="[335,488,774,800]" country="Brazil" name="Mato Grosso" pageId="29" pageNumber="30">Mato Grosso</collectingRegion>
State,
<collectingCountry id="DCB97661FFD8D421FDE4FCD5FC8CFCA1" box="[568,643,773,799]" name="Brazil" pageId="29" pageNumber="30">Brazil</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A41136F1FFD8D421FF61FC99FD58FC37" blockId="29.[189,1398,841,905]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="84BE11DFFFD8D421FF61FC99FD58FC37" pageId="29" pageNumber="30">
<taxonomicName id="63AE4D72FFD8D421FF61FC99FFA1FCDD" ID-CoL="J5PK" baseAuthorityName="Boulenger" baseAuthorityYear="1892" box="[189,430,841,867]" class="Actinopterygii" family="Characidae" genus="Astyanacinus" kingdom="Animalia" order="Characiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="moorii">
<emphasis id="96DAEAE3FFD8D421FF61FC99FFA1FCDD" box="[189,430,841,867]" italics="true" pageId="29" pageNumber="30">Astyanacinus moorii</emphasis>
</taxonomicName>
–
<treatmentCitation id="250F10E0FFD8D421FE17FC99FCDEFCDA" author="" box="[459,721,841,868]" page="435" pageId="29" pageNumber="30">
<bibRefCitation id="C03F4B00FFD8D421FE17FC99FC99FCDD" author="Eigenmann C." box="[459,662,841,867]" editor="Scott W. B." journalOrPublisher="Indiana University, Bloomington, USA" pageId="29" pageNumber="30" pagination="375 - 511" publicationUrl="https://doi.org/10.5962/bhl.title.2097" refId="ref19165" refString="Eigenmann C. 1910. Catalogue of freshwater fishes of tropical and South temperate America. In: Scott W. B. (ed.) Reports of the Princeton University Expedition to Patagonia, 1896 - 1899. Volume 3 - Zoology: 375 - 511, Indiana University, Bloomington, USA. https: // doi. org / 10.5962 / bhl. title. 2097" title="Catalogue of freshwater fishes of tropical and South temperate America" type="book chapter" volumeTitle="Reports of the Princeton University Expedition to Patagonia, 1896 - 1899. Volume 3 - Zoology" year="1910">Eigenmann 1910</bibRefCitation>
: 435
</treatmentCitation>
;
<treatmentCitation id="250F10E0FFD8D421FD00FC99FDA5FCDD" author="" box="[732,938,841,867]" page="41" pageId="29" pageNumber="30">
<bibRefCitation id="C03F4B00FFD8D421FD00FC99FD70FCDD" author="Pearson N. E." box="[732,895,841,867]" journalOrPublisher="Indiana University Studies" pageId="29" pageNumber="30" part="11" refId="ref20211" refString="Pearson N. E. 1924. The Fishes of the Eastern Slope of the Andes. Indiana University Studies 11, Indiana University, Bloomington, USA." title="The Fishes of the Eastern Slope of the Andes." type="journal article" year="1924">Pearson 1924</bibRefCitation>
: 41
</treatmentCitation>
. (
<collectingRegion id="666AF813FFD8D421FC61FC99FDF9FCDD" box="[957,1014,841,867]" country="Bolivia" name="Beni" pageId="29" pageNumber="30">Beni</collectingRegion>
River Basin at Tumupasa; Popoi River, Upper
<collectingRegion id="666AF813FFD8D421FE56FCBEFFCAFC36" box="[394,453,878,904]" country="Bolivia" name="Beni" pageId="29" pageNumber="30">Beni</collectingRegion>
; Ixiamas; Huachi; Rurrenabaque).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="ECB4657AFFD8D421FF61FC61FEFAFA9A" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="A41136F1FFD8D421FF61FC61FF39FC75" blockId="29.[189,310,945,971]" box="[189,310,945,971]" pageId="29" pageNumber="30">
<heading id="FF59819DFFD8D421FF61FC61FF39FC75" bold="true" box="[189,310,945,971]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="96DAEAE3FFD8D421FF61FC61FF39FC75" bold="true" box="[189,310,945,971]" pageId="29" pageNumber="30">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFD8D421FF61FC32FEFAFA9A" blockId="29.[189,1399,993,1316]" pageId="29" pageNumber="30">
<taxonomicName id="63AE4D72FFD8D421FF61FC32FF8EFC45" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" baseAuthorityName="Boulenger" baseAuthorityYear="1892" box="[189,385,994,1020]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="moorii" status="comb. nov.">
<emphasis id="96DAEAE3FFD8D421FF61FC32FF8EFC45" box="[189,385,994,1020]" italics="true" pageId="29" pageNumber="30">Astyanax moorii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFD8D421FE50FC31FC1EFC42" box="[396,529,993,1020]" pageId="29" pageNumber="30">comb. nov.</taxonomicNameLabel>
is a member of the
<taxonomicName id="63AE4D72FFD8D421FCD0FC31FD7BFC45" authorityName="Eigenmann in Eigenmann &amp; Ogle" authorityYear="1907" box="[780,884,993,1019]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="orthodus">
<emphasis id="96DAEAE3FFD8D421FCD0FC31FD7BFC45" box="[780,884,993,1019]" italics="true" pageId="29" pageNumber="30">orthodus</emphasis>
</taxonomicName>
species-group of
<taxonomicName id="63AE4D72FFD8D421FB93FC32FAB4FC42" authorityName="Baird &amp; Girard" authorityYear="1854" box="[1103,1211,994,1020]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
<emphasis id="96DAEAE3FFD8D421FB93FC32FAB4FC42" box="[1103,1211,994,1020]" italics="true" pageId="29" pageNumber="30">Astyanax</emphasis>
</taxonomicName>
, differing from other species of the group in having an inconspicuous spot in the caudal peduncle (vs conspicuous peduncle spot) and by the length of the upper jaw more than 47% HL (vs less than 47% HL), and from
<taxonomicName id="63AE4D72FFD8D421FF61FB81FF49FBD5" authorityName="Eigenmann in Eigenmann &amp; Ogle" authorityYear="1907" box="[189,326,1105,1131]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="orthodus">
<emphasis id="96DAEAE3FFD8D421FF61FB81FF49FBD5" box="[189,326,1105,1131]" italics="true" pageId="29" pageNumber="30">A. orthodus</emphasis>
</taxonomicName>
,
<taxonomicName id="63AE4D72FFD8D421FE89FB81FFEFFBD5" authorityName="Myers" authorityYear="1942" box="[341,480,1105,1131]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="superbus">
<emphasis id="96DAEAE3FFD8D421FE89FB81FFEFFBD5" box="[341,480,1105,1131]" italics="true" pageId="29" pageNumber="30">A. superbus</emphasis>
</taxonomicName>
,
<taxonomicName id="63AE4D72FFD8D421FE33FB81FC72FBD5" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" baseAuthorityName="Torres-Mejia, Hernandez &amp; Senechal" baseAuthorityYear="2012" box="[495,637,1105,1131]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="yariguies">
<emphasis id="96DAEAE3FFD8D421FE33FB81FC72FBD5" box="[495,637,1105,1131]" italics="true" pageId="29" pageNumber="30">A. yariguies</emphasis>
</taxonomicName>
and
<taxonomicName id="63AE4D72FFD8D421FD6BFB81FD4AFBD5" authority="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega, 2018" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" box="[695,837,1105,1131]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="gandhiae" status="sp. nov.">
<emphasis id="96DAEAE3FFD8D421FD6BFB81FD4AFBD5" box="[695,837,1105,1131]" italics="true" pageId="29" pageNumber="30">A. gandhiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFD8D421FC91FB81FDA7FBD5" box="[845,936,1105,1131]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
in the interorbital distance being 27.4– 31.4% HL (vs&gt; 32% HL); it is distinguished from
<taxonomicName id="63AE4D72FFD8D421FCF6FBA6FDB3FB2E" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" box="[810,956,1142,1168]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="bopiensis" status="nom. nov.">
<emphasis id="96DAEAE3FFD8D421FCF6FBA6FDB3FB2E" box="[810,956,1142,1168]" italics="true" pageId="29" pageNumber="30">A. bopiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFD8D421FC19FBA6FA33FB2E" box="[965,1084,1142,1168]" pageId="29" pageNumber="30">nom. nov.</taxonomicNameLabel>
and
<taxonomicName id="63AE4D72FFD8D421FBA5FBA6FAF3FB2E" box="[1145,1276,1142,1168]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="gandhia">
<emphasis id="96DAEAE3FFD8D421FBA5FBA6FAF3FB2E" box="[1145,1276,1142,1168]" italics="true" pageId="29" pageNumber="30">A. gandhia</emphasis>
</taxonomicName>
e sp. nov.,
<taxonomicName id="63AE4D72FFD8D421FF61FB4BFF4AFB0B" authorityName="Eigenmann in Eigenmann &amp; Ogle" authorityYear="1907" box="[189,325,1179,1205]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="orthodus">
<emphasis id="96DAEAE3FFD8D421FF61FB4BFF4AFB0B" box="[189,325,1179,1205]" italics="true" pageId="29" pageNumber="30">A. orthodus</emphasis>
</taxonomicName>
and
<taxonomicName id="63AE4D72FFD8D421FEA2FB4BFFF8FB0B" authority="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega, 2018" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" box="[382,503,1179,1205]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="embera" status="sp. nov.">
<emphasis id="96DAEAE3FFD8D421FEA2FB4BFFF8FB0B" box="[382,503,1179,1205]" italics="true" pageId="29" pageNumber="30">A. embera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFD8D421FE22FB4BFC57FB0B" box="[510,600,1179,1205]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
by the distance between the dorsal and pectoral fins 37.4–40.3% SL (vs more than 40.4 SL); from
<emphasis id="96DAEAE3FFD8D421FDC7FB10FCA9FB64" box="[539,678,1216,1242]" italics="true" pageId="29" pageNumber="30">A. villwocki</emphasis>
by the absence of a dark band on the side surface of the body (vs present) and from
<taxonomicName id="63AE4D72FFD8D421FE18FB35FC5AFB41" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" box="[452,597,1253,1279]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="bopiensis" status="nom. nov.">
<emphasis id="96DAEAE3FFD8D421FE18FB35FC5AFB41" box="[452,597,1253,1279]" italics="true" pageId="29" pageNumber="30">A. bopiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFD8D421FD81FB35FCDCFB41" box="[605,723,1253,1279]" pageId="29" pageNumber="30">nom. nov.</taxonomicNameLabel>
by the maxillary length 34.0–37.9% HL (vs 46.7–58.9% HL).
</paragraph>
</subSubSection>
<subSubSection id="ECB4657AFFD8D421FF61FA9DFCC7FA03" pageId="29" pageNumber="30" type="etymology">
<paragraph id="A41136F1FFD8D421FF61FA9DFF4BFAD9" blockId="29.[189,324,1357,1383]" box="[189,324,1357,1383]" pageId="29" pageNumber="30">
<heading id="FF59819DFFD8D421FF61FA9DFF4BFAD9" bold="true" box="[189,324,1357,1383]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="96DAEAE3FFD8D421FF61FA9DFF4BFAD9" bold="true" box="[189,324,1357,1383]" pageId="29" pageNumber="30">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFD8D421FF61FAAEFCC7FA03" blockId="29.[189,1399,1406,1469]" pageId="29" pageNumber="30">
The specific epithet
<taxonomicName id="63AE4D72FFD8D421FE6CFAAEFFF1FA29" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" baseAuthorityName="Boulenger" baseAuthorityYear="1892" box="[432,510,1406,1431]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="moorii">
<emphasis id="96DAEAE3FFD8D421FE6CFAAEFFF1FA29" box="[432,510,1406,1431]" italics="true" pageId="29" pageNumber="30">moorii</emphasis>
</taxonomicName>
refers to the collector Mr. Spencer le Marchant Moore, who collected the
<typeStatus id="7B158853FFD8D421FF61FA73FEFFFA03" box="[189,240,1443,1469]" pageId="29" pageNumber="30">type</typeStatus>
material of
<taxonomicName id="63AE4D72FFD8D421FEA2FA73FC30FA03" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" baseAuthorityName="Boulenger" baseAuthorityYear="1892" box="[382,575,1443,1469]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="moorii" status="comb. nov.">
<emphasis id="96DAEAE3FFD8D421FEA2FA73FC30FA03" box="[382,575,1443,1469]" italics="true" pageId="29" pageNumber="30">Astyanax moorii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFD8D421FD9AFA73FCC7FA03" box="[582,712,1443,1469]" pageId="29" pageNumber="30">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="ECB4657AFFD8D421FF61FA36FDCCF855" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="A41136F1FFD8D421FF61FA36FF63F9BE" blockId="29.[189,364,1510,1536]" box="[189,364,1510,1536]" pageId="29" pageNumber="30">
<heading id="FF59819DFFD8D421FF61FA36FF63F9BE" bold="true" box="[189,364,1510,1536]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="96DAEAE3FFD8D421FF61FA36FF63F9BE" bold="true" box="[189,364,1510,1536]" pageId="29" pageNumber="30">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFD8D421FF36F9C7FCCEF98F" blockId="29.[189,1399,1558,1659]" box="[234,705,1558,1585]" pageId="29" pageNumber="30">
<emphasis id="96DAEAE3FFD8D421FF36F9C7FF69F98F" bold="true" box="[234,358,1559,1585]" pageId="29" pageNumber="30">Lectotype</emphasis>
(examined from photograph)
</paragraph>
<paragraph id="A41136F1FFD8D421FF61F9ECFCFFF9C5" blockId="29.[189,1399,1558,1659]" pageId="29" pageNumber="30">
<materialsCitation id="14C63CACFFD8D421FF61F9ECFCE3F9C5" ID-GBIF-Occurrence="1806442558" collectionCode="BMNH" country="Brazil" location="Upper Paraguay Chapala Plateau" pageId="29" pageNumber="30" specimenCode="BMNH 1892-4-20-51-2 2" specimenCount="1" stateProvince="Mato Grosso" typeStatus="lectotype">
<collectingCountry id="DCB97661FFD8D421FF61F9ECFF23F9E8" box="[189,300,1596,1622]" name="Brazil" pageId="29" pageNumber="30">BRAZIL</collectingCountry>
:
<specimenCount id="B2A8FD78FFD8D421FEE7F9ECFFC8F9EB" box="[315,455,1595,1622]" pageId="29" pageNumber="30" type="generic">1 specimen</specimenCount>
,
<quantity id="63569B14FFD8D421FE0AF9EBFC4CF9E8" box="[470,579,1595,1622]" metricMagnitude="-2" metricUnit="m" metricValue="6.52" pageId="29" pageNumber="30" unit="mm" value="65.2">65.2 mm</quantity>
SL;
<location id="A171602AFFD8D421FD58F9ECFA19F9EB" LSID="urn:lsid:plazi:treatment:2C0787E7FFD8D423FDDFFDA8FD46FA9E:A171602AFFD8D421FD58F9ECFA19F9EB" box="[644,1046,1595,1622]" country="Brazil" name="Upper Paraguay Chapala Plateau" pageId="29" pageNumber="30" stateProvince="Mato Grosso">Upper Paraguay Chapala Plateau</location>
,
<collectingRegion id="666AF813FFD8D421FBFAF9ECFACCF9E8" box="[1062,1219,1596,1622]" country="Brazil" name="Mato Grosso" pageId="29" pageNumber="30">Mato Grosso</collectingRegion>
State (BMNH 1892.4.20.51; photo
<specimenCode id="F4089E8AFFD8D421FE6DF9B1FCEDF9C5" box="[433,738,1632,1659]" pageId="29" pageNumber="30">BMNH 1892-4-20-51-2 2</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="A41136F1FFD8D421FF36F974FF98F900" blockId="29.[189,1399,1700,1801]" box="[234,407,1700,1726]" pageId="29" pageNumber="30">
<emphasis id="96DAEAE3FFD8D421FF36F974FF98F900" bold="true" box="[234,407,1700,1726]" pageId="29" pageNumber="30">Paralectotype</emphasis>
</paragraph>
<paragraph id="A41136F1FFD8D421FF61F919FCFFF8B6" blockId="29.[189,1399,1700,1801]" pageId="29" pageNumber="30">
<materialsCitation id="14C63CACFFD8D421FF61F919FCE3F8B6" ID-GBIF-Occurrence="1806442561" collectionCode="BMNH" country="Brazil" location="Upper Paraguay Chapala Plateau" pageId="29" pageNumber="30" specimenCode="BMNH 1892-4-20-51-2 1" specimenCount="1" stateProvince="Mato Grosso" typeStatus="paralectotype">
<collectingCountry id="DCB97661FFD8D421FF61F919FF23F95D" box="[189,300,1737,1763]" name="Brazil" pageId="29" pageNumber="30">BRAZIL</collectingCountry>
:
<specimenCount id="B2A8FD78FFD8D421FEE7F919FFC8F95D" box="[315,455,1737,1763]" pageId="29" pageNumber="30" type="generic">1 specimen</specimenCount>
, SL unknown;
<location id="A171602AFFD8D421FD58F919FA19F95D" LSID="urn:lsid:plazi:treatment:2C0787E7FFD8D423FDDFFDA8FD46FA9E:A171602AFFD8D421FD58F919FA19F95D" box="[644,1046,1737,1763]" country="Brazil" name="Upper Paraguay Chapala Plateau" pageId="29" pageNumber="30" stateProvince="Mato Grosso">Upper Paraguay Chapala Plateau</location>
,
<collectingRegion id="666AF813FFD8D421FBFAF919FACCF95D" box="[1062,1219,1737,1763]" country="Brazil" name="Mato Grosso" pageId="29" pageNumber="30">Mato Grosso</collectingRegion>
State (BMNH 1892.4.20.52; photo
<specimenCode id="F4089E8AFFD8D421FE6DF93EFCD0F8B6" box="[433,735,1774,1801]" pageId="29" pageNumber="30">BMNH 1892-4-20-51-2 1</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="A41136F1FFD8D421FF61F8E1FFA5F8F5" blockId="29.[189,426,1841,1867]" box="[189,426,1841,1867]" pageId="29" pageNumber="30">
<heading id="FF59819DFFD8D421FF61F8E1FFA5F8F5" bold="true" box="[189,426,1841,1867]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="96DAEAE3FFD8D421FF61F8E1FFA5F8F5" bold="true" box="[189,426,1841,1867]" pageId="29" pageNumber="30">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFD8D421FF61F8B2FDCCF855" blockId="29.[189,1399,1889,2027]" pageId="29" pageNumber="30">
<materialsCitation id="14C63CACFFD8D421FF61F8B2FB7DF81F" ID-GBIF-Occurrence="1806442536" collectionCode="MNRJ" country="Brazil" county="Manso Hydro-Electric Plant" location="Cuiaba River" municipality="Cuiaba River Basin" pageId="29" pageNumber="30" specimenCode="MNRJ 29284" specimenCount="2" stateProvince="Mato Grosso">
<collectingCountry id="DCB97661FFD8D421FF61F8B2FF23F8C2" box="[189,300,1890,1916]" name="Brazil" pageId="29" pageNumber="30">BRAZIL</collectingCountry>
: State of
<collectingRegion id="666AF813FFD8D421FE78F8B2FC4AF8C2" box="[420,581,1890,1916]" country="Brazil" name="Mato Grosso" pageId="29" pageNumber="30">Mato Grosso</collectingRegion>
:
<specimenCount id="B2A8FD78FFD8D421FD88F8B2FCE7F8C5" box="[596,744,1889,1916]" pageId="29" pageNumber="30" type="generic">2 specimens</specimenCount>
(sex unknown),
<quantity id="63569B14FFD8D421FC66F8B1FA67F8C2" box="[954,1128,1889,1916]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="6.47" metricValueMin="5.029999999999999" pageId="29" pageNumber="30" unit="mm" value="57.5" valueMax="64.7" valueMin="50.3">50.3–64.7 mm</quantity>
SL, stream (&quot;riacho&quot;) near
<collectingCounty id="4D704E7DFFD8D421FF24F857FC48F81E" box="[248,583,1926,1953]" pageId="29" pageNumber="30">Manso Hydro-Electric Plant</collectingCounty>
,
<collectingMunicipality id="4475AC8BFFD8D421FD88F856FD31F81E" box="[596,830,1926,1952]" pageId="29" pageNumber="30">
<location id="A171602AFFD8D421FD88F856FCFFF81E" LSID="urn:lsid:plazi:treatment:2C0787E7FFD8D423FDDFFDA8FD46FA9E:A171602AFFD8D421FD88F856FCFFF81E" box="[596,752,1926,1952]" country="Brazil" county="Manso Hydro-Electric Plant" municipality="Cuiaba River Basin" name="Cuiaba River" pageId="29" pageNumber="30" stateProvince="Mato Grosso">
Cuiabá
<location id="A171602AFFD8D421FD73F856FCFFF81E" LSID="urn:lsid:plazi:treatment:2C0787E7FFD8D423FDDFFDA8FD46FA9E:A171602AFFD8D421FD73F856FCFFF81E" box="[687,752,1926,1952]" country="Brazil" county="Manso Hydro-Electric Plant" municipality="Cuiaba River Basin" name="River Basin" pageId="29" pageNumber="30" stateProvince="Mato Grosso">River</location>
</location>
Basin
</collectingMunicipality>
, in the region of
<location id="A171602AFFD8D421FBD5F856FABCF81F" LSID="urn:lsid:plazi:treatment:2C0787E7FFD8D423FDDFFDA8FD46FA9E:A171602AFFD8D421FBD5F856FABCF81F" box="[1033,1203,1926,1953]" country="Brazil" county="Manso Hydro-Electric Plant" municipality="Cuiaba River Basin" name="Fazenda Nova" pageId="29" pageNumber="30" stateProvince="Mato Grosso">Fazenda Nova</location>
(
<specimenCode id="F4089E8AFFD8D421FB1FF857FB65F81F" box="[1219,1386,1927,1953]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="29" pageNumber="30">MNRJ 29284</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="14C63CACFFD8D421FF61F87CFDCFF855" ID-GBIF-Occurrence="1806442547" collectionCode="MNRJ" country="Brazil" county="Forte Stream" location="Cuiaba River Basin" municipality="Coxipo-Acu River" pageId="29" pageNumber="30" specimenCode="MNRJ 1190" specimenCount="3" stateProvince="Mato Grosso">
<specimenCount id="B2A8FD78FFD8D421FF61F87CFF42F878" box="[189,333,1964,1990]" pageId="29" pageNumber="30" type="generic">3 specimens</specimenCount>
(sex unknown), 43.2–59.0 mm SL,
<collectingCounty id="4D704E7DFFD8D421FD2BF87CFD9FF878" box="[759,912,1964,1990]" pageId="29" pageNumber="30">Forte Stream</collectingCounty>
(&quot;ribeirão&quot;), tributary of the
<collectingMunicipality id="4475AC8BFFD8D421FB39F87CFF0FF855" pageId="29" pageNumber="30">Coxipó-Açu River</collectingMunicipality>
,
<location id="A171602AFFD8D421FED0F801FFFAF855" LSID="urn:lsid:plazi:treatment:2C0787E7FFD8D423FDDFFDA8FD46FA9E:A171602AFFD8D421FED0F801FFFAF855" box="[268,501,2001,2027]" country="Brazil" county="Forte Stream" municipality="Coxipo-Acu River" name="Cuiaba River Basin" pageId="29" pageNumber="30" stateProvince="Mato Grosso">
<location id="A171602AFFD8D421FED0F801FFA8F855" LSID="urn:lsid:plazi:treatment:2C0787E7FFD8D423FDDFFDA8FD46FA9E:A171602AFFD8D421FED0F801FFA8F855" box="[268,423,2001,2027]" country="Brazil" county="Forte Stream" municipality="Coxipo-Acu River" name="Cuiaba River" pageId="29" pageNumber="30" stateProvince="Mato Grosso">
Cuiabá
<location id="A171602AFFD8D421FEBAF801FFA8F855" LSID="urn:lsid:plazi:treatment:2C0787E7FFD8D423FDDFFDA8FD46FA9E:A171602AFFD8D421FEBAF801FFA8F855" box="[358,423,2001,2027]" country="Brazil" county="Forte Stream" municipality="Coxipo-Acu River" name="River Basin" pageId="29" pageNumber="30" stateProvince="Mato Grosso">River</location>
</location>
Basin
</location>
,
<location id="A171602AFFD8D421FDDCF801FD1DF855" LSID="urn:lsid:plazi:treatment:2C0787E7FFD8D423FDDFFDA8FD46FA9E:A171602AFFD8D421FDDCF801FD1DF855" box="[512,786,2001,2027]" country="Brazil" county="Forte Stream" municipality="Coxipo-Acu River" name="Municipality of Cuiaba" pageId="29" pageNumber="30" stateProvince="Mato Grosso">Municipality of Cuiabá</location>
(
<specimenCode id="F4089E8AFFD8D421FCFDF801FDB8F855" box="[801,951,2001,2027]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="29" pageNumber="30">MNRJ 1190</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="ECB4657AFFDBD423FF61FEDBFB40FB10" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="description">
<paragraph id="A41136F1FFDBD422FF61FEDBFF41FE9B" blockId="30.[189,334,267,293]" box="[189,334,267,293]" pageId="30" pageNumber="31">
<heading id="FF59819DFFDBD422FF61FEDBFF41FE9B" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="96DAEAE3FFDBD422FF61FEDBFF41FE9B" bold="true" box="[189,334,267,293]" pageId="30" pageNumber="31">Description</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFDBD422FF61FEE9FC8EFE5F" blockId="30.[189,1398,313,481]" pageId="30" pageNumber="31">Body compressed, greatest body depth at or anterior to dorsal-fin origin. Mouth terminal. Anterior half of dorsal profile of head slightly straight; profile between posterior margin of supraoccipital spine and dorsal-fin origin as between last dorsal-fin ray and adipose-fin origin is slightly convex. Dorsal and ventral margins of caudal peduncle straight. Ventral profile convex from tip of snout to pelvic-fin insertion, feature visible in both sexes.</paragraph>
<paragraph id="A41136F1FFDBD422FF61FDD8FDA5FD35" blockId="30.[189,1398,520,652]" pageId="30" pageNumber="31">Premaxillary teeth in two series; outer series with three to five tricuspid teeth, so the outer lateral tooth; inner tooth row with five pentacuspid teeth. Maxilla with three to four teeth in anterior-most part of ventral margin. Dentary with four anterior teeth pentacuspid, followed laterally by 6–10 smaller teeth of increasing posterior inclination, reducing from tri- to unicuspid.</paragraph>
<paragraph id="A41136F1FFDBD422FF61FD63FFBFFBB4" blockId="30.[189,1399,690,1034]" pageId="30" pageNumber="31">
Pored scales of lateral line 41(1), 44(2), 45(1), 46(1), 47(1), 50(1), (n = 7), transverse scales from lateral line to origin of dorsal fin 8(5) a 9(2) (n = 7), scales from lateral line to origin of anal fin 6(1), 7(5), 8(1) (n = 7), scales from lateral line to insertion of pelvic fin 6(5), 7(2), (n = 7). Predorsal midline covered with bilobed medial scales for more than ¾ of its length, naked anteriorly. Rays of dorsal fin
<date id="D0101031FFDBD422FAD2FCCCFB32FC88" box="[1294,1341,796,822]" pageId="30" pageNumber="31">iii 9</date>
(n = 7); first simple ray small, only visible in cleared and stained specimens; second simple ray about half length of third simple ray. Distal margin of dorsal fin slightly convex. Origin of adipose fin anterior to vertical line through insertion of last ray of anal fin. Rays of pectoral fin
<date id="D0101031FFDBD422FBD0FC56FA32FC1E" box="[1036,1085,902,928]" pageId="30" pageNumber="31" value="2012-01">i 12</date>
(1),
<date id="D0101031FFDBD422FBB5FC56FA98FC1F" box="[1129,1175,902,929]" pageId="30" pageNumber="31" value="2013-01">i 13</date>
(5),
<date id="D0101031FFDBD422FB1AFC56FAF7FC1E" box="[1222,1272,902,928]" pageId="30" pageNumber="31" value="2014-01">i 14</date>
(1). Pelvicfin rays
<date id="D0101031FFDBD422FEC3FC79FF4DFC7A" box="[287,322,937,964]" pageId="30" pageNumber="31">i 6</date>
(1),
<date id="D0101031FFDBD422FEADFC79FF9CFC7A" box="[369,403,937,964]" pageId="30" pageNumber="31">i 7</date>
(6). Rays of anal fin iii(2), iv(v) and 28(1), 29(4), 30(1) and 32(1), first simple rays only visible in cleared and stained material. Origin of anal fin posterior to vertical line through insertion of last dorsal-fin ray.
</paragraph>
<paragraph id="A41136F1FFDBD422FF61FBE1FDA4FAA0" blockId="30.[189,1399,1073,1311]" pageId="30" pageNumber="31">Caudal-fin rays10–9, dorsal lobe with 11(5), 13(2) procurrent rays, ventral with 10(7) procurrent radios. Upper 10(5) principal rays of caudal fin associated with four dorsal hypurals, next 9(2) associated with three ventral hypurals and last principal ray associated with last hemal spine. Total vertebrae 37(3), including those of the Weberian apparatus: precaudal centra 15(2) y 16(1), last two without true pleural rib. Caudal centra 19(2), 20(1). Epineurals 36(1), 38(2), 40(1), 41(1) posterior-most epineural may not reach anterior surface of urostyle; epipleurals 20(5). Caudal skeleton with seven hypurals. First three hypurals with swollen anterior margins in contact with urostyle.</paragraph>
<caption id="F0D16679FFDBD422FF61F859FEF1F854" httpUri="https://zenodo.org/record/1169585/files/figure.png" pageId="30" pageNumber="31" startId="30.[189,232,1929,1955]" targetBox="[189,1398,1393,1892]" targetPageId="30">
<paragraph id="A41136F1FFDBD422FF61F859FEF1F854" blockId="30.[189,1399,1929,2026]" pageId="30" pageNumber="31">
<emphasis id="96DAEAE3FFDBD422FF61F859FF14F81D" bold="true" box="[189,283,1929,1955]" pageId="30" pageNumber="31">Fig. 13.</emphasis>
<taxonomicName id="63AE4D72FFDBD422FEFFF85AFCC9F81A" authority="(Boulenger, 1892)" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" baseAuthorityName="Boulenger" baseAuthorityYear="1892" box="[291,710,1929,1956]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="moorii">
<emphasis id="96DAEAE3FFDBD422FEFFF85AFFEAF81D" box="[291,485,1930,1956]" italics="true" pageId="30" pageNumber="31">Astyanax moorii</emphasis>
(Boulenger, 1892)
</taxonomicName>
, 64.7 mm SL, Creek near the Manso Hydroelectric Plant, Cuiabá River drainage, Fazenda Nova region, Mato Grosso State, Brazil (MNRJ 29284). Scale bar = 1 cm.
</paragraph>
</caption>
<paragraph id="A41136F1FFDAD423FF61FEDBFFE9FE9B" blockId="31.[189,486,267,293]" box="[189,486,267,293]" pageId="31" pageNumber="32">
<heading id="FF59819DFFDAD423FF61FEDBFFE9FE9B" bold="true" box="[189,486,267,293]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="96DAEAE3FFDAD423FF61FEDBFFE9FE9B" bold="true" box="[189,486,267,293]" pageId="31" pageNumber="32">Pigmentation in alcohol</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFDAD423FF61FEEAFCFAFE5C" blockId="31.[189,1399,314,482]" pageId="31" pageNumber="32">Sides of body yellowish, without reticulated pattern, with silver midlateral stripe present from humeral region to caudal-peduncle base, overlain by series of chevron-shaped marks formed by dark lines along myosepta between myotomes extending from dorsal region of coelomic cavity to caudal peduncle; pigmented muscle septae forming chevrons not coinciding with scale rows. Chevrons with distal extensions both in juvenile and adult specimens.</paragraph>
<paragraph id="A41136F1FFDAD423FF61FDD9FA54FC38" blockId="31.[189,1399,521,903]" pageId="31" pageNumber="32">
Dorsal region of head and body chestnut brown. Sides of cranium and ventral surface of body light brown, not silvery. Melanophores of humeral region forming two (anterior and posterior) humeral spots. The anterior spot is formed by two layers of pigment, with brown melanophores distributed in thin superficial layer of the epithelium (Layer 1,
<figureCitation id="3C952A74FFDAD423FD1DFDA4FD0AFD30" box="[705,773,628,654]" captionStart="Fig.1" captionStartId="4.[189,232,1859,1885]" captionTargetBox="[189,1398,540,1837]" captionTargetId="figure@4.[189,1398,540,1837]" captionTargetPageId="4" captionText="Fig.1.A.DistributionofpigmentinthehumeralregionofAstyanaxspecies.B.A.bimaculatus(Linnaeus, 1758).C.A.caucanus(Steindachner,1879).D.A.mexicanus(DeFilippi,1853).E.A.courensisBertaco, Carvalho &amp; Jerep, 2010 (modified from Bertaco et al. 2010). The anterior humeral spot consists of two overlapping layers of pigment identified by numbers 1 and 2 (see Román-Valencia et al. 2015: fig. 1): 1. Diffuse vertical bar. 2. Horizontal spot with concentrated pigments." httpUri="https://zenodo.org/record/1169561/files/figure.png" pageId="31" pageNumber="32">Fig. 1</figureCitation>
); deeper layer that with dark melanophores (Layer 2,
<figureCitation id="3C952A74FFDAD423FF61FD47FF0BFD0F" box="[189,260,663,689]" captionStart="Fig.1" captionStartId="4.[189,232,1859,1885]" captionTargetBox="[189,1398,540,1837]" captionTargetId="figure@4.[189,1398,540,1837]" captionTargetPageId="4" captionText="Fig.1.A.DistributionofpigmentinthehumeralregionofAstyanaxspecies.B.A.bimaculatus(Linnaeus, 1758).C.A.caucanus(Steindachner,1879).D.A.mexicanus(DeFilippi,1853).E.A.courensisBertaco, Carvalho &amp; Jerep, 2010 (modified from Bertaco et al. 2010). The anterior humeral spot consists of two overlapping layers of pigment identified by numbers 1 and 2 (see Román-Valencia et al. 2015: fig. 1): 1. Diffuse vertical bar. 2. Horizontal spot with concentrated pigments." httpUri="https://zenodo.org/record/1169561/files/figure.png" pageId="31" pageNumber="32">Fig. 1</figureCitation>
). Layer 2 forming polygon-shaped spot, with two parts, forming vertices on margins, usually forming four-sided spot located from second to fifth or sixth scale of lateral series. Posterior humeral spot situated two scales posterior to anterior humeral spot, rectangle-shaped, inconspicuous and covering two to three scales above lateral-line. Individual scales on sides of body without spots or dots. Caudalpeduncle spot rhomboid with short anterior extension over silver lateral stripe not surpassing anal-fin origin, dark pigment of caudal peduncle stripe continued on to middle caudal-fin rays with dark pigment present on tips of remaining rays. Pectoral, pelvic, dorsal and anal fins hyaline.
</paragraph>
<paragraph id="A41136F1FFDAD423FF61FC7EFFBFFC76" blockId="31.[189,432,942,968]" box="[189,432,942,968]" pageId="31" pageNumber="32">
<heading id="FF59819DFFDAD423FF61FC7EFFBFFC76" bold="true" box="[189,432,942,968]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="96DAEAE3FFDAD423FF61FC7EFFBFFC76" bold="true" box="[189,432,942,968]" pageId="31" pageNumber="32">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFDAD423FF61FC0DFD9DFC49" blockId="31.[189,914,989,1015]" box="[189,914,989,1015]" pageId="31" pageNumber="32">No sexual dimorphism was observed, no hooks found on fins.</paragraph>
<paragraph id="A41136F1FFDAD423FF61FBCEFFC2FB87" blockId="31.[189,461,1054,1081]" box="[189,461,1054,1081]" pageId="31" pageNumber="32">
<heading id="FF59819DFFDAD423FF61FBCEFFC2FB87" bold="true" box="[189,461,1054,1081]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="96DAEAE3FFDAD423FF61FBCEFFC2FB87" bold="true" box="[189,461,1054,1081]" pageId="31" pageNumber="32">Taxonomic comments</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFDAD423FF61FB9DFB40FB10" blockId="31.[189,1399,1101,1199]" pageId="31" pageNumber="32">
This species is transferred from
<taxonomicName id="63AE4D72FFDAD423FD91FB9EFCE6FBD6" baseAuthorityName="D'Agosta" baseAuthorityYear="2011" box="[589,745,1102,1128]" class="Actinopterygii" family="Characidae" genus="Astyanacinus" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="32" phylum="Chordata" rank="genus">
<emphasis id="96DAEAE3FFDAD423FD91FB9EFCE6FBD6" box="[589,745,1102,1128]" italics="true" pageId="31" pageNumber="32">Astyanacinus</emphasis>
</taxonomicName>
to
<taxonomicName id="63AE4D72FFDAD423FCC4FB9EFD8BFBD6" authorityName="Baird &amp; Girard" authorityYear="1854" box="[792,900,1102,1128]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="32" phylum="Chordata" rank="genus">
<emphasis id="96DAEAE3FFDAD423FCC4FB9EFD8BFBD6" box="[792,900,1102,1128]" italics="true" pageId="31" pageNumber="32">Astyanax</emphasis>
</taxonomicName>
because it shares the anteriorly directed V-shaped chevrons along myomere junctions with members of the
<taxonomicName id="63AE4D72FFDAD423FBC4FBA1FA8FFB35" authorityName="Eigenmann in Eigenmann &amp; Ogle" authorityYear="1907" box="[1048,1152,1137,1163]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="orthodus">
<emphasis id="96DAEAE3FFDAD423FBC4FBA1FA8FFB35" box="[1048,1152,1137,1163]" italics="true" pageId="31" pageNumber="32">orthodus</emphasis>
</taxonomicName>
species-group. The conspicuous polygon-shaped humeral spot is also similar (
<figureCitation id="3C952A74FFDAD423FCB0FB44FDBEFB11" box="[876,945,1172,1199]" captionStart="Fig.1" captionStartId="4.[189,232,1859,1885]" captionTargetBox="[189,1398,540,1837]" captionTargetId="figure@4.[189,1398,540,1837]" captionTargetPageId="4" captionText="Fig.1.A.DistributionofpigmentinthehumeralregionofAstyanaxspecies.B.A.bimaculatus(Linnaeus, 1758).C.A.caucanus(Steindachner,1879).D.A.mexicanus(DeFilippi,1853).E.A.courensisBertaco, Carvalho &amp; Jerep, 2010 (modified from Bertaco et al. 2010). The anterior humeral spot consists of two overlapping layers of pigment identified by numbers 1 and 2 (see Román-Valencia et al. 2015: fig. 1): 1. Diffuse vertical bar. 2. Horizontal spot with concentrated pigments." httpUri="https://zenodo.org/record/1169561/files/figure.png" pageId="31" pageNumber="32">Fig. 1</figureCitation>
), consisting of dark melanophores.
</paragraph>
</subSubSection>
<subSubSection id="ECB4657AFFDAD423FF61FB06FD46FA9E" box="[189,841,1238,1312]" pageId="31" pageNumber="32" type="distribution">
<paragraph id="A41136F1FFDAD423FF61FB06FF59FB4E" blockId="31.[189,342,1238,1264]" box="[189,342,1238,1264]" pageId="31" pageNumber="32">
<heading id="FF59819DFFDAD423FF61FB06FF59FB4E" bold="true" box="[189,342,1238,1264]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="96DAEAE3FFDAD423FF61FB06FF59FB4E" bold="true" box="[189,342,1238,1264]" pageId="31" pageNumber="32">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFDAD423FF61FAD6FD46FA9E" blockId="31.[189,841,1285,1312]" box="[189,841,1285,1312]" pageId="31" pageNumber="32">
Known from the
<collectingRegion id="666AF813FFDAD423FE59FAD5FC22FA9E" box="[389,557,1285,1312]" country="Paraguay" name="Alto Paraguay" pageId="31" pageNumber="32">
Alto
<collectingCountry id="DCB97661FFDAD423FE1CFAD6FC22FA9E" box="[448,557,1286,1312]" name="Paraguay" pageId="31" pageNumber="32">Paraguay</collectingCountry>
</collectingRegion>
River in
<collectingCountry id="DCB97661FFDAD423FD47FAD5FCECFAA1" box="[667,739,1285,1311]" name="Brazil" pageId="31" pageNumber="32">Brazil</collectingCountry>
(
<figureCitation id="3C952A74FFDAD423FD28FAD5FD36FAA1" box="[756,825,1285,1311]" captionStart="Fig. 4" captionStartId="8.[189,232,2000,2026]" captionTargetBox="[189,1392,764,1921]" captionTargetId="figure@8.[151,1417,755,1932]" captionTargetPageId="8" captionText="Fig. 4.Distribution of speciesof the Astyanaxorthodus species-group(symbol includestypelocalities)." httpUri="https://zenodo.org/record/1169567/files/figure.png" pageId="31" pageNumber="32">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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