treatments-xml/data/2C/07/87/2C0787E7FFD7D429FE64FC00FFF8FA93.xml

<document id="DBEABEF76E5CBDC03EADFACEB10E3D70" ID-CLB-Dataset="30789" ID-DOI="10.5852/ejt.2018.402" ID-GBIF-Dataset="8f1510f6-1b79-467f-9e8d-f193ac82cc4b" ID-Zenodo-Dep="1169559" ID-ZooBank="A3CE68AA-C5C6-40B7-B57C-6EF6D949149B" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1518136191129" checkinUser="plazi" docAuthor="Ruiz-C, Raquel I., Román-Valencia, César, Taphorn, Donald C., Buckup, Paulo A. &amp; Ortega, Hernán" docDate="2018" docId="2C0787E7FFD7D429FE64FC00FFF8FA93" docLanguage="en" docName="ejt-402_Ruiz-C_Roman-Valencia.pdf.imf" docOrigin="European Journal of Taxonomy 402" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Astyanax orthodus Eigenmann in Eigenmann &amp; Ogle 1907" docType="treatment" docVersion="17" lastPageNumber="22" masterDocId="D03EFF9FFFC5D43CFFDCFFD0FE0FFFBE" masterDocTitle="Revision of the Astyanax orthodus species-group (Teleostei: Characidae) with descriptions of three new species" masterLastPageNumber="45" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1547298528796" originalUpdateUser="yanell" pageNumber="19" updateTime="1698520653973" updateUser="ExternalLinkService">
<mods:mods id="F19F56611D666E5A35BB68D70AB51241" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A6F8547E9EC830624CDE5126FF08C144">
<mods:title id="24C09C1515F0CBA6DDA5FE171D99C344">Revision of the Astyanax orthodus species-group (Teleostei: Characidae) with descriptions of three new species</mods:title>
</mods:titleInfo>
<mods:name id="F43EAC058F622B490EB82BAA3DEE2DEE" type="personal">
<mods:role id="9FA6D8C10755ED492453C20C1AA9AEC4">
<mods:roleTerm id="28879F2272D3D75AEF493B843304AB26">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C136AC389B565A9B6C31297B883793F9">Ruiz-C, Raquel I.</mods:namePart>
</mods:name>
<mods:name id="294347F8E2184BBF8B7242B725B5FBB4" type="personal">
<mods:role id="2881EAE8AD9B39B2AA40E2F4A4494602">
<mods:roleTerm id="E730E931591158DD7BD8EA72016FD828">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4D4647AAFAB4CD7E017E7A22358A89E0">Román-Valencia, César</mods:namePart>
</mods:name>
<mods:name id="25F701D93B2D3BCFB7081F8FECD3B663" type="personal">
<mods:role id="7FE741EAC5A5E8E04907A1B1A804B5C1">
<mods:roleTerm id="A5BB4E1E8D1C636200CCC6335340B6D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="15D9393A410D9FCE91E114EA12ADEA80">Taphorn, Donald C.</mods:namePart>
</mods:name>
<mods:name id="13A3B5EFD0A1AF9A1BB3D58E9DADB0A0" type="personal">
<mods:role id="C77EC746C8DABF75018041DF02D10079">
<mods:roleTerm id="66526A88AF77D4B2B1D22CCA17D59539">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="847CC1313997D65F96042D4C2A4A8096">Buckup, Paulo A.</mods:namePart>
</mods:name>
<mods:name id="8B4FB3DB623FCAAB29B6B211BEBEA528" type="personal">
<mods:role id="4386814AF24DD5A1A91D41144D8942A5">
<mods:roleTerm id="8A6D2EDF00907255DC8B8CC1F58C2E47">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="92905A364DF5940846B3F65F0E7D89F0">Ortega, Hernán</mods:namePart>
</mods:name>
<mods:typeOfResource id="841F7DC02712D8E462738BBC5DADCCE2">text</mods:typeOfResource>
<mods:relatedItem id="EC865471A17FBC50AC348C657142B918" type="host">
<mods:titleInfo id="2FBFAAFDA0B3C304EA1908E54F5B6895">
<mods:title id="196C97340B71EF5885A13CD537ACF639">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="BECAC05EFFE76873D06DDAB8AF6965F0">
<mods:date id="1DF41DC8A213235FBD489EE670A5584C">2018</mods:date>
<mods:detail id="4A1752D35509891EEDA4250D1F6FC5DB" type="pubDate">
<mods:number id="7F8D9A33CD368946F3352A04D85C76CF">2018-02-08</mods:number>
</mods:detail>
<mods:detail id="2DF904396EC144C44C5619F8C5412BC2" type="volume">
<mods:number id="E1DFA64FD88B80CDF156092CE64B74B4">402</mods:number>
</mods:detail>
<mods:extent id="FA553C01518D5E193FE8FBD726C10385" unit="page">
<mods:start id="42BFA4420B8C64E4B154AFE1A610DF90">1</mods:start>
<mods:end id="DBB797BBD354D2710F17F179D6FF4028">45</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5999EF5CD40138ECC91EE1CFB818FC10">journal article</mods:classification>
<mods:identifier id="A24CD481880DEEEFA7C17D71F1B26C4A" type="CLB-Dataset">30789</mods:identifier>
<mods:identifier id="DF463E56D00F8011532A1E09172358B2" type="DOI">10.5852/ejt.2018.402</mods:identifier>
<mods:identifier id="CBDC32526C814C3610DDA987A2020561" type="GBIF-Dataset">8f1510f6-1b79-467f-9e8d-f193ac82cc4b</mods:identifier>
<mods:identifier id="C36157FBC4E3C794A76B9EEE7F93BFBF" type="Zenodo-Dep">1169559</mods:identifier>
<mods:identifier id="35AF9242FC1B599A8FAD375331A75AB8" type="ZooBank">A3CE68AA-C5C6-40B7-B57C-6EF6D949149B</mods:identifier>
</mods:mods>
<treatment id="2C0787E7FFD7D429FE64FC00FFF8FA93" ID-DOI="http://doi.org/10.5281/zenodo.5622618" ID-GBIF-Taxon="140562869" ID-Zenodo-Dep="5622618" LSID="urn:lsid:plazi:treatment:2C0787E7FFD7D429FE64FC00FFF8FA93" httpUri="http://treatment.plazi.org/id/2C0787E7FFD7D429FE64FC00FFF8FA93" lastPageId="21" lastPageNumber="22" pageId="18" pageNumber="19">
<subSubSection id="ECB4657AFFD7D42EFE64FC00FA73FC54" box="[440,1148,976,1003]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="A41136F1FFD7D42EFE64FC00FA73FC54" blockId="18.[440,1148,976,1041]" box="[440,1148,976,1003]" pageId="18" pageNumber="19">
<heading id="FF59819DFFD7D42EFE64FC00FA73FC54" box="[440,1148,976,1003]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="19" reason="2">
<taxonomicName id="63AE4D72FFD7D42EFE64FC00FA73FC54" ID-CoL="J5TZ" authority="Eigenmann in Eigenmann &amp; Ogle, 1907" authorityName="Eigenmann in Eigenmann &amp; Ogle" authorityYear="1907" box="[440,1148,976,1003]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="orthodus">
<emphasis id="96DAEAE3FFD7D42EFE64FC00FC93FC55" bold="true" box="[440,668,976,1003]" italics="true" pageId="18" pageNumber="19">Astyanax orthodus</emphasis>
Eigenmann in 
<bibRefCitation id="C03F4B00FFD7D42EFC92FC00FA73FC54" author="Eigenmann C. H. &amp; Ogle F." box="[846,1148,976,1002]" journalOrPublisher="Proceedings of the United States National Museum" pageId="18" pageNumber="19" pagination="1 - 36" part="33" publicationUrl="https://doi.org/10.5479/si.00963801.33-1556.1" refId="ref19281" refString="Eigenmann C. H. &amp; Ogle F. 1907. An annotated list of characin fishes in the United States National Museum and the Museum of Indiana University, with descriptions of new species. Proceedings of the United States National Museum 33: 1 - 36. https: // doi. org / 10.5479 / si. 00963801.33 - 1556.1" title="An annotated list of characin fishes in the United States National Museum and the Museum of Indiana University, with descriptions of new species" type="journal article" year="1907">Eigenmann &amp; Ogle, 1907</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ECB4657AFFD7D42EFD47FC26FD96FBAE" box="[667,921,1014,1041]" pageId="18" pageNumber="19" type="description">
<paragraph id="A41136F1FFD7D42EFD47FC26FD96FBAE" blockId="18.[440,1148,976,1041]" box="[667,921,1014,1041]" pageId="18" pageNumber="19">
<figureCitation id="3C952A74FFD7D42EFD47FC26FCF4FBAE" box="[667,763,1014,1040]" captionStart="Fig. 3" captionStartId="6.[189,232,1964,1990]" captionTargetBox="[462,1020,848,1930]" captionTargetId="figure@6.[446,1142,837,1938]" captionTargetPageId="6" captionText="Fig. 3. Shape of caudal-peduncle spot in species of the Astyanax orthodus species-group. A. Astyanax orthodus Eigenmann, 1907. B. Astyanax embera sp. nov. Scale bar = 1 cm." httpUri="https://zenodo.org/record/1169565/files/figure.png" pageId="18" pageNumber="19">Figs 3A</figureCitation>
, 
<figureCitation id="3C952A74FFD7D42EFCDBFC26FD17FBAE" box="[775,792,1014,1040]" captionStart="Fig. 4" captionStartId="8.[189,232,2000,2026]" captionTargetBox="[189,1392,764,1921]" captionTargetId="figure@8.[151,1417,755,1932]" captionTargetPageId="8" captionText="Fig. 4.Distribution of speciesof the Astyanaxorthodus species-group(symbol includestypelocalities)." httpUri="https://zenodo.org/record/1169567/files/figure.png" pageId="18" pageNumber="19">4</figureCitation>
, 
<figureCitation id="3C952A74FFD7D42EFCF9FC26FD3BFBAE" box="[805,820,1014,1040]" captionStart="Fig. 8" captionStartId="19.[189,232,1930,1956]" captionTargetBox="[189,1397,871,1894]" captionTargetId="figure@19.[189,1397,871,1894]" captionTargetPageId="19" captionText="Fig. 8. Astyanax orthodus Eigenmann, 1907. A. Holotype, 88.3 mm SL, Truando River, Atrato River Basin, Chocó, Pacific coast of Colombia (USNM 55655). B. 32.5 mm SL,Vereda Sautatá,Atrato River drainage, Chocó, Pacific coast of Colombia (IAVH 7146). Scale bars = 1 cm." httpUri="https://zenodo.org/record/1169575/files/figure.png" pageId="18" pageNumber="19">8</figureCitation>
, 
<tableCitation id="E92C034AFFD7D42EFC9EFC26FD96FBAE" box="[834,921,1014,1040]" captionStart="Table 2" captionStartId="17.[145,214,609,635]" captionTargetPageId="17" captionText="Table 2. Morphometric data for some species of the Astyanax orthodus species-group." httpUri="http://table.plazi.org/id/F0D16679FFD4D42DFF4DFDB1FA88FDC5" pageId="18" pageNumber="19" tableUuid="F0D16679FFD4D42DFF4DFDB1FA88FDC5">Table 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="ECB4657AFFD7D42EFF61FBEBFDB9FBC4" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="A41136F1FFD7D42EFF61FBEBFDB9FBC4" blockId="18.[189,1399,1082,1147]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="84BE11DFFFD7D42EFF61FBEBFDB9FBC4" pageId="18" pageNumber="19">
<taxonomicName id="63AE4D72FFD7D42EFF61FBEBFDC2FBEB" ID-CoL="J5TZ" authority="Eigenmann in Eigenmann &amp; Ogle, 1907: 27" authorityName="Eigenmann in Eigenmann &amp; Ogle" authorityPageNumber="27" authorityYear="1907" box="[189,973,1082,1109]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="orthodus">
<emphasis id="96DAEAE3FFD7D42EFF61FBEBFF92FBEA" box="[189,413,1082,1109]" italics="true" pageId="18" pageNumber="19">Astyanax orthodus</emphasis>
Eigenmann in 
<treatmentCitation id="250F10E0FFD7D42EFD81FBEAFDC2FBEB" author="" box="[605,973,1082,1109]" page="27" pageId="18" pageNumber="19">
<bibRefCitation id="C03F4B00FFD7D42EFD81FBEAFD92FBEB" author="Eigenmann C. H. &amp; Ogle F." box="[605,925,1082,1109]" journalOrPublisher="Proceedings of the United States National Museum" pageId="18" pageNumber="19" pagination="1 - 36" part="33" publicationUrl="https://doi.org/10.5479/si.00963801.33-1556.1" refId="ref19281" refString="Eigenmann C. H. &amp; Ogle F. 1907. An annotated list of characin fishes in the United States National Museum and the Museum of Indiana University, with descriptions of new species. Proceedings of the United States National Museum 33: 1 - 36. https: // doi. org / 10.5479 / si. 00963801.33 - 1556.1" title="An annotated list of characin fishes in the United States National Museum and the Museum of Indiana University, with descriptions of new species" type="journal article" year="1907">Eigenmann &amp; Ogle, 1907</bibRefCitation>
: 27
</treatmentCitation>
</taxonomicName>
, fig. 23. Original description, 
<typeStatus id="7B158853FFD7D42EFA98FBEBFB78FBEB" box="[1348,1399,1083,1109]" pageId="18" pageNumber="19">type</typeStatus>
locality: 
<location id="A171602AFFD7D42EFE8DFBB0FFB9FBC4" LSID="urn:lsid:plazi:treatment:2C0787E7FFD7D429FE64FC00FFF8FA93:A171602AFFD7D42EFE8DFBB0FFB9FBC4" box="[337,438,1120,1146]" country="Colombia" municipality="Rio Atrato River Basin" name="Truando" pageId="18" pageNumber="19" stateProvince="Choco">Truando</location>
, 
<collectingMunicipality id="4475AC8BFFD7D42EFE1EFBB0FCDCFBC4" box="[450,723,1120,1147]" pageId="18" pageNumber="19">Rio Atrato River Basin</collectingMunicipality>
, 
<collectingRegion id="666AF813FFD7D42EFD03FBB0FD20FBC4" box="[735,815,1120,1146]" country="Colombia" name="Choco" pageId="18" pageNumber="19">Chocó</collectingRegion>
, 
<collectingCountry id="DCB97661FFD7D42EFCE6FBB0FDBDFBC4" box="[826,946,1120,1146]" name="Colombia" pageId="18" pageNumber="19">Colombia</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="ECB4657AFFD7D42EFF61FB74FC49F894" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="A41136F1FFD7D42EFF61FB74FF39FB00" blockId="18.[189,310,1188,1214]" box="[189,310,1188,1214]" pageId="18" pageNumber="19">
<heading id="FF59819DFFD7D42EFF61FB74FF39FB00" bold="true" box="[189,310,1188,1214]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="96DAEAE3FFD7D42EFF61FB74FF39FB00" bold="true" box="[189,310,1188,1214]" pageId="18" pageNumber="19">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFD7D42EFF61FB06FC49F894" blockId="18.[189,1399,1238,1834]" pageId="18" pageNumber="19">
<taxonomicName id="63AE4D72FFD7D42EFF61FB06FF90FB4E" authorityName="Eigenmann in Eigenmann &amp; Ogle" authorityYear="1907" box="[189,415,1238,1264]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="orthodus">
<emphasis id="96DAEAE3FFD7D42EFF61FB06FF90FB4E" box="[189,415,1238,1264]" italics="true" pageId="18" pageNumber="19">Astyanax orthodus</emphasis>
</taxonomicName>
is a member of the 
<taxonomicName id="63AE4D72FFD7D42EFD6CFB06FD17FB4E" authorityName="Eigenmann in Eigenmann &amp; Ogle" authorityYear="1907" box="[688,792,1238,1264]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="orthodus">
<emphasis id="96DAEAE3FFD7D42EFD6CFB06FD17FB4E" box="[688,792,1238,1264]" italics="true" pageId="18" pageNumber="19">orthodus</emphasis>
</taxonomicName>
species-group (as defined above) differing from 
<emphasis id="96DAEAE3FFD7D42EFF61FB2CFF44FAA8" box="[189,331,1276,1302]" italics="true" pageId="18" pageNumber="19">A. villwocki</emphasis>
, 
<taxonomicName id="63AE4D72FFD7D42EFE82FB2CFFE3FAA8" authorityName="Myers" authorityYear="1942" box="[350,492,1276,1302]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="superbus">
<emphasis id="96DAEAE3FFD7D42EFE82FB2CFFE3FAA8" box="[350,492,1276,1302]" italics="true" pageId="18" pageNumber="19">A. superbus</emphasis>
</taxonomicName>
, 
<taxonomicName id="63AE4D72FFD7D42EFE22FB2CFC9DFAA8" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" box="[510,658,1276,1302]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="bopiensis" status="nom. nov.">
<emphasis id="96DAEAE3FFD7D42EFE22FB2CFC9DFAA8" box="[510,658,1276,1302]" italics="true" pageId="18" pageNumber="19">A. bopiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFD7D42EFD40FB2CFD1AFAA8" box="[668,789,1276,1302]" pageId="18" pageNumber="19">nom. nov.</taxonomicNameLabel>
, 
<emphasis id="96DAEAE3FFD7D42EFCFBFB2CFD36FAA8" bold="true" box="[807,825,1276,1302]" italics="true" pageId="18" pageNumber="19">A</emphasis>
. 
<taxonomicName id="63AE4D72FFD7D42EFC97FB2CFDB7FAA8" authority="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega, 2018" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" box="[843,952,1276,1302]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="gandhiae" status="sp. nov.">
<emphasis id="96DAEAE3FFD7D42EFC97FB2CFDB7FAA8" box="[843,952,1276,1302]" italics="true" pageId="18" pageNumber="19">gandhiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFD7D42EFC1FFB2CFA2DFAA8" box="[963,1058,1276,1302]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="63AE4D72FFD7D42EFBBEFB2CFAFCFAA8" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" baseAuthorityName="Torres-Mejia, Hernandez &amp; Senechal" baseAuthorityYear="2012" box="[1122,1267,1276,1302]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="yariguies">
<emphasis id="96DAEAE3FFD7D42EFBBEFB2CFAFCFAA8" box="[1122,1267,1276,1302]" italics="true" pageId="18" pageNumber="19">A. yariguies</emphasis>
</taxonomicName>
by having a conspicuous, polygonal spot on the caudal peduncle that does not extend anteriorly to vertical line through posterior tip of anal fin (vs spot extending anteriorly as a stripe beyond vertical line through posterior tip of anal fin). It further differs from 
<taxonomicName id="63AE4D72FFD7D42EFD29FABEFD8EFA36" authorityName="Myers" authorityYear="1942" box="[757,897,1390,1416]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="superbus">
<emphasis id="96DAEAE3FFD7D42EFD29FABEFD8EFA36" box="[757,897,1390,1416]" italics="true" pageId="18" pageNumber="19">A. superbus</emphasis>
</taxonomicName>
by the absence of brown marks along the sides of body (vs presence of a series of longitudinal brown marks along the sides of body). It differs from 
<taxonomicName id="63AE4D72FFD7D42EFF23FA6AFF92FA6A" authority="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega, 2018" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" box="[255,413,1466,1492]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="boliviensis" status="sp. nov.">
<emphasis id="96DAEAE3FFD7D42EFF23FA6AFF92FA6A" box="[255,413,1466,1492]" italics="true" pageId="18" pageNumber="19">A. boliviensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFD7D42EFE7BFA6AFC0DFA6A" box="[423,514,1466,1492]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
in having shorter, simple extension of the chevrons (vs chevron extension elongate and branched distally). It differs from 
<taxonomicName id="63AE4D72FFD7D42EFCDFFA30FD9BFA44" authority="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega, 2018" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" box="[771,916,1504,1530]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="gandhiae" status="sp. nov.">
<emphasis id="96DAEAE3FFD7D42EFCDFFA30FD9BFA44" box="[771,916,1504,1530]" italics="true" pageId="18" pageNumber="19">A. gandhiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFD7D42EFC7CFA30FDF1FA44" box="[928,1022,1504,1530]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
in having a reticulated pattern predominant over upper region of coelomic cavity (vs without reticulated pattern predominant over upper region of coelomic cavity). It differs from 
<taxonomicName id="63AE4D72FFD7D42EFCDAF9FCFD8FF9F8" authority="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega, 2018" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" box="[774,896,1580,1606]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="embera" status="sp. nov.">
<emphasis id="96DAEAE3FFD7D42EFCDAF9FCFD8FF9F8" box="[774,896,1580,1606]" italics="true" pageId="18" pageNumber="19">A. embera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFD7D42EFC55F9FCFDEAF9F8" box="[905,997,1580,1606]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
in the position, size and shape of the spot on the caudal peduncle, which does not extend to the dorsal and ventral margins of the caudal peduncle (vs caudal peduncle extending as a fan-shaped blotch that reaches, or almost reaches, the dorsal and ventral margins of the caudal peduncle, 
<figureCitation id="3C952A74FFD7D42EFCCFF94DFD62F906" box="[787,877,1693,1720]" captionStart="Fig. 3" captionStartId="6.[189,232,1964,1990]" captionTargetBox="[462,1020,848,1930]" captionTargetId="figure@6.[446,1142,837,1938]" captionTargetPageId="6" captionText="Fig. 3. Shape of caudal-peduncle spot in species of the Astyanax orthodus species-group. A. Astyanax orthodus Eigenmann, 1907. B. Astyanax embera sp. nov. Scale bar = 1 cm." httpUri="https://zenodo.org/record/1169565/files/figure.png" pageId="18" pageNumber="19">Fig. 3A</figureCitation>
). The new species is distinct from 
<taxonomicName id="63AE4D72FFD7D42EFAD4F94EFB79F909" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" baseAuthorityName="Boulenger" baseAuthorityYear="1892" box="[1288,1398,1694,1720]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="moorii" status="comb. nov.">
<emphasis id="96DAEAE3FFD7D42EFAD4F94EFB79F909" box="[1288,1398,1694,1720]" italics="true" pageId="18" pageNumber="19">A. moorii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFD7D42EFF61F913FF31F960" box="[189,318,1731,1758]" pageId="18" pageNumber="19">comb. nov.</taxonomicNameLabel>
in having the distance between the dorsal and pectoral fins more than 40.4% HL (vs less than 40.3% HL), interorbital distance more than 32% HL (vs less 32% HL) and upper jaw length less than 31% HL (vs more than 45% HL).
</paragraph>
</subSubSection>
<subSubSection id="ECB4657AFFD7D42EFF61F884FA50F855" pageId="18" pageNumber="19" type="etymology">
<paragraph id="A41136F1FFD7D42EFF61F884FF4BF8D0" blockId="18.[189,324,1876,1902]" box="[189,324,1876,1902]" pageId="18" pageNumber="19">
<heading id="FF59819DFFD7D42EFF61F884FF4BF8D0" bold="true" box="[189,324,1876,1902]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="96DAEAE3FFD7D42EFF61F884FF4BF8D0" bold="true" box="[189,324,1876,1902]" pageId="18" pageNumber="19">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFD7D42EFF61F855FA50F855" blockId="18.[189,1398,1925,2027]" pageId="18" pageNumber="19">
The specific epithet 
<taxonomicName id="63AE4D72FFD7D42EFE77F855FC1CF821" authorityName="Eigenmann in Eigenmann &amp; Ogle" authorityYear="1907" box="[427,531,1925,1951]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="orthodus">
<emphasis id="96DAEAE3FFD7D42EFE77F855FC1CF821" box="[427,531,1925,1951]" italics="true" pageId="18" pageNumber="19">orthodus</emphasis>
</taxonomicName>
is latinized from Greek ‘ 
<emphasis id="96DAEAE3FFD7D42EFCE0F855FDA7F821" box="[828,936,1925,1951]" italics="true" pageId="18" pageNumber="19">orthodon</emphasis>
’, meaning ‘straight tooth’, in reference to the straight posterior surface of the inner premaxillary teeth and the cusps, which Eigenmann (in 
<bibRefCitation id="C03F4B00FFD7D42EFF61F801FFEBF855" author="Eigenmann C. H. &amp; Ogle F." box="[189,484,2001,2027]" journalOrPublisher="Proceedings of the United States National Museum" pageId="18" pageNumber="19" pagination="1 - 36" part="33" publicationUrl="https://doi.org/10.5479/si.00963801.33-1556.1" refId="ref19281" refString="Eigenmann C. H. &amp; Ogle F. 1907. An annotated list of characin fishes in the United States National Museum and the Museum of Indiana University, with descriptions of new species. Proceedings of the United States National Museum 33: 1 - 36. https: // doi. org / 10.5479 / si. 00963801.33 - 1556.1" title="An annotated list of characin fishes in the United States National Museum and the Museum of Indiana University, with descriptions of new species" type="journal article" year="1907">Eigenmann &amp; Ogle 1907</bibRefCitation>
) used to distinguish 
<taxonomicName id="63AE4D72FFD7D42EFD04F801FD6FF855" authorityName="Eigenmann in Eigenmann &amp; Ogle" authorityYear="1907" box="[728,864,2001,2027]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="orthodus">
<emphasis id="96DAEAE3FFD7D42EFD04F801FD6FF855" box="[728,864,2001,2027]" italics="true" pageId="18" pageNumber="19">A. orthodus</emphasis>
</taxonomicName>
from 
<taxonomicName id="63AE4D72FFD7D42EFC74F801FA57F855" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[936,1112,2001,2027]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="bimaculatus">
<emphasis id="96DAEAE3FFD7D42EFC74F801FA57F855" box="[936,1112,2001,2027]" italics="true" pageId="18" pageNumber="19">A. bimaculatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="ECB4657AFFD6D42FFF61FEDBFB36FC85" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="A41136F1FFD6D42FFF61FEDBFFA5FE9B" blockId="19.[189,426,267,293]" box="[189,426,267,293]" pageId="19" pageNumber="20">
<heading id="FF59819DFFD6D42FFF61FEDBFFA5FE9B" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="96DAEAE3FFD6D42FFF61FEDBFFA5FE9B" bold="true" box="[189,426,267,293]" pageId="19" pageNumber="20">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFD6D42FFF36FEEAFCB8FEEA" blockId="19.[189,1398,313,411]" box="[234,695,313,340]" pageId="19" pageNumber="20">
<emphasis id="96DAEAE3FFD6D42FFF36FEEAFF53FEEA" bold="true" box="[234,348,314,340]" pageId="19" pageNumber="20">Holotype</emphasis>
(examined from photograph)
</paragraph>
<paragraph id="A41136F1FFD6D42FFF61FE8DFA21FE25" blockId="19.[189,1398,313,411]" pageId="19" pageNumber="20">
<materialsCitation id="14C63CACFFD6D42FFF61FE8DFA21FE25" ID-GBIF-Occurrence="1806442538" collectionCode="USNM" country="Colombia" location="Urabá, Truandó River, lower Atrato River Basin, Caribbean coast" municipality="Rio Sucio Municipality" pageId="19" pageNumber="20" specimenCode="USNM 55655" specimenCount="1" stateProvince="Department of Chocó" typeStatus="holotype">
<collectingCountry id="DCB97661FFD6D42FFF61FE8DFF51FEC9" box="[189,350,349,375]" name="Colombia" pageId="19" pageNumber="20">COLOMBIA</collectingCountry>
: 
<quantity id="63569B14FFD6D42FFEB1FE8DFFD4FEC9" box="[365,475,349,375]" metricMagnitude="-2" metricUnit="m" metricValue="8.83" pageId="19" pageNumber="20" unit="mm" value="88.3">88.3 mm</quantity>
SL, NW 
<collectingCountry id="DCB97661FFD6D42FFD84FE8CFCC0FEC8" box="[600,719,348,374]" name="Colombia" pageId="19" pageNumber="20">Colombia</collectingCountry>
, 
<collectingRegion id="666AF813FFD6D42FFD03FE8DFDE5FEC8" box="[735,1002,348,375]" pageId="19" pageNumber="20">Department of Chocó</collectingRegion>
, 
<collectingMunicipality id="4475AC8BFFD6D42FFC25FE8CFB17FEC8" box="[1017,1304,348,374]" pageId="19" pageNumber="20">Rio Sucio Municipality</collectingMunicipality>
, 
<location id="A171602AFFD6D42FFAFBFE8CFD69FE24" LSID="urn:lsid:plazi:treatment:2C0787E7FFD7D429FE64FC00FFF8FA93:A171602AFFD6D42FFAFBFE8CFD69FE24" country="Colombia" municipality="Rio Sucio Municipality" name="Urabá, Truandó River, lower Atrato River Basin, Caribbean coast" pageId="19" pageNumber="20" stateProvince="Department of Chocó">Urabá, Truandó River, lower Atrato River Basin, Caribbean coast</location>
(
<specimenCode id="F4089E8AFFD6D42FFCA9FE50FA2EFE25" box="[885,1057,384,411]" pageId="19" pageNumber="20">USNM 55655</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="A41136F1FFD6D42FFF36FE11FFA9FE65" blockId="19.[189,1399,449,828]" box="[234,422,449,475]" pageId="19" pageNumber="20">
<emphasis id="96DAEAE3FFD6D42FFF36FE11FFA9FE65" bold="true" box="[234,422,449,475]" pageId="19" pageNumber="20">Other material</emphasis>
</paragraph>
<paragraph id="A41136F1FFD6D42FFF61FE35FB36FC85" blockId="19.[189,1399,449,828]" pageId="19" pageNumber="20">
<materialsCitation id="14C63CACFFD6D42FFF61FE35FF61FD9C" ID-GBIF-Occurrence="1806442559" collectionCode="AMNH" country="Colombia" location="Atrato River" municipality="Atrato River" pageId="19" pageNumber="20" specimenCode="AMNH 5370" specimenCount="2" specimenCount-female="2" stateProvince="Choco">
<collectingCountry id="DCB97661FFD6D42FFF61FE35FF51FE41" box="[189,350,485,511]" name="Colombia" pageId="19" pageNumber="20">COLOMBIA</collectingCountry>
: 
<collectingRegion id="666AF813FFD6D42FFEBAFE34FFB8FE40" box="[358,439,484,510]" country="Colombia" name="Choco" pageId="19" pageNumber="20">Chocó</collectingRegion>
: 
<specimenCount id="B2A8FD78FFD6D42FFE1CFE35FC0EFE41" box="[448,513,485,511]" pageId="19" pageNumber="20" type="female">2 ♀♀</specimenCount>
, 
<quantity id="63569B14FFD6D42FFDD0FE34FCBBFE41" box="[524,692,484,511]" metricMagnitude="-2" metricUnit="m" metricValue="6.885" metricValueMax="7.159999999999999" metricValueMin="6.609999999999999" pageId="19" pageNumber="20" unit="mm" value="68.85" valueMax="71.6" valueMin="66.1">66.1–71.6 mm</quantity>
SL, 
<collectingMunicipality id="4475AC8BFFD6D42FFD39FE35FD79FE40" box="[741,886,484,511]" pageId="19" pageNumber="20">Atrato River</collectingMunicipality>
Basin, Truandó River, tributary 
<location id="A171602AFFD6D42FFB39FE35FB79FE40" LSID="urn:lsid:plazi:treatment:2C0787E7FFD7D429FE64FC00FFF8FA93:A171602AFFD6D42FFB39FE35FB79FE40" box="[1253,1398,484,511]" country="Colombia" municipality="Atrato River" name="Atrato River" pageId="19" pageNumber="20" stateProvince="Choco">Atrato River</location>
(
<specimenCode id="F4089E8AFFD6D42FFF19FDD8FF68FD9C" box="[197,359,519,546]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="19" pageNumber="20">AMNH 5370</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="14C63CACFFD6D42FFEA5FDD8FF0AFDF8" ID-GBIF-Occurrence="1806442518" country="Colombia" county="River Basin" location="Atrato River" municipality="Yuto River" pageId="19" pageNumber="20" specimenCount="2" specimenCount-female="2" stateProvince="Choco">
<specimenCount id="B2A8FD78FFD6D42FFEA5FDD8FFB4FD9C" box="[377,443,520,546]" pageId="19" pageNumber="20" type="female">2 ♀♀</specimenCount>
, 
<quantity id="63569B14FFD6D42FFE14FDD7FC20FD9C" box="[456,559,519,546]" metricMagnitude="-2" metricUnit="m" metricValue="8.69" pageId="19" pageNumber="20" unit="mm" value="86.9">86.9 mm</quantity>
SL, Yuto, Atrato 
<collectingCounty id="4D704E7DFFD6D42FFD25FDD8FD89FD9C" box="[761,902,520,546]" pageId="19" pageNumber="20">River Basin</collectingCounty>
, 
<collectingMunicipality id="4475AC8BFFD6D42FFC53FDD8FA1FFD9C" box="[911,1040,520,546]" pageId="19" pageNumber="20">Yuto River</collectingMunicipality>
, tributary 
<location id="A171602AFFD6D42FFB58FDD8FB19FD9C" LSID="urn:lsid:plazi:treatment:2C0787E7FFD7D429FE64FC00FFF8FA93:A171602AFFD6D42FFB58FDD8FB19FD9C" box="[1156,1302,520,546]" country="Colombia" county="River Basin" municipality="Yuto River" name="Atrato River" pageId="19" pageNumber="20" stateProvince="Choco">Atrato River</location>
(IAvHP 6494)
</materialsCitation>
; 
<materialsCitation id="14C63CACFFD6D42FFEC8FDFBFDAFFDD7" ID-GBIF-Occurrence="1806442550" country="Colombia" county="Rio Sucio municipality" location="vereda Sautatá, Atrato River Basin, Tendal Creek" municipality="River Basin" pageId="19" pageNumber="20" specimenCount="20" specimenCount-female="18" specimenCount-male="2" stateProvince="Choco">
<specimenCount id="B2A8FD78FFD6D42FFEC8FDFBFF66FDFB" box="[276,361,555,581]" pageId="19" pageNumber="20" type="female">16 ♀♀</specimenCount>
, 
<quantity id="63569B14FFD6D42FFEA6FDFBFC39FDFB" box="[378,566,555,581]" metricMagnitude="-2" metricUnit="m" metricValue="6.65" metricValueMax="10.07" metricValueMin="3.2299999999999995" pageId="19" pageNumber="20" unit="mm" value="66.5" valueMax="100.7" valueMin="32.3">32.3–100.7 mm</quantity>
SL, 
<specimenCount id="B2A8FD78FFD6D42FFDA8FDFBFCB4FDFB" box="[628,699,555,581]" pageId="19" pageNumber="20" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="B2A8FD78FFD6D42FFD17FDFBFD1EFDFB" box="[715,785,555,581]" pageId="19" pageNumber="20" type="female">2 ♀♀</specimenCount>
, C&amp;S, 
<quantity id="63569B14FFD6D42FFCB3FDFBFA13FDFB" box="[879,1052,555,582]" metricMagnitude="-2" metricUnit="m" metricValue="5.959999999999999" metricValueMax="6.779999999999999" metricValueMin="5.14" pageId="19" pageNumber="20" unit="mm" value="59.599999999999994" valueMax="67.8" valueMin="51.4">51.4–67.8 mm</quantity>
SL, 
<collectingCounty id="4D704E7DFFD6D42FFB87FDFBFB7CFDFB" box="[1115,1395,555,581]" pageId="19" pageNumber="20">Rio Sucio municipality</collectingCounty>
, 
<location id="A171602AFFD6D42FFF61FD9EFCFDFDD6" LSID="urn:lsid:plazi:treatment:2C0787E7FFD7D429FE64FC00FFF8FA93:A171602AFFD6D42FFF61FD9EFCFDFDD6" box="[189,754,590,616]" country="Colombia" county="Rio Sucio municipality" municipality="River Basin" name="vereda Sautatá, Atrato River Basin, Tendal Creek" pageId="19" pageNumber="20" stateProvince="Choco">vereda Sautatá, Atrato River Basin,Tendal Creek</location>
(IAvHP 7146)
</materialsCitation>
; 
<materialsCitation id="14C63CACFFD6D42FFC77FD9EFA32FD11" ID-GBIF-Occurrence="1806442504" country="Colombia" county="Atrato River" elevation="161" latitude="7.8022223" location="Tendal Creek" longLatPrecision="21" longitude="-77.172775" municipality="Sautata Strema" pageId="19" pageNumber="20" specimenCount="11" specimenCount-female="11" stateProvince="Choco">
<specimenCount id="B2A8FD78FFD6D42FFC77FD9EFDF5FDD7" box="[939,1018,590,617]" pageId="19" pageNumber="20" type="female">11 ♀♀</specimenCount>
, 
<quantity id="63569B14FFD6D42FFBDBFD9EFABFFDD7" box="[1031,1200,590,617]" metricMagnitude="-2" metricUnit="m" metricValue="4.465" metricValueMax="5.1899999999999995" metricValueMin="3.7399999999999998" pageId="19" pageNumber="20" unit="mm" value="44.65" valueMax="51.9" valueMin="37.4">37.4–51.9 mm</quantity>
SL, 
<collectingCounty id="4D704E7DFFD6D42FFB38FD9EFB79FDD6" box="[1252,1398,590,616]" pageId="19" pageNumber="20">Atrato River</collectingCounty>
Basin, Rio Sucio Municipality, 
<collectingMunicipality id="4475AC8BFFD6D42FFDEEFDA1FCECFD35" box="[562,739,625,651]" pageId="19" pageNumber="20">Sautata Strema</collectingMunicipality>
(“vereda”), 
<location id="A171602AFFD6D42FFCAAFDA1FA1BFD35" LSID="urn:lsid:plazi:treatment:2C0787E7FFD7D429FE64FC00FFF8FA93:A171602AFFD6D42FFCAAFDA1FA1BFD35" box="[886,1044,625,651]" country="Colombia" county="Atrato River" latitude="7.8022223" longLatPrecision="21" longitude="-77.172775" municipality="Sautata Strema" name="Tendal Creek" pageId="19" pageNumber="20" stateProvince="Choco">Tendal Creek</location>
(“quebrada”), 
<location id="A171602AFFD6D42FFB19FDA2FFBDFD10" LSID="urn:lsid:plazi:treatment:2C0787E7FFD7D429FE64FC00FFF8FA93:A171602AFFD6D42FFB19FDA2FFBDFD10" country="Colombia" county="Atrato River" latitude="7.8022223" longLatPrecision="21" longitude="-77.172775" municipality="Sautata Strema" name="Parque Natural Nacional Los Katios" pageId="19" pageNumber="20" stateProvince="Choco">Parque Natural Nacional Los Katios</location>
, 
<geoCoordinate id="C19A5036FFD6D42FFE61FD43FC5CFD11" box="[445,595,659,687]" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="15" value="7.8022223">07°48′08″ N</geoCoordinate>
, 
<geoCoordinate id="C19A5036FFD6D42FFD83FD43FCF6FD11" box="[607,761,659,687]" direction="west" orientation="longitude" pageId="19" pageNumber="20" precision="15" value="-77.172775">77°10′22″ W</geoCoordinate>
, 
<elevation id="2F83D1C2FFD6D42FFCD9FD44FD82FD10" box="[773,909,660,687]" metricMagnitude="2" metricUnit="m" metricValue="1.61" pageId="19" pageNumber="20" unit="m" value="161.0">
<quantity id="63569B14FFD6D42FFCD9FD44FD40FD11" box="[773,847,660,687]" metricMagnitude="2" metricUnit="m" metricValue="1.61" pageId="19" pageNumber="20" unit="m" value="161.0">161 m</quantity>
a.s.l.
</elevation>
(IAvHP 7208)
</materialsCitation>
; 
<materialsCitation id="14C63CACFFD6D42FFB96FD45FDD1FD4B" ID-GBIF-Occurrence="1806442530" collectionCode="IAvHP" country="Colombia" county="rio Sucio Municipality" location="Atrato River Basin, Tendal Creek, Parque Natural Nacional Los Katios" municipality="Sautata Stream (“ vereda ”)" pageId="19" pageNumber="20" specimenCode="7209" specimenCount="10" specimenCount-female="10" stateProvince="Choco">
<specimenCount id="B2A8FD78FFD6D42FFB96FD45FA82FD11" box="[1098,1165,661,687]" pageId="19" pageNumber="20" type="female">8 ♀♀</specimenCount>
, 
<quantity id="63569B14FFD6D42FFB47FD44FB49FD11" box="[1179,1350,660,687]" metricMagnitude="-2" metricUnit="m" metricValue="5.435" metricValueMax="7.609999999999999" metricValueMin="3.26" pageId="19" pageNumber="20" unit="mm" value="54.349999999999994" valueMax="76.1" valueMin="32.6">32.6–76.1 mm</quantity>
SL, 
<specimenCount id="B2A8FD78FFD6D42FFF61FD68FF0FFD6C" box="[189,256,696,722]" pageId="19" pageNumber="20" type="female">2 ♀♀</specimenCount>
C&amp;S, 
<quantity id="63569B14FFD6D42FFE8DFD67FFF2FD6C" box="[337,509,695,722]" metricMagnitude="-2" metricUnit="m" metricValue="5.904999999999999" metricValueMax="6.13" metricValueMin="5.68" pageId="19" pageNumber="20" unit="mm" value="59.05" valueMax="61.3" valueMin="56.8">56.8–61.3 mm</quantity>
SL, 
<collectingCounty id="4D704E7DFFD6D42FFDEAFD68FD4DFD6C" box="[566,834,696,722]" pageId="19" pageNumber="20">rio Sucio Municipality</collectingCounty>
, 
<collectingMunicipality id="4475AC8BFFD6D42FFC92FD68FA89FD6D" box="[846,1158,696,723]" pageId="19" pageNumber="20">Sautata Stream (“vereda”)</collectingMunicipality>
, 
<location id="A171602AFFD6D42FFB4CFD68FD27FD4B" LSID="urn:lsid:plazi:treatment:2C0787E7FFD7D429FE64FC00FFF8FA93:A171602AFFD6D42FFB4CFD68FD27FD4B" country="Colombia" county="rio Sucio Municipality" municipality="Sautata Stream (“ vereda ”)" name="Atrato River Basin, Tendal Creek, Parque Natural Nacional Los Katios" pageId="19" pageNumber="20" stateProvince="Choco">Atrato River Basin, Tendal Creek, Parque Natural Nacional Los Katios</location>
(
<collectionCode id="C2BFAE34FFD6D42FFCE6FD0BFD83FD4B" box="[826,908,731,757]" pageId="19" pageNumber="20">IAvHP</collectionCode>
<specimenCode id="F4089E8AFFD6D42FFC4BFD0BFDD8FD4B" box="[919,983,731,757]" pageId="19" pageNumber="20">7209</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="14C63CACFFD6D42FFC32FD0BFF69FC85" ID-GBIF-Occurrence="1806442541" country="Colombia" county="Sucio River" location="Atrato River" municipality="Sautata Stream" pageId="19" pageNumber="20" specimenCount="2" specimenCount-female="2" stateProvince="Choco">
<specimenCount id="B2A8FD78FFD6D42FFC32FD0BFA3AFD4B" box="[1006,1077,731,757]" pageId="19" pageNumber="20" type="female">2 ♀♀</specimenCount>
, 82,6– 
<quantity id="63569B14FFD6D42FFB54FD0BFAFAFD4B" box="[1160,1269,731,757]" metricMagnitude="-2" metricUnit="m" metricValue="9.94" pageId="19" pageNumber="20" unit="mm" value="99.4">99.4 mm</quantity>
SL, 
<collectingCounty id="4D704E7DFFD6D42FFAE8FD0BFF0FFCA6" pageId="19" pageNumber="20">Sucio River</collectingCounty>
, 
<collectingMunicipality id="4475AC8BFFD6D42FFED6FD2EFFB5FCA6" box="[266,442,766,792]" pageId="19" pageNumber="20">Sautata Stream</collectingMunicipality>
(“vereda”), 
<location id="A171602AFFD6D42FFD9BFD2EFCD6FCA6" LSID="urn:lsid:plazi:treatment:2C0787E7FFD7D429FE64FC00FFF8FA93:A171602AFFD6D42FFD9BFD2EFCD6FCA6" box="[583,729,766,792]" country="Colombia" county="Sucio River" municipality="Sautata Stream" name="Atrato River" pageId="19" pageNumber="20" stateProvince="Choco">Atrato River</location>
Basin,Tendal Creek, 
<location id="A171602AFFD6D42FFC0EFD2FFB78FCA6" LSID="urn:lsid:plazi:treatment:2C0787E7FFD7D429FE64FC00FFF8FA93:A171602AFFD6D42FFC0EFD2FFB78FCA6" box="[978,1399,766,793]" country="Colombia" county="Sucio River" municipality="Sautata Stream" name="Parque Natural Nacional Los Katios" pageId="19" pageNumber="20" stateProvince="Choco">Parque Natural Nacional Los Katios</location>
(IAvHP 7210)
</materialsCitation>
; 
<materialsCitation id="14C63CACFFD6D42FFEAEFCF1FB36FC85" ID-GBIF-Occurrence="1806442556" box="[370,1337,801,828]" collectionCode="IUQ" country="Colombia" location="Nati River" municipality="River Basin" pageId="19" pageNumber="20" specimenCode="IUQ 1319" specimenCount="1" specimenCount-male="1" stateProvince="Choco">
<specimenCount id="B2A8FD78FFD6D42FFEAEFCF1FFAFFC82" box="[370,416,801,828]" pageId="19" pageNumber="20" type="male">1 ♂</specimenCount>
, 
<quantity id="63569B14FFD6D42FFE71FCF1FC1AFC82" box="[429,533,801,828]" metricMagnitude="-2" metricUnit="m" metricValue="7.15" pageId="19" pageNumber="20" unit="mm" value="71.5">71.5 mm</quantity>
SL, Acandí, Atrato 
<collectingMunicipality id="4475AC8BFFD6D42FFD23FCF1FD81FC85" box="[767,910,801,827]" pageId="19" pageNumber="20">River Basin</collectingMunicipality>
, tributary of 
<location id="A171602AFFD6D42FFBFAFCF1FAAFFC85" LSID="urn:lsid:plazi:treatment:2C0787E7FFD7D429FE64FC00FFF8FA93:A171602AFFD6D42FFBFAFCF1FAAFFC85" box="[1062,1184,801,827]" country="Colombia" municipality="River Basin" name="Nati River" pageId="19" pageNumber="20" stateProvince="Choco">Nati River</location>
(
<specimenCode id="F4089E8AFFD6D42FFB6CFCF1FB23FC85" box="[1200,1324,801,827]" collectionCode="IUQ" country="Colombia" httpUri="http://grbio.org/cool/8xqv-ibyz" name="Laboratorio de Ictiologia" pageId="19" pageNumber="20">IUQ 1319</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="F0D16679FFD6D42FFF61F85AFA4BF855" httpUri="https://zenodo.org/record/1169575/files/figure.png" pageId="19" pageNumber="20" startId="19.[189,232,1930,1956]" targetBox="[189,1397,871,1894]" targetPageId="19">
<paragraph id="A41136F1FFD6D42FFF61F85AFA4BF855" blockId="19.[189,1398,1930,2028]" pageId="19" pageNumber="20">
<emphasis id="96DAEAE3FFD6D42FFF61F85AFF03F81A" bold="true" box="[189,268,1930,1956]" pageId="19" pageNumber="20">Fig. 8.</emphasis>
<taxonomicName id="63AE4D72FFD6D42FFECAF85BFCDEF81A" authority="Eigenmann, 1907" authorityName="Eigenmann in Eigenmann &amp; Ogle" authorityYear="1907" box="[278,721,1930,1957]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="orthodus">
<emphasis id="96DAEAE3FFD6D42FFECAF85BFFFCF81A" box="[278,499,1930,1957]" italics="true" pageId="19" pageNumber="20">Astyanax orthodus</emphasis>
Eigenmann, 1907
</taxonomicName>
. 
<emphasis id="96DAEAE3FFD6D42FFD00F85AFCFEF81A" bold="true" box="[732,753,1930,1956]" pageId="19" pageNumber="20">A</emphasis>
. Holotype, 88.3 mm SL, Truando River, Atrato River Basin, Chocó, Pacific coast of Colombia (USNM 55655). 
<emphasis id="96DAEAE3FFD6D42FFCBAF87EFD75F876" bold="true" box="[870,890,1966,1992]" pageId="19" pageNumber="20">B</emphasis>
. 32.5 mm SL, Vereda Sautatá, Atrato River drainage, Chocó, Pacific coast of Colombia (IAVH 7146). Scale bars = 1 cm.
</paragraph>
</caption>
<subSubSection id="ECB4657AFFD1D429FF61FEDBFC7AFBB9" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
<paragraph id="A41136F1FFD1D428FF61FEDBFF41FE9B" blockId="20.[189,334,267,293]" box="[189,334,267,293]" pageId="20" pageNumber="21">
<heading id="FF59819DFFD1D428FF61FEDBFF41FE9B" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="96DAEAE3FFD1D428FF61FEDBFF41FE9B" bold="true" box="[189,334,267,293]" pageId="20" pageNumber="21">Description</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFD1D428FF61FEE9FF23FE5E" blockId="20.[189,1399,313,480]" pageId="20" pageNumber="21">Body compressed, greatest body depth at or anterior to dorsal-fin origin. Mouth terminal. Dorsal anterior profile of head sigmoid, concave between snout tip and posterior margin of supraoccipital crest, convex between supraoccipital and dorsal fin, convex between last dorsal-fin ray and adipose-fin origin. Dorsal and ventral caudal peduncle margins straight. Ventral profile convex from tip of snout to pelvic-fin insertion.</paragraph>
<paragraph id="A41136F1FFD1D428FF61FDD8FDBCFD10" blockId="20.[189,1399,520,687]" pageId="20" pageNumber="21">Premaxillary teeth in two series; outer series with four tricuspid teeth covering three internal teeth of inner series; inner row with five pentacuspid teeth. Maxilla long, of same width along entire length, with 2–3 tricuspid teeth set in anterior most part of ventral margin. Dentary with anterior four teeth pentacuspid, followed laterally by 8–10 smaller teeth of increasing posterior inclination and reducing from tri to unicuspid, total number of lateral teeth quite variable.</paragraph>
<paragraph id="A41136F1FFD1D428FF61FD06FFE4FBEE" blockId="20.[189,1399,726,1105]" pageId="20" pageNumber="21">
Pored lateral-line scales 35(1), 37(6), 38(10), 39(4), 40(6), 41(1), 42(1) (n = 29), scales between lateral line and origin of dorsal fin 7(27), 8(3) (n = 30), scales between lateral line and origin of anal fin 8 (4), 9(24) (n = 28), scales between lateral line and pelvic-fin insertion 6(27), 7(3) (n = 30). Dorsal-fin rays 
<date id="D0101031FFD1D428FF61FCEFFEE0FCE4" box="[189,239,831,858]" pageId="20" pageNumber="21">iii 9</date>
(30), predorsal midline covered with medial, bilobed scales for more that ¾ of its length, naked anteriorly. Postdorsal profile convex, from last dorsal-fin ray to adipose-fin origin; first simple ray small, only visible in cleared and stained specimens; second simple ray about half length of third simple ray. Distal margin of dorsal fin slightly convex. Adipose-fin origin anterior to vertical through insertion of last anal-fin ray. Pectoral-fin rays 
<date id="D0101031FFD1D428FD8DFC1CFC70FC58" box="[593,639,972,998]" pageId="20" pageNumber="21" value="2011-01">i 11</date>
ii (15), 
<date id="D0101031FFD1D428FD3FFC1CFD1EFC58" box="[739,785,972,998]" pageId="20" pageNumber="21" value="2012-01">i 12</date>
(7) (n = 22). Anal-fin rays 
<date id="D0101031FFD1D428FB8EFC1CFAA2FC58" box="[1106,1197,972,998]" pageId="20" pageNumber="21" value="1924-04" valueMax="1924-05-31" valueMin="1924-04-24">iv–v 24</date>
(1), 26(1), 27(6), 28(8), 29(5), 30(3), 31(1) (n = 27). Anal-fin origin posterior to vertical through insertion of last dorsalfin ray (
<tableCitation id="E92C034AFFD1D428FEC7FBC3FF7AFB93" box="[283,373,1043,1069]" captionStart="Table 1" captionStartId="11.[145,214,599,625]" captionTargetPageId="11" captionText="Table 1. Morphometric data for some members of the Astyanax orthodus species-group." httpUri="http://table.plazi.org/id/F0D16679FFCED437FF4DFD87FA92FDCF" pageId="20" pageNumber="21" tableUuid="F0D16679FFCED437FF4DFD87FA92FDCF">Table 1</tableCitation>
). Caudal fin with 10 (6) leading principal rays in dorsal and ventral lobes, each lobes with 8(6) procurrent rays.
</paragraph>
<paragraph id="A41136F1FFD1D428FF61FBA7FD3EFAA0" blockId="20.[189,1398,1143,1310]" pageId="20" pageNumber="21">Total vertebrae 34(4), 35(2), including those of Weberian apparatus: 16(4)–17(2) precaudal centra, last three without ribs; caudal centra 18(4)–19(2). Epipleurals 20(4)–21(2). Epineurals 30(4), 31(2); posterior most epineural may reach anterior surface of urostyle. Hypurals 7(6), first dorsal hypural with anterior margin swollen, without projections that articulate with the urostyle; second and third hypurals with anterior margin swollen and contacting urostyle.</paragraph>
<caption id="F0D16679FFD1D428FF61F892FC78F857" httpUri="https://zenodo.org/record/1169577/files/figure.png" pageId="20" pageNumber="21" startId="20.[189,232,1858,1884]" targetBox="[221,1398,1332,1820]" targetPageId="20">
<paragraph id="A41136F1FFD1D428FF61F892FC78F857" blockId="20.[189,1399,1858,2025]" pageId="20" pageNumber="21">
<emphasis id="96DAEAE3FFD1D428FF61F892FF1FF8E2" bold="true" box="[189,272,1858,1884]" pageId="20" pageNumber="21">Fig. 9.</emphasis>
Squamation of pectoral-fin base, ventral view. 
<emphasis id="96DAEAE3FFD1D428FCBEF892FD78F8E2" bold="true" box="[866,887,1858,1884]" pageId="20" pageNumber="21">A</emphasis>
. 
<taxonomicName id="63AE4D72FFD1D428FC50F892FB7EF8E3" authority="(Linnaeus, 1758)" authorityName="Linnaeus" authorityYear="1758" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[908,1393,1858,1885]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="bimaculatus">
<emphasis id="96DAEAE3FFD1D428FC50F892FA9BF8E2" box="[908,1172,1858,1884]" italics="true" pageId="20" pageNumber="21">Astyanax bimaculatus</emphasis>
(Linnaeus, 1758)
</taxonomicName>
. 
<emphasis id="96DAEAE3FFD1D428FF61F8B5FEDEF8C1" bold="true" box="[189,209,1893,1919]" pageId="20" pageNumber="21">B</emphasis>
. 
<taxonomicName id="63AE4D72FFD1D428FF38F8B6FDF0F83E" authority="(Torres-Mejia, Hernandez &amp; Senechal, 2012)" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" baseAuthorityName="Torres-Mejia, Hernandez &amp; Senechal" baseAuthorityYear="2012" box="[228,1023,1893,1920]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="yariguies" status="comb. nov.">
<emphasis id="96DAEAE3FFD1D428FF38F8B6FFC6F8C1" box="[228,457,1894,1920]" italics="true" pageId="20" pageNumber="21">Astyanax yariguies</emphasis>
(Torres-Mejía, Hernández &amp; Senechal, 2012)
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFD1D428FBD0F8B5FA9CF83E" box="[1036,1171,1893,1920]" pageId="20" pageNumber="21">comb. nov.</taxonomicNameLabel>
In A, the pore of pectoral-fin is in epithelial tissue, covered or circumscribed by three scales indicated by the curved line, while in B, the pore of the base of pectoral-fin is part of a scale circumscribed by four scales, indicated by the curved line. Scale bars = 1 cm.
</paragraph>
</caption>
<paragraph id="A41136F1FFD0D429FF61FEDBFFE9FE9B" blockId="21.[189,486,267,293]" box="[189,486,267,293]" pageId="21" pageNumber="22">
<heading id="FF59819DFFD0D429FF61FEDBFFE9FE9B" bold="true" box="[189,486,267,293]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="96DAEAE3FFD0D429FF61FEDBFFE9FE9B" bold="true" box="[189,486,267,293]" pageId="21" pageNumber="22">Pigmentation in alcohol</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFD0D429FF61FEEAFA6AFCCF" blockId="21.[189,1399,314,882]" pageId="21" pageNumber="22">
Sides of body yellowish, with reticulated pattern predominant over upper region of coelomic cavity, silvery stripe present from humeral region to caudal-peduncle base, overlain by series of chevronshaped marks formed by dark lines along myosepta between myotomes extending from dorsal region of coelomic cavity to caudal peduncle; pigmented muscle septae forming chevrons not coinciding with scale rows. Chevrons present in juveniles, immature specimens and adults, without distal extensions. Dorsal region of head and body chestnut brown. Sides of head and ventral surface of body light brown, not silvery. Melanophores of humeral region forming two spots. Anterior spot formed by two layers of pigment: brown melanophores distributed in a thin superficial layer of the epithelium (Layer 1, 
<figureCitation id="3C952A74FFD0D429FAFDFDE7FB6BFDEF" box="[1313,1380,567,593]" captionStart="Fig.1" captionStartId="4.[189,232,1859,1885]" captionTargetBox="[189,1398,540,1837]" captionTargetId="figure@4.[189,1398,540,1837]" captionTargetPageId="4" captionText="Fig.1.A.DistributionofpigmentinthehumeralregionofAstyanaxspecies.B.A.bimaculatus(Linnaeus, 1758).C.A.caucanus(Steindachner,1879).D.A.mexicanus(DeFilippi,1853).E.A.courensisBertaco, Carvalho &amp; Jerep, 2010 (modified from Bertaco et al. 2010). The anterior humeral spot consists of two overlapping layers of pigment identified by numbers 1 and 2 (see Román-Valencia et al. 2015: fig. 1): 1. Diffuse vertical bar. 2. Horizontal spot with concentrated pigments." httpUri="https://zenodo.org/record/1169561/files/figure.png" pageId="21" pageNumber="22">Fig. 1</figureCitation>
), deeper layer of dark melanophores (Layer 2, 
<figureCitation id="3C952A74FFD0D429FD16FD8BFD02FDCB" box="[714,781,603,629]" captionStart="Fig.1" captionStartId="4.[189,232,1859,1885]" captionTargetBox="[189,1398,540,1837]" captionTargetId="figure@4.[189,1398,540,1837]" captionTargetPageId="4" captionText="Fig.1.A.DistributionofpigmentinthehumeralregionofAstyanaxspecies.B.A.bimaculatus(Linnaeus, 1758).C.A.caucanus(Steindachner,1879).D.A.mexicanus(DeFilippi,1853).E.A.courensisBertaco, Carvalho &amp; Jerep, 2010 (modified from Bertaco et al. 2010). The anterior humeral spot consists of two overlapping layers of pigment identified by numbers 1 and 2 (see Román-Valencia et al. 2015: fig. 1): 1. Diffuse vertical bar. 2. Horizontal spot with concentrated pigments." httpUri="https://zenodo.org/record/1169561/files/figure.png" pageId="21" pageNumber="22">Fig. 1</figureCitation>
). Layer 2 forming a polygon-shaped spot, consisting of two groups of melanophores that do not precisely overlap, forming four-sided spot, extending from third to sixth or seventh scale of lateral line. Posterior humeral spot situated two or three scales posterior to the anterior humeral spot, arc- or sigmoid-shaped, inconspicuous, covering two to three scales above lateral line. Scales on sides of body with spots or dots. Spot on caudal peduncle pentagonal, located over lateral axis of peduncle, prolonged on posterior tips of middle caudal-fin rays, not extending to dorsal and ventral margins of caudal peduncle. Pectoral fins mostly hyaline, melanophores present on distal tips of pelvic-fin rays and interradial membranes of dorsal, caudal and anal fins.
</paragraph>
<paragraph id="A41136F1FFD0D429FF61FC49FFBFFC0D" blockId="21.[189,432,921,947]" box="[189,432,921,947]" pageId="21" pageNumber="22">
<heading id="FF59819DFFD0D429FF61FC49FFBFFC0D" bold="true" box="[189,432,921,947]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="96DAEAE3FFD0D429FF61FC49FFBFFC0D" bold="true" box="[189,432,921,947]" pageId="21" pageNumber="22">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFD0D429FF61FC19FC7AFBB9" blockId="21.[189,1399,969,1031]" pageId="21" pageNumber="22">Sexually dimorphic characters observed in one male, consisting of 7 to 12 small hooks located on distal portions of all branched anal-fin rays.</paragraph>
</subSubSection>
<subSubSection id="ECB4657AFFD0D429FF61FBFFFFF8FA93" pageId="21" pageNumber="22" type="distribution">
<paragraph id="A41136F1FFD0D429FF61FBFFFF59FBF7" blockId="21.[189,342,1071,1097]" box="[189,342,1071,1097]" pageId="21" pageNumber="22">
<heading id="FF59819DFFD0D429FF61FBFFFF59FBF7" bold="true" box="[189,342,1071,1097]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="96DAEAE3FFD0D429FF61FBFFFF59FBF7" bold="true" box="[189,342,1071,1097]" pageId="21" pageNumber="22">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFD0D429FF61FB8FFFF8FA93" blockId="21.[189,1399,1119,1325]" pageId="21" pageNumber="22">
<taxonomicName id="63AE4D72FFD0D429FF61FB8FFF96FBC7" authorityName="Eigenmann in Eigenmann &amp; Ogle" authorityYear="1907" box="[189,409,1119,1145]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="orthodus">
<emphasis id="96DAEAE3FFD0D429FF61FB8FFF96FBC7" box="[189,409,1119,1145]" italics="true" pageId="21" pageNumber="22">Astyanax orthodus</emphasis>
</taxonomicName>
has been reported from the Pacific slopes of 
<collectingRegion id="666AF813FFD0D429FC6BFB8FFA02FBC7" box="[951,1037,1119,1145]" country="Paraguay" name="Central" pageId="21" pageNumber="22">Central</collectingRegion>
America (
<bibRefCitation id="C03F4B00FFD0D429FB57FB8FFB61FBC7" author="Angulo A. &amp; Garita-Alvarado C. A. &amp; Bussing W. A. &amp; Lopez M. I." box="[1163,1390,1119,1145]" journalOrPublisher="Check List" pageId="21" pageNumber="22" pagination="987 - 1019" part="9" publicationUrl="https://doi.org/10.15560/9.5.987" refId="ref18780" refString="Angulo A., Garita-Alvarado C. A., Bussing W. A. &amp; Lopez M. I. 2013. Annotated checklist of the freshwater fishes of continental and insular Costa Rica: additions and nomenclatural revisions. Check List 9: 987 - 1019. https: // doi. org / 10.15560 / 9.5.987" title="Annotated checklist of the freshwater fishes of continental and insular Costa Rica: additions and nomenclatural revisions" type="journal article" year="2013">
Angulo 
<emphasis id="96DAEAE3FFD0D429FB31FBB0FB26FBC7" box="[1261,1321,1119,1145]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2013
</bibRefCitation>
) and 
<collectingCountry id="DCB97661FFD0D429FF2FFB53FF68FB23" box="[243,359,1155,1181]" name="Colombia" pageId="21" pageNumber="22">Colombia</collectingCountry>
in South America (
<bibRefCitation id="C03F4B00FFD0D429FD82FB53FD9EFB23" author="Torres-Mejia M. &amp; Hernandez E. &amp; Senechal V." box="[606,913,1155,1181]" journalOrPublisher="Copeia" pageId="21" pageNumber="22" pagination="501 - 506" publicationUrl="https://doi.org/10.1643/CI-10-160" refId="ref20676" refString="Torres-Mejia M., Hernandez E. &amp; Senechal V. 2012. A new species of Astyanacinus (Characiformes: Characidae) from the Rio Magdalena System, Colombia. Copeia 2012: 501 - 506. https: // doi. org / 10.1643 / CI- 10 - 160" title="A new species of Astyanacinus (Characiformes: Characidae) from the Rio Magdalena System, Colombia" type="book chapter" year="2012">
Torres-Mejía 
<emphasis id="96DAEAE3FFD0D429FCD9FB54FD4AFB23" box="[773,837,1155,1181]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2012
</bibRefCitation>
). However, based on the results of this study, 
<taxonomicName id="63AE4D72FFD0D429FED0FB77FF97FB7F" authorityName="Eigenmann in Eigenmann &amp; Ogle" authorityYear="1907" box="[268,408,1191,1217]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="orthodus">
<emphasis id="96DAEAE3FFD0D429FED0FB77FF97FB7F" box="[268,408,1191,1217]" italics="true" pageId="21" pageNumber="22">A. orthodus</emphasis>
</taxonomicName>
occurs only in the lower Atrato River Basin, which flows into the Caribbean Sea in the Truando Department of north-western 
<collectingCountry id="DCB97661FFD0D429FD0BFB1BFD44FB5B" box="[727,843,1227,1253]" name="Colombia" pageId="21" pageNumber="22">Colombia</collectingCountry>
(
<figureCitation id="3C952A74FFD0D429FC81FB1BFDABFB5B" box="[861,932,1227,1253]" captionStart="Fig. 4" captionStartId="8.[189,232,2000,2026]" captionTargetBox="[189,1392,764,1921]" captionTargetId="figure@8.[151,1417,755,1932]" captionTargetPageId="8" captionText="Fig. 4.Distribution of speciesof the Astyanaxorthodus species-group(symbol includestypelocalities)." httpUri="https://zenodo.org/record/1169567/files/figure.png" pageId="21" pageNumber="22">Fig. 4</figureCitation>
). Populations identified as 
<taxonomicName id="63AE4D72FFD0D429FB30FB1BFB79FB5B" authorityName="Eigenmann in Eigenmann &amp; Ogle" authorityYear="1907" box="[1260,1398,1227,1253]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="orthodus">
<emphasis id="96DAEAE3FFD0D429FB30FB1BFB79FB5B" box="[1260,1398,1227,1253]" italics="true" pageId="21" pageNumber="22">A. orthodus</emphasis>
</taxonomicName>
outside of the Atrato River Basin and similar to 
<taxonomicName id="63AE4D72FFD0D429FD36FB3FFD6DFAB7" authority="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega, 2018" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" box="[746,866,1263,1289]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="embera" status="sp. nov.">
<emphasis id="96DAEAE3FFD0D429FD36FB3FFD6DFAB7" box="[746,866,1263,1289]" italics="true" pageId="21" pageNumber="22">A. embera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFD0D429FCB5FB3FFDCAFAB7" box="[873,965,1263,1289]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
, described herein, represent an as yet unrecognized new species.
</paragraph>
</subSubSection>
</treatment>
</document>