treatments-xml/data/25/36/27/253627CA174B5C69B8B6231C187A0709.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.994.56810" ID-GBIF-Dataset="8639369d-0b63-4725-aa0c-aed78224bbe5" ID-PMC="PMC7686225" ID-Pensoft-Pub="1313-2970-994-1" ID-Pensoft-UUID="11A23FB601685F7ABFCC123875393447" ID-PubMed="33273881" ID-ZooBank="DB1879C919044320BE674D9F863F6775" ModsDocID="1313-2970-994-1" checkinTime="1605693934925" checkinUser="pensoft" docAuthor="Ng, Peter K. L." docDate="2020" docId="253627CA174B5C69B8B6231C187A0709" docLanguage="en" docName="ZooKeys 994: 1-34" docOrigin="ZooKeys 994" docSource="http://dx.doi.org/10.3897/zookeys.994.56810" docTitle="Johora booliati Ng 2020, sp. nov." docType="treatment" docUuid="8A3C9173-A7A5-4559-92D7-A548453C6688" docUuidSource="ZooBank" docVersion="4" id="11A23FB601685F7ABFCC123875393447" lastPageNumber="1" masterDocId="11A23FB601685F7ABFCC123875393447" masterDocTitle="Revision of the freshwater crabs of the Johora tahanensis (Bott, 1966) species group (Crustacea, Brachyura, Potamidae), with a key to the genus" masterLastPageNumber="34" masterPageNumber="1" pageNumber="1" updateTime="1668169916828" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the freshwater crabs of the Johora tahanensis (Bott, 1966) species group (Crustacea, Brachyura, Potamidae), with a key to the genus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ng, Peter K. L.</mods:namePart>
<mods:affiliation>Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Singapore</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5946-0608</mods:nameIdentifier>
<mods:nameIdentifier type="email">peterng@nus.edu.sg</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>994</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>34</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.994.56810</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.994.56810</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-994-1</mods:identifier>
<mods:identifier type="ZooBank">DB1879C919044320BE674D9F863F6775</mods:identifier>
<mods:identifier type="Pensoft-UUID">11A23FB601685F7ABFCC123875393447</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="169094345" LSID="urn:lsid:zoobank.org:act:8A3C9173-A7A5-4559-92D7-A548453C6688" httpUri="http://treatment.plazi.org/id/253627CA174B5C69B8B6231C187A0709" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="253627CA-174B-5C69-B8B6-231C187A0709" authority="Ng, 2020" authorityName="Ng" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora booliati" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="booliati" status="sp. nov.">Johora booliati</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">Figures 1F</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">, 2F</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">, 3F</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Left third maxilliped A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure4" httpUri="https://binary.pensoft.net/fig/476315" pageId="0" pageNumber="1">, 4F</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Anterior thoracic sternum and pleon A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure5" httpUri="https://binary.pensoft.net/fig/476316" pageId="0" pageNumber="1">, 5F</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">, 10E-I</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">, 12F</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pleon A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure13" httpUri="https://binary.pensoft.net/fig/476324" pageId="0" pageNumber="1">, 13F</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Sternopleonal cavity and vulvae A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure14" httpUri="https://binary.pensoft.net/fig/476325" pageId="0" pageNumber="1">, 14F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Bott" authorityYear="1966" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="tahanensis">Johora tahanensis</taxonomicName>
- Takeda &amp; Ng, 1992: 107 (part); Ng 1995: 249 (part) (not 
<taxonomicName class="Malacostraca" family="Potamidae" genus="Stoliczia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stoliczia (Johora) johorensis subsp. tahanensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subSpecies" species="johorensis" subGenus="Johora" subSpecies="tahanensis">Stoliczia (Johora) johorensis tahanensis</taxonomicName>
Bott, 1966).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: male (34.6 
<normalizedToken originalValue="×">x</normalizedToken>
30.0 mm) (ZRC 2020.0072), stream next to house, under rocks, in old farmland and secondary forest, Bukit Tinggi village, west side of northbound Karak Highway, ca. 480 m a.s.l., Bentong District, Pahang, 
<geoCoordinate degrees="3" direction="north" minutes="22" orientation="latitude" precision="1" seconds="3.0396" value="3.367511">3°22'3.0396&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="101" direction="east" minutes="48" orientation="longitude" precision="1" seconds="50.994" value="101.81417">101°48'50.994&quot;E</geoCoordinate>
, coll. July 2016. Paratype: 1 female (40.4 
<normalizedToken originalValue="×">x</normalizedToken>
34.4 mm) (ZRC 1995.270), in shallow stream with leaf litter, The Gap, 
<normalizedToken originalValue="Fraser’s">Fraser's</normalizedToken>
Hill, 853 m a.s.l., Pahang, ca. 
<geoCoordinate degrees="3" direction="north" minutes="41" orientation="latitude" precision="15" seconds="29" value="3.6913888">3°41'29&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="101" direction="east" minutes="44" orientation="longitude" precision="15" seconds="56" value="101.748886">101°44'56&quot;E</geoCoordinate>
, coll. Lim KKP, 1 June 1990; 1 juvenile male (10.5 
<normalizedToken originalValue="×">x</normalizedToken>
9.1 mm) (ZRC 2020.0364), under rocks, first stream on road towards Raub, descending from 
<normalizedToken originalValue="Fraser’s">Fraser's</normalizedToken>
Hill, Pahang, coll. Lai JCY, 6 December 2015. All locations in Peninsular Malaysia.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Adult carapace width to length ratio 1.15-1.17 (Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1F</figureCitation>
, 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2F</figureCitation>
, 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">12F</figureCitation>
); dorsal surface gently convex in frontal view, not inflated (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3F</figureCitation>
); frontal margin almost straight (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2F</figureCitation>
); suborbital, pterygostomial and sub-branchial regions rugose, pterygostomial region covered with dense setae (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3F</figureCitation>
); epigastric cristae distinct, distinctly anterior to sharp postorbital cristae, postorbital cristae with lateral edges low, joining lateral margin through oblique striae (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2F</figureCitation>
); external orbital tooth separated from epibranchial tooth by distinct cleft, epibranchial tooth sharp, distinct (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2F</figureCitation>
); anterolateral margin gently convex (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2F</figureCitation>
); posterolateral margin gently sinuous to almost straight, gently converging towards sinuous posterior carapace margin with shallow median indentation (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2F</figureCitation>
); posterior margin of epistome with triangular median triangle, lateral margin obliquely sloping (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3F</figureCitation>
); outer surfaces of third maxillipeds with dense, long stiff setae; ischium subrectangular, with shallow median oblique groove (Figs 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Frontal view of cephalothorax A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure3" httpUri="https://binary.pensoft.net/fig/476314" pageId="0" pageNumber="1">3F</figureCitation>
, 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Left third maxilliped A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure4" httpUri="https://binary.pensoft.net/fig/476315" pageId="0" pageNumber="1">4F</figureCitation>
); ambulatory legs not elongate, length to width ratio of merus of fourth ambulatory leg 2.7-2.8 (Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Overall dorsal view A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure1" httpUri="https://binary.pensoft.net/fig/476312" pageId="0" pageNumber="1">1F</figureCitation>
, 
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Overall dorsal view A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1">12F</figureCitation>
); G1 subterminal segment with broad proximal part, tapering relatively suddenly to slender distal part, with low shelf-like structure on outer margin; terminal segment gently curved outwards (from median part of sternum), less than half length of subterminal segment, surfaces with numerous short setae (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10E-H</figureCitation>
); G2 slightly longer than G1, distal segment long, about half length of basal segment (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10I</figureCitation>
). Female pleon ovate; somite 3 less wide than somite 4, somites 4-6 progressively narrower; telson semi-circular (Fig. 
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Female pleon A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure13" httpUri="https://binary.pensoft.net/fig/476324" pageId="0" pageNumber="1">13F</figureCitation>
). Vulvae large, on anterior half of sternite 6, slightly pushing into suture with sternite 5, lateral sternal vulvar cover semi-circular (Fig. 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Sternopleonal cavity and vulvae A Johora intermedia Ng, 1986, female (17.9 x 14.7 mm) (ZRC 2001.1002), Selangor B J. tahanensis (Bott, 1966), female (24.4 x 21.0 mm) (ZRC 1984.6765), Pahang C J. thoi Ng, 1990, female (31.7 x 25.5 mm) (ZRC 2001.1167), Terengganu D J. hoiseni Ng &amp; Takeda, 1992, paratype female (22.1 x 19.0 mm) (ZRC 1984.6675), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 x 17.4 mm) (ZRC 2006.0053), Thailand F J. booliati sp. nov., paratype female (40.4 x 34.4 mm) (ZRC 1995.270), Pahang G J. erici sp. nov., paratype female (32.8 x 26.9 mm) (ZRC 1995.268), Perak H J. michaeli sp. nov., paratype female (29.9 x 25.0 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure14" httpUri="https://binary.pensoft.net/fig/476325" pageId="0" pageNumber="1">14F</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.994.56810.figure12" httpUri="https://binary.pensoft.net/fig/476323" pageId="0" pageNumber="1" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
Overall dorsal view 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName baseAuthorityName="Ng" baseAuthorityYear="1986" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora intermedia" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis italics="true" pageId="0" pageNumber="1">Johora intermedia</emphasis>
</taxonomicName>
Ng, 1986, female (17.9 
<normalizedToken originalValue="×">x</normalizedToken>
14.7 mm) (ZRC 2001.1002), Selangor 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
<taxonomicName lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
</taxonomicName>
(Bott, 1966), female (24.4 
<normalizedToken originalValue="×">x</normalizedToken>
21.0 mm) (ZRC 1984.6765), Pahang 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
<taxonomicName lsidName="J. thoi" pageId="0" pageNumber="1" rank="species" species="thoi">
<emphasis italics="true" pageId="0" pageNumber="1">J. thoi</emphasis>
</taxonomicName>
Ng, 1990, female (31.7 
<normalizedToken originalValue="×">x</normalizedToken>
25.5 mm) (ZRC 2001.1167), Terengganu 
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
<taxonomicName lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
<emphasis italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
</taxonomicName>
Ng &amp; Takeda, 1992, paratype female (22.1 
<normalizedToken originalValue="×">x</normalizedToken>
19.0 mm) (ZRC 1984.6675), Kelantan 
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
<taxonomicName lsidName="J. thaiana" pageId="0" pageNumber="1" rank="species" species="thaiana">
<emphasis italics="true" pageId="0" pageNumber="1">J. thaiana</emphasis>
</taxonomicName>
Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype female (21.0 
<normalizedToken originalValue="×">x</normalizedToken>
17.4 mm) (ZRC 2006.0053), Thailand 
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
sp. nov., paratype female (40.4 
<normalizedToken originalValue="×">x</normalizedToken>
34.4 mm) (ZRC 1995.270), Pahang 
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
sp. nov., paratype female (32.8 
<normalizedToken originalValue="×">x</normalizedToken>
26.9 mm) (ZRC 1995.268), Perak 
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
<taxonomicName lsidName="J. michaeli" pageId="0" pageNumber="1" rank="species" species="michaeli">
<emphasis italics="true" pageId="0" pageNumber="1">J. michaeli</emphasis>
</taxonomicName>
sp. nov., paratype female (29.9 
<normalizedToken originalValue="×">x</normalizedToken>
25.0 mm) (ZRC 2010.0047), Terengganu.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The species is named after an old friend and mentor, the late Dr Lim Boo Liat (1926-2020), one of 
<normalizedToken originalValue="Malaysia’s">Malaysia's</normalizedToken>
most eminent biologists and naturalists. In a remarkable career spanning 70 years, he has contributed immensely to biodiversity and conservation science in Malaysia; more importantly, he has inspired, mentored, and trained many generations of biologists, many who now lead science in Malaysia and Singapore.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
The large adult size of 
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
sp. nov. (carapace width in excess of 25 mm) with the pronounced epibranchial teeth allies them with 
<taxonomicName lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
</taxonomicName>
s. str., 
<taxonomicName lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
<emphasis italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="J. thaiana" pageId="0" pageNumber="1" rank="species" species="thaiana">
<emphasis italics="true" pageId="0" pageNumber="1">J. thaiana</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="J. thoi" pageId="0" pageNumber="1" rank="species" species="thoi">
<emphasis italics="true" pageId="0" pageNumber="1">J. thoi</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="J. tiomanensis" pageId="0" pageNumber="1" rank="species" species="tiomanensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. tiomanensis</emphasis>
</taxonomicName>
, and 
<taxonomicName lsidName="J. counsilmani" pageId="0" pageNumber="1" rank="species" species="counsilmani">
<emphasis italics="true" pageId="0" pageNumber="1">J. counsilmani</emphasis>
</taxonomicName>
(the last two being endemic to Pulau Tioman, Peninsular Malaysia).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Ng" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora booliati" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">Johora booliati</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
resemble 
<taxonomicName lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
<emphasis italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
</taxonomicName>
, and 
<taxonomicName lsidName="J. thaiana" pageId="0" pageNumber="1" rank="species" species="thaiana">
<emphasis italics="true" pageId="0" pageNumber="1">J. thaiana</emphasis>
</taxonomicName>
in the carapace shape and general features. Their G1 structures, however, are different. In 
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
, the G1 subterminal segment is proportionately more elongate and the terminal segment is relatively shorter (Figs 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10E-H</figureCitation>
, 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A-D Johora erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak E-H J. erici sp. nov., male (41.2 x 33.8 mm) (ZRC 1995.0269), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure11" httpUri="https://binary.pensoft.net/fig/476322" pageId="0" pageNumber="1">11A-C, E-G</figureCitation>
), and significantly, the terminal segment has only scattered short setae (versus subterminal segment shorter with the terminal segment longer and the surface of the latter more densely covered with short setae in 
<taxonomicName lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
</taxonomicName>
(Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">8A-C</figureCitation>
). Both 
<taxonomicName lsidName="J. hoiseni" pageId="0" pageNumber="1" rank="species" species="hoiseni">
<emphasis italics="true" pageId="0" pageNumber="1">J. hoiseni</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="J. thaiana" pageId="0" pageNumber="1" rank="species" species="thaiana">
<emphasis italics="true" pageId="0" pageNumber="1">J. thaiana</emphasis>
</taxonomicName>
have relatively straight G1 terminal segments which are prominently setose (Figs 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-D Johora tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang E-H J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view) E-H after Ng and Takeda (1992: fig. 3 B, D-F). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure8" httpUri="https://binary.pensoft.net/fig/476319" pageId="0" pageNumber="1">8E-G</figureCitation>
, 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10A-C</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Ng" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora booliati" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">Johora booliati</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName lsidName="J. tiomanensis" pageId="0" pageNumber="1" rank="species" species="tiomanensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. tiomanensis</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="J. counsilmani" pageId="0" pageNumber="1" rank="species" species="counsilmani">
<emphasis italics="true" pageId="0" pageNumber="1">J. counsilmani</emphasis>
</taxonomicName>
by their carapace being relatively more quadrate in shape (width to length ratio less than 1.2) and the adult carapace is distinctly wider than long (width to length ratio 1.3-1.4) in 
<taxonomicName lsidName="J. tiomanensis" pageId="0" pageNumber="1" rank="species" species="tiomanensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. tiomanensis</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="J. counsilmani" pageId="0" pageNumber="1" rank="species" species="counsilmani">
<emphasis italics="true" pageId="0" pageNumber="1">J. counsilmani</emphasis>
</taxonomicName>
. In addition, the epigastric cristae of 
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
are distinctly anterior to the postorbital cristae and separated by a gap with the entire postorbital cristae sharp (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2F, G</figureCitation>
, 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Johora erici sp. nov., male (41.2 x 33.8 mm) (ZRC 1995.0269), Kelantan A overall dorsal view B dorsal view of carapace C frontal view of cephalothorax D anterior thoracic sternum and pleon." figureDoi="10.3897/zookeys.994.56810.figure7" httpUri="https://binary.pensoft.net/fig/476318" pageId="0" pageNumber="1">7B</figureCitation>
) while in 
<taxonomicName lsidName="J. tiomanensis" pageId="0" pageNumber="1" rank="species" species="tiomanensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. tiomanensis</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="J. counsilmani" pageId="0" pageNumber="1" rank="species" species="counsilmani">
<emphasis italics="true" pageId="0" pageNumber="1">J. counsilmani</emphasis>
</taxonomicName>
, the epigastric cristae are only slightly anterior of the postorbital cristae and separated only by striae (cf. 
<bibRefCitation author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="167 - 174" refId="B20" refString="Ng, PKL, Tan, LWH, 1984. A new freshwater crab, Stoliczia (Johora) tiomanensis n. sp. (Decapoda: Brachyura: Potamidae) from Pulau Tioman, West Malaysia. Malayan Nature Journal 37 (3): 167 - 174" title="A new freshwater crab, Stoliczia (Johora) tiomanensis n. sp. (Decapoda: Brachyura: Potamidae) from Pulau Tioman, West Malaysia." volume="37" year="1984">Ng and Tan 1984</bibRefCitation>
: figs 1A, 2A; 
<bibRefCitation author="Ng, PKL" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="1" pagination="149 - 162" refId="B8" refString="Ng, PKL, 1985. Freshwater decapod crustaceans from Pulau Tioman, West Malaysia. Zoologische Mededelingen 59 (14): 149 - 162" title="Freshwater decapod crustaceans from Pulau Tioman, West Malaysia." volume="59" year="1985">Ng 1985</bibRefCitation>
: fig. 3). Most significantly, the G1 terminal segments of 
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
are proportionately stouter and less curved (Figs 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10E-H</figureCitation>
, 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A-D Johora erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak E-H J. erici sp. nov., male (41.2 x 33.8 mm) (ZRC 1995.0269), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure11" httpUri="https://binary.pensoft.net/fig/476322" pageId="0" pageNumber="1">11A-C, E-G</figureCitation>
); in 
<taxonomicName lsidName="J. tiomanensis" pageId="0" pageNumber="1" rank="species" species="tiomanensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. tiomanensis</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="J. counsilmani" pageId="0" pageNumber="1" rank="species" species="counsilmani">
<emphasis italics="true" pageId="0" pageNumber="1">J. counsilmani</emphasis>
</taxonomicName>
, it is distinctly slenderer, more elongate and strongly curved (cf. 
<bibRefCitation author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="167 - 174" refId="B20" refString="Ng, PKL, Tan, LWH, 1984. A new freshwater crab, Stoliczia (Johora) tiomanensis n. sp. (Decapoda: Brachyura: Potamidae) from Pulau Tioman, West Malaysia. Malayan Nature Journal 37 (3): 167 - 174" title="A new freshwater crab, Stoliczia (Johora) tiomanensis n. sp. (Decapoda: Brachyura: Potamidae) from Pulau Tioman, West Malaysia." volume="37" year="1984">Ng and Tan 1984</bibRefCitation>
: fig. 3a-c; 
<bibRefCitation author="Ng, PKL" journalOrPublisher="Zoologische Mededelingen" pageId="0" pageNumber="1" pagination="149 - 162" refId="B8" refString="Ng, PKL, 1985. Freshwater decapod crustaceans from Pulau Tioman, West Malaysia. Zoologische Mededelingen 59 (14): 149 - 162" title="Freshwater decapod crustaceans from Pulau Tioman, West Malaysia." volume="59" year="1985">Ng 1985</bibRefCitation>
: figs 2a, c, e, 4a-c). 
<taxonomicName authorityName="Ng" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora booliati" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">Johora booliati</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
differ from 
<taxonomicName lsidName="J. thoi" pageId="0" pageNumber="1" rank="species" species="thoi">
<emphasis italics="true" pageId="0" pageNumber="1">J. thoi</emphasis>
</taxonomicName>
in possessing slightly shorter ambulatory legs (length to width ratio of merus of fourth leg 2.7-2.8 versus 3.0-3.2), the lateral carapace surfaces are relatively more rugose, and the distinctly shorter and gently curved G1 terminal segment (Figs 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10E-H</figureCitation>
, 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A-D Johora erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak E-H J. erici sp. nov., male (41.2 x 33.8 mm) (ZRC 1995.0269), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure11" httpUri="https://binary.pensoft.net/fig/476322" pageId="0" pageNumber="1">11A-C, E-G</figureCitation>
); in 
<taxonomicName lsidName="J. thoi" pageId="0" pageNumber="1" rank="species" species="thoi">
<emphasis italics="true" pageId="0" pageNumber="1">J. thoi</emphasis>
</taxonomicName>
, this structure is diagnostic, being very elongate and straight (Fig. 
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A-D Johora thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249), Terengganu E-H J. thoi Ng, 1990, male (21.1 x 17.7 mm) (ZRC 2001.1167), Terengganu I-L J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2020.0361), Terengganu A, E, I left G 1 (ventral view) B, F, J distal part of left G 1 (ventral view) C, G, K distal part of left G 1 (dorsal view) D, H, L left G 2 (ventral view) A-D after Ng (1990: fig. 2 A, C-E). Scale bars: 1.0 mm (A, D, E, H, I, L); 0.5 mm (B, C, F, G, J, K)." figureDoi="10.3897/zookeys.994.56810.figure9" httpUri="https://binary.pensoft.net/fig/476320" pageId="0" pageNumber="1">9A-C, E-G</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
The G1 structure of 
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
closely resembles that of 
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
, with the terminal segments similar in shape and proportions. The G1 subterminal segments of the two species, however, differ in form, with that of 
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
forming a shelf-like structure along the outer margin (Fig. 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10E</figureCitation>
) while in 
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
, the tapering of the segment is gradual, and no shelf is visible (Fig. 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A-D Johora erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak E-H J. erici sp. nov., male (41.2 x 33.8 mm) (ZRC 1995.0269), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure11" httpUri="https://binary.pensoft.net/fig/476322" pageId="0" pageNumber="1">11A, E</figureCitation>
). The carapaces of the two species are different in adult males and females. In 
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
, the carapace has a more quadrate form because the anterolateral margin is only gently convex, even in the largest male and the posterolateral margin is straighter and gently converging, with the posterior carapace margin medially indented (even in females) (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2F</figureCitation>
). In 
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
, the carapace appears more ovate, with the anterolateral margin distinctly convex, and the posterolateral margin is medially concave and strongly converging, and the posterior carapace margin is entire (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal view of carapace A Johora intermedia Ng, 1986, holotype male (19.8 x 15.1 mm) (ZRC 1984.6529), Pahang B J. tahanensis (Bott, 1966), male (30.7 x 25.5 mm) (ZRC 1984.6795), Pahang C J. thoi Ng, 1990, holotype male (41.2 x 33.3 mm) (ZRC 1989.2249) Terengganu D J. hoiseni Ng &amp; Takeda, 1992, holotype male (25.1 x 21.1 mm) (ZRC 1984.6673), Kelantan E J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand F J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang G J. erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak H J. michaeli sp. nov., holotype male (22.7 x 19.2 mm) (ZRC 2010.0047), Terengganu." figureDoi="10.3897/zookeys.994.56810.figure2" httpUri="https://binary.pensoft.net/fig/476313" pageId="0" pageNumber="1">2G</figureCitation>
, 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Johora erici sp. nov., male (41.2 x 33.8 mm) (ZRC 1995.0269), Kelantan A overall dorsal view B dorsal view of carapace C frontal view of cephalothorax D anterior thoracic sternum and pleon." figureDoi="10.3897/zookeys.994.56810.figure7" httpUri="https://binary.pensoft.net/fig/476318" pageId="0" pageNumber="1">7B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
The G1 structure of 
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
is superficially similar to that of 
<taxonomicName lsidName="J. murphyi" pageId="0" pageNumber="1" rank="species" species="murphyi">
<emphasis italics="true" pageId="0" pageNumber="1">J. murphyi</emphasis>
</taxonomicName>
from southern Peninsular Malaysia, notably in the shape of the G1 terminal and subterminal segments (
<bibRefCitation author="Ng, PKL" journalOrPublisher="Journal of the Singapore National Academy of Science" pageId="0" pageNumber="1" pagination="36 - 44" refId="B10" refString="Ng, PKL, 1986b. Preliminary descriptions of 17 new freshwater crabs of the genera Geosesarma, Parathelphusa, Johora and Stoliczia (Crustacea Decapoda, Brachyura) from South East Asia. Journal of the Singapore National Academy of Science 15: 36 - 44" title="Preliminary descriptions of 17 new freshwater crabs of the genera Geosesarma, Parathelphusa, Johora and Stoliczia (Crustacea Decapoda, Brachyura) from South East Asia." volume="15" year="1986 b">Ng 1986b</bibRefCitation>
: fig. 14a, b; 
<bibRefCitation author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="13 - 44" refId="B11" refString="Ng, PKL, 1987. A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae). Malayan Nature Journal 41: 13 - 44" title="A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae)." volume="41" year="1987">1987</bibRefCitation>
: fig. 6D, E; 
<bibRefCitation author="Ng, PKL" journalOrPublisher="Department of Zoology, National University of Singapore, Shinglee Press" pageId="0" pageNumber="1" refId="B12" refString="Ng, PKL, 1988. The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press" title="The Freshwater Crabs of Peninsular Malaysia and Singapore." year="1988">1988</bibRefCitation>
: fig. 16D, E), but differs in having the distal part of the terminal segment more rounded and less sharply tapering (Figs 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10E-H</figureCitation>
, 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A-D Johora erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak E-H J. erici sp. nov., male (41.2 x 33.8 mm) (ZRC 1995.0269), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure11" httpUri="https://binary.pensoft.net/fig/476322" pageId="0" pageNumber="1">11A-C, E-G</figureCitation>
). The G1 of 
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
is also similar to 
<taxonomicName lsidName="J. intermedia" pageId="0" pageNumber="1" rank="species" species="intermedia">
<emphasis italics="true" pageId="0" pageNumber="1">J. intermedia</emphasis>
</taxonomicName>
, which has a wide distribution in central Peninsular Malaysia. The G1 terminal segment of 
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
, however, is relatively stouter and less curved, and the subterminal segment narrows along the distal third to form a neck-like structure with the terminal segment (Figs 
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. A-D Johora thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005, paratype male (22.2 x 18.6 mm) (ZRC 2006.0052), Thailand E-I J. booliati sp. nov., holotype male (34.6 x 30.0 mm) (ZRC 2020.0072), Pahang A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, H distal part of left G 1 (dorsal view) G distal part of left G 1 (mesial view) D, I left G 2 (ventral view). Scale bas: 1.0 mm (A, D, E, I); 0.5 mm (B, C, F-H)." figureDoi="10.3897/zookeys.994.56810.figure10" httpUri="https://binary.pensoft.net/fig/476321" pageId="0" pageNumber="1">10E-H</figureCitation>
, 
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A-D Johora erici sp. nov., holotype male (38.3 x 32.0 mm) (ZRC 2020.0360), Perak E-H J. erici sp. nov., male (41.2 x 33.8 mm) (ZRC 1995.0269), Kelantan A, E left G 1 (ventral view) B, F distal part of left G 1 (ventral view) C, G distal part of left G 1 (dorsal view) D, H left G 2 (ventral view). Scale bars: 1.0 mm (A, D, E, H); 0.5 mm (B, C, F, G)." figureDoi="10.3897/zookeys.994.56810.figure11" httpUri="https://binary.pensoft.net/fig/476322" pageId="0" pageNumber="1">11A-C, E-G</figureCitation>
) (versus the G1 terminal segment is more curved and tapers to a slender tip with only the distalmost part of the subterminal segment distinctly narrowed to form a broad cleft in 
<taxonomicName lsidName="J. intermedia" pageId="0" pageNumber="1" rank="species" species="intermedia">
<emphasis italics="true" pageId="0" pageNumber="1">J. intermedia</emphasis>
</taxonomicName>
, cf. 
<bibRefCitation author="Ng, PKL" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="1" pagination="13 - 44" refId="B11" refString="Ng, PKL, 1987. A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae). Malayan Nature Journal 41: 13 - 44" title="A revision of the Malayan freshwater crabs of the genus Johora Bott, 1966 stat. nov. (Decapoda: Brachyura: Potamidae)." volume="41" year="1987">Ng 1987</bibRefCitation>
: figs 3A-D, G-J, 4A-D, F, G, I, J; 
<bibRefCitation author="Ng, PKL" journalOrPublisher="Department of Zoology, National University of Singapore, Shinglee Press" pageId="0" pageNumber="1" refId="B12" refString="Ng, PKL, 1988. The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press" title="The Freshwater Crabs of Peninsular Malaysia and Singapore." year="1988">1988</bibRefCitation>
: fig. 15D-F; 
<bibRefCitation author="Ng, PKL" journalOrPublisher="Bulletin of the National Science Museum, Tokyo, Series A" pageId="0" pageNumber="1" pagination="103 - 116" refId="B19" refString="Ng, PKL, Takeda, M, 1992. On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia. Bulletin of the National Science Museum, Tokyo, Series A 18 (3): 103 - 116" title="On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia." volume="18" year="1992">Ng and Takeda 1992</bibRefCitation>
: fig. 1A-E). It is also noteworthy that both 
<taxonomicName lsidName="J. murphyi" pageId="0" pageNumber="1" rank="species" species="murphyi">
<emphasis italics="true" pageId="0" pageNumber="1">J. murphyi</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="J. intermedia" pageId="0" pageNumber="1" rank="species" species="intermedia">
<emphasis italics="true" pageId="0" pageNumber="1">J. intermedia</emphasis>
</taxonomicName>
are smaller species than 
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="J. erici" pageId="0" pageNumber="1" rank="species" species="erici">
<emphasis italics="true" pageId="0" pageNumber="1">J. erici</emphasis>
</taxonomicName>
, with adults not reaching carapace widths of 25 mm.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Ng, PKL" journalOrPublisher="Bulletin of the National Science Museum, Tokyo, Series A" pageId="0" pageNumber="1" pagination="103 - 116" refId="B19" refString="Ng, PKL, Takeda, M, 1992. On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia. Bulletin of the National Science Museum, Tokyo, Series A 18 (3): 103 - 116" title="On some freshwater crabs (Crustacea, Brachyura, Potamidae, Parathelphusidae and Grapsidae) from Peninsular Malaysia." volume="18" year="1992">Ng and Takeda (1992</bibRefCitation>
: 107) had specimens from 
<normalizedToken originalValue="Fraser’s">Fraser's</normalizedToken>
Hill as well as adjacent areas (Sungei Gumut, Peretak, Selangor, 
<geoCoordinate degrees="3" direction="north" minutes="36" orientation="latitude" precision="1" seconds="53.3" value="3.6148055">3°36'53.3&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="101" direction="east" minutes="44" orientation="longitude" precision="1" seconds="40.4" value="101.74455">101°44'40.4&quot;E</geoCoordinate>
; Sungei Sum [probably Sungei Sum Sum], near Genting Highlands, Pahang, 
<geoCoordinate degrees="3" direction="north" minutes="20" orientation="latitude" precision="1" seconds="42.2" value="3.3450556">3°20'42.2&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="101" direction="east" minutes="51" orientation="longitude" precision="1" seconds="12.0" value="101.85333">101°51'12.0&quot;E</geoCoordinate>
; Ulu Teranum, Teras, Pahang, 
<geoCoordinate degrees="3" direction="north" minutes="44" orientation="latitude" precision="1" seconds="12.6" value="3.7368333">3°44'12.6&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="101" direction="east" minutes="47" orientation="longitude" precision="1" seconds="29.5" value="101.79153">101°47'29.5&quot;E</geoCoordinate>
) which they identified as &quot; 
<taxonomicName lsidName="J. tahanensis" pageId="0" pageNumber="1" rank="species" species="tahanensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. tahanensis</emphasis>
</taxonomicName>
&quot;. On the basis of geography, they are probably all 
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
as defined here.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Ng" authorityYear="2020" class="Malacostraca" family="Potamidae" genus="Johora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Johora booliati" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">Johora booliati</emphasis>
</taxonomicName>
sp. nov. is known from highland streams in the central highlands of Pahang (Genting Highlands, Bukit Tinggi and 
<normalizedToken originalValue="Fraser’s">Fraser's</normalizedToken>
Hill) (Fig. 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Distribution of species of the Johora tahanensis species group 1 J. tahanensis (Bott, 1966) 2 J. thoi Ng, 1990 3 J. hoiseni Ng &amp; Takeda, 1992 4 J. thaiana Leelawathanagoon, Lheknim &amp; Ng, 2005 5 J. booliati sp. nov. 6 J. erici sp. nov. 7 J. michaeli sp. nov. Map data 2020 Google." figureDoi="10.3897/zookeys.994.56810.figure15" httpUri="https://binary.pensoft.net/fig/476326" pageId="0" pageNumber="1">15</figureCitation>
). The distribution of 
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
overlaps with those of 
<taxonomicName lsidName="J. intermedia" pageId="0" pageNumber="1" rank="species" species="intermedia">
<emphasis italics="true" pageId="0" pageNumber="1">J. intermedia</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="J. gapensis" pageId="0" pageNumber="1" rank="species" species="gapensis">
<emphasis italics="true" pageId="0" pageNumber="1">J. gapensis</emphasis>
</taxonomicName>
, and we can expect the taxa to be found together. One juvenile male of 
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
(10.5 
<normalizedToken originalValue="×">x</normalizedToken>
9.1 mm, ZRC 2020.0364) was in fact collected with two adult males of 
<taxonomicName lsidName="J. intermedia" pageId="0" pageNumber="1" rank="species" species="intermedia">
<emphasis italics="true" pageId="0" pageNumber="1">J. intermedia</emphasis>
</taxonomicName>
at 
<normalizedToken originalValue="Fraser’s">Fraser's</normalizedToken>
Hill (ZRC 2016.003).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="conservation">
<paragraph pageId="0" pageNumber="1">Conservation.</paragraph>
<paragraph pageId="0" pageNumber="1">
The conservation status for 
<taxonomicName lsidName="J. booliati" pageId="0" pageNumber="1" rank="species" species="booliati">
<emphasis italics="true" pageId="0" pageNumber="1">J. booliati</emphasis>
</taxonomicName>
is not known as its actual distribution is not known. For the moment, it is known only from a 30 km stretch of hills along the Central Highlands of Peninsular Malaysia, in the area of 
<normalizedToken originalValue="Fraser’s">Fraser's</normalizedToken>
Hill, Genting Highlands and Bukit Tinggi. This area is not protected, and in this context, the species should be categorised as vulnerable for the time being (see 
<bibRefCitation DOI="https://doi.org/10.1016/j.biocon.2009.02.038" author="Cumberlidge, N" journalOrPublisher="Biological Conservation" pageId="0" pageNumber="1" pagination="1665 - 1673" refId="B3" refString="Cumberlidge, N, Ng, PKL, Yeo, DCJ, Magalhaes, C, Campos, MR, Alvarez, F, Naruse, T, Daniels, SR, Esser, LJ, Attipoe, FYK, Clotilde-Ba, F-L, Darwall, W, Mcivor, A, Collen, B, Ram, M, 2009. Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges. Biological Conservation 142: 1665 - 1673, DOI: https://doi.org/10.1016/j.biocon.2009.02.038" title="Freshwater crabs and the biodiversity crisis: importance, threats, status, and conservation challenges." url="https://doi.org/10.1016/j.biocon.2009.02.038" volume="142" year="2009">Cumberlidge et al. 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>