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<mods:title id="BB972D799D43BD703D1E800D8E49998E">A collection of crabs (Crustacea, Brachyura) from the southwestern coast of India, with a discussion of the systematic position of Nectopanope Wood-Mason in Wood-Mason &amp; Alcock, 1891 (Euryplacidae)</mods:title>
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<mods:namePart id="30E3B56805472BFBB2B8AF8F6A98C153">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="1861238962ECA5B6E29F5E1960BB6CF4">Priyaja, P.</mods:namePart>
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<mods:namePart id="F3AECA4AF23083AB38EA5E1041870F10">Kumar, A. Biju</mods:namePart>
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<treatment id="1BE10941E4EDBE63E400EDBC0E5F9137" ID-GBIF-Taxon="154126660" LSID="urn:lsid:plazi:treatment:1BE10941E4EDBE63E400EDBC0E5F9137" httpUri="http://treatment.plazi.org/id/1BE10941E4EDBE63E400EDBC0E5F9137" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
<subSubSection id="7DC65CB5CAB7894C6107CA15726C030A" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="CA5E0707304A9F67371635E4F5C3B537" pageId="8" pageNumber="9">
<taxonomicName id="A4D6BC904BF3FB8D64B2D56AA08B9138" ID-CoL="6LPVN" authority="Rathbun, 1902" authorityName="Rathbun" authorityYear="1902" class="Malacostraca" family="Euryplacidae" genus="Henicoplax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Henicoplax maldivensis" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="maldivensis">Henicoplax maldivensis (Rathbun, 1902)</taxonomicName>
Figs 5, 7
<normalizedToken id="FAC122870B195246E5B4B9C4030AD93C" originalValue="F–H">F-H</normalizedToken>
</paragraph>
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<subSubSection id="AC8E14593CEE07D6DF2C23978E714953" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="2B687E19FF3131D51C606EDA94A8EA34" pageId="8" pageNumber="9">
<taxonomicName id="0FD00DE9A79AD6B872DC76FAF2081D09" class="Malacostraca" family="Euryplacidae" genus="Goneplax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Goneplax maldivensis" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="maldivensis">Goneplax maldivensis</taxonomicName>
Rathbun, 1902: 124, figs 3-5;
<bibRefCitation id="2AC150DAF1D071EFA0282D982FC15ADF" pageId="8" pageNumber="9">Guinot 1969</bibRefCitation>
: 518;
<bibRefCitation id="53CC8AEB2783B6F031EF2B8E8E5538C1" author="Castro, P" journalOrPublisher="Zoosystema" pageId="14" pageNumber="15" pagination="609 - 773" title="A reappraisal of the family Goneplacidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura) and revision of the subfamily Goneplacinae, with the description of ten new genera and eighteen new species." volume="29" year="2007">Castro 2007</bibRefCitation>
: 686, 687 [list].
</paragraph>
<paragraph id="190898E2A705695E49517CBCCEBD8C0F" pageId="8" pageNumber="9">
&quot;?[
<taxonomicName id="64D8ADAECE738CD3F0E84147EBDFB449" class="Malacostraca" family="Euryplacidae" genus="Goneplax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Goneplax" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Goneplax</taxonomicName>
]
<taxonomicName id="13643962AA85E02FA5A29EF3F5C86518" family="Euryplacidae" genus="Pilumnoidea" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pilumnoidea maldivensis" order="Decapoda" pageId="8" pageNumber="9" rank="species" species="maldivensis">maldivensis</taxonomicName>
&quot;:
<bibRefCitation id="489C27E41EE7003ADD083B894781A514" pageId="8" pageNumber="9">Guinot 1971</bibRefCitation>
: 1081 [list].
</paragraph>
<paragraph id="099BAA2982FF3ECCE3F5D09647B00430" pageId="8" pageNumber="9">
<taxonomicName id="52DB28FAC0514997654CFEF663236AE5" class="Malacostraca" family="Euryplacidae" genus="Otmaroplax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Otmaroplax maldivensis" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="maldivensis">Otmaroplax maldivensis</taxonomicName>
:
<normalizedToken id="F8B6DE6638B94CAF36CFDBFCDE835DE5" originalValue="Števčić">Stevcic</normalizedToken>
2005: 134 [genus name nomen nudum]
</paragraph>
<paragraph id="FA89D47CA0E7A8A0A8D46DC27784C06E" pageId="8" pageNumber="9">
&quot;
<taxonomicName id="B1364FEBD8FBC919648F10B84394E710" class="Malacostraca" family="Euryplacidae" genus="Heteroplax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heteroplax" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Heteroplax</taxonomicName>
&quot;
<taxonomicName id="1EB23EFC0F3837E40956621038075D02" family="Euryplacidae" genus="Pilumnoidea" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pilumnoidea maldivensis" order="Decapoda" pageId="8" pageNumber="9" rank="species" species="maldivensis">maldivensis</taxonomicName>
:
<bibRefCitation id="F076ED32113959B043909AE84AF118D9" author="Ng, PKL" journalOrPublisher="Raffles Bulletin of Zoology, Supplement" pageId="14" pageNumber="15" pagination="1 - 286" title="Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world." volume="17" year="2008">Ng et al. 2008</bibRefCitation>
: 78, 79 [in list].
</paragraph>
<paragraph id="9985DE61231385A183D95D426EEC1C33" pageId="8" pageNumber="9">
<taxonomicName id="0717C26A6B9153AD2DE1064409A9A630" class="Malacostraca" family="Euryplacidae" genus="Henicoplax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Henicoplax maldivensis" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="maldivensis">Henicoplax maldivensis</taxonomicName>
:
<bibRefCitation id="94E75D196B7CFA92FBEEDA464BFA757A" author="Castro, P" journalOrPublisher="Zootaxa" pageId="14" pageNumber="15" pagination="1 - 130" title="Revision of the family Euryplacidae Stimpson, 1871 (Crustacea: Decapoda: Brachyura: Goneplacoidea)." url="https://doi.org/10.11646/zootaxa.2375.1.1" volume="2375" year="2010">Castro and Ng 2010</bibRefCitation>
: 61, figs 22
<normalizedToken id="18A1FDE0B1DB75323B298F609C699FE3" originalValue="A–E">A-E</normalizedToken>
, 24
<normalizedToken id="9FB9F49ABC4AEBB7600A25DB63EBE2F0" originalValue="D–F">D-F</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection id="B88D8F8FE68BC5A96EEBFDC8A24474E2" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="6177F75076EEBD45342A5E801C0F812C" pageId="8" pageNumber="9">Material examined.</paragraph>
<paragraph id="BA2800253BF5B8CB218778A4389C5B5E" pageId="8" pageNumber="9">
1 male (9.9
<normalizedToken id="3B249DE0991097478A4A488AC6190CFD" originalValue="×">x</normalizedToken>
6.3 mm),
<geoCoordinate id="FFA55C7EB15A5FCDF2E08E2EBA0B002E" direction="north" orientation="latitude" precision="1" value="7.4663">7°27.978'N</geoCoordinate>
,
<geoCoordinate id="E01DD342F9E07752F70EC94451E47FC1" direction="east" orientation="longitude" precision="1" value="77.538284">77°32.297'E</geoCoordinate>
, 100 m.
</paragraph>
</subSubSection>
<subSubSection id="04AB0D36EE0A9CCA3F03680DA2F1E880" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="remarks">
<paragraph id="49656C0CCD4893FCD3546C06689AC15D" pageId="8" pageNumber="9">Remarks.</paragraph>
<paragraph id="C1D553F7754B2EC411A89DA11E2CD67B" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<taxonomicName id="12B1FE26CA2CFD2AD6D7E5C0166C063D" class="Malacostraca" family="Euryplacidae" genus="Henicoplax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Henicoplax" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Henicoplax</taxonomicName>
Castro &amp; Ng, 2010, was established for Indo-West Pacific species that had been previously classified in
<taxonomicName id="DB9604D078DBFFB5669471CCB658576B" class="Malacostraca" family="Goneplacidae" genus="Goneplax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Goneplax" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Goneplax</taxonomicName>
Leach, 1814, or
<taxonomicName id="199373FABEE46089BE6A9173183F2011" class="Malacostraca" family="Euryplacidae" genus="Heteroplax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heteroplax" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Heteroplax</taxonomicName>
<pageBreakToken id="F76DDE8B0A28794273F88F6737D274BC" pageId="9" pageNumber="10" start="start">Stimpson</pageBreakToken>
, 1858. Five species are currently recognised:
<taxonomicName id="017C4DA92CDFB22ECC33BF4B17FFC1F6" lsidName="H. eriochir" pageId="9" pageNumber="10" rank="species" species="eriochir">H. eriochir</taxonomicName>
Castro &amp; Ng, 2010,
<taxonomicName id="18BB8DE55EC6CC70658502EFA4ABE8FE" lsidName="H. maldivensis" pageId="9" pageNumber="10" rank="species" species="maldivensis">H. maldivensis</taxonomicName>
(Rathbun, 1902) [type species],
<taxonomicName id="CFFD3F00BF115139233B0F0DD76732C3" lsidName="H. nitida" pageId="9" pageNumber="10" rank="species" species="nitida">H. nitida</taxonomicName>
(Miers, 1879a),
<taxonomicName id="277AAA1B5B128252E2AB80BAA13A075C" lsidName="H. pilimeles" pageId="9" pageNumber="10" rank="species" species="pilimeles">H. pilimeles</taxonomicName>
Castro &amp; Ng, 2010, and
<taxonomicName id="1D7ACADF287903486F68B848092C2CCF" lsidName="H. trachydactylus" pageId="9" pageNumber="10" rank="species" species="trachydactylus">H. trachydactylus</taxonomicName>
Castro &amp; Ng, 2010.
</paragraph>
<paragraph id="7B5FAF458DA1A5BA57F87EC2C1A3CD62" pageId="10" pageNumber="11">
<pageBreakToken id="20C846B17D3FC28B711F9EF97F120D0B" pageId="10" pageNumber="11" start="start">The</pageBreakToken>
present specimen is clearly
<taxonomicName id="59FC1E83473C4CEC7ABF6D30ACF6A242" lsidName="H. maldivensis" pageId="10" pageNumber="11" rank="species" species="maldivensis">H. maldivensis</taxonomicName>
as redescribed and figured at length by
<bibRefCitation id="F4B080CE6E91755D5BEB25F477DCFA69" author="Castro, P" journalOrPublisher="Zootaxa" pageId="14" pageNumber="15" pagination="1 - 130" title="Revision of the family Euryplacidae Stimpson, 1871 (Crustacea: Decapoda: Brachyura: Goneplacoidea)." url="https://doi.org/10.11646/zootaxa.2375.1.1" volume="2375" year="2010">Castro and Ng (2010</bibRefCitation>
: 61). The species was previously known only from the holotype male (7.4
<normalizedToken id="A3B68CED1D6643CA806A7EC9564D58D7" originalValue="×">x</normalizedToken>
4.8 mm) obtained from Gan Island in Addu Atoll in the Maldives. The present male differs from the type male in having the frontal margin slightly more sinuous and the cleft between the external orbital tooth and the anterolateral tooth is more U-shaped (Fig. 5A, B) (versus frontal margin almost straight and the lateral carapace cleft being V-shaped in the holotype; cf.
<bibRefCitation id="079AE27C8907E68A8703397EC1C1C07E" author="Castro, P" journalOrPublisher="Zootaxa" pageId="14" pageNumber="15" pagination="1 - 130" title="Revision of the family Euryplacidae Stimpson, 1871 (Crustacea: Decapoda: Brachyura: Goneplacoidea)." url="https://doi.org/10.11646/zootaxa.2375.1.1" volume="2375" year="2010">Castro and Ng 2010</bibRefCitation>
: fig. 22A); and while the G1 shape is similar, the distal spination is relatively less pronounced and the tip is sharply tapering (Fig. 7F, G) (versus distal half with relatively more spines and the tip is subtruncate in the holotype; cf.
<bibRefCitation id="8B3D543188D6FCAE4980AC3D3E066288" author="Castro, P" journalOrPublisher="Zootaxa" pageId="14" pageNumber="15" pagination="1 - 130" title="Revision of the family Euryplacidae Stimpson, 1871 (Crustacea: Decapoda: Brachyura: Goneplacoidea)." url="https://doi.org/10.11646/zootaxa.2375.1.1" volume="2375" year="2010">Castro and Ng 2010</bibRefCitation>
: fig. 24D, E). The differences are not substantial and can easily be explained by variation and size, the present male (9.9
<normalizedToken id="208BF3C33534B1375817B0EA22BED88C" originalValue="×">x</normalizedToken>
6.3 mm) being larger than the type (7.4
<normalizedToken id="63EE255DF025C3D7E57A2F24C9267491" originalValue="×">x</normalizedToken>
4.8 mm).
</paragraph>
<caption id="74A489C4FD64EE1A6F100D39A7931518" pageId="10" pageNumber="11">
<paragraph id="7CA0AD4FAC9E4ABF6A37099B6931DA46" pageId="10" pageNumber="11">
Figure 5.
<taxonomicName id="5711B2B84068B3D50A0410F9FE91942F" class="Malacostraca" family="Euryplacidae" genus="Henicoplax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Henicoplax maldivensis" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="maldivensis">Henicoplax maldivensis</taxonomicName>
(Rathbun, 1902), male (9.9
<normalizedToken id="A96337933E218C91EC358CBE434F3F6A" originalValue="×">x</normalizedToken>
6.3 mm), India. A overall dorsal habitus B dorsal view of carapace C frontal view of cephalothorax D left third maxilliped E anterior thoracic sternum and pleon F thoracic sternum and pleon G posterior thoracic sternum and pleon H outer view of chelae.
</paragraph>
</caption>
<paragraph id="A2066414A73761C1C5F682E282635F52" pageId="10" pageNumber="11">
<bibRefCitation id="541816E430833A5919A023D267CD0314" author="Castro, P" journalOrPublisher="Zootaxa" pageId="14" pageNumber="15" pagination="1 - 130" title="Revision of the family Euryplacidae Stimpson, 1871 (Crustacea: Decapoda: Brachyura: Goneplacoidea)." url="https://doi.org/10.11646/zootaxa.2375.1.1" volume="2375" year="2010">Castro and Ng (2010)</bibRefCitation>
showed that records of &quot;
<taxonomicName id="37D265AD3FC3FCEF1C29EB125D6952F8" lsidName="H. nitida" pageId="10" pageNumber="11" rank="species" species="nitida">H. nitida</taxonomicName>
&quot; from the Andaman Sea should be referred to a new species,
<taxonomicName id="E4B32F46A56CA17DC98A1D9A5ED6DC11" lsidName="H. pilimenes" pageId="10" pageNumber="11" rank="species" species="pilimenes">H. pilimenes</taxonomicName>
; and indicated that true
<taxonomicName id="7A5DD05BD8FCE69C5CABC669C37ABBE1" lsidName="H. nitida" pageId="10" pageNumber="11" rank="species" species="nitida">H. nitida</taxonomicName>
should be restricted to East Asia. The records of &quot;
<taxonomicName id="CBD05212AA6BC655DE3C2412B026DB60" lsidName="H. nitida" pageId="10" pageNumber="11" rank="species" species="nitida">H. nitida</taxonomicName>
&quot; from the Gulf of Martaban (south of Myanmar) and off Madras in India by
<bibRefCitation id="509C8FA258673A170B3D66F6E35FA1CC" author="Henderson, JR" journalOrPublisher="Zoologische Mededelingen" pageId="14" pageNumber="15" url="https://doi.org/10.1111/j.1096-3642.1893.tb00653.x" year="1893">Henderson (1893</bibRefCitation>
: 379) are almost certainly not this species. The Myanmar material is probably
<taxonomicName id="8768E4BEBD98382E5AA690D08438B275" lsidName="H. pilimenes" pageId="10" pageNumber="11" rank="species" species="pilimenes">H. pilimenes</taxonomicName>
, while that from India may be this species or even
<taxonomicName id="6569CCA7A5790E7D00F5416E3D787D84" lsidName="H. maldivensis" pageId="10" pageNumber="11" rank="species" species="maldivensis">H. maldivensis</taxonomicName>
instead. Specimens will need to be re-examined to be certain.
</paragraph>
<paragraph id="0A05FB2181F9179818800D8F04B70CC4" pageId="10" pageNumber="11">
<taxonomicName id="98D13AEF0074B2FB8FFA1EDC54DFB9C5" class="Malacostraca" family="Euryplacidae" genus="Heteroplax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heteroplax maldivensis" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="maldivensis">Heteroplax maldivensis</taxonomicName>
is thus far known for certain only from the Maldives (
<bibRefCitation id="710396F2F6D4E7991737E32B151DBD10" author="Rathbun, MJ" journalOrPublisher="Bulletin of the Museum of Comparative Zoology, Harvard College" pageId="14" pageNumber="15" pagination="123 - 138" title="Crabs from the Maldive Islands." volume="39" year="1902">Rathbun 1902</bibRefCitation>
;
<bibRefCitation id="184CD9C98D6153489F3EC3B2A151E73E" author="Castro, P" journalOrPublisher="Zootaxa" pageId="14" pageNumber="15" pagination="1 - 130" title="Revision of the family Euryplacidae Stimpson, 1871 (Crustacea: Decapoda: Brachyura: Goneplacoidea)." url="https://doi.org/10.11646/zootaxa.2375.1.1" volume="2375" year="2010">Castro and Ng 2010</bibRefCitation>
) and the present specimen represents the first confirmed presence of this species in India.
</paragraph>
</subSubSection>
</treatment>
</document>
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