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<document id="D0C7FF2B691935FF4113E332FF133202" ID-DOI="10.11646/zootaxa.3846.1.1" ID-GBIF-Dataset="01a9d979-7b10-4c36-b1af-4acc2c0cb9a7" ID-ISSN="1175-5326" ID-Zenodo-Dep="286792" ID-ZooBank="97D4A04A-D75E-45CC-8A70-3EB3A4E94D9B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460464243854" checkinUser="plazi" docAuthor="Zwick, Peter" docDate="2014" docId="113287AD9B52FFD7FF13FBEBFE3FF828" docLanguage="en" docName="zt03846p041.pdf" docOrigin="Zootaxa 3846 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calvarium (Calvariellum) bellendenker Zwick, 2014, n. sp." docType="treatment" docVersion="6" lastPageNumber="12" masterDocId="ED0BFFD59B59FFDCFF84FFF7FFB5FFF3" masterDocTitle="Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer" masterLastPageNumber="41" masterPageNumber="1" pageNumber="12" updateTime="1698304595582" updateUser="plazi">
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<mods:title id="2AC30A54B7D711918487CD709A5EB5D1">Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer</mods:title>
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<mods:name id="F1DBBA6BF25776AA42413908C0FF0371" type="personal">
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<mods:namePart id="DFB493B26147664341EF2CB4DF9AC09B">Zwick, Peter</mods:namePart>
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<mods:date id="7110C9C3ADE5CB49EE02DE3E37A9FA6F">2014</mods:date>
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<mods:number id="709CE57FB717C5C671796A7B17D59626">3846</mods:number>
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<treatment id="113287AD9B52FFD7FF13FBEBFE3FF828" ID-DOI="http://doi.org/10.5281/zenodo.6126813" ID-GBIF-Taxon="127669616" ID-Zenodo-Dep="6126813" LSID="urn:lsid:plazi:treatment:113287AD9B52FFD7FF13FBEBFE3FF828" httpUri="http://treatment.plazi.org/id/113287AD9B52FFD7FF13FBEBFE3FF828" lastPageNumber="12" pageId="11" pageNumber="12">
<subSubSection id="D18165309B52FFD7FF13FBEBFE9AFBA4" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="992436BB9B52FFD7FF13FBEBFD77FBC5" blockId="11.[151,706,1052,1111]" box="[151,706,1052,1078]" pageId="11" pageNumber="12">
<heading id="C26C81D79B52FFD7FF13FBEBFD77FBC5" bold="true" box="[151,706,1052,1078]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="ABEFEAA99B52FFD7FF13FBEBFD77FBC5" bold="true" box="[151,706,1052,1078]" pageId="11" pageNumber="12">
<taxonomicName id="5E9B4D389B52FFD7FF13FBEBFDC5FBC5" box="[151,624,1052,1078]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bellendenker" status="sp. nov." subGenus="Calvariellum">
<emphasis id="ABEFEAA99B52FFD7FF13FBEBFDC5FBC5" bold="true" box="[151,624,1052,1078]" italics="true" pageId="11" pageNumber="12">Calvarium (Calvariellum) bellendenker</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="B0DC57D29B52FFD7FD04FBEBFD77FBC5" box="[640,706,1052,1078]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="992436BB9B52FFD7FF13FBC9FE9AFBA4" blockId="11.[151,706,1052,1111]" box="[151,303,1086,1111]" pageId="11" pageNumber="12">
(
<figureCitation id="01A02A3E9B52FFD7FF1BFBC9FE92FBA4" box="[159,295,1086,1111]" captionStart="FIGURES 29 37" captionStartId="10.[151,264,1707,1730]" captionTargetBox="[246,1345,203,1671]" captionTargetId="figure@10.[228,1358,197,1686]" captionTargetPageId="10" captionText="FIGURES 29 37. Calvarium (Calvariellum) spp., males. C. cochlearifer, n. sp.: 29, T 8 and S 8, superimposed; 30, penis; 31, T 9, S 9 and tegmen (tn) with parameres (p) superimposed. C. bellendenker, n. sp.: 32, T 8; 33, S 8; 34, T 9; 35, S 9; 36, penis; 37, tegmen and parameres. Figures for a given species to the same scale." httpUri="https://zenodo.org/record/286799/files/figure.png" pageId="11" pageNumber="12">
Figs. 32
<emphasis id="ABEFEAA99B52FFD7FF7EFBC9FEBFFBA4" bold="true" box="[250,266,1086,1111]" pageId="11" pageNumber="12"></emphasis>
37
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D18165309B52FFD7FF13FB71FEC5FAA0" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="992436BB9B52FFD7FF13FB71FCBAFB30" blockId="11.[151,1437,1158,2011]" pageId="11" pageNumber="12">
<emphasis id="ABEFEAA99B52FFD7FF13FB71FE71FB6C" bold="true" box="[151,452,1158,1183]" pageId="11" pageNumber="12">
<typeStatus id="462088199B52FFD7FF13FB71FF67FB6C" box="[151,210,1158,1183]" pageId="11" pageNumber="12">Type</typeStatus>
material.
<typeStatus id="462088199B52FFD7FECAFB71FE0AFB6C" box="[334,447,1158,1183]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="8F9DFD329B52FFD7FE49FB70FE45FB6C" box="[461,496,1159,1183]" pageId="11" pageNumber="12" type="generic">1♂</specimenCount>
<emphasis id="ABEFEAA99B52FFD7FE7CFB71FDDBFB6C" bold="true" box="[504,622,1158,1183]" pageId="11" pageNumber="12">
<typeStatus id="462088199B52FFD7FE7CFB71FDDBFB6C" box="[504,622,1158,1183]" pageId="11" pageNumber="12" type="paratype">paratypes</typeStatus>
</emphasis>
<specimenCount id="8F9DFD329B52FFD7FDF3FB70FD2BFB6C" box="[631,670,1159,1183]" pageId="11" pageNumber="12" type="generic">3♂</specimenCount>
: “Russell R. at Bellenden Ker Landing [
<geoCoordinate id="FCAF507C9B52FFD7FBEDFB71FB6DFB6C" box="[1129,1240,1158,1183]" direction="south" orientation="latitude" pageId="11" pageNumber="12" precision="5" value="-17.2726">17.2726S</geoCoordinate>
,
<geoCoordinate id="FCAF507C9B52FFD7FB62FB71FAD6FB6C" box="[1254,1379,1158,1183]" direction="east" orientation="longitude" pageId="11" pageNumber="12" precision="5" value="145.9485">145.9485E</geoCoordinate>
]
<quantity id="5E639B5E9B52FFD7FAF5FB71FA22FB6C" box="[1393,1431,1158,1183]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="12" unit="m" value="5.0">5m</quantity>
,
<date id="ED25107B9B52FFD7FF13FB5CFEC0FB30" box="[151,373,1194,1219]" pageId="11" pageNumber="12" value="1981-10-24" valueMax="1981-11-09" valueMin="1981-10-24">
24 Oct
<emphasis id="ABEFEAA99B52FFD7FF66FB5DFF47FB30" bold="true" box="[226,242,1194,1219]" pageId="11" pageNumber="12"></emphasis>
9 Nov.1981
</date>
Earthwatch\ Qld.Museum” (
<collectionCode id="FF8AAE7E9B52FFD7FD3CFB5CFCB6FB30" LSID="urn:lsid:biocol.org:col:32981" box="[696,771,1195,1219]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="11" pageNumber="12">ANIC</collectionCode>
).
</paragraph>
<paragraph id="992436BB9B52FFD7FF43FB39FE14FAF8" blockId="11.[151,1437,1158,2011]" pageId="11" pageNumber="12">
<emphasis id="ABEFEAA99B52FFD7FF43FB39FDF4FB14" bold="true" box="[199,577,1230,1255]" pageId="11" pageNumber="12">Additional material studied. 1♀</emphasis>
(presumed) with the same data as the
<typeStatus id="462088199B52FFD7FC63FB38FB90FB14" box="[999,1061,1231,1255]" pageId="11" pageNumber="12">types</typeStatus>
, plus “
<collectionCode id="FF8AAE7E9B52FFD7FBF4FB38FB02FB14" LSID="urn:lsid:biocol.org:col:32981" box="[1136,1207,1231,1255]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="11" pageNumber="12">ANIC</collectionCode>
<taxonomicName id="5E9B4D389B52FFD7FB3AFB39FF75FAF9" authority="Voucher No." authorityName="Voucher No." class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="order">Coleoptera Voucher No.</taxonomicName>
83-0582” (
<collectionCode id="FF8AAE7E9B52FFD7FEC4FB04FE20FAF8" box="[320,405,1267,1291]" pageId="11" pageNumber="12">QMSB</collectionCode>
).
</paragraph>
<paragraph id="992436BB9B52FFD7FF43FAE1FEC5FAA0" blockId="11.[151,1437,1158,2011]" pageId="11" pageNumber="12">
<emphasis id="ABEFEAA99B52FFD7FF43FAE1FE93FADC" bold="true" box="[199,294,1302,1327]" pageId="11" pageNumber="12">Habitus</emphasis>
. Externally indistiguishable from the congeners.
<collectionCode id="FF8AAE7E9B52FFD7FCFDFAE0FC28FADD" box="[889,925,1303,1326]" pageId="11" pageNumber="12">BL</collectionCode>
<quantity id="5E639B5E9B52FFD7FC2FFAE0FB8BFADD" box="[939,1086,1302,1327]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="11" pageNumber="12" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">
1.9
<emphasis id="ABEFEAA99B52FFD7FC4AFAE1FC6BFADC" bold="true" box="[974,990,1302,1327]" pageId="11" pageNumber="12"></emphasis>
2.1 mm
</quantity>
. Uniformly chocolate brown, appendages lighter.
</paragraph>
</subSubSection>
<subSubSection id="D18165309B52FFD7FF43FAA9FB63F861" pageId="11" pageNumber="12" type="description">
<paragraph id="992436BB9B52FFD7FF43FAA9FB8BFA4C" blockId="11.[151,1437,1158,2011]" pageId="11" pageNumber="12">
<emphasis id="ABEFEAA99B52FFD7FF43FAA9FEB1FA84" bold="true" box="[199,260,1374,1399]" pageId="11" pageNumber="12">Male</emphasis>
. T8 short, strongly transverse, apodemes not continuing onto plate. Only sides of caudal margin well defined and sclerotized, middle membranous with numerous microtrichia in closely packed groups of 4 (3
<emphasis id="ABEFEAA99B52FFD7FABCFA75FAFDFA68" bold="true" box="[1336,1352,1410,1435]" pageId="11" pageNumber="12"></emphasis>
5) (
<figureCitation id="01A02A3E9B52FFD7FAEAFA75FF02FA4C" captionStart="FIGURES 29 37" captionStartId="10.[151,264,1707,1730]" captionTargetBox="[246,1345,203,1671]" captionTargetId="figure@10.[228,1358,197,1686]" captionTargetPageId="10" captionText="FIGURES 29 37. Calvarium (Calvariellum) spp., males. C. cochlearifer, n. sp.: 29, T 8 and S 8, superimposed; 30, penis; 31, T 9, S 9 and tegmen (tn) with parameres (p) superimposed. C. bellendenker, n. sp.: 32, T 8; 33, S 8; 34, T 9; 35, S 9; 36, penis; 37, tegmen and parameres. Figures for a given species to the same scale." httpUri="https://zenodo.org/record/286799/files/figure.png" pageId="11" pageNumber="12">Fig. 32</figureCitation>
). S8 almost unpigmented and difficult to see, approximately triangular (
<figureCitation id="01A02A3E9B52FFD7FC59FA51FB84FA4C" box="[989,1073,1446,1471]" captionStart="FIGURES 29 37" captionStartId="10.[151,264,1707,1730]" captionTargetBox="[246,1345,203,1671]" captionTargetId="figure@10.[228,1358,197,1686]" captionTargetPageId="10" captionText="FIGURES 29 37. Calvarium (Calvariellum) spp., males. C. cochlearifer, n. sp.: 29, T 8 and S 8, superimposed; 30, penis; 31, T 9, S 9 and tegmen (tn) with parameres (p) superimposed. C. bellendenker, n. sp.: 32, T 8; 33, S 8; 34, T 9; 35, S 9; 36, penis; 37, tegmen and parameres. Figures for a given species to the same scale." httpUri="https://zenodo.org/record/286799/files/figure.png" pageId="11" pageNumber="12">Fig. 33</figureCitation>
).
</paragraph>
<paragraph id="992436BB9B52FFD7FF13FA3CFE5FF9D8" blockId="11.[151,1437,1158,2011]" pageId="11" pageNumber="12">
T9 without plate, the lateral struts loosely connected by a transverse barely sclerotized crescent, apices curved mediad, with delicate apical microtrichia (
<figureCitation id="01A02A3E9B52FFD7FD0DFA19FD56F9F4" box="[649,739,1518,1543]" captionStart="FIGURES 29 37" captionStartId="10.[151,264,1707,1730]" captionTargetBox="[246,1345,203,1671]" captionTargetId="figure@10.[228,1358,197,1686]" captionTargetPageId="10" captionText="FIGURES 29 37. Calvarium (Calvariellum) spp., males. C. cochlearifer, n. sp.: 29, T 8 and S 8, superimposed; 30, penis; 31, T 9, S 9 and tegmen (tn) with parameres (p) superimposed. C. bellendenker, n. sp.: 32, T 8; 33, S 8; 34, T 9; 35, S 9; 36, penis; 37, tegmen and parameres. Figures for a given species to the same scale." httpUri="https://zenodo.org/record/286799/files/figure.png" pageId="11" pageNumber="12">Fig. 34</figureCitation>
). S9 oval, with narrow, handle-like base. Colourless, distal portion finely pilose (
<figureCitation id="01A02A3E9B52FFD7FE0DF9E5FE68F9D8" box="[393,477,1554,1579]" captionStart="FIGURES 29 37" captionStartId="10.[151,264,1707,1730]" captionTargetBox="[246,1345,203,1671]" captionTargetId="figure@10.[228,1358,197,1686]" captionTargetPageId="10" captionText="FIGURES 29 37. Calvarium (Calvariellum) spp., males. C. cochlearifer, n. sp.: 29, T 8 and S 8, superimposed; 30, penis; 31, T 9, S 9 and tegmen (tn) with parameres (p) superimposed. C. bellendenker, n. sp.: 32, T 8; 33, S 8; 34, T 9; 35, S 9; 36, penis; 37, tegmen and parameres. Figures for a given species to the same scale." httpUri="https://zenodo.org/record/286799/files/figure.png" pageId="11" pageNumber="12">Fig. 35</figureCitation>
).
</paragraph>
<paragraph id="992436BB9B52FFD7FF43F9C1FE46F948" blockId="11.[151,1437,1158,2011]" pageId="11" pageNumber="12">
Penis with wide triangularly excised pala with sharp anterolateral corners.
<collectingCountry id="E18C762B9B52FFD7FB95F9C1FBF4F9BC" box="[1041,1089,1590,1615]" name="Palau" pageId="11" pageNumber="12">Pala</collectingCountry>
shorter than the apical portion of penis. Trigonium with wide base, waisted, with claw-shaped centema. The flat parameroids very slender, curving around trigonium at some distance. Their widened axe-shaped tips turn mediad and meet, forming a frame around the trigonium (
<figureCitation id="01A02A3E9B52FFD7FE17F955FE52F948" box="[403,487,1698,1723]" captionStart="FIGURES 29 37" captionStartId="10.[151,264,1707,1730]" captionTargetBox="[246,1345,203,1671]" captionTargetId="figure@10.[228,1358,197,1686]" captionTargetPageId="10" captionText="FIGURES 29 37. Calvarium (Calvariellum) spp., males. C. cochlearifer, n. sp.: 29, T 8 and S 8, superimposed; 30, penis; 31, T 9, S 9 and tegmen (tn) with parameres (p) superimposed. C. bellendenker, n. sp.: 32, T 8; 33, S 8; 34, T 9; 35, S 9; 36, penis; 37, tegmen and parameres. Figures for a given species to the same scale." httpUri="https://zenodo.org/record/286799/files/figure.png" pageId="11" pageNumber="12">Fig. 36</figureCitation>
).
</paragraph>
<paragraph id="992436BB9B52FFD7FF43F930FE51F8F0" blockId="11.[151,1437,1158,2011]" pageId="11" pageNumber="12">
Tegmen and parameres shaped like a slender Y, parameres basally abruptly widened and then tapering to sharply pointed tips (
<figureCitation id="01A02A3E9B52FFD7FE07F91DFE63F8F0" box="[387,470,1770,1795]" captionStart="FIGURES 29 37" captionStartId="10.[151,264,1707,1730]" captionTargetBox="[246,1345,203,1671]" captionTargetId="figure@10.[228,1358,197,1686]" captionTargetPageId="10" captionText="FIGURES 29 37. Calvarium (Calvariellum) spp., males. C. cochlearifer, n. sp.: 29, T 8 and S 8, superimposed; 30, penis; 31, T 9, S 9 and tegmen (tn) with parameres (p) superimposed. C. bellendenker, n. sp.: 32, T 8; 33, S 8; 34, T 9; 35, S 9; 36, penis; 37, tegmen and parameres. Figures for a given species to the same scale." httpUri="https://zenodo.org/record/286799/files/figure.png" pageId="11" pageNumber="12">Fig. 37</figureCitation>
).
</paragraph>
<paragraph id="992436BB9B52FFD7FF43F8F9FC49F8D4" blockId="11.[151,1437,1158,2011]" box="[199,1020,1806,1831]" pageId="11" pageNumber="12">
<emphasis id="ABEFEAA99B52FFD7FF43F8F9FEABF8D4" bold="true" box="[199,286,1806,1831]" pageId="11" pageNumber="12">Female</emphasis>
(presumed). As for genus. No prehensor or vulvar sclerite found.
</paragraph>
<paragraph id="992436BB9B52FFD7FF43F8C5FB63F861" blockId="11.[151,1437,1158,2011]" pageId="11" pageNumber="12">
<emphasis id="ABEFEAA99B52FFD7FF43F8C5FF4AF8B8" bold="true" box="[199,255,1842,1867]" pageId="11" pageNumber="12">Note</emphasis>
. The caudal contact between the ends of the parameroids distantly resembles the structure of
<taxonomicName id="5E9B4D389B52FFD7FF13F8A1FE87F89D" box="[151,306,1878,1902]" class="Insecta" family="Scirtidae" genus="Austrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="ABEFEAA99B52FFD7FF13F8A1FE87F89D" box="[151,306,1878,1902]" italics="true" pageId="11" pageNumber="12">Austrocyphon</emphasis>
</taxonomicName>
spp. (Zwick 2013c). The female is very probably conspecific with the males with which it was taken. However, it cannot unambiguously be identified. The female is therefore assigned no
<typeStatus id="462088199B52FFD7FBD3F88CFB32F860" box="[1111,1159,1915,1939]" pageId="11" pageNumber="12">type</typeStatus>
status.
</paragraph>
</subSubSection>
<subSubSection id="D18165309B52FFD7FF43F869FE3FF828" pageId="11" pageNumber="12" type="etymology">
<paragraph id="992436BB9B52FFD7FF43F869FE3FF828" blockId="11.[151,1437,1158,2011]" pageId="11" pageNumber="12">
<emphasis id="ABEFEAA99B52FFD7FF43F869FEF3F844" bold="true" box="[199,326,1950,1975]" pageId="11" pageNumber="12">Etymology</emphasis>
. Named after the
<typeStatus id="462088199B52FFD7FD8AF868FD8BF844" box="[526,574,1951,1975]" pageId="11" pageNumber="12">type</typeStatus>
locality over which Mt Bellenden Ker peaks, the second highest mountain in Queensland (
<quantity id="5E639B5E9B52FFD7FEAEF835FEC8F828" box="[298,381,1986,2011]" metricMagnitude="3" metricUnit="m" metricValue="1.557" pageId="11" pageNumber="12" unit="m" value="1557.0">1557m</quantity>
).
</paragraph>
</subSubSection>
</treatment>
</document>