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<document id="AAF4A9E136BEAEBA560C073E0248CB02" ID-DOI="10.11646/zootaxa.3846.1.1" ID-GBIF-Dataset="01a9d979-7b10-4c36-b1af-4acc2c0cb9a7" ID-ISSN="1175-5326" ID-Zenodo-Dep="286792" ID-ZooBank="97D4A04A-D75E-45CC-8A70-3EB3A4E94D9B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460464243854" checkinUser="plazi" docAuthor="Zwick, Peter" docDate="2014" docId="113287AD9B4AFFCFFF13FCAAFBD5F806" docLanguage="en" docName="zt03846p041.pdf" docOrigin="Zootaxa 3846 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ypsiloncyphon virgulifer Zwick, 2014, n. sp." docType="treatment" docUuid="A6313C20-29E3-47A8-907B-5EA0B1E31DD0" docUuidSource="ZooBank" docVersion="7" lastPageNumber="20" masterDocId="ED0BFFD59B59FFDCFF84FFF7FFB5FFF3" masterDocTitle="Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer" masterLastPageNumber="41" masterPageNumber="1" pageNumber="20" updateTime="1698304595582" updateUser="plazi">
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<mods:title id="E59AC9CB1DAFA0D1B4E900B7F5F9D422">Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer</mods:title>
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<mods:namePart id="9170A482F5F7B442535BCF01222735B6">Zwick, Peter</mods:namePart>
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<mods:date id="0D5CCE5686AD8F6AB188AA39D5E7FBE7">2014</mods:date>
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<mods:number id="2FF9908BC4B5F8EAA87DA6E0B8900B29">3846</mods:number>
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<treatment id="113287AD9B4AFFCFFF13FCAAFBD5F806" ID-DOI="http://doi.org/10.5281/zenodo.6126829" ID-GBIF-Taxon="127669637" ID-Zenodo-Dep="6126829" LSID="urn:lsid:zoobank.org:act:A6313C20-29E3-47A8-907B-5EA0B1E31DD0" httpUri="http://treatment.plazi.org/id/113287AD9B4AFFCFFF13FCAAFBD5F806" lastPageNumber="20" pageId="19" pageNumber="20">
<subSubSection id="D18165309B4AFFCFFF13FCAAFE9BFC6B" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="992436BB9B4AFFCFFF13FCAAFDA5FC84" blockId="19.[151,528,861,920]" box="[151,528,861,887]" pageId="19" pageNumber="20">
<heading id="C26C81D79B4AFFCFFF13FCAAFDA5FC84" bold="true" box="[151,528,861,887]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<emphasis id="ABEFEAA99B4AFFCFFF13FCAAFDA5FC84" bold="true" box="[151,528,861,887]" pageId="19" pageNumber="20">
<emphasis id="ABEFEAA99B4AFFCFFF13FCAAFE72FC84" bold="true" box="[151,455,861,887]" italics="true" pageId="19" pageNumber="20">
<taxonomicName id="5E9B4D389B4AFFCFFF13FCAAFE74FC84" box="[151,449,861,887]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="virgulifer" status="sp. nov.">Ypsiloncyphon virgulifer</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="B0DC57D29B4AFFCFFE4AFCAAFDA5FC84" box="[462,528,861,887]" pageId="19" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="992436BB9B4AFFCFFF13FC88FE9BFC6B" blockId="19.[151,528,861,920]" box="[151,302,895,920]" pageId="19" pageNumber="20">
(
<figureCitation id="01A02A3E9B4AFFCFFF1BFC88FE93FC6B" box="[159,294,895,920]" captionStart="FIGURES 58 61" captionStartId="20.[151,264,1331,1354]" captionTargetBox="[165,1414,202,1298]" captionTargetId="figure@20.[160,1427,195,1306]" captionTargetPageId="20" captionText="FIGURES 58 61. Ypsiloncyphon virgulifer, n. sp., male: 58, T 8, T 9 and S 9, superimposed; 59, penis, ventral view; 60, oblique dorsolateral view of same, trigonium tip bent ventrad; 61, tegmen and parameres. Pilosity and surface structure of T 8 and T 9 shown only diagrammatically in small selected areas. All to same scale." httpUri="https://zenodo.org/record/286804/files/figure.png" pageId="19" pageNumber="20">Figs. 5861</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D18165309B4AFFCFFF13FC30FC01FA9E" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="992436BB9B4AFFCFFF13FC30FD7FFB66" blockId="19.[151,1437,967,2037]" pageId="19" pageNumber="20">
<emphasis id="ABEFEAA99B4AFFCFFF13FC30FEF6FC13" bold="true" box="[151,323,967,992]" pageId="19" pageNumber="20">
<typeStatus id="462088199B4AFFCFFF13FC30FF67FC13" box="[151,210,967,992]" pageId="19" pageNumber="20">Type</typeStatus>
material
</emphasis>
.
<emphasis id="ABEFEAA99B4AFFCFFED2FC30FE76FC13" bold="true" box="[342,451,967,992]" pageId="19" pageNumber="20">
<typeStatus id="462088199B4AFFCFFED2FC30FE76FC13" box="[342,451,967,992]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
</emphasis>
♂: “
<emphasis id="ABEFEAA99B4AFFCFFD81FC30FD9AFC13" bold="true" box="[517,559,967,992]" pageId="19" pageNumber="20">WA</emphasis>
: Cadjeput Rockhole E of Woodstock Station
<geoCoordinate id="FCAF507C9B4AFFCFFBD2FC30FB60FC13" box="[1110,1237,967,992]" direction="south" orientation="latitude" pageId="19" pageNumber="20" precision="15" value="-21.531944">21°31'55''S</geoCoordinate>
,
<geoCoordinate id="FCAF507C9B4AFFCFFB63FC30FAC1FC13" box="[1255,1396,967,992]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="15" value="119.14916">119°08'57''E</geoCoordinate>
<date id="ED25107B9B4AFFCFFA04FC3FFEF8FBF7" pageId="19" pageNumber="20" value="1988-09-29">29 September 1988</date>
B.P.Hanich et al. \ BPH 29-11AI UV fluorescent light at night 6:008:
<specimenCount id="8F9DFD329B4AFFCFFBDFFC1BFB21FBF7" box="[1115,1172,1004,1029]" pageId="19" pageNumber="20" type="generic">45 p.</specimenCount>
m. \ Western Australian Museum Entomology Reg. no. 42898 \ C.H.S.Watts det.
<taxonomicName id="5E9B4D389B4AFFCFFC92FBF8FCDAFBDB" box="[790,879,1039,1064]" class="Insecta" family="Scirtidae" genus="Cyphon" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="41" phylum="Arthropoda" rank="genus">Cyphon</taxonomicName>
sp.” (
<collectionCode id="FF8AAE7E9B4AFFCFFC35FBE7FBA2FBDB" box="[945,1047,1040,1064]" pageId="19" pageNumber="20">WAMW</collectionCode>
).
<emphasis id="ABEFEAA99B4AFFCFFBADFBF8FB14FBDB" bold="true" box="[1065,1185,1039,1064]" pageId="19" pageNumber="20">
<typeStatus id="462088199B4AFFCFFBADFBF8FB14FBDB" box="[1065,1185,1039,1064]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
</emphasis>
(all
<collectionCode id="FF8AAE7E9B4AFFCFFB50FBE7FAAFFBDB" LSID="urn:lsid:biocol.org:col:32981" box="[1236,1306,1040,1064]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="19" pageNumber="20">ANIC</collectionCode>
)
<emphasis id="ABEFEAA99B4AFFCFFAADFBF8FAFBFBDB" bold="true" box="[1321,1358,1039,1064]" pageId="19" pageNumber="20">NT</emphasis>
:
<specimenCount id="8F9DFD329B4AFFCFFAD9FBE7FA36FBDB" box="[1373,1411,1040,1064]" pageId="19" pageNumber="20" type="generic">1♂</specimenCount>
: “ Port Keat N.T.
<geoCoordinate id="FCAF507C9B4AFFCFFEC5FBC3FE3AFBBE" box="[321,399,1076,1101]" direction="south" orientation="latitude" pageId="19" pageNumber="20" precision="555" value="-14.06">14.06S</geoCoordinate>
<geoCoordinate id="FCAF507C9B4AFFCFFE12FBC3FE41FBBF" box="[406,500,1076,1100]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="555" value="129.33">129.33E</geoCoordinate>
<date id="ED25107B9B4AFFCFFE7FFBC3FD37FBBE" box="[507,642,1076,1101]" pageId="19" pageNumber="20" value="1968-08-19">19.viii.1968</date>
M.Mendum”;
<specimenCount id="8F9DFD329B4AFFCFFCAEFBC3FCE5FBBF" box="[810,848,1076,1100]" pageId="19" pageNumber="20" type="generic">1♂</specimenCount>
: “
<geoCoordinate id="FCAF507C9B4AFFCFFCE2FBC3FC03FBBE" box="[870,950,1076,1101]" direction="south" orientation="latitude" pageId="19" pageNumber="20" precision="555" value="-16.35">16.35S</geoCoordinate>
<geoCoordinate id="FCAF507C9B4AFFCFFC3AFBC3FBAEFBBF" box="[958,1051,1076,1100]" direction="south" orientation="latitude" pageId="19" pageNumber="20" precision="555" value="-131.17">131.17S</geoCoordinate>
GPS Sullivan Ck Gregory NP NT
<date id="ED25107B9B4AFFCFFF13FBAFFE9EFB83" box="[151,299,1111,1136]" pageId="19" pageNumber="20" value="1994-07-21">21 July 1994</date>
T.Weir A.Roach deep billabong rocky edges“;
<emphasis id="ABEFEAA99B4AFFCFFCCCFBA0FCC7FB83" bold="true" box="[840,882,1111,1136]" pageId="19" pageNumber="20">WA</emphasis>
:
<specimenCount id="8F9DFD329B4AFFCFFC02FBAFFC19FB83" box="[902,940,1112,1136]" pageId="19" pageNumber="20" type="generic">1♂</specimenCount>
: “Crossing Pool, Millstream [
<geoCoordinate id="FCAF507C9B4AFFCFFA84FBAFFA29FB83" box="[1280,1436,1112,1136]" direction="south" orientation="latitude" pageId="19" pageNumber="20" precision="1" value="-21.574816">21° 34.489' S</geoCoordinate>
<geoCoordinate id="FCAF507C9B4AFFCFFF13FB8BFEF5FB67" box="[151,320,1148,1173]" direction="east" orientation="longitude" pageId="19" pageNumber="20" precision="1" value="117.084435">117° 05.066' E</geoCoordinate>
], W.A..
<date id="ED25107B9B4AFFCFFE18FB8BFD93FB67" box="[412,550,1148,1172]" pageId="19" pageNumber="20" value="1970-10-22">22.Oct.1970</date>
D.H.Colless”.
</paragraph>
<paragraph id="992436BB9B4AFFCFFF43FB68FCA4FB2E" blockId="19.[151,1437,967,2037]" pageId="19" pageNumber="20">
<emphasis id="ABEFEAA99B4AFFCFFF43FB68FE98FB4B" bold="true" box="[199,301,1183,1208]" pageId="19" pageNumber="20">Habitus.</emphasis>
<collectionCode id="FF8AAE7E9B4AFFCFFEB3FB57FEEEFB44" box="[311,347,1184,1207]" pageId="19" pageNumber="20">BL</collectionCode>
<quantity id="5E639B5E9B4AFFCFFEE1FB57FDBBFB44" box="[357,526,1184,1208]" metricMagnitude="-3" metricUnit="m" metricValue="1.67" metricValueMax="1.9" metricValueMin="1.44" pageId="19" pageNumber="20" unit="mm" value="1.67" valueMax="1.9" valueMin="1.44">1.441.90 mm</quantity>
,
<collectionCode id="FF8AAE7E9B4AFFCFFD99FB57FDF5FB4B" box="[541,576,1184,1208]" pageId="19" pageNumber="20">BL</collectionCode>
/BW ~1.55, oval. Pronotal front angles projecting little, rounded. Sides of pronotum regularly arched, base barely wider than front.
</paragraph>
<paragraph id="992436BB9B4AFFCFFF43FB1FFC01FA9E" blockId="19.[151,1437,967,2037]" pageId="19" pageNumber="20">Punctures on head fine, granular especially behind eyes. Pronotal punctures large, granular, increasingly so towards the sides where raised little pearls are about 1.5 diameters apart. Punctation on scutellum obsolete. Elytra shining, with normal punctures, basal ones coarse, caudally increasingly finer and denser. Dorsal side blackish brown, antennae yellowish, legs with brownish femora, distally lighter.</paragraph>
</subSubSection>
<subSubSection id="D18165309B4AFFCFFF43FA80FBD5F806" pageId="19" pageNumber="20" type="description">
<paragraph id="992436BB9B4AFFCFFF43FA80FC98F943" blockId="19.[151,1437,967,2037]" pageId="19" pageNumber="20">
<emphasis id="ABEFEAA99B4AFFCFFF43FA80FEB1FA63" bold="true" box="[199,260,1399,1424]" pageId="19" pageNumber="20">Male</emphasis>
. Caudal margin of T8 approximately pentagonal, with a few irregularly arranged setae and grouped microtrichia, anterolaterally with some sharp raised granulations. Apodemes barely longer than the plate, straight, medially constricted, base pointed. T9 medially membranous, the lightly slcerotized sides with small spicules and elongate asperities. Apodemes more than twice longer than those of of T8, strong, straight. S9 strongly sclerotized, with loop-like base supporting two closely appressed stiff, caudally pointed sclerite rods (
<figureCitation id="01A02A3E9B4AFFCFFB01F9F0FB6FF9D3" box="[1157,1242,1543,1568]" captionStart="FIGURES 58 61" captionStartId="20.[151,264,1331,1354]" captionTargetBox="[165,1414,202,1298]" captionTargetId="figure@20.[160,1427,195,1306]" captionTargetPageId="20" captionText="FIGURES 58 61. Ypsiloncyphon virgulifer, n. sp., male: 58, T 8, T 9 and S 9, superimposed; 59, penis, ventral view; 60, oblique dorsolateral view of same, trigonium tip bent ventrad; 61, tegmen and parameres. Pilosity and surface structure of T 8 and T 9 shown only diagrammatically in small selected areas. All to same scale." httpUri="https://zenodo.org/record/286804/files/figure.png" pageId="19" pageNumber="20">Fig. 58</figureCitation>
). Penis (
<figureCitation id="01A02A3E9B4AFFCFFABDF9F0FF01F9B6" captionStart="FIGURES 58 61" captionStartId="20.[151,264,1331,1354]" captionTargetBox="[165,1414,202,1298]" captionTargetId="figure@20.[160,1427,195,1306]" captionTargetPageId="20" captionText="FIGURES 58 61. Ypsiloncyphon virgulifer, n. sp., male: 58, T 8, T 9 and S 9, superimposed; 59, penis, ventral view; 60, oblique dorsolateral view of same, trigonium tip bent ventrad; 61, tegmen and parameres. Pilosity and surface structure of T 8 and T 9 shown only diagrammatically in small selected areas. All to same scale." httpUri="https://zenodo.org/record/286804/files/figure.png" pageId="19" pageNumber="20">Figs. 59, 60</figureCitation>
) long, truncate base parallel for about ¼ of total penis length, in side view at a distinct angle to distal part. Trigonium wide, band-like, ending in a sharp sickle-shaped centema (apex bent back in
<figureCitation id="01A02A3E9B4AFFCFFB26F9B8FB4CF99B" box="[1186,1273,1615,1640]" captionStart="FIGURES 58 61" captionStartId="20.[151,264,1331,1354]" captionTargetBox="[165,1414,202,1298]" captionTargetId="figure@20.[160,1427,195,1306]" captionTargetPageId="20" captionText="FIGURES 58 61. Ypsiloncyphon virgulifer, n. sp., male: 58, T 8, T 9 and S 9, superimposed; 59, penis, ventral view; 60, oblique dorsolateral view of same, trigonium tip bent ventrad; 61, tegmen and parameres. Pilosity and surface structure of T 8 and T 9 shown only diagrammatically in small selected areas. All to same scale." httpUri="https://zenodo.org/record/286804/files/figure.png" pageId="19" pageNumber="20">Fig. 60</figureCitation>
). Parameroids arising from protruding ear-like bases, little longer than trigonium. Distally, the parameroids resemble open tubes. They support a subterminal sclerotized sinuous appendage.
</paragraph>
<paragraph id="992436BB9B4AFFCFFF43F94AFD28F8EE" blockId="19.[151,1437,967,2037]" pageId="19" pageNumber="20">
Tegmen and parameres are a narrow Y-shaped sclerite with truncate, enlarged, sclerotized base forking into 2 strong rods. Caudally, they are increasingly less sclerotized, the distal third appears to be longitudinally rolled, ending in a transparent pointed apex (
<figureCitation id="01A02A3E9B4AFFCFFDB9F8F3FD25F8EE" box="[573,656,1796,1821]" captionStart="FIGURES 58 61" captionStartId="20.[151,264,1331,1354]" captionTargetBox="[165,1414,202,1298]" captionTargetId="figure@20.[160,1427,195,1306]" captionTargetPageId="20" captionText="FIGURES 58 61. Ypsiloncyphon virgulifer, n. sp., male: 58, T 8, T 9 and S 9, superimposed; 59, penis, ventral view; 60, oblique dorsolateral view of same, trigonium tip bent ventrad; 61, tegmen and parameres. Pilosity and surface structure of T 8 and T 9 shown only diagrammatically in small selected areas. All to same scale." httpUri="https://zenodo.org/record/286804/files/figure.png" pageId="19" pageNumber="20">Fig. 61</figureCitation>
).
</paragraph>
<paragraph id="992436BB9B4AFFCFFF43F8D0FE17F8B3" blockId="19.[151,1437,967,2037]" box="[199,418,1831,1856]" pageId="19" pageNumber="20">
<emphasis id="ABEFEAA99B4AFFCFFF43F8D0FE90F8B3" bold="true" box="[199,293,1831,1856]" pageId="19" pageNumber="20">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="992436BB9B4AFFCFFF43F8BBFBD5F806" blockId="19.[151,1437,967,2037]" pageId="19" pageNumber="20">
<emphasis id="ABEFEAA99B4AFFCFFF43F8BBFE07F896" bold="true" box="[199,434,1868,1893]" pageId="19" pageNumber="20">Note and etymology</emphasis>
. By its genitalia and the shape of the mesoventral process
<taxonomicName id="5E9B4D389B4AFFCFFBBFF8BAFB09F897" box="[1083,1212,1868,1892]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="virgulifer">
<emphasis id="ABEFEAA99B4AFFCFFBBFF8BAFB09F897" box="[1083,1212,1868,1892]" italics="true" pageId="19" pageNumber="20">Y. virgulifer</emphasis>
</taxonomicName>
is closely related to the other Australian species, even though these have an asymmetrical S9 and simple parameroids. Because the female is unknown,
<taxonomicName id="5E9B4D389B4AFFCFFEF8F862FDB4F85F" box="[380,513,1940,1964]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="virgulifer">
<emphasis id="ABEFEAA99B4AFFCFFEF8F862FDB4F85F" box="[380,513,1940,1964]" italics="true" pageId="19" pageNumber="20">Y. virgulifer</emphasis>
</taxonomicName>
is only tentatively included in
<emphasis id="ABEFEAA99B4AFFCFFCE4F862FCC6F85F" box="[864,883,1941,1964]" italics="true" pageId="19" pageNumber="20">Y.</emphasis>
species group 3. The localities are spread over a wide range in the Northwest of the continent, from the Hamersley Ranges to the Kimberleys SW of Darwin. Name derived from Latin,
<emphasis id="ABEFEAA99B4AFFCFFE46F82BFDA4F807" box="[450,529,2012,2036]" italics="true" pageId="19" pageNumber="20">virgula</emphasis>
, branchlet, and
<emphasis id="ABEFEAA99B4AFFCFFD41F82BFD4FF807" box="[709,762,2012,2036]" italics="true" pageId="19" pageNumber="20">ferre</emphasis>
, to carry: carrier of a branchlet.
</paragraph>
</subSubSection>
</treatment>
</document>