136 lines
17 KiB
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136 lines
17 KiB
XML
<document id="B99BC1DB182C38BDA924314962C4563C" ID-DOI="10.5281/zenodo.213542" ID-GBIF-Dataset="094b3cc3-9dc5-4862-8033-e1ade495d594" ID-ISSN="1175-5326" ID-Zenodo-Dep="213542" ID-ZooBank="A6E5477A-FBB3-4CF7-9151-F3F4A96FCC1F" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460535516969" checkinUser="plazi" docAuthor="Kania, Iwona & Krzemiński, Wiesław" docDate="2012" docId="041087A82908501352A7B069FEBCFB95" docLanguage="en" docName="zt03495p056.pdf" docOrigin="Zootaxa 3495" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Palaeopoecilostola eocenica Kania & Krzemiński, 2012, sp. nov." docType="treatment" docVersion="7" lastPageNumber="45" masterDocId="F829FFD0290950105230B566FFEEFB62" masterDocTitle="A new species of Palaeopoecilostola Meunier, 1899 (Diptera: Limoniidae) from the Eocene Baltic amber" masterLastPageNumber="56" masterPageNumber="42" pageNumber="43" updateTime="1698316298331" updateUser="plazi">
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<mods:title id="F47BAC2355EA8A0B6E051D6074E8F45A">A new species of Palaeopoecilostola Meunier, 1899 (Diptera: Limoniidae) from the Eocene Baltic amber</mods:title>
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<mods:name id="B0CAF5041D55D0416C54A9453FE699B6" type="personal">
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<mods:namePart id="6080CF996C5DB07D238CDD3857331F82">Kania, Iwona</mods:namePart>
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<mods:namePart id="21C8214437BBBBD95712470FE453BEA7">Krzemiński, Wiesław</mods:namePart>
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<mods:typeOfResource id="2DDE72A5027A6BCC34653E3256D23E93">text</mods:typeOfResource>
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<mods:title id="4BE80A58A9E5A9ED344FFD2D57190573">Zootaxa</mods:title>
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<mods:part id="B709CA6EF26E9287B027DB10C6C83942">
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<mods:date id="5145C451D541D3F3A509632732F1E9E1">2012</mods:date>
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<mods:number id="52134DC65C147E638F7651C5DF7CC275">3495</mods:number>
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<mods:start id="A0014B7A3EAE5D190EEB6F985B255998">42</mods:start>
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<mods:identifier id="DF02B76EDDD0929A1E0AE2CEDE10475A" type="DOI">10.5281/zenodo.213542</mods:identifier>
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<mods:identifier id="236B9D5E836536554D51C0EDF8BBE5E8" type="GBIF-Dataset">094b3cc3-9dc5-4862-8033-e1ade495d594</mods:identifier>
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<mods:identifier id="C6CB8045DC8B01267D313C484041D133" type="ISSN">1175-5326</mods:identifier>
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<treatment id="041087A82908501352A7B069FEBCFB95" ID-DOI="http://doi.org/10.5281/zenodo.6494030" ID-GBIF-Taxon="119527568" ID-Zenodo-Dep="6494030" LSID="urn:lsid:plazi:treatment:041087A82908501352A7B069FEBCFB95" httpUri="http://treatment.plazi.org/id/041087A82908501352A7B069FEBCFB95" lastPageId="3" lastPageNumber="45" pageId="1" pageNumber="43">
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<subSubSection id="C4A365352908501152A7B069FEE4FE28" pageId="1" pageNumber="43" type="nomenclature">
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<paragraph id="8C0636BE2908501152A7B069FDA1FE4B" blockId="1.[151,591,1295,1354]" box="[151,591,1295,1321]" pageId="1" pageNumber="43">
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<heading id="D74E81D22908501152A7B069FDA1FE4B" bold="true" box="[151,591,1295,1321]" fontSize="11" level="1" pageId="1" pageNumber="43" reason="1">
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<emphasis id="BECDEAAC2908501152A7B069FDA1FE4B" bold="true" box="[151,591,1295,1321]" pageId="1" pageNumber="43">
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<taxonomicName id="4BB94D3D2908501152A7B069FE04FE4B" box="[151,490,1295,1321]" class="Insecta" family="Limoniidae" genus="Palaeopoecilostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="43" phylum="Arthropoda" rank="species" species="eocenica" status="sp. nov.">
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<emphasis id="BECDEAAC2908501152A7B069FE04FE4B" bold="true" box="[151,490,1295,1321]" italics="true" pageId="1" pageNumber="43">Palaeopoecilostola eocenica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A5FE57D72908501153C1B069FDA1FE4B" box="[497,591,1295,1321]" pageId="1" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8C0636BE2908501152A7B054FEE4FE28" blockId="1.[151,591,1295,1354]" box="[151,266,1330,1354]" pageId="1" pageNumber="43">
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(
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<figureCitation id="14822A3B2908501152AFB054FEEFFE28" box="[159,257,1330,1354]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 8" captionStartId-0="2.[151,264,1571,1594]" captionStartId-1="3.[151,264,1907,1930]" captionTargetBox-0="[158,1428,182,1542]" captionTargetBox-1="[160,1414,338,1838]" captionTargetId-0="figure@2.[151,1436,171,1556]" captionTargetId-1="figure@3.[151,1436,319,1862]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 – 4. Palaeopoecilostola eocenica sp. nov., holotype 1. Lateral view. 2. Antennae. 3. Hypopygium. 4. Wing venation." captionText-1="FIGURES 5 – 8. Palaeopoecilostola eocenica sp. nov., holotype 1. Antennae. 2. Palpi. 3. Wing venation. 4. Hypopygium." httpUri-0="https://zenodo.org/record/213543/files/figure.png" httpUri-1="https://zenodo.org/record/213544/files/figure.png" pageId="1" pageNumber="43">Figs 1–8</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C4A365352908501152A7B013FD74FE9B" pageId="1" pageNumber="43" type="diagnosis">
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<paragraph id="8C0636BE2908501152A7B013FD74FE9B" blockId="1.[151,1436,1397,2033]" pageId="1" pageNumber="43">
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<emphasis id="BECDEAAC2908501152A7B013FEE4FEEC" bold="true" box="[151,266,1397,1422]" italics="true" pageId="1" pageNumber="43">Diagnosis</emphasis>
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. Antennae 16-segmented, covered by long but not numerous setae, the base of each flagellomeres with two very elongated setae and rarely appearing shorter setae, usually reaching more than half the length of the elongated setae; outer dististylus wide and short, strongly expanded at the end, with short and thick setae; inner dististylus wide at the base, tapered at the top.
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</paragraph>
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</subSubSection>
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<subSubSection id="C4A365352908501252F7B363FCD1FC83" lastPageId="2" lastPageNumber="44" pageId="1" pageNumber="43" type="description">
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<paragraph id="8C0636BE2908501152F7B363FF05FD97" blockId="1.[151,1436,1397,2033]" pageId="1" pageNumber="43">
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<emphasis id="BECDEAAC2908501152F7B363FEA3FD7C" bold="true" box="[199,333,1541,1566]" italics="true" pageId="1" pageNumber="43">Description</emphasis>
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. The body length: 12.5 mm.; wing length 6.9 mm, width 4.1 mm, antennae 2.6 mm long. Head (
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<figureCitation id="14822A3B2908501152AFB34FFF08FD23" box="[159,230,1577,1601]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1571,1594]" captionTargetBox="[158,1428,182,1542]" captionTargetId="figure@2.[151,1436,171,1556]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Palaeopoecilostola eocenica sp. nov., holotype 1. Lateral view. 2. Antennae. 3. Hypopygium. 4. Wing venation." httpUri="https://zenodo.org/record/213543/files/figure.png" pageId="1" pageNumber="43">Fig. 1</figureCitation>
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): slender with characteristic huge eyes; antennae (Figs 1,2,5) shorter than mid-length of the abdomen, usually pale; shape almost cylindrical, small; pedicel laterally swollen, almost cylindrical in shape; antennal flagellomeres narrow, elongate, cylindrical, covered by tiny, wispy setae, each flagellomere with two elongated setae, additionally, a few 2–4 setae are visible on all flagellomeres, the same length or slightly longer than the length of the flagellomere on which they occur; the last segment almost as long as the sub-apical one, slightly shorter.
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</paragraph>
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<paragraph id="8C0636BE2908501152F7B267FB97FC7B" blockId="1.[151,1436,1397,2033]" box="[199,1145,1793,1818]" pageId="1" pageNumber="43">
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Palpi (
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<figureCitation id="14822A3B290850115320B267FEBBFC7B" box="[272,341,1793,1817]" captionStart="FIGURES 5 – 8" captionStartId="3.[151,264,1907,1930]" captionTargetBox="[160,1414,338,1838]" captionTargetId="figure@3.[151,1436,319,1862]" captionTargetPageId="3" captionText="FIGURES 5 – 8. Palaeopoecilostola eocenica sp. nov., holotype 1. Antennae. 2. Palpi. 3. Wing venation. 4. Hypopygium." httpUri="https://zenodo.org/record/213544/files/figure.png" pageId="1" pageNumber="43">Fig. 6</figureCitation>
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): 4-segmented, the last segment subtly longer than the penultimate one.
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</paragraph>
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<paragraph id="8C0636BE2908501152F7B243FD75FC93" blockId="1.[151,1436,1397,2033]" pageId="1" pageNumber="43">Wings (Figs 1,4,7): strongly dark, stigma clearly separated; Sc long, ending a little bit past bifurcation of vein Rs into R2+3+4 and R5; Rs slightly longer than R2+3+4; R2+3+4 two times as long as R3; R1 ending opposite proximal ¼ of R3 length; r-r (R2) almost before the end of R1; R4 twice as long as R3; discal cell small, rectangular; M1 about 1/3 longer than petiola, cross vein m-cu behind the midpoint of discal cell base and approximately as long as d-cell base; A1 straight and long; A2 sinuous for half of its length, the distal part of this vein strongly curved towards the wing’s edge.</paragraph>
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<caption id="D8C66636290B501252A7B345FF18FD3A" httpUri="https://zenodo.org/record/213543/files/figure.png" pageId="2" pageNumber="44" targetBox="[158,1428,182,1542]" targetPageId="2">
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<paragraph id="8C0636BE290B501252A7B345FF18FD3A" blockId="2.[151,1436,1571,1624]" pageId="2" pageNumber="44">
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<emphasis id="BECDEAAC290B501252A7B345FED6FD5B" bold="true" box="[151,312,1571,1594]" pageId="2" pageNumber="44">FIGURES 1–4</emphasis>
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.
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<emphasis id="BECDEAAC290B5012537AB342FC8CFD5B" bold="true" box="[330,866,1572,1594]" pageId="2" pageNumber="44">
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<taxonomicName id="4BB94D3D290B5012537AB342FD9AFD58" box="[330,628,1572,1594]" class="Insecta" family="Limoniidae" genus="Palaeopoecilostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="44" phylum="Arthropoda" rank="species" species="eocenica" status="sp. nov.">
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<emphasis id="BECDEAAC290B5012537AB342FD9AFD58" bold="true" box="[330,628,1572,1594]" italics="true" pageId="2" pageNumber="44">Palaeopoecilostola eocenica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A5FE57D7290B5012504FB342FD38FD5B" box="[639,726,1572,1593]" pageId="2" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype 1.
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</emphasis>
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Lateral view.
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<emphasis id="BECDEAAC290B50125630B342FBE3FD5B" bold="true" box="[1024,1037,1572,1593]" pageId="2" pageNumber="44">2</emphasis>
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. Antennae.
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<emphasis id="BECDEAAC290B501256A1B342FB73FD5B" bold="true" box="[1169,1181,1572,1593]" pageId="2" pageNumber="44">3</emphasis>
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. Hypopygium.
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<emphasis id="BECDEAAC290B50125776B342FABDFD5B" bold="true" box="[1350,1363,1572,1593]" pageId="2" pageNumber="44">4</emphasis>
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. Wing venation.
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</paragraph>
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</caption>
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<paragraph id="8C0636BE290B501252F7B3E3FE40FD87" blockId="2.[151,1437,1669,2018]" pageId="2" pageNumber="44">
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Hypopygium (
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<figureCitation id="14822A3B290B5012535DB3E3FE37FDFF" box="[365,473,1669,1693]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1571,1594]" captionTargetBox="[158,1428,182,1542]" captionTargetId="figure@2.[151,1436,171,1556]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Palaeopoecilostola eocenica sp. nov., holotype 1. Lateral view. 2. Antennae. 3. Hypopygium. 4. Wing venation." httpUri="https://zenodo.org/record/213543/files/figure.png" pageId="2" pageNumber="44">Figs 1, 3</figureCitation>
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,
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<figureCitation id="14822A3B290B501253DAB3E3FE14FDFF" box="[490,506,1669,1693]" captionStart="FIGURES 5 – 8" captionStartId="3.[151,264,1907,1930]" captionTargetBox="[160,1414,338,1838]" captionTargetId="figure@3.[151,1436,319,1862]" captionTargetPageId="3" captionText="FIGURES 5 – 8. Palaeopoecilostola eocenica sp. nov., holotype 1. Antennae. 2. Palpi. 3. Wing venation. 4. Hypopygium." httpUri="https://zenodo.org/record/213544/files/figure.png" pageId="2" pageNumber="44">8</figureCitation>
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): elongated, basistylus covered by wispy, elongate setae; outer dististylus wide, strongly expanded at the end, rather short with short and thick setae at outer edge; inner dististylus expanded at the base, narrow at the apex.
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</paragraph>
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<paragraph id="8C0636BE290B501252F7B397FCD1FC83" blockId="2.[151,1437,1669,2018]" pageId="2" pageNumber="44">
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<emphasis id="BECDEAAC290B501252F7B397FE5EFC6B" bold="true" box="[199,432,1777,1802]" italics="true" pageId="2" pageNumber="44">Age and occurence.</emphasis>
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Baltic amber, Upper Eocene. The Baltic amber is aged within the range of 38–47 Ma (
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<bibRefCitation id="E8284B4F290B501252AEB273FE8CFC4F" author="Ritzkowski" box="[158,354,1813,1837]" pageId="2" pageNumber="44" refString="Ritzkowski, S. (1997) K-Ar-Altersbestimmung der Bernsteinfuhrenden Sedimente des Sammlandes (Palaogen, Bezirk Kaliningrad). Metalla (Sonderheft) 66, 19 - 23." type="journal article" year="1997">Ritzkowski 1997</bibRefCitation>
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;
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<bibRefCitation id="E8284B4F290B5012535DB273FD8BFC4F" author="Perkovsky" box="[365,613,1813,1837]" pageId="2" pageNumber="44" refString="Perkovsky, E. E., Rasnitsyn, A. P., Vlaskin, A. P. & Taraschuk, M. V. (2007) A comparative analysis of the Baltic and Rovno amber arthropod faunas: representative samples. African Invertebrates, 48 (1), 229 - 245." type="journal article" year="2007">Perkovsky et al. 2007</bibRefCitation>
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). Absolute dating analyses of glauconites from
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<collectingCountry id="F4AE762E290B50125648B273FB22FC4F" box="[1144,1228,1813,1837]" name="Zambia" pageId="2" pageNumber="44">Sambia</collectingCountry>
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Peninsula showed that the “blue earth” formation (amber bearing Prussian Formation) is allocated to the Middle Eocene (Lutetian: 44.1 ± 1.1 Ma) and is thus significantly older than previously assumed (
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<bibRefCitation id="E8284B4F290B501251E0B23BFB9AFC17" author="Wappler" box="[976,1140,1885,1909]" pageId="2" pageNumber="44" refString="Wappler, T. (2003) Die Insekten aus dem Mittel-Eozan des Eckfelder Maares, Vulkaneifel. Mainzer Naturwissenschaftliches Archiv 27, i - viii + 1 - 234." type="journal article" year="2003">Wappler 2003</bibRefCitation>
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,
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<bibRefCitation id="E8284B4F290B501256B1B23BFB50FC17" author="Wappler" box="[1153,1214,1885,1909]" pageId="2" pageNumber="44" refString="Wappler, T. (2005) The age of Baltic amber: could Eckfeld resolve this problem? Fossils X 3, Insects - Arthropods - Amber. Programme and Abstracts, 3 rd International Congress of Palaeoentomology with 2 nd International Meeting on Palaeoarthropodology and 2 nd World Congress on Amber and its Inclusions, 7 th to 11 th February 2005, Pretoria, South Africa, 53." type="journal article" year="2005">2005</bibRefCitation>
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). Also Weitschat &
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<bibRefCitation id="E8284B4F290B501252A7B2E7FEA7FCFB" box="[151,329,1921,1945]" pageId="2" pageNumber="44" refString="Wetischat, W. & Wichard, W. (2010) Baltic amber, pp. 80 - 115 In: Penney, D. (Ed.), Biodiversity of fossils in amber from major world deposits. Siri Scientific Press, Manchester, UK." type="book chapter">Wichard (2010)</bibRefCitation>
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suggested older age of the Baltic amber. However, assumptions on the Middle Eocene (or older) age of Baltic amber was argued by
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<bibRefCitation id="E8284B4F290B50125014B2C3FCC2FCDF" box="[548,812,1957,1981]" pageId="2" pageNumber="44" refString="Perkovsky, E. E., Rasnitsyn, A. P., Vlaskin, A. P. & Taraschuk, M. V. (2007) A comparative analysis of the Baltic and Rovno amber arthropod faunas: representative samples. African Invertebrates, 48 (1), 229 - 245." type="journal article">Perkovsky et al. (2007)</bibRefCitation>
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, and the Upper Eocene (Bartonian/Priabonian: 37.7 ± 3 Ma) age of Prussian Formation is preferred by these authors.
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</paragraph>
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</subSubSection>
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<subSubSection id="C4A36535290A501352F7B5F1FEBCFB95" pageId="3" pageNumber="45" type="etymology">
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<paragraph id="8C0636BE290A501352F7B5F1FEBCFB95" blockId="3.[151,1436,151,247]" pageId="3" pageNumber="45">
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<emphasis id="BECDEAAC290A501352F7B5F1FEA9FBD2" bold="true" box="[199,327,151,176]" italics="true" pageId="3" pageNumber="45">Etymology.</emphasis>
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Specific epithet after the geological period—the Eocene.
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<emphasis id="BECDEAAC290A501352F7B5DDFE42FBB6" bold="true" box="[199,428,187,212]" italics="true" pageId="3" pageNumber="45">Material examined.</emphasis>
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<typeStatus id="5302881C290A50135387B5DAFDCCFBB6" box="[439,546,188,212]" pageId="3" pageNumber="45" type="holotype">Holotype</typeStatus>
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, male. No. 19931, Coll. Museum of the Earth Polish Academy of Sciences, Warsaw,
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<collectingCountry id="F4AE762E290A501352CDB5B9FEA0FB95" box="[253,334,223,247]" name="Poland" pageId="3" pageNumber="45">Poland</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |