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<mods:title id="3472400CDCD7FEA9E4902555FFF6C6B6">Taxonomy and nomenclature of Kalanchoe tomentosa (Crassulaceae subfam. Kalanchooideae), a variable, completely hairy species from Madagascar</mods:title>
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<treatmentCitation id="0A9810E0FFB0FF91FF01FB38FDCFA4A0" author="Baker, J. G." box="[136,597,1172,1199]" page=")" pageId="3" pageNumber="292" year="1882">
<taxonomicName id="4C394D72FFB0FF91FF01FB38FDCFA4A0" ID-CoL="6NJZK" ID-ENA="80922" authority="Baker (1882: 110)" authorityName="Baker" authorityPageNumber="110" authorityYear="1882" box="[136,597,1172,1199]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="292" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="B94DEAE3FFB0FF91FF01FB38FE18A4A0" bold="true" box="[136,386,1172,1199]" italics="true" pageId="3" pageNumber="292">Kalanchoe tomentosa</emphasis>
<bibRefCitation id="EFA84B00FFB0FF91FE00FB38FDCFA4A0" author="Baker, J. G." box="[393,597,1172,1198]" pageId="3" pageNumber="292" pagination="109 - 114" refId="ref5465" refString="Baker, J. G. (1882) Contributions to the flora of central Madagascar (continued from p. 70) and (to be continued). Journal of Botany, British and Foreign. London 20: 109 - 114. https: // doi. org / 10.1111 / j. 1095 - 8339.1884. tb 00548. x" type="journal article" year="1882">Baker (1882: 110)</bibRefCitation>
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(
<figureCitation id="13022A74FFB0FF91FDEDFB38FC99A4A0" box="[612,771,1172,1198]" captionStart="FIGURE 1" captionStartId="1.[136,235,1649,1673]" captionTargetBox="[163,1424,201,1614]" captionTargetId="figure-15@1.[151,1436,189,1626]" captionTargetPageId="1" captionText="FIGURE 1. A. Portrait of John Gilbert Baker (1834–1920) who published the name Kalanchoe tomentosa in 1882. Reproduced from Journal of Botany, British and Foreign. London, volume 39 (1901). Photograph: Berthold Carl Seemann. Public Domain. http://www.plantillustrations.org/illustration.php?id_illustration=176522. B. Portrait of Reverend Richard Baron (1847–1907) who collected the material on which Baker based the name K. tomentosa. Credit of the photograph is to Fiangonan’ I Jesoa Kristy Eto Madagasikara (FJKM), which is the lineal descendent of the London Missionary Society, Antananarivo, which employed Baron. Permission was granted to Dr Laurence J. Dorr by FJKM to disassemble a framed collection of photographs of 19th century missionaries to obtain individual portraits for use in Dorr (1997). Dr Dorr kindly supplied the image to the author of this paper. C. Kalanchoe tomentosa is a sparsely- to many-leaved, entirely hairy, sparsely branched, small to medium-sized, terrestrial, succulent shrub that can reach a height of about 1 m. Note the hard, stiff aerial roots that often develop on the stem and branches. D. The hairy flowers of K. tomentosa are borne erect to erectly spreading to spreading, and are hairy throughout. Flowers are campanulate to sometimes very slightly urceolate, yellowish brown to greenish and often sometimes purple-infused. The corolla lobes are light to deep purple on a greenish background, and adaxially velvety rather than hairy. Photographs C and D taken by Gideon F. Smith." figureDoi="http://doi.org/10.5281/zenodo.8297143" httpUri="https://zenodo.org/record/8297143/files/figure.png" pageId="3" pageNumber="292">Figs 1C &amp; 1D</figureCitation>
)
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<subSubSection id="C323657AFFB0FF91FF01FB70FCC1A5A4" pageId="3" pageNumber="292" type="multiple">
<paragraph id="8B8636F1FFB0FF91FF01FB70FCC1A5A4" blockId="3.[136,1452,1244,2063]" pageId="3" pageNumber="292">
<emphasis id="B94DEAE3FFB0FF91FF01FB70FEACA4F8" box="[136,310,1244,1270]" italics="true" pageId="3" pageNumber="292">Also treated in</emphasis>
:— 
<bibRefCitation id="EFA84B00FFB0FF91FEDEFB70FDD9A4F8" author="Baillon, M. H." box="[343,579,1244,1271]" pageId="3" pageNumber="292" pagination="465 - 472" refId="ref5310" refString="Baillon, M. H. (1885 [seance du 18 fevrier]) Liste des plantes de Madagascar (suite de la page 464). [Kalanchoe § Kitchingia]. Bulletin Mensuel de la Societe Linneenne de Paris (Paris) 1 (59): 465 - 472. [https: // www. biodiversitylibrary. org / item / 41447 # page / 60 / mode / 1 up]" type="journal article" year="1885">Baillon (1885: 468)</bibRefCitation>
; 
<bibRefCitation id="EFA84B00FFB0FF91FDDDFB71FCB2A4F8" author="Hamet, R." box="[596,808,1244,1271]" pageId="3" pageNumber="292" pagination="17 - 48" refId="ref6659" refString="Hamet, R. (1908) Monographie du genre Kalanchoe. Suite et fin. Bulletin de l'Herbier Boissier ser. 2 8: 17 - 48. [https: // www. biodiversitylibrary. org / item / 104945 # page / 31 / mode / 1 up]" type="journal article" year="1908">Hamet (1908: 31)</bibRefCitation>
; Raymond-Hamet &amp; 
<bibRefCitation id="EFA84B00FFB0FF91FBAEFB70FA3DA4F8" author="Perrier de la Bathie" box="[1063,1447,1244,1271]" pageId="3" pageNumber="292" pagination="63 - 117" refId="ref7867" refString="Raymond-Hamet &amp; Perrier de la Bathie, [J. M.] H. [A.] (1915) Troisieme contribution a l'etude des Crassulacees Malgaches. Annales du Musee Colonial de Marseille ser. 3 3: 63 - 117. [https: // www. biodiversitylibrary. org / item / 22992 # page / 85 / mode / 1 up]" type="journal article" year="1915">Perrier de la Bâthie (1915: 102)</bibRefCitation>
; Raymond-Hamet &amp; 
<bibRefCitation id="EFA84B00FFB0FF91FEE5FAACFD20A514" author="Marnier-Lapostolle, J." box="[364,698,1280,1306]" pageId="3" pageNumber="292" pagination="1 - 110" refId="ref7802" refString="Raymond-Hamet &amp; Marnier-Lapostolle, J. (1964) Le genre Kalanchoe au Jardin Botanique &quot; Les Cedres &quot;. Archives du Museum National d'Histoire Naturelle, Septieme Serie, Tome VIII: 1 - 110; Plates I - XXXVII. [https: // www. biodiversitylibrary. org / item / 280857 # page / 21 / mode / 1 up]" type="journal article" year="1964">Marnier-Lapostolle (1964: 37</bibRefCitation>
, Planche XI, Figures 29–30, Planche XII, Figure 31); 
<bibRefCitation id="EFA84B00FFB0FF91FA91FAACFF24A531" author="Maire, R." pageId="3" pageNumber="292" pagination="251 - 252" refId="ref7230" refString="Maire, R. (1977) Kalanchoe Adanson (1763) Rosales: Crassulaceae. In: P. [A.] Quezel, Flore de l'Afrique du Nord (Maroc, Algerie, Tunisie, Tripolitaine, Cyrenaique et Sahara). Vol. XIV. Rhoeadales: Cruciferae fin Resedaceae. Sarraceniales: Droseraceae. Rosales: Crassulaceae. Editions Paul Lechevalier editeur S. A. R. L., 19, rue Augereau, 19, Paris, pp. 239 - 241, 251 - 252, 263. [https: // archive. org / details / flore-afrique-du-nord / Vol % 2014 % 20 Flore % 20 de % 20 l % 27 Afrique % 20 Du % 20 Nord % 20 Maroc % 20 Algerie % 20 Tunisie % 20 Tripolitaine % 20 Cyrenaique % 20 Et % 20 Sahara % 20 R-Maire- 1977 / page / n 119 / mode / 2 up]" type="journal article" year="1977">Maire (1977: 253)</bibRefCitation>
; 
<bibRefCitation id="EFA84B00FFB0FF91FF44FA88FE6EA531" author="Jacobsen, H." box="[205,500,1316,1343]" pageId="3" pageNumber="292" refId="ref6810" refString="Jacobsen, H. (1954) Handbuch der sukkulenten Pflanzen. Beschreibung und Kultur der Sukkulenten mit Ausnahme der Cactaceae. Band II. Fockea bis Zygophyllum. [Kalanchoe on pp. 810 - 861.] VEB Gustav Fischer Verlag, Jena, pp. 615 - 1124." type="book" year="1954">Jacobsen (1954: 853–854</bibRefCitation>
, Abb. 357, 796); 
<bibRefCitation id="EFA84B00FFB0FF91FD4BFA88FC2BA530" author="Jacobsen, H." box="[706,945,1316,1342]" pageId="3" pageNumber="292" refId="ref6862" refString="Jacobsen, H. (1977) Lexicon of succulent plants. Short descriptions, habitats and synonymy of succulent plants other than Cactaceae. Revised and enlarged 2 nd edn. [Kalanchoe on pp. 281 - 291.] Blandford Press, Poole, Dorset, 682 pp." type="book" year="1977">Jacobsen (1977: 290</bibRefCitation>
, Plate 106, 
<figureCitation id="13022A74FFB0FF91FBB5FA88FB06A530" box="[1084,1180,1316,1342]" captionStart="FIGURE 1" captionStartId="1.[136,235,1649,1673]" captionTargetBox="[163,1424,201,1614]" captionTargetId="figure-15@1.[151,1436,189,1626]" captionTargetPageId="1" captionText="FIGURE 1. A. Portrait of John Gilbert Baker (1834–1920) who published the name Kalanchoe tomentosa in 1882. Reproduced from Journal of Botany, British and Foreign. London, volume 39 (1901). Photograph: Berthold Carl Seemann. Public Domain. http://www.plantillustrations.org/illustration.php?id_illustration=176522. B. Portrait of Reverend Richard Baron (1847–1907) who collected the material on which Baker based the name K. tomentosa. Credit of the photograph is to Fiangonan’ I Jesoa Kristy Eto Madagasikara (FJKM), which is the lineal descendent of the London Missionary Society, Antananarivo, which employed Baron. Permission was granted to Dr Laurence J. Dorr by FJKM to disassemble a framed collection of photographs of 19th century missionaries to obtain individual portraits for use in Dorr (1997). Dr Dorr kindly supplied the image to the author of this paper. C. Kalanchoe tomentosa is a sparsely- to many-leaved, entirely hairy, sparsely branched, small to medium-sized, terrestrial, succulent shrub that can reach a height of about 1 m. Note the hard, stiff aerial roots that often develop on the stem and branches. D. The hairy flowers of K. tomentosa are borne erect to erectly spreading to spreading, and are hairy throughout. Flowers are campanulate to sometimes very slightly urceolate, yellowish brown to greenish and often sometimes purple-infused. The corolla lobes are light to deep purple on a greenish background, and adaxially velvety rather than hairy. Photographs C and D taken by Gideon F. Smith." figureDoi="http://doi.org/10.5281/zenodo.8297143" httpUri="https://zenodo.org/record/8297143/files/figure.png" pageId="3" pageNumber="292">Figure 1</figureCitation>
); 
<bibRefCitation id="EFA84B00FFB0FF91FB31FA88FA3CA530" author="Jacobsen, H." box="[1208,1446,1316,1342]" pageId="3" pageNumber="292" refId="ref6914" refString="Jacobsen, H. (1986) A handbook of succulent plants. Descriptions, synonyms and cultural details for succulents other than Cactaceae. Vol. II. Ficus to Zygophyllum. [Kalanchoe on pp. 603 - 633.] Blandford Press, Poole, Dorset, pp. 465 - 864 of 1380 pp." type="book" year="1986">Jacobsen (1986: 631</bibRefCitation>
, Figures 414 and 894); 
<bibRefCitation id="EFA84B00FFB0FF91FE19FAE4FCCDA56C" author="Boiteau, P. &amp; Allorge-Boiteau, L." box="[400,855,1352,1379]" pageId="3" pageNumber="292" refId="ref6051" refString="Boiteau, P. &amp; Allorge-Boiteau, L. (1995) Kalanchoe (Crassulacees) de Madagascar. Systematique, ecophysiologie et phytochimie. ICSN, CNRS, 91198 Gif-sur-Yvette, Editions Karthala, Paris, 252 pp." type="book" year="1995">Boiteau &amp; Allorge-Boiteau (1995: 166)</bibRefCitation>
; 
<bibRefCitation id="EFA84B00FFB0FF91FCECFAE4FB81A56C" author="Rauh, W." box="[869,1051,1352,1379]" pageId="3" pageNumber="292" refId="ref7716" refString="Rauh, W. (1995) Succulent and xerophytic plants of Madagascar. Vol. 1 [of two volumes]. Strawberry Press, Mill Valley, California, 343 pp." type="book" year="1995">Rauh (1995: 92</bibRefCitation>
, 93, Figure 223, 116, 219, Figures 609–610, 220, Figures 611–617); 
<bibRefCitation id="EFA84B00FFB0FF91FD86FAC0FD61A589" author="Brickell, C." box="[527,763,1388,1415]" pageId="3" pageNumber="292" refId="ref6141" refString="Brickell, C. (Editor-in-Chief) (1998) The Royal Horticultural Society A - Z encyclopedia of garden plants. Corrected reprint. Dorling Kindersley, London, 1080 pp." type="book" year="1998">Brickell (1998: 578)</bibRefCitation>
; 
<bibRefCitation id="EFA84B00FFB0FF91FC83FAC0FC6DA589" author="Brickell, C." box="[778,1015,1388,1415]" pageId="3" pageNumber="292" refId="ref6174" refString="Brickell, C. (Editor-in-Chief) (2003) The Royal Horticultural Society A - Z encyclopedia of garden plants. Vol. 2. K - Z. Revised edition. Dorling Kindersley, London, pp. 593 - 1128." type="book" year="2003">Brickell (2003: 594)</bibRefCitation>
; 
<bibRefCitation id="EFA84B00FFB0FF91FB8CFAC1FAB3A588" author="Bryant, G. &amp; Bryant, K. &amp; Rutherford, B. &amp; Vogan, R." box="[1029,1321,1388,1415]" pageId="3" pageNumber="292" refId="ref6218" refString="Bryant, G., Bryant, K., Rutherford, B. &amp; Vogan, R. (2005) Cacti &amp; Succulents. Kalanchoe. In: Barnard, L., Edwards, A., Etherington, K., Imwold, D., King, E., Malone, M. &amp; Parker, J. (Eds.) The ultimate plant book. New Holland, London, 1024 pp." type="book" year="2005">
Bryant 
<emphasis id="B94DEAE3FFB0FF91FBD2FAC1FB15A588" box="[1115,1167,1388,1414]" italics="true" pageId="3" pageNumber="292">et al</emphasis>
. (2005: 741)
</bibRefCitation>
; 
<bibRefCitation id="EFA84B00FFB0FF91FABEFAC0FE95A5A4" author="Descoings, B." pageId="3" pageNumber="292" pagination="143 - 181" refId="ref6332" refString="Descoings, B. (2003) Kalanchoe. In: Eggli, U. (Ed.) Illustrated handbook of succulent plants: Crassulaceae. Springer Verlag, Berlin, pp. 143 - 181." type="book chapter" year="2003">Descoings (2003: 178)</bibRefCitation>
; 
<bibRefCitation id="EFA84B00FFB0FF91FE93FA3CFDBAA5A4" author="Smith, G. F. &amp; Figueiredo, E. &amp; Van Wyk, A. E." box="[282,544,1424,1451]" pageId="3" pageNumber="292" refId="ref8834" refString="Smith, G. F., Figueiredo, E. &amp; Van Wyk, A. E. (2019) Kalanchoe (Crassulaceae) in southern Africa. Classification, biology, and cultivation. Academic Press, an imprint of Elsevier, London, San Diego, Cambridge (U. S. A.), and Oxford, 328 pp. https: // doi. org / 10.1016 / C 2017 - 0 - 00602 - X" type="book" year="2019">
Smith 
<emphasis id="B94DEAE3FFB0FF91FEEDFA3DFE0FA5A4" box="[356,405,1424,1450]" italics="true" pageId="3" pageNumber="292">et al</emphasis>
. (2019: 299
</bibRefCitation>
, Figures 12.30.1–12.30.10).
</paragraph>
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<subSubSection id="C323657AFFB0FF91FF34FA19FD8BA6A9" pageId="3" pageNumber="292" type="materials_examined">
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<emphasis id="B94DEAE3FFB0FF91FF34FA19FF6BA5C0" box="[189,241,1461,1486]" italics="true" pageId="3" pageNumber="292">Type</emphasis>
:— 
<materialsCitation id="3B513CACFFB0FF91FE9BFA18FAEBA618" collectingDate="1880-07" collectionCode="R, K, P" country="Madagascar" county="Baron" location="Central Madagascar" municipality="Betsileo-land" pageId="3" pageNumber="292" specimenCode="K001040417, P00374199" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F32E7661FFB0FF91FE9BFA18FE7AA5C0" box="[274,480,1460,1486]" name="Madagascar" pageId="3" pageNumber="292">MADAGASCAR</collectingCountry>
. 
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, chiefly in 
<collectingMunicipality id="6BE2AC8BFFB0FF91FCC8FA18FC43A5C0" box="[833,985,1460,1486]" pageId="3" pageNumber="292">Betsileo-land</collectingMunicipality>
, recd. 
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<collectingDate id="EFC3E9D9FFB0FF91FB96FA18FB17A5C0" box="[1055,1165,1460,1486]" pageId="3" pageNumber="292" value="1880-07">July 1880</collectingDate>
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, [
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<collectionCode id="ED28AE34FFB0FF91FB29FA19FB2BA5C0" box="[1184,1201,1461,1486]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="292">R</collectionCode>
</emphasis>
.] 
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<collectingCounty id="62E74E7DFFB0FF91FB4EFA19FA96A5C0" box="[1223,1292,1461,1486]" pageId="3" pageNumber="292">Baron</collectingCounty>
247
</emphasis>
(
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<collectionCode id="ED28AE34FFB0FF91FF01FA75FF06A5FD" box="[136,156,1497,1523]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="3" pageNumber="292" type="Herbarium">K</collectionCode>
barcode 
<specimenCode id="DB9F9E8AFFB0FF91FE8CFA74FE05A5FC" box="[261,415,1496,1522]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="3" pageNumber="292" type="Herbarium">K001040417</specimenCode>
!, [image available at http://specimens.kew.org/herbarium/ 
<specimenCode id="DB9F9E8AFFB0FF91FBAEFA74FB20A5FC" box="[1063,1210,1496,1522]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="3" pageNumber="292" type="Herbarium">K001040417</specimenCode>
; not yet online] (
<figureCitation id="13022A74FFB0FF91FAF7FA74FF02A618" captionStart="FIGURE 2" captionStartId="4.[136,235,1923,1947]" captionTargetBox="[152,1435,193,1899]" captionTargetId="figure-18@4.[151,1435,189,1899]" captionTargetPageId="4" captionText="FIGURE 2. Holotype, [R.]Baron 247, K K001040417 [http://specimens.kew.org/herbarium/K001040417; not yet available online], of the name Kalanchoe tomentosa. “Central Madagascar, chiefly in Betsileo-land, recd. July 1880, [R.] Baron 247. Reproduced with the kind permission of the Director and the Board of Trustees, Royal Botanic Gardens, Kew." figureDoi="http://doi.org/10.5281/zenodo.8297145" httpUri="https://zenodo.org/record/8297145/files/figure.png" pageId="3" pageNumber="292">Fig. 2</figureCitation>
); isotype 
<collectionCode id="ED28AE34FFB0FF91FE8FFA51FE8FA619" box="[262,277,1533,1559]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="292" type="Herbarium">P</collectionCode>
barcode 
<specimenCode id="DB9F9E8AFFB0FF91FEF2FA50FD9BA618" box="[379,513,1532,1558]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="3" pageNumber="292" type="Herbarium">P00374199</specimenCode>
! [image available at http://coldb.mnhn.fr/catalognumber/mnhn/p/p00374199])
</materialsCitation>
.
</paragraph>
<paragraph id="8B8636F1FFB0FF91FF34F98DFAD6A651" blockId="3.[136,1452,1244,2063]" pageId="3" pageNumber="292">
<emphasis id="B94DEAE3FFB0FF91FF34F98DFE89A634" box="[189,275,1569,1594]" italics="true" pageId="3" pageNumber="292">Epitype</emphasis>
:— 
<materialsCitation id="3B513CACFFB0FF91FEBAF98CFADFA650" collectingDate="1885-11" collectionCode="R, K" country="Madagascar" location="Central Madagascar. Received" pageId="3" pageNumber="292" specimenCode="K000232863" specimenCount="1">
<collectingCountry id="F32E7661FFB0FF91FEBAF98CFD9BA634" box="[307,513,1568,1594]" name="Madagascar" pageId="3" pageNumber="292">MADAGASCAR</collectingCountry>
. 
<location id="8EE6602AFFB0FF91FD80F98CFCC2A634" LSID="urn:lsid:plazi:treatment:039087E7FFB0FF95FF01FB38FF59A266:8EE6602AFFB0FF91FD80F98CFCC2A634" box="[521,856,1568,1594]" country="Madagascar" name="Central Madagascar. Received" pageId="3" pageNumber="292">Central Madagascar. Received</location>
<date id="FF871031FFB0FF91FCD5F98DFC4AA635" box="[860,976,1568,1595]" pageId="3" pageNumber="292" value="1885-11">
<collectingDate id="EFC3E9D9FFB0FF91FCD5F98DFC4AA635" box="[860,976,1568,1595]" pageId="3" pageNumber="292" value="1885-11">Nov.1885</collectingDate>
</date>
. [
<emphasis id="B94DEAE3FFB0FF91FC68F98DFC68A634" box="[993,1010,1569,1594]" italics="true" pageId="3" pageNumber="292">
<collectionCode id="ED28AE34FFB0FF91FC68F98DFC68A634" box="[993,1010,1569,1594]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="292">R</collectionCode>
</emphasis>
.] 
<emphasis id="B94DEAE3FFB0FF91FB8FF98DFB1DA634" box="[1030,1159,1568,1594]" italics="true" pageId="3" pageNumber="292">Baron 3560</emphasis>
(
<collectionCode id="ED28AE34FFB0FF91FB1AF98CFB33A634" box="[1171,1193,1568,1594]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="3" pageNumber="292" type="Herbarium">K</collectionCode>
barcode 
<specimenCode id="DB9F9E8AFFB0FF91FA80F98CFA3EA634" box="[1289,1444,1568,1594]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="3" pageNumber="292" type="Herbarium">K000232863</specimenCode>
!, [image available at http://specimens.kew.org/herbarium/ 
<specimenCode id="DB9F9E8AFFB0FF91FC89F9E8FC0FA650" box="[768,917,1604,1630]" collectionCode="K" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="3" pageNumber="292" type="Herbarium">K000232863</specimenCode>
]) (
<figureCitation id="13022A74FFB0FF91FC3BF9E8FC6FA650" box="[946,1013,1604,1630]" captionStart="FIGURE 3" captionStartId="5.[136,235,1923,1947]" captionTargetBox="[179,1425,202,1889]" captionTargetId="figure-15@5.[151,1435,189,1899]" captionTargetPageId="5" captionText="FIGURE 3. Epitype, [R.] Baron 3560, K K000232863 [http://specimens.kew.org/herbarium/K000232863], designated in this paper, in support of the holotype of the name Kalanchoe tomentosa. “Central Madagascar. Received Nov. 1885.” Reproduced with the kind permission of the Director and the Board of Trustees, Royal Botanic Gardens, Kew." figureDoi="http://doi.org/10.5281/zenodo.8297147" httpUri="https://zenodo.org/record/8297147/files/figure.png" pageId="3" pageNumber="292">Fig. 3</figureCitation>
), 
<emphasis id="B94DEAE3FFB0FF91FB85F9E8FADFA650" bold="true" box="[1036,1349,1604,1630]" pageId="3" pageNumber="292">here designated as epitype</emphasis>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8636F1FFB0FF91FF34F9C5FD8BA6A9" blockId="3.[136,1452,1244,2063]" pageId="3" pageNumber="292">
<emphasis id="B94DEAE3FFB0FF91FF34F9C5FE28A68C" box="[189,434,1641,1666]" italics="true" pageId="3" pageNumber="292">Nom. utique rej. prop</emphasis>
.:— 
<taxonomicName id="4C394D72FFB0FF91FE54F9C4FC45A68C" authority="Blanco (1845: 221)" authorityName="Blanco" authorityPageNumber="221" authorityYear="1845" box="[477,991,1640,1667]" class="Magnoliopsida" family="Crassulaceae" genus="Bryophyllum" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="292" phylum="Tracheophyta" rank="species" species="triangulare">
<emphasis id="B94DEAE3FFB0FF91FE54F9C4FD6FA68C" box="[477,757,1640,1666]" italics="true" pageId="3" pageNumber="292">Bryophyllum triangulare</emphasis>
<bibRefCitation id="EFA84B00FFB0FF91FD77F9C4FC45A68C" author="Blanco, F. M." box="[766,991,1640,1667]" pageId="3" pageNumber="292" refId="ref5984" refString="Blanco, F. M. (1845) Flora de Filipinas, segun el sistema sexual de Linneo: [...] segunda impresion, corregida y aumentada por el mismo autor. D. Miguel Sanchez, Manila, 619 pp. [https: // www. biodiversitylibrary. org / page / 32925947 # page / 282 / mode / 1 up]" type="book" year="1845">Blanco (1845: 221)</bibRefCitation>
</taxonomicName>
. (So proposed by Smith 2023: 435–436, under 
<bibRefCitation id="EFA84B00FFB0FF91FF47F920FE32A6A8" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[206,424,1676,1703]" pageId="3" pageNumber="292" refId="ref9375" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Koeltz Botanical Books, Glash ¸ tten. [Regnum Vegetabile 159], 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland 
<emphasis id="B94DEAE3FFB0FF91FEA4F921FEC4A6A8" box="[301,350,1676,1702]" italics="true" pageId="3" pageNumber="292">et al</emphasis>
. 2018
</bibRefCitation>
: Art. 56).
</paragraph>
</subSubSection>
<subSubSection id="C323657AFFB0FF94FF34F91CFB0AA63A" lastPageId="6" lastPageNumber="295" pageId="3" pageNumber="292" type="discussion">
<paragraph id="8B8636F1FFB0FF91FF34F91CFBE5A800" blockId="3.[136,1452,1244,2063]" pageId="3" pageNumber="292">
<bibRefCitation id="EFA84B00FFB0FF91FF34F91CFE0CA6C4" author="Blanco, F. M." box="[189,406,1712,1739]" pageId="3" pageNumber="292" refId="ref5984" refString="Blanco, F. M. (1845) Flora de Filipinas, segun el sistema sexual de Linneo: [...] segunda impresion, corregida y aumentada por el mismo autor. D. Miguel Sanchez, Manila, 619 pp. [https: // www. biodiversitylibrary. org / page / 32925947 # page / 282 / mode / 1 up]" type="book" year="1845">Blanco (1845: 221)</bibRefCitation>
provided the following description for what is today known as 
<taxonomicName id="4C394D72FFB0FF91FBD1F91DFB71A6C4" authorityName="Baker" authorityYear="1882" box="[1112,1259,1713,1738]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="292" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="B94DEAE3FFB0FF91FBD1F91DFB71A6C4" box="[1112,1259,1713,1738]" italics="true" pageId="3" pageNumber="292">K. tomentosa</emphasis>
</taxonomicName>
: “ 
<taxonomicName id="4C394D72FFB0FF91FA8FF91CFEBFA6E0" authorityName="Blanco" authorityYear="1845" class="Magnoliopsida" family="Crassulaceae" genus="Bryophyllum" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="292" phylum="Tracheophyta" rank="species" species="triangulare">BRYOPHYLLUM TRIANGULARE</taxonomicName>
. BRIOFILO TRIANGULAR. Hojas sesiles, opuestas, muy carnosas, lanceoladas, y de tres ángulos como de figura de bayoneta, enterisimas. Cal. como en la especie anterior. Cor. muy larga, con cuatro ángulos, y el limbo con cuatro lacinias. Éstam. ocho: los cuatro, mas altos que los otros. Gérmen con cuatro lóbulos. Estigmas globosos. Nect. cuatro glándulas en la base del gérmen. Espec. nueva.” [English: “ 
<taxonomicName id="4C394D72FFB0FF91FCFAF8ECFB10A754" authorityName="Blanco" authorityYear="1845" box="[883,1162,1856,1882]" class="Magnoliopsida" family="Crassulaceae" genus="Bryophyllum" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="292" phylum="Tracheophyta" rank="species" species="triangulare">
<emphasis id="B94DEAE3FFB0FF91FCFAF8ECFB10A754" box="[883,1162,1856,1882]" italics="true" pageId="3" pageNumber="292">Bryophyllum triangulare</emphasis>
</taxonomicName>
. Triangular bryophyllum. 
<emphasis id="B94DEAE3FFB0FF91FF01F8C9FF4CA770" box="[136,214,1893,1918]" italics="true" pageId="3" pageNumber="292">Leaves</emphasis>
sessile, opposite, very fleshy, lanceolate, with three angles like the shape of a bayonet, entire. 
<emphasis id="B94DEAE3FFB0FF91FAA0F8C8FAF0A770" box="[1321,1386,1892,1918]" italics="true" pageId="3" pageNumber="292">Calyx</emphasis>
as in preceding species [
<taxonomicName id="4C394D72FFB0FF91FEEEF824FCCDA7AC" authority="Blanco (1845: 220)" authorityName="Blanco" authorityPageNumber="220" authorityYear="1845" box="[359,855,1928,1955]" class="Magnoliopsida" family="Crassulaceae" genus="Bryophyllum" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="292" phylum="Tracheophyta" rank="species" species="serratum">
<emphasis id="B94DEAE3FFB0FF91FEEEF824FDFDA7AC" box="[359,615,1928,1954]" italics="true" pageId="3" pageNumber="292">Bryophyllum serratum</emphasis>
<bibRefCitation id="EFA84B00FFB0FF91FDFBF824FCCDA7AC" author="Blanco, F. M." box="[626,855,1928,1955]" pageId="3" pageNumber="292" refId="ref5984" refString="Blanco, F. M. (1845) Flora de Filipinas, segun el sistema sexual de Linneo: [...] segunda impresion, corregida y aumentada por el mismo autor. D. Miguel Sanchez, Manila, 619 pp. [https: // www. biodiversitylibrary. org / page / 32925947 # page / 282 / mode / 1 up]" type="book" year="1845">Blanco (1845: 220)</bibRefCitation>
</taxonomicName>
, where the calyx is described as: “Cal. muy corto, hendido muy profundamente en cuatro partes.” [English: Calyx very short, very deeply divided into four parts.]]. Corolla very long, with four angles, the limb divided into four lobes. Stamens eight, four longer than the others. Ovary with four lobes. Stigmas globose. Nectar glands four at the base of the ovary. New spec.”].
</paragraph>
<caption id="DF466679FFB7FF96FF01F82FFBFFA7EC" ID-DOI="http://doi.org/10.5281/zenodo.8297145" ID-Zenodo-Dep="8297145" httpUri="https://zenodo.org/record/8297145/files/figure.png" pageId="4" pageNumber="293" startId="4.[136,235,1923,1947]" targetBox="[152,1435,193,1899]" targetPageId="4" targetType="figure">
<paragraph id="8B8636F1FFB7FF96FF01F82FFBFFA7EC" blockId="4.[136,1451,1923,2019]" pageId="4" pageNumber="293">
<emphasis id="B94DEAE3FFB7FF96FF01F82FFE98A795" bold="true" box="[136,258,1923,1947]" pageId="4" pageNumber="293">FIGURE 2.</emphasis>
Holotype, [
<emphasis id="B94DEAE3FFB7FF96FEFEF828FE1CA795" box="[375,390,1924,1947]" italics="true" pageId="4" pageNumber="293">R</emphasis>
.] 
<emphasis id="B94DEAE3FFB7FF96FE10F828FD98A795" box="[409,514,1923,1947]" italics="true" pageId="4" pageNumber="293">Baron 247</emphasis>
, K K001040417 [http://specimens.kew.org/herbarium/K001040417; not yet available online], of the name 
<taxonomicName id="4C394D72FFB7FF96FE82F80BFE72A7B1" authorityName="Baker" authorityYear="1882" box="[267,488,1959,1983]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="293" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="B94DEAE3FFB7FF96FE82F80BFE72A7B1" box="[267,488,1959,1983]" italics="true" pageId="4" pageNumber="293">Kalanchoe tomentosa</emphasis>
</taxonomicName>
. “Central Madagascar, chiefly in Betsileo-land, recd. July 1880, [
<emphasis id="B94DEAE3FFB7FF96FB19F804FB05A7B1" box="[1168,1183,1960,1983]" italics="true" pageId="4" pageNumber="293">R</emphasis>
.] 
<emphasis id="B94DEAE3FFB7FF96FB3FF804FAB9A7B1" box="[1206,1315,1959,1983]" italics="true" pageId="4" pageNumber="293">Baron 247</emphasis>
. Reproduced with the kind permission of the Director and the Board of Trustees, Royal Botanic Gardens, Kew.
</paragraph>
</caption>
<caption id="DF466679FFB6FF97FF01F82FFBABA7EC" ID-DOI="http://doi.org/10.5281/zenodo.8297147" ID-Zenodo-Dep="8297147" httpUri="https://zenodo.org/record/8297147/files/figure.png" pageId="5" pageNumber="294" startId="5.[136,235,1923,1947]" targetBox="[179,1425,202,1889]" targetPageId="5" targetType="figure">
<paragraph id="8B8636F1FFB6FF97FF01F82FFBABA7EC" blockId="5.[136,1452,1923,2019]" pageId="5" pageNumber="294">
<emphasis id="B94DEAE3FFB6FF97FF01F82FFE92A795" bold="true" box="[136,264,1923,1947]" pageId="5" pageNumber="294">FIGURE 3.</emphasis>
Epitype, [
<emphasis id="B94DEAE3FFB6FF97FEF3F828FE13A795" box="[378,393,1924,1947]" italics="true" pageId="5" pageNumber="294">R</emphasis>
.] 
<emphasis id="B94DEAE3FFB6FF97FE2AF828FD84A795" box="[419,542,1923,1947]" italics="true" pageId="5" pageNumber="294">Baron 3560</emphasis>
, K K000232863 [http://specimens.kew.org/herbarium/K000232863], designated in this paper, in support of the holotype of the name 
<taxonomicName id="4C394D72FFB6FF97FDD8F80BFCB7A7B1" authorityName="Baker" authorityYear="1882" box="[593,813,1959,1983]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="5" pageNumber="294" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="B94DEAE3FFB6FF97FDD8F80BFCB7A7B1" box="[593,813,1959,1983]" italics="true" pageId="5" pageNumber="294">Kalanchoe tomentosa</emphasis>
</taxonomicName>
. “Central Madagascar. Received Nov. 1885.” Reproduced with the kind permission of the Director and the Board of Trustees, Royal Botanic Gardens, Kew.
</paragraph>
</caption>
<paragraph id="8B8636F1FFB5FF94FF34FF32FC96A12B" blockId="6.[136,1452,158,2093]" pageId="6" pageNumber="295">
The name 
<taxonomicName id="4C394D72FFB5FF94FEBFFF33FE4CA0B6" authorityName="Blanco" authorityYear="1845" box="[310,470,158,184]" class="Magnoliopsida" family="Crassulaceae" genus="Bryophyllum" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="295" phylum="Tracheophyta" rank="species" species="triangulare">
<emphasis id="B94DEAE3FFB5FF94FEBFFF33FE4CA0B6" box="[310,470,158,184]" italics="true" pageId="6" pageNumber="295">B. triangulare</emphasis>
</taxonomicName>
, which applies to material at present known as 
<taxonomicName id="4C394D72FFB5FF94FC7CFF33FB10A0B6" authorityName="Baker" authorityYear="1882" box="[1013,1162,159,184]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="295" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="B94DEAE3FFB5FF94FC7CFF33FB10A0B6" box="[1013,1162,159,184]" italics="true" pageId="6" pageNumber="295">K. tomentosa</emphasis>
</taxonomicName>
, predates the latter by 37 years and, under 
<bibRefCitation id="EFA84B00FFB5FF94FEC0FF6EFDB4A0D2" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[329,558,194,221]" pageId="6" pageNumber="295" refId="ref9375" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Koeltz Botanical Books, Glash ¸ tten. [Regnum Vegetabile 159], 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland 
<emphasis id="B94DEAE3FFB5FF94FE21FF6FFE40A0D2" box="[424,474,194,220]" italics="true" pageId="6" pageNumber="295">et al</emphasis>
. (2018
</bibRefCitation>
: Art. 11.4), the epithet ‘ 
<taxonomicName id="4C394D72FFB5FF94FCB2FF6EFC20A0D2" authorityName="Blanco" authorityYear="1845" box="[827,954,194,220]" class="Magnoliopsida" family="Crassulaceae" genus="Bryophyllum" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="295" phylum="Tracheophyta" rank="species" species="triangulare">
<emphasis id="B94DEAE3FFB5FF94FCB2FF6EFC20A0D2" box="[827,954,194,220]" italics="true" pageId="6" pageNumber="295">triangulare</emphasis>
</taxonomicName>
’ should have been adopted by 
<bibRefCitation id="EFA84B00FFB5FF94FA91FF6EFF20A10E" author="Baker, J. G." pageId="6" pageNumber="295" pagination="109 - 114" refId="ref5465" refString="Baker, J. G. (1882) Contributions to the flora of central Madagascar (continued from p. 70) and (to be continued). Journal of Botany, British and Foreign. London 20: 109 - 114. https: // doi. org / 10.1111 / j. 1095 - 8339.1884. tb 00548. x" type="journal article" year="1882">Baker (1882: 110)</bibRefCitation>
when the species is included in 
<taxonomicName id="4C394D72FFB5FF94FDA6FF4AFD32A10E" authorityName="Adanson" authorityYear="1763" box="[559,680,230,256]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="295" phylum="Tracheophyta" rank="genus">
<emphasis id="B94DEAE3FFB5FF94FDA6FF4AFD32A10E" box="[559,680,230,256]" italics="true" pageId="6" pageNumber="295">Kalanchoe</emphasis>
</taxonomicName>
. However, the name 
<taxonomicName id="4C394D72FFB5FF94FC17FF4BFBA7A10E" authorityName="Blanco" authorityYear="1845" box="[926,1085,230,256]" class="Magnoliopsida" family="Crassulaceae" genus="Bryophyllum" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="295" phylum="Tracheophyta" rank="species" species="triangulare">
<emphasis id="B94DEAE3FFB5FF94FC17FF4BFBA7A10E" box="[926,1085,230,256]" italics="true" pageId="6" pageNumber="295">B. triangulare</emphasis>
</taxonomicName>
has been proposed for rejection (
<bibRefCitation id="EFA84B00FFB5FF94FF06FEA6FEF1A12A" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[143,363,266,293]" pageId="6" pageNumber="295" refId="ref9375" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Koeltz Botanical Books, Glash ¸ tten. [Regnum Vegetabile 159], 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland 
<emphasis id="B94DEAE3FFB5FF94FF79FEA7FEBBA12A" box="[240,289,266,292]" italics="true" pageId="6" pageNumber="295">et al</emphasis>
. 2018
</bibRefCitation>
: Art. 56) (see Smith 2023: 435–436).
</paragraph>
<paragraph id="8B8636F1FFB5FF94FF34FE82FED8A24B" blockId="6.[136,1452,158,2093]" pageId="6" pageNumber="295">
Earlier, 
<bibRefCitation id="EFA84B00FFB5FF94FE91FE82FDB6A146" author="Blanco, F. M." box="[280,556,302,328]" pageId="6" pageNumber="295" refId="ref5925" refString="Blanco, F. M. (1837) Flora de Filipinas, segun el sistema sexual de Linneo: [...]. En la imprenta de Sto. Thomas [Tomas] por D. Candido Lopez, Manila, 887 pp. https: // doi. org / 10.5962 / bhl. title. 120031" type="book" year="1837">Blanco (1837: 382–383)</bibRefCitation>
provided exactly the same description for 
<taxonomicName id="4C394D72FFB5FF94FB98FE82FF24A162" authority="Blanco (1837: 382)" authorityName="Blanco" authorityPageNumber="382" authorityYear="1837" class="Magnoliopsida" family="Crassulaceae" genus="Cotyledon" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="295" phylum="Tracheophyta" rank="species" species="lanceolata">
<emphasis id="B94DEAE3FFB5FF94FB98FE82FA99A146" box="[1041,1283,302,328]" italics="true" pageId="6" pageNumber="295">Cotyledon lanceolata</emphasis>
<bibRefCitation id="EFA84B00FFB5FF94FA82FE82FF24A162" author="Blanco, F. M." pageId="6" pageNumber="295" refId="ref5925" refString="Blanco, F. M. (1837) Flora de Filipinas, segun el sistema sexual de Linneo: [...]. En la imprenta de Sto. Thomas [Tomas] por D. Candido Lopez, Manila, 887 pp. https: // doi. org / 10.5962 / bhl. title. 120031" type="book" year="1837">Blanco (1837: 382)</bibRefCitation>
</taxonomicName>
, with the sole exception that “otros” was given as “otro” in the 1837 work. In 
<bibRefCitation id="EFA84B00FFB5FF94FBBBFEFEFA97A162" author="Blanco, F. M." box="[1074,1293,338,364]" pageId="6" pageNumber="295" refId="ref5925" refString="Blanco, F. M. (1837) Flora de Filipinas, segun el sistema sexual de Linneo: [...]. En la imprenta de Sto. Thomas [Tomas] por D. Candido Lopez, Manila, 887 pp. https: // doi. org / 10.5962 / bhl. title. 120031" type="book" year="1837">Blanco (1837: 383)</bibRefCitation>
the following note was provided: “ 
<emphasis id="B94DEAE3FFB5FF94FEF8FEDBFE3DA19E" box="[369,423,375,400]" italics="true" pageId="6" pageNumber="295">Nota</emphasis>
. Difiere de aquella especie, en que no tiene aserraduras en el apice de la hoja. He visto toda via otra especie con las hojas ternadas y carnosas.” [English: “Note. It differs from that species [
<taxonomicName id="4C394D72FFB5FF94FB57FE36FEF2A1D6" authority="Blanco (1837: 382)" authorityName="Blanco" authorityPageNumber="382" authorityYear="1837" class="Magnoliopsida" family="Crassulaceae" genus="Cotyledon" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="295" phylum="Tracheophyta" rank="species" species="serrata">
<emphasis id="B94DEAE3FFB5FF94FB57FE36FA31A1BA" box="[1246,1451,410,436]" italics="true" pageId="6" pageNumber="295">Cotyledon serrata</emphasis>
<bibRefCitation id="EFA84B00FFB5FF94FF01FE12FEF2A1D6" author="Blanco, F. M." box="[136,360,446,472]" pageId="6" pageNumber="295" refId="ref5925" refString="Blanco, F. M. (1837) Flora de Filipinas, segun el sistema sexual de Linneo: [...]. En la imprenta de Sto. Thomas [Tomas] por D. Candido Lopez, Manila, 887 pp. https: // doi. org / 10.5962 / bhl. title. 120031" type="book" year="1837">Blanco (1837: 382)</bibRefCitation>
</taxonomicName>
] in that it does not have serrations at the apex of the leaf. 
<collectionCode id="ED28AE34FFB5FF94FB85FE13FB8FA1D7" box="[1036,1045,447,473]" country="Romania" lsid="urn:lsid:biocol.org:col:14415" name="&amp;quot;Alexandru Ioan Cuza&amp;quot; University" pageId="6" pageNumber="295" type="Herbarium">I</collectionCode>
have seen yet another species with ternate and fleshy leaves.”] However, the name 
<taxonomicName id="4C394D72FFB5FF94FD2BFE4EFC09A1F2" authorityName="Blanco" authorityYear="1837" box="[674,915,482,508]" class="Magnoliopsida" family="Crassulaceae" genus="Cotyledon" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="295" phylum="Tracheophyta" rank="species" species="lanceolata">
<emphasis id="B94DEAE3FFB5FF94FD2BFE4EFC09A1F2" box="[674,915,482,508]" italics="true" pageId="6" pageNumber="295">Cotyledon lanceolata</emphasis>
</taxonomicName>
published by 
<bibRefCitation id="EFA84B00FFB5FF94FBBBFE4EFA96A1F2" author="Blanco, F. M." box="[1074,1292,482,508]" pageId="6" pageNumber="295" refId="ref5925" refString="Blanco, F. M. (1837) Flora de Filipinas, segun el sistema sexual de Linneo: [...]. En la imprenta de Sto. Thomas [Tomas] por D. Candido Lopez, Manila, 887 pp. https: // doi. org / 10.5962 / bhl. title. 120031" type="book" year="1837">Blanco (1837: 382)</bibRefCitation>
was preceded by 
<taxonomicName id="4C394D72FFB5FF94FF22FDABFE6FA22F" authority="Forsskal (1775" authorityName="Forsskal" authorityYear="1775" box="[171,501,518,545]" class="Magnoliopsida" family="Crassulaceae" genus="Cotyledon" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="295" phylum="Tracheophyta" rank="species" species="lanceolata">
<emphasis id="B94DEAE3FFB5FF94FF22FDABFED8A22E" box="[171,322,518,545]" italics="true" pageId="6" pageNumber="295">C. lanceolata</emphasis>
<bibRefCitation id="EFA84B00FFB5FF94FEC0FDAAFE6FA22F" author="Forsskal, P." box="[329,501,518,545]" pageId="6" pageNumber="295" refId="ref6599" refString="Forsskal, P. (1775) Flora AEgyptiaco-Arabica. Sive descriptiones plantarum, Quas per AEgyptum inferiorem et Arabiam felicem detexit […]. Ex officina M ˆ lleri, aulae Typographi. Prostat apud Heineck et Faber, Hauniae, 220 pp. https: // doi. org / 10.5962 / bhl. title. 6625" type="book" year="1775">Forsskål (1775</bibRefCitation>
</taxonomicName>
: CXI &amp; 89) and the former is therefore an illegitimate later homonym (
<bibRefCitation id="EFA84B00FFB5FF94FA9AFDAAFF5EA24A" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. - H. &amp; Li, D. - Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." pageId="6" pageNumber="295" refId="ref9375" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. - H., Li, D. - Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Koeltz Botanical Books, Glash ¸ tten. [Regnum Vegetabile 159], 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland 
<emphasis id="B94DEAE3FFB5FF94FAFAFDABFA3FA22E" box="[1395,1445,518,544]" italics="true" pageId="6" pageNumber="295">et al</emphasis>
. 2018
</bibRefCitation>
: Art. 53.1).
</paragraph>
<paragraph id="8B8636F1FFB5FF94FF34FDE2FCF0A2BE" blockId="6.[136,1452,158,2093]" pageId="6" pageNumber="295">
<bibRefCitation id="EFA84B00FFB5FF94FF34FDE2FEC6A267" author="Merrill, E. D." box="[189,348,590,617]" pageId="6" pageNumber="295" refId="ref7396" refString="Merrill, E. D. (1905) A review of the identifications of the species described in Blanco's Flora Filipinas. Department of the Interior, Bureau of Government Laboratories, Bureau of Public Printing, Manila, 132 pp. https: // doi. org / 10.5962 / bhl. title. 58554" type="book" year="1905">Merrill (1905</bibRefCitation>
, 
<bibRefCitation id="EFA84B00FFB5FF94FEE7FDE2FE33A266" author="Merrill, E. D." box="[366,425,590,616]" pageId="6" pageNumber="295" refId="ref7453" refString="Merrill, E. D. (1918 [15 June]) Species blancoanae. A critical revision of the Philippine species of plants described by Blanco and by Llanos. Department of Agriculture and Natural Resources, Bureau of Science, Publication No. 12. Bureau of Printing, Manila, 423 pp. https: // doi. org / 10.5962 / bhl. title. 2116" type="book" year="1918">1918</bibRefCitation>
, 
<bibRefCitation id="EFA84B00FFB5FF94FE30FDE2FE68A266" author="Merrill, E. D." box="[441,498,590,616]" pageId="6" pageNumber="295" refId="ref7524" refString="Merrill, E. D. (1923 [2 June]) An enumeration of Philippine flowering plants. Vol. 2. Department of Agriculture and Natural Resources, Bureau of Science, Publication No. 18. Vol. 2. Bureau of Printing, Manila, 530 pp. [https: // www. biodiversitylibrary. org / item / 104550 # page / 223 / mode / 1 up]" type="book" year="1923">1923</bibRefCitation>
) provide additional information on the identification of the species described in 
<bibRefCitation id="EFA84B00FFB5FF94FF01FDDEFEB9A282" author="Blanco, F. M." box="[136,291,626,652]" pageId="6" pageNumber="295" refId="ref5925" refString="Blanco, F. M. (1837) Flora de Filipinas, segun el sistema sexual de Linneo: [...]. En la imprenta de Sto. Thomas [Tomas] por D. Candido Lopez, Manila, 887 pp. https: // doi. org / 10.5962 / bhl. title. 120031" type="book" year="1837">Blanco (1837</bibRefCitation>
, 
<bibRefCitation id="EFA84B00FFB5FF94FEA7FDDEFEFDA283" author="Blanco, F. M." box="[302,359,626,653]" pageId="6" pageNumber="295" refId="ref5984" refString="Blanco, F. M. (1845) Flora de Filipinas, segun el sistema sexual de Linneo: [...] segunda impresion, corregida y aumentada por el mismo autor. D. Miguel Sanchez, Manila, 619 pp. [https: // www. biodiversitylibrary. org / page / 32925947 # page / 282 / mode / 1 up]" type="book" year="1845">1845</bibRefCitation>
) as well as in the further work based on Blanco’s 
<emphasis id="B94DEAE3FFB5FF94FC11FDDEFBFFA282" box="[920,1125,626,652]" italics="true" pageId="6" pageNumber="295">
Flora de 
<collectingCountry id="F32E7661FFB5FF94FB89FDDEFBFFA282" box="[1024,1125,626,652]" name="Philippines" pageId="6" pageNumber="295">Filipinas</collectingCountry>
</emphasis>
published under the name of 
<bibRefCitation id="EFA84B00FFB5FF94FF01FD3AFE03A2BE" author="Fernandez-Villar, C." box="[136,409,662,688]" pageId="6" pageNumber="295" refId="ref6468" refString="Fernandez-Villar, C. (1880) Flora de Filipinas por el P. Fr. Manuel Blanco agustino calzado adicionada con el manuscrito inedito del P. Fr. Ignacio Mercado las obras del P. Fr. Antonio Llanos y de un apendice con todas las nuevas investigaciones botanicas referentes al archipielago Filipino. Tomo Cuarto. Gran Edicion hecha a expensas de la provincia de agustinos calzados de Filipinas bajo la direccion cientifica y literaria de los pp. agustinos calzados Fr. Andres Naves. Establecimiento Tipografico, de Plana y C. a, Manila, 375 + I - VI pp. [https: // www. biodiversitylibrary. org / page / 59644890 # page / 290 / mode / 1 up]" type="book" year="1880">Fernández-Villar (1880)</bibRefCitation>
, 35 years after Blanco (1778–1845) died.
</paragraph>
<paragraph id="8B8636F1FFB5FF94FF34FD16FD1BA36A" blockId="6.[136,1452,158,2093]" pageId="6" pageNumber="295">
<emphasis id="B94DEAE3FFB5FF94FF34FD16FE30A2DA" bold="true" box="[189,426,698,724]" pageId="6" pageNumber="295">Nomenclature notes</emphasis>
:—The only collection cited by 
<bibRefCitation id="EFA84B00FFB5FF94FC8CFD16FC54A2DA" author="Baker, J. G." box="[773,974,698,724]" pageId="6" pageNumber="295" pagination="109 - 114" refId="ref5465" refString="Baker, J. G. (1882) Contributions to the flora of central Madagascar (continued from p. 70) and (to be continued). Journal of Botany, British and Foreign. London 20: 109 - 114. https: // doi. org / 10.1111 / j. 1095 - 8339.1884. tb 00548. x" type="journal article" year="1882">Baker (1882: 110)</bibRefCitation>
in the protologue of the name 
<taxonomicName id="4C394D72FFB5FF94FA90FD17FA31A2DA" authorityName="Baker" authorityYear="1882" box="[1305,1451,699,724]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="295" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="B94DEAE3FFB5FF94FA90FD17FA31A2DA" box="[1305,1451,699,724]" italics="true" pageId="6" pageNumber="295">K. tomentosa</emphasis>
</taxonomicName>
is “ 
<emphasis id="B94DEAE3FFB5FF94FF24FD73FF68A2F6" box="[173,242,735,760]" italics="true" pageId="6" pageNumber="295">Baron</emphasis>
247!”, which originated from “Central Madagascar”. The specimen 
<collectionCode id="ED28AE34FFB5FF94FC62FD72FC65A2F6" box="[1003,1023,734,760]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="6" pageNumber="295" type="Herbarium">K</collectionCode>
001040417 [http://specimens.kew.org/ herbarium/ 
<collectionCode id="ED28AE34FFB5FF94FE8FFCAEFE80A312" box="[262,282,770,796]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="6" pageNumber="295" type="Herbarium">K</collectionCode>
001040417; not yet online] held at Herb. 
<collectionCode id="ED28AE34FFB5FF94FD66FCAFFC99A313" box="[751,771,771,797]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="6" pageNumber="295" type="Herbarium">K</collectionCode>
carries a label stating: “ 
<collectingCountry id="F32E7661FFB5FF94FB9FFCAEFB7EA312" box="[1046,1252,770,796]" name="Madagascar" pageId="6" pageNumber="295">MADAGASCAR</collectingCountry>
: chiefly Betsileo-land. Coll. Rev. 
<collectionCode id="ED28AE34FFB5FF94FECAFC8BFEC3A34F" box="[323,345,807,833]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="295">R</collectionCode>
. Baron. Recd. July, 1880.”, and this is the 
<typeStatus id="54828853FFB5FF94FCCCFC8AFC3DA34E" box="[837,935,806,832]" pageId="6" pageNumber="295">holotype</typeStatus>
of the name 
<taxonomicName id="4C394D72FFB5FF94FBC9FC8BFB4CA34E" authorityName="Baker" authorityYear="1882" box="[1088,1238,807,832]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="295" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="B94DEAE3FFB5FF94FBC9FC8BFB4CA34E" box="[1088,1238,807,832]" italics="true" pageId="6" pageNumber="295">K. tomentosa</emphasis>
</taxonomicName>
. It consists of two inflorescences and flower fragments (
<figureCitation id="13022A74FFB5FF94FDA7FCE6FDE9A36A" box="[558,627,842,868]" captionStart="FIGURE 2" captionStartId="4.[136,235,1923,1947]" captionTargetBox="[152,1435,193,1899]" captionTargetId="figure-18@4.[151,1435,189,1899]" captionTargetPageId="4" captionText="FIGURE 2. Holotype, [R.]Baron 247, K K001040417 [http://specimens.kew.org/herbarium/K001040417; not yet available online], of the name Kalanchoe tomentosa. “Central Madagascar, chiefly in Betsileo-land, recd. July 1880, [R.] Baron 247. Reproduced with the kind permission of the Director and the Board of Trustees, Royal Botanic Gardens, Kew." figureDoi="http://doi.org/10.5281/zenodo.8297145" httpUri="https://zenodo.org/record/8297145/files/figure.png" pageId="6" pageNumber="295">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8B8636F1FFB5FF94FF34FCC2FECEA416" blockId="6.[136,1452,158,2093]" pageId="6" pageNumber="295">
Herb. 
<collectionCode id="ED28AE34FFB5FF94FE8DFCC3FE89A387" box="[260,275,879,905]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="6" pageNumber="295" type="Herbarium">P</collectionCode>
holds a specimen labelled “ 
<collectionCode id="ED28AE34FFB5FF94FDC4FCC2FD07A386" box="[589,669,878,904]" pageId="6" pageNumber="295">HERB</collectionCode>
. 
<collectionCode id="ED28AE34FFB5FF94FD21FCC2FD72A386" box="[680,744,878,904]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13677" lsid="urn:lsid:biocol.org:col:13677" name="Muskingum College" pageId="6" pageNumber="295" type="Herbarium">MUS</collectionCode>
. 
<collectionCode id="ED28AE34FFB5FF94FD7AFCC2FCDAA386" box="[755,832,878,904]" pageId="6" pageNumber="295">PARIS</collectionCode>
<taxonomicName id="4C394D72FFB5FF94FCCEFCC2FBECA386" authority="Bak." authorityName="Baker" authorityYear="1882" box="[839,1142,878,905]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="295" phylum="Tracheophyta" rank="species" species="tomentosa">Kalanchoe tomentosa Bak.</taxonomicName>
247 
<location id="8EE6602AFFB5FF94FB27FCC3FB68A387" LSID="urn:lsid:plazi:treatment:039087E7FFB0FF95FF01FB38FF59A266:8EE6602AFFB5FF94FB27FCC3FB68A387" box="[1198,1266,879,905]" country="Madagascar" municipality="The" name="Baron" pageId="6" pageNumber="295">Baron</location>
<collectingCountry id="F32E7661FFB5FF94FB73FCC2FA1CA386" box="[1274,1414,878,904]" name="Madagascar" pageId="6" pageNumber="295">Madagascar</collectingCountry>
.” 
<collectionCode id="ED28AE34FFB5FF94FA10FCC2FA36A386" box="[1433,1452,878,904]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="6" pageNumber="295" type="Herbarium">A</collectionCode>
smaller red-lettered label has “ 
<collectionCode id="ED28AE34FFB5FF94FE56FC3EFDB0A3A2" box="[479,554,914,940]" pageId="6" pageNumber="295">TYPE</collectionCode>
” printed on it, with “iso-” added to this label in ink. 
<collectingMunicipality id="6BE2AC8BFFB5FF94FBF5FC3EFB32A3A2" box="[1148,1192,914,940]" pageId="6" pageNumber="295">The</collectingMunicipality>
specimen consists of a piece of an inflorescence that is enclosed in a small paper capsule attached to the specimen and is undated (see http:// coldb.mnhn.fr/catalognumber/mnhn/p/p00374199, 
<collectionCode id="ED28AE34FFB5FF94FD40FC77FD42A3FB" box="[713,728,987,1013]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="6" pageNumber="295" type="Herbarium">P</collectionCode>
<specimenCode id="DB9F9E8AFFB5FF94FD6BFC76FCFCA3FA" box="[738,870,986,1012]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="6" pageNumber="295" type="Herbarium">P00374199</specimenCode>
). This is a duplicate of the 
<typeStatus id="54828853FFB5FF94FB21FC76FA90A3FA" box="[1192,1290,986,1012]" pageId="6" pageNumber="295">holotype</typeStatus>
and therefore indeed an isotype.
</paragraph>
<paragraph id="8B8636F1FFB5FF94FF34FB8FFB10A51A" blockId="6.[136,1452,158,2093]" pageId="6" pageNumber="295">
<bibRefCitation id="EFA84B00FFB5FF94FF34FB8FFE19A432" author="Hamet, R." box="[189,387,1058,1085]" pageId="6" pageNumber="295" pagination="17 - 48" refId="ref6659" refString="Hamet, R. (1908) Monographie du genre Kalanchoe. Suite et fin. Bulletin de l'Herbier Boissier ser. 2 8: 17 - 48. [https: // www. biodiversitylibrary. org / item / 104945 # page / 31 / mode / 1 up]" type="journal article" year="1908">Hamet (1908: 31)</bibRefCitation>
stated: “[
<collectionCode id="ED28AE34FFB5FF94FE62FB8EFD98A432" box="[491,514,1058,1084]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="295">R</collectionCode>
. Baron, 
<superScript id="7C4C9BB9FFB5FF94FDD7FB8FFDEEA43E" attach="left" box="[606,628,1058,1085]" fontSize="6" pageId="6" pageNumber="295">no</superScript>
3560.—Echantillon authentique!]”, and also cited two further specimens, “[
<collectionCode id="ED28AE34FFB5FF94FF15FBEAFF29A46E" box="[156,179,1094,1120]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="295">R</collectionCode>
. Baron, 
<superScript id="7C4C9BB9FFB5FF94FE90FBEBFEB5A45A" attach="left" box="[281,303,1094,1121]" fontSize="6" pageId="6" pageNumber="295">no</superScript>
247]” and “[D. Cowan!]”. The specimen 
<emphasis id="B94DEAE3FFB5FF94FC97FBEBFC3CA46E" box="[798,934,1094,1120]" italics="true" pageId="6" pageNumber="295">Baron 3560</emphasis>
(
<collectionCode id="ED28AE34FFB5FF94FC30FBEAFC77A46E" box="[953,1005,1094,1120]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="6" pageNumber="295" type="Herbarium">K K</collectionCode>
000232863) from Central 
<collectingCountry id="F32E7661FFB5FF94FA97FBEAFA3DA46E" box="[1310,1447,1094,1120]" name="Madagascar" pageId="6" pageNumber="295">Madagascar</collectingCountry>
, received in 
<date id="FF871031FFB5FF94FE83FBC6FE58A48B" box="[266,450,1130,1157]" pageId="6" pageNumber="295" value="1885-11">November 1885</date>
, is databased and labelled as 
<typeStatus id="54828853FFB5FF94FC8EFBC7FCADA48B" box="[775,823,1131,1157]" pageId="6" pageNumber="295">type</typeStatus>
at Herb. 
<collectionCode id="ED28AE34FFB5FF94FC17FBC7FC2FA48B" box="[926,949,1131,1157]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="6" pageNumber="295" type="Herbarium">K</collectionCode>
, but it is not part of the original material as it was received on a later date. It cannot take precedence over 
<emphasis id="B94DEAE3FFB5FF94FCA2FB23FC5AA4A6" box="[811,960,1166,1192]" italics="true" pageId="6" pageNumber="295">
<collectionCode id="ED28AE34FFB5FF94FCA2FB23FCA5A4A6" box="[811,831,1167,1192]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="295">R</collectionCode>
. Baron 247
</emphasis>
, which is the 
<typeStatus id="54828853FFB5FF94FBD4FB23FB17A4A7" box="[1117,1165,1167,1193]" pageId="6" pageNumber="295">type</typeStatus>
collection. The reference by 
<bibRefCitation id="EFA84B00FFB5FF94FF23FB1FFEEAA4C2" author="Hamet, R." box="[170,368,1202,1229]" pageId="6" pageNumber="295" pagination="17 - 48" refId="ref6659" refString="Hamet, R. (1908) Monographie du genre Kalanchoe. Suite et fin. Bulletin de l'Herbier Boissier ser. 2 8: 17 - 48. [https: // www. biodiversitylibrary. org / item / 104945 # page / 31 / mode / 1 up]" type="journal article" year="1908">Hamet (1908: 31)</bibRefCitation>
to the specimen “ 
<collectionCode id="ED28AE34FFB5FF94FDBEFB1EFDD4A4C2" box="[567,590,1202,1228]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="295">R</collectionCode>
. Baron, 
<superScript id="7C4C9BB9FFB5FF94FD20FB1FFD25A4CE" attach="left" box="[681,703,1202,1229]" fontSize="6" pageId="6" pageNumber="295">no</superScript>
3560” as the “Echantillon authentique!” of the name 
<taxonomicName id="4C394D72FFB5FF94FA91FB1FFA31A4C2" authorityName="Baker" authorityYear="1882" box="[1304,1451,1203,1228]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="295" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="B94DEAE3FFB5FF94FA91FB1FFA31A4C2" box="[1304,1451,1203,1228]" italics="true" pageId="6" pageNumber="295">K. tomentosa</emphasis>
</taxonomicName>
has no nomenclatural standing as these two words are not equivalent to “ 
<typeStatus id="54828853FFB5FF94FC5FFB7AFB96A4FE" box="[982,1036,1238,1264]" pageId="6" pageNumber="295">type</typeStatus>
”; they only refer to a representative specimen and, in this case, additionally, not to the 
<typeStatus id="54828853FFB5FF94FD35FB57FD76A51B" box="[700,748,1275,1301]" pageId="6" pageNumber="295">type</typeStatus>
material cited by 
<bibRefCitation id="EFA84B00FFB5FF94FC3EFB56FB1CA51A" author="Baker, J. G." box="[951,1158,1274,1300]" pageId="6" pageNumber="295" pagination="109 - 114" refId="ref5465" refString="Baker, J. G. (1882) Contributions to the flora of central Madagascar (continued from p. 70) and (to be continued). Journal of Botany, British and Foreign. London 20: 109 - 114. https: // doi. org / 10.1111 / j. 1095 - 8339.1884. tb 00548. x" type="journal article" year="1882">Baker (1882: 110)</bibRefCitation>
.
</paragraph>
<paragraph id="8B8636F1FFB5FF94FF34FAB3FEE5A58E" blockId="6.[136,1452,158,2093]" pageId="6" pageNumber="295">
However, the specimen 
<emphasis id="B94DEAE3FFB5FF94FE45FAB3FDCBA536" box="[460,593,1310,1336]" italics="true" pageId="6" pageNumber="295">Baron 3560</emphasis>
(
<collectionCode id="ED28AE34FFB5FF94FDE9FAB2FDECA536" box="[608,630,1310,1336]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="6" pageNumber="295" type="Herbarium">K</collectionCode>
K000232863) is a considerably more complete specimen and, unlike the 
<typeStatus id="54828853FFB5FF94FF01FAEEFF77A552" box="[136,237,1346,1372]" pageId="6" pageNumber="295">holotype</typeStatus>
, consists of three leafy terminal stem segments, as well as two inflorescences (
<figureCitation id="13022A74FFB5FF94FBD6FAEEFB39A552" box="[1119,1187,1346,1372]" captionStart="FIGURE 3" captionStartId="5.[136,235,1923,1947]" captionTargetBox="[179,1425,202,1889]" captionTargetId="figure-15@5.[151,1435,189,1899]" captionTargetPageId="5" captionText="FIGURE 3. Epitype, [R.] Baron 3560, K K000232863 [http://specimens.kew.org/herbarium/K000232863], designated in this paper, in support of the holotype of the name Kalanchoe tomentosa. “Central Madagascar. Received Nov. 1885.” Reproduced with the kind permission of the Director and the Board of Trustees, Royal Botanic Gardens, Kew." figureDoi="http://doi.org/10.5281/zenodo.8297147" httpUri="https://zenodo.org/record/8297147/files/figure.png" pageId="6" pageNumber="295">Fig. 3</figureCitation>
). This specimen is here designated as epitype.
</paragraph>
<paragraph id="8B8636F1FFB5FF94FF34FA27FB0AA63A" blockId="6.[136,1452,158,2093]" pageId="6" pageNumber="295">
In the horticultural trade, 
<taxonomicName id="4C394D72FFB5FF94FE69FA27FDEFA5AA" authorityName="Baker" authorityYear="1882" box="[480,629,1419,1444]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="295" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis id="B94DEAE3FFB5FF94FE69FA27FDEFA5AA" box="[480,629,1419,1444]" italics="true" pageId="6" pageNumber="295">K. tomentosa</emphasis>
</taxonomicName>
is often in error referred to as “ 
<taxonomicName id="4C394D72FFB5FF94FC69FA26FB3FA5AA" authorityName="Baker" authorityYear="1895" box="[992,1189,1418,1444]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="295" phylum="Tracheophyta" rank="species" species="pilosa">
<emphasis id="B94DEAE3FFB5FF94FC69FA26FB3FA5AA" box="[992,1189,1418,1444]" italics="true" pageId="6" pageNumber="295">Kalanchoe pilosa</emphasis>
</taxonomicName>
[Hort.]”, a designation not validly published. This designation is not to be confused with 
<taxonomicName id="4C394D72FFB5FF94FC08FA03FB5FA5C6" authority="Baker (1895: 107)" authorityName="Baker" authorityPageNumber="107" authorityYear="1895" box="[897,1221,1454,1481]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="295" phylum="Tracheophyta" rank="species" species="pilosa">
<emphasis id="B94DEAE3FFB5FF94FC08FA03FC7DA5C6" box="[897,999,1454,1480]" italics="true" pageId="6" pageNumber="295">K. pilosa</emphasis>
<bibRefCitation id="EFA84B00FFB5FF94FC78FA02FB5FA5C6" author="Baker, J. G." box="[1009,1221,1454,1481]" pageId="6" pageNumber="295" pagination="288 - 293" refId="ref5721" refString="Baker, J. G. (1895 [November]) CCCCLXXXVII. - Diagnoses africanae, IX. [404. Kalanchoe pilosa Baker on p. 289.] Bulletin of Miscellaneous Information, Kew 1895 (107): 288 - 293. [https: // www. biodiversitylibrary. org / item / 127378 # page / 295 / mode / 1 up]" type="journal article" year="1895">Baker (1895: 107)</bibRefCitation>
</taxonomicName>
, a validly published name based on material from Africa collected “[…] west of Lake Tanganyika […]” (
<bibRefCitation id="EFA84B00FFB5FF94FBCFFA7EFA92A5E2" author="Baker, J. G." box="[1094,1288,1490,1517]" pageId="6" pageNumber="295" pagination="288 - 293" refId="ref5721" refString="Baker, J. G. (1895 [November]) CCCCLXXXVII. - Diagnoses africanae, IX. [404. Kalanchoe pilosa Baker on p. 289.] Bulletin of Miscellaneous Information, Kew 1895 (107): 288 - 293. [https: // www. biodiversitylibrary. org / item / 127378 # page / 295 / mode / 1 up]" type="journal article" year="1895">Baker 1895: 107</bibRefCitation>
), i.e., likely in the present-day 
<collectingCountry id="F32E7661FFB5FF94FEB3FA5AFD06A61E" box="[314,668,1526,1552]" name="Democratic Republic of the Congo" pageId="6" pageNumber="295">Democratic Republic of Congo</collectingCountry>
. The name 
<taxonomicName id="4C394D72FFB5FF94FC92FA5BFCE5A61E" authorityName="Baker" authorityYear="1895" box="[795,895,1526,1552]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="295" phylum="Tracheophyta" rank="species" species="pilosa">
<emphasis id="B94DEAE3FFB5FF94FC92FA5BFCE5A61E" box="[795,895,1526,1552]" italics="true" pageId="6" pageNumber="295">K. pilosa</emphasis>
</taxonomicName>
(of Baker, not of “Hort.”) is, at present, generally included in the synonymy of 
<taxonomicName id="4C394D72FFB5FF94FE46F9B7FDFCA63A" authorityName="Forsskal" authorityYear="1775" box="[463,614,1562,1588]" class="Magnoliopsida" family="Crassulaceae" genus="Cotyledon" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="295" phylum="Tracheophyta" rank="species" species="lanceolata">
<emphasis id="B94DEAE3FFB5FF94FE46F9B7FDFCA63A" box="[463,614,1562,1588]" italics="true" pageId="6" pageNumber="295">K. lanceolata</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFA84B00FFB5FF94FDFCF9B6FC83A63B" author="Forsskal, P." box="[629,793,1562,1589]" pageId="6" pageNumber="295" refId="ref6599" refString="Forsskal, P. (1775) Flora AEgyptiaco-Arabica. Sive descriptiones plantarum, Quas per AEgyptum inferiorem et Arabiam felicem detexit […]. Ex officina M ˆ lleri, aulae Typographi. Prostat apud Heineck et Faber, Hauniae, 220 pp. https: // doi. org / 10.5962 / bhl. title. 6625" type="book" year="1775">Forsskål 1775</bibRefCitation>
: CXI &amp; 89) 
<bibRefCitation id="EFA84B00FFB5FF94FC2DF9B7FB16A63A" author="Persoon, C. H." box="[932,1164,1562,1589]" pageId="6" pageNumber="295" refId="ref7632" refString="Persoon, C. H. (1805 [1 April - 15 June]) Synopsis plantarum, seu enchiridium botanicum, complectens enumerationem systematicam specierum hucusque cognitorum. Pars prima. Apud Carol. Frid. Cramerum, Parisiis lutetiorum [Paris] et apud J. G. Cottam, Tubingae [London], 546 pp. [https: // www. biodiversitylibrary. org / item / 11125 # page / 458 / mode / 1 up]" type="book" year="1805">Persoon (1805: 446)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C323657AFFB5FF95FF34F992FF59A266" lastPageId="7" lastPageNumber="296" pageId="6" pageNumber="295" type="description">
<paragraph id="8B8636F1FFB5FF95FF34F992FF59A266" blockId="6.[136,1452,158,2093]" lastBlockId="7.[136,1452,158,616]" lastPageId="7" lastPageNumber="296" pageId="6" pageNumber="295">
<emphasis id="B94DEAE3FFB5FF94FF34F992FE5AA656" bold="true" box="[189,448,1598,1624]" pageId="6" pageNumber="295">Amplified description</emphasis>
:—Perennial, sparsely- to many-leaved, entirely hairy, sparsely branched, small to mediumsized, terrestrial, succulent shrub, to 
<quantity id="4CC19B14FFB5FF94FDA7F9CEFDC1A673" box="[558,603,1634,1661]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="295" unit="m" value="1.0">1 m</quantity>
tall. 
<emphasis id="B94DEAE3FFB5FF94FD11F9CFFC96A672" box="[664,780,1635,1660]" italics="true" pageId="6" pageNumber="295">Tomentum</emphasis>
variable, usually dense, fine and short to coarse and quite long, erectly spreading, white, brown, or reddish brown. 
<emphasis id="B94DEAE3FFB5FF94FC9CF92AFC07A6AE" box="[789,925,1670,1696]" italics="true" pageId="6" pageNumber="295">Aerial roots</emphasis>
sometimes developing on stem and branches, hard, stiff. 
<emphasis id="B94DEAE3FFB5FF94FE8EF907FEA7A6CB" box="[263,317,1707,1733]" italics="true" pageId="6" pageNumber="295">Stem</emphasis>
branched, basally somewhat woody and in time covered in longitudinally flaking, yellowish bark, erect to arched to leaning, ± round in cross-section, scars of abscised leaves smooth, rather obscure. 
<emphasis id="B94DEAE3FFB5FF94FB66F963FAA7A6E6" box="[1263,1341,1743,1768]" italics="true" pageId="6" pageNumber="295">Leaves</emphasis>
alternate, often appearing rosulate towards branch apex, sessile, dull light green, whitish green, dull bluish green, to mid- to dark green and golden yellow-infused, distinctly succulent, erect to erectly spreading; 
<emphasis id="B94DEAE3FFB5FF94FBAEF8BAFBE9A73E" box="[1063,1139,1814,1840]" italics="true" pageId="6" pageNumber="295">petiole</emphasis>
absent, leaves not clasping the stem; 
<emphasis id="B94DEAE3FFB5FF94FF7CF896FEA9A75A" box="[245,307,1850,1876]" italics="true" pageId="6" pageNumber="295">blade</emphasis>
2.5–8.0 × 1.0– 
<quantity id="4CC19B14FFB5FF94FE54F896FDB6A75B" box="[477,556,1850,1877]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="295" unit="cm" value="2.5">2.5 cm</quantity>
, obovate, ovate, oblong, elongated-lanceolate or subcylindrical, concave above, convex below; 
<emphasis id="B94DEAE3FFB5FF94FEB0F8F2FEF6A776" box="[313,364,1886,1912]" italics="true" pageId="6" pageNumber="295">base</emphasis>
cuneate; 
<emphasis id="B94DEAE3FFB5FF94FE57F8F3FDA3A776" box="[478,569,1887,1912]" italics="true" pageId="6" pageNumber="295">margins</emphasis>
often somewhat upcurved, entire but hairy, from ± mid-leaf to apex usually ± coarsely crenate with rounded, harmless, light to reddish to dark brown teeth, teeth rarely lacking; 
<emphasis id="B94DEAE3FFB5FF94FA80F82FFAA4A792" box="[1289,1342,1923,1948]" italics="true" pageId="6" pageNumber="295">apex</emphasis>
roundedobtuse, brown-tipped. 
<emphasis id="B94DEAE3FFB5FF94FE0CF80AFD81A7CE" box="[389,539,1958,1984]" italics="true" pageId="6" pageNumber="295">Inflorescence</emphasis>
variable in length, to 
<quantity id="4CC19B14FFB5FF94FC98F80AFCC8A7CF" box="[785,850,1958,1985]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="295" unit="m" value="0.7">0.7 m</quantity>
long, erect to leaning, few- to many-flowered on few short branches, usually apically dense, corymbose; 
<emphasis id="B94DEAE3FFB5FF94FD45F866FCB2A7EA" box="[716,808,1994,2020]" italics="true" pageId="6" pageNumber="295">pedicels</emphasis>
<quantity id="4CC19B14FFB5FF94FCA6F866FC01A7EB" box="[815,923,1994,2021]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="6" pageNumber="295" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 mm</quantity>
long, slender. 
<emphasis id="B94DEAE3FFB5FF94FBC8F866FB07A7EA" box="[1089,1181,1994,2020]" italics="true" pageId="6" pageNumber="295">Flowers</emphasis>
<quantity id="4CC19B14FFB5FF94FB2DF866FA8AA7EB" box="[1188,1296,1994,2021]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.6" metricValueMin="0.8" pageId="6" pageNumber="295" unit="mm" value="12.0" valueMax="16.0" valueMin="8.0">8–16 mm</quantity>
long, erect to erectly spreading to spreading, hairy throughout, weakly diurnal; 
<emphasis id="B94DEAE3FFB5FF94FCFFF842FC2BA806" box="[886,945,2030,2056]" italics="true" pageId="6" pageNumber="295">calyx</emphasis>
consisting of 4 sepals; 
<emphasis id="B94DEAE3FFB5FF94FB48F842FA9DA806" box="[1217,1287,2030,2056]" italics="true" pageId="6" pageNumber="295">sepals</emphasis>
3–7 × 1.0– 
<quantity id="4CC19B14FFB5FF94FA01F842FF2CA823" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="295" unit="mm" value="2.5">2.5 mm</quantity>
, yellowish brown to greenish, often somewhat purplish-infused along margins and at tip, ± deltoid to elongatedtriangular, ± separate, basally fused for ± 
<quantity id="4CC19B14FFB4FF95FDDFFF32FD02A0B7" box="[598,664,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="296" unit="mm" value="1.0">1 mm</quantity>
, obtuse-tipped; 
<emphasis id="B94DEAE3FFB4FF95FCC0FF32FC55A0B6" box="[841,975,158,184]" italics="true" pageId="7" pageNumber="296">corolla tube</emphasis>
<quantity id="4CC19B14FFB4FF95FC5CFF32FBA5A0B7" box="[981,1087,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.5" metricValueMin="0.7" pageId="7" pageNumber="296" unit="mm" value="11.0" valueMax="15.0" valueMin="7.0">7–15 mm</quantity>
long, campanulate to sometimes very slightly urceolate, yellowish brown to greenish, often sometimes somewhat to strongly purple-infused; 
<emphasis id="B94DEAE3FFB4FF95FAD2FF6EFF59A10E" italics="true" pageId="7" pageNumber="296">corolla lobes</emphasis>
3–6 × 
<quantity id="4CC19B14FFB4FF95FE9FFF4AFEE3A10F" box="[278,377,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="7" pageNumber="296" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">4–8 mm</quantity>
, light to deep purple on greenish background, adaxially not hairy, velvety, deltoid to nearly round, erectly spreading to horizontally recurved, obtuse to slightly indented at tip, mucronate. 
<emphasis id="B94DEAE3FFB4FF95FB20FEA7FA9CA12B" box="[1193,1286,267,293]" italics="true" pageId="7" pageNumber="296">Stamens</emphasis>
in two whorls, one inserted low down at ± level of ovaries, the other in middle of corolla tube, included; 
<emphasis id="B94DEAE3FFB4FF95FB13FE82FA9BA146" box="[1178,1281,302,328]" italics="true" pageId="7" pageNumber="296">filaments</emphasis>
<quantity id="4CC19B14FFB4FF95FA83FE82FAF0A147" box="[1290,1386,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="7" pageNumber="296" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
long, conspicuously thickened below, thinner above, light green; 
<emphasis id="B94DEAE3FFB4FF95FCA5FEFEFC1AA162" box="[812,896,338,364]" italics="true" pageId="7" pageNumber="296">anthers</emphasis>
± 
<quantity id="4CC19B14FFB4FF95FC16FEFEFC7BA163" box="[927,993,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="296" unit="mm" value="1.0">1 mm</quantity>
long, yellow, ovate to hastate, visible at mouth. 
<emphasis id="B94DEAE3FFB4FF95FF55FEDAFE8DA19E" box="[220,279,374,400]" italics="true" pageId="7" pageNumber="296">Pistil</emphasis>
consisting of 4 carpels; 
<emphasis id="B94DEAE3FFB4FF95FDA8FEDAFDE9A19E" box="[545,627,374,400]" italics="true" pageId="7" pageNumber="296">carpels</emphasis>
<quantity id="4CC19B14FFB4FF95FDF0FEDAFD78A19F" box="[633,738,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="11.0" metricValueMin="6.0" pageId="7" pageNumber="296" unit="mm" value="8.5" valueMax="11.0" valueMin="6.0">6–11 mm</quantity>
long; 
<emphasis id="B94DEAE3FFB4FF95FCAFFEDAFCFEA19E" box="[806,868,374,400]" italics="true" pageId="7" pageNumber="296">styles</emphasis>
1.5–2.0 mm long; 
<emphasis id="B94DEAE3FFB4FF95FBBDFEDBFB10A19E" box="[1076,1162,375,400]" italics="true" pageId="7" pageNumber="296">stigmas</emphasis>
yellowish white, capitate; 
<emphasis id="B94DEAE3FFB4FF95FF01FE37FE86A1BA" box="[136,284,410,436]" italics="true" pageId="7" pageNumber="296">nectar scales</emphasis>
square to vertically rectangular, ± 1–2 × 
<quantity id="4CC19B14FFB4FF95FD7DFE36FCCCA1BB" box="[756,854,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="7" pageNumber="296" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
. 
<emphasis id="B94DEAE3FFB4FF95FCEBFE36FC5EA1BA" box="[866,964,410,436]" italics="true" pageId="7" pageNumber="296">Follicles</emphasis>
(6–) 
<quantity id="4CC19B14FFB4FF95FC72FE36FBECA1BB" box="[1019,1142,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="7" pageNumber="296" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
long, light to mid-green at first, later light brownish white, brittle, grass spikelet-like, for long tightly enveloped in dry remains of calyx. 
<emphasis id="B94DEAE3FFB4FF95FAE5FE12FA36A1D6" box="[1388,1452,446,472]" italics="true" pageId="7" pageNumber="296">Seeds</emphasis>
(0.75–)1.00–1.25(–1.50) mm long, light to dark reddish brown to dark brown to nearly black, cylindrical to bananashaped-curved to slightly to distinctly D-shaped, obscurely to distinctly longitudinally striped. 
<emphasis id="B94DEAE3FFB4FF95FB3BFDAAFA3EA22E" box="[1202,1444,518,544]" italics="true" pageId="7" pageNumber="296">Chromosome number</emphasis>
: 2 
<emphasis id="B94DEAE3FFB4FF95FF1FFD87FF3EA24A" box="[150,164,555,580]" italics="true" pageId="7" pageNumber="296">n</emphasis>
= 36 (
<bibRefCitation id="EFA84B00FFB4FF95FF66FD86FE56A24B" author="Baldwin, J. T. Jr." box="[239,460,554,581]" pageId="7" pageNumber="296" pagination="572 - 579" refId="ref5791" refString="Baldwin, J. T. Jr. (1938 [October]) Kalanchoe: the genus and its chromosomes. American Journal of Botany 25 (8): 572 - 579. https: // doi. org / 10.1002 / j. 1537 - 2197.1938. tb 09263. x" type="journal article" year="1938">Baldwin 1938: 576</bibRefCitation>
[in error as 2 
<emphasis id="B94DEAE3FFB4FF95FDE5FD87FDE0A24A" box="[620,634,555,580]" italics="true" pageId="7" pageNumber="296">n</emphasis>
= 34], 
<bibRefCitation id="EFA84B00FFB4FF95FD47FD86FCECA24A" author="Uhl, Ch. H." box="[718,886,554,580]" pageId="7" pageNumber="296" pagination="695 - 697" refId="ref9543" refString="Uhl, Ch. H. (1948) Cytotaxonomic studies in the subfamilies Crassuloideae, Kalanchoideae, and Cotyledonoideae of the Crassulaceae. American Journal of Botany 35 (10): 695 - 697. https: // doi. org / 10.1002 / j. 1537 - 2197.1948. tb 08138. x" type="journal article" year="1948">Uhl 1948: 701</bibRefCitation>
, 
<bibRefCitation id="EFA84B00FFB4FF95FC0DFD86FB51A24A" author="Sharma, A. K. &amp; Ghosh, S." box="[900,1227,554,580]" pageId="7" pageNumber="296" pagination="313 - 336" refId="ref8007" refString="Sharma, A. K. &amp; Ghosh, S. (1967) Cytotaxonomy of Crassulaceae. Biologisches Zentralblatt 86 (supplement): 313 - 336." type="journal article" year="1967">Sharma &amp; Ghosh 1967: 319</bibRefCitation>
, Smith 2022a: 162, 172).
</paragraph>
</subSubSection>
</treatment>
</document>