875 lines
157 KiB
XML
875 lines
157 KiB
XML
<document ID-DOI="10.11646/zootaxa.5244.5.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7663766" ID-ZooBank="B55396DA-15DB-4CDF-81FD-78686A0EC3A2" checkinTime="1677051835230" checkinUser="plazi" docAuthor="Ng, Peter K. L. & Rahayu, Dwi Listyo" docDate="2023" docId="039087CD7E07732047C6FAA7F6E7FCE3" docLanguage="en" docName="zootaxa.5244.5.2.pdf" docOrigin="Zootaxa 5244 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5244.5.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lophoplax sculpta" docType="treatment" docVersion="3" lastPageNumber="436" masterDocId="FFA9FFB57E0573284751FF9DF768FFDA" masterDocTitle="Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura)" masterLastPageNumber="454" masterPageNumber="428" pageNumber="430" updateTime="1677083936602" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title>Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura)</mods:title>
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<mods:namePart>Ng, Peter K. L.</mods:namePart>
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<mods:affiliation>Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore. peterng @ nus. edu. sg; https: // orcid. org / 0000 - 0001 - 5946 - 0608</mods:affiliation>
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<mods:namePart>Rahayu, Dwi Listyo</mods:namePart>
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<mods:affiliation>Research Center for Marine and Land Bioindustry, National Research and Innovation Agency, Dusun Teluk Kodek, Pemenang, Lombok Utara 83756, NTB, Indonesia. dwilistyo @ yahoo. com; https: // orcid. org / 0000 - 0003 - 4342 - 9045</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<taxonomicName ID-CoL="3W3WT" authority="(Stimpson, 1858)" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[151,593,1338,1365]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="sculpta">
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<emphasis bold="true" box="[151,375,1338,1365]" italics="true" pageId="2" pageNumber="430">Lophoplax sculpta</emphasis>
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(
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<bibRefCitation author="Stimpson, W." box="[391,584,1338,1364]" pageId="2" pageNumber="430" pagination="93 - 110" refId="ref18446" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 (39 - 56). https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
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(
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<figureCitation box="[159,292,1375,1401]" captionStart="FIGURE 1" captionStartId="3.[151,250,1724,1748]" captionTargetBox="[151,1435,561,1701]" captionTargetId="figure-203@3.[151,1435,561,1701]" captionTargetPageId="3" captionText="FIGURE 1. Colour in life. A, Lophoplax sculpta (Stimpson, 1858), male (10.2 × 8.2 mm) (RUMF-ZC-5207), Okinawa [photograph:T.Maenosono]; B, L.sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines [photograph: T.-Y. Chan]; C, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok [photograph: A. Anker]; D, L. vermiculata (A. Milne-Edwards, 1873), male, New Caledonia (not examined) [photograph: G. Paulay]; E, L. vermiculata (A. Milne-Edwards, 1873), male (14.1 × 10.8 mm) (UF 388), Guam [photograph: G. Paulay]; F, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]." figureDoi="http://doi.org/10.5281/zenodo.7663770" httpUri="https://zenodo.org/record/7663770/files/figure.png" pageId="2" pageNumber="430">Figs. 1A, B</figureCitation>
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,
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<figureCitation box="[304,373,1375,1401]" captionStart="FIGURE 2" captionStartId="5.[151,250,1775,1799]" captionTargetBox="[151,1435,309,1752]" captionTargetId="figure-91@5.[151,1435,309,1752]" captionTargetPageId="5" captionText="FIGURE 2. Overall dorsal view. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; D, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; E, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; F, Lophoplax vermiculata, subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa; H, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663772" httpUri="https://zenodo.org/record/7663772/files/figure.png" pageId="2" pageNumber="430">2A, B</figureCitation>
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,
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<figureCitation box="[385,454,1375,1401]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="2" pageNumber="430">3A–C</figureCitation>
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,
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<figureCitation box="[465,604,1374,1401]" captionStart="FIGURE 5" captionStartId="10.[151,250,1835,1859]" captionTargetBox="[151,1435,237,1812]" captionTargetId="figure-47@10.[151,1435,237,1812]" captionTargetPageId="10" captionText="FIGURE 5. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A–F, frontal view of cephalothorax; G–J, right third maxilliped; K, L, left third maxilliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663782" httpUri="https://zenodo.org/record/7663782/files/figure.png" pageId="2" pageNumber="430">5A, B, G, H</figureCitation>
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,
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<figureCitation box="[615,684,1375,1401]" captionStart="FIGURE 6" captionStartId="11.[151,250,1726,1750]" captionTargetBox="[151,1436,813,1703]" captionTargetId="figure-327@11.[151,1436,813,1703]" captionTargetPageId="11" captionText="FIGURE 6. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu.A, B, D, E, dorsal view of right cheliped; C, dorsal view of left cheliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663784" httpUri="https://zenodo.org/record/7663784/files/figure.png" pageId="2" pageNumber="430">6A, B</figureCitation>
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,
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<figureCitation box="[696,834,1375,1401]" captionStart="FIGURE 7" captionStartId="12.[151,250,1823,1847]" captionTargetBox="[229,1359,181,1799]" captionTargetId="figure-17@12.[229,1359,181,1799]" captionTargetPageId="12" captionText="FIGURE 7. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, B, right chela; C–F, left chela; G–L, right P4." figureDoi="http://doi.org/10.5281/zenodo.7663790" httpUri="https://zenodo.org/record/7663790/files/figure.png" pageId="2" pageNumber="430">7A, B, G, H</figureCitation>
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,
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<figureCitation box="[845,914,1375,1401]" captionStart="FIGURE 8" captionStartId="13.[151,250,1105,1129]" captionTargetBox="[151,1435,181,1081]" captionTargetId="figure-25@13.[151,1435,181,1081]" captionTargetPageId="13" captionText="FIGURE 8. A, B, Lophoplax sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, female thoracic sternum and pleon; B–D, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.7663793" httpUri="https://zenodo.org/record/7663793/files/figure.png" pageId="2" pageNumber="430">8A, B</figureCitation>
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<figureCitation box="[926,997,1375,1401]" captionStart="FIGURE 9" captionStartId="14.[151,250,1509,1533]" captionTargetBox="[176,1410,181,1485]" captionTargetId="figure-17@14.[176,1410,181,1485]" captionTargetPageId="14" captionText="FIGURE 9. A–D, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia. A, right lateral view of posterior part of sub-branchial region; B, E–G, anterior male thoracic sternum and pleon; C, male pleonal somites 1–4; D, male pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.7663797" httpUri="https://zenodo.org/record/7663797/files/figure.png" pageId="2" pageNumber="430">9A–D</figureCitation>
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<figureCitation box="[1008,1089,1375,1401]" captionStart="FIGURE 10" captionStartId="15.[151,247,1934,1958]" captionTargetBox="[227,1360,463,1902]" captionTargetId="figure-158@15.[212,1374,453,1911]" captionTargetPageId="15" captionText="FIGURE 10.A–E, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; F–J, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok. A, B, pleon; B, G, left G1 (ventral view); C, H, distal part of left G1 (ventral view); D, I, left G1 (dorsal view); E, J, left G2. Scales:A, F = 1.0 mm; B, E, G, J = 0.5 mm; C, D, H, I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7663799" httpUri="https://zenodo.org/record/7663799/files/figure.png" pageId="2" pageNumber="430">10A–E</figureCitation>
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)
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<taxonomicName authority="Stimpson, 1858: 93" authorityName="Stimpson" authorityPageNumber="93" authorityYear="1858" box="[151,567,1443,1467]" class="Malacostraca" family="Pilumnidae" genus="Pilumnoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="sculpta">
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<emphasis box="[151,360,1443,1467]" italics="true" pageId="2" pageNumber="430">Pilumnoplax sculpta</emphasis>
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<bibRefCitation author="Stimpson, W." box="[366,567,1443,1467]" pageId="2" pageNumber="430" pagination="93 - 110" refId="ref18446" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 (39 - 56). https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson, 1858: 93</bibRefCitation>
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;
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<treatmentCitation author="Miers, E. J." box="[578,748,1443,1467]" page="226" pageId="2" pageNumber="430" year="1886">
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<bibRefCitation author="Miers, E. J." box="[578,748,1443,1467]" pageId="2" pageNumber="430" pagination="1873 - 76" refId="ref16506" refString="Miers, E. J. (1886) Part II. Report on the Brachyura collected by H. M. S. Challenger during the years 1873 - 76. In: Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 1876 under the command of Captain George S. Nares, N. R., F. R. S. and the late Captain Frank Tourle Thomson, R. N. prepared under the Superintendence of the late Sir C. Wyville Thomson, Knt., F. R. S. & c. Regius Professor of Natural history in the University of Edinburgh of the civilian scientific staff on board and now of John Murray one of the naturalists of the Expedition. Zoology, Published by Order of Her Majesty's Government. Vol. 17. HMSO, London, Edinburgh and Dublin, pp. i - l + 1 - 362, pls. 1 - 29." type="book chapter" year="1886">Miers 1886: 226</bibRefCitation>
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;
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<treatmentCitation author="Stimpson, W." box="[758,952,1443,1467]" page="91" pageId="2" pageNumber="430" year="1907">
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<bibRefCitation author="Stimpson, W." box="[758,952,1443,1467]" pageId="2" pageNumber="430" pagination="1 - 240" refId="ref18518" refString="Stimpson, W. (1907) Report on the Crustacea (Brachyura and Anomura) collected by the North Pacific Exploring Expedition 1853 - 1856. Smithsonian Miscellaneous Collections, Part of 49 (1717), 1 - 240, 26 pls. https: // doi. org / 10.5962 / bhl. title. 51448" type="journal article" year="1907">Stimpson 1907: 91</bibRefCitation>
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, pl. 11 fig. 3.
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<taxonomicName ID-CoL="3W3WT" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[151,336,1474,1498]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="430" phylum="Arthropoda" rank="species" species="sculpta">Lophoplax sculpta</taxonomicName>
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—
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</emphasis>
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<treatmentCitation author="Tesch, J. J." box="[363,536,1474,1498]" page="199" pageId="2" pageNumber="430" year="1918">
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<bibRefCitation author="Tesch, J. J." box="[363,536,1474,1498]" pageId="2" pageNumber="430" pagination="149 - 295" refId="ref18770" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch 1918: 199</bibRefCitation>
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</treatmentCitation>
|
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; Sakai 1939: 567;
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<treatmentCitation author="Serene, R." box="[730,900,1474,1498]" page="91" pageId="2" pageNumber="430" year="1968">
|
||
<bibRefCitation author="Serene, R." box="[730,900,1474,1498]" pageId="2" pageNumber="430" pagination="33 - 120" refId="ref18274" refString="Serene, R. (1968) The Brachyura of the Indo Pacific Region. In: Prodromus for a Check List of the Non-planctonic Marine Fauna of South East Asia. Special Publication of the Singapore National Academy of Science, No. 1, 33 - 120." type="book chapter" year="1968">Serène 1968: 91</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation author="Serene, R. & Lohavanijaya, P." box="[913,1256,1474,1498]" page="69" pageId="2" pageNumber="430" year="1973">
|
||
<bibRefCitation author="Serene, R. & Lohavanijaya, P." box="[913,1256,1474,1498]" pageId="2" pageNumber="430" pagination="1 - 186" refId="ref18374" refString="Serene, R. & Lohavanijaya, P. (1973) The Brachyura (Crustacea: Decapoda) collected by the Naga Expedition, including a review of the Homolidae. Naga Report. Scientific Results of Marine Investigations of the South China Sea and the Gulf of Thailand, 4 (4), 1 - 186, figs. 1 - 186, pls. 1 - 21." type="journal article" year="1973">Serène & Lohavanijaya 1973: 69</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 75; Serène
|
||
<emphasis box="[1384,1437,1474,1498]" italics="true" pageId="2" pageNumber="430">et al.</emphasis>
|
||
1974: 24; Serène
|
||
<emphasis box="[378,429,1504,1528]" italics="true" pageId="2" pageNumber="430">et al.</emphasis>
|
||
1976: 19;
|
||
<treatmentCitation author="Sakai, T." box="[538,705,1504,1529]" page="304" pageId="2" pageNumber="430" year="1976">
|
||
<bibRefCitation author="Sakai, T." box="[538,705,1504,1529]" pageId="2" pageNumber="430" refId="ref18220" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. In three volumes. Kodansha Ltd., Tokyo, xxix + 773 pp. (English text), 461 pp. (Japanese text), 16 pp, 251 pls. (plates volume)." type="book" year="1976">Sakai 1976: 304</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 541;
|
||
<treatmentCitation author="Takeda, M." box="[767,950,1504,1529]" page="120" pageId="2" pageNumber="430" year="1977">
|
||
<bibRefCitation author="Takeda, M." box="[767,950,1504,1529]" pageId="2" pageNumber="430" pagination="120 - 123" refId="ref18577" refString="Takeda, M. (1977) Rediscovery of Lophoplax sculpta (Stimpson, 1858) (Crustacea, Decapoda, Brachyura). Zoological Magazine, 86, 120 - 123." type="journal article" year="1977">Takeda 1977: 120</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 1;
|
||
<treatmentCitation author="Miyake, S." box="[1028,1217,1504,1529]" page="231" pageId="2" pageNumber="430" year="1983">
|
||
<bibRefCitation author="Miyake, S." box="[1028,1217,1504,1529]" pageId="2" pageNumber="430" pagination="1 - 64" refId="ref16794" refString="Miyake, S. (1983) Brachyura (Crabs). Japanese crustacean decapods and stomatopods in Color. Vol. 2. 1 st Edition. Hoikusha, Osaka, viii + 277 pp., pls. 1 - 64, unnumbered fig." type="book chapter" year="1983">Miyake 1983: 231</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation author="Dai, A. - Y. & Yang, S. - L. & Song, Y. - Z. & Chen, G. - X." box="[1227,1432,1504,1529]" page="379" pageId="2" pageNumber="430" year="1986">
|
||
<bibRefCitation author="Dai, A. - Y. & Yang, S. - L. & Song, Y. - Z. & Chen, G. - X." box="[1227,1432,1504,1529]" pageId="2" pageNumber="430" refId="ref14730" refString="Dai, A. - Y., Yang, S. - L., Song, Y. - Z. & Chen, G. - X. (1986) Crabs of the China Seas. China Ocean Press, Beijing, 11 + 642 pp. [in Chinese]" type="book" year="1986">
|
||
Dai
|
||
<emphasis box="[1270,1322,1504,1528]" italics="true" pageId="2" pageNumber="430">et al.</emphasis>
|
||
1986: 379
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 200 (1, 2), pl. 55 (4);
|
||
<treatmentCitation author="Ng, P. K. L." box="[464,595,1535,1560]" page="79" pageId="2" pageNumber="430" year="1987">
|
||
<bibRefCitation author="Ng, P. K. L." box="[464,595,1535,1560]" pageId="2" pageNumber="430" pagination="69 - 111" refId="ref16944" refString="Ng, P. K. L. (1987) The Indo-Pacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). Indo-Malayan Zoology, 1987, 4 (1), 69 - 111, pl. 1." type="journal article" year="1987">Ng 1987: 79</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 100, 101;
|
||
<treatmentCitation author="Nagai, S. & Nomura, K." box="[709,999,1535,1559]" page="196" pageId="2" pageNumber="430" year="1988">
|
||
<bibRefCitation author="Nagai, S. & Nomura, K." box="[709,999,1535,1559]" pageId="2" pageNumber="430" refId="ref16841" refString="Nagai, S. & Nomura, K. (1988) Crustacea (Brachyura), [The guide book of marine organisms of Okinawa]. Vol. 7. Shinsei Tosho, Okinawa, 250 pp. [in Japanese]" type="book" year="1988">Nagai & Nomura 1988: 196</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation author="Dai, A. - Y. & Yang, S. - L." box="[1010,1243,1535,1560]" page="409" pageId="2" pageNumber="430" year="1991">
|
||
<bibRefCitation author="Dai, A. - Y. & Yang, S. - L." box="[1010,1243,1535,1560]" pageId="2" pageNumber="430" refId="ref14687" refString="Dai, A. - Y. & Yang, S. - L. (1991) Crabs of the China Seas. China Ocean Press, Beijing and Springer-Verlag, Berlin, 21 + 608 pp., 74 pls." type="book" year="1991">Dai & Yang 1991: 409</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 200 (1, 2), pl. 55 (4);
|
||
<treatmentCitation author="Nomura, K. & Nagai, S. & Asakura, A. & Komai, T." box="[273,511,1566,1590]" page="17" pageId="2" pageNumber="430" year="1996">
|
||
<bibRefCitation author="Nomura, K. & Nagai, S. & Asakura, A. & Komai, T." box="[273,511,1566,1590]" pageId="2" pageNumber="430" pagination="7 - 21" refId="ref17885" refString="Nomura, K., Nagai, S., Asakura, A. & Komai, T. (1996) A preliminary list of shallow water decapod Crustacea in the Kerama Group, the Ryukyu Archipelago. Bulletin of the Biogeographical Society of Japan, 51, 7 - 21." type="journal article" year="1996">
|
||
Nomura
|
||
<emphasis box="[362,414,1566,1590]" italics="true" pageId="2" pageNumber="430">et al.</emphasis>
|
||
1996: 17
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation author="Sakai, K." box="[521,676,1566,1590]" page="22" pageId="2" pageNumber="430" year="2003">
|
||
<bibRefCitation author="Sakai, K." box="[521,676,1566,1590]" pageId="2" pageNumber="430" pagination="13 - 30" refId="ref18132" refString="Sakai, K. (2003) The Japanese names for the Japanese crabs. Taxa, 15, 13 - 30. [in Japanese]" type="journal article" year="2003">Sakai 2003: 22</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation author="Marumura, M. & Kosaka, A." box="[686,999,1566,1590]" page="47" pageId="2" pageNumber="430" year="2003">
|
||
<bibRefCitation author="Marumura, M. & Kosaka, A." box="[686,999,1566,1590]" pageId="2" pageNumber="430" pagination="1 - 8" refId="ref16330" refString="Marumura, M. & Kosaka, A. (2003) Catalogue of Brachyuran and Anomuran crabs Collection Donated by the late Mr. Seiji Nagai to the Wakayama Prefectural Museum of Natural History. Wakayama Prefectural Museum of Natural History, Kainan, Wakayama Prefecture, 73 pp., pls. 1 - 8." type="book chapter" year="2003">Marumura & Kosaka 2003: 47</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation author="Ng, P. K. L. & D. Guinot & P. J. F. Davie" box="[1009,1207,1566,1590]" page="143" pageId="2" pageNumber="430" year="2008">
|
||
<bibRefCitation author="Ng, P. K. L. & D. Guinot & P. J. F. Davie" box="[1009,1207,1566,1590]" pageId="2" pageNumber="430" pagination="1 - 286" refId="ref17397" refString="Ng, P. K. L., D. Guinot & P. J. F. Davie, 2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
|
||
Ng
|
||
<emphasis box="[1046,1098,1566,1590]" italics="true" pageId="2" pageNumber="430">et al.</emphasis>
|
||
2008: 143
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 111;
|
||
<treatmentCitation author="Castro, P. & Ng, P. K. L." page="25" pageId="2" pageNumber="430" year="2010">
|
||
<bibRefCitation author="Castro, P. & Ng, P. K. L." pageId="2" pageNumber="430" pagination="1 - 130" refId="ref14630" refString="Castro, P. & Ng, P. K. L. (2010) Revision of the family Euryplacidae Stimpson, 1871 (Crustacea: Decapoda: Brachyura: Goneplacoidea). Zootaxa, 2375 (1), 1 - 130. https: // doi. org / 10.11646 / zootaxa. 2375.1.1" type="journal article" year="2010">Castro & Ng 2010: 25</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation author="Naruse, T." box="[302,472,1597,1621]" page="31" pageId="2" pageNumber="430" year="2010">
|
||
<bibRefCitation author="Naruse, T." box="[302,472,1597,1621]" pageId="2" pageNumber="430" pagination="1 - 72" refId="ref16885" refString="Naruse, T. (2010) Catalogue of Crustacean specimens deposited in the university museum (Fujukan), University of the Ryukyus. Catalogue of materials deposited in the university museum (Fujukan), University of the Ryukyus, 3, 1 - 72. https: // doi. org / 10.1017 / CBO 9780511703546.002" type="journal article" year="2010">Naruse 2010: 31</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation author="Maenosono, T." box="[483,699,1597,1621]" page="30" pageId="2" pageNumber="430" year="2019">
|
||
<bibRefCitation author="Maenosono, T." box="[483,699,1597,1621]" pageId="2" pageNumber="430" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019: 30</bibRefCitation>
|
||
</treatmentCitation>
|
||
, figs. 8A, 9, 13A, 14I, J.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="430" type="materials_examined">
|
||
<paragraph blockId="2.[151,1437,1666,2017]" pageId="2" pageNumber="430">
|
||
<emphasis bold="true" box="[151,328,1666,1693]" pageId="2" pageNumber="430">Type material.</emphasis>
|
||
<materialsCitation collectingDate="2008-01-06" collectorName="T. Maenosono" country="Japan" location="Okinawa" pageId="2" pageNumber="430" specimenCount="1" stateProvince="Okinawa" typeStatus="neotype">
|
||
<emphasis box="[339,431,1667,1692]" italics="true" pageId="2" pageNumber="430">
|
||
<typeStatus box="[339,431,1667,1692]" pageId="2" pageNumber="430">Neotype</typeStatus>
|
||
</emphasis>
|
||
(here designated): male (7.8 ×
|
||
<quantity box="[802,893,1667,1693]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="2" pageNumber="430" unit="mm" value="6.3">6.3 mm</quantity>
|
||
) (RUMF-ZC-8127), Minatagawa, Urasoe City,
|
||
<collectingRegion box="[151,255,1703,1729]" country="Japan" name="Okinawa" pageId="2" pageNumber="430">Okinawa</collectingRegion>
|
||
,
|
||
<collectingCountry box="[266,333,1703,1729]" name="Japan" pageId="2" pageNumber="430">Japan</collectingCountry>
|
||
, coll.
|
||
<collectorName box="[399,563,1703,1729]" pageId="2" pageNumber="430">T. Maenosono</collectorName>
|
||
,
|
||
<date box="[573,748,1703,1729]" pageId="2" pageNumber="430" value="2008-01-06">
|
||
<collectingDate box="[573,748,1703,1729]" pageId="2" pageNumber="430" value="2008-01-06">6 January 2008</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,1666,2017]" pageId="2" pageNumber="430">
|
||
<emphasis bold="true" box="[199,504,1738,1764]" pageId="2" pageNumber="430">Other material examined</emphasis>
|
||
.
|
||
<materialsCitation collectingDate="2009-08-22" collectionCode="ZRC" collectorName="T. Maenosono" country="Japan" location="Ryukyu Islands" municipality="Onna Village" pageId="2" pageNumber="430" specimenCode="ZRC 2022.777" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa">
|
||
<emphasis bold="true" box="[520,593,1739,1765]" pageId="2" pageNumber="430">
|
||
<collectingCountry box="[520,593,1739,1765]" name="Japan" pageId="2" pageNumber="430">Japan</collectingCountry>
|
||
</emphasis>
|
||
—
|
||
<specimenCount box="[625,697,1739,1765]" pageId="2" pageNumber="430" type="male">1 male</specimenCount>
|
||
(7.8 ×
|
||
<quantity box="[784,873,1739,1765]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="2" pageNumber="430" unit="mm" value="6.2">6.2 mm</quantity>
|
||
) (
|
||
<specimenCode box="[897,1072,1739,1765]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="430">ZRC 2022.777</specimenCode>
|
||
),
|
||
<collectorName box="[1092,1157,1739,1765]" pageId="2" pageNumber="430">Ukaji</collectorName>
|
||
,
|
||
<collectingMunicipality box="[1170,1323,1739,1765]" pageId="2" pageNumber="430">Onna Village</collectingMunicipality>
|
||
,
|
||
<collectingRegion box="[1335,1436,1739,1765]" country="Japan" name="Okinawa" pageId="2" pageNumber="430">Okinawa</collectingRegion>
|
||
<collectorName box="[151,222,1775,1801]" pageId="2" pageNumber="430">Island</collectorName>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:039087CD7E07732047C6FAA7F6E7FCE3:8EE660007E07732A47BEF972F6CCF8D3" box="[239,420,1775,1801]" country="Japan" municipality="Onna Village" name="Ryukyu Islands" pageId="2" pageNumber="430" stateProvince="Okinawa">Ryukyu Islands</location>
|
||
, coll.
|
||
<collectorName box="[497,665,1775,1801]" pageId="2" pageNumber="430">T. Maenosono</collectorName>
|
||
,
|
||
<date box="[680,872,1775,1801]" pageId="2" pageNumber="430" value="2009-08-22">
|
||
<collectingDate box="[680,872,1775,1801]" pageId="2" pageNumber="430" value="2009-08-22">22 August 2009</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation collectingDate="2004-06-10" collectingDateMax="2004-06-12" collectingDateMin="2004-06-10" collectionCode="ZRC" collectorName="Panglao & Expedition" country="Philippines" elevation="5" location="Gak-Ang Island" pageId="2" pageNumber="430" specimenCode="ZRC 2019.1611" specimenCount="1" specimenCount-female="1" stateProvince="Bohol">
|
||
<emphasis bold="true" box="[887,1021,1774,1800]" pageId="2" pageNumber="430">
|
||
<collectingCountry box="[887,1021,1774,1800]" name="Philippines" pageId="2" pageNumber="430">Philippines</collectingCountry>
|
||
</emphasis>
|
||
—
|
||
<specimenCount box="[1053,1150,1775,1801]" pageId="2" pageNumber="430" type="female">1 female</specimenCount>
|
||
(10.4 ×
|
||
<quantity box="[1261,1353,1774,1801]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="430" unit="mm" value="8.5">8.5 mm</quantity>
|
||
) (
|
||
<specimenCode collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="430">ZRC 2019.1611</specimenCode>
|
||
), station M18, sandy bottom with seagrass,
|
||
<location LSID="urn:lsid:plazi:treatment:039087CD7E07732047C6FAA7F6E7FCE3:8EE660007E07732A4457F88EF4D5F8F7" box="[774,957,1811,1837]" country="Philippines" name="Gak-Ang Island" pageId="2" pageNumber="430" stateProvince="Bohol">Gak-Ang Island</location>
|
||
,
|
||
<collectorName box="[970,1063,1811,1837]" pageId="2" pageNumber="430">Panglao</collectorName>
|
||
,
|
||
<collectingRegion box="[1076,1147,1811,1837]" country="Philippines" name="Bohol" pageId="2" pageNumber="430">Bohol</collectingRegion>
|
||
,
|
||
<quantity box="[1160,1234,1811,1837]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="2" pageNumber="430" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">
|
||
<elevation box="[1160,1234,1811,1837]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="2" pageNumber="430" unit="m" value="0.5" valueMax="1.0" valueMin="0.0">0–1 m</elevation>
|
||
</quantity>
|
||
, coll. PANGLAO 2004
|
||
<collectorName box="[214,340,1847,1873]" pageId="2" pageNumber="430">Expedition</collectorName>
|
||
,
|
||
<date box="[351,546,1847,1873]" pageId="2" pageNumber="430" value="2004-06-10" valueMax="2004-06-12" valueMin="2004-06-10">
|
||
<collectingDate box="[351,546,1847,1873]" pageId="2" pageNumber="430" value="2004-06-10" valueMax="2004-06-12" valueMin="2004-06-10">10–12 June 2004</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="3" lastPageNumber="431" pageId="2" pageNumber="430" type="diagnosis">
|
||
<paragraph blockId="2.[151,1437,1666,2017]" lastBlockId="3.[151,1437,151,537]" lastPageId="3" lastPageNumber="431" pageId="2" pageNumber="430">
|
||
<emphasis bold="true" box="[199,314,1882,1908]" pageId="2" pageNumber="430">Diagnosis</emphasis>
|
||
. Carapace subquadrate (
|
||
<figureCitation box="[592,716,1883,1909]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="2" pageNumber="430">Fig. 3A–C</figureCitation>
|
||
); epigastric, protogastric, hepatic, cardiac and intestinal areolets all glabrous, prominent, smooth; epigastric areolets relatively wide, ovate, not fused with protogastric areolets (
|
||
<figureCitation captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="2" pageNumber="430">Fig. 3A–C</figureCitation>
|
||
); protogastric areolet prominent, transversely ovate to weakly crescent-shaped, positioned transversely (
|
||
<figureCitation captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="2" pageNumber="430">Fig. 3A–C</figureCitation>
|
||
); cardiac areolet transverse in position (
|
||
<figureCitation box="[680,806,1991,2017]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="2" pageNumber="430">Fig. 3A–C</figureCitation>
|
||
); hepatic areolet swollen, prominent, sometimes with a median depression, proportionately short, reaching only to base of first anterolateral tooth (
|
||
<figureCitation box="[1198,1324,151,177]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="3" pageNumber="431">Fig. 3A–C</figureCitation>
|
||
); external orbital tooth acutely triangular (
|
||
<figureCitation box="[526,654,187,213]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="3" pageNumber="431">Fig. 3A–C</figureCitation>
|
||
); anterolateral teeth sharply pointed (
|
||
<figureCitation box="[1093,1222,187,213]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="3" pageNumber="431">Fig. 3A–C</figureCitation>
|
||
); last anterolateral tooth rounded, at junction between antero- and posterolateral margins, partially fused with 1 or 2 partially fused rounded to subtruncate tubercles each often with median depression (
|
||
<figureCitation box="[947,1082,259,285]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="3" pageNumber="431">Figs. 3A–C</figureCitation>
|
||
,
|
||
<figureCitation box="[1096,1133,259,285]" captionStart="FIGURE 9" captionStartId="14.[151,250,1509,1533]" captionTargetBox="[176,1410,181,1485]" captionTargetId="figure-17@14.[176,1410,181,1485]" captionTargetPageId="14" captionText="FIGURE 9. A–D, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia. A, right lateral view of posterior part of sub-branchial region; B, E–G, anterior male thoracic sternum and pleon; C, male pleonal somites 1–4; D, male pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.7663797" httpUri="https://zenodo.org/record/7663797/files/figure.png" pageId="3" pageNumber="431">9A</figureCitation>
|
||
); epistome relatively wide longitudinally (
|
||
<figureCitation box="[323,442,294,321]" captionStart="FIGURE 5" captionStartId="10.[151,250,1835,1859]" captionTargetBox="[151,1435,237,1812]" captionTargetId="figure-47@10.[151,1435,237,1812]" captionTargetPageId="10" captionText="FIGURE 5. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A–F, frontal view of cephalothorax; G–J, right third maxilliped; K, L, left third maxilliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663782" httpUri="https://zenodo.org/record/7663782/files/figure.png" pageId="3" pageNumber="431">Fig. 5A, B</figureCitation>
|
||
); median lobe of posterior margin of the epistome broadly triangular, with lateral margins deeply concave (
|
||
<figureCitation box="[337,456,330,357]" captionStart="FIGURE 5" captionStartId="10.[151,250,1835,1859]" captionTargetBox="[151,1435,237,1812]" captionTargetId="figure-47@10.[151,1435,237,1812]" captionTargetPageId="10" captionText="FIGURE 5. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A–F, frontal view of cephalothorax; G–J, right third maxilliped; K, L, left third maxilliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663782" httpUri="https://zenodo.org/record/7663782/files/figure.png" pageId="3" pageNumber="431">Fig. 5A, B</figureCitation>
|
||
). Third maxilliped with merus quadrate, ischium subrectangular (
|
||
<figureCitation box="[1181,1303,330,357]" captionStart="FIGURE 5" captionStartId="10.[151,250,1835,1859]" captionTargetBox="[151,1435,237,1812]" captionTargetId="figure-47@10.[151,1435,237,1812]" captionTargetPageId="10" captionText="FIGURE 5. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A–F, frontal view of cephalothorax; G–J, right third maxilliped; K, L, left third maxilliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663782" httpUri="https://zenodo.org/record/7663782/files/figure.png" pageId="3" pageNumber="431">Fig. 5G, H</figureCitation>
|
||
); dorsal and lateral surfaces of carpus of cheliped with 5 distinct smooth tubercles (
|
||
<figureCitation box="[950,1073,367,393]" captionStart="FIGURE 6" captionStartId="11.[151,250,1726,1750]" captionTargetBox="[151,1436,813,1703]" captionTargetId="figure-327@11.[151,1436,813,1703]" captionTargetPageId="11" captionText="FIGURE 6. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu.A, B, D, E, dorsal view of right cheliped; C, dorsal view of left cheliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663784" httpUri="https://zenodo.org/record/7663784/files/figure.png" pageId="3" pageNumber="431">Fig. 6A, B</figureCitation>
|
||
). Cheliped fingers not distinctly bent, pollex subparallel with the ventral margin of palm (
|
||
<figureCitation box="[785,905,403,429]" captionStart="FIGURE 7" captionStartId="12.[151,250,1823,1847]" captionTargetBox="[229,1359,181,1799]" captionTargetId="figure-17@12.[229,1359,181,1799]" captionTargetPageId="12" captionText="FIGURE 7. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, B, right chela; C–F, left chela; G–L, right P4." figureDoi="http://doi.org/10.5281/zenodo.7663790" httpUri="https://zenodo.org/record/7663790/files/figure.png" pageId="3" pageNumber="431">Fig. 7A, B</figureCitation>
|
||
). P2–P5 not prominently elongate (
|
||
<figureCitation box="[1300,1432,403,429]" captionStart="FIGURE 2" captionStartId="5.[151,250,1775,1799]" captionTargetBox="[151,1435,309,1752]" captionTargetId="figure-91@5.[151,1435,309,1752]" captionTargetPageId="5" captionText="FIGURE 2. Overall dorsal view. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; D, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; E, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; F, Lophoplax vermiculata, subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa; H, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663772" httpUri="https://zenodo.org/record/7663772/files/figure.png" pageId="3" pageNumber="431">Figs. 2A, B</figureCitation>
|
||
,
|
||
<figureCitation box="[151,224,439,465]" captionStart="FIGURE 7" captionStartId="12.[151,250,1823,1847]" captionTargetBox="[229,1359,181,1799]" captionTargetId="figure-17@12.[229,1359,181,1799]" captionTargetPageId="12" captionText="FIGURE 7. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, B, right chela; C–F, left chela; G–L, right P4." figureDoi="http://doi.org/10.5281/zenodo.7663790" httpUri="https://zenodo.org/record/7663790/files/figure.png" pageId="3" pageNumber="431">7G, H</figureCitation>
|
||
); outer surface of the P3–P5 merus, carpus and propodus relatively smooth to slightly rugose (
|
||
<figureCitation box="[1298,1422,439,465]" captionStart="FIGURE 6" captionStartId="11.[151,250,1726,1750]" captionTargetBox="[151,1436,813,1703]" captionTargetId="figure-327@11.[151,1436,813,1703]" captionTargetPageId="11" captionText="FIGURE 6. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu.A, B, D, E, dorsal view of right cheliped; C, dorsal view of left cheliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663784" httpUri="https://zenodo.org/record/7663784/files/figure.png" pageId="3" pageNumber="431">Fig. 6A, B</figureCitation>
|
||
). Male pleonal somites 4–6 transversely narrow, telson slightly longer than broad, triangular (
|
||
<figureCitation box="[1185,1319,475,501]" captionStart="FIGURE 9" captionStartId="14.[151,250,1509,1533]" captionTargetBox="[176,1410,181,1485]" captionTargetId="figure-17@14.[176,1410,181,1485]" captionTargetPageId="14" captionText="FIGURE 9. A–D, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia. A, right lateral view of posterior part of sub-branchial region; B, E–G, anterior male thoracic sternum and pleon; C, male pleonal somites 1–4; D, male pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.7663797" httpUri="https://zenodo.org/record/7663797/files/figure.png" pageId="3" pageNumber="431">Figs. 9B, D</figureCitation>
|
||
,
|
||
<figureCitation box="[1330,1382,475,501]" captionStart="FIGURE 10" captionStartId="15.[151,247,1934,1958]" captionTargetBox="[227,1360,463,1902]" captionTargetId="figure-158@15.[212,1374,453,1911]" captionTargetPageId="15" captionText="FIGURE 10.A–E, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; F–J, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok. A, B, pleon; B, G, left G1 (ventral view); C, H, distal part of left G1 (ventral view); D, I, left G1 (dorsal view); E, J, left G2. Scales:A, F = 1.0 mm; B, E, G, J = 0.5 mm; C, D, H, I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7663799" httpUri="https://zenodo.org/record/7663799/files/figure.png" pageId="3" pageNumber="431">10A</figureCitation>
|
||
). G1 strongly sinuous, distal part long, gently curved, elongate (
|
||
<figureCitation box="[809,947,511,537]" captionStart="FIGURE 10" captionStartId="15.[151,247,1934,1958]" captionTargetBox="[227,1360,463,1902]" captionTargetId="figure-158@15.[212,1374,453,1911]" captionTargetPageId="15" captionText="FIGURE 10.A–E, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; F–J, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok. A, B, pleon; B, G, left G1 (ventral view); C, H, distal part of left G1 (ventral view); D, I, left G1 (dorsal view); E, J, left G2. Scales:A, F = 1.0 mm; B, E, G, J = 0.5 mm; C, D, H, I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7663799" httpUri="https://zenodo.org/record/7663799/files/figure.png" pageId="3" pageNumber="431">Fig. 10B–D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7663770" ID-Zenodo-Dep="7663770" httpUri="https://zenodo.org/record/7663770/files/figure.png" pageId="3" pageNumber="431" startId="3.[151,250,1724,1748]" targetBox="[151,1435,561,1701]" targetPageId="3">
|
||
<paragraph blockId="3.[151,1437,1724,1928]" pageId="3" pageNumber="431">
|
||
<emphasis bold="true" box="[151,280,1724,1748]" pageId="3" pageNumber="431">FIGURE 1.</emphasis>
|
||
Colour in life. A,
|
||
<taxonomicName authority="(Stimpson, 1858)" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[488,874,1724,1748]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="431" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis box="[488,680,1724,1748]" italics="true" pageId="3" pageNumber="431">Lophoplax sculpta</emphasis>
|
||
(
|
||
<bibRefCitation author="Stimpson, W." box="[698,866,1724,1748]" pageId="3" pageNumber="431" pagination="93 - 110" refId="ref18446" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 (39 - 56). https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male (10.2 × 8.2 mm) (RUMF-ZC-5207), Okinawa [photograph: T. Maenosono]; B,
|
||
<taxonomicName authority="(Stimpson, 1858)" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[476,754,1760,1784]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="431" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis box="[476,573,1760,1784]" italics="true" pageId="3" pageNumber="431">L. sculpta</emphasis>
|
||
(
|
||
<bibRefCitation author="Stimpson, W." box="[585,747,1760,1784]" pageId="3" pageNumber="431" pagination="93 - 110" refId="ref18446" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 (39 - 56). https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines [photograph: T.-Y. Chan]; C,
|
||
<taxonomicName authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[321,427,1796,1820]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="431" phylum="Arthropoda" rank="species" species="sordida" status="n. sp.">
|
||
<emphasis box="[321,427,1796,1820]" italics="true" pageId="3" pageNumber="431">L. sordida</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[437,498,1796,1820]" pageId="3" pageNumber="431">
|
||
<taxonomicNameLabel box="[437,498,1796,1820]" pageId="3" pageNumber="431" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok [photograph: A. Anker]; D,
|
||
<taxonomicName authority="(A. Milne-Edwards, 1873)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="431" phylum="Arthropoda" rank="species" species="vermiculata">
|
||
<emphasis italics="true" pageId="3" pageNumber="431">L. vermiculata</emphasis>
|
||
(A.
|
||
<bibRefCitation author="Milne-Edwards, A." box="[320,545,1832,1856]" pageId="3" pageNumber="431" pagination="155 - 332" refId="ref16688" refString="Milne-Edwards, A. (1873) Recherches sur la faune carcinologique de la Nouvelle-Caledonie. Deuxieme Partie. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 9, 155 - 332, pls. 4 - 18. https: // doi. org / 10.5962 / bhl. title. 10410" type="journal article" year="1873">Milne-Edwards, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male, New Caledonia (not examined) [photograph: G. Paulay]; E,
|
||
<taxonomicName authority="(A. Milne-Edwards, 1873)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="431" phylum="Arthropoda" rank="species" species="vermiculata">
|
||
<emphasis box="[1247,1396,1832,1856]" italics="true" pageId="3" pageNumber="431">L. vermiculata</emphasis>
|
||
(A.
|
||
<bibRefCitation author="Milne-Edwards, A." box="[151,375,1868,1892]" pageId="3" pageNumber="431" pagination="155 - 332" refId="ref16688" refString="Milne-Edwards, A. (1873) Recherches sur la faune carcinologique de la Nouvelle-Caledonie. Deuxieme Partie. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 9, 155 - 332, pls. 4 - 18. https: // doi. org / 10.5962 / bhl. title. 10410" type="journal article" year="1873">Milne-Edwards, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male (14.1 × 10.8 mm) (UF 388), Guam [photograph: G. Paulay]; F,
|
||
<taxonomicName authority="(A. Milne-Edwards, 1873)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="431" phylum="Arthropoda" rank="species" species="vermiculata">
|
||
<emphasis box="[1081,1228,1868,1892]" italics="true" pageId="3" pageNumber="431">L. vermiculata</emphasis>
|
||
(A.
|
||
<bibRefCitation author="Milne-Edwards, A." pageId="3" pageNumber="431" pagination="155 - 332" refId="ref16688" refString="Milne-Edwards, A. (1873) Recherches sur la faune carcinologique de la Nouvelle-Caledonie. Deuxieme Partie. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 9, 155 - 332, pls. 4 - 18. https: // doi. org / 10.5962 / bhl. title. 10410" type="journal article" year="1873">Milne-Edwards, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono].
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="4" pageNumber="432" type="description">
|
||
<paragraph blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="432">
|
||
<emphasis bold="true" box="[199,430,150,177]" pageId="4" pageNumber="432">Description of male</emphasis>
|
||
. Carapace subquadrate, slightly broader than long; dorsal surface gently convex in anterior half, relatively flatter on posterior half; regions distinct; surface entirely covered by short, thick setae and scattered longer plumose setae, forming spongy tomentum except for smooth, glabrous areolets on symmetrical epigastric, protogastric, hepatic, cardiac and intestinal regions; epigastric region relatively wide, longitudinally subrectangular, not fused with protogastric areolets; protogastric region transversely ovate to weakly crescent-shape, positioned transversely; cardiac region longitudinally narrow, wide, positioned transversely, medially separated by shallow setose depression; intestinal region longitudinally narrow, very wide, reaching across most of posterior margin, medially separated by shallow weakly setose part (sometimes not easily discernible); hepatic region obliquely ovate, swollen, smooth, often with median depression, adjacent to external orbital and first anterolateral teeth (
|
||
<figureCitation box="[1301,1432,439,465]" captionStart="FIGURE 2" captionStartId="5.[151,250,1775,1799]" captionTargetBox="[151,1435,309,1752]" captionTargetId="figure-91@5.[151,1435,309,1752]" captionTargetPageId="5" captionText="FIGURE 2. Overall dorsal view. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; D, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; E, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; F, Lophoplax vermiculata, subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa; H, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663772" httpUri="https://zenodo.org/record/7663772/files/figure.png" pageId="4" pageNumber="432">Figs. 2A, B</figureCitation>
|
||
,
|
||
<figureCitation box="[151,223,475,501]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="4" pageNumber="432">3A–C</figureCitation>
|
||
). Frontal margin slightly produced, gently deflexed, with 2 low lobes separated by shallow concavity; lateral lobe very low, almost undiscernible (
|
||
<figureCitation box="[577,711,511,537]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="4" pageNumber="432">Figs. 3A–C</figureCitation>
|
||
,
|
||
<figureCitation box="[725,797,510,537]" captionStart="FIGURE 5" captionStartId="10.[151,250,1835,1859]" captionTargetBox="[151,1435,237,1812]" captionTargetId="figure-47@10.[151,1435,237,1812]" captionTargetPageId="10" captionText="FIGURE 5. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A–F, frontal view of cephalothorax; G–J, right third maxilliped; K, L, left third maxilliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663782" httpUri="https://zenodo.org/record/7663782/files/figure.png" pageId="4" pageNumber="432">5A, B</figureCitation>
|
||
). Anterolateral margin gently convex, weakly separated from posterolateral margin; external orbital tooth acutely triangular, directed anteriorly, tip sharply pointed to gently rounded; first anterolateral tooth broadly triangular, sharply pointed; second anterolateral tooth smaller than first, tip sharp; 2 large rounded to subtruncate tubercles at junction between antero- and posterolateral margin, tip rounded to flattened, often with median depression (
|
||
<figureCitation box="[644,779,655,681]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="4" pageNumber="432">Figs. 3A–C</figureCitation>
|
||
). Posterolateral margin gently convex, surface granulated; margin gently converging to sinuous posterior carapace margin (
|
||
<figureCitation box="[874,998,691,717]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="4" pageNumber="432">Fig. 3A–C</figureCitation>
|
||
). Posterior part of sub-branchial region with distinct rounded ridge, forming channel with margin, connected to end of posterolateral margin, at junction with metabranchial region. Suborbital, subhepatic, pterygostomial and sub-branchial regions smooth (
|
||
<figureCitation box="[1299,1422,762,789]" captionStart="FIGURE 5" captionStartId="10.[151,250,1835,1859]" captionTargetBox="[151,1435,237,1812]" captionTargetId="figure-47@10.[151,1435,237,1812]" captionTargetPageId="10" captionText="FIGURE 5. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A–F, frontal view of cephalothorax; G–J, right third maxilliped; K, L, left third maxilliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663782" httpUri="https://zenodo.org/record/7663782/files/figure.png" pageId="4" pageNumber="432">Fig. 5A, B</figureCitation>
|
||
). Orbit transverse, large; supraorbital margin gently sinuous, entire; suborbital margin gently convex, entire, with low tooth on inner angle; eyes large with short peduncle and large cornea (
|
||
<figureCitation box="[987,1120,835,861]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="4" pageNumber="432">Figs. 3A–C</figureCitation>
|
||
,
|
||
<figureCitation box="[1131,1200,834,861]" captionStart="FIGURE 5" captionStartId="10.[151,250,1835,1859]" captionTargetBox="[151,1435,237,1812]" captionTargetId="figure-47@10.[151,1435,237,1812]" captionTargetPageId="10" captionText="FIGURE 5. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A–F, frontal view of cephalothorax; G–J, right third maxilliped; K, L, left third maxilliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663782" httpUri="https://zenodo.org/record/7663782/files/figure.png" pageId="4" pageNumber="432">5A, B</figureCitation>
|
||
). Antenna with basal article quadrate, fused with epistome but sutures visible, article 3 slightly shorter than basal article; article 4 as long as article 5, flagellum short, not entering orbit; antennule folding transversely (
|
||
<figureCitation box="[1111,1235,906,933]" captionStart="FIGURE 5" captionStartId="10.[151,250,1835,1859]" captionTargetBox="[151,1435,237,1812]" captionTargetId="figure-47@10.[151,1435,237,1812]" captionTargetPageId="10" captionText="FIGURE 5. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A–F, frontal view of cephalothorax; G–J, right third maxilliped; K, L, left third maxilliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663782" httpUri="https://zenodo.org/record/7663782/files/figure.png" pageId="4" pageNumber="432">Fig. 5A, B</figureCitation>
|
||
). Epistome wider longitudinally, posterior margin with wide triangular median lobe with median fissure, separated from lateral part by short fissure; lateral margin prominently concave (
|
||
<figureCitation box="[717,839,978,1005]" captionStart="FIGURE 5" captionStartId="10.[151,250,1835,1859]" captionTargetBox="[151,1435,237,1812]" captionTargetId="figure-47@10.[151,1435,237,1812]" captionTargetPageId="10" captionText="FIGURE 5. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A–F, frontal view of cephalothorax; G–J, right third maxilliped; K, L, left third maxilliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663782" httpUri="https://zenodo.org/record/7663782/files/figure.png" pageId="4" pageNumber="432">Fig. 5A, B</figureCitation>
|
||
). Endostome with low oblique ridge on each half.
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="432">
|
||
Third maxilliped relatively short; merus quadrate, anteroexternal angle weakly auriculiform; ischium subrectangular with shallow oblique median sulcus; exopod relatively broad, tip not reaching distal edge of merus, with long flagellum (
|
||
<figureCitation box="[387,512,1086,1113]" captionStart="FIGURE 5" captionStartId="10.[151,250,1835,1859]" captionTargetBox="[151,1435,237,1812]" captionTargetId="figure-47@10.[151,1435,237,1812]" captionTargetPageId="10" captionText="FIGURE 5. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A–F, frontal view of cephalothorax; G–J, right third maxilliped; K, L, left third maxilliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663782" httpUri="https://zenodo.org/record/7663782/files/figure.png" pageId="4" pageNumber="432">Fig. 5G, H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="432">
|
||
Chelipeds almost symmetrical (
|
||
<figureCitation box="[552,672,1123,1149]" captionStart="FIGURE 2" captionStartId="5.[151,250,1775,1799]" captionTargetBox="[151,1435,309,1752]" captionTargetId="figure-91@5.[151,1435,309,1752]" captionTargetPageId="5" captionText="FIGURE 2. Overall dorsal view. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; D, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; E, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; F, Lophoplax vermiculata, subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa; H, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663772" httpUri="https://zenodo.org/record/7663772/files/figure.png" pageId="4" pageNumber="432">Fig. 2A, B</figureCitation>
|
||
), relatively short, covered with same pubescence on carapace except for areolets; basis-ischium lined with small sharp granules on ventral margin; merus short, unarmed, outer surface with 4 transverse tubercles, proximal ones largest; carpus with sharp inner distal tooth, dorsal surface with 1 large oblique tubercle and 2 smaller ones, outer lateral margin with 2 low rounded lobes, basally fused in part, proximal one larger; chela not enlarged, outer surface granulate, fingers longer than palm, dorsal margin of dactylus serrate; propodus not bent downwards, almost subparallel with ventral margin of palm (
|
||
<figureCitation box="[1044,1176,1303,1329]" captionStart="FIGURE 6" captionStartId="11.[151,250,1726,1750]" captionTargetBox="[151,1436,813,1703]" captionTargetId="figure-327@11.[151,1436,813,1703]" captionTargetPageId="11" captionText="FIGURE 6. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu.A, B, D, E, dorsal view of right cheliped; C, dorsal view of left cheliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663784" httpUri="https://zenodo.org/record/7663784/files/figure.png" pageId="4" pageNumber="432">Figs. 6A, B</figureCitation>
|
||
,
|
||
<figureCitation box="[1188,1257,1303,1329]" captionStart="FIGURE 7" captionStartId="12.[151,250,1823,1847]" captionTargetBox="[229,1359,181,1799]" captionTargetId="figure-17@12.[229,1359,181,1799]" captionTargetPageId="12" captionText="FIGURE 7. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, B, right chela; C–F, left chela; G–L, right P4." figureDoi="http://doi.org/10.5281/zenodo.7663790" httpUri="https://zenodo.org/record/7663790/files/figure.png" pageId="4" pageNumber="432">7A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="432">
|
||
Ambulatory legs not prominently elongate, covered with same pubescence on carapace except for swollen surfaces; outer surface weakly eroded; P3 longest, P5 shortest; merus with subdorsal longitudinal swelling, forming low dorsal and ventral cristae; carpus and propodus with prominent median longitudinal swelling, outer surface almost smooth to gently rugose; dactylus gently curved along most of length, tip hooked, glabrous (
|
||
<figureCitation box="[1203,1337,1447,1473]" captionStart="FIGURE 2" captionStartId="5.[151,250,1775,1799]" captionTargetBox="[151,1435,309,1752]" captionTargetId="figure-91@5.[151,1435,309,1752]" captionTargetPageId="5" captionText="FIGURE 2. Overall dorsal view. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; D, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; E, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; F, Lophoplax vermiculata, subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa; H, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663772" httpUri="https://zenodo.org/record/7663772/files/figure.png" pageId="4" pageNumber="432">Figs. 2A, B</figureCitation>
|
||
,
|
||
<figureCitation box="[1350,1424,1447,1473]" captionStart="FIGURE 7" captionStartId="12.[151,250,1823,1847]" captionTargetBox="[229,1359,181,1799]" captionTargetId="figure-17@12.[229,1359,181,1799]" captionTargetPageId="12" captionText="FIGURE 7. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, B, right chela; C–F, left chela; G–L, right P4." figureDoi="http://doi.org/10.5281/zenodo.7663790" httpUri="https://zenodo.org/record/7663790/files/figure.png" pageId="4" pageNumber="432">7G, H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="432">
|
||
Thoracic sternum relatively smooth, setose; sternites 1 and 2 completely fused to form triangular structure; sternite 2 and 3 separated by distinctly convex suture; sternites 3 and 4 fused but shallow oblique lateral and submedian depression demarcating sternites visible; sternopleonal cavity reaching imaginary line connecting proximal edges of coxae of chelipeds; sternite 8 exposed when pleon closed, visible as subquadrate plate (
|
||
<figureCitation box="[1120,1243,1591,1617]" captionStart="FIGURE 9" captionStartId="14.[151,250,1509,1533]" captionTargetBox="[176,1410,181,1485]" captionTargetId="figure-17@14.[176,1410,181,1485]" captionTargetPageId="14" captionText="FIGURE 9. A–D, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia. A, right lateral view of posterior part of sub-branchial region; B, E–G, anterior male thoracic sternum and pleon; C, male pleonal somites 1–4; D, male pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.7663797" httpUri="https://zenodo.org/record/7663797/files/figure.png" pageId="4" pageNumber="432">Fig. 9B–D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="432">
|
||
Pleon triangular, transversely narrow, especially along somites 4–6; all somites and telson free; somite 1 widest, reaching to P5 coxae; somite 2 less than somites 1 and 3; somite 3 wide, reaching condyles of P5 coxae; somites 4 and 5 trapezoidal with gently concave margins; somite 6 rectangular; telson triangular, slightly longer than wide with rounded tip and gently sinuous lateral margins (
|
||
<figureCitation box="[742,875,1735,1761]" captionStart="FIGURE 9" captionStartId="14.[151,250,1509,1533]" captionTargetBox="[176,1410,181,1485]" captionTargetId="figure-17@14.[176,1410,181,1485]" captionTargetPageId="14" captionText="FIGURE 9. A–D, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia. A, right lateral view of posterior part of sub-branchial region; B, E–G, anterior male thoracic sternum and pleon; C, male pleonal somites 1–4; D, male pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.7663797" httpUri="https://zenodo.org/record/7663797/files/figure.png" pageId="4" pageNumber="432">Figs. 9B–D</figureCitation>
|
||
,
|
||
<figureCitation box="[886,938,1735,1761]" captionStart="FIGURE 10" captionStartId="15.[151,247,1934,1958]" captionTargetBox="[227,1360,463,1902]" captionTargetId="figure-158@15.[212,1374,453,1911]" captionTargetPageId="15" captionText="FIGURE 10.A–E, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; F–J, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok. A, B, pleon; B, G, left G1 (ventral view); C, H, distal part of left G1 (ventral view); D, I, left G1 (dorsal view); E, J, left G2. Scales:A, F = 1.0 mm; B, E, G, J = 0.5 mm; C, D, H, I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7663799" httpUri="https://zenodo.org/record/7663799/files/figure.png" pageId="4" pageNumber="432">10A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,150,2013]" box="[199,1418,1771,1797]" pageId="4" pageNumber="432">
|
||
G1 strongly sinuous, distal part long, gently curved, elongate (
|
||
<figureCitation box="[898,1036,1771,1797]" captionStart="FIGURE 10" captionStartId="15.[151,247,1934,1958]" captionTargetBox="[227,1360,463,1902]" captionTargetId="figure-158@15.[212,1374,453,1911]" captionTargetPageId="15" captionText="FIGURE 10.A–E, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; F–J, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok. A, B, pleon; B, G, left G1 (ventral view); C, H, distal part of left G1 (ventral view); D, I, left G1 (dorsal view); E, J, left G2. Scales:A, F = 1.0 mm; B, E, G, J = 0.5 mm; C, D, H, I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7663799" httpUri="https://zenodo.org/record/7663799/files/figure.png" pageId="4" pageNumber="432">Fig. 10B–D</figureCitation>
|
||
); G2 short spatuliform (
|
||
<figureCitation box="[1304,1406,1771,1797]" captionStart="FIGURE 10" captionStartId="15.[151,247,1934,1958]" captionTargetBox="[227,1360,463,1902]" captionTargetId="figure-158@15.[212,1374,453,1911]" captionTargetPageId="15" captionText="FIGURE 10.A–E, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; F–J, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok. A, B, pleon; B, G, left G1 (ventral view); C, H, distal part of left G1 (ventral view); D, I, left G1 (dorsal view); E, J, left G2. Scales:A, F = 1.0 mm; B, E, G, J = 0.5 mm; C, D, H, I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7663799" httpUri="https://zenodo.org/record/7663799/files/figure.png" pageId="4" pageNumber="432">Fig. 10E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="432">
|
||
<emphasis bold="true" box="[199,297,1807,1833]" pageId="4" pageNumber="432">Females</emphasis>
|
||
. Adult female specimens have a longitudinally ovate pleon that covers most of thoracic sternum, telson semicircular (
|
||
<figureCitation box="[304,394,1843,1869]" captionStart="FIGURE 8" captionStartId="13.[151,250,1105,1129]" captionTargetBox="[151,1435,181,1081]" captionTargetId="figure-25@13.[151,1435,181,1081]" captionTargetPageId="13" captionText="FIGURE 8. A, B, Lophoplax sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, female thoracic sternum and pleon; B–D, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.7663793" httpUri="https://zenodo.org/record/7663793/files/figure.png" pageId="4" pageNumber="432">Fig. 8A</figureCitation>
|
||
); vulva large, ovate, obliquely positioned, occupying proximal three-quarters of sternite 6, no operculum (
|
||
<figureCitation box="[286,374,1879,1905]" captionStart="FIGURE 8" captionStartId="13.[151,250,1105,1129]" captionTargetBox="[151,1435,181,1081]" captionTargetId="figure-25@13.[151,1435,181,1081]" captionTargetPageId="13" captionText="FIGURE 8. A, B, Lophoplax sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa; D, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, female thoracic sternum and pleon; B–D, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.7663793" httpUri="https://zenodo.org/record/7663793/files/figure.png" pageId="4" pageNumber="432">Fig. 8B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,150,2013]" pageId="4" pageNumber="432">
|
||
<emphasis bold="true" box="[199,357,1914,1940]" pageId="4" pageNumber="432">Colour in life</emphasis>
|
||
. Background colour of carapace and pereopods white, with brown setae; epigastric areolets red; protogastric areolets bright to pale red; hepatic areolets white to orange; cardiac areolets pale yellow to dirty white (
|
||
<figureCitation box="[159,281,1987,2013]" captionStart="FIGURE 1" captionStartId="3.[151,250,1724,1748]" captionTargetBox="[151,1435,561,1701]" captionTargetId="figure-203@3.[151,1435,561,1701]" captionTargetPageId="3" captionText="FIGURE 1. Colour in life. A, Lophoplax sculpta (Stimpson, 1858), male (10.2 × 8.2 mm) (RUMF-ZC-5207), Okinawa [photograph:T.Maenosono]; B, L.sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines [photograph: T.-Y. Chan]; C, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok [photograph: A. Anker]; D, L. vermiculata (A. Milne-Edwards, 1873), male, New Caledonia (not examined) [photograph: G. Paulay]; E, L. vermiculata (A. Milne-Edwards, 1873), male (14.1 × 10.8 mm) (UF 388), Guam [photograph: G. Paulay]; F, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]." figureDoi="http://doi.org/10.5281/zenodo.7663770" httpUri="https://zenodo.org/record/7663770/files/figure.png" pageId="4" pageNumber="432">Fig. 1A, B</figureCitation>
|
||
;
|
||
<bibRefCitation author="Maenosono, T." box="[294,491,1987,2013]" pageId="4" pageNumber="432" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
|
||
: fig. 8A).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="8" lastPageNumber="436" pageId="5" pageNumber="433" type="discussion">
|
||
<paragraph blockId="5.[151,1437,151,285]" pageId="5" pageNumber="433">
|
||
<emphasis bold="true" box="[199,308,151,177]" pageId="5" pageNumber="433">Remarks</emphasis>
|
||
. The taxonomy of
|
||
<taxonomicName baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[525,636,151,177]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis box="[525,636,151,177]" italics="true" pageId="5" pageNumber="433">L. sculpta</emphasis>
|
||
</taxonomicName>
|
||
has been reviewed by
|
||
<bibRefCitation author="Tesch, J. J." box="[896,1045,151,177]" pageId="5" pageNumber="433" pagination="149 - 295" refId="ref18770" refString="Tesch, J. J. (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1, 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch (1918)</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Takeda, M." box="[1058,1220,151,177]" pageId="5" pageNumber="433" pagination="120 - 123" refId="ref18577" refString="Takeda, M. (1977) Rediscovery of Lophoplax sculpta (Stimpson, 1858) (Crustacea, Decapoda, Brachyura). Zoological Magazine, 86, 120 - 123." type="journal article" year="1977">Takeda (1977)</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Ng, P. K. L." box="[1277,1396,151,177]" pageId="5" pageNumber="433" pagination="69 - 111" refId="ref16944" refString="Ng, P. K. L. (1987) The Indo-Pacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). Indo-Malayan Zoology, 1987, 4 (1), 69 - 111, pl. 1." type="journal article" year="1987">Ng (1987)</bibRefCitation>
|
||
. Its original placement in
|
||
<taxonomicName authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[403,737,186,213]" class="Malacostraca" family="Pilumnidae" genus="Pilumnoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="433" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[403,546,187,213]" italics="true" pageId="5" pageNumber="433">Pilumnoplax</emphasis>
|
||
<bibRefCitation author="Stimpson, W." box="[556,737,186,213]" pageId="5" pageNumber="433" pagination="93 - 110" refId="ref18446" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 (39 - 56). https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
|
||
</taxonomicName>
|
||
, is not tenable as that genus is now a junior synonym of the euryplacid
|
||
<taxonomicName authority="De Haan, 1835" authorityName="De Haan" authorityYear="1835" box="[277,543,222,249]" class="Malacostraca" family="Euryplacidae" genus="Eucrate" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="433" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[277,366,223,248]" italics="true" pageId="5" pageNumber="433">Eucrate</emphasis>
|
||
De Haan, 1835
|
||
</taxonomicName>
|
||
(cf.
|
||
<bibRefCitation author="Castro, P. & Ng, P. K. L." box="[596,809,223,249]" pageId="5" pageNumber="433" pagination="1 - 130" refId="ref14630" refString="Castro, P. & Ng, P. K. L. (2010) Revision of the family Euryplacidae Stimpson, 1871 (Crustacea: Decapoda: Brachyura: Goneplacoidea). Zootaxa, 2375 (1), 1 - 130. https: // doi. org / 10.11646 / zootaxa. 2375.1.1" type="journal article" year="2010">Castro & Ng 2010</bibRefCitation>
|
||
). The present specimens agree very well with what was described and figured in
|
||
<bibRefCitation author="Takeda, M." box="[429,591,259,285]" pageId="5" pageNumber="433" pagination="120 - 123" refId="ref18577" refString="Takeda, M. (1977) Rediscovery of Lophoplax sculpta (Stimpson, 1858) (Crustacea, Decapoda, Brachyura). Zoological Magazine, 86, 120 - 123." type="journal article" year="1977">Takeda (1977)</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Maenosono, T." box="[646,861,259,285]" pageId="5" pageNumber="433" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono (2019)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7663772" ID-Zenodo-Dep="7663772" httpUri="https://zenodo.org/record/7663772/files/figure.png" pageId="5" pageNumber="433" startId="5.[151,250,1775,1799]" targetBox="[151,1435,309,1752]" targetPageId="5">
|
||
<paragraph blockId="5.[151,1437,1775,2014]" pageId="5" pageNumber="433">
|
||
<emphasis bold="true" box="[151,277,1775,1799]" pageId="5" pageNumber="433">FIGURE 2.</emphasis>
|
||
Overall dorsal view. A,
|
||
<taxonomicName authority="(Stimpson, 1858)" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[535,915,1775,1799]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis box="[535,725,1775,1799]" italics="true" pageId="5" pageNumber="433">Lophoplax sculpta</emphasis>
|
||
(
|
||
<bibRefCitation author="Stimpson, W." box="[741,907,1775,1799]" pageId="5" pageNumber="433" pagination="93 - 110" refId="ref18446" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 (39 - 56). https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B,
|
||
<taxonomicName authority="(Stimpson, 1858)" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[285,569,1811,1835]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis box="[285,384,1811,1835]" italics="true" pageId="5" pageNumber="433">L. sculpta</emphasis>
|
||
(
|
||
<bibRefCitation author="Stimpson, W." box="[397,561,1811,1835]" pageId="5" pageNumber="433" pagination="93 - 110" refId="ref18446" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 (39 - 56). https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C,
|
||
<taxonomicName authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[1171,1274,1811,1835]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="sordida" status="n. sp.">
|
||
<emphasis box="[1171,1274,1811,1835]" italics="true" pageId="5" pageNumber="433">L. sordida</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1280,1337,1811,1835]" pageId="5" pageNumber="433">
|
||
<taxonomicNameLabel box="[1280,1337,1811,1835]" pageId="5" pageNumber="433" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; D,
|
||
<taxonomicName authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[666,769,1847,1871]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="sordida" status="n. sp.">
|
||
<emphasis box="[666,769,1847,1871]" italics="true" pageId="5" pageNumber="433">L. sordida</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[776,833,1847,1871]" pageId="5" pageNumber="433">
|
||
<taxonomicNameLabel box="[776,833,1847,1871]" pageId="5" pageNumber="433" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; E,
|
||
<taxonomicName authority="(A. Milne-Edwards, 1873)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[180,608,1883,1907]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="vermiculata">
|
||
<emphasis box="[180,328,1883,1907]" italics="true" pageId="5" pageNumber="433">L. vermiculata</emphasis>
|
||
(A.
|
||
<bibRefCitation author="Milne-Edwards, A." box="[375,600,1883,1907]" pageId="5" pageNumber="433" pagination="155 - 332" refId="ref16688" refString="Milne-Edwards, A. (1873) Recherches sur la faune carcinologique de la Nouvelle-Caledonie. Deuxieme Partie. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 9, 155 - 332, pls. 4 - 18. https: // doi. org / 10.5962 / bhl. title. 10410" type="journal article" year="1873">Milne-Edwards, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; F,
|
||
<taxonomicName baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="vermiculata">
|
||
<emphasis italics="true" pageId="5" pageNumber="433">Lophoplax vermiculata</emphasis>
|
||
</taxonomicName>
|
||
, subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G,
|
||
<taxonomicName authority="(A. Milne-Edwards, 1873)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[879,1300,1919,1943]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="vermiculata">
|
||
<emphasis box="[879,1026,1919,1943]" italics="true" pageId="5" pageNumber="433">L. vermiculata</emphasis>
|
||
(A.
|
||
<bibRefCitation author="Milne-Edwards, A." box="[1069,1293,1919,1943]" pageId="5" pageNumber="433" pagination="155 - 332" refId="ref16688" refString="Milne-Edwards, A. (1873) Recherches sur la faune carcinologique de la Nouvelle-Caledonie. Deuxieme Partie. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 9, 155 - 332, pls. 4 - 18. https: // doi. org / 10.5962 / bhl. title. 10410" type="journal article" year="1873">Milne-Edwards, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa; H,
|
||
<taxonomicName authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[605,720,1956,1979]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="433" phylum="Arthropoda" rank="species" species="pannosa" status="n. sp.">
|
||
<emphasis box="[605,720,1956,1979]" italics="true" pageId="5" pageNumber="433">L. pannosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[728,786,1955,1979]" pageId="5" pageNumber="433">
|
||
<taxonomicNameLabel box="[728,786,1955,1979]" pageId="5" pageNumber="433" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7663776" ID-Zenodo-Dep="7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="6" pageNumber="434" startId="6.[151,250,1744,1768]" targetBox="[153,1433,181,1720]" targetPageId="6">
|
||
<paragraph blockId="6.[151,1437,1744,1913]" pageId="6" pageNumber="434">
|
||
<emphasis bold="true" box="[151,274,1744,1769]" pageId="6" pageNumber="434">FIGURE 3.</emphasis>
|
||
Dorsal view of carapace. A,
|
||
<taxonomicName authority="(Stimpson, 1858)" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[564,935,1745,1769]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis box="[564,751,1745,1769]" italics="true" pageId="6" pageNumber="434">Lophoplax sculpta</emphasis>
|
||
(
|
||
<bibRefCitation author="Stimpson, W." box="[764,927,1745,1769]" pageId="6" pageNumber="434" pagination="93 - 110" refId="ref18446" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 (39 - 56). https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B,
|
||
<taxonomicName authority="(Stimpson, 1858)" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[287,573,1781,1805]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis box="[287,387,1781,1805]" italics="true" pageId="6" pageNumber="434">L. sculpta</emphasis>
|
||
(
|
||
<bibRefCitation author="Stimpson, W." box="[401,565,1781,1805]" pageId="6" pageNumber="434" pagination="93 - 110" refId="ref18446" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 (39 - 56). https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C,
|
||
<taxonomicName authority="(Stimpson, 1858)" baseAuthorityName="Stimpson" baseAuthorityYear="1858" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis italics="true" pageId="6" pageNumber="434">L. sculpta</emphasis>
|
||
(
|
||
<bibRefCitation author="Stimpson, W." box="[237,401,1817,1841]" pageId="6" pageNumber="434" pagination="93 - 110" refId="ref18446" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 (39 - 56). https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D,
|
||
<taxonomicName authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[1012,1115,1817,1841]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="sordida" status="n. sp.">
|
||
<emphasis box="[1012,1115,1817,1841]" italics="true" pageId="6" pageNumber="434">L. sordida</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1121,1178,1817,1841]" pageId="6" pageNumber="434">
|
||
<taxonomicNameLabel box="[1121,1178,1817,1841]" pageId="6" pageNumber="434" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E,
|
||
<taxonomicName authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[502,605,1853,1877]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="sordida" status="n. sp.">
|
||
<emphasis box="[502,605,1853,1877]" italics="true" pageId="6" pageNumber="434">L. sordida</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[612,670,1853,1877]" pageId="6" pageNumber="434">
|
||
<taxonomicNameLabel box="[612,670,1853,1877]" pageId="6" pageNumber="434" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F,
|
||
<taxonomicName authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[1306,1409,1853,1877]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="434" phylum="Arthropoda" rank="species" species="sordida" status="n. sp.">
|
||
<emphasis box="[1306,1409,1853,1877]" italics="true" pageId="6" pageNumber="434">L. sordida</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="6" pageNumber="434">
|
||
<taxonomicNameLabel pageId="6" pageNumber="434" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="7.[151,1436,150,393]" pageId="7" pageNumber="435">
|
||
<taxonomicName baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[199,406,151,177]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis box="[199,406,151,177]" italics="true" pageId="7" pageNumber="435">Lophoplax sculpta</emphasis>
|
||
</taxonomicName>
|
||
was described from at least
|
||
<specimenCount box="[717,839,151,177]" pageId="7" pageNumber="435" type="female">one female</specimenCount>
|
||
specimen (cf.
|
||
<bibRefCitation author="Stimpson, W." box="[1000,1212,150,177]" pageId="7" pageNumber="435" pagination="93 - 110" refId="ref18446" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 (39 - 56). https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson 1858: 93</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Stimpson, W." box="[1223,1321,151,177]" pageId="7" pageNumber="435" pagination="1 - 240" refId="ref18518" refString="Stimpson, W. (1907) Report on the Crustacea (Brachyura and Anomura) collected by the North Pacific Exploring Expedition 1853 - 1856. Smithsonian Miscellaneous Collections, Part of 49 (1717), 1 - 240, 26 pls. https: // doi. org / 10.5962 / bhl. title. 51448" type="journal article" year="1907">1907: 91</bibRefCitation>
|
||
) collected from Ousima, Ryukyu Islands, in
|
||
<collectingCountry box="[548,615,187,213]" name="Japan" pageId="7" pageNumber="435">Japan</collectingCountry>
|
||
, with one measurement provided: 7.1 by
|
||
<quantity box="[1103,1195,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="7" pageNumber="435" unit="mm" value="5.6">5.6 mm</quantity>
|
||
. The specimen is no longer extant (cf.
|
||
<bibRefCitation author="Evans, A. C." box="[351,486,223,249]" pageId="7" pageNumber="435" pagination="399 - 411" refId="ref15266" refString="Evans, A. C. (1967) Syntypes of Decapoda described by William Stimpson and James Dana in the collections of the British Museum (Natural History). Journal of Natural History, 1, 399 - 411. https: // doi. org / 10.1080 / 00222936700770391" type="journal article" year="1967">Evans 1967</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Deiss, W. A. & Manning, R. B." box="[498,764,223,249]" pageId="7" pageNumber="435" pagination="79 - 85" refId="ref15141" refString="Deiss, W. A. & Manning, R. B. (1981) The fate of the invertebrate collections of the North Pacific Exploring Expedition, 1853 - 1856. In: Wheeler, A. & Price, J. H. (Eds.), History in the Service of Systematics. Society for the Bibliography of Natural History, London, pp. 79 - 85. https: // doi. org / 10.3366 / anh. 1981.012" type="book chapter" year="1981">Deiss & Manning 1981</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Manning, R. B." box="[777,944,223,249]" pageId="7" pageNumber="435" pagination="109 - 117" refId="ref16215" refString="Manning, R. B. (1993) The scientific contributions of William Stimpson, an early American naturalist and taxonomist. In: Truesdale, F. (Ed.), History of Carcinology. Crustacean Issues 8. A. A. Balkema, Rotterdam, Brookfield, pp. 109 - 117. https: // doi. org / 10.1201 / 9781003077633 - 8" type="book chapter" year="1993">Manning 1993</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Manning, R. B. & Reed, K. J." box="[956,1220,223,249]" pageId="7" pageNumber="435" pagination="283 - 293" refId="ref16285" refString="Manning, R. B. & Reed, K. J. (2006) Decapod crustaceans deposited in the Zoological Museum of Copenhagen by William Stimpson in 1859. Raffles Bulletin of Zoology, 54 (2), 283 - 293." type="journal article" year="2006">Manning & Reed 2006</bibRefCitation>
|
||
). While the species is distinctive, the descriptions of two new allied species from
|
||
<collectingCountry box="[839,948,259,285]" name="Indonesia" pageId="7" pageNumber="435">Indonesia</collectingCountry>
|
||
and
|
||
<collectingCountry box="[1001,1093,259,285]" name="Vanuatu" pageId="7" pageNumber="435">Vanuatu</collectingCountry>
|
||
requires stability in this name, and a
|
||
<typeStatus box="[224,313,295,321]" pageId="7" pageNumber="435">neotype</typeStatus>
|
||
is therefore needed. We therefore designate the male specimen examined here as the
|
||
<typeStatus box="[1314,1403,295,321]" pageId="7" pageNumber="435">neotype</typeStatus>
|
||
of
|
||
<taxonomicName authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[151,567,330,357]" class="Malacostraca" family="Pilumnidae" genus="Pilumnoplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis box="[151,383,331,357]" italics="true" pageId="7" pageNumber="435">Pilumnoplax sculpta</emphasis>
|
||
<bibRefCitation author="Stimpson, W." box="[390,567,330,357]" pageId="7" pageNumber="435" pagination="93 - 110" refId="ref18446" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 (39 - 56). https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[583,683,331,357]" captionStart="FIGURE 2" captionStartId="5.[151,250,1775,1799]" captionTargetBox="[151,1435,309,1752]" captionTargetId="figure-91@5.[151,1435,309,1752]" captionTargetPageId="5" captionText="FIGURE 2. Overall dorsal view. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; D, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; E, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; F, Lophoplax vermiculata, subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa; H, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663772" httpUri="https://zenodo.org/record/7663772/files/figure.png" pageId="7" pageNumber="435">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation box="[695,732,331,357]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="7" pageNumber="435">3A</figureCitation>
|
||
,
|
||
<figureCitation box="[744,781,330,357]" captionStart="FIGURE 5" captionStartId="10.[151,250,1835,1859]" captionTargetBox="[151,1435,237,1812]" captionTargetId="figure-47@10.[151,1435,237,1812]" captionTargetPageId="10" captionText="FIGURE 5. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A–F, frontal view of cephalothorax; G–J, right third maxilliped; K, L, left third maxilliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663782" httpUri="https://zenodo.org/record/7663782/files/figure.png" pageId="7" pageNumber="435">5A</figureCitation>
|
||
,
|
||
<figureCitation box="[793,830,331,357]" captionStart="FIGURE 6" captionStartId="11.[151,250,1726,1750]" captionTargetBox="[151,1436,813,1703]" captionTargetId="figure-327@11.[151,1436,813,1703]" captionTargetPageId="11" captionText="FIGURE 6. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu.A, B, D, E, dorsal view of right cheliped; C, dorsal view of left cheliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663784" httpUri="https://zenodo.org/record/7663784/files/figure.png" pageId="7" pageNumber="435">6A</figureCitation>
|
||
,
|
||
<figureCitation box="[842,914,331,357]" captionStart="FIGURE 7" captionStartId="12.[151,250,1823,1847]" captionTargetBox="[229,1359,181,1799]" captionTargetId="figure-17@12.[229,1359,181,1799]" captionTargetPageId="12" captionText="FIGURE 7. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, B, right chela; C–F, left chela; G–L, right P4." figureDoi="http://doi.org/10.5281/zenodo.7663790" httpUri="https://zenodo.org/record/7663790/files/figure.png" pageId="7" pageNumber="435">7A, G</figureCitation>
|
||
,
|
||
<figureCitation box="[926,997,331,357]" captionStart="FIGURE 9" captionStartId="14.[151,250,1509,1533]" captionTargetBox="[176,1410,181,1485]" captionTargetId="figure-17@14.[176,1410,181,1485]" captionTargetPageId="14" captionText="FIGURE 9. A–D, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia. A, right lateral view of posterior part of sub-branchial region; B, E–G, anterior male thoracic sternum and pleon; C, male pleonal somites 1–4; D, male pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.7663797" httpUri="https://zenodo.org/record/7663797/files/figure.png" pageId="7" pageNumber="435">9A–D</figureCitation>
|
||
,
|
||
<figureCitation box="[1009,1093,331,357]" captionStart="FIGURE 10" captionStartId="15.[151,247,1934,1958]" captionTargetBox="[227,1360,463,1902]" captionTargetId="figure-158@15.[212,1374,453,1911]" captionTargetPageId="15" captionText="FIGURE 10.A–E, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; F–J, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok. A, B, pleon; B, G, left G1 (ventral view); C, H, distal part of left G1 (ventral view); D, I, left G1 (dorsal view); E, J, left G2. Scales:A, F = 1.0 mm; B, E, G, J = 0.5 mm; C, D, H, I = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7663799" httpUri="https://zenodo.org/record/7663799/files/figure.png" pageId="7" pageNumber="435">10A–E</figureCitation>
|
||
). It is in good condition and is also from the Ryukyu Islands.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7663778" ID-Zenodo-Dep="7663778" httpUri="https://zenodo.org/record/7663778/files/figure.png" pageId="7" pageNumber="435" startId="7.[151,250,1454,1478]" targetBox="[151,1436,417,1430]" targetPageId="7">
|
||
<paragraph blockId="7.[151,1437,1454,1586]" pageId="7" pageNumber="435">
|
||
<emphasis bold="true" box="[151,278,1454,1478]" pageId="7" pageNumber="435">FIGURE 4.</emphasis>
|
||
Dorsal view of carapace. A,
|
||
<taxonomicName authority="(A. Milne-Edwards, 1873)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[587,1023,1454,1478]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="vermiculata">
|
||
<emphasis box="[587,738,1454,1478]" italics="true" pageId="7" pageNumber="435">L. vermiculata</emphasis>
|
||
(A.
|
||
<bibRefCitation author="Milne-Edwards, A." box="[789,1016,1454,1478]" pageId="7" pageNumber="435" pagination="155 - 332" refId="ref16688" refString="Milne-Edwards, A. (1873) Recherches sur la faune carcinologique de la Nouvelle-Caledonie. Deuxieme Partie. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 9, 155 - 332, pls. 4 - 18. https: // doi. org / 10.5962 / bhl. title. 10410" type="journal article" year="1873">Milne-Edwards, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; B,
|
||
<taxonomicName authority="(A. Milne-Edwards, 1873)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[477,913,1490,1514]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="vermiculata">
|
||
<emphasis box="[477,627,1490,1514]" italics="true" pageId="7" pageNumber="435">L. vermiculata</emphasis>
|
||
(A.
|
||
<bibRefCitation author="Milne-Edwards, A." box="[678,905,1490,1514]" pageId="7" pageNumber="435" pagination="155 - 332" refId="ref16688" refString="Milne-Edwards, A. (1873) Recherches sur la faune carcinologique de la Nouvelle-Caledonie. Deuxieme Partie. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 9, 155 - 332, pls. 4 - 18. https: // doi. org / 10.5962 / bhl. title. 10410" type="journal article" year="1873">Milne-Edwards, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; C,
|
||
<taxonomicName authority="(A. Milne-Edwards, 1873)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[262,700,1526,1550]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="vermiculata">
|
||
<emphasis box="[262,413,1526,1550]" italics="true" pageId="7" pageNumber="435">L. vermiculata</emphasis>
|
||
(A.
|
||
<bibRefCitation author="Milne-Edwards, A." box="[465,693,1526,1550]" pageId="7" pageNumber="435" pagination="155 - 332" refId="ref16688" refString="Milne-Edwards, A. (1873) Recherches sur la faune carcinologique de la Nouvelle-Caledonie. Deuxieme Partie. Nouvelles Archives du Museum d'Histoire naturelle, Paris, 9, 155 - 332, pls. 4 - 18. https: // doi. org / 10.5962 / bhl. title. 10410" type="journal article" year="1873">Milne-Edwards, 1873</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa [photograph: T. Maenosono]; D,
|
||
<taxonomicName authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[322,435,1563,1586]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="pannosa" status="n. sp.">
|
||
<emphasis box="[322,435,1563,1586]" italics="true" pageId="7" pageNumber="435">L. pannosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[441,498,1562,1586]" pageId="7" pageNumber="435">
|
||
<taxonomicNameLabel box="[441,498,1562,1586]" pageId="7" pageNumber="435" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="7.[151,1437,1632,2019]" lastBlockId="8.[151,1437,151,825]" lastPageId="8" lastPageNumber="436" pageId="7" pageNumber="435">
|
||
<taxonomicName baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[199,405,1632,1658]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis box="[199,405,1632,1658]" italics="true" pageId="7" pageNumber="435">Lophoplax sculpta</emphasis>
|
||
</taxonomicName>
|
||
most closely resembles
|
||
<taxonomicName baseAuthorityName="Deb" baseAuthorityYear="1989" box="[671,836,1632,1658]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="andamanica">
|
||
<emphasis box="[671,836,1632,1658]" italics="true" pageId="7" pageNumber="435">L. andamanica</emphasis>
|
||
</taxonomicName>
|
||
in carapace features, but as already discussed at length by
|
||
<bibRefCitation author="Trivedi, J. & Patel, K. & Mitra, S. & Ng, P. K. L." box="[186,418,1668,1695]" pageId="7" pageNumber="435" pagination="595 - 600" refId="ref18894" refString="Trivedi, J., Patel, K., Mitra, S. & Ng, P. K. L. (2022) On the identity of Myopilumnus andamanicus Deb, 1989 (Crustacea: Decapoda: Brachyura: Pilumnidae) from India. Zootaxa, 5194 (4), 595 - 600. https: // doi. org / 10.11646 / zootaxa. 5194.4.8" type="journal article" year="2022">
|
||
Trivedi
|
||
<emphasis box="[276,333,1668,1694]" italics="true" pageId="7" pageNumber="435">et al.</emphasis>
|
||
(2022)
|
||
</bibRefCitation>
|
||
,
|
||
<taxonomicName baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[430,541,1668,1694]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis box="[430,541,1668,1694]" italics="true" pageId="7" pageNumber="435">L. sculpta</emphasis>
|
||
</taxonomicName>
|
||
can easily be separated by the epigastric areolet being more ovate (
|
||
<figureCitation box="[1307,1428,1668,1695]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="7" pageNumber="435">Fig. 3A–C</figureCitation>
|
||
) (versus epigastric areolet quadrilateral in shape in
|
||
<taxonomicName baseAuthorityName="Deb" baseAuthorityYear="1989" box="[732,902,1704,1730]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="andamanica">
|
||
<emphasis box="[732,902,1704,1730]" italics="true" pageId="7" pageNumber="435">L. andamanica</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation author="Trivedi, J. & Patel, K. & Mitra, S. & Ng, P. K. L." box="[919,1137,1704,1731]" pageId="7" pageNumber="435" pagination="595 - 600" refId="ref18894" refString="Trivedi, J., Patel, K., Mitra, S. & Ng, P. K. L. (2022) On the identity of Myopilumnus andamanicus Deb, 1989 (Crustacea: Decapoda: Brachyura: Pilumnidae) from India. Zootaxa, 5194 (4), 595 - 600. https: // doi. org / 10.11646 / zootaxa. 5194.4.8" type="journal article" year="2022">
|
||
Trivedi
|
||
<emphasis box="[1010,1068,1704,1730]" italics="true" pageId="7" pageNumber="435">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
: fig. 1A); the protogastric areolet is transversely ovate to weakly crescent-shaped in
|
||
<taxonomicName baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[799,909,1740,1766]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="sculpta">
|
||
<emphasis box="[799,909,1740,1766]" italics="true" pageId="7" pageNumber="435">L. sculpta</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[924,1044,1740,1767]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="7" pageNumber="435">Fig. 3A–C</figureCitation>
|
||
) (versus distinctly crescent-shaped in
|
||
<taxonomicName baseAuthorityName="Deb" baseAuthorityYear="1989" box="[181,349,1776,1802]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="andamanica">
|
||
<emphasis box="[181,349,1776,1802]" italics="true" pageId="7" pageNumber="435">L. andamanica</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation author="Trivedi, J. & Patel, K. & Mitra, S. & Ng, P. K. L." box="[364,577,1776,1803]" pageId="7" pageNumber="435" pagination="595 - 600" refId="ref18894" refString="Trivedi, J., Patel, K., Mitra, S. & Ng, P. K. L. (2022) On the identity of Myopilumnus andamanicus Deb, 1989 (Crustacea: Decapoda: Brachyura: Pilumnidae) from India. Zootaxa, 5194 (4), 595 - 600. https: // doi. org / 10.11646 / zootaxa. 5194.4.8" type="journal article" year="2022">
|
||
Trivedi
|
||
<emphasis box="[453,510,1776,1802]" italics="true" pageId="7" pageNumber="435">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
: fig. 1A); the hepatic region is proportionately shorter, reaching only to first anterolateral tooth (
|
||
<figureCitation box="[373,494,1812,1839]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="7" pageNumber="435">Fig. 3A–C</figureCitation>
|
||
) (versus distinctly longer, reaching to second anterolateral tooth in
|
||
<taxonomicName baseAuthorityName="Deb" baseAuthorityYear="1989" box="[1259,1428,1812,1838]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="andamanica">
|
||
<emphasis box="[1259,1428,1812,1838]" italics="true" pageId="7" pageNumber="435">L. andamanica</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation author="Trivedi, J. & Patel, K. & Mitra, S. & Ng, P. K. L." box="[151,368,1848,1875]" pageId="7" pageNumber="435" pagination="595 - 600" refId="ref18894" refString="Trivedi, J., Patel, K., Mitra, S. & Ng, P. K. L. (2022) On the identity of Myopilumnus andamanicus Deb, 1989 (Crustacea: Decapoda: Brachyura: Pilumnidae) from India. Zootaxa, 5194 (4), 595 - 600. https: // doi. org / 10.11646 / zootaxa. 5194.4.8" type="journal article" year="2022">
|
||
Trivedi
|
||
<emphasis box="[241,299,1848,1874]" italics="true" pageId="7" pageNumber="435">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
: fig. 1A); the tips of the external orbital and anterolateral teeth are sharp (
|
||
<figureCitation box="[1216,1339,1848,1875]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="7" pageNumber="435">Fig. 3A–C</figureCitation>
|
||
) (versus first three anterolateral teeth proportionately broader with the blunt tips in
|
||
<taxonomicName baseAuthorityName="Deb" baseAuthorityYear="1989" box="[991,1159,1884,1910]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="andamanica">
|
||
<emphasis box="[991,1159,1884,1910]" italics="true" pageId="7" pageNumber="435">L. andamanica</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation author="Trivedi, J. & Patel, K. & Mitra, S. & Ng, P. K. L." box="[1174,1387,1884,1911]" pageId="7" pageNumber="435" pagination="595 - 600" refId="ref18894" refString="Trivedi, J., Patel, K., Mitra, S. & Ng, P. K. L. (2022) On the identity of Myopilumnus andamanicus Deb, 1989 (Crustacea: Decapoda: Brachyura: Pilumnidae) from India. Zootaxa, 5194 (4), 595 - 600. https: // doi. org / 10.11646 / zootaxa. 5194.4.8" type="journal article" year="2022">
|
||
Trivedi
|
||
<emphasis box="[1263,1320,1884,1910]" italics="true" pageId="7" pageNumber="435">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
: fig. 1A); the epistome is proportionately slightly wider (
|
||
<figureCitation box="[728,846,1920,1947]" captionStart="FIGURE 5" captionStartId="10.[151,250,1835,1859]" captionTargetBox="[151,1435,237,1812]" captionTargetId="figure-47@10.[151,1435,237,1812]" captionTargetPageId="10" captionText="FIGURE 5. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A–F, frontal view of cephalothorax; G–J, right third maxilliped; K, L, left third maxilliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663782" httpUri="https://zenodo.org/record/7663782/files/figure.png" pageId="7" pageNumber="435">Fig. 5A, B</figureCitation>
|
||
) (versus proportionately narrower in
|
||
<taxonomicName baseAuthorityName="Deb" baseAuthorityYear="1989" box="[1261,1428,1920,1946]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="435" phylum="Arthropoda" rank="species" species="andamanica">
|
||
<emphasis box="[1261,1428,1920,1946]" italics="true" pageId="7" pageNumber="435">L. andamanica</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation author="Trivedi, J. & Patel, K. & Mitra, S. & Ng, P. K. L." box="[151,371,1956,1983]" pageId="7" pageNumber="435" pagination="595 - 600" refId="ref18894" refString="Trivedi, J., Patel, K., Mitra, S. & Ng, P. K. L. (2022) On the identity of Myopilumnus andamanicus Deb, 1989 (Crustacea: Decapoda: Brachyura: Pilumnidae) from India. Zootaxa, 5194 (4), 595 - 600. https: // doi. org / 10.11646 / zootaxa. 5194.4.8" type="journal article" year="2022">
|
||
Trivedi
|
||
<emphasis box="[243,302,1956,1982]" italics="true" pageId="7" pageNumber="435">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
: fig. 1B); the outer surface of the carpus of the cheliped has 2 large tubercles on the dorsal surface and 2 tubercles on the outer margin (
|
||
<figureCitation box="[668,792,1992,2019]" captionStart="FIGURE 6" captionStartId="11.[151,250,1726,1750]" captionTargetBox="[151,1436,813,1703]" captionTargetId="figure-327@11.[151,1436,813,1703]" captionTargetPageId="11" captionText="FIGURE 6. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu.A, B, D, E, dorsal view of right cheliped; C, dorsal view of left cheliped (laterally transposed for comparative purposes)." figureDoi="http://doi.org/10.5281/zenodo.7663784" httpUri="https://zenodo.org/record/7663784/files/figure.png" pageId="7" pageNumber="435">Fig. 6A, B</figureCitation>
|
||
) (versus outer surface and margin with 6–8 smaller and more dispersed tubercles in
|
||
<taxonomicName baseAuthorityName="Deb" baseAuthorityYear="1989" box="[462,629,151,177]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="436" phylum="Arthropoda" rank="species" species="andamanica">
|
||
<emphasis box="[462,629,151,177]" italics="true" pageId="8" pageNumber="436">L. andamanica</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation author="Trivedi, J. & Patel, K. & Mitra, S. & Ng, P. K. L." box="[643,852,151,177]" pageId="8" pageNumber="436" pagination="595 - 600" refId="ref18894" refString="Trivedi, J., Patel, K., Mitra, S. & Ng, P. K. L. (2022) On the identity of Myopilumnus andamanicus Deb, 1989 (Crustacea: Decapoda: Brachyura: Pilumnidae) from India. Zootaxa, 5194 (4), 595 - 600. https: // doi. org / 10.11646 / zootaxa. 5194.4.8" type="journal article" year="2022">
|
||
Trivedi
|
||
<emphasis box="[731,786,151,177]" italics="true" pageId="8" pageNumber="436">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
: fig. 1C); the cheliped fingers are not distinctly bent, with the pollex subparallel with the ventral margin of palm (
|
||
<figureCitation box="[843,966,187,213]" captionStart="FIGURE 7" captionStartId="12.[151,250,1823,1847]" captionTargetBox="[229,1359,181,1799]" captionTargetId="figure-17@12.[229,1359,181,1799]" captionTargetPageId="12" captionText="FIGURE 7. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, B, right chela; C–F, left chela; G–L, right P4." figureDoi="http://doi.org/10.5281/zenodo.7663790" httpUri="https://zenodo.org/record/7663790/files/figure.png" pageId="8" pageNumber="436">Fig. 7A, B</figureCitation>
|
||
) (versus fingers sharply bent downwards in
|
||
<taxonomicName baseAuthorityName="Deb" baseAuthorityYear="1989" box="[180,348,223,249]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="436" phylum="Arthropoda" rank="species" species="andamanica">
|
||
<emphasis box="[180,348,223,249]" italics="true" pageId="8" pageNumber="436">L. andamanica</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation author="Trivedi, J. & Patel, K. & Mitra, S. & Ng, P. K. L." box="[362,574,223,249]" pageId="8" pageNumber="436" pagination="595 - 600" refId="ref18894" refString="Trivedi, J., Patel, K., Mitra, S. & Ng, P. K. L. (2022) On the identity of Myopilumnus andamanicus Deb, 1989 (Crustacea: Decapoda: Brachyura: Pilumnidae) from India. Zootaxa, 5194 (4), 595 - 600. https: // doi. org / 10.11646 / zootaxa. 5194.4.8" type="journal article" year="2022">
|
||
Trivedi
|
||
<emphasis box="[451,507,223,249]" italics="true" pageId="8" pageNumber="436">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
: fig. 1D); and the outer surface of the P3–P5 merus, carpus and propodus are relatively smoother (
|
||
<figureCitation box="[383,505,259,285]" captionStart="FIGURE 7" captionStartId="12.[151,250,1823,1847]" captionTargetBox="[229,1359,181,1799]" captionTargetId="figure-17@12.[229,1359,181,1799]" captionTargetPageId="12" captionText="FIGURE 7. A, G, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, H, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, I, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; D, J, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; E, K, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; F, L, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu. A, B, right chela; C–F, left chela; G–L, right P4." figureDoi="http://doi.org/10.5281/zenodo.7663790" httpUri="https://zenodo.org/record/7663790/files/figure.png" pageId="8" pageNumber="436">Fig. 7G, H</figureCitation>
|
||
) (versus surfaces distinctly more eroded in
|
||
<taxonomicName baseAuthorityName="Deb" baseAuthorityYear="1989" box="[994,1162,259,285]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="436" phylum="Arthropoda" rank="species" species="andamanica">
|
||
<emphasis box="[994,1162,259,285]" italics="true" pageId="8" pageNumber="436">L. andamanica</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation author="Trivedi, J. & Patel, K. & Mitra, S. & Ng, P. K. L." box="[1176,1387,259,285]" pageId="8" pageNumber="436" pagination="595 - 600" refId="ref18894" refString="Trivedi, J., Patel, K., Mitra, S. & Ng, P. K. L. (2022) On the identity of Myopilumnus andamanicus Deb, 1989 (Crustacea: Decapoda: Brachyura: Pilumnidae) from India. Zootaxa, 5194 (4), 595 - 600. https: // doi. org / 10.11646 / zootaxa. 5194.4.8" type="journal article" year="2022">
|
||
Trivedi
|
||
<emphasis box="[1265,1321,259,285]" italics="true" pageId="8" pageNumber="436">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
: fig. 1F–H). It is also useful to note that
|
||
<taxonomicName baseAuthorityName="Deb" baseAuthorityYear="1989" box="[554,722,295,321]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="436" phylum="Arthropoda" rank="species" species="andamanica">
|
||
<emphasis box="[554,722,295,321]" italics="true" pageId="8" pageNumber="436">L. andamanica</emphasis>
|
||
</taxonomicName>
|
||
is only known from the Andamans in the eastern Indian Ocean and is the only species representing the genus from that ocean thus far.
|
||
</paragraph>
|
||
<paragraph blockId="8.[151,1437,151,825]" pageId="8" pageNumber="436">
|
||
The Philippine specimen (ZRC 2019.1611) differs somewhat from the Japanese specimen and what has been reported by
|
||
<bibRefCitation author="Takeda, M." box="[350,514,403,429]" pageId="8" pageNumber="436" pagination="120 - 123" refId="ref18577" refString="Takeda, M. (1977) Rediscovery of Lophoplax sculpta (Stimpson, 1858) (Crustacea, Decapoda, Brachyura). Zoological Magazine, 86, 120 - 123." type="journal article" year="1977">Takeda (1977)</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Maenosono, T." box="[571,785,403,429]" pageId="8" pageNumber="436" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono (2019)</bibRefCitation>
|
||
in that the carapace is proportionately slightly wider, the protogastric areolets are weakly crescent-shaped rather than ovate, and the two large tubercles at the junction between the antero- and posterolateral margins are prominently larger (
|
||
<figureCitation box="[841,928,475,501]" captionStart="FIGURE 2" captionStartId="5.[151,250,1775,1799]" captionTargetBox="[151,1435,309,1752]" captionTargetId="figure-91@5.[151,1435,309,1752]" captionTargetPageId="5" captionText="FIGURE 2. Overall dorsal view. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; D, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; E, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; F, Lophoplax vermiculata, subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa; H, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663772" httpUri="https://zenodo.org/record/7663772/files/figure.png" pageId="8" pageNumber="436">Fig. 2B</figureCitation>
|
||
,
|
||
<figureCitation box="[939,972,475,501]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="8" pageNumber="436">3C</figureCitation>
|
||
vs.
|
||
<figureCitation box="[1016,1103,475,501]" captionStart="FIGURE 2" captionStartId="5.[151,250,1775,1799]" captionTargetBox="[151,1435,309,1752]" captionTargetId="figure-91@5.[151,1435,309,1752]" captionTargetPageId="5" captionText="FIGURE 2. Overall dorsal view. A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; C, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; D, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; E, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia; F, Lophoplax vermiculata, subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), female (13.7 × 10.5 mm) (ZRC 2022.775), Okinawa; H, L. pannosa n. sp., holotype ovigerous female (7.4 × 5.7 mm) (ZRC 2019.1190), Vanuatu." figureDoi="http://doi.org/10.5281/zenodo.7663772" httpUri="https://zenodo.org/record/7663772/files/figure.png" pageId="8" pageNumber="436">Fig. 2A</figureCitation>
|
||
,
|
||
<figureCitation box="[1114,1152,475,501]" captionStart="FIGURE 3" captionStartId="6.[151,250,1744,1768]" captionTargetBox="[153,1433,181,1720]" captionTargetId="figure-17@6.[153,1433,181,1720]" captionTargetPageId="6" captionText="FIGURE 3. Dorsal view of carapace.A, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; B, L. sculpta (Stimpson, 1858), male (7.8 × 6.2 mm) (ZRC 2022.777), Okinawa [photograph: T. Maenosono]; C, L. sculpta (Stimpson, 1858), female (10.4 × 8.5 mm) (ZRC 2019.1611), Philippines; D, L. sordida n. sp., paratype male (6.1 × 4.8 mm) (ZRC 2022.91), Lombok; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. sordida n. sp., paratype male (7.0 × 6.1 mm) (ZRC 2022.718), Lombok." figureDoi="http://doi.org/10.5281/zenodo.7663776" httpUri="https://zenodo.org/record/7663776/files/figure.png" pageId="8" pageNumber="436">3A</figureCitation>
|
||
). The two tubercles at the junction between the lateral margins are also slightly larger and not arranged in a linear manner like the Japanese specimens (
|
||
<figureCitation box="[286,378,547,573]" captionStart="FIGURE 9" captionStartId="14.[151,250,1509,1533]" captionTargetBox="[176,1410,181,1485]" captionTargetId="figure-17@14.[176,1410,181,1485]" captionTargetPageId="14" captionText="FIGURE 9. A–D, Lophoplax sculpta (Stimpson, 1858), neotype male (7.8 × 6.3 mm) (RUMF-ZC-8127), Okinawa; E, L. sordida n. sp., holotype male (8.2 × 6.5 mm) (MZB Cru 5449), Lombok; F, L. vermiculata (A. Milne-Edwards, 1873), subadult male (5.6 × 4.5 mm) (ZRC 2013.1047), Guam; G, L. vermiculata (A. Milne-Edwards, 1873), lectotype male (10.6 × 8.2 mm) (ZRC 2022.696), New Caledonia. A, right lateral view of posterior part of sub-branchial region; B, E–G, anterior male thoracic sternum and pleon; C, male pleonal somites 1–4; D, male pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.7663797" httpUri="https://zenodo.org/record/7663797/files/figure.png" pageId="8" pageNumber="436">Fig. 9A</figureCitation>
|
||
), appearing as if there are three tubercles rather than two. The differences, however, do not appear significant at the species level and can be explained by intraspecific variation.
|
||
</paragraph>
|
||
<paragraph blockId="8.[151,1437,151,825]" box="[199,1230,619,645]" pageId="8" pageNumber="436">
|
||
For comparisons with
|
||
<taxonomicName authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[450,565,619,645]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="436" phylum="Arthropoda" rank="species" species="sordida" status="n. sp.">
|
||
<emphasis box="[450,565,619,645]" italics="true" pageId="8" pageNumber="436">L. sordida</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[571,634,619,645]" pageId="8" pageNumber="436">
|
||
<taxonomicNameLabel box="[571,634,619,645]" pageId="8" pageNumber="436" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName authority="Ng & Rahayu, 2023" authorityName="Ng & Rahayu" authorityYear="2023" box="[689,814,619,644]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="436" phylum="Arthropoda" rank="species" species="pannosa" status="n. sp.">
|
||
<emphasis box="[689,814,619,644]" italics="true" pageId="8" pageNumber="436">L. pannosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[820,883,619,645]" pageId="8" pageNumber="436">
|
||
<taxonomicNameLabel box="[820,883,619,645]" pageId="8" pageNumber="436" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, see remarks for these species.
|
||
</paragraph>
|
||
<paragraph blockId="8.[151,1437,151,825]" pageId="8" pageNumber="436">
|
||
<emphasis bold="true" box="[199,291,654,680]" pageId="8" pageNumber="436">Habitat</emphasis>
|
||
. From the available data, the species occurs in shallow waters outside reefs. In intertidal areas, it occurs under rocks, often with seagrass nearby. One specimen (ZRC 2019.1611) was collected from a dense seagrass patch with a sandy substrate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="436" type="distribution">
|
||
<paragraph blockId="8.[151,1437,151,825]" pageId="8" pageNumber="436">
|
||
<emphasis bold="true" box="[199,345,762,788]" pageId="8" pageNumber="436">Distribution</emphasis>
|
||
. Known from the Ryukyus,
|
||
<collectingCountry box="[677,744,763,789]" name="Japan" pageId="8" pageNumber="436">Japan</collectingCountry>
|
||
; and
|
||
<collectingCountry box="[809,935,763,789]" name="Philippines" pageId="8" pageNumber="436">Philippines</collectingCountry>
|
||
(
|
||
<bibRefCitation author="Stimpson, W." box="[952,1128,762,789]" pageId="8" pageNumber="436" pagination="93 - 110" refId="ref18446" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 93 - 110 (39 - 56). https: // doi. org / 10.5962 / bhl. title. 51447" type="journal article" year="1858">Stimpson 1858</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Takeda, M." box="[1142,1291,763,789]" pageId="8" pageNumber="436" pagination="120 - 123" refId="ref18577" refString="Takeda, M. (1977) Rediscovery of Lophoplax sculpta (Stimpson, 1858) (Crustacea, Decapoda, Brachyura). Zoological Magazine, 86, 120 - 123." type="journal article" year="1977">Takeda 1977</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Maenosono, T." pageId="8" pageNumber="436" pagination="19 - 44" refId="ref16091" refString="Maenosono, T. (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48, 19 - 44. [in Japanese with English abstract]" type="journal article" year="2019">Maenosono 2019</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Ng, P. K. L. & D. Guinot & P. J. F. Davie" box="[222,387,799,825]" pageId="8" pageNumber="436" pagination="1 - 286" refId="ref17397" refString="Ng, P. K. L., D. Guinot & P. J. F. Davie, 2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
|
||
Ng
|
||
<emphasis box="[263,319,799,825]" italics="true" pageId="8" pageNumber="436">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |