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41 KiB
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510 lines
41 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.151.50538" ID-GBIF-Dataset="a62873a2-c4e1-4c6d-9baa-5c5fb94e7348" ID-PMC="PMC7329918" ID-Pensoft-Pub="1314-2003-151-107" ID-Pensoft-UUID="42D1EB2DF0545744B92BFCCCB395C45B" ID-PubMed="32655282" ModsDocID="1314-2003-151-107" checkinTime="1593036331120" checkinUser="pensoft" docAuthor="Sylvester, Steven P., Cuta-Alarcon, Lia E., Bravo-Pedraza, William J. & Soreng, Robert J." docDate="2020" docId="9FED202A658955E48A38CF45E18212D9" docLanguage="en" docName="PhytoKeys 151: 107-160" docOrigin="PhytoKeys 151" docSource="http://dx.doi.org/10.3897/phytokeys.151.50538" docTitle="Agrostis tolucensis Kunth, Nov. Gen. Sp. [H. B. K.] 1: 135. 1816" docType="treatment" docVersion="4" id="42D1EB2DF0545744B92BFCCCB395C45B" lastPageNumber="107" masterDocId="42D1EB2DF0545744B92BFCCCB395C45B" masterDocTitle="Agrostis and Podagrostis (Agrostidinae, Poaceae) from paramos of Boyaca, Colombia: synoptic taxonomy including a key to Colombian species" masterLastPageNumber="160" masterPageNumber="107" pageNumber="107" updateTime="1668139837814" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Agrostis and Podagrostis (Agrostidinae, Poaceae) from paramos of Boyaca, Colombia: synoptic taxonomy including a key to Colombian species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sylvester, Steven P.</mods:namePart>
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<mods:affiliation>College of Biology and the Environment, Nanjing Forestry University, Long Pan Road No. 159, Nanjing, 210037, China & Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 AE, UK</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5577-8782</mods:nameIdentifier>
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<mods:nameIdentifier type="email">steven_sylvester@hotmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cuta-Alarcon, Lia E.</mods:namePart>
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<mods:affiliation>Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bravo-Pedraza, William J.</mods:namePart>
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<mods:affiliation>Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Soreng, Robert J.</mods:namePart>
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<mods:affiliation>Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington DC 20560, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-8358-4915</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>151</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>107</mods:start>
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<mods:end>160</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.151.50538</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.151.50538</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-151-107</mods:identifier>
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<mods:identifier type="Pensoft-UUID">42D1EB2DF0545744B92BFCCCB395C45B</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="164730850" LSID="urn:lsid:plazi:treatment:9FED202A658955E48A38CF45E18212D9" httpUri="http://treatment.plazi.org/id/9FED202A658955E48A38CF45E18212D9" lastPageNumber="107" pageId="0" pageNumber="107">
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<subSubSection pageId="0" pageNumber="107" type="nomenclature">
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<paragraph pageId="0" pageNumber="107">
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<taxonomicName LSID="9FED202A-6589-55E4-8A38-CF45E18212D9" authority="Kunth, Nov. Gen. Sp. [H. B. K.] 1: 135. 1816" authorityName="Kunth, Nov. Gen. Sp. [H. B. K.] 1: 135." authorityYear="1816" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis tolucensis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tolucensis">Agrostis tolucensis Kunth, Nov. Gen. Sp. [H.B.K.] 1: 135. 1816</taxonomicName>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Agrostis tolucensis A spikelet, glumes in lateral view, floret in dorsal view, detached and raised above the glumes B whole plant. Images of Cuta-Alarcon 376 (FMB)." figureDoi="10.3897/phytokeys.151.50538.figure10" httpUri="https://binary.pensoft.net/fig/424848" pageId="0" pageNumber="107">Fig. 10</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="107" type="reference_group">
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<paragraph pageId="0" pageNumber="107">
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis tolucensis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tolucensis">Agrostis tolucensis</taxonomicName>
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Willd. ex Steud., Syn. Pl. Glumac. 1: 164. 1855[1854], nom. inval.
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</paragraph>
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<paragraph pageId="0" pageNumber="107">
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=
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<taxonomicName authorityName="Kunth" authorityYear="1833" baseAuthorityName="J.Presl" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis glomerata" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="glomerata">Agrostis glomerata</taxonomicName>
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(J. Presl) Kunth, Enum. Pl. [Kunth] 1: 219. 1833.
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<taxonomicName authorityName="J.Presl" authorityYear="1830" class="Liliopsida" family="Poaceae" genus="Vilfa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Vilfa glomerata" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="glomerata">Vilfa glomerata</taxonomicName>
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J. Presl, Reliq. Haenk. 1 (4-5): 239. 1830. Type: Peru. [
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<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
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] [Hab. in montanis Peruviae huanoccensibus], [1791; 1891 written on W isotype], T.P.X. Haenke s.n. [#196 written on W isotype] (holotype: [not found]; isotypes: HAL (HAL0106916 [image!]), PRC (PRC450953 [image!]), US (US00589472 fragm.), W (W0025326 [image!])).
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</paragraph>
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<paragraph pageId="0" pageNumber="107">
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=
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis hoffmannii" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="hoffmannii">Agrostis hoffmannii</taxonomicName>
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Mez, Repert. Spec. Nov. Regni Veg. 18: 3. 1922. Type: Costa Rica. Irazu (holotype: [not found]; isotypes: [not found]).
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</paragraph>
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<paragraph pageId="0" pageNumber="107">
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=
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis nana subsp. var. var. andicola" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="variety" species="nana" subSpecies="var." variety="andicola">Agrostis nana var. andicola</taxonomicName>
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Pilg., Bot. Jahrb. Syst. 37 (5): 505. 1906. Type: Ecuador. Chimborazo: Chimborazo, N-Seite,
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<normalizedToken originalValue="Páramo-Region">Paramo-Region</normalizedToken>
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, 4500 m alt., July 1903, H. Meyer 146 (syntypes: JE (JE00020226 [image!]; JE00020228 [image!])); Napo: Antisana, in frigidis alpinis, 4600 m alt., July 1903, H. Meyer 145 (syntypes: JE (JE00020225 [image!]; JE00020227 [image!])).
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</paragraph>
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<paragraph pageId="0" pageNumber="107">
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=
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis nana subsp. var. var. aristata" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="variety" species="nana" subSpecies="var." variety="aristata">Agrostis nana var. aristata</taxonomicName>
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Griseb., Abh.
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<normalizedToken originalValue="Königl">Koenigl</normalizedToken>
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. Ges. Wiss.
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<normalizedToken originalValue="Göttingen">Goettingen</normalizedToken>
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24: 294. 1879. Type: Argentina. Salta: Umgebungen des Nevado del Castillo [around the Nevado del Castillo], 10-15000 ft. [3048-4572 m alt.], 19-23 Mar. 1873, P.G. Lorentz & G.H.E.W. Hieronymus 82 (holotype: GOET (GOET006541 [image!]); isotypes: B, BAA (BAA00000724 [image!] fragm. ex B; BAA00000723 [image!] fragm. ex B), CORD (CORD00004693 [image!]), W (W19160036588 [image!]; W19160036664 [image!])).
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</paragraph>
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<paragraph pageId="0" pageNumber="107">
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=
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis virescens" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="virescens">Agrostis virescens</taxonomicName>
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Kunth, Nov. Gen. Sp. [H.B.K.] 1: 135. 1816. Type: Mexico. Toluca, Sep., F.W.H.A. Humboldt & A.J.A. Bonpland s.n (holotype: P (P00669395 [image!]); isotypes: LE-TRIN (LE-TRIN1668.01; LE-TRIN1668.02), P (P00136912 [image!])).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="107" type="materials_examined">
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<paragraph pageId="0" pageNumber="107">Type.</paragraph>
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<paragraph pageId="0" pageNumber="107">Mexico. [prope Toluca et Islahuaca], F.W.H.A. Humboldt & A.J.A. Bonpland s.n. (holotype: P (P00669394 [image!]); isotypes: B (B-W1704), BAA (BAA00000235 [image!] fragm. ex P), BM, K (microfiche), LE-TRIN (LE-TRIN1660.01), P (P00136914 [image!]; P00136915 [image!]; P00740426 [image!]), US (US00156505! fragm. ex P)).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="107" type="description">
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<paragraph pageId="0" pageNumber="107">Description.</paragraph>
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<paragraph pageId="0" pageNumber="107">
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<emphasis bold="true" pageId="0" pageNumber="107">Perennial herbs</emphasis>
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, tussock-forming or laxly to densely tufted, usually with short ascending rhizomes.
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<emphasis bold="true" pageId="0" pageNumber="107">Tillers</emphasis>
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extravaginal and intravaginal, with cataphylls usually present.
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<emphasis bold="true" pageId="0" pageNumber="107">Culms</emphasis>
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(3-)5.5-51(-80) cm tall, erect or arching, fairly firm, with 0(-2) nodes exerted at flowering, smooth or rarely scaberulous.
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<emphasis bold="true" pageId="0" pageNumber="107">Leaves</emphasis>
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mostly basal or more-or-less evenly spread along the culm, glabrous, smooth or scabrous;
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<emphasis bold="true" pageId="0" pageNumber="107">ligules</emphasis>
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2-4(-6.2) mm long, of upper culm usually longer than those of lower culm and tillers, truncate to triangular, moderately to strongly decurrent with the sheath, abaxial surface usually scabrous, rarely scaberulous or smooth;
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<emphasis bold="true" pageId="0" pageNumber="107">blades</emphasis>
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2.5-19 cm long, 1-3(-5) mm wide, at least in the upper culm filiform, folded or flat, usually lax to sometimes firm, sometimes basal and tiller blades involute or convolute and firm to rigid, smooth throughout or scabrous only on the margin and sometimes veins, apices blunt or slightly naviculate-acute.
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<emphasis bold="true" pageId="0" pageNumber="107">Panicles</emphasis>
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(1-)2-15 cm long, 0.1-1.5 cm wide, moderately to densely congested, sub-spikelike or spikelike, sometimes interrupted towards the base, subincluded in the basal foliage or slightly to greatly exerted, lateral branches with spikelets almost to the base, upper lateral branches short and held close to the central inflorescence axis, central axis and panicle branches scabrous;
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<emphasis bold="true" pageId="0" pageNumber="107">pedicels</emphasis>
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0.7-3(-4.5) mm long, usually shorter than their spikelets, not or slightly dilated at their apex, scabrous.
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<emphasis bold="true" pageId="0" pageNumber="107">Spikelets</emphasis>
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(not including awn, if present) 2-3(-3.6) mm long;
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<emphasis bold="true" pageId="0" pageNumber="107">glumes</emphasis>
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equal or subequal, 1-veined, lower glume keel and surface usually scabrous at least in the distal half, infrequently smooth throughout, upper glume keel and surface scabrous for almost their entirety to at least in the distal half or surface sometimes smooth throughout, apices acute or acuminate;
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<emphasis bold="true" pageId="0" pageNumber="107">floret</emphasis>
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<normalizedToken originalValue="½">1/2</normalizedToken>
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-2/3(-3/4) the length of the glumes;
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<emphasis bold="true" pageId="0" pageNumber="107">calluses</emphasis>
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lightly pilose with 2 sparse tufts of short hairs on the lateral sides;
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<emphasis bold="true" pageId="0" pageNumber="107">lemmas</emphasis>
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1.4-2.8 mm long, glabrous, smooth, 5-veined, apex truncate, denticulate, usually awned, awn 2-3.5 mm long, inserted in the lower, middle, or upper 1/3 of the dorsal keel, exerted from the glumes, geniculate, twisted, very rarely muticous or with a short straight awn, <1 mm long, inserted in the upper half of the lemma (see notes below regarding
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<taxonomicName lsidName="A. glomerata" pageId="0" pageNumber="107" rank="species" species="glomerata">
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<emphasis italics="true" pageId="0" pageNumber="107">A. glomerata</emphasis>
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</taxonomicName>
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);
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<emphasis bold="true" pageId="0" pageNumber="107">paleas</emphasis>
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absent or 0.1-0.2 mm long, <
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<normalizedToken originalValue="¼">1/4</normalizedToken>
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the length of the lemma;
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<emphasis bold="true" pageId="0" pageNumber="107">rachilla</emphasis>
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absent;
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<emphasis bold="true" pageId="0" pageNumber="107">anthers</emphasis>
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0.5-1 mm long.
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</paragraph>
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<caption doi="10.3897/phytokeys.151.50538.figure10" httpUri="https://binary.pensoft.net/fig/424848" pageId="0" pageNumber="107" start="Figure 10" startId="F10">
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<paragraph pageId="0" pageNumber="107">
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<emphasis bold="true" pageId="0" pageNumber="107">Figure 10.</emphasis>
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis tolucensis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tolucensis">
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<emphasis italics="true" pageId="0" pageNumber="107">Agrostis tolucensis</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="107">A</emphasis>
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spikelet, glumes in lateral view, floret in dorsal view, detached and raised above the glumes
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<emphasis bold="true" pageId="0" pageNumber="107">B</emphasis>
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whole plant. Images of Cuta-Alarcon 376 (FMB).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="107" type="distribution">
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<paragraph pageId="0" pageNumber="107">Distribution and ecology.</paragraph>
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<paragraph pageId="0" pageNumber="107">Mexico, Guatemala, Costa Rica, Panama, Colombia, Venezuela, Ecuador, Peru, Bolivia, Chile, Argentina. High-elevation open grasslands and forests, 2700-4900 m alt.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="107" type="materials_examined">
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<paragraph pageId="0" pageNumber="107">Other specimens examined.</paragraph>
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<paragraph pageId="0" pageNumber="107">
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<emphasis bold="true" pageId="0" pageNumber="107">Colombia</emphasis>
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. Sin loc., 1760-1808, J.C. Mutis 5521 (US1561906).
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<emphasis bold="true" pageId="0" pageNumber="107">
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<normalizedToken originalValue="Boyacá">Boyaca</normalizedToken>
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</emphasis>
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: Munic. Arcabuco,
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<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
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El Valle,
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<geoCoordinate degrees="5.7445833" direction="north" orientation="latitude" precision="1" value="5.744583">5.7445833N</geoCoordinate>
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,
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<geoCoordinate degrees="73.37118" direction="west" orientation="longitude" precision="1" value="-73.37118">73.37118W</geoCoordinate>
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, 3744 m alt., 16 Nov. 2017, S.P. Sylvester 3081 (FMB, K, US); Munic. Arcabuco,
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<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
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El Valle, just below high point of ridge to the W of site 14,
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<geoCoordinate degrees="5.75436" direction="north" orientation="latitude" precision="1" value="5.75436">5.75436N</geoCoordinate>
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,
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<geoCoordinate degrees="73.38722" direction="west" orientation="longitude" precision="1" value="-73.38722">73.38722W</geoCoordinate>
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, 3747 m alt.,
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<normalizedToken originalValue="páramo">paramo</normalizedToken>
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seco dominado por
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<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Espeletia barclayana" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="barclayana">
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<emphasis italics="true" pageId="0" pageNumber="107">Espeletia barclayana</emphasis>
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</taxonomicName>
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[Cuatrec.], 16 Nov. 2017, S.P. Sylvester 3083 (K, US); Munic.
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<normalizedToken originalValue="Belén">Belen</normalizedToken>
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,
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<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
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de La Rusia, border with Santander, Boqueron El Consuelo, unprotected private land,
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<geoCoordinate degrees="6.03598" direction="north" orientation="latitude" precision="1" value="6.03598">6.03598N</geoCoordinate>
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,
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<geoCoordinate degrees="72.57056" direction="west" orientation="longitude" precision="1" value="-72.57056">72.57056W</geoCoordinate>
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, 3975 m alt.,
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<normalizedToken originalValue="páramo">paramo</normalizedToken>
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with limited grazing by horses and rodents with
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<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Espeletia brachyaxiantha" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="brachyaxiantha">
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<emphasis italics="true" pageId="0" pageNumber="107">Espeletia brachyaxiantha</emphasis>
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</taxonomicName>
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[S.
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<normalizedToken originalValue="Díaz">Diaz</normalizedToken>
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] and
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<taxonomicName lsidName="E. annemariana" pageId="0" pageNumber="107" rank="species" species="annemariana">
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<emphasis italics="true" pageId="0" pageNumber="107">E. annemariana</emphasis>
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</taxonomicName>
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[Cuatrec.], steep slope on rock outcrop, 23 Nov. 2017, M. Vorontsova 2260 (FMB, K, SI, US); Munic.
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<normalizedToken originalValue="Belén">Belen</normalizedToken>
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,
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<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
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de La Rusia, near
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<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
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Le Consuelo, unprotected private land, somewhat disturbed
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<normalizedToken originalValue="páramo">paramo</normalizedToken>
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on top of the ridge,
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<geoCoordinate degrees="6.02416" direction="north" orientation="latitude" precision="1" value="6.02416">6.02416N</geoCoordinate>
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,
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<geoCoordinate degrees="72.57242" direction="west" orientation="longitude" precision="1" value="-72.57242">72.57242W</geoCoordinate>
|
|
, 3894 m alt., grazed by horses and rodents, with
|
|
<taxonomicName authorityName="Cuatrec" authorityYear="1973" class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Espeletia boyacensis" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="boyacensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">Espeletia boyacensis</emphasis>
|
|
</taxonomicName>
|
|
, 22 Nov. 2017, M. Vorontsova 2231 (K, UPTC, US); Munic. Chiscas,
|
|
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
|
de Chacaritas, asociado a rocas de 4 m de altura,
|
|
<geoCoordinate degrees="6.62227" direction="north" orientation="latitude" precision="1" value="6.62227">6.62227N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="72.3904" direction="west" orientation="longitude" precision="5" value="-72.3904">72.3904W</geoCoordinate>
|
|
, 4192 m alt., 4 Mar. 2018, S.P. Sylvester 3115 (US); Munic. Chiscas,
|
|
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
|
de Chacaritas,
|
|
<normalizedToken originalValue="límites">limites</normalizedToken>
|
|
entre
|
|
<normalizedToken originalValue="páramo">paramo</normalizedToken>
|
|
y
|
|
<normalizedToken originalValue="superpáramo">superparamo</normalizedToken>
|
|
,
|
|
<geoCoordinate degrees="6.62865" direction="north" orientation="latitude" precision="1" value="6.62865">6.62865N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="72.3944" direction="west" orientation="longitude" precision="5" value="-72.3944">72.3944W</geoCoordinate>
|
|
, 4064 m alt., 4 Mar. 2018, S.P. Sylvester 3104 (FMB, K, US); Munic. Chiscas,
|
|
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
|
El
|
|
<normalizedToken originalValue="Peñon">Penon</normalizedToken>
|
|
, Chiscas,
|
|
<geoCoordinate degrees="6.63012" direction="north" orientation="latitude" precision="1" value="6.63012">6.63012N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="72.40073" direction="west" orientation="longitude" precision="1" value="-72.40073">72.40073W</geoCoordinate>
|
|
, 4172 m alt., 5 Mar. 2018, S.P. Sylvester 3152 (UPTC); Munic. Chiscas,
|
|
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
|
El
|
|
<normalizedToken originalValue="Peñon">Penon</normalizedToken>
|
|
, Chiscas,
|
|
<geoCoordinate degrees="6.62876" direction="north" orientation="latitude" precision="1" value="6.62876">6.62876N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="72.40283" direction="west" orientation="longitude" precision="1" value="-72.40283">72.40283W</geoCoordinate>
|
|
, 4255 m alt.,
|
|
<normalizedToken originalValue="vegetación">vegetacion</normalizedToken>
|
|
de pajonal frailejonal,
|
|
<normalizedToken originalValue="páramo">paramo</normalizedToken>
|
|
<normalizedToken originalValue="húmedo">humedo</normalizedToken>
|
|
, 5 Mar. 2018, S.P. Sylvester 3153 (FMB, K, US); Munic. Mongua,
|
|
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
|
de
|
|
<normalizedToken originalValue="Ocetá">Oceta</normalizedToken>
|
|
, Valle de Laguna Negra,
|
|
<geoCoordinate degrees="5.69525" direction="north" orientation="latitude" precision="1" value="5.69525">5.69525N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="72.79133" direction="west" orientation="longitude" precision="1" value="-72.79133">72.79133W</geoCoordinate>
|
|
, 3694 m alt., 29 Nov. 2017, L.E.
|
|
<normalizedToken originalValue="Cuta-Alarcón">Cuta-Alarcon</normalizedToken>
|
|
355 (FMB, K, UPTC, US); Munic. Mongui, Vereda Vallado, Sector La Pedrisca,
|
|
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
|
de
|
|
<normalizedToken originalValue="Ocetá">Oceta</normalizedToken>
|
|
,
|
|
<geoCoordinate degrees="5.69969" direction="north" orientation="latitude" precision="1" value="5.69969">5.69969N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="72.80891" direction="west" orientation="longitude" precision="1" value="-72.80891">72.80891W</geoCoordinate>
|
|
, 3751 m alt., 30 Nov. 2017, L.E.
|
|
<normalizedToken originalValue="Cuta-Alarcón">Cuta-Alarcon</normalizedToken>
|
|
376 (FMB, US).
|
|
<emphasis bold="true" pageId="0" pageNumber="107">Cauca</emphasis>
|
|
: Macizo Colombiano, Valle de las Papas, alrededores de Valencia, [
|
|
<geoCoordinate degrees="1.8831" direction="north" orientation="latitude" precision="5" value="1.8831">1.8831N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="76.6828" direction="west" orientation="longitude" precision="5" value="-76.6828">76.6828W</geoCoordinate>
|
|
], 2910 m alt., 11 Sep.-1 Oct. 1958, Idrobo, Pinto & Bischler 3682A (US2540528).
|
|
<emphasis bold="true" pageId="0" pageNumber="107">Cundinamarca</emphasis>
|
|
:
|
|
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
|
de Chisaca, [
|
|
<geoCoordinate degrees="4.2747" direction="north" orientation="latitude" precision="5" value="4.2747">4.2747N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="74.2006" direction="west" orientation="longitude" precision="5" value="-74.2006">74.2006W</geoCoordinate>
|
|
], 3750-3962 m alt., 5 Oct. 1966, T.R. Soderstrom 1312 (US3136708).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="107" type="notes">
|
|
<paragraph pageId="0" pageNumber="107">Notes.</paragraph>
|
|
<paragraph pageId="0" pageNumber="107">
|
|
Although in Bolivia (
|
|
<bibRefCitation author="Renvoize, SA" journalOrPublisher="Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Mexico" pageId="0" pageNumber="107" refId="B29" refString="Renvoize, SA, 1998. Gramineas de Bolivia. Continental Printing, Belgium." title="Gramineas de Bolivia. Continental Printing, Belgium." year="1998">Renvoize 1998</bibRefCitation>
|
|
) and austral South America (
|
|
<bibRefCitation author="Rugolo de Agrasar, ZE" editor="Zuloaga, FO" journalOrPublisher="Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina" pageId="0" pageNumber="107" pagination="102 - 125" refId="B30" refString="Rugolo de Agrasar, ZE, 2012. Agrostis L. In: Zuloaga, FO, Rugolo de Agrasar, ZE, Anton, AM, Eds., Flora Argentina. Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina: 102 - 125" title="Agrostis L." volumeTitle="Flora Argentina." year="2012">
|
|
<normalizedToken originalValue="Rúgolo">Rugolo</normalizedToken>
|
|
de Agrasar 2012
|
|
</bibRefCitation>
|
|
)
|
|
<taxonomicName lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
|
|
</taxonomicName>
|
|
is mentioned to be notably rhizomatous, lateral tending rhizomes were not notable on specimens from the
|
|
<normalizedToken originalValue="páramos">paramos</normalizedToken>
|
|
of
|
|
<normalizedToken originalValue="Boyacá">Boyaca</normalizedToken>
|
|
, which had short, ascending rhizomes/pseudostolons and formed small tussocks or were laxly to densely tufted. Type specimens and original material of
|
|
<taxonomicName lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
|
|
</taxonomicName>
|
|
and other species now considered synonyms of
|
|
<taxonomicName lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
|
|
</taxonomicName>
|
|
, i.e.
|
|
<taxonomicName authorityName="Kunth" authorityYear="1833" baseAuthorityName="J.Presl" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis glomerata" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="glomerata">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">Agrostis glomerata</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="A. nana subsp. var. var. andicola" pageId="0" pageNumber="107" rank="variety" species="nana" subSpecies="var." variety="andicola">A. nana var. andicola</taxonomicName>
|
|
,
|
|
<taxonomicName lsidName="A. nana subsp. var. var. aristata" pageId="0" pageNumber="107" rank="variety" species="nana" subSpecies="var." variety="aristata">A. nana var. aristata</taxonomicName>
|
|
, and
|
|
<taxonomicName lsidName="A. virescens" pageId="0" pageNumber="107" rank="species" species="virescens">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. virescens</emphasis>
|
|
</taxonomicName>
|
|
, also had either short ascending rhizomes or lacked notable rhizomes, but with all having obvious extravaginal cataphyllous shoots.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="107">
|
|
Discrepancy was sometimes also found in the form of the leaf blades, with specimens encountered in
|
|
<normalizedToken originalValue="Boyacá">Boyaca</normalizedToken>
|
|
, Colombia, and Venezuela sometimes having more rigid, convolute to folded, often recurved, basal leaf blades instead of the lax and filiform leaf blades more common to this species. These specimens did usually still have flat upper culm blades to help differentiate them from e.g.
|
|
<taxonomicName lsidName="A. laegaardii" pageId="0" pageNumber="107" rank="species" species="laegaardii">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. laegaardii</emphasis>
|
|
</taxonomicName>
|
|
(see notes on similar species below). While
|
|
<bibRefCitation author="Renvoize, SA" journalOrPublisher="Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Mexico" pageId="0" pageNumber="107" refId="B29" refString="Renvoize, SA, 1998. Gramineas de Bolivia. Continental Printing, Belgium." title="Gramineas de Bolivia. Continental Printing, Belgium." year="1998">Renvoize (1998)</bibRefCitation>
|
|
and
|
|
<bibRefCitation author="Rugolo de Agrasar, ZE" editor="Zuloaga, FO" journalOrPublisher="Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina" pageId="0" pageNumber="107" pagination="102 - 125" refId="B30" refString="Rugolo de Agrasar, ZE, 2012. Agrostis L. In: Zuloaga, FO, Rugolo de Agrasar, ZE, Anton, AM, Eds., Flora Argentina. Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina: 102 - 125" title="Agrostis L." volumeTitle="Flora Argentina." year="2012">
|
|
<normalizedToken originalValue="Rúgolo">Rugolo</normalizedToken>
|
|
de Agrasar (2012)
|
|
</bibRefCitation>
|
|
state the blades in
|
|
<taxonomicName lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
|
|
</taxonomicName>
|
|
to be flat, folded, or filiform and generally lax, the type material also has fairly firm blades which are narrow and folded to slightly convolute and sometimes cylindrical in outline.
|
|
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis nana" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="nana">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">Agrostis nana</emphasis>
|
|
</taxonomicName>
|
|
, here considered a synonym of
|
|
<taxonomicName lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
|
|
</taxonomicName>
|
|
, also has more convolute, curved and rigid leaf blades.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="107" type="similar species">
|
|
<paragraph pageId="0" pageNumber="107">Similar species.</paragraph>
|
|
<paragraph pageId="0" pageNumber="107">
|
|
See notes under
|
|
<taxonomicName lsidName="A. foliata" pageId="0" pageNumber="107" rank="species" species="foliata">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. foliata</emphasis>
|
|
</taxonomicName>
|
|
for how to differentiate from that species.
|
|
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis breviculmis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="breviculmis">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">Agrostis breviculmis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName lsidName="A. laegaardii" pageId="0" pageNumber="107" rank="species" species="laegaardii">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. laegaardii</emphasis>
|
|
</taxonomicName>
|
|
bear similarities in having mainly basal, narrow, usually convolute, rigid, leaf blades and densely congested spikelike panicles that are usually short (<5 cm long), and spikelets with an absent or reduced palea <
|
|
<normalizedToken originalValue="¼">1/4</normalizedToken>
|
|
the length of the lemma.
|
|
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis tolucensis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tolucensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">Agrostis tolucensis</emphasis>
|
|
</taxonomicName>
|
|
principally differs from these by the leaf blades being usually laxer and folded or flat, at least in the upper culm. However, as noted above, specimens can be found with convolute basal blades that makes distinguishing these species more difficult. In these instances,
|
|
<taxonomicName lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
|
|
</taxonomicName>
|
|
can be differentiated from
|
|
<taxonomicName lsidName="A. breviculmis" pageId="0" pageNumber="107" rank="species" species="breviculmis">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. breviculmis</emphasis>
|
|
</taxonomicName>
|
|
by its larger spikelets 2-3(-3.6) mm long (vs. 1.5-2.1(-2.5 in Bolivia?;
|
|
<bibRefCitation author="Renvoize, SA" journalOrPublisher="Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Mexico" pageId="0" pageNumber="107" refId="B29" refString="Renvoize, SA, 1998. Gramineas de Bolivia. Continental Printing, Belgium." title="Gramineas de Bolivia. Continental Printing, Belgium." year="1998">Renvoize 1998</bibRefCitation>
|
|
) mm long in
|
|
<taxonomicName lsidName="A. breviculmis" pageId="0" pageNumber="107" rank="species" species="breviculmis">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. breviculmis</emphasis>
|
|
</taxonomicName>
|
|
), usually the presence of a well-developed geniculate and twisted awn inserted dorsally usually in the lower third of the lemma (lemmas very rarely muticous or with a short straight weak awn, <1 mm long, inserted in the upper
|
|
<normalizedToken originalValue="½">1/2</normalizedToken>
|
|
; see notes on
|
|
<taxonomicName lsidName="A. glomerata" pageId="0" pageNumber="107" rank="species" species="glomerata">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. glomerata</emphasis>
|
|
</taxonomicName>
|
|
below) (vs. lemmas muticous or with a short straight weak awn inserted above the middle in
|
|
<taxonomicName lsidName="A. breviculmis" pageId="0" pageNumber="107" rank="species" species="breviculmis">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. breviculmis</emphasis>
|
|
</taxonomicName>
|
|
), panicles generally wider, 1-15 mm wide (vs. 0.5-2(-6) mm wide in
|
|
<taxonomicName lsidName="A. breviculmis" pageId="0" pageNumber="107" rank="species" species="breviculmis">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. breviculmis</emphasis>
|
|
</taxonomicName>
|
|
), and short prickle hairs on the glume keels (vs. coarse and shiny in
|
|
<taxonomicName lsidName="A. breviculmis" pageId="0" pageNumber="107" rank="species" species="breviculmis">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. breviculmis</emphasis>
|
|
</taxonomicName>
|
|
). While
|
|
<taxonomicName lsidName="A. laegaardii" pageId="0" pageNumber="107" rank="species" species="laegaardii">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. laegaardii</emphasis>
|
|
</taxonomicName>
|
|
has characters of often larger spikelets and dorsally inserted awn,
|
|
<taxonomicName lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
|
|
</taxonomicName>
|
|
can be differentiated from
|
|
<taxonomicName lsidName="A. laegaardii" pageId="0" pageNumber="107" rank="species" species="laegaardii">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. laegaardii</emphasis>
|
|
</taxonomicName>
|
|
by its extravaginal cataphyllous shoots and distinct ascending rhizomes with rhizome internodes usually> 7 mm long (vs. purely intravaginal innovations, without cataphyllous shoots or distinct ascending rhizomes, rhizome internodes <2 mm long in
|
|
<taxonomicName lsidName="A. laegaardii" pageId="0" pageNumber="107" rank="species" species="laegaardii">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. laegaardii</emphasis>
|
|
</taxonomicName>
|
|
), generally wider panicles, 1-15 mm wide (vs. 0.5-2(-6) mm wide in
|
|
<taxonomicName lsidName="A. laegaardii" pageId="0" pageNumber="107" rank="species" species="laegaardii">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. laegaardii</emphasis>
|
|
</taxonomicName>
|
|
), and short prickle hairs on the glume keels (vs. coarse and shiny in
|
|
<taxonomicName lsidName="A. laegaardii" pageId="0" pageNumber="107" rank="species" species="laegaardii">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. laegaardii</emphasis>
|
|
</taxonomicName>
|
|
). Specimens can sometimes have laxer inflorescences that could lead to confusion with
|
|
<taxonomicName lsidName="A. mertensii" pageId="0" pageNumber="107" rank="species" species="mertensii">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. mertensii</emphasis>
|
|
</taxonomicName>
|
|
, but this species does not form small tussocks.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="107">
|
|
<taxonomicName authorityName="Trin" authorityYear="1841" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis meyenii" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="meyenii">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">Agrostis meyenii</emphasis>
|
|
</taxonomicName>
|
|
, a species known from drier high-elevation puna grassland and pampa of Argentina, Chile and Bolivia (
|
|
<bibRefCitation author="Renvoize, SA" journalOrPublisher="Alismataceae a Cyperaceae. Universidad Nacional Autonoma de Mexico, Mexico" pageId="0" pageNumber="107" refId="B29" refString="Renvoize, SA, 1998. Gramineas de Bolivia. Continental Printing, Belgium." title="Gramineas de Bolivia. Continental Printing, Belgium." year="1998">Renvoize 1998</bibRefCitation>
|
|
;
|
|
<bibRefCitation author="Rugolo de Agrasar, ZE" editor="Zuloaga, FO" journalOrPublisher="Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina" pageId="0" pageNumber="107" pagination="102 - 125" refId="B30" refString="Rugolo de Agrasar, ZE, 2012. Agrostis L. In: Zuloaga, FO, Rugolo de Agrasar, ZE, Anton, AM, Eds., Flora Argentina. Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina: 102 - 125" title="Agrostis L." volumeTitle="Flora Argentina." year="2012">
|
|
<normalizedToken originalValue="Rúgolo">Rugolo</normalizedToken>
|
|
de Agrasar 2012
|
|
</bibRefCitation>
|
|
), is similar in its overall appearance, usually being tufted and with rhizomes, having similar ligules and filiform or flat leaf blades, and having a condensed spikelike panicle with spikelets of similar size.
|
|
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis tolucensis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="tolucensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">Agrostis tolucensis</emphasis>
|
|
</taxonomicName>
|
|
can usually be differentiated from
|
|
<taxonomicName lsidName="A. meyenii" pageId="0" pageNumber="107" rank="species" species="meyenii">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. meyenii</emphasis>
|
|
</taxonomicName>
|
|
by the presence of an awn inserted in the lower third of the lemma, 2-3.5 mm long, twisted and bent and exerted from the glumes (vs. muticous or, if awn present, inserted in the middle or upper third of the lemma, to 1.2 mm long, straight or slightly flexuous in
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<taxonomicName lsidName="A. meyenii" pageId="0" pageNumber="107" rank="species" species="meyenii">
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<emphasis italics="true" pageId="0" pageNumber="107">A. meyenii</emphasis>
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</taxonomicName>
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). However, specimens akin to
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<taxonomicName lsidName="A. glomerata" pageId="0" pageNumber="107" rank="species" species="glomerata">
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<emphasis italics="true" pageId="0" pageNumber="107">A. glomerata</emphasis>
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</taxonomicName>
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, a species described from Peru and here considered a synonym of
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<taxonomicName lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
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<emphasis italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
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</taxonomicName>
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, can sometimes be found in Colombia, albeit not in the Cordillera Oriental, which have muticous lemmas or with a short straight awn inserted in the upper half of the lemma (e.g. Idrobo 3882a, Mutis 5521). These can be differentiated from
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<taxonomicName lsidName="A. meyenii" pageId="0" pageNumber="107" rank="species" species="meyenii">
|
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<emphasis italics="true" pageId="0" pageNumber="107">A. meyenii</emphasis>
|
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</taxonomicName>
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by the plants being generally taller, 20-60 cm tall, condensed panicles often> 10 cm long that are often interrupted and with the central inflorescence axis notably wider compared to the lateral branches, and the pedicels, panicle branches, and sometimes central inflorescence axis, being notably scabrous. The blades of these are variable and can be filiform to flat and to 5 mm wide. The characteristic of notably scabrous pedicels, panicle branches, and sometimes central inflorescence axis of
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<taxonomicName lsidName="A. tolucensis" pageId="0" pageNumber="107" rank="species" species="tolucensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="107">A. tolucensis</emphasis>
|
|
</taxonomicName>
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|
are considered to be key in differentiating these from
|
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<taxonomicName lsidName="A. meyenii" pageId="0" pageNumber="107" rank="species" species="meyenii">
|
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<emphasis italics="true" pageId="0" pageNumber="107">A. meyenii</emphasis>
|
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</taxonomicName>
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, which are usually smooth or exceptionally lightly scaberulous.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |