317 lines
42 KiB
XML
317 lines
42 KiB
XML
<document id="A9DCAEC82BC80C17CFE40C73F7E5BE42" ID-DOI="10.5281/zenodo.209356" ID-GBIF-Dataset="e370e9be-a92c-4197-990f-d12eef45a506" ID-ISSN="1175-5326" ID-Zenodo-Dep="209356" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460481483668" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2012" docId="DB6E0B263E0CFFE7FF73FE57FAB0B851" docLanguage="en" docName="zt03313p068.pdf" docOrigin="Zootaxa 3313" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Allohoraeomorphus Franz" docType="treatment" docVersion="9" lastPageNumber="64" masterDocId="2757735E3E0DFFE5FFE4FFC7FFEEBA46" masterDocTitle="Taxonomic status of Allohoraeomorphus Franz (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="68" masterPageNumber="62" pageNumber="63" updateTime="1698307848069" updateUser="plazi">
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<mods:title id="6467D99FBD8843F65611424FFD2C5CC9">Taxonomic status of Allohoraeomorphus Franz (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="460E46531D11CA5CFDC31507F199EBA7">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="96D13068DC20D124F351921C6B2F46B9">Zootaxa</mods:title>
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<mods:part id="FEAB0E7BD108736B22ECB13B0E2E5F0B">
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<mods:date id="F4F5752C3AEF88C24AE2A741961EBA5A">2012</mods:date>
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<mods:number id="7D97E6310947CF663C31389AC0E8C740">3313</mods:number>
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<mods:identifier id="4DADFC2CD5D12D3F73A9A7F184E75C8B" type="DOI">10.5281/zenodo.209356</mods:identifier>
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<treatment id="DB6E0B263E0CFFE7FF73FE57FAB0B851" ID-DOI="http://doi.org/10.5281/zenodo.6178623" ID-GBIF-Taxon="119517431" ID-Zenodo-Dep="6178623" LSID="urn:lsid:plazi:treatment:DB6E0B263E0CFFE7FF73FE57FAB0B851" httpUri="http://treatment.plazi.org/id/DB6E0B263E0CFFE7FF73FE57FAB0B851" lastPageId="2" lastPageNumber="64" pageId="1" pageNumber="63">
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<subSubSection id="1BDDE9BB3E0CFFE4FF73FE57FDA1BBEC" box="[151,591,400,427]" pageId="1" pageNumber="63" type="nomenclature">
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<paragraph id="5378BA303E0CFFE4FF73FE57FDA1BBEC" blockId="1.[151,591,400,427]" box="[151,591,400,427]" pageId="1" pageNumber="63">
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<heading id="08300D5C3E0CFFE4FF73FE57FDA1BBEC" bold="true" box="[151,591,400,427]" fontSize="11" level="1" pageId="1" pageNumber="63" reason="1">
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<emphasis id="61B366223E0CFFE4FF73FE57FDA1BBEC" bold="true" box="[151,591,400,427]" pageId="1" pageNumber="63">
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Status of
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<taxonomicName id="94C7C1B33E0CFFE4FEEFFE56FDA1BBEC" ID-CoL="8XJG9" authority="Franz" authorityName="Franz" box="[267,591,400,427]" class="Insecta" family="Staphylinidae" genus="Allohoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="genus">
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<emphasis id="61B366223E0CFFE4FEEFFE56FE10BBED" bold="true" box="[267,510,401,427]" italics="true" pageId="1" pageNumber="63">Allohoraeomorphus</emphasis>
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Franz
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="1BDDE9BB3E0CFFE7FF73FE1FFAB0B851" lastPageId="2" lastPageNumber="64" pageId="1" pageNumber="63" type="description">
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<paragraph id="5378BA303E0CFFE4FF73FE1FFE0AB9D9" blockId="1.[151,1437,471,2008]" pageId="1" pageNumber="63">
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<bibRefCitation id="3756C7C13E0CFFE4FF73FE1FFEDABBA9" author="Franz" box="[151,308,471,495]" pageId="1" pageNumber="63" refString="Franz, H. (1986 a) Zweiter Beitrag zur Kenntnis der Scydmaenidenfauna der Fiji-Inseln. Entomologische Blatter, 82, 147 - 178." type="journal article" year="1986" yearSuffix="a">Franz (1986a)</bibRefCitation>
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treated
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<taxonomicName id="94C7C1B33E0CFFE4FE69FE1FFD83BBB6" box="[397,621,472,496]" class="Insecta" family="Staphylinidae" genus="Allohoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="genus">
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<emphasis id="61B366223E0CFFE4FE69FE1FFD83BBB6" box="[397,621,472,496]" italics="true" pageId="1" pageNumber="63">Allohoraeomorphus</emphasis>
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</taxonomicName>
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as closely related to
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<taxonomicName id="94C7C1B33E0CFFE4FCB1FE1FFBE3BBB6" box="[853,1037,472,496]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="genus">
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<emphasis id="61B366223E0CFFE4FCB1FE1FFBE3BBB6" box="[853,1037,472,496]" italics="true" pageId="1" pageNumber="63">Horaeomorphus</emphasis>
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</taxonomicName>
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and his diagnosis was based on differences found in the structure of mesoventrite and pronotal base. Diagnostic characters of
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<taxonomicName id="94C7C1B33E0CFFE4FB7AFE3BFABBB852" box="[1182,1365,508,532]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="genus">
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<emphasis id="61B366223E0CFFE4FB7AFE3BFABBB852" box="[1182,1365,508,532]" italics="true" pageId="1" pageNumber="63">Horaeomorphus</emphasis>
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</taxonomicName>
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originally given by
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<bibRefCitation id="3756C7C13E0CFFE4FEA8FDD8FDFDB871" author="Schaufuss" box="[332,531,543,567]" pageId="1" pageNumber="63" refString="Schaufuss, L. W. (1889) Neue Scydmaeniden im Museum Ludwig Salvator. Berliner Entomologische Zeitschrift, 33, 1 - 42." type="journal article" year="1889">Schaufuss (1889)</bibRefCitation>
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were recently redefined by
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<bibRefCitation id="3756C7C13E0CFFE4FC80FDD8FBDAB871" author="Jaloszynski" box="[868,1076,543,567]" pageId="1" pageNumber="63" refString="Jaloszynski P. (2002) First record of the genus Horaeomorphus Schaufuss (Coleoptera, Scydmaenidae) from Japan, with description of a new species. Bulletin of National Science Museum Tokyo, 28 (4), 223 - 232." type="journal article" year="2002">Jałoszyński (2002</bibRefCitation>
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,
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<bibRefCitation id="3756C7C13E0CFFE4FBA0FDD8FB91B871" author="Jaloszynski" box="[1092,1151,543,567]" pageId="1" pageNumber="63" refString="Jaloszynski, P. (2003) Taxonomical notes on Southeast Asiatic species of Horaeomorphus Schaufuss (Coleoptera, Scydmaenidae), with description of a new species from Taiwan. Bulletin of National Science Museum Tokyo, 29 (2), 107 - 117." type="journal article" year="2003">2003</bibRefCitation>
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,
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<bibRefCitation id="3756C7C13E0CFFE4FB6BFDD8FB3BB871" author="Jaloszynski" box="[1167,1237,543,567]" pageId="1" pageNumber="63" refString="Jaloszynski P. (2004 a) The first record of Horaeomorphus Schaufuss (Coleoptera, Scydmaenidae) from the Philippines, with description of H. blattnyi sp. nov. Revue suisse de Zoologie, 111 (4), 785 - 789." type="journal article" year="2004" yearSuffix="a">2004a</bibRefCitation>
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) and include the typical slender and constricted body form (the "ant-like" appearance); the occipital part of head ("neck" region) broad; antennae gradually thickened distally, distinctly 11-segmented; the pronotum without lateral carinae, with ante-basal foveae; the prosternal process narrow, weakly separating procoxae; the mesoventral process distinct but weakly projecting ventrally; each elytron with two small basal foveae (in intact specimens concealed by the pronotal base); femora clavate; the aedeagus with free parameres and a complex endophallus. Additionally, nearly all species of
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<taxonomicName id="94C7C1B33E0CFFE4FEEAFD3FFE2BB956" box="[270,453,760,784]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="genus">
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<emphasis id="61B366223E0CFFE4FEEAFD3FFE2BB956" box="[270,453,760,784]" italics="true" pageId="1" pageNumber="63">Horaeomorphus</emphasis>
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</taxonomicName>
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have a pair of lateral pits on the frons, and some species show a distinct sexual dimorphism in the development of metatrochanters. The hind trochanters of males are often strongly elongated and forming a long rod-like projection running along femur, the trochanter in some cases exceeds half length of metafemur. Among
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<taxonomicName id="94C7C1B33E0CFFE4FF12FCA3FE72B93A" box="[246,412,868,892]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="subFamily" subFamily="Scydmaeninae">Scydmaeninae</taxonomicName>
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, only a single species of
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<taxonomicName id="94C7C1B33E0CFFE4FD2FFCA3FBF0B93A" authority="(Jaloszynski 2004b)" baseAuthorityName="Jaloszynski" baseAuthorityYear="2004" box="[715,1054,868,892]" class="Insecta" family="Staphylinidae" genus="Syndicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="genus">
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<emphasis id="61B366223E0CFFE4FD2FFCA3FCC1B93A" box="[715,815,868,892]" italics="true" pageId="1" pageNumber="63">Syndicus</emphasis>
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(
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<bibRefCitation id="3756C7C13E0CFFE4FCA5FCA3FBF8B93A" author="Jaloszynski" box="[833,1046,868,892]" pageId="1" pageNumber="63" refString="Jaloszynski, P. (2004 b) Revision of scydmaenid beetles of the genus Syndicus Motschulsky (Coleoptera, Scydmaenidae). National Science Museum Monographs No. 25, National Science Museum, Tokyo, 108 pp." type="book" year="2004" yearSuffix="b">Jałoszyński 2004b</bibRefCitation>
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)
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</taxonomicName>
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and
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<taxonomicName id="94C7C1B33E0CFFE4FBB8FCA3FB00B93A" box="[1116,1262,868,892]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="calcarifer">
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<emphasis id="61B366223E0CFFE4FBB8FCA3FB00B93A" box="[1116,1262,868,892]" italics="true" pageId="1" pageNumber="63">A. calcarifer</emphasis>
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</taxonomicName>
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have similarly shaped male metatrochanters.
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</paragraph>
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<paragraph id="5378BA303E0CFFE4FF23FC6BFBDBBEF9" blockId="1.[151,1437,471,2008]" pageId="1" pageNumber="63">
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Examination of a male
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<typeStatus id="8C7C04923E0CFFE4FE2FFC6AFDC2B983" box="[459,556,941,965]" pageId="1" pageNumber="63" type="paratype">paratype</typeStatus>
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of
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<taxonomicName id="94C7C1B33E0CFFE4FDB5FC6BFD31B982" box="[593,735,940,964]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="calcarifer">
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<emphasis id="61B366223E0CFFE4FDB5FC6BFD31B982" box="[593,735,940,964]" italics="true" pageId="1" pageNumber="63">A. calcarifer</emphasis>
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</taxonomicName>
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and direct comparison with
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<taxonomicName id="94C7C1B33E0CFFE4FBF9FC6AFB29B982" box="[1053,1223,940,964]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="eumicroides">
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<emphasis id="61B366223E0CFFE4FBF9FC6AFB29B982" box="[1053,1223,940,964]" italics="true" pageId="1" pageNumber="63">H. eumicroides</emphasis>
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</taxonomicName>
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revealed a number of characters shared by these two taxa, and only minor differences.
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<taxonomicName id="94C7C1B33E0CFFE4FC4AFC17FAE8B9AE" box="[942,1286,976,1000]" class="Insecta" family="Staphylinidae" genus="Allohoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="calcarifer">
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<emphasis id="61B366223E0CFFE4FC4AFC17FAE8B9AE" box="[942,1286,976,1000]" italics="true" pageId="1" pageNumber="63">Allohoraeomorphus calcarifer</emphasis>
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</taxonomicName>
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(
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<figureCitation id="CBFCA6B53E0CFFE4FAFDFC08FAB3B9A1" box="[1305,1373,975,999]" captionStart="FIGURE 1" captionStartId="3.[151,250,1975,1998]" captionText="FIGURE 1. Male habitus of Horaeomorphus calcarifer comb. n. Scale bar: 1 mm." pageId="1" pageNumber="63">Fig. 1</figureCitation>
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) is a large, darkly pigmented species, in body shape resembling
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<taxonomicName id="94C7C1B33E0CFFE4FCCFFC32FB81BE4A" authority="Franz, 1992" authorityName="Franz" authorityYear="1992" box="[811,1135,1012,1036]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="sarawakensis">
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<emphasis id="61B366223E0CFFE4FCCFFC32FC0ABE4A" box="[811,996,1012,1036]" italics="true" pageId="1" pageNumber="63">H. sarawakensis</emphasis>
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<bibRefCitation id="3756C7C13E0CFFE4FC0EFC32FB81BE4A" author="Franz" box="[1002,1135,1012,1036]" pageId="1" pageNumber="63" refString="Franz, H. (1992) Scydmaeniden aus Malaysia und Sulawesi (Coleoptera: Scydmaenidae). Koleopterologische Rundschau, 62, 69 - 76." type="journal article" year="1992">Franz, 1992</bibRefCitation>
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</taxonomicName>
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(from W and E
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<collectingCountry id="2BD0FAA03E0CFFE4FAC0FC33FA61BE4A" box="[1316,1423,1012,1036]" name="Malaysia" pageId="1" pageNumber="63">Malaysia</collectingCountry>
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),
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<taxonomicName id="94C7C1B33E0CFFE4FF73FBDEFE01BE69" authority="Franz, 1984" authorityName="Franz" authorityYear="1984" box="[151,495,1047,1072]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="valdepunctatus">
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<emphasis id="61B366223E0CFFE4FF73FBDEFE8CBE76" box="[151,354,1048,1072]" italics="true" pageId="1" pageNumber="63">H. valdepunctatus</emphasis>
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<bibRefCitation id="3756C7C13E0CFFE4FE8DFBDEFE01BE69" author="Franz" box="[361,495,1047,1072]" pageId="1" pageNumber="63" refString="Franz, H. (1984) Beitrag zur Kenntnis der Scydmaeniden von Sumatra und benachbarter Gebiete. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Abteilung I, 193, 89 - 142." type="journal article" year="1984">Franz, 1984</bibRefCitation>
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</taxonomicName>
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(W
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<collectingCountry id="2BD0FAA03E0CFFE4FDC4FBD0FD65BE69" box="[544,651,1047,1071]" name="Malaysia" pageId="1" pageNumber="63">Malaysia</collectingCountry>
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),
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<taxonomicName id="94C7C1B33E0CFFE4FD79FBDEFC1CBE69" authority="Franz, 1992" authorityName="Franz" authorityYear="1992" box="[669,1010,1047,1072]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="punctatissimus">
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<emphasis id="61B366223E0CFFE4FD79FBDEFC8BBE76" box="[669,869,1049,1072]" italics="true" pageId="1" pageNumber="63">H. punctatissimus</emphasis>
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<bibRefCitation id="3756C7C13E0CFFE4FC88FBDEFC1CBE69" author="Franz" box="[876,1010,1047,1072]" pageId="1" pageNumber="63" refString="Franz, H. (1992) Scydmaeniden aus Malaysia und Sulawesi (Coleoptera: Scydmaenidae). Koleopterologische Rundschau, 62, 69 - 76." type="journal article" year="1992">Franz, 1992</bibRefCitation>
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</taxonomicName>
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(E
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<collectingCountry id="2BD0FAA03E0CFFE4FBFEFBD0FB6DBE69" box="[1050,1155,1047,1071]" name="Malaysia" pageId="1" pageNumber="63">Malaysia</collectingCountry>
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,
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<collectingCountry id="2BD0FAA03E0CFFE4FB6AFBD0FAEEBE69" box="[1166,1280,1047,1071]" name="Indonesia" pageId="1" pageNumber="63">Indonesia</collectingCountry>
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),
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<taxonomicName id="94C7C1B33E0CFFE4FAF6FBDEFE6CBE12" authority="Jaloszynski, 2009" authorityName="Jaloszynski" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="fakfakensis">
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<emphasis id="61B366223E0CFFE4FAF6FBDEFF5BBE12" italics="true" pageId="1" pageNumber="63">H. fakfakensis</emphasis>
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<bibRefCitation id="3756C7C13E0CFFE4FF5FFBFBFE6CBE12" author="Jaloszynski" box="[187,386,1084,1108]" pageId="1" pageNumber="63" refString="Jaloszynski P. (2009) Four new species of Horaeomorphus Schaufuss from Oriental Region (Coleoptera: Scydmaenidae). Raffles Bulletin of Zoology, 57 (2), 297 - 303." type="journal article" year="2009">Jałoszyński, 2009</bibRefCitation>
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</taxonomicName>
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(New
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<collectingCountry id="2BD0FAA03E0CFFE4FE2AFBFBFDCFBE12" box="[462,545,1084,1108]" name="Guinea-Bissau" pageId="1" pageNumber="63">Guinea</collectingCountry>
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) and
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<taxonomicName id="94C7C1B33E0CFFE4FDBAFBFAFBB6BE12" authority="Jaloszynski, 2009" authorityName="Jaloszynski" authorityYear="2009" box="[606,1112,1084,1108]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="samosirensis">
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<emphasis id="61B366223E0CFFE4FDBAFBFAFCFEBE12" box="[606,784,1085,1108]" italics="true" pageId="1" pageNumber="63">H. samosirensis</emphasis>
|
||
<bibRefCitation id="3756C7C13E0CFFE4FCF3FBFBFC30BE12" author="Jaloszynski" box="[791,990,1084,1108]" pageId="1" pageNumber="63" refString="Jaloszynski P. (2009) Four new species of Horaeomorphus Schaufuss from Oriental Region (Coleoptera: Scydmaenidae). Raffles Bulletin of Zoology, 57 (2), 297 - 303." type="journal article" year="2009">Jałoszyński, 2009</bibRefCitation>
|
||
(Sumatra)
|
||
</taxonomicName>
|
||
. All these species have strikingly large pronotum, in dorsal view nearly circular and nearly as broad as elytra. Most species of
|
||
<taxonomicName id="94C7C1B33E0CFFE4FB00FBA7FA75BE3E" box="[1252,1435,1120,1144]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="61B366223E0CFFE4FB00FBA7FA75BE3E" box="[1252,1435,1120,1144]" italics="true" pageId="1" pageNumber="63">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
are brown, but such unusually dark, nearly black body pigmentation as that of
|
||
<taxonomicName id="94C7C1B33E0CFFE4FBD6FB43FB2CBEDA" box="[1074,1218,1156,1180]" class="Insecta" family="Staphylinidae" genus="Allohoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="calcarifer">
|
||
<emphasis id="61B366223E0CFFE4FBD6FB43FB2CBEDA" box="[1074,1218,1156,1180]" italics="true" pageId="1" pageNumber="63">A. calcarifer</emphasis>
|
||
</taxonomicName>
|
||
can also be found within this genus, e.g. in
|
||
<taxonomicName id="94C7C1B33E0CFFE4FE54FB6EFCEDBEF9" authority="Jaloszynski, 2004" authorityName="Jaloszynski" authorityYear="2004" box="[432,771,1191,1216]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="blattnyi">
|
||
<emphasis id="61B366223E0CFFE4FE54FB6EFDC7BE86" box="[432,553,1192,1216]" italics="true" pageId="1" pageNumber="63">H. blattnyi</emphasis>
|
||
<bibRefCitation id="3756C7C13E0CFFE4FDCBFB60FCEDBEF9" author="Jaloszynski" box="[559,771,1191,1215]" pageId="1" pageNumber="63" refString="Jaloszynski P. (2004 a) The first record of Horaeomorphus Schaufuss (Coleoptera, Scydmaenidae) from the Philippines, with description of H. blattnyi sp. nov. Revue suisse de Zoologie, 111 (4), 785 - 789." type="journal article" year="2004" yearSuffix="a">Jałoszyński, 2004a</bibRefCitation>
|
||
</taxonomicName>
|
||
(from Luzon,
|
||
<collectingCountry id="2BD0FAA03E0CFFE4FC42FB60FBC4BEF9" box="[934,1066,1191,1215]" name="Philippines" pageId="1" pageNumber="63">Philippines</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5378BA303E0CFFE4FF23FB0BFE33BF09" blockId="1.[151,1437,471,2008]" pageId="1" pageNumber="63">
|
||
The lack of posterolateral pronotal corners, mentioned by Franz (1986) as one of two diagnostic characters of
|
||
<taxonomicName id="94C7C1B33E0CFFE4FF73FB37FE99BF4E" box="[151,375,1264,1288]" class="Insecta" family="Staphylinidae" genus="Allohoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="61B366223E0CFFE4FF73FB37FE99BF4E" box="[151,375,1264,1288]" italics="true" pageId="1" pageNumber="63">Allohoraeomorphus</emphasis>
|
||
</taxonomicName>
|
||
, was apparently misinterpreted. The pronotum of
|
||
<taxonomicName id="94C7C1B33E0CFFE4FC40FB37FBDEBF4E" box="[932,1072,1264,1288]" class="Insecta" family="Staphylinidae" genus="Allohoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="calcarifer">
|
||
<emphasis id="61B366223E0CFFE4FC40FB37FBDEBF4E" box="[932,1072,1264,1288]" italics="true" pageId="1" pageNumber="63">A. calcarifer</emphasis>
|
||
</taxonomicName>
|
||
, although with strongly rounded sides, has distinct, obtuse hind angles. This character is developed to a similar extent as in all species of
|
||
<taxonomicName id="94C7C1B33E0CFFE4FAD9FAD2FF17BF16" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="61B366223E0CFFE4FAD9FAD2FF17BF16" italics="true" pageId="1" pageNumber="63">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
with broad pronota.
|
||
</paragraph>
|
||
<paragraph id="5378BA303E0CFFE4FF23FA9BFD8EBC0B" blockId="1.[151,1437,471,2008]" pageId="1" pageNumber="63">
|
||
The other diagnostic character mentioned by Franz (1986), the shape of mesoventral process, which is weakly (
|
||
<taxonomicName id="94C7C1B33E0CFFE4FF44FA47FE6EBFDE" box="[160,384,1408,1432]" class="Insecta" family="Staphylinidae" genus="Allohoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="61B366223E0CFFE4FF44FA47FE6EBFDE" box="[160,384,1408,1432]" italics="true" pageId="1" pageNumber="63">Allohoraeomorphus</emphasis>
|
||
</taxonomicName>
|
||
) or strongly (
|
||
<taxonomicName id="94C7C1B33E0CFFE4FDFEFA47FD3FBFDE" box="[538,721,1408,1432]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="61B366223E0CFFE4FDFEFA47FD3FBFDE" box="[538,721,1408,1432]" italics="true" pageId="1" pageNumber="63">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
) expanded ventrally, is problematic. In
|
||
<taxonomicName id="94C7C1B33E0CFFE4FB6AFA47FAF4BFDE" box="[1166,1306,1408,1432]" class="Insecta" family="Staphylinidae" genus="Allohoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="calcarifer">
|
||
<emphasis id="61B366223E0CFFE4FB6AFA47FAF4BFDE" box="[1166,1306,1408,1432]" italics="true" pageId="1" pageNumber="63">A. calcarifer</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="CBFCA6B53E0CFFE4FACEFAB8FA7CBFD1" box="[1322,1426,1407,1431]" captionStart="FIGURES 2 – 5" captionStartId="4.[151,264,1930,1953]" captionTargetBox="[254,1373,198,1882]" captionTargetId="figure@4.[188,1391,193,1890]" captionTargetPageId="4" captionText="FIGURES 2 – 5. Body venter of Horaeomorphus calcarifer comb. n. (2, 3) (abdomen removed) and Horaeomorphus eumicroides Schaufuss (4, 5). Abbreviations: bs — prothoracic basisternum; hy — prothoracic hypomeron; msp — mesoventral process; msv — mesoventrite; mtp — metaventral process; mtv — metaventrite; pp — prosternal process. Scale bars: 1 mm." httpUri="https://zenodo.org/record/209357/files/figure.png" pageId="1" pageNumber="63">Figs. 2, 3</figureCitation>
|
||
) this process (msp) is indeed shorter and lower (i.e., less expanded ventrally) than in
|
||
<taxonomicName id="94C7C1B33E0CFFE4FBADFA62FB1ABFFA" box="[1097,1268,1444,1468]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="eumicroides">
|
||
<emphasis id="61B366223E0CFFE4FBADFA62FB1ABFFA" box="[1097,1268,1444,1468]" italics="true" pageId="1" pageNumber="63">H. eumicroides</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="CBFCA6B53E0CFFE4FAE7FA63FA80BFFA" box="[1283,1390,1444,1468]" captionStart="FIGURES 2 – 5" captionStartId="4.[151,264,1930,1953]" captionTargetBox="[254,1373,198,1882]" captionTargetId="figure@4.[188,1391,193,1890]" captionTargetPageId="4" captionText="FIGURES 2 – 5. Body venter of Horaeomorphus calcarifer comb. n. (2, 3) (abdomen removed) and Horaeomorphus eumicroides Schaufuss (4, 5). Abbreviations: bs — prothoracic basisternum; hy — prothoracic hypomeron; msp — mesoventral process; msv — mesoventrite; mtp — metaventral process; mtv — metaventrite; pp — prosternal process. Scale bars: 1 mm." httpUri="https://zenodo.org/record/209357/files/figure.png" pageId="1" pageNumber="63">Figs. 4, 5</figureCitation>
|
||
), or in all other species of Asian
|
||
<taxonomicName id="94C7C1B33E0CFFE4FE39FA0FFD7ABFA6" box="[477,660,1480,1504]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="61B366223E0CFFE4FE39FA0FFD7ABFA6" box="[477,660,1480,1504]" italics="true" pageId="1" pageNumber="63">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
previously studied by the author. However, the general shape of the mesoventral process and the structure of the entire mesoventrite in
|
||
<taxonomicName id="94C7C1B33E0CFFE4FC6FFA2BFBF6BC42" box="[907,1048,1516,1540]" class="Insecta" family="Staphylinidae" genus="Allohoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="calcarifer">
|
||
<emphasis id="61B366223E0CFFE4FC6FFA2BFBF6BC42" box="[907,1048,1516,1540]" italics="true" pageId="1" pageNumber="63">A. calcarifer</emphasis>
|
||
</taxonomicName>
|
||
and in
|
||
<taxonomicName id="94C7C1B33E0CFFE4FB89FA2AFAF6BC42" box="[1133,1304,1516,1540]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="eumicroides">
|
||
<emphasis id="61B366223E0CFFE4FB89FA2AFAF6BC42" box="[1133,1304,1516,1540]" italics="true" pageId="1" pageNumber="63">H. eumicroides</emphasis>
|
||
</taxonomicName>
|
||
are similar. The difference in the shape of mesoventral processes is too small to unambiguously define the length and height that could be diagnostic for either genus.
|
||
</paragraph>
|
||
<paragraph id="5378BA303E0CFFE4FF23F990FDC3BD2A" blockId="1.[151,1437,471,2008]" pageId="1" pageNumber="63">
|
||
The diagnostic characters used by
|
||
<bibRefCitation id="3756C7C13E0CFFE4FDB6F99EFD1FBC29" author="Franz" box="[594,753,1623,1648]" pageId="1" pageNumber="63" refString="Franz, H. (1986 a) Zweiter Beitrag zur Kenntnis der Scydmaenidenfauna der Fiji-Inseln. Entomologische Blatter, 82, 147 - 178." type="journal article" year="1986" yearSuffix="a">Franz (1986a)</bibRefCitation>
|
||
to define
|
||
<taxonomicName id="94C7C1B33E0CFFE4FC83F99FFBA9BC36" box="[871,1095,1624,1648]" class="Insecta" family="Staphylinidae" genus="Allohoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="61B366223E0CFFE4FC83F99FFBA9BC36" box="[871,1095,1624,1648]" italics="true" pageId="1" pageNumber="63">Allohoraeomorphus</emphasis>
|
||
</taxonomicName>
|
||
are insufficient to distinguish this genus from
|
||
<taxonomicName id="94C7C1B33E0CFFE4FEB2F9BBFDE3BCD2" box="[342,525,1660,1684]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="61B366223E0CFFE4FEB2F9BBFDE3BCD2" box="[342,525,1660,1684]" italics="true" pageId="1" pageNumber="63">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
. However, two additional differences were found during the present study. As seen in
|
||
<figureCitation id="CBFCA6B53E0CFFE4FF09F958FEB6BCF1" box="[237,344,1695,1719]" captionStart="FIGURES 2 – 5" captionStartId="4.[151,264,1930,1953]" captionTargetBox="[254,1373,198,1882]" captionTargetId="figure@4.[188,1391,193,1890]" captionTargetPageId="4" captionText="FIGURES 2 – 5. Body venter of Horaeomorphus calcarifer comb. n. (2, 3) (abdomen removed) and Horaeomorphus eumicroides Schaufuss (4, 5). Abbreviations: bs — prothoracic basisternum; hy — prothoracic hypomeron; msp — mesoventral process; msv — mesoventrite; mtp — metaventral process; mtv — metaventrite; pp — prosternal process. Scale bars: 1 mm." httpUri="https://zenodo.org/record/209357/files/figure.png" pageId="1" pageNumber="63">Figs. 2–5</figureCitation>
|
||
, the prothoracic basisternum (bs) of the compared species differs in length—that in
|
||
<taxonomicName id="94C7C1B33E0CFFE4FAEAF967FA75BCFE" box="[1294,1435,1696,1720]" class="Insecta" family="Staphylinidae" genus="Allohoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="calcarifer">
|
||
<emphasis id="61B366223E0CFFE4FAEAF967FA75BCFE" box="[1294,1435,1696,1720]" italics="true" pageId="1" pageNumber="63">A. calcarifer</emphasis>
|
||
</taxonomicName>
|
||
is longer than procoxae, while that of
|
||
<taxonomicName id="94C7C1B33E0CFFE4FDDFF902FD0ABC9A" box="[571,740,1732,1756]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="eumicroides">
|
||
<emphasis id="61B366223E0CFFE4FDDFF902FD0ABC9A" box="[571,740,1732,1756]" italics="true" pageId="1" pageNumber="63">H. eumicroides</emphasis>
|
||
</taxonomicName>
|
||
is distinctly shorter. Also the prosternal process (pp) of
|
||
<taxonomicName id="94C7C1B33E0CFFE4FAB6F903FF0DBD46" class="Insecta" family="Staphylinidae" genus="Allohoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="calcarifer">
|
||
<emphasis id="61B366223E0CFFE4FAB6F903FF0DBD46" italics="true" pageId="1" pageNumber="63">A. calcarifer</emphasis>
|
||
</taxonomicName>
|
||
is longer than that of
|
||
<taxonomicName id="94C7C1B33E0CFFE4FE3EF92EFD6BBD46" box="[474,645,1768,1792]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="eumicroides">
|
||
<emphasis id="61B366223E0CFFE4FE3EF92EFD6BBD46" box="[474,645,1768,1792]" italics="true" pageId="1" pageNumber="63">H. eumicroides</emphasis>
|
||
</taxonomicName>
|
||
. However, it is difficult to define a clear-cut limit between two genera on the basis of a shorter or longer basisternum alone. Such length differences in
|
||
<taxonomicName id="94C7C1B33E0CFFE4FBD7F8CBFB37BD62" box="[1075,1241,1804,1828]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="subFamily" subFamily="Scydmaeninae">Scydmaeninae</taxonomicName>
|
||
, if not correlated with other significantly different structures, are generally treated as a morphological diversity within a genus, and not as generic diagnostic characters.
|
||
</paragraph>
|
||
<paragraph id="5378BA303E0CFFE7FF23F8BFFEC6BBEA" blockId="1.[151,1437,471,2008]" lastBlockId="2.[151,1436,151,536]" lastPageId="2" lastPageNumber="64" pageId="1" pageNumber="63">
|
||
Another difference between
|
||
<taxonomicName id="94C7C1B33E0CFFE4FDE3F8BFFD7BBDD6" box="[519,661,1912,1936]" class="Insecta" family="Staphylinidae" genus="Allohoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="calcarifer">
|
||
<emphasis id="61B366223E0CFFE4FDE3F8BFFDF6BDD6" box="[519,536,1912,1936]" italics="true" pageId="1" pageNumber="63">A</emphasis>
|
||
.
|
||
<emphasis id="61B366223E0CFFE4FDC3F8BFFD7BBDD6" box="[551,661,1912,1936]" italics="true" pageId="1" pageNumber="63">calcarifer</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="94C7C1B33E0CFFE4FD29F8BEFC99BDD6" box="[717,887,1912,1936]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="eumicroides">
|
||
<emphasis id="61B366223E0CFFE4FD29F8BEFC99BDD6" box="[717,887,1912,1936]" italics="true" pageId="1" pageNumber="63">H. eumicroides</emphasis>
|
||
</taxonomicName>
|
||
is the lack of pits on the head of the former species. The frons of nearly all species of
|
||
<taxonomicName id="94C7C1B33E0CFFE4FDB4F85BFCE9BDF2" box="[592,775,1948,1972]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="61B366223E0CFFE4FDB4F85BFCE9BDF2" box="[592,775,1948,1972]" italics="true" pageId="1" pageNumber="63">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
, including
|
||
<taxonomicName id="94C7C1B33E0CFFE4FC6DF85AFBDABDF2" box="[905,1076,1948,1972]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="eumicroides">
|
||
<emphasis id="61B366223E0CFFE4FC6DF85AFBDABDF2" box="[905,1076,1948,1972]" italics="true" pageId="1" pageNumber="63">H. eumicroides</emphasis>
|
||
</taxonomicName>
|
||
, bears variously developed, but usually well-visible lateral pits, each located at the posteromesal margin of supraantennal tubercle (illustrated in
|
||
<bibRefCitation id="3756C7C13E0FFFE7FF73FF50FE8FBAE9" author="Jaloszynski" box="[151,353,151,175]" pageId="2" pageNumber="64" refString="Jaloszynski, P. (2003) Taxonomical notes on Southeast Asiatic species of Horaeomorphus Schaufuss (Coleoptera, Scydmaenidae), with description of a new species from Taiwan. Bulletin of National Science Museum Tokyo, 29 (2), 107 - 117." type="journal article" year="2003">Jałoszyński, 2003</bibRefCitation>
|
||
; figs. 2A, B). This character was included in the generic diagnosis (
|
||
<bibRefCitation id="3756C7C13E0FFFE7FBBCFF50FAF0BAE9" author="Jaloszynski" box="[1112,1310,151,175]" pageId="2" pageNumber="64" refString="Jaloszynski P. (2002) First record of the genus Horaeomorphus Schaufuss (Coleoptera, Scydmaenidae) from Japan, with description of a new species. Bulletin of National Science Museum Tokyo, 28 (4), 223 - 232." type="journal article" year="2002">Jałoszyński 2002</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="3756C7C13E0FFFE7FACDFF50FA8BBAE9" author="Jaloszynski" box="[1321,1381,151,175]" pageId="2" pageNumber="64" refString="Jaloszynski, P. (2003) Taxonomical notes on Southeast Asiatic species of Horaeomorphus Schaufuss (Coleoptera, Scydmaenidae), with description of a new species from Taiwan. Bulletin of National Science Museum Tokyo, 29 (2), 107 - 117." type="journal article" year="2003">2003</bibRefCitation>
|
||
), but subsequent revisions revealed a significant extent of variability in diameter and depth not only between, but also within species. For instance, the pits in
|
||
<taxonomicName id="94C7C1B33E0FFFE7FDABFF26FCE6BABE" box="[591,776,224,248]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="species" species="sarawakensis">
|
||
<emphasis id="61B366223E0FFFE7FDABFF26FCE6BABE" box="[591,776,224,248]" italics="true" pageId="2" pageNumber="64">H. sarawakensis</emphasis>
|
||
</taxonomicName>
|
||
are shallow and in some individuals can be barely discernible (
|
||
<bibRefCitation id="3756C7C13E0FFFE7FF36FEC3FE4DBB5A" author="Jaloszynski" box="[210,419,260,284]" pageId="2" pageNumber="64" refString="Jaloszynski P. (2006) Revision of Horaeomorphus Schaufuss (Coleoptera, Scydmaenidae) of East Malaysia, Singapore and Sunda Islands. Genus, 17 (1), 19 - 66." type="journal article" year="2006">Jałoszyński, 2006</bibRefCitation>
|
||
). It seems that the depth of pits correlates with the development of supraantennal tuber-cles—in species with weakly raised tubercles the pits are shallow or indistinct. The supraantennal tubercles are weakly raised in
|
||
<taxonomicName id="94C7C1B33E0FFFE7FEB7FE8BFE31BB22" box="[339,479,332,356]" class="Insecta" family="Staphylinidae" genus="Allohoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="species" species="calcarifer">
|
||
<emphasis id="61B366223E0FFFE7FEB7FE8BFE31BB22" box="[339,479,332,356]" italics="true" pageId="2" pageNumber="64">A. calcarifer</emphasis>
|
||
</taxonomicName>
|
||
, and additionally its frons and vertex are more convex than in Asian
|
||
<taxonomicName id="94C7C1B33E0FFFE7FB39FE8BFA7ABB22" box="[1245,1428,332,356]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="61B366223E0FFFE7FB39FE8BFA7ABB22" box="[1245,1428,332,356]" italics="true" pageId="2" pageNumber="64">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
. The presence or lack of pits may be misleading and should not be treated as a primary diagnostic character of
|
||
<taxonomicName id="94C7C1B33E0FFFE7FA82FEB6FECCBBEA" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="61B366223E0FFFE7FA82FEB6FECCBBEA" italics="true" pageId="2" pageNumber="64">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5378BA303E0FFFE7FF23FE7FFAB0B851" blockId="2.[151,1436,151,536]" pageId="2" pageNumber="64">
|
||
In conclusion, the previously listed unique characters of
|
||
<taxonomicName id="94C7C1B33E0FFFE7FCDAFE7FFBF0BB96" box="[830,1054,440,464]" class="Insecta" family="Staphylinidae" genus="Allohoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="61B366223E0FFFE7FCDAFE7FFBF0BB96" box="[830,1054,440,464]" italics="true" pageId="2" pageNumber="64">Allohoraeomorphus</emphasis>
|
||
</taxonomicName>
|
||
were found to have little diagnostic value, and the observed differences between
|
||
<taxonomicName id="94C7C1B33E0FFFE7FD52FE1BFCADBBB2" box="[694,835,476,500]" class="Insecta" family="Staphylinidae" genus="Allohoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="species" species="calcarifer">
|
||
<emphasis id="61B366223E0FFFE7FD52FE1BFCADBBB2" box="[694,835,476,500]" italics="true" pageId="2" pageNumber="64">A. calcarifer</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="94C7C1B33E0FFFE7FC98FE1AFBC9BBB2" box="[892,1063,476,500]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="species" species="eumicroides">
|
||
<emphasis id="61B366223E0FFFE7FC98FE1AFBC9BBB2" box="[892,1063,476,500]" italics="true" pageId="2" pageNumber="64">H. eumicroides</emphasis>
|
||
</taxonomicName>
|
||
can be treated as variable within
|
||
<taxonomicName id="94C7C1B33E0FFFE7FF73FDC7FEA1B85E" box="[151,335,512,536]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="61B366223E0FFFE7FF73FDC7FEA1B85E" box="[151,335,512,536]" italics="true" pageId="2" pageNumber="64">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
. Therefore,
|
||
<taxonomicName id="94C7C1B33E0FFFE7FE3DFDC7FD57B85E" box="[473,697,512,536]" class="Insecta" family="Staphylinidae" genus="Allohoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="61B366223E0FFFE7FE3DFDC7FD57B85E" box="[473,697,512,536]" italics="true" pageId="2" pageNumber="64">Allohoraeomorphus</emphasis>
|
||
</taxonomicName>
|
||
is placed as a junior objective synonym of
|
||
<taxonomicName id="94C7C1B33E0FFFE7FB7AFDC7FABBB85E" box="[1182,1365,512,536]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="61B366223E0FFFE7FB7AFDC7FABBB85E" box="[1182,1365,512,536]" italics="true" pageId="2" pageNumber="64">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |