332 lines
42 KiB
XML
332 lines
42 KiB
XML
<document id="00F9AAABA1776E841B16487CF0AEEB55" ID-DOI="10.1649/0010" ID-ISSN="1938-4394" ID-Zenodo-Dep="10110906" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630265353800" checkinUser="valdenar" docAuthor="López-Guerrero, Irma" docDate="2007" docId="CC6C1721FFB0FFCC1569FA8AFC19FD6A" docLanguage="en" docName="0010-065X_2007_61_463_ARNSOM_2.0.CO_2.pdf" docOrigin="The Coleopterists Bulletin 61 (3)" docSource="http://dx.doi.org/10.5281/zenodo.5326738" docStyle="DocumentStyle:76939A7658C5F80CF4BB6818E728552A.2:TheColeopBull.2003-2009.journal_article.1cover" docStyleId="76939A7658C5F80CF4BB6818E728552A" docStyleName="TheColeopBull.2003-2009.journal_article.1cover" docStyleVersion="2" docTitle="Megaceras briansaltini Ratcliffe 2007, new species" docType="treatment" docVersion="4" lastPageNumber="467" masterDocId="30556F59FFB1FFC91402FF87FFA3FF81" masterDocTitle="Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits" masterLastPageNumber="467" masterPageNumber="463" pageNumber="463" updateTime="1700023696164" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="6756BAF9B307B8567AEF86993A9DB4BB">
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<mods:title id="1E6F1C7364A25E7ACDFD02E31B3B95AB">Figs. 30 – 33. Attavicinus monstrosus. 30 in Comparison of Mouthpart Morphology of Three Species of Mexican Oniticellini (Coleoptera: Scarabaeidae: Scarabaeinae) in Relation to Their Trophic Habits</mods:title>
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<mods:name id="3AE985B228A0706BAAB9B3D67D324F68" type="personal">
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<mods:roleTerm id="B43E31A528794982BFBB2FD77B314FE8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C3654B44A841079090925DACC94D3C14">López-Guerrero, Irma</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="7C5F36534E639CE81BB18D82CCB916A7">text</mods:typeOfResource>
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<mods:titleInfo id="E058E4B4E0BF09B1DC63F506613CEB28">
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<mods:title id="1906339102A0D0A61D0EA517D40DEBF8">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="295DE1CD74BF8BE1961740E8A0EF9A74">
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<mods:date id="03865875073A4DD16D8250811B7A480E">2007</mods:date>
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<mods:detail id="8A94B0155AE1F08AD9F0C32A16C9CDBA" type="volume">
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<mods:number id="337D3B21BF748D218CEB452FF2CA2774">61</mods:number>
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</mods:detail>
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<mods:detail id="5580A4B36CB6CBDA492BDF85CBFC8E2C" type="issue">
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<mods:number id="D182C1EADE90041402584B30890B4BA6">3</mods:number>
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<mods:start id="3C551ED23816447C89AE3115E0CF3814">463</mods:start>
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<mods:end id="F77D4D3AAA9BD4D714FBE6E417F7F6BC">467</mods:end>
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<mods:location id="D8465D2FE229ED4C377FD3BFFECD5AD8">
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<mods:url id="524B51A178CF746D3A3F490C18F89094">http://dx.doi.org/10.5281/zenodo.5326738</mods:url>
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<mods:classification id="C8C0F8B7C895B6E88548F0BB51931421">journal article</mods:classification>
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<mods:identifier id="05A44A9D0B65B84BAD008B95CF59AF23" type="DOI">10.1649/0010</mods:identifier>
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<mods:identifier id="64FB7ACB69A85FD92714903B9B30B156" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="9BC8B5559A1FE59708E063EA12E4A13E" type="Zenodo-Dep">10110906</mods:identifier>
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</mods:mods>
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<treatment id="CC6C1721FFB0FFCC1569FA8AFC19FD6A" LSID="urn:lsid:plazi:treatment:CC6C1721FFB0FFCC1569FA8AFC19FD6A" httpUri="http://treatment.plazi.org/id/CC6C1721FFB0FFCC1569FA8AFC19FD6A" lastPageId="5" pageId="1">
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<subSubSection id="0CDFF5BCFFB0FFC81569FA8AFC93FAA2" box="[363,816,1293,1315]" pageId="1" type="nomenclature">
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<paragraph id="447AA637FFB0FFC81569FA8AFC93FAA2" blockId="1.[363,816,1293,1343]" box="[363,816,1293,1315]" pageId="1">
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<heading id="1F32115BFFB0FFC81569FA8AFC93FAA2" box="[363,816,1293,1315]" centered="true" fontSize="9" level="2" pageId="1" reason="2">
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<emphasis id="76B17A25FFB0FFC81569FA8AFC93FAA2" bold="true" box="[363,816,1293,1315]" pageId="1">
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<taxonomicName id="83C5DDB4FFB0FFC81569FA8AFD12FAA2" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="2007" box="[363,689,1293,1315]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="briansaltini" status="sp. nov.">
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<emphasis id="76B17A25FFB0FFC81569FA8AFDF3FAA2" bold="true" box="[363,592,1293,1315]" italics="true" pageId="1">Megaceras briansaltini</emphasis>
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Ratcliffe
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</taxonomicName>
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,
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<taxonomicNameLabel id="6D82C75EFFB0FFC816BCFA8AFC93FAA2" box="[702,816,1293,1315]" pageId="1" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0CDFF5BCFFB0FFC815DFFAAEFD1DFABE" box="[477,702,1321,1343]" pageId="1" type="description">
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<paragraph id="447AA637FFB0FFC815DFFAAEFD1DFABE" blockId="1.[363,816,1293,1343]" box="[477,702,1321,1343]" pageId="1">
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(
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<figureCitation id="DCFEBAB2FFB0FFC815E7FAAEFD31FABE" box="[485,658,1321,1343]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="2.[167,206,1044,1063]" captionStartId-1="3.[196,235,1373,1392]" captionTargetBox-0="[273,849,198,1022]" captionTargetBox-1="[184,996,197,1348]" captionTargetId-0="figure-196@2.[272,850,194,1022]" captionTargetId-1="figure-1@3.[181,998,194,1350]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figs. 1–2. Megaceras saltini, holotype, dorsal and lateral views, respectively. Photos courtesy of J. Saltin." captionText-1="Figs. 3–10. Dorsal view of head of (3) M. saltini and (4) M. morpheus showing form of mandibles. Lateral view of pygidium of (5) M. saltini and (6) M. morpheus showing surface convexity. Parameres (caudal and lateral views) of (7–8) M. saltini and (9–10) M. morpheus." figureDoi-0="http://doi.org/10.5281/zenodo.10129326" figureDoi-1="http://doi.org/10.5281/zenodo.10129328" httpUri-0="https://zenodo.org/record/10129326/files/figure.png" httpUri-1="https://zenodo.org/record/10129328/files/figure.png" pageId="1">Figs. 1–3, 5, 7–8</figureCitation>
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,
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<figureCitation id="DCFEBAB2FFB0FFC8169CFAAEFD15FABE" box="[670,694,1321,1343]" captionStart="Fig" captionStartId="4.[167,198,667,686]" captionTargetBox="[146,976,195,641]" captionTargetId="figure-360@4.[146,976,194,645]" captionTargetPageId="4" captionText="Fig. 11. Lateral view of head of Dim and M. saltini showing similarity in horn configuration (‘‘the Dim Effect’’). Dim character º Disney Enterprises, Inc. and Pixar. Used by permission from Disney Enterprises, Inc." figureDoi="http://doi.org/10.5281/zenodo.10129330" httpUri="https://zenodo.org/record/10129330/files/figure.png" pageId="1">11</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="0CDFF5BCFFB0FFCD14C5FAD6FE52FC71" lastPageId="4" pageId="1" type="materials_examined">
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<paragraph id="447AA637FFB0FFC814C5FAD6FE78FA53" blockId="1.[175,1005,1361,1570]" pageId="1">
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<emphasis id="76B17A25FFB0FFC814C5FAD6FEC5FAE6" bold="true" box="[199,358,1361,1383]" pageId="1">Type Material.</emphasis>
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<materialsCitation id="F4ADAC6AFFB0FFC81576FAD6FE6FFA36" collectingDate="2006-06" collectorName="Coll. J. - P. & Saltin' & Calabaza" country="Peru" elevation="2000" location="Junin" pageId="1" specimenCount="2" stateProvince="Junin" typeStatus="holotype">
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<typeStatus id="9B7E1895FFB0FFC81576FAD6FE77FAE6" box="[372,468,1361,1383]" pageId="1" type="holotype">Holotype</typeStatus>
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male, labeled ‘‘DYNASTINAE, Dynastinae sp.,
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<collectingCountry id="3CD2E6A7FFB0FFC814ADFAEBFF54FA03" box="[175,247,1388,1410]" name="Peru" pageId="1">PERU</collectingCountry>
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: Calabaza (5street from Satipo to Huancayo),
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<date id="307B80F7FFB0FFC816F1FAEBFCEAFA03" box="[755,841,1388,1410]" pageId="1" value="2006-06">
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<collectingDate id="203F791FFFB0FFC816F1FAEBFCEAFA03" box="[755,841,1388,1410]" pageId="1" value="2006-06">VI-2006</collectingDate>
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</date>
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, JPSCOLLNO: DYN/0/P00E,
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<collectorName id="E930C3E1FFB0FFC8154AFA01FDA2FA1D" box="[328,513,1414,1436]" pageId="1">Coll. J.-P. Saltin’</collectorName>
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’, ‘‘
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<collectorName id="E930C3E1FFB0FFC8161EFA01FDDCFA1D" box="[540,639,1414,1436]" pageId="1">Calabaza</collectorName>
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<quantity id="833D0BD2FFB0FFC8168AFA01FCA5FA1D" box="[648,774,1414,1436]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="1" unit="m" value="2000.0" valueMax="2200.0" valueMin="1800.0">
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<elevation id="CFE84104FFB0FFC8168AFA01FCA5FA1D" box="[648,774,1414,1436]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="1" unit="m" value="2000.0" valueMax="2200.0" valueMin="1800.0">1800-2200m</elevation>
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</quantity>
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,
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<collectingRegion id="860168D5FFB0FFC81711FA01FCE8FA1D" box="[787,843,1414,1436]" country="Peru" name="Junin" pageId="1">Junin</collectingRegion>
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<collectingCountry id="3CD2E6A7FFB0FFC81757FA01FC3FFA1D" box="[853,924,1414,1436]" name="Peru" pageId="1">PERU</collectingCountry>
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, 2006’’, and my red
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<typeStatus id="9B7E1895FFB0FFC81536FA26FE2DFA36" box="[308,398,1441,1463]" pageId="1" type="holotype">holotype</typeStatus>
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label
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</materialsCitation>
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.
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<materialsCitation id="F4ADAC6AFFB0FFC815DBFA26FE74FA53" collectorName="Lincoln, NE" country="Peru" location="University of Nebraska State Museum" pageId="1" specimenCount="1" stateProvince="Junin" typeStatus="holotype">
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<typeStatus id="9B7E1895FFB0FFC815DBFA26FD9AFA36" box="[473,569,1441,1463]" pageId="1" type="holotype">Holotype</typeStatus>
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deposited at the
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<location id="411AF0ECFFB0FFC816F4FA26FEE4FA53" LSID="urn:lsid:plazi:treatment:CC6C1721FFB0FFCC1569FA8AFC19FD6A:411AF0ECFFB0FFC816F4FA26FEE4FA53" country="Peru" name="University of Nebraska State Museum" pageId="1" stateProvince="Junin">University of Nebraska State Museum</location>
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,
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<collectorName id="E930C3E1FFB0FFC81551FA3BFE74FA53" box="[339,471,1468,1490]" pageId="1">Lincoln, NE</collectorName>
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</materialsCitation>
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.
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</paragraph>
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||
<paragraph id="447AA637FFB0FFC814C5FA50FC4FF9A3" blockId="1.[175,1005,1361,1570]" pageId="1">
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<emphasis id="76B17A25FFB0FFC814C5FA50FE89FA6C" bold="true" box="[199,298,1495,1517]" pageId="1">
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||
<typeStatus id="9B7E1895FFB0FFC814C5FA50FE85FA6C" box="[199,294,1495,1517]" pageId="1" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Length 34.5 mm; width across humeri 14.9 mm. Color of dorsum piceous, weakly shining; venter reddish brown.
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<emphasis id="76B17A25FFB0FFC81699FA75FD72F986" box="[667,721,1522,1543]" italics="true" pageId="1">Head</emphasis>
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: Dorsal surface completely occupied by stout, recurved horn (
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<figureCitation id="DCFEBAB2FFB0FFC81617F98BFDF7F9A3" box="[533,596,1548,1570]" captionStart="Figs" captionStartId="2.[167,206,1044,1063]" captionTargetBox="[273,849,198,1022]" captionTargetId="figure-196@2.[272,850,194,1022]" captionTargetPageId="2" captionText="Figs. 1–2. Megaceras saltini, holotype, dorsal and lateral views, respectively. Photos courtesy of J. Saltin." figureDoi="http://doi.org/10.5281/zenodo.10129326" httpUri="https://zenodo.org/record/10129326/files/figure.png" pageId="1">Fig. 2</figureCitation>
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); horn expanded at base and extending
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</paragraph>
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<paragraph id="447AA637FFB0FFC8163CF9B9FDFDF9D0" blockId="1.[574,606,1598,1617]" box="[574,606,1598,1617]" pageId="1">463</paragraph>
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<paragraph id="447AA637FFB3FFCD1490FFF9FE52FC71" blockId="2.[146,178,126,145]" lastBlockId="4.[146,976,773,1570]" lastPageId="4" pageId="2">
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464 from clypeal apex to occiput and from eye canthus to eye canthus; surface of horn minutely alutaceous, sides with sparse, moderately large punctures; anterior face with small, sparse punctures; posterior margin just below apex with small swelling (suggestive of a subapical tooth). Clypeus with apex broadly rounded, feebly emarginate at center. Interocular width equals 4.9 transverse eye diameters. Antenna with 10 segments, club subequal in length to segments 2–7. Mandible with large, angulate basal lobe and narrow, acute apical tooth (
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<figureCitation id="DCFEBAB2FFB3FFCB1711FA9EFCF0FAAE" box="[787,851,1305,1327]" captionStart="Figs" captionStartId="3.[196,235,1373,1392]" captionTargetBox="[184,996,197,1348]" captionTargetId="figure-1@3.[181,998,194,1350]" captionTargetPageId="3" captionText="Figs. 3–10. Dorsal view of head of (3) M. saltini and (4) M. morpheus showing form of mandibles. Lateral view of pygidium of (5) M. saltini and (6) M. morpheus showing surface convexity. Parameres (caudal and lateral views) of (7–8) M. saltini and (9–10) M. morpheus." figureDoi="http://doi.org/10.5281/zenodo.10129328" httpUri="https://zenodo.org/record/10129328/files/figure.png" pageId="2">Fig. 3</figureCitation>
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).
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<emphasis id="76B17A25FFB3FFCB1767FA9EFC6BFAAF" box="[869,968,1305,1326]" italics="true" pageId="2">Pronotum</emphasis>
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: Surface strongly aciculate, minutely shagreened, punctate in anterior angle and on lateral margin; punctures in anterior angle mostly large, extending obliquely onto disc, those along lateral margin large, becoming confluent and rugose anteriorly. Base with very slender marginal bead. Lateral margins arcuate, widest just before middle. Disc at center with raised, bituberculate prominence, tubercles separated from one another by distance equal to transverse eye diameter.
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<emphasis id="76B17A25FFB3FFCB178BFA3CFC6BFA51" box="[905,968,1467,1488]" italics="true" pageId="2">Elytra</emphasis>
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: Surface strongly aciculate, minutely shagreened, rugopunctate at apices. Sutural stria strongly impressed; disc at center and just mesad of humerus with weakly impressed stria. Sides vaguely wrinkled. Lateral margin with strong bead. 466
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<emphasis id="76B17A25FFB5FFCD1490FC82FF52FC9B" box="[146,241,773,794]" italics="true" pageId="4">Pygidium</emphasis>
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: Surface weakly aciculate, minutely alutaceous. Base with transverse row of large, moderately dense, setigerous punctures; setae long, reddish brown. In lateral view, surface strongly convex in basal fourth, nearly flat elsewhere (
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<figureCitation id="DCFEBAB2FFB5FFCD1498FCD2FF7AFCEA" box="[154,217,853,875]" captionStart="Figs" captionStartId="3.[196,235,1373,1392]" captionTargetBox="[184,996,197,1348]" captionTargetId="figure-1@3.[181,998,194,1350]" captionTargetPageId="3" captionText="Figs. 3–10. Dorsal view of head of (3) M. saltini and (4) M. morpheus showing form of mandibles. Lateral view of pygidium of (5) M. saltini and (6) M. morpheus showing surface convexity. Parameres (caudal and lateral views) of (7–8) M. saltini and (9–10) M. morpheus." figureDoi="http://doi.org/10.5281/zenodo.10129328" httpUri="https://zenodo.org/record/10129328/files/figure.png" pageId="4">Fig. 5</figureCitation>
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).
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<emphasis id="76B17A25FFB5FFCD14EFFCD2FEBEFCEB" box="[237,285,853,874]" italics="true" pageId="4">Legs</emphasis>
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: Protibia tridentate, teeth subequally spaced. Meso- and metatibia each with 2 transversely oblique carinae. Metatibia at apex with large, narrowly rounded lobe.
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<emphasis id="76B17A25FFB5FFCD1536FC0DFED4FC1E" box="[308,375,906,927]" italics="true" pageId="4">Venter</emphasis>
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: Prosternal process long, subconical. Pro-, meso-, and metasternum with long, reddish brown setae. Metasternum either side of middle nearly completely punctate; punctures dense, small; metasternum at center impunctate.
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<emphasis id="76B17A25FFB5FFCD1511FC5DFEDDFC6E" box="[275,382,986,1007]" italics="true" pageId="4">Parameres</emphasis>
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:
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<figureCitation id="DCFEBAB2FFB5FFCD158EFC5DFE4DFC71" box="[396,494,986,1008]" captionStart="Figs" captionStartId="3.[196,235,1373,1392]" captionTargetBox="[184,996,197,1348]" captionTargetId="figure-1@3.[181,998,194,1350]" captionTargetPageId="3" captionText="Figs. 3–10. Dorsal view of head of (3) M. saltini and (4) M. morpheus showing form of mandibles. Lateral view of pygidium of (5) M. saltini and (6) M. morpheus showing surface convexity. Parameres (caudal and lateral views) of (7–8) M. saltini and (9–10) M. morpheus." figureDoi="http://doi.org/10.5281/zenodo.10129328" httpUri="https://zenodo.org/record/10129328/files/figure.png" pageId="4">Figs. 7–8</figureCitation>
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.
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</paragraph>
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</subSubSection>
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||
<caption id="10BAF6BFFFB3FFCB14A5FB93FEF2FBBE" ID-DOI="http://doi.org/10.5281/zenodo.10129326" ID-Zenodo-Dep="10129326" httpUri="https://zenodo.org/record/10129326/files/figure.png" pageId="2" startId="2.[167,206,1044,1063]" targetBox="[273,849,198,1022]" targetPageId="2">
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||
<paragraph id="447AA637FFB3FFCB14A5FB93FEF2FBBE" blockId="2.[146,975,1044,1087]" pageId="2">
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||
<emphasis id="76B17A25FFB3FFCB14A5FB93FEA1FBA6" bold="true" box="[167,258,1044,1063]" pageId="2">Figs. 1–2.</emphasis>
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<taxonomicName id="83C5DDB4FFB3FFCB151AFB92FE1BFBA9" box="[280,440,1045,1064]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="saltini">
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<emphasis id="76B17A25FFB3FFCB151AFB92FE1BFBA9" box="[280,440,1045,1064]" italics="true" pageId="2">Megaceras saltini</emphasis>
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</taxonomicName>
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, holotype, dorsal and lateral views, respectively. Photos courtesy of J. Saltin.
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</paragraph>
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||
</caption>
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||
<caption id="10BAF6BFFFB2FFCA14C6FADAFC4FFA21" ID-DOI="http://doi.org/10.5281/zenodo.10129328" ID-Zenodo-Dep="10129328" httpUri="https://zenodo.org/record/10129328/files/figure.png" pageId="3" startId="3.[196,235,1373,1392]" targetBox="[184,996,197,1348]" targetPageId="3">
|
||
<paragraph id="447AA637FFB2FFCA14C6FADAFC4FFA21" blockId="3.[175,1004,1373,1440]" pageId="3">
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||
<emphasis id="76B17A25FFB2FFCA14C6FADAFE89FAF1" bold="true" box="[196,298,1373,1392]" pageId="3">Figs. 3–10.</emphasis>
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Dorsal view of head of (
|
||
<emphasis id="76B17A25FFB2FFCA161CFADAFD8AFAF1" bold="true" box="[542,553,1373,1392]" pageId="3">3</emphasis>
|
||
)
|
||
<taxonomicName id="83C5DDB4FFB2FFCA1634FADAFD2EFAF1" box="[566,653,1373,1392]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="saltini">
|
||
<emphasis id="76B17A25FFB2FFCA1634FADAFD2EFAF1" box="[566,653,1373,1392]" italics="true" pageId="3">M. saltini</emphasis>
|
||
</taxonomicName>
|
||
and (
|
||
<emphasis id="76B17A25FFB2FFCA16C0FADAFD6EFAF1" bold="true" box="[706,717,1373,1392]" pageId="3">4</emphasis>
|
||
)
|
||
<taxonomicName id="83C5DDB4FFB2FFCA16D8FADAFCF3FAF1" authorityName="Burmeister" authorityYear="1847" box="[730,848,1373,1392]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="morpheus">
|
||
<emphasis id="76B17A25FFB2FFCA16D8FADAFCF3FAF1" box="[730,848,1373,1392]" italics="true" pageId="3">M. morpheus</emphasis>
|
||
</taxonomicName>
|
||
showing form of mandibles. Lateral view of pygidium of (
|
||
<emphasis id="76B17A25FFB2FFCA1621FAF2FD8DFA09" bold="true" box="[547,558,1397,1416]" pageId="3">5</emphasis>
|
||
)
|
||
<taxonomicName id="83C5DDB4FFB2FFCA1639FAF2FD31FA09" box="[571,658,1397,1416]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="saltini">
|
||
<emphasis id="76B17A25FFB2FFCA1639FAF2FD31FA09" box="[571,658,1397,1416]" italics="true" pageId="3">M. saltini</emphasis>
|
||
</taxonomicName>
|
||
and (
|
||
<emphasis id="76B17A25FFB2FFCA16C5FAF2FD71FA09" bold="true" box="[711,722,1397,1416]" pageId="3">6</emphasis>
|
||
)
|
||
<taxonomicName id="83C5DDB4FFB2FFCA16DDFAF2FCF7FA09" authorityName="Burmeister" authorityYear="1847" box="[735,852,1397,1416]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="morpheus">
|
||
<emphasis id="76B17A25FFB2FFCA16DDFAF2FCF7FA09" box="[735,852,1397,1416]" italics="true" pageId="3">M. morpheus</emphasis>
|
||
</taxonomicName>
|
||
showing surface convexity. Parameres (caudal and lateral views) of (
|
||
<emphasis id="76B17A25FFB2FFCA1681FA0AFD01FA21" bold="true" box="[643,674,1421,1440]" pageId="3">7–8</emphasis>
|
||
)
|
||
<taxonomicName id="83C5DDB4FFB2FFCA16ADFA0AFCA5FA21" box="[687,774,1421,1440]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="saltini">
|
||
<emphasis id="76B17A25FFB2FFCA16ADFA0AFCA5FA21" box="[687,774,1421,1440]" italics="true" pageId="3">M. saltini</emphasis>
|
||
</taxonomicName>
|
||
and (
|
||
<emphasis id="76B17A25FFB2FFCA1738FA0AFCC6FA21" bold="true" box="[826,869,1421,1440]" pageId="3">9–10</emphasis>
|
||
)
|
||
<taxonomicName id="83C5DDB4FFB2FFCA1770FA0AFC45FA21" authorityName="Burmeister" authorityYear="1847" box="[882,998,1421,1440]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="morpheus">
|
||
<emphasis id="76B17A25FFB2FFCA1770FA0AFC45FA21" box="[882,998,1421,1440]" italics="true" pageId="3">M. morpheus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="10BAF6BFFFB5FFCD14A5FD1CFD8BFD5F" ID-DOI="http://doi.org/10.5281/zenodo.10129330" ID-Zenodo-Dep="10129330" httpUri="https://zenodo.org/record/10129330/files/figure.png" pageId="4" startId="4.[167,198,667,686]" targetBox="[146,976,195,641]" targetPageId="4">
|
||
<paragraph id="447AA637FFB5FFCD14A5FD1CFD8BFD5F" blockId="4.[146,975,667,734]" pageId="4">
|
||
<emphasis id="76B17A25FFB5FFCD14A5FD1CFF4CFD2F" bold="true" box="[167,239,667,686]" pageId="4">Fig. 11.</emphasis>
|
||
Lateral view of head of Dim and
|
||
<taxonomicName id="83C5DDB4FFB5FFCD1661FD1BFD63FD2E" box="[611,704,668,687]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="saltini">
|
||
<emphasis id="76B17A25FFB5FFCD1661FD1BFD63FD2E" box="[611,704,668,687]" italics="true" pageId="4">M. saltini</emphasis>
|
||
</taxonomicName>
|
||
showing similarity in horn configuration (‘‘the Dim Effect’’). Dim character º Disney Enterprises, Inc. and Pixar. Used by permission from Disney Enterprises, Inc.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="0CDFF5BCFFB5FFCD14A8FC72FE35FBDA" pageId="4" type="etymology">
|
||
<paragraph id="447AA637FFB5FFCD14A8FC72FE35FBDA" blockId="4.[146,976,773,1570]" pageId="4">
|
||
<emphasis id="76B17A25FFB5FFCD14A8FC72FEBEFB8A" bold="true" box="[170,285,1013,1035]" pageId="4">Etymology.</emphasis>
|
||
At the request of Jochen-P. Saltin, who graciously donated the specimen for description, this species is named in honor of his son, Brian. Brian has sustained, without complaint, his father’s passion for beetles and is now himself studying biology.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0CDFF5BCFFB5FFCD14A8FBE7FF6AFB47" pageId="4" type="distribution">
|
||
<paragraph id="447AA637FFB5FFCD14A8FBE7FF6AFB47" blockId="4.[146,976,773,1570]" pageId="4">
|
||
<emphasis id="76B17A25FFB5FFCD14A8FBE7FE8BFBF7" bold="true" box="[170,296,1120,1142]" pageId="4">Distribution.</emphasis>
|
||
<taxonomicName id="83C5DDB4FFB5FFCD1533FBE6FDB5FBF7" authorityName="Ratcliffe" authorityYear="2007" box="[305,534,1121,1142]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="briansaltini">
|
||
<emphasis id="76B17A25FFB5FFCD1533FBE6FDB5FBF7" box="[305,534,1121,1142]" italics="true" pageId="4">Megaceras briansaltini</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<typeStatus id="9B7E1895FFB5FFCD1719FBE7FCE4FBF7" box="[795,839,1120,1142]" pageId="4">type</typeStatus>
|
||
locality near Calabaza (on the road from Satipo to Huancayo), District of Pampa in the Department of
|
||
<collectingRegion id="860168D5FFB5FFCD1537FB12FED3FB2A" box="[309,368,1173,1195]" country="Peru" name="Junin" pageId="4">Junin</collectingRegion>
|
||
, Province of Satipo on the eastern slopes of the Andes in
|
||
<collectingCountry id="3CD2E6A7FFB5FFCD1490FB37FF66FB47" box="[146,197,1200,1222]" name="Peru" pageId="4">Peru</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0CDFF5BCFFB5FFCC14A8FB4BFC19FD6A" lastPageId="5" pageId="4" type="diagnosis">
|
||
<paragraph id="447AA637FFB5FFCD14A8FB4BFDD5FAB0" blockId="4.[146,976,773,1570]" pageId="4">
|
||
<emphasis id="76B17A25FFB5FFCD14A8FB4BFEB1FB63" bold="true" box="[170,274,1228,1250]" pageId="4">Diagnosis.</emphasis>
|
||
<taxonomicName id="83C5DDB4FFB5FFCD151EFB4BFDA0FB60" authorityName="Ratcliffe" authorityYear="2007" box="[284,515,1228,1249]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="briansaltini">
|
||
<emphasis id="76B17A25FFB5FFCD151EFB4BFDA0FB60" box="[284,515,1228,1249]" italics="true" pageId="4">Megaceras briansaltini</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="83C5DDB4FFB5FFCD16CDFB4BFC6CFB63" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[719,975,1228,1250]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="morpheus">
|
||
<emphasis id="76B17A25FFB5FFCD16CDFB4BFCF6FB60" box="[719,853,1228,1249]" italics="true" pageId="4">M. morpheus</emphasis>
|
||
Burmeister
|
||
</taxonomicName>
|
||
and will key to this species in Endrödi (1976, 1985).
|
||
<taxonomicName id="83C5DDB4FFB5FFCD16A6FB61FC2BFB7A" authorityName="Ratcliffe" authorityYear="2007" box="[676,904,1254,1275]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="briansaltini">
|
||
<emphasis id="76B17A25FFB5FFCD16A6FB61FC2BFB7A" box="[676,904,1254,1275]" italics="true" pageId="4">Megaceras briansaltini</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="83C5DDB4FFB5FFCD14C9FA86FEECFA97" authorityName="Burmeister" authorityYear="1847" box="[203,335,1281,1302]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="morpheus">
|
||
<emphasis id="76B17A25FFB5FFCD14C9FA86FEECFA97" box="[203,335,1281,1302]" italics="true" pageId="4">M. morpheus</emphasis>
|
||
</taxonomicName>
|
||
in the form of the parameres, head horn, dorsal surface of the clypeus, teeth of the mandibles, and pygidium.
|
||
</paragraph>
|
||
<paragraph id="447AA637FFB5FFCD14A8FAB0FD44FA1D" blockId="4.[146,976,773,1570]" pageId="4">
|
||
The parameres, in caudal view, of
|
||
<taxonomicName id="83C5DDB4FFB5FFCD1624FAB0FD63FACD" authorityName="Ratcliffe" authorityYear="2007" box="[550,704,1335,1356]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="briansaltini">
|
||
<emphasis id="76B17A25FFB5FFCD1624FAB0FD63FACD" box="[550,704,1335,1356]" italics="true" pageId="4">M. briansaltini</emphasis>
|
||
</taxonomicName>
|
||
are subquadrate at their apices (
|
||
<figureCitation id="DCFEBAB2FFB5FFCD14E2FAD6FE83FAE6" box="[224,288,1361,1383]" captionStart="Figs" captionStartId="3.[196,235,1373,1392]" captionTargetBox="[184,996,197,1348]" captionTargetId="figure-1@3.[181,998,194,1350]" captionTargetPageId="3" captionText="Figs. 3–10. Dorsal view of head of (3) M. saltini and (4) M. morpheus showing form of mandibles. Lateral view of pygidium of (5) M. saltini and (6) M. morpheus showing surface convexity. Parameres (caudal and lateral views) of (7–8) M. saltini and (9–10) M. morpheus." figureDoi="http://doi.org/10.5281/zenodo.10129328" httpUri="https://zenodo.org/record/10129328/files/figure.png" pageId="4">Fig. 7</figureCitation>
|
||
), whereas they are narrowly rounded in
|
||
<taxonomicName id="83C5DDB4FFB5FFCD16CFFAD5FCF0FAE6" authorityName="Burmeister" authorityYear="1847" box="[717,851,1362,1383]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="morpheus">
|
||
<emphasis id="76B17A25FFB5FFCD16CFFAD5FCF0FAE6" box="[717,851,1362,1383]" italics="true" pageId="4">M. morpheus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="DCFEBAB2FFB5FFCD1766FAD6FC00FAE6" box="[868,931,1361,1383]" captionStart="Figs" captionStartId="3.[196,235,1373,1392]" captionTargetBox="[184,996,197,1348]" captionTargetId="figure-1@3.[181,998,194,1350]" captionTargetPageId="3" captionText="Figs. 3–10. Dorsal view of head of (3) M. saltini and (4) M. morpheus showing form of mandibles. Lateral view of pygidium of (5) M. saltini and (6) M. morpheus showing surface convexity. Parameres (caudal and lateral views) of (7–8) M. saltini and (9–10) M. morpheus." figureDoi="http://doi.org/10.5281/zenodo.10129328" httpUri="https://zenodo.org/record/10129328/files/figure.png" pageId="4">Fig. 9</figureCitation>
|
||
). In lateral view, the subapical, lateroventral depression of the parameres is wide in
|
||
<taxonomicName id="83C5DDB4FFB5FFCD17B3FAEBFEA1FA1D" authorityName="Ratcliffe" authorityYear="2007" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="briansaltini">
|
||
<emphasis id="76B17A25FFB5FFCD17B3FAEBFEA1FA1D" italics="true" pageId="4">M. briansaltini</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="DCFEBAB2FFB5FFCD1510FA01FEECFA1D" box="[274,335,1414,1436]" captionStart="Figs" captionStartId="3.[196,235,1373,1392]" captionTargetBox="[184,996,197,1348]" captionTargetId="figure-1@3.[181,998,194,1350]" captionTargetPageId="3" captionText="Figs. 3–10. Dorsal view of head of (3) M. saltini and (4) M. morpheus showing form of mandibles. Lateral view of pygidium of (5) M. saltini and (6) M. morpheus showing surface convexity. Parameres (caudal and lateral views) of (7–8) M. saltini and (9–10) M. morpheus." figureDoi="http://doi.org/10.5281/zenodo.10129328" httpUri="https://zenodo.org/record/10129328/files/figure.png" pageId="4">Fig. 8</figureCitation>
|
||
) and narrow in
|
||
<taxonomicName id="83C5DDB4FFB5FFCD15F9FA00FD23FA1D" authorityName="Burmeister" authorityYear="1847" box="[507,640,1415,1436]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="morpheus">
|
||
<emphasis id="76B17A25FFB5FFCD15F9FA00FD23FA1D" box="[507,640,1415,1436]" italics="true" pageId="4">M. morpheus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="DCFEBAB2FFB5FFCD168DFA01FD7FFA1D" box="[655,732,1414,1436]" captionStart="Figs" captionStartId="3.[196,235,1373,1392]" captionTargetBox="[184,996,197,1348]" captionTargetId="figure-1@3.[181,998,194,1350]" captionTargetPageId="3" captionText="Figs. 3–10. Dorsal view of head of (3) M. saltini and (4) M. morpheus showing form of mandibles. Lateral view of pygidium of (5) M. saltini and (6) M. morpheus showing surface convexity. Parameres (caudal and lateral views) of (7–8) M. saltini and (9–10) M. morpheus." figureDoi="http://doi.org/10.5281/zenodo.10129328" httpUri="https://zenodo.org/record/10129328/files/figure.png" pageId="4">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="447AA637FFB5FFCC14A8FA25FC4EFF73" blockId="4.[146,976,773,1570]" lastBlockId="5.[175,1005,193,747]" lastPageId="5" pageId="4">
|
||
The form of the head horn in
|
||
<taxonomicName id="83C5DDB4FFB5FFCD15E2FA25FDD5FA36" authorityName="Ratcliffe" authorityYear="2007" box="[480,630,1442,1463]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="briansaltini">
|
||
<emphasis id="76B17A25FFB5FFCD15E2FA25FDD5FA36" box="[480,630,1442,1463]" italics="true" pageId="4">M. briansaltini</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="DCFEBAB2FFB5FFCD1687FA25FD6EFA39" box="[645,717,1442,1464]" captionStart="Figs" captionStartId="2.[167,206,1044,1063]" captionTargetBox="[273,849,198,1022]" captionTargetId="figure-196@2.[272,850,194,1022]" captionTargetPageId="2" captionText="Figs. 1–2. Megaceras saltini, holotype, dorsal and lateral views, respectively. Photos courtesy of J. Saltin." figureDoi="http://doi.org/10.5281/zenodo.10129326" httpUri="https://zenodo.org/record/10129326/files/figure.png" pageId="4">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="DCFEBAB2FFB5FFCD16DBFA25FD52FA39" box="[729,753,1442,1464]" captionStart="Fig" captionStartId="4.[167,198,667,686]" captionTargetBox="[146,976,195,641]" captionTargetId="figure-360@4.[146,976,194,645]" captionTargetPageId="4" captionText="Fig. 11. Lateral view of head of Dim and M. saltini showing similarity in horn configuration (‘‘the Dim Effect’’). Dim character º Disney Enterprises, Inc. and Pixar. Used by permission from Disney Enterprises, Inc." figureDoi="http://doi.org/10.5281/zenodo.10129330" httpUri="https://zenodo.org/record/10129330/files/figure.png" pageId="4">11</figureCitation>
|
||
) is unlike that of any other dynastine species with which I am familiar, because it is so swollen at its base... so much so that it obscures or encompasses most of the top of the head. The clypeus is not visible except at its extreme apex. In
|
||
<taxonomicName id="83C5DDB4FFB5FFCD16CEFA75FCF3F986" authorityName="Burmeister" authorityYear="1847" box="[716,848,1522,1543]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="morpheus">
|
||
<emphasis id="76B17A25FFB5FFCD16CEFA75FCF3F986" box="[716,848,1522,1543]" italics="true" pageId="4">M. morpheus</emphasis>
|
||
</taxonomicName>
|
||
, conversely, the dorsal surface of the clypeus is clearly evident anterior to the base of the horn. Because of the unusual nature of the horn, the possibility remains that the horn configuration is a monstrosity, but additional material is needed to ascertain this.
|
||
</paragraph>
|
||
<paragraph id="447AA637FFB4FFCC14C5FF70FE82FD97" blockId="5.[175,1005,193,747]" pageId="5">
|
||
The form of the horn is startlingly similar to that of Dim, the blue rhinoceros beetle in the Disney/Pixar animated motion picture,
|
||
<emphasis id="76B17A25FFB4FFCC16E4FE96FCCDFEA7" box="[742,878,273,294]" italics="true" pageId="5">A Bug’s Life</emphasis>
|
||
(
|
||
<figureCitation id="DCFEBAB2FFB4FFCC1782FE96FC68FEA6" box="[896,971,273,295]" captionStart="Fig" captionStartId="4.[167,198,667,686]" captionTargetBox="[146,976,195,641]" captionTargetId="figure-360@4.[146,976,194,645]" captionTargetPageId="4" captionText="Fig. 11. Lateral view of head of Dim and M. saltini showing similarity in horn configuration (‘‘the Dim Effect’’). Dim character º Disney Enterprises, Inc. and Pixar. Used by permission from Disney Enterprises, Inc." figureDoi="http://doi.org/10.5281/zenodo.10129330" httpUri="https://zenodo.org/record/10129330/files/figure.png" pageId="5">Fig. 11</figureCitation>
|
||
). I know of no dynastine head horn that has ever had the shape of the one seen in
|
||
<taxonomicName id="83C5DDB4FFB4FFCC17CCFEABFEBCFEDD" authorityName="Ratcliffe" authorityYear="2007" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="briansaltini">
|
||
<emphasis id="76B17A25FFB4FFCC17CCFEABFEBCFEDD" italics="true" pageId="5">M. briansaltini</emphasis>
|
||
</taxonomicName>
|
||
, and so its resemblance to a movie character seems like a case of nature mimicking art... or what could be referred to as ‘‘the Dim Effect.’’ There are numerous examples of art mimicking nature (paintings, sculpture, etc.), but that cannot be the case here, because there had never been a known rhinoceros beetle in nature upon which the creators of Dim could have used as a model for the head horn. In my experience, then, Dim was the first ‘‘rhinoceros beetle’’ to display such a horn, and the discovery of
|
||
<taxonomicName id="83C5DDB4FFB4FFCC165AFE61FD4FFE7A" authorityName="Ratcliffe" authorityYear="2007" box="[600,748,486,507]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="briansaltini">
|
||
<emphasis id="76B17A25FFB4FFCC165AFE61FD4FFE7A" box="[600,748,486,507]" italics="true" pageId="5">M. briansaltini</emphasis>
|
||
</taxonomicName>
|
||
, a real rhinoceros beetle, came later.
|
||
</paragraph>
|
||
<paragraph id="447AA637FFB4FFCC14C5FD9CFD29FD01" blockId="5.[175,1005,193,747]" pageId="5">
|
||
The teeth of the mandibles differ between
|
||
<taxonomicName id="83C5DDB4FFB4FFCC1678FD9CFCACFDB1" authorityName="Ratcliffe" authorityYear="2007" box="[634,783,539,560]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="briansaltini">
|
||
<emphasis id="76B17A25FFB4FFCC1678FD9CFCACFDB1" box="[634,783,539,560]" italics="true" pageId="5">M. briansaltini</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="83C5DDB4FFB4FFCC1746FD9CFC6BFDB1" authorityName="Burmeister" authorityYear="1847" box="[836,968,539,560]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="morpheus">
|
||
<emphasis id="76B17A25FFB4FFCC1746FD9CFC6BFDB1" box="[836,968,539,560]" italics="true" pageId="5">M. morpheus</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="83C5DDB4FFB4FFCC14ADFDB1FEE7FDCA" authorityName="Ratcliffe" authorityYear="2007" box="[175,324,566,587]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="briansaltini">
|
||
<emphasis id="76B17A25FFB4FFCC14ADFDB1FEE7FDCA" box="[175,324,566,587]" italics="true" pageId="5">M. briansaltini</emphasis>
|
||
</taxonomicName>
|
||
, the basal lobe of the mandible is large, obtusely rounded, weakly bilobed, while the apical tooth is narrow and acute (
|
||
<figureCitation id="DCFEBAB2FFB4FFCC16C8FDD7FCAAFDE7" box="[714,777,592,614]" captionStart="Figs" captionStartId="3.[196,235,1373,1392]" captionTargetBox="[184,996,197,1348]" captionTargetId="figure-1@3.[181,998,194,1350]" captionTargetPageId="3" captionText="Figs. 3–10. Dorsal view of head of (3) M. saltini and (4) M. morpheus showing form of mandibles. Lateral view of pygidium of (5) M. saltini and (6) M. morpheus showing surface convexity. Parameres (caudal and lateral views) of (7–8) M. saltini and (9–10) M. morpheus." figureDoi="http://doi.org/10.5281/zenodo.10129328" httpUri="https://zenodo.org/record/10129328/files/figure.png" pageId="5">Fig. 3</figureCitation>
|
||
). In
|
||
<taxonomicName id="83C5DDB4FFB4FFCC1738FDD7FC1CFDE4" authorityName="Burmeister" authorityYear="1847" box="[826,959,592,613]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="morpheus">
|
||
<emphasis id="76B17A25FFB4FFCC1738FDD7FC1CFDE4" box="[826,959,592,613]" italics="true" pageId="5">M. morpheus</emphasis>
|
||
</taxonomicName>
|
||
, the basal and apical teeth are both acute (
|
||
<figureCitation id="DCFEBAB2FFB4FFCC163CFDEDFDDDFD01" box="[574,638,618,640]" captionStart="Figs" captionStartId="3.[196,235,1373,1392]" captionTargetBox="[184,996,197,1348]" captionTargetId="figure-1@3.[181,998,194,1350]" captionTargetPageId="3" captionText="Figs. 3–10. Dorsal view of head of (3) M. saltini and (4) M. morpheus showing form of mandibles. Lateral view of pygidium of (5) M. saltini and (6) M. morpheus showing surface convexity. Parameres (caudal and lateral views) of (7–8) M. saltini and (9–10) M. morpheus." figureDoi="http://doi.org/10.5281/zenodo.10129328" httpUri="https://zenodo.org/record/10129328/files/figure.png" pageId="5">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="447AA637FFB4FFCC14C5FD02FC19FD6A" blockId="5.[175,1005,193,747]" pageId="5">
|
||
Lastly, the form of the male pygidium differs between the species. In lateral view, the pygidium of
|
||
<taxonomicName id="83C5DDB4FFB4FFCC1599FD27FD91FD34" authorityName="Ratcliffe" authorityYear="2007" box="[411,562,672,693]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="briansaltini">
|
||
<emphasis id="76B17A25FFB4FFCC1599FD27FD91FD34" box="[411,562,672,693]" italics="true" pageId="5">M. briansaltini</emphasis>
|
||
</taxonomicName>
|
||
is strongly convex and protuberant in the basal fourth and nearly flat elsewhere (
|
||
<figureCitation id="DCFEBAB2FFB4FFCC1657FD3DFD36FD51" box="[597,661,698,720]" captionStart="Figs" captionStartId="3.[196,235,1373,1392]" captionTargetBox="[184,996,197,1348]" captionTargetId="figure-1@3.[181,998,194,1350]" captionTargetPageId="3" captionText="Figs. 3–10. Dorsal view of head of (3) M. saltini and (4) M. morpheus showing form of mandibles. Lateral view of pygidium of (5) M. saltini and (6) M. morpheus showing surface convexity. Parameres (caudal and lateral views) of (7–8) M. saltini and (9–10) M. morpheus." figureDoi="http://doi.org/10.5281/zenodo.10129328" httpUri="https://zenodo.org/record/10129328/files/figure.png" pageId="5">Fig. 5</figureCitation>
|
||
). In
|
||
<taxonomicName id="83C5DDB4FFB4FFCC16CFFD3DFCF6FD4E" authorityName="Burmeister" authorityYear="1847" box="[717,853,698,719]" class="Insecta" family="Dynastidae" genus="Megaceras" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="morpheus">
|
||
<emphasis id="76B17A25FFB4FFCC16CFFD3DFCF6FD4E" box="[717,853,698,719]" italics="true" pageId="5">M. morpheus</emphasis>
|
||
</taxonomicName>
|
||
, however, the pygidium is normally and evenly convex, not strongly protuberant (
|
||
<figureCitation id="DCFEBAB2FFB4FFCC176DFD52FC0EFD6A" box="[879,941,725,747]" captionStart="Figs" captionStartId="3.[196,235,1373,1392]" captionTargetBox="[184,996,197,1348]" captionTargetId="figure-1@3.[181,998,194,1350]" captionTargetPageId="3" captionText="Figs. 3–10. Dorsal view of head of (3) M. saltini and (4) M. morpheus showing form of mandibles. Lateral view of pygidium of (5) M. saltini and (6) M. morpheus showing surface convexity. Parameres (caudal and lateral views) of (7–8) M. saltini and (9–10) M. morpheus." figureDoi="http://doi.org/10.5281/zenodo.10129328" httpUri="https://zenodo.org/record/10129328/files/figure.png" pageId="5">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |