treatments-xml/data/CC/16/87/CC168784FFA9FFC902E8306AFE47FA30.xml

<document id="902CE8C60921FA788DB7B444C7A1EF54" ID-DOI="10.5281/zenodo.6392005" ID-ISSN="1942-1354" ID-Zenodo-Dep="6392005" ID-ZooBank="53EFD9C2-FF9C-4636-B86E-0717AE67EF4A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648505177194" checkinUser="felipe" docAuthor="Solís, Ángel &amp; Kohlmann, Bert" docDate="2022" docId="CC168784FFA9FFC902E8306AFE47FA30" docLanguage="en" docName="InsectaMundi.2022.0918.1-12.pdf" docOrigin="Insecta Mundi 2022 (918)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Dichotomius (Selenocopris) woodruffi Solis and Kohlmann 2022, new species" docType="treatment" docVersion="3" lastPageNumber="8" masterDocId="302FFFFCFFADFFC00228343AFFAEFFBF" masterDocTitle="Dichotomius woodruffi, a new Dichotomius species of the agenor group from Costa Rica and Nicaragua (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="12" masterPageNumber="1" pageNumber="3" updateTime="1699308889974" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="F62F10C2DC4245FCC360A8CFA7661F16" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="50F122D5600E077EC20C405EF38C4E32">
<mods:title id="E5FE84B8FE327B8BEDE7F79BE058ECA4">Dichotomius woodruffi, a new Dichotomius species of the agenor group from Costa Rica and Nicaragua (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="9EBA51E6A8C3B22FB94B1497F6F090B2" type="personal">
<mods:role id="C3B2658E0D5C94C886E2980C49B43F4F">
<mods:roleTerm id="A6739934DF0B374F122100BD816F8047">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6E2FC4133A2BEF9E548ED497A5817984">Solís, Ángel</mods:namePart>
<mods:affiliation id="C1A3B09FB56598BB8865E09CD588AC04">BioAlfa Barcoding Project Santo Domingo de Heredia, Costa Rica angelsoliscr @ gmail. com</mods:affiliation>
</mods:name>
<mods:name id="1712CEC60B877B3D9B8085C6C72DF42A" type="personal">
<mods:role id="0948E2BA69E88F421FAB366CBF0D1E9D">
<mods:roleTerm id="2EF65174577C2F9187C27CCEE7EBB9D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="97D8F534D8E9E0EC4B3380D65FA5CC15">Kohlmann, Bert</mods:namePart>
<mods:affiliation id="417195A9EE21839E5BD7083D0D94BC41">Finca Las Cusingas Buenos Aires de Jiménez, Costa Rica</mods:affiliation>
</mods:name>
<mods:typeOfResource id="8B643893C66720C13439598C5A510C20">text</mods:typeOfResource>
<mods:relatedItem id="F19F80F60FC287E2A6A53FBF2D0D78CF" type="host">
<mods:titleInfo id="18E33C4497371CF27FB7A97526F1B640">
<mods:title id="88650E070757899D013A03F2F1304177">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="4198E3F9F2FB808A347C7F6A96CDB48C">
<mods:date id="C5497D90823464141580DF4D4F379EC1">2022</mods:date>
<mods:detail id="FBAADB6607D26D82A2AD568DD32E2235" type="pubDate">
<mods:number id="E2F8DC127E00E4C7EED0352251B38000">2022-02-24</mods:number>
</mods:detail>
<mods:detail id="DED1A54968D814566DADD9315EE64AB9" type="volume">
<mods:number id="0069B96AE5F7E799D5646BADC3002EC1">2022</mods:number>
</mods:detail>
<mods:detail id="A632C2C6C0648E0519646D52E3F01EC9" type="issue">
<mods:number id="5FAA2F3F38660FEF35B9FE3EA0393A6D">918</mods:number>
</mods:detail>
<mods:extent id="185AB01FCF077D89BBEE17AAD608F3E7" unit="page">
<mods:start id="2B4B933BCD7143BD54CAA46D7DBDD121">1</mods:start>
<mods:end id="265FB3B870573DC1B4A98251334BA3B9">12</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B372721D981C4D94D232FB6B462E3AB6">journal article</mods:classification>
<mods:identifier id="5321BC729E75A0D489AC33F6EBDC6006" type="DOI">10.5281/zenodo.6392005</mods:identifier>
<mods:identifier id="5393B0A07A994A9039385E64FCF36AFF" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="B3B7CC59AD6186004CA344AAB987D078" type="Zenodo-Dep">6392005</mods:identifier>
<mods:identifier id="71C5EF3086D4C95021B3DAF46D47E040" type="ZooBank">53EFD9C2-FF9C-4636-B86E-0717AE67EF4A</mods:identifier>
</mods:mods>
<treatment id="CC168784FFA9FFC902E8306AFE47FA30" ID-DOI="http://doi.org/10.5281/zenodo.6392007" ID-Zenodo-Dep="6392007" LSID="urn:lsid:plazi:treatment:CC168784FFA9FFC902E8306AFE47FA30" httpUri="http://treatment.plazi.org/id/CC168784FFA9FFC902E8306AFE47FA30" lastPageId="9" lastPageNumber="8" pageId="4" pageNumber="3">
<subSubSection id="0CA56519FFA9FFC402E8306AFBE6FBD2" box="[192,1096,1104,1133]" pageId="4" pageNumber="3" type="nomenclature">
<paragraph id="44003692FFA9FFC402E8306AFBE6FBD2" blockId="4.[192,1096,1104,1166]" box="[192,1096,1104,1133]" pageId="4" pageNumber="3">
<heading id="1F4881FEFFA9FFC402E8306AFBE6FBD2" box="[192,1096,1104,1133]" fontSize="12" level="2" pageId="4" pageNumber="3" reason="2">
<emphasis id="76CBEA80FFA9FFC402E8306AFBE6FBD2" bold="true" box="[192,1096,1104,1133]" pageId="4" pageNumber="3">
<taxonomicName id="83BF4D11FFA9FFC402E8306AFC08FBD2" ID-CoL="8VR2L" authority="Solis and Kohlmann" authorityName="Solis and Kohlmann" authorityYear="2022" box="[192,934,1104,1133]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov." subGenus="Selenocopris">
<emphasis id="76CBEA80FFA9FFC402E8306AFECFFBD2" bold="true" box="[192,353,1104,1133]" italics="true" pageId="4" pageNumber="3">Dichotomius</emphasis>
(
<emphasis id="76CBEA80FFA9FFC4035A306AFDA1FBD2" bold="true" box="[370,527,1104,1133]" italics="true" pageId="4" pageNumber="3">Selenocopris</emphasis>
) 
<emphasis id="76CBEA80FFA9FFC40008306AFD34FBD2" bold="true" box="[544,666,1104,1133]" italics="true" pageId="4" pageNumber="3">woodruffi</emphasis>
Solís and Kohlmann
</taxonomicName>
, 
<taxonomicNameLabel id="6DF857FBFFA9FFC4019B306BFBE6FBD2" box="[947,1096,1104,1133]" pageId="4" pageNumber="3" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0CA56519FFA9FFC402E8304EFECCFB31" box="[192,354,1140,1166]" pageId="4" pageNumber="3" type="description">
<paragraph id="44003692FFA9FFC402E8304EFECCFB31" blockId="4.[192,1096,1104,1166]" box="[192,354,1140,1166]" pageId="4" pageNumber="3">
(
<figureCitation id="DC842A17FFA9FFC402E0304EFE9CFB31" box="[200,306,1140,1166]" captionStart-0="Figures 1–6" captionStart-1="Figures 7–12" captionStartId-0="6.[193,278,1833,1859]" captionStartId-1="8.[192,279,1799,1825]" captionTargetBox-0="[229,1405,221,1803]" captionTargetBox-1="[248,1385,213,1769]" captionTargetId-0="figure-18@6.[229,1405,199,1806]" captionTargetId-1="figure-84@8.[248,1385,213,1769]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionText-0="Figures 1–6. Dichotomius woodruffi, new species. 1) Dorsal habitus, holotype, male. Scale bar equals 1 cm. 2) Dorsal habitus, allotype, female. Scale bar equals 1 cm. 3) Frontal view, holotype, male. 4) Frontal view, allotype, female. 5) Ventral clypeal process spiniform, holotype male. 6) Anterior pronotal border (frontal view), holotype male." captionText-1="Figures 7–12. Dichotomius woodruffi, new species, and D. centralis. 7–11) D. woodruffi. 7) Anterior pronotal sulcus (lateral-oblique view), holotype male. 8) Male pygidium twice as broad as long, slightly convex, holotype male. 9) Lateral view of the aedeagus, holotype male. 10) Ventral view of the aedeagus, holotype male. 11) Lamella copulatrix, holotype male. 12) D. centralis, elytral stria crenulate." figureDoi-0="http://doi.org/10.5281/zenodo.6392009" figureDoi-1="http://doi.org/10.5281/zenodo.6392011" httpUri-0="https://zenodo.org/record/6392009/files/figure.png" httpUri-1="https://zenodo.org/record/6392011/files/figure.png" pageId="4" pageNumber="3">Fig. 1–11</figureCitation>
, 
<figureCitation id="DC842A17FFA9FFC40316304EFEF4FB31" box="[318,346,1140,1166]" captionStart="Figure 19" captionStartId="11.[192,268,1866,1892]" captionTargetBox="[192,1440,1091,1837]" captionTargetId="figure-407@11.[192,1440,1091,1837]" captionTargetPageId="11" captionText="Figure 19. Known distribution of Dichotomius agenor, D. centralis, and D. woodruffi new species in Nicaragua, Costa Rica, and Panama." figureDoi="http://doi.org/10.5281/zenodo.6392015" httpUri="https://zenodo.org/record/6392015/files/figure.png" pageId="4" pageNumber="3">19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="0CA56519FFA9FFC402C830A7FC6DFB4D" pageId="4" pageNumber="3" type="reference_group">
<paragraph id="44003692FFA9FFC402C830A7FEBCFB6C" blockId="4.[224,1441,1181,1266]" pageId="4" pageNumber="3">
<treatmentCitationGroup id="64AF11BCFFA9FFC402C830A7FEBCFB6C" pageId="4" pageNumber="3">
<taxonomicName id="83BF4D11FFA9FFC402C830A7FDEEFB0A" ID-CoL="6CS36" authority="(Harold, 1869)" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[224,576,1181,1205]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="agenor">
<emphasis id="76CBEA80FFA9FFC402C830A7FE08FB0A" box="[224,422,1181,1205]" italics="true" pageId="4" pageNumber="3">Dichotomius agenor</emphasis>
(Harold, 1869)
</taxonomicName>
– 
<treatmentCitation id="C51E1083FFA9FFC4007F30A7FC24FB0A" author="Kohlmann B &amp; Solis A." box="[599,906,1181,1205]" page="345" pageId="4" pageNumber="3" year="1997">
<bibRefCitation id="202E4B63FFA9FFC4007F30A7FC24FB0A" author="Kohlmann B &amp; Solis A." box="[599,906,1181,1205]" pageId="4" pageNumber="3" pagination="343 - 382" refId="ref8326" refString="Kohlmann B, Solis A. 1997. El genero Dichotomius (Coleoptera: Scarabaeidae) en Costa Rica. Giornale Italiano di Entomologia 8: 343 - 382." type="journal article" year="1997">Kohlmann and Solís 1997: 345</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="C51E1083FFA9FFC401BB30A7FB21FB0A" author="Kohlmann B &amp; Solis A &amp; Elle O &amp; Soto X &amp; Russo R." box="[915,1167,1181,1205]" page="30" pageId="4" pageNumber="3" year="2007">
<bibRefCitation id="202E4B63FFA9FFC401BB30A7FB21FB0A" author="Kohlmann B &amp; Solis A &amp; Elle O &amp; Soto X &amp; Russo R." box="[915,1167,1181,1205]" pageId="4" pageNumber="3" pagination="1 - 34" refId="ref8356" refString="Kohlmann B, Solis A, Elle O, Soto X, Russo R. 2007. Biodiversity, conservation, and hotspot atlas of Costa Rica: a dung beetle perspective (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 1457: 1 - 34." type="journal article" year="2007">Kohlmann et al. 2007: 30</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="C51E1083FFA9FFC406B030A7FEA3FB6C" author="Solis A &amp; Kohlmann B." page="6" pageId="4" pageNumber="3" year="2012">
<bibRefCitation id="202E4B63FFA9FFC406B030A7FEA3FB6C" author="Solis A &amp; Kohlmann B." pageId="4" pageNumber="3" pagination="1 - 32" refId="ref8704" refString="Solis A, Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae (Coleoptera: Scarabaeidae) of Costa Rica. Zootaxa 3482: 1 - 32." type="journal article" year="2012">Solís and Kohlmann 2012: 6</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="44003692FFA9FFC402C830E0FC6DFB4D" blockId="4.[224,1441,1181,1266]" box="[224,963,1242,1266]" pageId="4" pageNumber="3">
<taxonomicName id="83BF4D11FFA9FFC402C830E0FC90FB4D" ID-CoL="7VQMQ" authority="Montoya-Molina and Vaz-de-Mello, 2021" authorityName="Montoya-Molina and Vaz-de-Mello" authorityYear="2021" box="[224,830,1242,1266]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="enioi">
<emphasis id="76CBEA80FFA9FFC402C830E0FE3BFB4D" box="[224,405,1242,1266]" italics="true" pageId="4" pageNumber="3">Dichotomius enioi</emphasis>
<bibRefCitation id="202E4B63FFA9FFC403B230E0FC90FB4D" author="Montoya-Molina S &amp; Vaz-de-Mello FZ" box="[410,830,1242,1266]" pageId="4" pageNumber="3" pagination="1 - 64" refId="ref8403" refString="Montoya-Molina S, Vaz-de-Mello FZ. 2021. Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae). European Journal of Taxonomy 734: 1 - 64." type="journal article" year="2021">Montoya-Molina and Vaz-de-Mello, 2021</bibRefCitation>
</taxonomicName>
(in part): 25.
</paragraph>
</subSubSection>
<subSubSection id="0CA56519FFA9FFC502E83138FF5AF982" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3" type="materials_examined">
<paragraph id="44003692FFA9FFC402E83138FD48FA81" blockId="4.[192,1440,1282,1345]" pageId="4" pageNumber="3">
<emphasis id="76CBEA80FFA9FFC402E83138FE2AFAA3" bold="true" box="[192,388,1282,1308]" pageId="4" pageNumber="3">Type depository.</emphasis>
<materialsCitation id="F4D73CCFFFA9FFC403A53138FD48FA81" collectionCode="MNCR, FSCA, ASC" pageId="4" pageNumber="3" specimenCount="216" specimenCount-female="92" specimenCount-male="124" typeStatus="holotype">
<typeStatus id="9B048830FFA9FFC403A53138FE5BFAA3" box="[397,501,1282,1308]" pageId="4" pageNumber="3" type="holotype">Holotype</typeStatus>
male, 
<typeStatus id="9B048830FFA9FFC4006A3138FD35FAA3" box="[578,667,1282,1308]" pageId="4" pageNumber="3" type="allotype">allotype</typeStatus>
female and 210 (
<specimenCount id="52B9FD1BFFA9FFC401483138FC3EFAA0" box="[864,912,1282,1311]" pageId="4" pageNumber="3" type="female">88♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA9FFC401B63138FC75FAA0" box="[926,987,1282,1311]" pageId="4" pageNumber="3" type="male">122♂</specimenCount>
) 
<typeStatus id="9B048830FFA9FFC401C53139FBF7FAA2" box="[1005,1113,1283,1309]" pageId="4" pageNumber="3" type="paratype">paratypes</typeStatus>
in 
<collectionCode id="22AEAE57FFA9FFC406A83138FB7AFAA3" box="[1152,1236,1282,1308]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="4" pageNumber="3">MNCR</collectionCode>
, 2 (
<specimenCount id="52B9FD1BFFA9FFC407293138FA8CFAA0" box="[1281,1314,1282,1311]" pageId="4" pageNumber="3" type="female">1♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA9FFC407183138FAFDFAA0" box="[1328,1363,1282,1311]" pageId="4" pageNumber="3" type="male">1♂</specimenCount>
) 
<typeStatus id="9B048830FFA9FFC4074D3139FF55FA80" pageId="4" pageNumber="3" type="paratype">paratypes</typeStatus>
in 
<collectionCode id="22AEAE57FFA9FFC40336311EFECDFA81" box="[286,355,1316,1342]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="4" pageNumber="3" type="Museum">FSCA</collectionCode>
, and 4 (
<specimenCount id="52B9FD1BFFA9FFC40392311EFE73FAFE" box="[442,477,1316,1345]" pageId="4" pageNumber="3" type="female">3♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA9FFC403C2311EFDA2FAFE" box="[490,524,1316,1345]" pageId="4" pageNumber="3" type="male">1♂</specimenCount>
) 
<typeStatus id="9B048830FFA9FFC40034311FFD26FA80" box="[540,648,1317,1343]" pageId="4" pageNumber="3" type="paratype">paratypes</typeStatus>
in 
<collectionCode id="22AEAE57FFA9FFC40084311EFD4CFA81" box="[684,738,1316,1342]" pageId="4" pageNumber="3">ASC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="44003692FFA9FFC402E83174FD02FA13" blockId="4.[191,1441,1358,1928]" pageId="4" pageNumber="3">
<emphasis id="76CBEA80FFA9FFC402E83174FEE4FAD7" bold="true" box="[192,330,1358,1384]" pageId="4" pageNumber="3">Type status.</emphasis>
<materialsCitation id="F4D73CCFFFA9FFC403673174FD02FA13" collectorName="Est. Q. Bonita. &amp; R. M. Guzman." country="Costa Rica" elevation="50" location="Puntarenas Prov." pageId="4" pageNumber="3" specimenCount="2" stateProvince="Puntarenas" typeStatus="holotype">
<typeStatus id="9B048830FFA9FFC403673174FE19FAD7" box="[335,439,1358,1384]" pageId="4" pageNumber="3" type="holotype">Holotype</typeStatus>
male, 
<emphasis id="76CBEA80FFA9FFC400283175FDDFFAD7" box="[512,625,1358,1384]" italics="true" pageId="4" pageNumber="3">type labels</emphasis>
: “ 
<collectingCountry id="3CA87602FFA9FFC400AE3174FC87FAD7" box="[646,809,1358,1384]" name="Costa Rica" pageId="4" pageNumber="3">COSTA RICA</collectingCountry>
: 
<emphasis id="76CBEA80FFA9FFC401193174FC57FAD7" bold="true" box="[817,1017,1358,1384]" pageId="4" pageNumber="3">
<collectingRegion id="867BF870FFA9FFC401193174FC57FAD7" box="[817,1017,1358,1384]" country="Costa Rica" name="Puntarenas" pageId="4" pageNumber="3">Puntarenas Prov.</collectingRegion>
</emphasis>
<collectorName id="E94A5344FFA9FFC401D63175FB31FAD7" box="[1022,1183,1358,1385]" pageId="4" pageNumber="3">Est. Q. Bonita.</collectorName>
<quantity id="83479B77FFA9FFC4068C3174FB70FAD6" box="[1188,1246,1358,1385]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="3" unit="m" value="50.0">
<elevation id="CF92D1A1FFA9FFC4068C3174FB70FAD6" box="[1188,1246,1358,1385]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="3" unit="m" value="50.0">50 m</elevation>
</quantity>
, agos 1993, 
<collectorName id="E94A5344FFA9FFC4074A3175FE89FA35" pageId="4" pageNumber="3">R. M. Guzmán.</collectorName>
L N 194500_469850 #2297. INBIO CRI001 969440”. “ 
<typeStatus id="9B048830FFA9FFC401A6314AFB8EFA35" box="[910,1056,1392,1418]" pageId="4" pageNumber="3" type="holotype">HOLOTYPE</typeStatus>
/ 
<taxonomicName id="83BF4D11FFA9FFC40615314AFB66FA35" authorityName="Hope" authorityYear="1838" box="[1085,1224,1392,1418]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="76CBEA80FFA9FFC40615314AFB66FA35" box="[1085,1224,1392,1418]" italics="true" pageId="4" pageNumber="3">Dichotomius</emphasis>
</taxonomicName>
/ 
<taxonomicName id="83BF4D11FFA9FFC406CA314AFAE2FA35" authority="Solís &amp; Kohlmann, 2022" authorityName="Solís &amp; Kohlmann" authorityYear="2022" box="[1250,1356,1392,1418]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov.">
<emphasis id="76CBEA80FFA9FFC406CA314AFAE2FA35" box="[1250,1356,1392,1418]" italics="true" pageId="4" pageNumber="3">woodruffi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6DF857FBFFA9FFC4077A314AFA21FA35" box="[1362,1423,1392,1418]" pageId="4" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
/ Solís &amp; Kohlmann ded. 2022 [printed, red]”.
</materialsCitation>
</paragraph>
<paragraph id="44003692FFA9FFC50328318EFACEFB3B" blockId="4.[191,1441,1358,1928]" lastBlockId="5.[191,1441,214,1597]" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3">
<emphasis id="76CBEA80FFA9FFC40328318EFEDCFA71" bold="true" box="[256,370,1460,1486]" pageId="4" pageNumber="3">
<typeStatus id="9B048830FFA9FFC40328318EFEDCFA71" box="[256,370,1460,1486]" pageId="4" pageNumber="3" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="52B9FD1BFFA9FFC403A9318EFE1DFA6F" box="[385,435,1460,1488]" pageId="4" pageNumber="3" type="female">89♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA9FFC403E8318EFE53FA6F" box="[448,509,1460,1488]" pageId="4" pageNumber="3" type="male">122♂</specimenCount>
). 
<materialsCitation id="F4D73CCFFFA9FFC5003C318EFAF2FB3B" collectingDate="1990-07-01" collectingDateMax="2019-11-27" collectingDateMin="1990-07-01" collectionCode="MNCR, ASC, FSAC" collectorName="Est. Q. Bonita. &amp; J. C. Saborio. &amp; Col. Rodriguez, G. &amp; Col. Lopez, E. &amp; Col. Azofeifa A. &amp; Col. Hernandez, B. &amp; Osa. P. N. Corcovado. Est. Agujas. &amp; Col. Bello, E. &amp; Col. Saborio, J. C. &amp; Col. Guzman, R. &amp; Col. Cardenas &amp; Col. Briceno &amp; Col. Porras Vega &amp; Col. Alvarado, F. &amp; Col. Pulido, A. &amp; P. N. La Cangreja &amp; N. de Bajo La Palma. &amp; Col. Solis, A." country="Costa Rica" county="Rio San Lorenzo" elevation="50" lastPageId="5" lastPageNumber="4" latitude="11.991915" location="Agujas" longLatPrecision="1" longitude="-86.322754" municipality="Rio San Lorenzo" pageId="4" pageNumber="3" specimenCount="1414" specimenCount-female="93" specimenCount-male="124" stateProvince="Managua" typeStatus="allotype">
<typeStatus id="9B048830FFA9FFC4003C318EFDD8FA71" box="[532,630,1460,1486]" pageId="4" pageNumber="3" type="allotype">Allotype</typeStatus>
, female, 
<emphasis id="76CBEA80FFA9FFC400F1318EFCE4FA71" box="[729,842,1460,1486]" italics="true" pageId="4" pageNumber="3">type labels</emphasis>
. “ 
<collectingCountry id="3CA87602FFA9FFC40149318EFBAAFA71" box="[865,1028,1460,1486]" name="Costa Rica" pageId="4" pageNumber="3">COSTA RICA</collectingCountry>
: 
<emphasis id="76CBEA80FFA9FFC40627318EFB79FA71" bold="true" box="[1039,1239,1460,1486]" pageId="4" pageNumber="3">Puntarenas Prov.</emphasis>
<collectorName id="E94A5344FFA9FFC406F5318EFAD1FA71" box="[1245,1407,1460,1486]" pageId="4" pageNumber="3">Est. Q. Bonita.</collectorName>
<quantity id="83479B77FFA9FFC407AE3189FF74FA4F" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="3" unit="m" value="50.0">
<elevation id="CF92D1A1FFA9FFC407AE3189FF74FA4F" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="3" unit="m" value="50.0">50 m</elevation>
</quantity>
, Res. Biol. Carara. 
<date id="30011052FFA9FFC4038D31ECFDBAFA4F" box="[421,532,1494,1520]" pageId="4" pageNumber="3" value="1992-05">
<collectingDate id="2045E9BAFFA9FFC4038D31ECFDBAFA4F" box="[421,532,1494,1520]" pageId="4" pageNumber="3" value="1992-05">May 1992</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA9FFC4003531ECFD0BFA50" box="[541,677,1493,1520]" pageId="4" pageNumber="3">J.C. Saborío.</collectorName>
L-N194500-469850. INBIO CRI000 796048”. “ 
<typeStatus id="9B048830FFA9FFC4068E31ECFA80FA4F" box="[1190,1326,1494,1520]" pageId="4" pageNumber="3" type="allotype">ALLOTYPE</typeStatus>
/ 
<taxonomicName id="83BF4D11FFA9FFC4071731EFFF5BF9AE" authorityName="Hope" authorityYear="1838" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="76CBEA80FFA9FFC4071731EFFF5BF9AE" italics="true" pageId="4" pageNumber="3">Dichotomius</emphasis>
</taxonomicName>
/ 
<taxonomicName id="83BF4D11FFA9FFC4032331CDFEDBF9AE" authority="Solís &amp; Kohlmann, 2022" authorityName="Solís &amp; Kohlmann" authorityYear="2022" box="[267,373,1527,1553]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov.">
<emphasis id="76CBEA80FFA9FFC4032331CDFEDBF9AE" box="[267,373,1527,1553]" italics="true" pageId="4" pageNumber="3">woodruffi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6DF857FBFFA9FFC4035331C2FE1BF9AD" box="[379,437,1528,1554]" pageId="4" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
/ Solís &amp; Kohlmann ded. 2022 [printed, red]”. 
<emphasis id="76CBEA80FFA9FFC4019231C2FB2FF9AD" bold="true" box="[954,1153,1528,1554]" pageId="4" pageNumber="3">Puntarenas Prov.</emphasis>
<collectorName id="E94A5344FFA9FFC406AF31C2FA86F9AE" box="[1159,1320,1527,1554]" pageId="4" pageNumber="3">Est. Q. Bonita.</collectorName>
<quantity id="83479B77FFA9FFC4070531CDFAC9F9AD" box="[1325,1383,1527,1554]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="3" unit="m" value="50.0">
<elevation id="CF92D1A1FFA9FFC4070531CDFAC9F9AD" box="[1325,1383,1527,1554]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="3" unit="m" value="50.0">50 m</elevation>
</quantity>
, Res. Biol. Carara. 
<date id="30011052FFA9FFC4037C3220FE6BF98C" box="[340,453,1561,1588]" pageId="4" pageNumber="3" value="1992-05">
<collectingDate id="2045E9BAFFA9FFC4037C3220FE6BF98C" box="[340,453,1561,1588]" pageId="4" pageNumber="3" value="1992-05">May 1992</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA9FFC403E73223FDF4F98C" box="[463,602,1561,1587]" pageId="4" pageNumber="3">J.C. Saborío.</collectorName>
L-N194500-469850. INBIO CRI000 796048”. “ 
<typeStatus id="9B048830FFA9FFC4064D3223FB43F98C" box="[1125,1261,1561,1587]" pageId="4" pageNumber="3" type="allotype">ALLOTYPE</typeStatus>
/ 
<taxonomicName id="83BF4D11FFA9FFC4072D3223FA3EF98C" authorityName="Hope" authorityYear="1838" box="[1285,1424,1561,1587]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="76CBEA80FFA9FFC4072D3223FA3EF98C" box="[1285,1424,1561,1587]" italics="true" pageId="4" pageNumber="3">Dichotomius</emphasis>
</taxonomicName>
/ 
<taxonomicName id="83BF4D11FFA9FFC402E93201FE85F9EA" authority="Solís &amp; Kohlmann, 2022" authorityName="Solís &amp; Kohlmann" authorityYear="2022" box="[193,299,1595,1621]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov.">
<emphasis id="76CBEA80FFA9FFC402E93201FE85F9EA" box="[193,299,1595,1621]" italics="true" pageId="4" pageNumber="3">woodruffi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6DF857FBFFA9FFC403193206FEC3F9E9" box="[305,365,1596,1622]" pageId="4" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
/ Solís &amp; Kohlmann ded. 2022 [printed, red]”. “ 
<collectingCountry id="3CA87602FFA9FFC401AF3201FB85F9EA" box="[903,1067,1595,1621]" name="Costa Rica" pageId="4" pageNumber="3">COSTA RICA</collectingCountry>
: 
<emphasis id="76CBEA80FFA9FFC4061E3201FAAFF9EA" bold="true" box="[1078,1281,1595,1621]" pageId="4" pageNumber="3">Guanacaste Prov.</emphasis>
Area de cons. Arenal. 
<collectingCounty id="AD614E1EFFA9FFC403333267FE79F9C7" box="[283,471,1629,1656]" pageId="4" pageNumber="3">Río San Lorenzo</collectingCounty>
, Tierras Morenas. 
<geoCoordinate id="218B5055FFA9FFC400813267FC97F9C7" box="[681,825,1629,1656]" degrees="10.610459" direction="north" orientation="latitude" pageId="4" pageNumber="3" precision="1" value="10.610459">10.610459 N</geoCoordinate>
, − 
<geoCoordinate id="218B5055FFA9FFC4017B3267FC46F9C7" box="[851,1000,1629,1656]" degrees="84.994969" direction="west" orientation="longitude" pageId="4" pageNumber="3" precision="1" value="-84.99497">84.994969 W</geoCoordinate>
. 
<date id="30011052FFA9FFC401DB3267FBDCF9C8" box="[1011,1138,1629,1655]" pageId="4" pageNumber="3" value="1994-07-01">
<collectingDate id="2045E9BAFFA9FFC401DB3267FBDCF9C8" box="[1011,1138,1629,1655]" pageId="4" pageNumber="3" value="1994-07-01">01 jul 1994</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA9FFC406553267FAE3F9C8" box="[1149,1357,1629,1655]" pageId="4" pageNumber="3">Col. Rodríguez, G.</collectorName>
” (
<specimenCount id="52B9FD1BFFA9FFC407413267FA3EF9C6" box="[1385,1424,1629,1657]" pageId="4" pageNumber="3" type="female">1 ♀</specimenCount>
); with same data as previous but “Tierras Morenas. 
<geoCoordinate id="218B5055FFA9FFC400CC3245FC2EF926" box="[740,896,1663,1690]" degrees="10.5710619" direction="north" orientation="latitude" pageId="4" pageNumber="3" precision="1" value="10.571062">10.5710619 N</geoCoordinate>
, − 
<geoCoordinate id="218B5055FFA9FFC401B03245FB85F926" box="[920,1067,1663,1690]" degrees="85.025949" direction="west" orientation="longitude" pageId="4" pageNumber="3" precision="1" value="-85.02595">85.025949 W</geoCoordinate>
. 
<date id="30011052FFA9FFC4061C3245FB6CF926" box="[1076,1218,1663,1689]" pageId="4" pageNumber="3" value="1994-05-01">
<collectingDate id="2045E9BAFFA9FFC4061C3245FB6CF926" box="[1076,1218,1663,1689]" pageId="4" pageNumber="3" value="1994-05-01">01 may 1994</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA9FFC406E23245FA39F926" box="[1226,1431,1663,1689]" pageId="4" pageNumber="3">Col. Rodríguez, G.</collectorName>
” (
<specimenCount id="52B9FD1BFFA9FFC402EF329BFF5CF902" box="[199,242,1697,1725]" pageId="4" pageNumber="3" type="female">4 ♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA9FFC402D6329BFE8EF902" box="[254,288,1697,1725]" pageId="4" pageNumber="3" type="male">2♂</specimenCount>
); with same data as previous but “ 
<collectingMunicipality id="A464ACE8FFA9FFC400B0329BFCFCF904" box="[664,850,1697,1723]" pageId="4" pageNumber="3">Río San Lorenzo</collectingMunicipality>
, Tierras Morenas. 
<geoCoordinate id="218B5055FFA9FFC40637329BFB00F904" box="[1055,1198,1697,1724]" degrees="10.610459" direction="north" orientation="latitude" pageId="4" pageNumber="3" precision="1" value="10.610459">10.610459 N</geoCoordinate>
, − 
<geoCoordinate id="218B5055FFA9FFC406EE329BFAF7F904" box="[1222,1369,1697,1723]" degrees="84.994969" direction="west" orientation="longitude" pageId="4" pageNumber="3" precision="1" value="-84.99497">84.994969 W</geoCoordinate>
. 
<date id="30011052FFA9FFC4074B329BFF56F962" pageId="4" pageNumber="3" value="1994-07-01">
<collectingDate id="2045E9BAFFA9FFC4074B329BFF56F962" pageId="4" pageNumber="3" value="1994-07-01">01 jul 1994</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA9FFC4032A32F9FE7FF962" box="[258,465,1731,1757]" pageId="4" pageNumber="3">Col. Rodríguez, G.</collectorName>
” (
<specimenCount id="52B9FD1BFFA9FFC403C232F9FD8CF960" box="[490,546,1731,1759]" pageId="4" pageNumber="3" type="male">10 ♂</specimenCount>
); with same data as previous but “Área de cons. Guanacaste. La Cruz, La Garita, Est Los Almendros. 
<geoCoordinate id="218B5055FFA9FFC403B732DEFD83F940" box="[415,557,1764,1791]" degrees="11.033856" direction="north" orientation="latitude" pageId="4" pageNumber="3" precision="1" value="11.033856">11.033856 N</geoCoordinate>
, − 
<geoCoordinate id="218B5055FFA9FFC4006C32DEFD79F940" box="[580,727,1764,1791]" degrees="85.524789" direction="west" orientation="longitude" pageId="4" pageNumber="3" precision="1" value="-85.52479">85.524789 W</geoCoordinate>
. 
<date id="30011052FFA9FFC400C832DFFCF5F940" box="[736,859,1764,1791]" pageId="4" pageNumber="3" value="1992-07-28">
<collectingDate id="2045E9BAFFA9FFC400C832DFFCF5F940" box="[736,859,1764,1791]" pageId="4" pageNumber="3" value="1992-07-28">28 jul 1992</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA9FFC4014C32DEFC55F940" box="[868,1019,1764,1791]" pageId="4" pageNumber="3">Col. López, E.</collectorName>
” (
<specimenCount id="52B9FD1BFFA9FFC4063C32DFFB97F8BE" box="[1044,1081,1765,1793]" pageId="4" pageNumber="3" type="male">1 ♂</specimenCount>
); with same data as previous but “ 
<emphasis id="76CBEA80FFA9FFC402E3333DFE3AF89E" bold="true" box="[203,404,1799,1825]" pageId="4" pageNumber="3">Puntarenas Prov.</emphasis>
Area de cons. Osa. Golfito, Pque Nal Corcovado, Est 
<location id="41606049FFA9FFC401C6333CFB93F89F" LSID="urn:lsid:plazi:treatment:CC168784FFA9FFC902E8306AFE47FA30:41606049FFA9FFC401C6333CFB93F89F" box="[1006,1085,1798,1824]" country="Costa Rica" county="Rio San Lorenzo" latitude="11.991915" longLatPrecision="1" longitude="-86.322754" municipality="Rio San Lorenzo" name="Agujas" pageId="4" pageNumber="3" stateProvince="Managua">Agujas</location>
, Las Quebraditas. 
<geoCoordinate id="218B5055FFA9FFC40724333CFA32F89E" box="[1292,1436,1798,1825]" degrees="8.5226319" direction="north" orientation="latitude" pageId="4" pageNumber="3" precision="1" value="8.522632">8.5226319 N</geoCoordinate>
, − 
<geoCoordinate id="218B5055FFA9FFC402E73312FEFBF8FC" box="[207,341,1832,1859]" degrees="83.48411" direction="west" orientation="longitude" pageId="4" pageNumber="3" precision="1" value="-83.48411">83.48411 W</geoCoordinate>
. 
<date id="30011052FFA9FFC403763312FE42F8FD" box="[350,492,1832,1859]" pageId="4" pageNumber="3" value="2002-05-03">
<collectingDate id="2045E9BAFFA9FFC403763312FE42F8FD" box="[350,492,1832,1859]" pageId="4" pageNumber="3" value="2002-05-03">03 may 2002</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA9FFC403DC3312FD0DF8FD" box="[500,675,1832,1858]" pageId="4" pageNumber="3">Col. Azofeifa A.</collectorName>
” (
<specimenCount id="52B9FD1BFFA9FFC400913312FD72F8FA" box="[697,732,1832,1861]" pageId="4" pageNumber="3" type="female">2♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA9FFC400C03312FCA1F8FA" box="[744,783,1832,1861]" pageId="4" pageNumber="3" type="male">1 ♂</specimenCount>
); with same data as previous but “La Bonanza, 
<locationDeviation id="55B22FB0FFA9FFC4073D3313FEDFF8DB" firstLocation="Agujas" pageId="4" pageNumber="3" secondLocation="Cerro Rincon.">entre Agujas y Cerro Rincón.</locationDeviation>
<geoCoordinate id="218B5055FFA9FFC4035D3370FDADF8DA" box="[373,515,1866,1893]" degrees="8.5304219" direction="north" orientation="latitude" pageId="4" pageNumber="3" precision="1" value="8.530422">8.5304219 N</geoCoordinate>
, − 
<geoCoordinate id="218B5055FFA9FFC400333370FD00F8DA" box="[539,686,1866,1893]" degrees="83.449049" direction="west" orientation="longitude" pageId="4" pageNumber="3" precision="1" value="-83.44905">83.449049 W</geoCoordinate>
. 
<date id="30011052FFA9FFC4009F3370FC94F8DB" box="[695,826,1866,1892]" pageId="4" pageNumber="3" value="2008-06-17">
<collectingDate id="2045E9BAFFA9FFC4009F3370FC94F8DB" box="[695,826,1866,1892]" pageId="4" pageNumber="3" value="2008-06-17">17 jun 2008</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA9FFC4016B3370FBBDF8DB" box="[835,1043,1866,1892]" pageId="4" pageNumber="3">Col. Hernández, B.</collectorName>
” (
<specimenCount id="52B9FD1BFFA9FFC406023370FBE0F8D8" box="[1066,1102,1866,1895]" pageId="4" pageNumber="3" type="female">2♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA9FFC406723370FB2EF8D8" box="[1114,1152,1866,1895]" pageId="4" pageNumber="3" type="male">3 ♂</specimenCount>
); with same data as previous but “ 
<collectorName id="E94A5344FFA9FFC4030C3356FD33F839" box="[292,669,1900,1927]" pageId="4" pageNumber="3">
Osa. P.N. Corcovado. Est. 
<location id="41606049FFA9FFC400613356FD37F839" LSID="urn:lsid:plazi:treatment:CC168784FFA9FFC902E8306AFE47FA30:41606049FFA9FFC400613356FD37F839" box="[585,665,1900,1926]" country="Costa Rica" county="Rio San Lorenzo" latitude="11.991915" longLatPrecision="1" longitude="-86.322754" municipality="Rio San Lorenzo" name="Agujas" pageId="4" pageNumber="3" stateProvince="Managua">Agujas</location>
.
</collectorName>
<geoCoordinate id="218B5055FFA9FFC4008B3356FC88F838" box="[675,806,1900,1927]" degrees="8.536614" direction="north" orientation="latitude" pageId="4" pageNumber="3" precision="1" value="8.536614">8.536614 N</geoCoordinate>
, − 
<geoCoordinate id="218B5055FFA9FFC401173356FC69F838" box="[831,967,1900,1927]" degrees="83.42551" direction="west" orientation="longitude" pageId="4" pageNumber="3" precision="1" value="-83.42551">83.42551 W</geoCoordinate>
. 
<date id="30011052FFA9FFC401F93356FBF6F839" box="[977,1112,1900,1926]" pageId="4" pageNumber="3" value="2000-04-12">
<collectingDate id="2045E9BAFFA9FFC401F93356FBF6F839" box="[977,1112,1900,1926]" pageId="4" pageNumber="3" value="2000-04-12">12 abr 2000</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA9FFC4064A3356FABAF839" box="[1122,1300,1900,1926]" pageId="4" pageNumber="3">Col. Azofeifa A.</collectorName>
” (
<specimenCount id="52B9FD1BFFA9FFC407073356FAF8F837" box="[1327,1366,1900,1928]" pageId="4" pageNumber="3" type="female">1 ♀</specimenCount>
); with same data as previous but “Area de cons. Pacifico Central. Garabito, Reserva Biol. Carara, Est. Quebrada Bonita. 
<geoCoordinate id="218B5055FFA8FFC502E834CDFEFEFEAD" box="[192,336,247,274]" degrees="9.7674529" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="1" value="9.767453">9.7674529 N</geoCoordinate>
, − 
<geoCoordinate id="218B5055FFA8FFC5034034C2FE53FEAD" box="[360,509,248,274]" degrees="84.608119" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="1" value="-84.608116">84.608119 W</geoCoordinate>
. 
<date id="30011052FFA8FFC5002F34C2FD2BFEAD" box="[519,645,248,274]" pageId="5" pageNumber="4" value="1990-07-01">
<collectingDate id="2045E9BAFFA8FFC5002F34C2FD2BFEAD" box="[519,645,248,274]" pageId="5" pageNumber="4" value="1990-07-01">01 jul 1990</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC500A634C2FCB3FEAD" box="[654,797,248,274]" pageId="5" pageNumber="4">Col. Bello, E.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC5011F34C2FCF6FEAB" box="[823,856,248,276]" pageId="5" pageNumber="4" type="female">1♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC5014D34C2FC20FEAB" box="[869,910,248,276]" pageId="5" pageNumber="4" type="male">2 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC5072434C2FA33FEAD" box="[1292,1437,248,274]" pageId="5" pageNumber="4" value="1992-05-01">
<collectingDate id="2045E9BAFFA8FFC5072434C2FA33FEAD" box="[1292,1437,248,274]" pageId="5" pageNumber="4" value="1992-05-01">01 may 1992</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC502E83523FED0FE8B" box="[192,382,281,308]" pageId="5" pageNumber="4">Col. Saborío, J.C.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC503B03523FE66FE89" box="[408,456,281,310]" pageId="5" pageNumber="4" type="female">15♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC503FE3523FDA3FE89" box="[470,525,281,310]" pageId="5" pageNumber="4" type="male">15 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC501B93520FBB4FE8B" box="[913,1050,281,308]" pageId="5" pageNumber="4" value="1992-06-01">
<collectingDate id="2045E9BAFFA8FFC501B93520FBB4FE8B" box="[913,1050,281,308]" pageId="5" pageNumber="4" value="1992-06-01">01 jun 1992</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC5060C3523FB4CFE8B" box="[1060,1250,281,308]" pageId="5" pageNumber="4">Col. Saborío, J.C.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC506D33520FAB1FE89" box="[1275,1311,282,310]" pageId="5" pageNumber="4" type="female">4♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC507053520FAF8FE89" box="[1325,1366,282,310]" pageId="5" pageNumber="4" type="male">8 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC503C63506FDC7FEE9" box="[494,617,315,342]" pageId="5" pageNumber="4" value="1992-07-01">
<collectingDate id="2045E9BAFFA8FFC503C63506FDC7FEE9" box="[494,617,315,342]" pageId="5" pageNumber="4" value="1992-07-01">01 jul 1992</collectingDate>
</date>
”. (
<specimenCount id="52B9FD1BFFA8FFC500AC3501FD09FEE7" box="[644,679,315,344]" pageId="5" pageNumber="4" type="female">6♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC5009C3501FD72FEE7" box="[692,732,315,344]" pageId="5" pageNumber="4" type="male">5 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC506703506FB78FEE9" box="[1112,1238,315,342]" pageId="5" pageNumber="4" value="1992-07-01">
<collectingDate id="2045E9BAFFA8FFC506703506FB78FEE9" box="[1112,1238,315,342]" pageId="5" pageNumber="4" value="1992-07-01">01 jul 1992</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC506F73501FA39FEE9" box="[1247,1431,315,342]" pageId="5" pageNumber="4">Col. Guzmán, R.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC502E03567FF5CFEC5" box="[200,242,349,378]" pageId="5" pageNumber="4" type="female">6 ♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC502D73567FE86FEC5" box="[255,296,349,378]" pageId="5" pageNumber="4" type="male">6 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC5008D3564FC85FEC7" box="[677,811,350,376]" pageId="5" pageNumber="4" value="1992-08-10">
<collectingDate id="2045E9BAFFA8FFC5008D3564FC85FEC7" box="[677,811,350,376]" pageId="5" pageNumber="4" value="1992-08-10">10 ago 1992</collectingDate>
</date>
.” (
<specimenCount id="52B9FD1BFFA8FFC501613564FCC4FEC5" box="[841,874,350,378]" pageId="5" pageNumber="4" type="female">1♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC5015F3564FC0EFEC5" box="[887,928,350,378]" pageId="5" pageNumber="4" type="male">2 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC507353564FA33FEC7" box="[1309,1437,350,376]" pageId="5" pageNumber="4" value="1992-09-01">
<collectingDate id="2045E9BAFFA8FFC507353564FA33FEC7" box="[1309,1437,350,376]" pageId="5" pageNumber="4" value="1992-09-01">01 set 1992</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC502E83545FED5FE26" box="[192,379,383,409]" pageId="5" pageNumber="4">Col. Saborío, J.C.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC503BD3545FE15FE23" box="[405,443,383,412]" pageId="5" pageNumber="4" type="male">1 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC5011D3545FC1AFE26" box="[821,948,383,410]" pageId="5" pageNumber="4" value="1992-09-02">
<collectingDate id="2045E9BAFFA8FFC5011D3545FC1AFE26" box="[821,948,383,410]" pageId="5" pageNumber="4" value="1992-09-02">02 set 1992</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC501953545FBDBFE26" box="[957,1141,383,409]" pageId="5" pageNumber="4">Col. Guzmán, R.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC506A43545FB01FE23" box="[1164,1199,383,412]" pageId="5" pageNumber="4" type="female">4♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC506943545FB4DFE23" box="[1212,1251,383,412]" pageId="5" pageNumber="4" type="male">3 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC50353359BFE50FE04" box="[379,510,417,444]" pageId="5" pageNumber="4" value="1992-10-18">
<collectingDate id="2045E9BAFFA8FFC50353359BFE50FE04" box="[379,510,417,444]" pageId="5" pageNumber="4" value="1992-10-18">18 oct 1992</collectingDate>
</date>
.” (
<specimenCount id="52B9FD1BFFA8FFC50033359BFD91FE01" box="[539,575,417,446]" pageId="5" pageNumber="4" type="female">3♀</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC501E9359BFBE0FE04" box="[961,1102,417,444]" pageId="5" pageNumber="4" value="1992-11-01">
<collectingDate id="2045E9BAFFA8FFC501E9359BFBE0FE04" box="[961,1102,417,444]" pageId="5" pageNumber="4" value="1992-11-01">01 nov 1992</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC50670359BFABBFE04" box="[1112,1301,417,443]" pageId="5" pageNumber="4">Col. Saborío, J.C.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC50707359BFAF7FE01" box="[1327,1369,417,446]" pageId="5" pageNumber="4" type="female">3 ♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC5074E359BFA3EFE01" box="[1382,1424,417,446]" pageId="5" pageNumber="4" type="male">1 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC5000035F9FD17FE62" box="[552,697,451,478]" pageId="5" pageNumber="4" value="1993-05-01">
<collectingDate id="2045E9BAFFA8FFC5000035F9FD17FE62" box="[552,697,451,478]" pageId="5" pageNumber="4" value="1993-05-01">01 may 1993</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC500EB35F9FC2EFE62" box="[707,896,451,477]" pageId="5" pageNumber="4">Col. Saborío, J.C.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC501B135F9FC6CFE5F" box="[921,962,451,480]" pageId="5" pageNumber="4" type="female">3 ♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC501E735F9FBA8FE5F" box="[975,1030,451,480]" pageId="5" pageNumber="4" type="male">17 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC507AC35F9FE8CFE40" pageId="5" pageNumber="4" value="1993-06-01">
<collectingDate id="2045E9BAFFA8FFC507AC35F9FE8CFE40" pageId="5" pageNumber="4" value="1993-06-01">01 jun 1993</collectingDate>
</date>
.” (
<specimenCount id="52B9FD1BFFA8FFC5031435DFFECBFDBD" box="[316,357,485,514]" pageId="5" pageNumber="4" type="male">2 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC500F535DFFCCDFE40" box="[733,867,485,511]" pageId="5" pageNumber="4" value="1993-06-01">
<collectingDate id="2045E9BAFFA8FFC500F535DFFCCDFE40" box="[733,867,485,511]" pageId="5" pageNumber="4" value="1993-06-01">01 jun 1993</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC5014335DFFB8DFE40" box="[875,1059,485,511]" pageId="5" pageNumber="4">Col. Guzmán, R.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC5061135DFFBCCFDBD" box="[1081,1122,485,514]" pageId="5" pageNumber="4" type="male">4 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC502DE363DFED9FD9E" box="[246,375,519,545]" pageId="5" pageNumber="4" value="1993-07-01">
<collectingDate id="2045E9BAFFA8FFC502DE363DFED9FD9E" box="[246,375,519,545]" pageId="5" pageNumber="4" value="1993-07-01">01 jul 1993</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC503AA363DFDEFFD9E" box="[386,577,519,545]" pageId="5" pageNumber="4">Col. Saborío, J.C.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC50072363DFDD1FD9C" box="[602,639,519,547]" pageId="5" pageNumber="4" type="female">2♀</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC5062D363DFB28FD9E" box="[1029,1158,519,545]" pageId="5" pageNumber="4" value="1993-07-01">
<collectingDate id="2045E9BAFFA8FFC5062D363DFB28FD9E" box="[1029,1158,519,545]" pageId="5" pageNumber="4" value="1993-07-01">01 jul 1993</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC506B9363DFAE2FD9E" box="[1169,1356,519,545]" pageId="5" pageNumber="4">Col. Guzmán, R.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC5074F363DFA3EFD9C" box="[1383,1424,519,547]" pageId="5" pageNumber="4" type="female">1 ♀</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC5000B3613FD05FDFC" box="[547,683,553,580]" pageId="5" pageNumber="4" value="1993-08-01">
<collectingDate id="2045E9BAFFA8FFC5000B3613FD05FDFC" box="[547,683,553,580]" pageId="5" pageNumber="4" value="1993-08-01">01 ago 1993</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC5009C3613FCC1FDFC" box="[692,879,553,579]" pageId="5" pageNumber="4">Col. Saborío, J.C.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC501AE3613FC04FDFA" box="[902,938,553,581]" pageId="5" pageNumber="4" type="female">2♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC5019E3613FC70FDFA" box="[950,990,553,581]" pageId="5" pageNumber="4" type="male">4 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC5077F3613FF56FDDA" pageId="5" pageNumber="4" value="1993-08-01">
<collectingDate id="2045E9BAFFA8FFC5077F3613FF56FDDA" pageId="5" pageNumber="4" value="1993-08-01">01 ago 1993</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC503293671FE16FDDA" box="[257,440,587,613]" pageId="5" pageNumber="4">Col. Guzmán, R.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC503E63671FE59FDD8" box="[462,503,587,615]" pageId="5" pageNumber="4" type="female">2 ♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC5002C3671FD85FDD8" box="[516,555,587,615]" pageId="5" pageNumber="4" type="male">9 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC5018A3671FBB0FDDA" box="[930,1054,587,613]" pageId="5" pageNumber="4" value="1993-09-01">
<collectingDate id="2045E9BAFFA8FFC5018A3671FBB0FDDA" box="[930,1054,587,613]" pageId="5" pageNumber="4" value="1993-09-01">01 set 1993</collectingDate>
</date>
.” (
<specimenCount id="52B9FD1BFFA8FFC506113671FBCFFDD8" box="[1081,1121,587,615]" pageId="5" pageNumber="4" type="female">2 ♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC506463671FB3AFDD8" box="[1134,1172,587,615]" pageId="5" pageNumber="4" type="male">2 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC5031D3657FE19FD38" box="[309,439,621,647]" pageId="5" pageNumber="4" value="1994-04-01">
<collectingDate id="2045E9BAFFA8FFC5031D3657FE19FD38" box="[309,439,621,647]" pageId="5" pageNumber="4" value="1994-04-01">01 abr 1994</collectingDate>
</date>
.” (
<specimenCount id="52B9FD1BFFA8FFC503FC3657FE54FD36" box="[468,506,621,649]" pageId="5" pageNumber="4" type="male">1 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC501593657FC43FD38" box="[881,1005,621,647]" pageId="5" pageNumber="4" value="1994-09-01">
<collectingDate id="2045E9BAFFA8FFC501593657FC43FD38" box="[881,1005,621,647]" pageId="5" pageNumber="4" value="1994-09-01">01 set 1994</collectingDate>
</date>
.” (
<specimenCount id="52B9FD1BFFA8FFC5062F3657FB81FD36" box="[1031,1071,621,649]" pageId="5" pageNumber="4" type="female">2 ♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC506143657FBCDFD36" box="[1084,1123,621,649]" pageId="5" pageNumber="4" type="male">1 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC502DD36B5FED5FD16" box="[245,379,655,681]" pageId="5" pageNumber="4" value="1994-10-01">
<collectingDate id="2045E9BAFFA8FFC502DD36B5FED5FD16" box="[245,379,655,681]" pageId="5" pageNumber="4" value="1994-10-01">01 oct 1994</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC503AD36B5FDEDFD16" box="[389,579,655,681]" pageId="5" pageNumber="4">Col. Saborío, J.C.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC5007636B5FD2FFD14" box="[606,641,655,683]" pageId="5" pageNumber="4" type="female">1♀</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC5062B36B5FB27FD16" box="[1027,1161,655,681]" pageId="5" pageNumber="4" value="1994-10-01">
<collectingDate id="2045E9BAFFA8FFC5062B36B5FB27FD16" box="[1027,1161,655,681]" pageId="5" pageNumber="4" value="1994-10-01">01 oct 1994</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC506BB36B5FAE3FD16" box="[1171,1357,655,681]" pageId="5" pageNumber="4">Col. Guzmán, R.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC5074E36B5FA3EFD14" box="[1382,1424,655,683]" pageId="5" pageNumber="4" type="female">8 ♀</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC5000B368BFD03FD74" box="[547,685,689,715]" pageId="5" pageNumber="4" value="1994-11-01">
<collectingDate id="2045E9BAFFA8FFC5000B368BFD03FD74" box="[547,685,689,715]" pageId="5" pageNumber="4" value="1994-11-01">01 nov 1994</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC5009E368BFCDEFD74" box="[694,880,689,715]" pageId="5" pageNumber="4">Col. Saborío, J.C.</collectorName>
” 
<specimenCount id="52B9FD1BFFA8FFC50157368BFC09FD72" box="[895,935,689,717]" pageId="5" pageNumber="4" type="female">1 ♀</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC50708368BFA33FD74" box="[1312,1437,688,715]" pageId="5" pageNumber="4" value="1995-07-01">
<collectingDate id="2045E9BAFFA8FFC50708368BFA33FD74" box="[1312,1437,688,715]" pageId="5" pageNumber="4" value="1995-07-01">01 jul 1995</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC5029736E9FED9FD52" box="[191,375,723,749]" pageId="5" pageNumber="4">Col. Guzmán, R.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC503B936E9FE1DFD50" box="[401,435,723,751]" pageId="5" pageNumber="4" type="female">1♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC5039736E9FE49FD50" box="[447,487,723,751]" pageId="5" pageNumber="4" type="male">1 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC5014A36E9FC46FD52" box="[866,1000,722,749]" pageId="5" pageNumber="4" value="1995-08-01">
<collectingDate id="2045E9BAFFA8FFC5014A36E9FC46FD52" box="[866,1000,722,749]" pageId="5" pageNumber="4" value="1995-08-01">01 ago 1995</collectingDate>
</date>
.” (
<specimenCount id="52B9FD1BFFA8FFC5062D36E9FB82FD50" box="[1029,1068,723,751]" pageId="5" pageNumber="4" type="female">1 ♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC5061036E9FBCEFD50" box="[1080,1120,723,751]" pageId="5" pageNumber="4" type="male">2 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC502DD36CFFE2BFCB1" box="[245,389,756,783]" pageId="5" pageNumber="4" value="1996-05-01">
<collectingDate id="2045E9BAFFA8FFC502DD36CFFE2BFCB1" box="[245,389,756,783]" pageId="5" pageNumber="4" value="1996-05-01">01 may 1996</collectingDate>
</date>
.” (
<specimenCount id="52B9FD1BFFA8FFC5038B36CFFE65FCAE" box="[419,459,757,785]" pageId="5" pageNumber="4" type="male">1 ♂</specimenCount>
); with same data as previous but “Area de cons. Tempisque. Lepanto, Karen Morgensen, Sendero Principal. 
<geoCoordinate id="218B5055FFA8FFC503B1372CFD84FC8E" box="[409,554,790,817]" degrees="9.8670949" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="1" value="9.867095">9.8670949 N</geoCoordinate>
, − 
<geoCoordinate id="218B5055FFA8FFC5006C372CFD63FC8E" box="[580,717,790,817]" degrees="85.05995" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="1" value="-85.05995">85.05995 W</geoCoordinate>
. 
<date id="30011052FFA8FFC500F1372DFCF2FC8E" box="[729,860,791,817]" pageId="5" pageNumber="4" value="2003-09-20">
<collectingDate id="2045E9BAFFA8FFC500F1372DFCF2FC8E" box="[729,860,791,817]" pageId="5" pageNumber="4" value="2003-09-20">20 set 2003</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC5014F372CFBA4FC8F" box="[871,1034,790,816]" pageId="5" pageNumber="4">Col. Cárdenas</collectorName>
, Yow.” (
<specimenCount id="52B9FD1BFFA8FFC5064C372DFB2BFC8C" box="[1124,1157,791,819]" pageId="5" pageNumber="4" type="female">1♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC506BB372DFB10FC8C" box="[1171,1214,791,819]" pageId="5" pageNumber="4" type="male">2 ♂</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC503743703FE4FFCEC" box="[348,481,824,851]" pageId="5" pageNumber="4" value="2003-06-23">
<collectingDate id="2045E9BAFFA8FFC503743703FE4FFCEC" box="[348,481,824,851]" pageId="5" pageNumber="4" value="2003-06-23">23 jun 2003</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC503C23702FDD9FCED" box="[490,631,824,850]" pageId="5" pageNumber="4">Col. Briceño</collectorName>
, Duvalier.” (
<specimenCount id="52B9FD1BFFA8FFC500D63703FCB0FCEA" box="[766,798,825,853]" pageId="5" pageNumber="4" type="female">1♀</specimenCount>
); with same data as previous but “ 
<date id="30011052FFA8FFC506BD3703FAB5FCEC" box="[1173,1307,824,851]" pageId="5" pageNumber="4" value="2003-06-22">
<collectingDate id="2045E9BAFFA8FFC506BD3703FAB5FCEC" box="[1173,1307,824,851]" pageId="5" pageNumber="4" value="2003-06-22">22 jun 2003</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC5070C3702FF56FCCA" pageId="5" pageNumber="4">Col. Porras Vega</collectorName>
, Wendy.” (
<specimenCount id="52B9FD1BFFA8FFC503443760FE3BFCC8" box="[364,405,858,887]" pageId="5" pageNumber="4" type="male">3 ♂</specimenCount>
); with same data as previous but “Send. Central. Est. San Miguel. 
<geoCoordinate id="218B5055FFA8FFC506463760FB53FCCA" box="[1134,1277,858,885]" degrees="9.5839859" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="1" value="9.583986">9.5839859 N</geoCoordinate>
, − 
<geoCoordinate id="218B5055FFA8FFC5073E3760FA32FCCA" box="[1302,1436,858,885]" degrees="85.11212" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="1" value="-85.11212">85.11212 W</geoCoordinate>
. 
<date id="30011052FFA8FFC502E83746FEEAFC29" box="[192,324,892,918]" pageId="5" pageNumber="4" value="1997-04-22">
<collectingDate id="2045E9BAFFA8FFC502E83746FEEAFC29" box="[192,324,892,918]" pageId="5" pageNumber="4" value="1997-04-22">22 abr 1997</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC503653746FDA9FC29" box="[333,519,892,918]" pageId="5" pageNumber="4">Col. Alvarado, F.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC500373746FDE8FC26" box="[543,582,892,921]" pageId="5" pageNumber="4" type="male">1 ♂</specimenCount>
); with same data as previous but “Area de cons. La Amistad Pacifico. Parcelas IDA. 
<geoCoordinate id="218B5055FFA8FFC502D737A4FE2CFC06" box="[255,386,926,953]" degrees="8.958083" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="1" value="8.958083">8.958083 N</geoCoordinate>
, − 
<geoCoordinate id="218B5055FFA8FFC503B537A4FD88FC06" box="[413,550,926,953]" degrees="83.05217" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="1" value="-83.05217">83.05217 W</geoCoordinate>
. 
<date id="30011052FFA8FFC5001937A4FD15FC07" box="[561,699,926,952]" pageId="5" pageNumber="4" value="2008-06-10">
<collectingDate id="2045E9BAFFA8FFC5001937A4FD15FC07" box="[561,699,926,952]" pageId="5" pageNumber="4" value="2008-06-10">10 jun 2008</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC500EE37A4FCC3FC07" box="[710,877,926,952]" pageId="5" pageNumber="4">Col. Pulido, A.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC501AE37A4FC1DFC04" box="[902,947,926,955]" pageId="5" pageNumber="4" type="male">2 ♂</specimenCount>
); with same data as previous but “ 
<emphasis id="76CBEA80FFA8FFC5071637A4FF52FC65" bold="true" pageId="5" pageNumber="4">San José Prov.</emphasis>
Area de cons. Pacifico Central. Est. Bijagual, 
<specimenCount id="52B9FD1BFFA8FFC500D637FAFCE5FC64" box="[766,843,960,987]" pageId="5" pageNumber="4" type="generic">
<quantity id="83479B77FFA8FFC500D637FAFCE6FC64" box="[766,840,960,987]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="4" unit="m" value="600.0">
<elevation id="CF92D1A1FFA8FFC500D637FAFCE6FC64" box="[766,840,960,987]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="4" unit="m" value="600.0">600 m</elevation>
</quantity>
.
</specimenCount>
<locationDeviation id="55B22FB0FFA8FFC5017A37FBFBBEFC65" box="[850,1040,960,987]" location="Bijagualito." pageId="5" pageNumber="4">N de Bijagualito.</locationDeviation>
<geoCoordinate id="218B5055FFA8FFC5063E37FAFB09FC64" box="[1046,1191,960,987]" degrees="9.7430889" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="1" value="9.743089">9.7430889 N</geoCoordinate>
, − 
<geoCoordinate id="218B5055FFA8FFC506E837FAFAF8FC64" box="[1216,1366,960,987]" degrees="84.544759" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="1" value="-84.54476">84.544759 W</geoCoordinate>
. 
<date id="30011052FFA8FFC5074937FAFF54FC42" pageId="5" pageNumber="4" value="1995-07-01">
<collectingDate id="2045E9BAFFA8FFC5074937FAFF54FC42" pageId="5" pageNumber="4" value="1995-07-01">01 jul 1995</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC5032B37D8FE11FC43" box="[259,447,994,1020]" pageId="5" pageNumber="4">Col. Saborío, J.C.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC503FF37D8FDAEFC41" box="[471,512,994,1022]" pageId="5" pageNumber="4" type="female">6 ♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC5002537D8FD9BFC41" box="[525,565,994,1022]" pageId="5" pageNumber="4" type="male">4 ♂</specimenCount>
); with same data as previous but “ 
<collectorName id="E94A5344FFA8FFC5019937D8FBDAFC43" box="[945,1140,994,1021]" pageId="5" pageNumber="4">P. N. La Cangreja</collectorName>
, Puriscal, Chires. 
<geoCoordinate id="218B5055FFA8FFC5071337D8FF79FBA1" degrees="9.690711" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="1" value="9.690711">9.690711 N</geoCoordinate>
, − 
<geoCoordinate id="218B5055FFA8FFC502D9303EFE28FBA1" box="[241,390,1028,1055]" degrees="84.377529" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="1" value="-84.377525">84.377529 W</geoCoordinate>
. 
<date id="30011052FFA8FFC503B9303EFDB7FBA0" box="[401,537,1028,1055]" pageId="5" pageNumber="4" value="2005-06-28">
<collectingDate id="2045E9BAFFA8FFC503B9303EFDB7FBA0" box="[401,537,1028,1055]" pageId="5" pageNumber="4" value="2005-06-28">28 jun 2005</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC5000B303EFD56FBA1" box="[547,760,1028,1054]" pageId="5" pageNumber="4">Col. Hernández, B.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC50139303EFC92FB9F" box="[785,828,1028,1056]" pageId="5" pageNumber="4" type="female">2 ♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC50162303EFCDAFB9F" box="[842,884,1028,1056]" pageId="5" pageNumber="4" type="male">3 ♂</specimenCount>
); with same data as previous but “ 
<quantity id="83479B77FFA8FFC506D0303EFAEEFBA1" box="[1272,1344,1028,1054]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="4" unit="m" value="800.0">
<elevation id="CF92D1A1FFA8FFC506D0303EFAEEFBA1" box="[1272,1344,1028,1054]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="4" unit="m" value="800.0">800 m</elevation>
</quantity>
al 
<collectorName id="E94A5344FFA8FFC5074B303EFEC8FBFF" pageId="5" pageNumber="4">
N. de 
<location id="41606049FFA8FFC502E9301CFEC8FBFF" LSID="urn:lsid:plazi:treatment:CC168784FFA9FFC902E8306AFE47FA30:41606049FFA8FFC502E9301CFEC8FBFF" box="[193,358,1062,1088]" country="Costa Rica" county="Rio San Lorenzo" latitude="11.991915" longLatPrecision="1" longitude="-86.322754" municipality="Rio San Lorenzo" name="Bajo La Palma." pageId="5" pageNumber="4" stateProvince="Managua">Bajo La Palma.</location>
</collectorName>
<geoCoordinate id="218B5055FFA8FFC50343301CFE57FBFF" box="[363,505,1062,1088]" degrees="9.7630379" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="1" value="9.763038">9.7630379 N</geoCoordinate>
, − 
<geoCoordinate id="218B5055FFA8FFC50038301CFD0AFBFF" box="[528,676,1062,1088]" degrees="84.244929" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="1" value="-84.24493">84.244929 W</geoCoordinate>
. 
<date id="30011052FFA8FFC50084301CFC94FBFE" box="[684,826,1062,1089]" pageId="5" pageNumber="4" value="1995-05-22">
<collectingDate id="2045E9BAFFA8FFC50084301CFC94FBFE" box="[684,826,1062,1089]" pageId="5" pageNumber="4" value="1995-05-22">22 may 1995</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC5016A301CFC61FBFF" box="[834,975,1062,1088]" pageId="5" pageNumber="4">Col. Solis, A.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC501CE301CFBA2FBFD" box="[998,1036,1062,1090]" pageId="5" pageNumber="4" type="male">3♂</specimenCount>
). All 
<typeStatus id="9B048830FFA8FFC50661301CFB1BFBFF" box="[1097,1205,1062,1088]" pageId="5" pageNumber="4" type="paratype">paratypes</typeStatus>
deposited in 
<collectionCode id="22AEAE57FFA8FFC50760301CFA33FBFF" box="[1352,1437,1062,1088]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="5" pageNumber="4">MNCR</collectionCode>
. “ 
<collectingCountry id="3CA87602FFA8FFC502E03072FEC1FBDD" box="[200,367,1096,1122]" name="Nicaragua" pageId="5" pageNumber="4">NICARAGUA</collectingCountry>
: 
<emphasis id="76CBEA80FFA8FFC503513072FD8AFBDD" bold="true" box="[377,548,1096,1122]" pageId="5" pageNumber="4">
<collectingRegion id="867BF870FFA8FFC503513072FE48FBDD" box="[377,486,1096,1122]" country="Nicaragua" name="Managua" pageId="5" pageNumber="4">Managua</collectingRegion>
Dep.
</emphasis>
El Crucero, Reserva privada El Bajo, Sendero Quebrada, 730– 
<specimenCount id="52B9FD1BFFA8FFC506F43072FA86FBDD" box="[1244,1320,1096,1123]" pageId="5" pageNumber="4" type="generic">
<quantity id="83479B77FFA8FFC506F43072FA8AFBDD" box="[1244,1316,1096,1123]" metricMagnitude="2" metricUnit="m" metricValue="5.97" pageId="5" pageNumber="4" unit="m" value="597.0">
<elevation id="CF92D1A1FFA8FFC506F43072FA8AFBDD" box="[1244,1316,1096,1123]" metricMagnitude="2" metricUnit="m" metricValue="5.97" pageId="5" pageNumber="4" unit="m" value="597.0">597 m</elevation>
</quantity>
.
</specimenCount>
<geoCoordinate id="218B5055FFA8FFC507063072FF79FB3B" degrees="11.991915" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="1" value="11.991915">11.991915 N</geoCoordinate>
, − 
<geoCoordinate id="218B5055FFA8FFC502D83053FE2AFB3B" box="[240,388,1129,1156]" degrees="86.322757" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="1" value="-86.322754">86.322757 W</geoCoordinate>
. 
<date id="30011052FFA8FFC503A63050FDECFB3B" box="[398,578,1130,1156]" pageId="5" pageNumber="4" value="2019-11-26" valueMax="2019-11-27" valueMin="2019-11-26">
<collectingDate id="2045E9BAFFA8FFC503A63050FDECFB3B" box="[398,578,1130,1156]" pageId="5" pageNumber="4" value="2019-11-26" valueMax="2019-11-27" valueMin="2019-11-26">26–27 nov 2019</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC5007A3050FC8BFB3B" box="[594,805,1130,1156]" pageId="5" pageNumber="4">Col. Hernández, B.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC501143050FCCEFB39" box="[828,864,1130,1158]" pageId="5" pageNumber="4" type="female">4♀</specimenCount>
, 
<specimenCount id="52B9FD1BFFA8FFC501453050FC3BFB39" box="[877,917,1130,1158]" pageId="5" pageNumber="4" type="male">2 ♂</specimenCount>
, 
<typeStatus id="9B048830FFA8FFC5018A3050FBA0FB3B" box="[930,1038,1130,1156]" pageId="5" pageNumber="4" type="paratype">paratypes</typeStatus>
deposited in 
<collectionCode id="22AEAE57FFA8FFC5068C3050FB79FB3B" box="[1188,1239,1130,1156]" pageId="5" pageNumber="4">ASC</collectionCode>
and 
<collectionCode id="22AEAE57FFA8FFC507263050FAFBFB3B" box="[1294,1365,1130,1156]" pageId="5" pageNumber="4">FSAC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="44003692FFA8FFC5032830B6FF5AF982" blockId="5.[191,1441,214,1597]" pageId="5" pageNumber="4">
<materialsCitation id="F4D73CCFFFA8FFC5032830B6FF5EF982" collectingDate="2004-11-14" collectingDateMax="2016-06" collectingDateMin="2004-11-14" collectionCode="MNCR" collectorName="A. Solis &amp; E Cantillano, J &amp; Col. Gamboa R. B. &amp; de Asentamiento &amp; Sansi &amp; C Godinez &amp; Coord. &amp; La Bonanza &amp; B. Hernandez &amp; M. Moraga &amp; La Cangreja &amp; Col. B. Hernandez" country="Costa Rica" county="Their" elevation="375" latitude="9.69916" location="Guanacaste. Camino del Aguacate" longLatPrecision="1" longitude="-84.3775" municipality="Area" pageId="5" pageNumber="4" specimenCount="707" specimenCount-male="1" stateProvince="Guanacaste">
Other examined material (
<specimenCount id="52B9FD1BFFA8FFC5000E30B1FD63FB19" box="[550,717,1163,1190]" pageId="5" pageNumber="4" type="generic">106 specimens</specimenCount>
, non-paratypes, material identified by AS, not mounted, and preserved in alcohol at 
<collectionCode id="22AEAE57FFA8FFC503B53094FE5AFB77" box="[413,500,1198,1224]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="5" pageNumber="4">MNCR</collectionCode>
). 
<collectingCounty id="AD614E1EFFA8FFC5002C3097FDEEFB78" box="[516,576,1197,1223]" pageId="5" pageNumber="4">Their</collectingCounty>
locality data is the following: “ 
<collectingCountry id="3CA87602FFA8FFC501B93094FB9DFB77" box="[913,1075,1198,1224]" name="Costa Rica" pageId="5" pageNumber="4">COSTA RICA</collectingCountry>
: 
<emphasis id="76CBEA80FFA8FFC506143097FAABFB77" bold="true" box="[1084,1285,1197,1224]" pageId="5" pageNumber="4">
<collectingRegion id="867BF870FFA8FFC506143097FAABFB77" box="[1084,1285,1197,1224]" country="Costa Rica" name="Guanacaste" pageId="5" pageNumber="4">Guanacaste Prov.</collectingRegion>
</emphasis>
<collectingMunicipality id="A464ACE8FFA8FFC507233094FAEFFB77" box="[1291,1345,1198,1224]" pageId="5" pageNumber="4">Area</collectingMunicipality>
de cons. 
<location id="41606049FFA8FFC5029730EAFDE7FB55" LSID="urn:lsid:plazi:treatment:CC168784FFA9FFC902E8306AFE47FA30:41606049FFA8FFC5029730EAFDE7FB55" box="[191,585,1231,1258]" country="Costa Rica" county="Their" latitude="9.69916" longLatPrecision="1" longitude="-84.3775" municipality="Area" name="Guanacaste. Camino del Aguacate" pageId="5" pageNumber="4" stateProvince="Guanacaste">Guanacaste. Camino del Aguacate</location>
, 
<location id="41606049FFA8FFC5007B30EAFD4FFB56" LSID="urn:lsid:plazi:treatment:CC168784FFA9FFC902E8306AFE47FA30:41606049FFA8FFC5007B30EAFD4FFB56" box="[595,737,1231,1258]" country="Costa Rica" county="Their" latitude="9.69916" longLatPrecision="1" longitude="-84.3775" municipality="Area" name="Est. La Perla" pageId="5" pageNumber="4" stateProvince="Guanacaste">Est. La Perla</location>
, 
<quantity id="83479B77FFA8FFC500C430F5FC9BFB55" box="[748,821,1231,1258]" metricMagnitude="2" metricUnit="m" metricValue="3.75" pageId="5" pageNumber="4" unit="m" value="375.0">
<elevation id="CF92D1A1FFA8FFC500C430F5FC9BFB55" box="[748,821,1231,1258]" metricMagnitude="2" metricUnit="m" metricValue="3.75" pageId="5" pageNumber="4" unit="m" value="375.0">375 m</elevation>
</quantity>
, 
<collectingMethod id="9DFE4E85FFA8FFC5016830F5FC2CFB56" box="[832,898,1231,1257]" pageId="5" pageNumber="4">Pitfall</collectingMethod>
T. 28 al 30 junio de 2017, 
<collectorName id="E94A5344FFA8FFC5068230EAFAACFB56" box="[1194,1282,1231,1258]" pageId="5" pageNumber="4">A. Solís</collectorName>
, 
<collectorName id="E94A5344FFA8FFC5072630EAFF62FAB3" pageId="5" pageNumber="4">E Cantillano, J</collectorName>
. Cortez, Coord. 
<geoCoordinate id="218B5055FFA8FFC503AD30CBFE57FAB4" box="[389,505,1265,1291]" degrees="10.771236" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="1" value="10.771236">10.771236</geoCoordinate>
, 
<geoCoordinate id="218B5055FFA8FFC5002C30CBFD27FAB3" box="[516,649,1265,1292]" degrees="85.428981" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="1" value="-85.42898">−85.428981</geoCoordinate>
” (25 unverified sex); with same data as previous but “ 
<location id="41606049FFA8FFC506CA30C8FE00FA91" LSID="urn:lsid:plazi:treatment:CC168784FFA9FFC902E8306AFE47FA30:41606049FFA8FFC506CA30C8FE00FA91" country="Costa Rica" county="Their" latitude="9.69916" longLatPrecision="1" longitude="-84.3775" municipality="Area" name="Bosque Humedo en Sector Santa Rosa" pageId="5" pageNumber="4" stateProvince="Guanacaste">Bosque Húmedo en Sector Santa Rosa</location>
, ACG, 
<quantity id="83479B77FFA8FFC503D73129FDE6FA91" box="[511,584,1299,1326]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="4" unit="m" value="300.0">
<elevation id="CF92D1A1FFA8FFC503D73129FDE6FA91" box="[511,584,1299,1326]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="4" unit="m" value="300.0">300 m</elevation>
</quantity>
, 
<collectingMethod id="9DFE4E85FFA8FFC5007B3129FD3BFA92" box="[595,661,1299,1325]" pageId="5" pageNumber="4">Pitfall</collectingMethod>
T. 8 al 10 de 
<date id="30011052FFA8FFC501033129FC0BFA92" box="[811,933,1299,1325]" pageId="5" pageNumber="4" value="2016-06">
<collectingDate id="2045E9BAFFA8FFC501033129FC0BFA92" box="[811,933,1299,1325]" pageId="5" pageNumber="4" value="2016-06">junio 2016</collectingDate>
</date>
, 
<collectorName id="E94A5344FFA8FFC50198312EFBA9FA92" box="[944,1031,1299,1326]" pageId="5" pageNumber="4">A. Solís</collectorName>
, Coord. 
<geoCoordinate id="218B5055FFA8FFC506403129FB72FA91" box="[1128,1244,1299,1326]" degrees="10.851171" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="1" value="10.851171">10.851171</geoCoordinate>
, 
<geoCoordinate id="218B5055FFA8FFC506CF3129FAC5FA91" box="[1255,1387,1299,1326]" degrees="85.606955" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="1" value="-85.60696">−85.606955</geoCoordinate>
” (35 unverified sex); with same data as previous but “Área de cons. Tempisque. Bagaces, Est. Palo Verde. 
<geoCoordinate id="218B5055FFA8FFC50768310FFA32FAEF" box="[1344,1436,1333,1360]" degrees="10.35" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="555" value="10.35">10.35 N</geoCoordinate>
, − 
<geoCoordinate id="218B5055FFA8FFC502E6316DFECCFACD" box="[206,354,1367,1394]" degrees="85.352779" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="1" value="-85.352776">85.352779 W</geoCoordinate>
. 
<date id="30011052FFA8FFC50343316DFE5AFACE" box="[363,500,1367,1394]" pageId="5" pageNumber="4" value="2004-11-14">
<collectingDate id="2045E9BAFFA8FFC50343316DFE5AFACE" box="[363,500,1367,1394]" pageId="5" pageNumber="4" value="2004-11-14">14 nov 2004</collectingDate>
</date>
. 
<collectorName id="E94A5344FFA8FFC503D5316DFD67FACE" box="[509,713,1367,1394]" pageId="5" pageNumber="4">Col. Gamboa R. B.</collectorName>
” (
<specimenCount id="52B9FD1BFFA8FFC500CA316DFCA6FACB" box="[738,776,1367,1396]" pageId="5" pageNumber="4" type="male">1 ♂</specimenCount>
); with same data as previous but “ 
<emphasis id="76CBEA80FFA8FFC506A9316DFAE6FACE" bold="true" box="[1153,1352,1367,1393]" pageId="5" pageNumber="4">
<collectingRegion id="867BF870FFA8FFC506A9316DFAE6FACE" box="[1153,1352,1367,1393]" country="Costa Rica" name="Puntarenas" pageId="5" pageNumber="4">Puntarenas Prov.</collectingRegion>
</emphasis>
<location id="41606049FFA8FFC507653162FED0FA2C" LSID="urn:lsid:plazi:treatment:CC168784FFA9FFC902E8306AFE47FA30:41606049FFA8FFC507653162FED0FA2C" country="Costa Rica" county="Their" latitude="9.69916" longLatPrecision="1" longitude="-84.3775" municipality="Area" name="Bosque de Asentamiento" pageId="5" pageNumber="4" stateProvince="Guanacaste">
Bosque 
<collectorName id="E94A5344FFA8FFC502E83143FED0FA2C" box="[192,382,1401,1427]" pageId="5" pageNumber="4">de Asentamiento</collectorName>
</location>
de INDER en 
<collectorName id="E94A5344FFA8FFC5000A3143FDF0FA2C" box="[546,606,1401,1427]" pageId="5" pageNumber="4">Sansi</collectorName>
, SINAC, 
<quantity id="83479B77FFA8FFC500ED3143FCA3FA2B" box="[709,781,1401,1428]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="4" unit="m" value="500.0">
<elevation id="CF92D1A1FFA8FFC500ED3143FCA3FA2B" box="[709,781,1401,1428]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="4" unit="m" value="500.0">500 m</elevation>
</quantity>
, 
<collectingMethod id="9DFE4E85FFA8FFC501303143FCF4FA2C" box="[792,858,1401,1427]" pageId="5" pageNumber="4">Pitfall</collectingMethod>
T. 13 al 
<date id="30011052FFA8FFC5019F3143FBFCFA2C" box="[951,1106,1401,1428]" pageId="5" pageNumber="4" value="2013-06-15">
<collectingDate id="2045E9BAFFA8FFC5019F3143FBFCFA2C" box="[951,1106,1401,1428]" pageId="5" pageNumber="4" value="2013-06-15">15 junio 2013</collectingDate>
</date>
, 
<collectorName id="E94A5344FFA8FFC506753143FB1FFA2C" box="[1117,1201,1401,1427]" pageId="5" pageNumber="4">A. Solís</collectorName>
y 
<collectorName id="E94A5344FFA8FFC506E33143FAE9FA2C" box="[1227,1351,1401,1427]" pageId="5" pageNumber="4">C Godínez</collectorName>
, 
<collectorName id="E94A5344FFA8FFC507793143FA0FFA2C" box="[1361,1441,1401,1427]" pageId="5" pageNumber="4">Coord.</collectorName>
<geoCoordinate id="218B5055FFA8FFC502E831A1FE89FA09" box="[192,295,1435,1462]" degrees="8.959002" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="1" value="8.959002">8.959002</geoCoordinate>
, 
<geoCoordinate id="218B5055FFA8FFC5031A31A1FE18FA09" box="[306,438,1435,1462]" degrees="83.052355" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="1" value="-83.05235">−83.052355</geoCoordinate>
” (20 unverified sex); with same data as previous but “ 
<collectorName id="E94A5344FFA8FFC5062531A1FB38FA09" box="[1037,1174,1435,1462]" pageId="5" pageNumber="4">La Bonanza</collectorName>
, sendero 
<locationDeviation id="55B22FB0FFA8FFC5072831A6FD30FA68" firstLocation="Agujas" pageId="5" pageNumber="4" secondLocation="Cerro Rincon, Parque Nacional Corcovado">entre Agujas y Cerro Rincón, Parque Nacional Corcovado</locationDeviation>
, 
<quantity id="83479B77FFA8FFC5008F3187FC82FA68" box="[679,812,1469,1495]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="5" pageNumber="4" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">
<elevation id="CF92D1A1FFA8FFC5008F3187FC82FA68" box="[679,812,1469,1495]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="5" pageNumber="4" unit="m" value="500.0" valueMax="600.0" valueMin="400.0">400 a 600 m</elevation>
</quantity>
, 
<collectingMethod id="9DFE4E85FFA8FFC5011D3187FCD9FA68" box="[821,887,1469,1495]" pageId="5" pageNumber="4">Pitfall</collectingMethod>
T. 17 al 
<date id="30011052FFA8FFC501E73187FBC9FA68" box="[975,1127,1469,1495]" pageId="5" pageNumber="4" value="2008-06-19">
<collectingDate id="2045E9BAFFA8FFC501E73187FBC9FA68" box="[975,1127,1469,1495]" pageId="5" pageNumber="4" value="2008-06-19">19 junio 2008</collectingDate>
</date>
, 
<collectorName id="E94A5344FFA8FFC506593187FAA6FA68" box="[1137,1288,1469,1495]" pageId="5" pageNumber="4">B. Hernández</collectorName>
y 
<collectorName id="E94A5344FFA8FFC507083187FA32FA68" box="[1312,1436,1469,1495]" pageId="5" pageNumber="4">M. Moraga</collectorName>
, 
<collectorName id="E94A5344FFA8FFC5029731E5FEA0FA46" box="[191,270,1503,1529]" pageId="5" pageNumber="4">Coord.</collectorName>
<geoCoordinate id="218B5055FFA8FFC5033D31E5FED2FA45" box="[277,380,1503,1530]" degrees="8.530277" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="1" value="8.530277">8.530277</geoCoordinate>
, 
<geoCoordinate id="218B5055FFA8FFC503AF31E5FE5FFA46" box="[391,497,1503,1529]" degrees="83.4488" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="5" value="-83.4488">−83.4488</geoCoordinate>
” (14 unverified sex); with same data as previous but “ 
<collectorName id="E94A5344FFA8FFC5067831E5FB72FA46" box="[1104,1244,1503,1529]" pageId="5" pageNumber="4">La Cangreja</collectorName>
, 
<location id="41606049FFA8FFC506CF31E5FE1AF9A4" LSID="urn:lsid:plazi:treatment:CC168784FFA9FFC902E8306AFE47FA30:41606049FFA8FFC506CF31E5FE1AF9A4" country="Costa Rica" county="Their" latitude="9.69916" longLatPrecision="1" longitude="-84.3775" municipality="Area" name="Zona Protectora Cerro de La Cangreja" pageId="5" pageNumber="4" stateProvince="Guanacaste">Zona Protectora Cerro de La Cangreja</location>
, 
<specimenCount id="52B9FD1BFFA8FFC50397323BFDA5F9A4" box="[447,523,1537,1563]" pageId="5" pageNumber="4" type="generic">
<quantity id="83479B77FFA8FFC50397323BFDA6F9A4" box="[447,520,1537,1563]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="4" unit="m" value="600.0">
<elevation id="CF92D1A1FFA8FFC50397323BFDA6F9A4" box="[447,520,1537,1563]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="4" unit="m" value="600.0">600 m</elevation>
</quantity>
.
</specimenCount>
<geoCoordinate id="218B5055FFA8FFC5003A323BFD28F9A4" box="[530,646,1537,1563]" degrees="9.69916" direction="north" orientation="latitude" pageId="5" pageNumber="4" precision="1" value="9.69916">9.69916 N</geoCoordinate>
, − 
<geoCoordinate id="218B5055FFA8FFC500B7323BFC88F9A4" box="[671,806,1537,1564]" degrees="84.37750" direction="west" orientation="longitude" pageId="5" pageNumber="4" precision="1" value="-84.3775">84.37750 W</geoCoordinate>
. 
<collectingMethod id="9DFE4E85FFA8FFC50119323BFCDDF9A4" box="[817,883,1537,1563]" pageId="5" pageNumber="4">Pitfall</collectingMethod>
T. 
<date id="30011052FFA8FFC501BD323BFBB4F9A3" box="[917,1050,1537,1564]" pageId="5" pageNumber="4" value="2005-06-29">
<collectingDate id="2045E9BAFFA8FFC501BD323BFBB4F9A3" box="[917,1050,1537,1564]" pageId="5" pageNumber="4" value="2005-06-29">29 jun 2005</collectingDate>
</date>
, 
<collectorName id="E94A5344FFA8FFC5060D323BFB5AF9A4" box="[1061,1268,1537,1563]" pageId="5" pageNumber="4">Col. B. Hernández</collectorName>
” (11 unverified sex)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0CA56519FFA8FFC502E83276FEC1F936" pageId="5" pageNumber="4" type="distribution">
<paragraph id="44003692FFA8FFC502E83276FEC1F936" blockId="5.[192,1440,1612,1673]" pageId="5" pageNumber="4">
<emphasis id="76CBEA80FFA8FFC502E83276FEF4F9D9" bold="true" box="[192,346,1612,1638]" pageId="5" pageNumber="4">Distribution.</emphasis>
<collectingCountry id="3CA87602FFA8FFC5034B3277FE73F9D8" box="[355,477,1613,1639]" name="Costa Rica" pageId="5" pageNumber="4">Costa Rica</collectingCountry>
(
<collectingRegion id="867BF870FFA8FFC503C53277FDDCF9D8" box="[493,626,1613,1639]" country="Costa Rica" name="Guanacaste" pageId="5" pageNumber="4">Guanacaste</collectingRegion>
, 
<collectingRegion id="867BF870FFA8FFC500573277FD51F9D8" box="[639,767,1613,1639]" country="Costa Rica" name="Puntarenas" pageId="5" pageNumber="4">Puntarenas</collectingRegion>
, and 
<collectingRegion id="867BF870FFA8FFC501173277FC33F9D8" box="[831,925,1613,1639]" country="Costa Rica" name="San Jose" pageId="5" pageNumber="4">San José</collectingRegion>
provinces) and 
<collectingCountry id="3CA87602FFA8FFC5067F3277FB64F9D8" box="[1111,1226,1613,1639]" name="Nicaragua" pageId="5" pageNumber="4">Nicaragua</collectingCountry>
(
<collectingRegion id="867BF870FFA8FFC506F23277FF52F936" country="Nicaragua" name="Managua" pageId="5" pageNumber="4">Managua department</collectingRegion>
) (
<figureCitation id="DC842A17FFA8FFC5033A3254FECCF936" box="[274,354,1646,1673]" captionStart="Figure 19" captionStartId="11.[192,268,1866,1892]" captionTargetBox="[192,1440,1091,1837]" captionTargetId="figure-407@11.[192,1440,1091,1837]" captionTargetPageId="11" captionText="Figure 19. Known distribution of Dichotomius agenor, D. centralis, and D. woodruffi new species in Nicaragua, Costa Rica, and Panama." figureDoi="http://doi.org/10.5281/zenodo.6392015" httpUri="https://zenodo.org/record/6392015/files/figure.png" pageId="5" pageNumber="4">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0CA56519FFA8FFC502E832A3FB9FF949" pageId="5" pageNumber="4" type="etymology">
<paragraph id="44003692FFA8FFC502E832A3FB9FF949" blockId="5.[191,1440,1688,1783]" pageId="5" pageNumber="4">
<emphasis id="76CBEA80FFA8FFC502E832A3FEEAF90C" bold="true" box="[192,324,1689,1715]" pageId="5" pageNumber="4">Etymology.</emphasis>
<taxonomicName id="83BF4D11FFA8FFC5036132A2FDEDF90D" authority="Solís &amp; Kohlmann, 2022" authorityName="Solís &amp; Kohlmann" authorityYear="2022" box="[329,579,1688,1714]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov.">
<emphasis id="76CBEA80FFA8FFC5036132A2FDEDF90D" box="[329,579,1688,1714]" italics="true" pageId="5" pageNumber="4">Dichotomius woodruffi</emphasis>
</taxonomicName>
<emphasis id="76CBEA80FFA8FFC5006032A3FD60F90D" bold="true" box="[584,718,1688,1715]" pageId="5" pageNumber="4">
<taxonomicNameLabel id="6DF857FBFFA8FFC5006032A3FD60F90D" box="[584,718,1688,1715]" pageId="5" pageNumber="4" rank="species">new species</taxonomicNameLabel>
</emphasis>
is an eponym after the late Robert Woodruff, a scarabaeodologist, known for his work on 
<collectingRegion id="867BF870FFA8FFC503963280FDA1F96B" box="[446,527,1722,1748]" country="United States of America" name="Florida" pageId="5" pageNumber="4">Florida</collectingRegion>
<taxonomicName id="83BF4D11FFA8FFC5003C3280FD09F96B" authorityName="Latreille" authorityYear="1802" box="[532,679,1722,1748]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
, especially dung beetles and the melolonthine genus 
<taxonomicName id="83BF4D11FFA8FFC506C53280FADDF96B" authorityName="Harris" authorityYear="1826" box="[1261,1395,1722,1748]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="76CBEA80FFA8FFC506C53280FADDF96B" box="[1261,1395,1722,1748]" italics="true" pageId="5" pageNumber="4">Phyllophaga</emphasis>
</taxonomicName>
. He was also interested in collecting postage stamps with scarabs depicted on them.
</paragraph>
</subSubSection>
<subSubSection id="0CA56519FFA8FFC702E8333CFAFCFE23" lastPageId="7" lastPageNumber="6" pageId="5" pageNumber="4" type="diagnosis">
<paragraph id="44003692FFA8FFC702E8333CFAFCFE23" blockId="5.[191,1440,1798,1927]" lastBlockId="7.[192,1440,214,412]" lastPageId="7" lastPageNumber="6" pageId="5" pageNumber="4">
<emphasis id="76CBEA80FFA8FFC502E8333CFE92F89F" bold="true" box="[192,316,1798,1824]" pageId="5" pageNumber="4">Diagnosis.</emphasis>
<taxonomicName id="83BF4D11FFA8FFC5036A333CFD93F89F" authority="Solís &amp; Kohlmann, 2022" authorityName="Solís &amp; Kohlmann" authorityYear="2022" box="[322,573,1798,1824]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov.">
<emphasis id="76CBEA80FFA8FFC5036A333CFD93F89F" box="[322,573,1798,1824]" italics="true" pageId="5" pageNumber="4">Dichotomius woodruffi</emphasis>
</taxonomicName>
<emphasis id="76CBEA80FFA8FFC5006B333CFD65F89F" bold="true" box="[579,715,1798,1824]" pageId="5" pageNumber="4">
<taxonomicNameLabel id="6DF857FBFFA8FFC5006B333CFD65F89F" box="[579,715,1798,1824]" pageId="5" pageNumber="4" rank="species">new species</taxonomicNameLabel>
</emphasis>
is separated from other species in the 
<taxonomicName id="83BF4D11FFA8FFC5065B333DFB70F89F" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[1139,1246,1799,1825]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="agenor">
<emphasis id="76CBEA80FFA8FFC5065B333DFB29F89F" box="[1139,1159,1799,1824]" italics="true" pageId="5" pageNumber="4">D</emphasis>
. 
<emphasis id="76CBEA80FFA8FFC506BC333DFB70F89F" box="[1172,1246,1799,1824]" italics="true" pageId="5" pageNumber="4">agenor</emphasis>
</taxonomicName>
species-group by the following combination of characters: Male interocular surface with lightly shagreened microsculpture (small, lightly impressed punctures) (
<figureCitation id="DC842A17FFA8FFC500243370FDE5F8DB" box="[524,587,1866,1892]" captionStart="Figures 1–6" captionStartId="6.[193,278,1833,1859]" captionTargetBox="[229,1405,221,1803]" captionTargetId="figure-18@6.[229,1405,199,1806]" captionTargetPageId="6" captionText="Figures 1–6. Dichotomius woodruffi, new species. 1) Dorsal habitus, holotype, male. Scale bar equals 1 cm. 2) Dorsal habitus, allotype, female. Scale bar equals 1 cm. 3) Frontal view, holotype, male. 4) Frontal view, allotype, female. 5) Ventral clypeal process spiniform, holotype male. 6) Anterior pronotal border (frontal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.6392009" httpUri="https://zenodo.org/record/6392009/files/figure.png" pageId="5" pageNumber="4">Fig. 1</figureCitation>
). Anterior pronotal border medially sinuate in major males (
<figureCitation id="DC842A17FFA8FFC506D93370FA9FF8DA" box="[1265,1329,1866,1893]" captionStart="Figures 1–6" captionStartId="6.[193,278,1833,1859]" captionTargetBox="[229,1405,221,1803]" captionTargetId="figure-18@6.[229,1405,199,1806]" captionTargetPageId="6" captionText="Figures 1–6. Dichotomius woodruffi, new species. 1) Dorsal habitus, holotype, male. Scale bar equals 1 cm. 2) Dorsal habitus, allotype, female. Scale bar equals 1 cm. 3) Frontal view, holotype, male. 4) Frontal view, allotype, female. 5) Ventral clypeal process spiniform, holotype male. 6) Anterior pronotal border (frontal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.6392009" httpUri="https://zenodo.org/record/6392009/files/figure.png" pageId="5" pageNumber="4">Fig. 6</figureCitation>
); anterior pronotal declivity with a medial sulcus (
<figureCitation id="DC842A17FFA8FFC500513356FD17F838" box="[633,697,1900,1927]" captionStart="Figures 7–12" captionStartId="8.[192,279,1799,1825]" captionTargetBox="[248,1385,213,1769]" captionTargetId="figure-84@8.[248,1385,213,1769]" captionTargetPageId="8" captionText="Figures 7–12. Dichotomius woodruffi, new species, and D. centralis. 7–11) D. woodruffi. 7) Anterior pronotal sulcus (lateral-oblique view), holotype male. 8) Male pygidium twice as broad as long, slightly convex, holotype male. 9) Lateral view of the aedeagus, holotype male. 10) Ventral view of the aedeagus, holotype male. 11) Lamella copulatrix, holotype male. 12) D. centralis, elytral stria crenulate." figureDoi="http://doi.org/10.5281/zenodo.6392011" httpUri="https://zenodo.org/record/6392011/files/figure.png" pageId="5" pageNumber="4">Fig. 7</figureCitation>
); male median tubercle conical, without apical emargination, twice as high as lateral tubercle; lateral tubercles with acute apex (
<figureCitation id="DC842A17FFAAFFC7017134ECFC34FF4F" box="[857,922,214,240]" captionStart="Figures 1–6" captionStartId="6.[193,278,1833,1859]" captionTargetBox="[229,1405,221,1803]" captionTargetId="figure-18@6.[229,1405,199,1806]" captionTargetPageId="6" captionText="Figures 1–6. Dichotomius woodruffi, new species. 1) Dorsal habitus, holotype, male. Scale bar equals 1 cm. 2) Dorsal habitus, allotype, female. Scale bar equals 1 cm. 3) Frontal view, holotype, male. 4) Frontal view, allotype, female. 5) Ventral clypeal process spiniform, holotype male. 6) Anterior pronotal border (frontal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.6392009" httpUri="https://zenodo.org/record/6392009/files/figure.png" pageId="7" pageNumber="6">Fig. 3</figureCitation>
). Female fronto-clypeal carina quadrituberculate (
<figureCitation id="DC842A17FFAAFFC702DE34C2FE96FEAD" box="[246,312,248,274]" captionStart="Figures 1–6" captionStartId="6.[193,278,1833,1859]" captionTargetBox="[229,1405,221,1803]" captionTargetId="figure-18@6.[229,1405,199,1806]" captionTargetPageId="6" captionText="Figures 1–6. Dichotomius woodruffi, new species. 1) Dorsal habitus, holotype, male. Scale bar equals 1 cm. 2) Dorsal habitus, allotype, female. Scale bar equals 1 cm. 3) Frontal view, holotype, male. 4) Frontal view, allotype, female. 5) Ventral clypeal process spiniform, holotype male. 6) Anterior pronotal border (frontal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.6392009" httpUri="https://zenodo.org/record/6392009/files/figure.png" pageId="7" pageNumber="6">Fig. 4</figureCitation>
). Ventral clypeal process spiniform (
<figureCitation id="DC842A17FFAAFFC700F934C2FCBDFEAC" box="[721,787,248,275]" captionStart="Figures 1–6" captionStartId="6.[193,278,1833,1859]" captionTargetBox="[229,1405,221,1803]" captionTargetId="figure-18@6.[229,1405,199,1806]" captionTargetPageId="6" captionText="Figures 1–6. Dichotomius woodruffi, new species. 1) Dorsal habitus, holotype, male. Scale bar equals 1 cm. 2) Dorsal habitus, allotype, female. Scale bar equals 1 cm. 3) Frontal view, holotype, male. 4) Frontal view, allotype, female. 5) Ventral clypeal process spiniform, holotype male. 6) Anterior pronotal border (frontal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.6392009" httpUri="https://zenodo.org/record/6392009/files/figure.png" pageId="7" pageNumber="6">Fig. 5</figureCitation>
). Row of dense uninterrupted ocellate punctures arranged along anterior and posterior margin of pronotum (
<figureCitation id="DC842A17FFAAFFC700DF3520FC97FE8A" box="[759,825,282,309]" captionStart="Figures 1–6" captionStartId="6.[193,278,1833,1859]" captionTargetBox="[229,1405,221,1803]" captionTargetId="figure-18@6.[229,1405,199,1806]" captionTargetPageId="6" captionText="Figures 1–6. Dichotomius woodruffi, new species. 1) Dorsal habitus, holotype, male. Scale bar equals 1 cm. 2) Dorsal habitus, allotype, female. Scale bar equals 1 cm. 3) Frontal view, holotype, male. 4) Frontal view, allotype, female. 5) Ventral clypeal process spiniform, holotype male. 6) Anterior pronotal border (frontal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.6392009" httpUri="https://zenodo.org/record/6392009/files/figure.png" pageId="7" pageNumber="6">Fig. 6</figureCitation>
). Interstriae with surface shagreened (
<figureCitation id="DC842A17FFAAFFC706CC3520FAECFE8B" box="[1252,1346,282,308]" captionStart="Figures 1–6" captionStartId="6.[193,278,1833,1859]" captionTargetBox="[229,1405,221,1803]" captionTargetId="figure-18@6.[229,1405,199,1806]" captionTargetPageId="6" captionText="Figures 1–6. Dichotomius woodruffi, new species. 1) Dorsal habitus, holotype, male. Scale bar equals 1 cm. 2) Dorsal habitus, allotype, female. Scale bar equals 1 cm. 3) Frontal view, holotype, male. 4) Frontal view, allotype, female. 5) Ventral clypeal process spiniform, holotype male. 6) Anterior pronotal border (frontal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.6392009" httpUri="https://zenodo.org/record/6392009/files/figure.png" pageId="7" pageNumber="6">Fig. 1–2</figureCitation>
). Pygidium twice as wide as long (
<figureCitation id="DC842A17FFAAFFC700283506FDEAFEE8" box="[512,580,316,343]" captionStart="Figures 7–12" captionStartId="8.[192,279,1799,1825]" captionTargetBox="[248,1385,213,1769]" captionTargetId="figure-84@8.[248,1385,213,1769]" captionTargetPageId="8" captionText="Figures 7–12. Dichotomius woodruffi, new species, and D. centralis. 7–11) D. woodruffi. 7) Anterior pronotal sulcus (lateral-oblique view), holotype male. 8) Male pygidium twice as broad as long, slightly convex, holotype male. 9) Lateral view of the aedeagus, holotype male. 10) Ventral view of the aedeagus, holotype male. 11) Lamella copulatrix, holotype male. 12) D. centralis, elytral stria crenulate." figureDoi="http://doi.org/10.5281/zenodo.6392011" httpUri="https://zenodo.org/record/6392011/files/figure.png" pageId="7" pageNumber="6">Fig. 8</figureCitation>
). Setigerous lateral areas on the anterior and lateral lobes of metasternum interrupted at the posteromedial mesocoxal margins. Ventral surface of profemur covered with well-impressed punctures, setigerous setae near the medial edge. Ventral surface of metafemur without setose punctures.
</paragraph>
</subSubSection>
<caption id="10C0661AFFABFFC602E93313FAD8F839" ID-DOI="http://doi.org/10.5281/zenodo.6392009" ID-Zenodo-Dep="6392009" httpUri="https://zenodo.org/record/6392009/files/figure.png" pageId="6" pageNumber="5" startId="6.[193,278,1833,1859]" targetBox="[229,1405,221,1803]" targetPageId="6">
<paragraph id="44003692FFABFFC602E93313FAD8F839" blockId="6.[192,1442,1832,1927]" pageId="6" pageNumber="5">
<emphasis id="76CBEA80FFABFFC602E93313FEE3F8FC" bold="true" box="[193,333,1833,1859]" pageId="6" pageNumber="5">Figures 1–6.</emphasis>
<taxonomicName id="83BF4D11FFABFFC6037A3313FDE6F8FC" authority="Solís &amp; Kohlmann, 2022" authorityName="Solís &amp; Kohlmann" authorityYear="2022" box="[338,584,1833,1859]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov.">
<emphasis id="76CBEA80FFABFFC6037A3313FDE6F8FC" box="[338,584,1833,1859]" italics="true" pageId="6" pageNumber="5">Dichotomius woodruffi</emphasis>
</taxonomicName>
, 
<emphasis id="76CBEA80FFABFFC6007C3313FD77F8FC" bold="true" box="[596,729,1833,1859]" pageId="6" pageNumber="5">
<taxonomicNameLabel id="6DF857FBFFABFFC6007C3313FD77F8FC" box="[596,729,1833,1859]" pageId="6" pageNumber="5" rank="species">new species</taxonomicNameLabel>
</emphasis>
. 
<emphasis id="76CBEA80FFABFFC600CD3312FD53F8FD" bold="true" box="[741,765,1832,1858]" pageId="6" pageNumber="5">1)</emphasis>
Dorsal habitus, holotype, male. Scale bar equals 1 cm. 
<emphasis id="76CBEA80FFABFFC607663312FACBF8FD" bold="true" box="[1358,1381,1832,1858]" pageId="6" pageNumber="5">2)</emphasis>
Dorsal habitus, allotype, female. Scale bar equals 1 cm. 
<emphasis id="76CBEA80FFABFFC600CE3370FD50F8DB" bold="true" box="[742,766,1866,1892]" pageId="6" pageNumber="5">3)</emphasis>
Frontal view, holotype, male. 
<emphasis id="76CBEA80FFABFFC606693370FBF7F8DB" bold="true" box="[1089,1113,1866,1892]" pageId="6" pageNumber="5">4)</emphasis>
Frontal view, allotype, female. 
<emphasis id="76CBEA80FFABFFC602E83356FF76F839" bold="true" box="[192,216,1900,1926]" pageId="6" pageNumber="5">5)</emphasis>
Ventral clypeal process spiniform, holotype male. 
<emphasis id="76CBEA80FFABFFC600D53356FCBBF839" bold="true" box="[765,789,1900,1926]" pageId="6" pageNumber="5">6)</emphasis>
Anterior pronotal border (frontal view), holotype male.
</paragraph>
</caption>
<subSubSection id="0CA56519FFAAFFC902E83591FB27FD9C" lastPageId="9" lastPageNumber="8" pageId="7" pageNumber="6" type="description">
<paragraph id="44003692FFAAFFC702E83591FA27FE7A" blockId="7.[190,1442,427,1927]" box="[192,1417,427,454]" pageId="7" pageNumber="6">
<emphasis id="76CBEA80FFAAFFC702E83591FDA5FE7A" bold="true" box="[192,523,427,453]" pageId="7" pageNumber="6">
Description. 
<typeStatus id="9B048830FFAAFFC703703591FE6BFE7A" box="[344,453,427,453]" pageId="7" pageNumber="6" type="holotype">Holotype</typeStatus>
male.
</emphasis>
Length 23.0 mm. Width 
<quantity id="83479B77FFAAFFC7010C3591FC24FE79" box="[804,906,427,454]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="7" pageNumber="6" unit="mm" value="13.5">13.5 mm</quantity>
. Dorsal surface black and shagreened (
<figureCitation id="DC842A17FFAAFFC707123591FAD7FE7A" box="[1338,1401,427,453]" captionStart="Figures 1–6" captionStartId="6.[193,278,1833,1859]" captionTargetBox="[229,1405,221,1803]" captionTargetId="figure-18@6.[229,1405,199,1806]" captionTargetPageId="6" captionText="Figures 1–6. Dichotomius woodruffi, new species. 1) Dorsal habitus, holotype, male. Scale bar equals 1 cm. 2) Dorsal habitus, allotype, female. Scale bar equals 1 cm. 3) Frontal view, holotype, male. 4) Frontal view, allotype, female. 5) Ventral clypeal process spiniform, holotype male. 6) Anterior pronotal border (frontal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.6392009" httpUri="https://zenodo.org/record/6392009/files/figure.png" pageId="7" pageNumber="6">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="44003692FFAAFFC7032835F7FB8BFD67" blockId="7.[190,1442,427,1927]" pageId="7" pageNumber="6">
Head and pronotum shiny, elytra and pygidium dull. Head with the anterior dorsal surface crosswise rugose, rugosity irregular. Transverse tubercle wide and flattened fronto-posteriorly (
<figureCitation id="DC842A17FFAAFFC7063835CAFBFEFDB5" box="[1040,1104,496,522]" captionStart="Figures 1–6" captionStartId="6.[193,278,1833,1859]" captionTargetBox="[229,1405,221,1803]" captionTargetId="figure-18@6.[229,1405,199,1806]" captionTargetPageId="6" captionText="Figures 1–6. Dichotomius woodruffi, new species. 1) Dorsal habitus, holotype, male. Scale bar equals 1 cm. 2) Dorsal habitus, allotype, female. Scale bar equals 1 cm. 3) Frontal view, holotype, male. 4) Frontal view, allotype, female. 5) Ventral clypeal process spiniform, holotype male. 6) Anterior pronotal border (frontal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.6392009" httpUri="https://zenodo.org/record/6392009/files/figure.png" pageId="7" pageNumber="6">Fig. 3</figureCitation>
), with a simple conical central process or horn and two lateral carinae whose profile in frontal view is acute at the lateral angles and sinuous in the upper part (
<figureCitation id="DC842A17FFAAFFC70358360EFE1CFDF1" box="[368,434,564,590]" captionStart="Figures 1–6" captionStartId="6.[193,278,1833,1859]" captionTargetBox="[229,1405,221,1803]" captionTargetId="figure-18@6.[229,1405,199,1806]" captionTargetPageId="6" captionText="Figures 1–6. Dichotomius woodruffi, new species. 1) Dorsal habitus, holotype, male. Scale bar equals 1 cm. 2) Dorsal habitus, allotype, female. Scale bar equals 1 cm. 3) Frontal view, holotype, male. 4) Frontal view, allotype, female. 5) Ventral clypeal process spiniform, holotype male. 6) Anterior pronotal border (frontal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.6392009" httpUri="https://zenodo.org/record/6392009/files/figure.png" pageId="7" pageNumber="6">Fig. 3</figureCitation>
). Punctures decrease in size and depth as one advances from the clypeus up the slope and the upper part of the transverse forehead, where they almost disappear. Vertical posterior area of the transverse tubercle shows punctures in the form of small transverse lines. Horizontal posterior area of the head shows small transverse punctures in the lateral area, central area devoid of punctures. Anterior clypeal border with two teeth separated by an obtuse indentation. Ventral clypeal process spiniform (
<figureCitation id="DC842A17FFAAFFC701FD3687FBB8FD67" box="[981,1046,701,728]" captionStart="Figures 1–6" captionStartId="6.[193,278,1833,1859]" captionTargetBox="[229,1405,221,1803]" captionTargetId="figure-18@6.[229,1405,199,1806]" captionTargetPageId="6" captionText="Figures 1–6. Dichotomius woodruffi, new species. 1) Dorsal habitus, holotype, male. Scale bar equals 1 cm. 2) Dorsal habitus, allotype, female. Scale bar equals 1 cm. 3) Frontal view, holotype, male. 4) Frontal view, allotype, female. 5) Ventral clypeal process spiniform, holotype male. 6) Anterior pronotal border (frontal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.6392009" httpUri="https://zenodo.org/record/6392009/files/figure.png" pageId="7" pageNumber="6">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="44003692FFAAFFC7032836DAFCCBFC3C" blockId="7.[190,1442,427,1927]" pageId="7" pageNumber="6">
Pronotum wider than long with a pronounced slope in the anterior area and with a slight central concavity, a slight elevation on each side and another slight bilobed elevation in the upper part of the concavity. The surface of the pronotum possesses fine, small, simple punctures in the central disc area than in the lateral area. Punctures near the anterior and posterior margin are ocellate. Anterior pronotal border medially sinuate (
<figureCitation id="DC842A17FFAAFFC706C6377CFA9EFCDE" box="[1262,1328,838,865]" captionStart="Figures 1–6" captionStartId="6.[193,278,1833,1859]" captionTargetBox="[229,1405,221,1803]" captionTargetId="figure-18@6.[229,1405,199,1806]" captionTargetPageId="6" captionText="Figures 1–6. Dichotomius woodruffi, new species. 1) Dorsal habitus, holotype, male. Scale bar equals 1 cm. 2) Dorsal habitus, allotype, female. Scale bar equals 1 cm. 3) Frontal view, holotype, male. 4) Frontal view, allotype, female. 5) Ventral clypeal process spiniform, holotype male. 6) Anterior pronotal border (frontal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.6392009" httpUri="https://zenodo.org/record/6392009/files/figure.png" pageId="7" pageNumber="6">Fig. 6</figureCitation>
); anterior pronotal declivity with a fine sulcus in its upper part (
<figureCitation id="DC842A17FFAAFFC7013D3752FCF9FC3C" box="[789,855,872,899]" captionStart="Figures 7–12" captionStartId="8.[192,279,1799,1825]" captionTargetBox="[248,1385,213,1769]" captionTargetId="figure-84@8.[248,1385,213,1769]" captionTargetPageId="8" captionText="Figures 7–12. Dichotomius woodruffi, new species, and D. centralis. 7–11) D. woodruffi. 7) Anterior pronotal sulcus (lateral-oblique view), holotype male. 8) Male pygidium twice as broad as long, slightly convex, holotype male. 9) Lateral view of the aedeagus, holotype male. 10) Ventral view of the aedeagus, holotype male. 11) Lamella copulatrix, holotype male. 12) D. centralis, elytral stria crenulate." figureDoi="http://doi.org/10.5281/zenodo.6392011" httpUri="https://zenodo.org/record/6392011/files/figure.png" pageId="7" pageNumber="6">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="44003692FFAAFFC7032837B1FA91FC1A" blockId="7.[190,1442,427,1927]" box="[256,1343,907,933]" pageId="7" pageNumber="6">Elytral striae with ocellate punctures. Interstriae with very small, almost imperceptible punctures.</paragraph>
<paragraph id="44003692FFAAFFC703283797FBAFFC56" blockId="7.[190,1442,427,1927]" pageId="7" pageNumber="6">
Pygidium twice as wide as long (
<figureCitation id="DC842A17FFAAFFC7004E3797FD08FC78" box="[614,678,941,967]" captionStart="Figures 7–12" captionStartId="8.[192,279,1799,1825]" captionTargetBox="[248,1385,213,1769]" captionTargetId="figure-84@8.[248,1385,213,1769]" captionTargetPageId="8" captionText="Figures 7–12. Dichotomius woodruffi, new species, and D. centralis. 7–11) D. woodruffi. 7) Anterior pronotal sulcus (lateral-oblique view), holotype male. 8) Male pygidium twice as broad as long, slightly convex, holotype male. 9) Lateral view of the aedeagus, holotype male. 10) Ventral view of the aedeagus, holotype male. 11) Lamella copulatrix, holotype male. 12) D. centralis, elytral stria crenulate." figureDoi="http://doi.org/10.5281/zenodo.6392011" httpUri="https://zenodo.org/record/6392011/files/figure.png" pageId="7" pageNumber="6">Fig. 8</figureCitation>
); with very small punctures throughout most of its surface except for a row of strongly impressed ocellate punctures bordering the basal margin.
</paragraph>
<paragraph id="44003692FFAAFFC7032837C8FCEBFBCC" blockId="7.[190,1442,427,1927]" pageId="7" pageNumber="6">
<emphasis id="76CBEA80FFAAFFC7032837C8FE3BFBB3" bold="true" box="[256,405,1010,1036]" pageId="7" pageNumber="6">Ventral area.</emphasis>
The entire ventral surface, as well as the dorsal surface, with shagreened micro-sculpture. Area of the hypomeron with the anterior and posterior third covered with setigerous punctures strongly impressed with long setae, central area smooth without setigerous punctures except for one, two or three rows of setigerous punctures near the lateral border (4 or 5 rows in 
<taxonomicName id="83BF4D11FFAAFFC700F63063FC9BFBCD" authorityName="Montoya-Molina and Vaz-de-Mello" authorityYear="2021" box="[734,821,1113,1139]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="enioi">
<emphasis id="76CBEA80FFAAFFC700F63063FD5CFBCD" box="[734,754,1113,1138]" italics="true" pageId="7" pageNumber="6">D</emphasis>
. 
<emphasis id="76CBEA80FFAAFFC700D73063FC9BFBCD" box="[767,821,1113,1138]" italics="true" pageId="7" pageNumber="6">enioi</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="44003692FFAAFFC703283040FC18FAA1" blockId="7.[190,1442,427,1927]" pageId="7" pageNumber="6">Prosternal surface lacking obvious punctures. Mesosternum with setaceous, ocellate punctures; setae minute. Mesepimeron and metepisternum densely covered with setigerous, ocellate punctures. Metasternum with lateral lobes covered with setigerous punctures, anterior central lobe with ocellate setigerous punctures strongly impressed in the anterolateral areas. Central and posterior area of metasternum, including the area adjacent to the posterior border of the mesocoxa, without setigerous punctures.</paragraph>
<paragraph id="44003692FFAAFFC70328311CFEBEFADD" blockId="7.[190,1442,427,1927]" pageId="7" pageNumber="6">Abdominal ventrites with one (more frequently two) row of ocellate punctures in the area near the anterior border.</paragraph>
<paragraph id="44003692FFAAFFC703283150FB4DFA76" blockId="7.[190,1442,427,1927]" pageId="7" pageNumber="6">Ventral surface of profemora with small and shallow punctures medially (surface appearing rugose), with large setigerous punctures very strongly impressed in the posterior distal region. Ventral area of the meso- and metafemora smooth, with minute punctures (almost invisible) except in the most distal region.</paragraph>
<paragraph id="44003692FFAAFFC7032831EBFED1F9B2" blockId="7.[190,1442,427,1927]" pageId="7" pageNumber="6">Posterior coxa with a row of setigerous annular punctures along the entire length and contiguous to the posterior border.</paragraph>
<paragraph id="44003692FFAAFFC70328322CFCE5F98F" blockId="7.[190,1442,427,1927]" box="[256,843,1557,1584]" pageId="7" pageNumber="6">
Parameres and lamella copulatrix as in 
<figureCitation id="DC842A17FFAAFFC7009D322FFCE9F98F" box="[693,839,1557,1584]" captionStart="Figures 7–12" captionStartId="8.[192,279,1799,1825]" captionTargetBox="[248,1385,213,1769]" captionTargetId="figure-84@8.[248,1385,213,1769]" captionTargetPageId="8" captionText="Figures 7–12. Dichotomius woodruffi, new species, and D. centralis. 7–11) D. woodruffi. 7) Anterior pronotal sulcus (lateral-oblique view), holotype male. 8) Male pygidium twice as broad as long, slightly convex, holotype male. 9) Lateral view of the aedeagus, holotype male. 10) Ventral view of the aedeagus, holotype male. 11) Lamella copulatrix, holotype male. 12) D. centralis, elytral stria crenulate." figureDoi="http://doi.org/10.5281/zenodo.6392011" httpUri="https://zenodo.org/record/6392011/files/figure.png" pageId="7" pageNumber="6">Figures 9–11</figureCitation>
.
</paragraph>
<paragraph id="44003692FFAAFFC70328320DFC07F964" blockId="7.[190,1442,427,1927]" pageId="7" pageNumber="6">
<emphasis id="76CBEA80FFAAFFC70328320DFE11F9EE" bold="true" box="[256,447,1591,1617]" pageId="7" pageNumber="6">
<typeStatus id="9B048830FFAAFFC70328320DFECBF9EE" box="[256,357,1591,1617]" pageId="7" pageNumber="6" type="allotype">Allotype</typeStatus>
female.
</emphasis>
Length 19.0 mm. Width 13.0 mm. As male except for the following characters: Head with the entire dorsal surface crosswise rugose, the rugosity is irregular and most strongly impressed on the clypeus (
<figureCitation id="DC842A17FFAAFFC702E13246FEA3F929" box="[201,269,1660,1686]" captionStart="Figures 1–6" captionStartId="6.[193,278,1833,1859]" captionTargetBox="[229,1405,221,1803]" captionTargetId="figure-18@6.[229,1405,199,1806]" captionTargetPageId="6" captionText="Figures 1–6. Dichotomius woodruffi, new species. 1) Dorsal habitus, holotype, male. Scale bar equals 1 cm. 2) Dorsal habitus, allotype, female. Scale bar equals 1 cm. 3) Frontal view, holotype, male. 4) Frontal view, allotype, female. 5) Ventral clypeal process spiniform, holotype male. 6) Anterior pronotal border (frontal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.6392009" httpUri="https://zenodo.org/record/6392009/files/figure.png" pageId="7" pageNumber="6">Fig. 2</figureCitation>
). Transverse, quadrate elevation in the surface of the forehead. Two central tubercles a little higher and connected to each other by a small, curved carina forming a concavity posteriorly (
<figureCitation id="DC842A17FFAAFFC7067532A4FB31F906" box="[1117,1183,1694,1721]" captionStart="Figures 1–6" captionStartId="6.[193,278,1833,1859]" captionTargetBox="[229,1405,221,1803]" captionTargetId="figure-18@6.[229,1405,199,1806]" captionTargetPageId="6" captionText="Figures 1–6. Dichotomius woodruffi, new species. 1) Dorsal habitus, holotype, male. Scale bar equals 1 cm. 2) Dorsal habitus, allotype, female. Scale bar equals 1 cm. 3) Frontal view, holotype, male. 4) Frontal view, allotype, female. 5) Ventral clypeal process spiniform, holotype male. 6) Anterior pronotal border (frontal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.6392009" httpUri="https://zenodo.org/record/6392009/files/figure.png" pageId="7" pageNumber="6">Fig. 4</figureCitation>
). Less developed males are like females in the characteristics of the dorsal area of the head.
</paragraph>
<paragraph id="44003692FFAAFFC7032832DEFDF9F8FE" blockId="7.[190,1442,427,1927]" pageId="7" pageNumber="6">
Pronotum without declivity as in the 
<typeStatus id="9B048830FFAAFFC7008332D9FCA3F942" box="[683,781,1763,1789]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
but with a slight elevation in the anterior discal area, like intermediately developed males and nearly absent in less-developed females. Punctures always more strongly impressed in less developed females.
</paragraph>
<paragraph id="44003692FFAAFFC903283373FC90FEE9" blockId="7.[190,1442,427,1927]" lastBlockId="9.[192,1442,214,547]" lastPageId="9" lastPageNumber="8" pageId="7" pageNumber="6">
<emphasis id="76CBEA80FFAAFFC703283373FD1DF8DC" bold="true" box="[256,691,1865,1891]" pageId="7" pageNumber="6">
Morphological variation (
<typeStatus id="9B048830FFAAFFC700063373FD09F8DC" box="[558,679,1865,1891]" pageId="7" pageNumber="6" type="paratype">paratypes</typeStatus>
).
</emphasis>
Length 17.0–23.0 mm. Width 
<quantity id="83479B77FFAAFFC706263370FB1DF8DB" box="[1038,1203,1866,1893]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.35" metricValueMin="1.05" pageId="7" pageNumber="6" unit="mm" value="12.0" valueMax="13.5" valueMin="10.5">10.5–13.5 mm</quantity>
. Anterior edge of the pronotum of males varies from very wide (majors) to less wide (minors). The anterior declivity of the pronotum is wider in major males and decreases in amplitude until it is lacking in intermediate males (like females). Pronotum may vary from finely to coarsely punctate; females have coarser punctures. Minor males have the anterior pronotal border parallel and not sinuate. Minor males resemble females, especially regarding the cephalic carina. Male forms vary from minors to intermediates to majors.
</paragraph>
<caption id="10C0661AFFA5FFC802E8333DFC3FF839" ID-DOI="http://doi.org/10.5281/zenodo.6392011" ID-Zenodo-Dep="6392011" httpUri="https://zenodo.org/record/6392011/files/figure.png" pageId="8" pageNumber="7" startId="8.[192,279,1799,1825]" targetBox="[248,1385,213,1769]" targetPageId="8">
<paragraph id="44003692FFA5FFC802E8333DFC3FF839" blockId="8.[191,1440,1799,1927]" pageId="8" pageNumber="7">
<emphasis id="76CBEA80FFA5FFC802E8333DFEF1F89E" bold="true" box="[192,351,1799,1825]" pageId="8" pageNumber="7">Figures 7–12.</emphasis>
<taxonomicName id="83BF4D11FFA5FFC80340333DFDC8F89E" authority="Solís &amp; Kohlmann, 2022" authorityName="Solís &amp; Kohlmann" authorityYear="2022" box="[360,614,1799,1825]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov.">
<emphasis id="76CBEA80FFA5FFC80340333DFDC8F89E" box="[360,614,1799,1825]" italics="true" pageId="8" pageNumber="7">Dichotomius woodruffi</emphasis>
</taxonomicName>
, 
<emphasis id="76CBEA80FFA5FFC8005D333DFD51F89E" bold="true" box="[629,767,1799,1825]" pageId="8" pageNumber="7">
<taxonomicNameLabel id="6DF857FBFFA5FFC8005D333DFD51F89E" box="[629,767,1799,1825]" pageId="8" pageNumber="7" rank="species">new species</taxonomicNameLabel>
</emphasis>
, and 
<taxonomicName id="83BF4D11FFA5FFC8016A3332FC6EF89E" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[834,960,1799,1826]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="centralis">
<emphasis id="76CBEA80FFA5FFC8016A3332FCF8F89E" box="[834,854,1800,1825]" italics="true" pageId="8" pageNumber="7">D</emphasis>
. 
<emphasis id="76CBEA80FFA5FFC8014D333DFC6EF89E" box="[869,960,1799,1825]" italics="true" pageId="8" pageNumber="7">centralis</emphasis>
</taxonomicName>
. 
<emphasis id="76CBEA80FFA5FFC801E7333DFBBFF89E" bold="true" box="[975,1041,1799,1825]" pageId="8" pageNumber="7">7–11)</emphasis>
<taxonomicName id="83BF4D11FFA5FFC806333332FB09F89E" authorityName="Solís &amp; Kohlmann" authorityYear="2022" box="[1051,1191,1799,1825]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="woodruffi">
<emphasis id="76CBEA80FFA5FFC806333332FB09F89E" box="[1051,1191,1799,1825]" italics="true" pageId="8" pageNumber="7">D. woodruffi</emphasis>
</taxonomicName>
. 
<emphasis id="76CBEA80FFA5FFC8069E333DFB61F89E" bold="true" box="[1206,1231,1799,1825]" pageId="8" pageNumber="7">7)</emphasis>
Anterior pronotal sulcus (lateral-oblique view), holotype male. 
<emphasis id="76CBEA80FFA5FFC8009F3312FD61F8FD" bold="true" box="[695,719,1832,1858]" pageId="8" pageNumber="7">8)</emphasis>
Male pygidium twice as broad as long, slightly convex, holotype male. 
<emphasis id="76CBEA80FFA5FFC803293370FEB7F8DB" bold="true" box="[257,281,1866,1892]" pageId="8" pageNumber="7">9)</emphasis>
Lateral view of the aedeagus, holotype male. 
<emphasis id="76CBEA80FFA5FFC8012D3370FC84F8DB" bold="true" box="[773,810,1866,1892]" pageId="8" pageNumber="7">10)</emphasis>
Ventral view of the aedeagus, holotype male. 
<emphasis id="76CBEA80FFA5FFC807343370FAECF8DB" bold="true" box="[1308,1346,1866,1892]" pageId="8" pageNumber="7">11)</emphasis>
Lamella copulatrix, holotype male. 
<emphasis id="76CBEA80FFA5FFC803C23356FDBEF839" bold="true" box="[490,528,1900,1926]" pageId="8" pageNumber="7">12)</emphasis>
<taxonomicName id="83BF4D11FFA5FFC8003E3357FD3CF839" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[534,658,1900,1927]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="centralis">
<emphasis id="76CBEA80FFA5FFC8003E3357FD84F839" box="[534,554,1901,1926]" italics="true" pageId="8" pageNumber="7">D</emphasis>
. 
<emphasis id="76CBEA80FFA5FFC8001F3356FD3CF839" box="[567,658,1900,1926]" italics="true" pageId="8" pageNumber="7">centralis</emphasis>
</taxonomicName>
, elytral stria crenulate.
</paragraph>
</caption>
<paragraph id="44003692FFA4FFC903283564FB27FD9C" blockId="9.[192,1442,214,547]" pageId="9" pageNumber="8">The hypomeron in the central area may lack punctures or possess a few ring-like punctures without setae. Intermediately developed males with apex of the central process bituberculate and in less developed males the transverse elevation of the forehead like that of females. Ventral profemur surface varies from finely to coarsely punctate; females tend to possess coarser punctures. On its left side, one very well-developed female specimen was found to have setigerous punctures of the medial and lateral edges of the metasternum continuous at the inner mesocoxal area, but not on its right-side specimen possessed a convex pygidium.</paragraph>
</subSubSection>
<subSubSection id="0CA56519FFA4FFC902E83609FE47FA30" pageId="9" pageNumber="8" type="biology_ecology">
<paragraph id="44003692FFA4FFC902E83609FC37FD2E" blockId="9.[192,1442,563,658]" pageId="9" pageNumber="8">
<emphasis id="76CBEA80FFA4FFC902E83609FE8DFDF2" bold="true" box="[192,291,563,589]" pageId="9" pageNumber="8">Ecology.</emphasis>
We collected this species in traps baited with human, pig, and horse dung. It is distributed in humid forests and piedmont forest areas along the Pacific slope in 
<collectingCountry id="3CA87602FFA4FFC90172366FFC7DFDD0" box="[858,979,597,623]" name="Costa Rica" pageId="9" pageNumber="8">Costa Rica</collectingCountry>
and southern 
<collectingCountry id="3CA87602FFA4FFC9065E366FFB42FDD0" box="[1142,1260,597,623]" name="Nicaragua" pageId="9" pageNumber="8">Nicaragua</collectingCountry>
. The species has been collected from April to November, from 
<quantity id="83479B77FFA4FFC900E8364DFC99FD2D" box="[704,823,631,658]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="8.0" metricValueMin="0.0" pageId="9" pageNumber="8" unit="m" value="400.0" valueMax="800.0" valueMin="0.0">0 to 800 m</quantity>
altitude.
</paragraph>
<paragraph id="44003692FFA4FFC902E8369BFC9FFCD9" blockId="9.[191,1441,673,870]" pageId="9" pageNumber="8">
<emphasis id="76CBEA80FFA4FFC902E8369BFEECFD04" bold="true" box="[192,322,673,699]" pageId="9" pageNumber="8">Chorology.</emphasis>
The known southern distribution limit of 
<taxonomicName id="83BF4D11FFA4FFC901353698FC06FD04" authority="Solís &amp; Kohlmann, 2022" authorityName="Solís &amp; Kohlmann" authorityYear="2022" box="[797,936,673,700]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov.">
<emphasis id="76CBEA80FFA4FFC901353698FC9EFD04" box="[797,816,674,699]" italics="true" pageId="9" pageNumber="8">D</emphasis>
. 
<emphasis id="76CBEA80FFA4FFC90116369BFC06FD04" box="[830,936,673,699]" italics="true" pageId="9" pageNumber="8">woodruffi</emphasis>
</taxonomicName>
<emphasis id="76CBEA80FFA4FFC90186369BFB98FD04" bold="true" box="[942,1078,673,699]" pageId="9" pageNumber="8">
<taxonomicNameLabel id="6DF857FBFFA4FFC90186369BFB98FD04" box="[942,1078,673,699]" pageId="9" pageNumber="8" rank="species">new species</taxonomicNameLabel>
</emphasis>
is in 
<collectingCountry id="3CA87602FFA4FFC9065B369BFB45FD04" box="[1139,1259,673,699]" name="Costa Rica" pageId="9" pageNumber="8">Costa Rica</collectingCountry>
near the border with 
<collectingCountry id="3CA87602FFA4FFC902D436FEFEF8FD61" box="[252,342,708,734]" name="Panama" pageId="9" pageNumber="8">Panama</collectingCountry>
(
<geoCoordinate id="218B5055FFA4FFC9034F36F9FE75FD61" box="[359,475,707,734]" degrees="8.9520000" direction="north" orientation="latitude" pageId="9" pageNumber="8" precision="1" value="8.952">8.9520000</geoCoordinate>
<geoCoordinate id="218B5055FFA4FFC903CD36F9FDD4FD61" box="[485,634,707,734]" degrees="83.0345000" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="1" value="-83.0345">−83.0345000</geoCoordinate>
), and the known northernmost distribution limit is at El Bajo Private Reserve near 
<collectingRegion id="867BF870FFA4FFC9037036DCFE6DFCBF" box="[344,451,742,768]" country="Nicaragua" name="Managua" pageId="9" pageNumber="8">Managua</collectingRegion>
, 
<collectingCountry id="3CA87602FFA4FFC903E536DFFDEEFD40" box="[461,576,741,767]" name="Nicaragua" pageId="9" pageNumber="8">Nicaragua</collectingCountry>
(
<geoCoordinate id="218B5055FFA4FFC9006636DFFD6CFD40" box="[590,706,741,767]" degrees="11.991928" direction="north" orientation="latitude" pageId="9" pageNumber="8" precision="1" value="11.991928">11.991928</geoCoordinate>
<geoCoordinate id="218B5055FFA4FFC900E236DFFCFFFCBF" box="[714,849,741,768]" degrees="86.322666" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="1" value="-86.32266">−86.322666</geoCoordinate>
). It is an uncommon species in the northern portion of its range, whereas it is prevalent in the southern portion and is allopatric with 
<taxonomicName id="83BF4D11FFA4FFC9061A3732FB03FC9E" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[1074,1197,775,802]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="centralis">
<emphasis id="76CBEA80FFA4FFC9061A3732FBE8FC9E" box="[1074,1094,776,801]" italics="true" pageId="9" pageNumber="8">D</emphasis>
. 
<emphasis id="76CBEA80FFA4FFC9067A373DFB03FC9E" box="[1106,1197,775,801]" italics="true" pageId="9" pageNumber="8">centralis</emphasis>
</taxonomicName>
. The known northern range limit for 
<taxonomicName id="83BF4D11FFA4FFC9034B3710FE60FCFC" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[355,462,810,836]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="agenor">
<emphasis id="76CBEA80FFA4FFC9034B3710FED9FCFC" box="[355,375,810,835]" italics="true" pageId="9" pageNumber="8">D</emphasis>
. 
<emphasis id="76CBEA80FFA4FFC903AC3710FE60FCFC" box="[388,462,810,835]" italics="true" pageId="9" pageNumber="8">agenor</emphasis>
</taxonomicName>
is 
<collectingRegion id="867BF870FFA4FFC903C33710FDFDFCFB" box="[491,595,810,836]" country="Panama" name="Veraguas" pageId="9" pageNumber="8">Veraguas</collectingRegion>
, 
<collectingCountry id="3CA87602FFA4FFC900743710FD18FCFB" box="[604,694,810,836]" name="Panama" pageId="9" pageNumber="8">Panama</collectingCountry>
(
<geoCoordinate id="218B5055FFA4FFC900EB3713FC99FCFC" box="[707,823,809,835]" degrees="8.1996830" direction="north" orientation="latitude" pageId="9" pageNumber="8" precision="1" value="8.199683">8.1996830</geoCoordinate>
<geoCoordinate id="218B5055FFA4FFC901143713FC7FFCFB" box="[828,977,809,836]" degrees="81.2343500" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="1" value="-81.23435">−81.2343500</geoCoordinate>
), thus separating populations of 
<taxonomicName id="83BF4D11FFA4FFC9071D3710FA0EFCFC" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[1333,1440,810,836]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="agenor">
<emphasis id="76CBEA80FFA4FFC9071D3710FAE7FCFC" box="[1333,1353,810,835]" italics="true" pageId="9" pageNumber="8">D</emphasis>
. 
<emphasis id="76CBEA80FFA4FFC9077E3710FA0EFCFC" box="[1366,1440,810,835]" italics="true" pageId="9" pageNumber="8">agenor</emphasis>
</taxonomicName>
and 
<taxonomicName id="83BF4D11FFA4FFC902D83776FED5FCD9" authority="Solís &amp; Kohlmann, 2022" authorityName="Solís &amp; Kohlmann" authorityYear="2022" box="[240,379,844,870]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov.">
<emphasis id="76CBEA80FFA4FFC902D83776FEAAFCDA" box="[240,260,844,869]" italics="true" pageId="9" pageNumber="8">D</emphasis>
. 
<emphasis id="76CBEA80FFA4FFC903393776FED5FCD9" box="[273,379,844,870]" italics="true" pageId="9" pageNumber="8">woodruffi</emphasis>
</taxonomicName>
<emphasis id="76CBEA80FFA4FFC903A93776FDA7FCDA" bold="true" box="[385,521,843,870]" pageId="9" pageNumber="8">
<taxonomicNameLabel id="6DF857FBFFA4FFC903A93776FDA7FCDA" box="[385,521,843,870]" pageId="9" pageNumber="8" rank="species">new species</taxonomicNameLabel>
</emphasis>
by approximately 
<quantity id="83479B77FFA4FFC900FE3776FC83FCD9" box="[726,813,844,870]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="8" unit="km" value="200.0">200 km</quantity>
.
</paragraph>
<paragraph id="44003692FFA4FFC902E8374CFE47FA30" blockId="9.[191,1442,886,1424]" pageId="9" pageNumber="8">
<emphasis id="76CBEA80FFA4FFC902E8374CFE1DFC2F" bold="true" box="[192,435,886,912]" pageId="9" pageNumber="8">Taxonomic remarks.</emphasis>
<bibRefCitation id="202E4B63FFA4FFC90394374CFC0AFC2F" author="Montoya-Molina S &amp; Vaz-de-Mello FZ" box="[444,932,886,912]" pageId="9" pageNumber="8" pagination="1 - 64" refId="ref8403" refString="Montoya-Molina S, Vaz-de-Mello FZ. 2021. Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae). European Journal of Taxonomy 734: 1 - 64." type="journal article" year="2021">Montoya-Molina and Vaz-de-Mello (2021)</bibRefCitation>
reported 
<specimenCount id="52B9FD1BFFA4FFC90630374CFB18FC2F" box="[1048,1206,886,912]" pageId="9" pageNumber="8" type="generic">six specimens</specimenCount>
of 
<taxonomicName id="83BF4D11FFA4FFC906F7374DFA97FC2F" authorityName="Montoya-Molina and Vaz-de-Mello" authorityYear="2021" box="[1247,1337,886,912]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="enioi">
<emphasis id="76CBEA80FFA4FFC906F7374DFB5DFC2F" box="[1247,1267,887,912]" italics="true" pageId="9" pageNumber="8">D</emphasis>
. 
<emphasis id="76CBEA80FFA4FFC9072B374DFA97FC2F" box="[1283,1337,887,912]" italics="true" pageId="9" pageNumber="8">enioi</emphasis>
</taxonomicName>
in Santa Rosa, 
<collectingRegion id="867BF870FFA4FFC9032F37A2FE5BFC0D" box="[263,501,920,946]" country="Costa Rica" name="Guanacaste" pageId="9" pageNumber="8">Guanacaste Province</collectingRegion>
and in the Quebrada Bonita Station in the Carara Biological Reserve in 
<collectingRegion id="867BF870FFA4FFC9076237A2FEF2FC6B" country="Costa Rica" name="Puntarenas" pageId="9" pageNumber="8">Puntarenas Province</collectingRegion>
along the Pacific coast of 
<collectingCountry id="3CA87602FFA4FFC900A93780FD53FC6B" box="[641,765,954,980]" name="Costa Rica" pageId="9" pageNumber="8">Costa Rica</collectingCountry>
. We examined specimens from these same localities, but we recorded the presence only of 
<taxonomicName id="83BF4D11FFA4FFC9002437E7FD39FC49" authority="Solís &amp; Kohlmann, 2022" authorityName="Solís &amp; Kohlmann" authorityYear="2022" box="[524,663,988,1015]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov.">
<emphasis id="76CBEA80FFA4FFC9002437E7FD8EFC49" box="[524,544,989,1014]" italics="true" pageId="9" pageNumber="8">D</emphasis>
. 
<emphasis id="76CBEA80FFA4FFC9000537E6FD39FC49" box="[557,663,988,1014]" italics="true" pageId="9" pageNumber="8">woodruffi</emphasis>
</taxonomicName>
<emphasis id="76CBEA80FFA4FFC900B437E6FC8DFC49" bold="true" box="[668,803,988,1014]" pageId="9" pageNumber="8">
<taxonomicNameLabel id="6DF857FBFFA4FFC900B437E6FC8DFC49" box="[668,803,988,1014]" pageId="9" pageNumber="8" rank="species">new species</taxonomicNameLabel>
</emphasis>
. (
<specimenCount id="52B9FD1BFFA4FFC9011037E6FC61FC49" box="[824,975,988,1015]" pageId="9" pageNumber="8" type="generic">35 specimens</specimenCount>
from Santa Rosa and 
<specimenCount id="52B9FD1BFFA4FFC906E837E6FACDFC49" box="[1216,1379,988,1015]" pageId="9" pageNumber="8" type="generic">152 specimens</specimenCount>
from Quebrada Bonita). We have recorded no specimens of 
<taxonomicName id="83BF4D11FFA4FFC9011037C5FC3CFBA7" authorityName="Montoya-Molina and Vaz-de-Mello" authorityYear="2021" box="[824,914,1023,1049]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="enioi">
<emphasis id="76CBEA80FFA4FFC9011037C5FCE2FBA7" box="[824,844,1023,1048]" italics="true" pageId="9" pageNumber="8">D</emphasis>
. 
<emphasis id="76CBEA80FFA4FFC9017437C5FC3CFBA7" box="[860,914,1023,1048]" italics="true" pageId="9" pageNumber="8">enioi</emphasis>
</taxonomicName>
in 
<collectingCountry id="3CA87602FFA4FFC9019337C4FB98FBA7" box="[955,1078,1022,1048]" name="Costa Rica" pageId="9" pageNumber="8">Costa Rica</collectingCountry>
or 
<collectingCountry id="3CA87602FFA4FFC9064937C4FB79FBA7" box="[1121,1239,1022,1048]" name="Nicaragua" pageId="9" pageNumber="8">Nicaragua</collectingCountry>
. Instead, we have recorded only 
<taxonomicName id="83BF4D11FFA4FFC9034C301BFE4EFB85" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[356,480,1056,1083]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="centralis">
<emphasis id="76CBEA80FFA4FFC9034C301BFED9FB85" box="[356,375,1057,1082]" italics="true" pageId="9" pageNumber="8">D</emphasis>
. 
<emphasis id="76CBEA80FFA4FFC903AD301AFE4EFB85" box="[389,480,1056,1082]" italics="true" pageId="9" pageNumber="8">centralis</emphasis>
</taxonomicName>
and 
<taxonomicName id="83BF4D11FFA4FFC90031301BFD0BFB85" authority="Solís &amp; Kohlmann, 2022" authorityName="Solís &amp; Kohlmann" authorityYear="2022" box="[537,677,1056,1083]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov.">
<emphasis id="76CBEA80FFA4FFC90031301BFD83FB85" box="[537,557,1057,1082]" italics="true" pageId="9" pageNumber="8">D</emphasis>
. 
<emphasis id="76CBEA80FFA4FFC90013301AFD0BFB85" box="[571,677,1056,1082]" italics="true" pageId="9" pageNumber="8">woodruffi</emphasis>
</taxonomicName>
<emphasis id="76CBEA80FFA4FFC90084301AFC9AFB85" bold="true" box="[684,820,1056,1082]" pageId="9" pageNumber="8">
<taxonomicNameLabel id="6DF857FBFFA4FFC90084301AFC9AFB85" box="[684,820,1056,1082]" pageId="9" pageNumber="8" rank="species">new species</taxonomicNameLabel>
</emphasis>
in 
<collectingCountry id="3CA87602FFA4FFC90172301AFC7DFB85" box="[858,979,1056,1082]" name="Costa Rica" pageId="9" pageNumber="8">Costa Rica</collectingCountry>
and 
<collectingCountry id="3CA87602FFA4FFC90624301AFBD1FB85" box="[1036,1151,1056,1082]" name="Nicaragua" pageId="9" pageNumber="8">Nicaragua</collectingCountry>
(
<figureCitation id="DC842A17FFA4FFC906A6301AFB70FB85" box="[1166,1246,1056,1082]" captionStart="Figure 19" captionStartId="11.[192,268,1866,1892]" captionTargetBox="[192,1440,1091,1837]" captionTargetId="figure-407@11.[192,1440,1091,1837]" captionTargetPageId="11" captionText="Figure 19. Known distribution of Dichotomius agenor, D. centralis, and D. woodruffi new species in Nicaragua, Costa Rica, and Panama." figureDoi="http://doi.org/10.5281/zenodo.6392015" httpUri="https://zenodo.org/record/6392015/files/figure.png" pageId="9" pageNumber="8">Fig. 19</figureCitation>
). Substantial evidence is here presented for the species concept (
<taxonomicName id="83BF4D11FFA4FFC900FA3079FCF2FBE3" authority="Solís &amp; Kohlmann, 2022" authorityName="Solís &amp; Kohlmann" authorityYear="2022" box="[722,860,1090,1117]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov.">
<emphasis id="76CBEA80FFA4FFC900FA3079FD48FBE3" box="[722,742,1091,1116]" italics="true" pageId="9" pageNumber="8">D</emphasis>
. 
<emphasis id="76CBEA80FFA4FFC900DA3078FCF2FBE3" box="[754,860,1090,1116]" italics="true" pageId="9" pageNumber="8">woodruffi</emphasis>
</taxonomicName>
<emphasis id="76CBEA80FFA4FFC901493079FC47FBE3" bold="true" box="[865,1001,1090,1117]" pageId="9" pageNumber="8">
<taxonomicNameLabel id="6DF857FBFFA4FFC901493079FC47FBE3" box="[865,1001,1090,1117]" pageId="9" pageNumber="8" rank="species">new species</taxonomicNameLabel>
</emphasis>
). As indicated earlier, 
<bibRefCitation id="202E4B63FFA4FFC906C93078FE78FBC0" author="Montoya-Molina S &amp; Vaz-de-Mello FZ" pageId="9" pageNumber="8" pagination="1 - 64" refId="ref8403" refString="Montoya-Molina S, Vaz-de-Mello FZ. 2021. Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae). European Journal of Taxonomy 734: 1 - 64." type="journal article" year="2021">Montoya-Molina and Vaz-de-Mello (2021)</bibRefCitation>
seem to have confused 
<taxonomicName id="83BF4D11FFA4FFC900F2305FFC9FFBC1" authorityName="Montoya-Molina and Vaz-de-Mello" authorityYear="2021" box="[730,817,1125,1151]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="enioi">
<emphasis id="76CBEA80FFA4FFC900F2305FFD40FBC1" box="[730,750,1125,1150]" italics="true" pageId="9" pageNumber="8">D</emphasis>
. 
<emphasis id="76CBEA80FFA4FFC900D3305FFC9FFBC1" box="[763,817,1125,1150]" italics="true" pageId="9" pageNumber="8">enioi</emphasis>
</taxonomicName>
specimens with 
<taxonomicName id="83BF4D11FFA4FFC901CF305FFBCCFBC1" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[999,1122,1124,1151]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="centralis">
<emphasis id="76CBEA80FFA4FFC901CF305FFC55FBC1" box="[999,1019,1125,1150]" italics="true" pageId="9" pageNumber="8">D</emphasis>
. 
<emphasis id="76CBEA80FFA4FFC9062F305EFBCCFBC1" box="[1031,1122,1124,1150]" italics="true" pageId="9" pageNumber="8">centralis</emphasis>
</taxonomicName>
. These species are separated based on the following characteristics, in the case of 
<taxonomicName id="83BF4D11FFA4FFC9013630BDFC76FB1E" authority=": Ventral" authorityName="Ventral" box="[798,984,1159,1185]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="enioi">
<emphasis id="76CBEA80FFA4FFC9013630BDFC9CFB1F" box="[798,818,1159,1184]" italics="true" pageId="9" pageNumber="8">D</emphasis>
. 
<emphasis id="76CBEA80FFA4FFC9016930BDFCD9FB1F" box="[833,887,1159,1184]" italics="true" pageId="9" pageNumber="8">enioi</emphasis>
: Ventral
</taxonomicName>
clypeal process truncate (
<figureCitation id="DC842A17FFA4FFC9072A30BDFAFDFB1D" box="[1282,1363,1159,1186]" captionStart="Figures 13–18" captionStartId="10.[192,279,1765,1791]" captionTargetBox="[260,1372,213,1736]" captionTargetId="figure-95@10.[260,1372,213,1736]" captionTargetPageId="10" captionText="Figures 13–18. Comparative structures of Dichotomius spp. 13) D. centralis anterior pronotal surface (lateraloblique view) subrugose, male pronotal anterior declivity with a medial carina. 14) D. agenor ventral clypeal process bilobed. 15) D. agenor female cephalic carina bituberculate. 16) D. enioi ventral clypeal process truncate. 17) D. enioi male anterior pronotal border simple, not medially sinuate. 18) D. enioi male pygidium less than twice as long as wide, convex." figureDoi="http://doi.org/10.5281/zenodo.6392013" httpUri="https://zenodo.org/record/6392013/files/figure.png" pageId="9" pageNumber="8">Fig. 16</figureCitation>
); male anterior pronotal border simple, not medially sinuate (
<figureCitation id="DC842A17FFA4FFC901343093FCC5FB7C" box="[796,875,1193,1219]" captionStart="Figures 13–18" captionStartId="10.[192,279,1765,1791]" captionTargetBox="[260,1372,213,1736]" captionTargetId="figure-95@10.[260,1372,213,1736]" captionTargetPageId="10" captionText="Figures 13–18. Comparative structures of Dichotomius spp. 13) D. centralis anterior pronotal surface (lateraloblique view) subrugose, male pronotal anterior declivity with a medial carina. 14) D. agenor ventral clypeal process bilobed. 15) D. agenor female cephalic carina bituberculate. 16) D. enioi ventral clypeal process truncate. 17) D. enioi male anterior pronotal border simple, not medially sinuate. 18) D. enioi male pygidium less than twice as long as wide, convex." figureDoi="http://doi.org/10.5281/zenodo.6392013" httpUri="https://zenodo.org/record/6392013/files/figure.png" pageId="9" pageNumber="8">Fig. 17</figureCitation>
); setigerous lateral areas on the anterior and lateral lobes of metasternum continuous at the posteromedial mesocoxal margins; male pygidium less than twice as long as width, strongly convex (
<figureCitation id="DC842A17FFA4FFC903CD30D7FD9AFAB8" box="[485,564,1261,1287]" captionStart="Figures 13–18" captionStartId="10.[192,279,1765,1791]" captionTargetBox="[260,1372,213,1736]" captionTargetId="figure-95@10.[260,1372,213,1736]" captionTargetPageId="10" captionText="Figures 13–18. Comparative structures of Dichotomius spp. 13) D. centralis anterior pronotal surface (lateraloblique view) subrugose, male pronotal anterior declivity with a medial carina. 14) D. agenor ventral clypeal process bilobed. 15) D. agenor female cephalic carina bituberculate. 16) D. enioi ventral clypeal process truncate. 17) D. enioi male anterior pronotal border simple, not medially sinuate. 18) D. enioi male pygidium less than twice as long as wide, convex." figureDoi="http://doi.org/10.5281/zenodo.6392013" httpUri="https://zenodo.org/record/6392013/files/figure.png" pageId="9" pageNumber="8">Fig. 18</figureCitation>
); it is distributed in 
<collectingCountry id="3CA87602FFA4FFC9013A30D7FC25FAB8" box="[786,907,1261,1287]" name="Guatemala" pageId="9" pageNumber="8">Guatemala</collectingCountry>
and 
<collectingCountry id="3CA87602FFA4FFC901E930D7FBB8FAB8" box="[961,1046,1261,1287]" name="Mexico" pageId="9" pageNumber="8">Mexico</collectingCountry>
. Whereas 
<taxonomicName id="83BF4D11FFA4FFC906A030D4FABDFAB8" authority="Solís &amp; Kohlmann, 2022" authorityName="Solís &amp; Kohlmann" authorityYear="2022" box="[1160,1299,1261,1288]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov.">
<emphasis id="76CBEA80FFA4FFC906A030D4FB32FAB8" box="[1160,1180,1262,1287]" italics="true" pageId="9" pageNumber="8">D</emphasis>
. 
<emphasis id="76CBEA80FFA4FFC9068130D7FABDFAB8" box="[1193,1299,1261,1287]" italics="true" pageId="9" pageNumber="8">woodruffi</emphasis>
</taxonomicName>
<emphasis id="76CBEA80FFA4FFC9073130D7FA0EFAB8" bold="true" box="[1305,1440,1261,1287]" pageId="9" pageNumber="8">
<taxonomicNameLabel id="6DF857FBFFA4FFC9073130D7FA0EFAB8" box="[1305,1440,1261,1287]" pageId="9" pageNumber="8" rank="species">new species</taxonomicNameLabel>
</emphasis>
is characterized by: Ventral clypeal process spiniform (
<figureCitation id="DC842A17FFA4FFC901183135FCDDFA95" box="[816,883,1295,1322]" captionStart="Figures 1–6" captionStartId="6.[193,278,1833,1859]" captionTargetBox="[229,1405,221,1803]" captionTargetId="figure-18@6.[229,1405,199,1806]" captionTargetPageId="6" captionText="Figures 1–6. Dichotomius woodruffi, new species. 1) Dorsal habitus, holotype, male. Scale bar equals 1 cm. 2) Dorsal habitus, allotype, female. Scale bar equals 1 cm. 3) Frontal view, holotype, male. 4) Frontal view, allotype, female. 5) Ventral clypeal process spiniform, holotype male. 6) Anterior pronotal border (frontal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.6392009" httpUri="https://zenodo.org/record/6392009/files/figure.png" pageId="9" pageNumber="8">Fig. 5</figureCitation>
); male anterior pronotal border medially sinuate (
<figureCitation id="DC842A17FFA4FFC902E2310BFEA4FAF3" box="[202,266,1329,1356]" captionStart="Figures 1–6" captionStartId="6.[193,278,1833,1859]" captionTargetBox="[229,1405,221,1803]" captionTargetId="figure-18@6.[229,1405,199,1806]" captionTargetPageId="6" captionText="Figures 1–6. Dichotomius woodruffi, new species. 1) Dorsal habitus, holotype, male. Scale bar equals 1 cm. 2) Dorsal habitus, allotype, female. Scale bar equals 1 cm. 3) Frontal view, holotype, male. 4) Frontal view, allotype, female. 5) Ventral clypeal process spiniform, holotype male. 6) Anterior pronotal border (frontal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.6392009" httpUri="https://zenodo.org/record/6392009/files/figure.png" pageId="9" pageNumber="8">Fig. 6</figureCitation>
); setigerous lateral areas on the anterior and lateral lobes of metasternum interrupted at the posteromedial mesocoxal margins; male pygidium more than twice as long as width, slightly convex (
<figureCitation id="DC842A17FFA4FFC906AB3169FB6AFAD2" box="[1155,1220,1363,1389]" captionStart="Figures 7–12" captionStartId="8.[192,279,1799,1825]" captionTargetBox="[248,1385,213,1769]" captionTargetId="figure-84@8.[248,1385,213,1769]" captionTargetPageId="8" captionText="Figures 7–12. Dichotomius woodruffi, new species, and D. centralis. 7–11) D. woodruffi. 7) Anterior pronotal sulcus (lateral-oblique view), holotype male. 8) Male pygidium twice as broad as long, slightly convex, holotype male. 9) Lateral view of the aedeagus, holotype male. 10) Ventral view of the aedeagus, holotype male. 11) Lamella copulatrix, holotype male. 12) D. centralis, elytral stria crenulate." figureDoi="http://doi.org/10.5281/zenodo.6392011" httpUri="https://zenodo.org/record/6392011/files/figure.png" pageId="9" pageNumber="8">Fig. 8</figureCitation>
); it is distributed in 
<collectingCountry id="3CA87602FFA4FFC902E8314CFE96FA30" box="[192,312,1397,1424]" name="Costa Rica" pageId="9" pageNumber="8">Costa Rica</collectingCountry>
and 
<collectingCountry id="3CA87602FFA4FFC90347314FFE4BFA30" box="[367,485,1397,1423]" name="Nicaragua" pageId="9" pageNumber="8">Nicaragua</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>