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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.430.8094" ID-PMC="PMC4141162" ID-Pensoft-Pub="1313-2970-430-1" ID-Pensoft-UUID="FFE6FF8CFFD43B66335AD2203774DB49" ID-PubMed="25152663" ID-Zenodo-Dep="578441" ID-ZooBank="86F760563B8B49FB9C86FAD0DB0CBE8C" ModsDocID="1313-2970-430-1" checkinTime="1451245551152" checkinUser="pensoft" docAuthor="Erwin, Terry L. &amp; Zamorano, Laura S." docDate="2014" docId="C7DF5510A9100961CCA1CB439B486E30" docLanguage="en" docName="ZooKeys 430: 1-108" docOrigin="ZooKeys 430" docPubDate="2014-08-01" docSource="http://dx.doi.org/10.3897/zookeys.430.8094" docTitle="Asklepia surinamensis Zamorano &amp; Erwin 2014, sp. n." docType="treatment" docUuid="EEC60AF5-9DBC-4304-AE84-901AFEF674FD" docUuidSource="ZooBank" docVersion="5" id="FFE6FF8CFFD43B66335AD2203774DB49" lastPageNumber="73" masterDocId="FFE6FF8CFFD43B66335AD2203774DB49" masterDocTitle="A synopsis of the tribe Lachnophorini, with a new genus of Neotropical distribution and a revision of the Neotropical genus Asklepia Liebke, 1938 (Insecta, Coleoptera, Carabidae)" masterLastPageNumber="108" masterPageNumber="1" pageNumber="69" updateTime="1668159067277" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A synopsis of the tribe Lachnophorini, with a new genus of Neotropical distribution and a revision of the Neotropical genus Asklepia Liebke, 1938 (Insecta, Coleoptera, Carabidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Erwin, Terry L.</mods:namePart>
<mods:affiliation>Hyper-diversity Group, Department of Entomology, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA</mods:affiliation>
<mods:nameIdentifier type="email">erwint@si.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zamorano, Laura S.</mods:namePart>
<mods:affiliation>Research Student, MRC- 187, National Museum of Natural History, Smithsonian Institution, Washington, P. O. Box 37012, DC 20013 - 7012, USA &amp; Laboratorio de Zoologia Acuatica LAZOEA, Departamento de Ciencias Biologicas, Universidad de los Andes, Colombia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-08-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>430</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>108</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.430.8094</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.430.8094</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-430-1</mods:identifier>
<mods:identifier type="ZooBank">86F760563B8B49FB9C86FAD0DB0CBE8C</mods:identifier>
<mods:identifier type="Pensoft-UUID">FFE6FF8CFFD43B66335AD2203774DB49</mods:identifier>
<mods:identifier type="Zenodo-Dep">578441</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152054762" LSID="urn:lsid:zoobank.org:act:EEC60AF5-9DBC-4304-AE84-901AFEF674FD" httpUri="http://treatment.plazi.org/id/C7DF5510A9100961CCA1CB439B486E30" lastPageId="72" lastPageNumber="73" pageId="68" pageNumber="69">
<subSubSection pageId="68" pageNumber="69" type="nomenclature">
<paragraph pageId="68" pageNumber="69">
<taxonomicName LSID="http://zoobank.org/EEC60AF5-9DBC-4304-AE84-901AFEF674FD" authority="Zamorano &amp; Erwin" authorityName="Zamorano &amp; Erwin" authorityYear="2014" class="Insecta" family="Carabidae" genus="Asklepia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Asklepia surinamensis" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="surinamensis" status="sp. n.">Asklepia surinamensis Zamorano &amp; Erwin</taxonomicName>
<taxonomicNameLabel pageId="68" pageNumber="69">sp. n.</taxonomicNameLabel>
Surinam pattern-wing beetle
<figureCitation captionStart="Figure 5355" captionStartId="F15" captionText="Figure 53 - 55. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 53 Asklepia strandi Liebke, 1938, Guyana, ABL = 3.00 mm (from Liebke, 1938: 114). Liebke's illustration of Asklepia strandi Liebke. The single specimen Liebke described was likely lost during World War II. We have seen no specimens from Guyana, nor any specimens matching this illustration from anywhere 54 Asklepia surinamensis Erwin &amp; Zamorano, sp. n. ADP 130040, l'Hermitage, Surinam River, Surinam 55 Asklepia vigilante Erwin &amp; Zamorano, sp. n. ADP 051642, Boca del Rio Samiria, Peru. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30645" pageId="68" pageNumber="69">Figs 54</figureCitation>
<figureCitation captionStart="Figure 6974" captionStartId="F19" captionText="Figure 69 - 74. 69 Asklepia kathleenae Erwin &amp; Zamorano, sp. n. ADP 132529, Belem, Brazil 70 Asklepia marchantaria Erwin &amp; Zamorano, sp. n. Adis 001103, Ilha de Marchantaria, Lago Camaleao, Brazil 71 Asklepia pakitza Erwin &amp; Zamorano, sp. n. ADP 132466, Pakitza, Peru 72 Asklepia paraguayensis Zamorano &amp; Erwin, sp. n. ADP 132769, San Lorenzo Paraguay 73 Asklepia surinamensis Zamorano &amp; Erwin, sp. n. ADP 132763, l'Hermitage, Surinam 74 Asklepia vigilante Erwin &amp; Zamorano, sp. n. ADP 067301, Boca del Rio Samiria, Peru." httpUri="https://binary.pensoft.net/fig/30649" pageId="68" pageNumber="69">, 73</figureCitation>
<figureCitation captionStart="Figure 78" captionStartId="F22" captionText="Figure 78. Distribution map for known localities of Asklepia species of the pulchripennis group." httpUri="https://binary.pensoft.net/fig/30652" pageId="68" pageNumber="69">, 78</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="68" pageNumber="69" type="holotype">
<paragraph pageId="68" pageNumber="69">Holotype.</paragraph>
<paragraph pageId="68" pageNumber="69">
<emphasis bold="true" pageId="68" pageNumber="69">Surinam</emphasis>
, Paramaribo,
<normalizedToken originalValue="lHermitage">l'Hermitage</normalizedToken>
, Surinam River,
<geoCoordinate degrees="5.8182" direction="north" orientation="latitude" precision="5" value="5.8182">5.8182°N</geoCoordinate>
,
<geoCoordinate degrees="55.1639" direction="west" orientation="longitude" precision="5" value="-55.1639">55.1639°W</geoCoordinate>
, 2m, 14 July 1969 (N. Nieser)(CMNH: ADP130040, male).
</paragraph>
</subSubSection>
<subSubSection pageId="69" pageNumber="70" type="derivation of specific epithet">
<paragraph pageId="69" pageNumber="70">
<pageBreakToken pageId="69" pageNumber="70" start="start">Derivation</pageBreakToken>
of specific epithet.
</paragraph>
<paragraph pageId="69" pageNumber="70">
The specific epithet,
<emphasis italics="true" pageId="69" pageNumber="70">surinamensis</emphasis>
, is a singular Latinized feminine noun in apposition, based on the name of the country in which these beetles are found.
</paragraph>
</subSubSection>
<subSubSection pageId="70" pageNumber="71" type="proposed english vernacular name">
<paragraph pageId="70" pageNumber="71">
<pageBreakToken pageId="70" pageNumber="71" start="start">Proposed</pageBreakToken>
english vernacular name.
</paragraph>
<paragraph pageId="70" pageNumber="71">Surinam pattern-wing beetle.</paragraph>
</subSubSection>
<subSubSection lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71" type="diagnosis">
<paragraph pageId="70" pageNumber="71">Diagnosis.</paragraph>
<paragraph lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71">
With the attributes of the genus
<taxonomicName authorityName="Liebke" authorityYear="1938" class="Insecta" family="Carabidae" genus="Asklepia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="71" pageNumber="72">
<pageBreakToken pageId="71" pageNumber="72" start="start">Asklepia</pageBreakToken>
</emphasis>
</taxonomicName>
as described by
<bibRefCitation author="Liebke, M" journalOrPublisher="Embrik Strand" pageId="81" pageNumber="82" pagination="37 - 141" refId="B54" refString="Liebke, M, 1938. Denkschrift ueber die Carabiden-Tribus Colliurini. Festschrift zum 60. Geburtstage von Profesor Dr. Embrik Strand 4: 37 - 141" title="Denkschrift ueber die Carabiden-Tribus Colliurini. Festschrift zum 60. Geburtstage von Profesor Dr." volume="4" year="1938">Liebke (1938)</bibRefCitation>
and as noted above under the generic diagnosis, and medium-size to large-size for the genus (SBL = 2.841-2.99 mm). Adults with head fuscous, prothorax fulvous, elytral maculae fulvous or slightly aurantiacus in some individuals; elytron fuscous with a triangular flavous macula in the upper right corner of the proximal basal quadrant, broad flavous macula ending in hook crossing from medial lateral quadrant to left half of medial proximal quadrant, triangular flavous macula crossing from the lower right corner of apical proximal quadrant to lower half of apical lateral quadrant, apical and lateral margin fulvous; metepisternum fuscous, metasternum fuscous laterally flavous medially, abdominal sterna with III-VI, and epipleuron fulvous, abdominal sternum VII fuscous; legs flavotestaceous; antennal scape, pedicel and antennomere 3 testaceous, antennomeres 4-6 deeply infuscated, 7-11 white. Dorsal surface devoid of microsculpture, surface luster very shiny. Pronotum markedly convex and globose with lateral margin effaced except just anterior to hind angle and there a feeble bead; hind angle moderately prominent; anterior angles feebly produced; median line feebly defined. Elytral interneurs evident as continuous rows of widely spaced coarse punctures.
</paragraph>
</subSubSection>
<subSubSection pageId="71" pageNumber="72" type="description">
<paragraph pageId="71" pageNumber="72">Description.</paragraph>
<paragraph pageId="71" pageNumber="72">
(
<figureCitation captionStart="Figure 5355" captionStartId="F15" captionText="Figure 53 - 55. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 53 Asklepia strandi Liebke, 1938, Guyana, ABL = 3.00 mm (from Liebke, 1938: 114). Liebke's illustration of Asklepia strandi Liebke. The single specimen Liebke described was likely lost during World War II. We have seen no specimens from Guyana, nor any specimens matching this illustration from anywhere 54 Asklepia surinamensis Erwin &amp; Zamorano, sp. n. ADP 130040, l'Hermitage, Surinam River, Surinam 55 Asklepia vigilante Erwin &amp; Zamorano, sp. n. ADP 051642, Boca del Rio Samiria, Peru. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30645" pageId="71" pageNumber="72">Fig. 54</figureCitation>
,
<figureCitation captionStart="Figure 6974" captionStartId="F19" captionText="Figure 69 - 74. 69 Asklepia kathleenae Erwin &amp; Zamorano, sp. n. ADP 132529, Belem, Brazil 70 Asklepia marchantaria Erwin &amp; Zamorano, sp. n. Adis 001103, Ilha de Marchantaria, Lago Camaleao, Brazil 71 Asklepia pakitza Erwin &amp; Zamorano, sp. n. ADP 132466, Pakitza, Peru 72 Asklepia paraguayensis Zamorano &amp; Erwin, sp. n. ADP 132769, San Lorenzo Paraguay 73 Asklepia surinamensis Zamorano &amp; Erwin, sp. n. ADP 132763, l'Hermitage, Surinam 74 Asklepia vigilante Erwin &amp; Zamorano, sp. n. ADP 067301, Boca del Rio Samiria, Peru." httpUri="https://binary.pensoft.net/fig/30649" pageId="71" pageNumber="72">73</figureCitation>
).
<emphasis bold="true" pageId="71" pageNumber="72">
<emphasis italics="true" pageId="71" pageNumber="72">Habitus</emphasis>
:
</emphasis>
<figureCitation captionStart="Figure 5355" captionStartId="F15" captionText="Figure 53 - 55. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 53 Asklepia strandi Liebke, 1938, Guyana, ABL = 3.00 mm (from Liebke, 1938: 114). Liebke's illustration of Asklepia strandi Liebke. The single specimen Liebke described was likely lost during World War II. We have seen no specimens from Guyana, nor any specimens matching this illustration from anywhere 54 Asklepia surinamensis Erwin &amp; Zamorano, sp. n. ADP 130040, l'Hermitage, Surinam River, Surinam 55 Asklepia vigilante Erwin &amp; Zamorano, sp. n. ADP 051642, Boca del Rio Samiria, Peru. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30645" pageId="71" pageNumber="72">Fig. 54</figureCitation>
).
<emphasis bold="true" pageId="71" pageNumber="72">
<emphasis italics="true" pageId="71" pageNumber="72">Size</emphasis>
:
</emphasis>
[See also
<tableCitation captionStart="Table 25" captionStartId="T25" captionText="Table 25. Asklepia surinamensis Zamorano &amp; Erwin, sp. n." httpUri="http://table.plazi.org/id/9CE08E80ED76D73837844A3D77ADC7B7" pageId="71" pageNumber="72" tableUuid="9CE08E80ED76D73837844A3D77ADC7B7">Table 25</tableCitation>
] Medium-size to large for the genus; ABL = 3.002-3.372 mm, SBL = 2.841-2.992 mm, TW (total width) 1.420-1.584 mm, LP = 0.575-0.626 mm, WP = 0.795-0.836 mm, LE = 1.709-1.803 mm.
<emphasis bold="true" pageId="71" pageNumber="72">
<emphasis italics="true" pageId="71" pageNumber="72">Color</emphasis>
:
</emphasis>
See diagnosis above.
<emphasis bold="true" pageId="71" pageNumber="72">
<emphasis italics="true" pageId="71" pageNumber="72">Luster</emphasis>
:
</emphasis>
See diagnosis above.
<emphasis bold="true" italics="true" pageId="71" pageNumber="72">Head</emphasis>
(
<figureCitation captionStart="Figure 5355" captionStartId="F15" captionText="Figure 53 - 55. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 53 Asklepia strandi Liebke, 1938, Guyana, ABL = 3.00 mm (from Liebke, 1938: 114). Liebke's illustration of Asklepia strandi Liebke. The single specimen Liebke described was likely lost during World War II. We have seen no specimens from Guyana, nor any specimens matching this illustration from anywhere 54 Asklepia surinamensis Erwin &amp; Zamorano, sp. n. ADP 130040, l'Hermitage, Surinam River, Surinam 55 Asklepia vigilante Erwin &amp; Zamorano, sp. n. ADP 051642, Boca del Rio Samiria, Peru. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30645" pageId="71" pageNumber="72">Fig. 54</figureCitation>
): as in description for genus above.
<emphasis bold="true" pageId="71" pageNumber="72">
<emphasis italics="true" pageId="71" pageNumber="72">Prothorax</emphasis>
.
</emphasis>
Pronotum (
<figureCitation captionStart="Figure 5355" captionStartId="F15" captionText="Figure 53 - 55. Digital Photo-illustrations, habitus, dorsal aspect of holotypes. 53 Asklepia strandi Liebke, 1938, Guyana, ABL = 3.00 mm (from Liebke, 1938: 114). Liebke's illustration of Asklepia strandi Liebke. The single specimen Liebke described was likely lost during World War II. We have seen no specimens from Guyana, nor any specimens matching this illustration from anywhere 54 Asklepia surinamensis Erwin &amp; Zamorano, sp. n. ADP 130040, l'Hermitage, Surinam River, Surinam 55 Asklepia vigilante Erwin &amp; Zamorano, sp. n. ADP 051642, Boca del Rio Samiria, Peru. Scale line = 1 mm." httpUri="https://binary.pensoft.net/fig/30645" pageId="71" pageNumber="72">Fig. 54</figureCitation>
) slightly broad, as wide as head across eyes (WH/WP, mean both sexes: 1.101), length about the same (LP/LH, mean both sexes: 1.093), wider than long (W/L, mean both sexes: 1.367); markedly cordiform and rounded, lateral margin effaced with seta at anterior third on slightly raised area; apex markedly constricted; anterior angle feebly produced, hind angle slightly produced and setose; median line feebly defined, apical transverse impressions punctate, punctures infuscated; surface smooth throughout.
<emphasis bold="true" pageId="71" pageNumber="72">
<emphasis italics="true" pageId="71" pageNumber="72">Pterothorax</emphasis>
.
</emphasis>
Normal for genus, see description for genus above. Elytra moderate convex; at apical third twice as wide as head across eyes (WH/TW, mean both sexes: 0.542) and pronotum (WP/TW, mean both sexes: 0.4923). Elytral interneurs evident as continuous rows of widely spaced coarse punctures, punctures infuscated, interneurs continuous along length of entire elytron. Hind wings fully developed.
<emphasis bold="true" pageId="71" pageNumber="72">
<emphasis italics="true" pageId="71" pageNumber="72">Legs</emphasis>
.
</emphasis>
Overall, normal for genus, see description for genus above.
<emphasis bold="true" pageId="71" pageNumber="72">
<emphasis italics="true" pageId="71" pageNumber="72">Abdominal sterna</emphasis>
.
</emphasis>
Overall, normal for genus, see description for genus above.
<emphasis bold="true" italics="true" pageId="71" pageNumber="72">Male genitalia</emphasis>
(
<figureCitation captionStart="Figure 6974" captionStartId="F19" captionText="Figure 69 - 74. 69 Asklepia kathleenae Erwin &amp; Zamorano, sp. n. ADP 132529, Belem, Brazil 70 Asklepia marchantaria Erwin &amp; Zamorano, sp. n. Adis 001103, Ilha de Marchantaria, Lago Camaleao, Brazil 71 Asklepia pakitza Erwin &amp; Zamorano, sp. n. ADP 132466, Pakitza, Peru 72 Asklepia paraguayensis Zamorano &amp; Erwin, sp. n. ADP 132769, San Lorenzo Paraguay 73 Asklepia surinamensis Zamorano &amp; Erwin, sp. n. ADP 132763, l'Hermitage, Surinam 74 Asklepia vigilante Erwin &amp; Zamorano, sp. n. ADP 067301, Boca del Rio Samiria, Peru." httpUri="https://binary.pensoft.net/fig/30649" pageId="71" pageNumber="72">Fig. 73</figureCitation>
, see
<figureCitation captionStart="Figure 5762" captionStartId="F17" captionText="Figure 57 - 62. Illustrations, male aedeagus, dorsal, ventral, left lateral aspects. 57 Asklepia geminata (Bates, 1871) ADP 109186, Rio Samiria, Boca Cano Ingles Camp, Peru 58 Asklepia campbellorum Zamorano &amp; Erwin, sp. n. ADP 109196, 20 km SW Manaus, Brazil 59 Asklepia demiti Erwin &amp; Zamorano, sp. n. ADP 132585, Rio Demiti, Brazil 60 Asklepia grammechrysea Zamorano &amp; Erwin, sp. n. ADP 052565, Rio Sucusari, Peru 61 Asklepia laetitia Zamorano &amp; Erwin, sp. n. ADP 109190, Leticia, Colombia. Legend: a apical area; bl basal lobe; bo basal orifice; lp left paramere; ml median lobe; om ostial membrane; oo ostial opening; rp right paramere; sh shaft; pb phallobase; ps phallobase shaft; ms medial spine; ds distal spine 62 Asklepia lebioides (Bates, 1871), comb. n., ADP 109208, Rio Demiti, Brazil. Scale line = 0.25 mm." httpUri="https://binary.pensoft.net/fig/30647" pageId="71" pageNumber="72">Fig. 61</figureCitation>
for attribute labels). Median lobe with phallobase moderate in length, about a fifth the length of shaft, basal opening moderate, oriented oblique to shaft. Shaft broad, slightly curved ventrally, dorsally sclerotized except for short ostium; in ventral aspect tapered toward broadly rounded apex, in lateral aspect, a rounded blunt apex. Left paramere very large and broad, right paramere moderately large and triangular; apex of left paramere lobate much, longer than right paramere about two-thirds the length of shaft (measured in left lateral aspect). Endophallus with one small median spine, and one very large distal spine.
<emphasis bold="true" pageId="71" pageNumber="72">
<emphasis italics="true" pageId="71" pageNumber="72">Female genitalia</emphasis>
.
</emphasis>
Not investigated, presumably similar to that of
<taxonomicName authorityName="Erwin &amp; Zamorano" authorityYear="2014" class="Insecta" family="Carabidae" genus="Asklepia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="demiti">
<emphasis italics="true" pageId="71" pageNumber="72">Asklepia demiti</emphasis>
</taxonomicName>
sp. n.
</paragraph>
</subSubSection>
<subSubSection pageId="71" pageNumber="72" type="dispersal potential">
<paragraph pageId="71" pageNumber="72">Dispersal potential.</paragraph>
<paragraph pageId="71" pageNumber="72">These beetles are macropterous and capable of flight; they are attracted to lights. They are moderately swift and agile runners.</paragraph>
</subSubSection>
<subSubSection pageId="71" pageNumber="72" type="distribution">
<paragraph pageId="71" pageNumber="72">Distribution.</paragraph>
<paragraph pageId="71" pageNumber="72">
(
<figureCitation captionStart="Figure 78" captionStartId="F22" captionText="Figure 78. Distribution map for known localities of Asklepia species of the pulchripennis group." httpUri="https://binary.pensoft.net/fig/30652" pageId="71" pageNumber="72">Fig. 78</figureCitation>
). This species has been found at only one location on the white-water system of the middle Surinam River drainage system. But that does not at all indicate its real distribution: as has been pointed out above, very small beetles are inadequately sampled, especially in the Neotropics.
</paragraph>
</subSubSection>
<subSubSection pageId="71" pageNumber="72" type="way of life">
<paragraph pageId="71" pageNumber="72">Way of life.</paragraph>
<paragraph pageId="71" pageNumber="72">
See
<bibRefCitation author="Erwin, TL" journalOrPublisher="Revista Peruana de Entomologia" pageId="79" pageNumber="80" pagination="1 - 85" refId="B22" refString="Erwin, TL, 1991. Natural history of the carabid beetles at the BIOLAT Rio Manu Biological Station, Pakitza, Peru. Revista Peruana de Entomologia 33: 1 - 85" title="Natural history of the carabid beetles at the BIOLAT Rio Manu Biological Station, Pakitza, Peru." volume="33" year="1991">Erwin (1991)</bibRefCitation>
for a general description. Adults of this species are active in July.
</paragraph>
</subSubSection>
<subSubSection pageId="72" pageNumber="73" type="other specimens examined">
<paragraph pageId="72" pageNumber="73">
<pageBreakToken pageId="72" pageNumber="73" start="start">Other</pageBreakToken>
specimens examined.
</paragraph>
<paragraph pageId="72" pageNumber="73">
<emphasis bold="true" pageId="72" pageNumber="73">Surinam</emphasis>
, Paramaribo,
<normalizedToken originalValue="lHermitage">l'Hermitage</normalizedToken>
, Surinam River,
<geoCoordinate degrees="5.8182" direction="north" orientation="latitude" precision="5" value="5.8182">5.8182°N</geoCoordinate>
,
<geoCoordinate degrees="55.1639" direction="west" orientation="longitude" precision="5" value="-55.1639">55.1639°W</geoCoordinate>
, 2m, 10 July 1969 (N. Nieser)(CMNH: ADP130042, ADP132761, female paratypes; ADP132763, ADP132759, ADP132757, ADP133569 male paratypes).
</paragraph>
</subSubSection>
</treatment>
</document>