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<document id="B4FB7DDCAEA74B65F8E699BFDE496788" ID-DOI="10.17161/jom.v0i73.6673" ID-ISSN="2325-4467" ID-Zenodo-Dep="8057340" ID-ZooBank="urn:lsid:zoobank.org:pub:CF6CC73C-1405-48C5-9C23-6E11DAD4D06B" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1687209556228" checkinUser="felipe" docAuthor="Engel, Michael S. &amp; Rasmussen, Claus" docDate="2017" docId="C773EE1C21662447882FFB13DC4E0D8F" docLanguage="en" docName="JMelittology.2017.73.1-16.pdf" docOrigin="Journal of Melitology 2017 (73)" docSource="http://dx.doi.org/10.17161/jom.v0i73.6673" docStyle="DocumentStyle:20C606B715423E98DFED429F4976A416.1:JMelittology.2013-.journal_article.0cover" docStyleId="20C606B715423E98DFED429F4976A416" docStyleName="JMelittology.2013-.journal_article.0cover" docStyleVersion="1" docTitle="Heterotrigona (Sahulotrigona) Engel &amp; Rasmussen, new subgenus" docType="treatment" docVersion="1" lastPageNumber="8" masterDocId="3B4A9664216424408940FF8BDD230C76" masterDocTitle="A new subgenus of Heterotrigona from New Guinea (Hymenoptera: Apidae)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="3" updateTime="1689065143053" updateUser="tatiana" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:titleInfo id="D5A0FAADF9AF22E8BDBA76A024454310">
<mods:title id="1565211F7E04FC1FCF19C7045C3CACE4">A new subgenus of Heterotrigona from New Guinea (Hymenoptera: Apidae)</mods:title>
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<mods:name id="54474EE4ABD2AFF80C3A1D19A9F1F1BB" type="personal">
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<mods:roleTerm id="9C9BEA2448C552DAC25A784EB17A8043">Author</mods:roleTerm>
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<mods:namePart id="A7B0977B77E6FE956DA96EDAFD0DA53A">Engel, Michael S.</mods:namePart>
<mods:affiliation id="CE911658E63B6E258F28C77CCD8D2EC0">Division of Entomology, Natural History Museum, and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045 - 4415, USA (msengel @ ku. edu). &amp; Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79 th Street, New York, New York 10024 - 5192, USA.</mods:affiliation>
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<mods:namePart id="58C42E939CBD56D8610376622A39BD5B">Rasmussen, Claus</mods:namePart>
<mods:affiliation id="A55A660024D044BDFC9415E19CF125FC">Department of Bioscience, Aarhus University, Ole Worms Allé 1, DK- 8000 Aarhus, Denmark (alrunen @ yahoo. com).</mods:affiliation>
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<mods:title id="6401A777A1351396E44F330209E1DD5B">Journal of Melitology</mods:title>
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<mods:date id="BE53B2BF493E854FAD2748CC8562F1ED">2017</mods:date>
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<mods:number id="1C5C338E45FB55D2C374E66E057C5BE2">2017-11-03</mods:number>
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<treatment id="C773EE1C21662447882FFB13DC4E0D8F" LSID="urn:lsid:plazi:treatment:C773EE1C21662447882FFB13DC4E0D8F" httpUri="http://treatment.plazi.org/id/C773EE1C21662447882FFB13DC4E0D8F" lastPageId="7" lastPageNumber="8" pageId="2" pageNumber="3">
<subSubSection id="07C00C8121662442882FFB13DEE108C7" box="[367,962,1176,1201]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="4F655F0A21662442882FFB13DEE108C7" blockId="2.[367,962,1176,1201]" box="[367,962,1176,1201]" pageId="2" pageNumber="3">
<heading id="142DE86621662442882FFB13DEE108C7" box="[367,962,1176,1201]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="3" reason="2">
<taxonomicName id="88DA248921662442882FFB13DE2F08C6" authority="Engel &amp; Rasmussen" authorityName="Engel &amp; Rasmussen" box="[367,780,1176,1201]" class="Insecta" family="Apidae" genus="Heterotrigona" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subGenus" subGenus="Sahulotrigona">
<emphasis id="7DAE831821662442882FFB13DF3A08C6" bold="true" box="[367,537,1176,1200]" italics="true" pageId="2" pageNumber="3">Sahulotrigona</emphasis>
Engel &amp; Rasmussen
</taxonomicName>
,
<taxonomicNameLabel id="669D3E63216624428A57FB12DEE108C7" box="[791,962,1177,1201]" pageId="2" pageNumber="3" rank="subGenus">new subgenus</taxonomicNameLabel>
</heading>
</paragraph>
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<paragraph id="4F655F0A2166244289AFFB30D96208A6" blockId="2.[239,1089,1210,1233]" box="[239,1089,1210,1233]" pageId="2" pageNumber="3">
ZooBank:
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urn:lsid:zoobank.org:act:
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</paragraph>
</subSubSection>
<subSubSection id="07C00C812166244289E0FB70DE1F0966" box="[160,828,1274,1297]" pageId="2" pageNumber="3" type="reference_group">
<paragraph id="4F655F0A2166244289E0FB70DE1F0966" blockId="2.[160,828,1274,1297]" box="[160,828,1274,1297]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="6FCA78242166244289E0FB70DE1F0966" box="[160,828,1274,1297]" pageId="2" pageNumber="3">
<treatmentCitation id="CE7B791B2166244289E0FB70DFBD0966" author="Tadauchi, O. &amp; M. Ito &amp; J. - I. Kojima &amp; M. J. Toda" box="[160,670,1274,1297]" page="245" pageId="2" pageNumber="3" year="1998">
<taxonomicName id="88DA24892166244289E0FB70DFBD0966" authority="Tadauchi et al., 1998: 245" authorityName="Tadauchi" authorityPageNumber="245" authorityYear="1998" box="[160,670,1274,1297]" class="Insecta" family="Apidae" genus="Trigona" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subGenus" subGenus="Pacificotrigona">
<emphasis id="7DAE83182166244289E0FB70DDCE0966" box="[160,237,1275,1296]" italics="true" pageId="2" pageNumber="3">Trigona</emphasis>
(
<emphasis id="7DAE83182166244289BBFB70DCAF0966" box="[251,396,1275,1296]" italics="true" pageId="2" pageNumber="3">Pacificotrigona</emphasis>
)
<bibRefCitation id="2B4B22FB2166244288DAFB70DFBD0966" author="Tadauchi, O. &amp; M. Ito &amp; J. - I. Kojima &amp; M. J. Toda" box="[410,670,1274,1297]" pageId="2" pageNumber="3" pagination="229 - 246" refId="ref9334" refString="Tadauchi, O., M. Ito, J. - I. Kojima, &amp; M. J. Toda. 1998. Species and subspecies described by the late professor Dr. Sh. F. Sakagami (Insecta: Hymenoptera). Natural History Bulletin of Ibaraki University 2: 229 - 246." type="journal article" year="1998">
Tadauchi
<emphasis id="7DAE8318216624428B42FB70DF090966" box="[514,554,1275,1296]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
., 1998: 245
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<emphasis id="7DAE8318216624428BE8FB70DE140966" box="[680,823,1275,1296]" italics="true" pageId="2" pageNumber="3">nomen nudum</emphasis>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="07C00C81216624428990FAB2DDD80907" pageId="2" pageNumber="3" type="type_taxon">
<paragraph id="4F655F0A216624428990FAB2DDD80907" blockId="2.[160,1169,1337,1777]" pageId="2" pageNumber="3">
<typeStatus id="9061E1A8216624428990FAB2DC250926" box="[208,262,1337,1360]" pageId="2" pageNumber="3">TYPE</typeStatus>
SPECIES:
<taxonomicName id="88DA2489216624428836FAB1DEB80927" authority="Engel" authorityName="Engel &amp; Rasmussen" authorityYear="2017" box="[374,923,1337,1361]" class="Insecta" family="Apidae" genus="Heterotrigona" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="paradisaea" subGenus="Sahulotrigona">
<emphasis id="7DAE8318216624428836FAB1DF330927" box="[374,528,1338,1361]" italics="true" pageId="2" pageNumber="3">Heterotrigona</emphasis>
(
<emphasis id="7DAE8318216624428B66FAB2DFE10927" box="[550,706,1337,1361]" italics="true" pageId="2" pageNumber="3">Sahulotrigona</emphasis>
)
<emphasis id="7DAE8318216624428B97FAB2DE690927" box="[727,842,1337,1361]" italics="true" pageId="2" pageNumber="3">paradisaea</emphasis>
Engel
</taxonomicName>
&amp; Rasmussen,
<taxonomicNameLabel id="669D3E63216624428D1DFAB2DDD40907" pageId="2" pageNumber="3" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<subSubSection id="07C00C81216624438990FAF2D95B0A01" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="4F655F0A216624438990FAF2D95B0A01" blockId="2.[160,1169,1337,1777]" lastBlockId="3.[160,1168,1535,1783]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
DIAGNOSIS: The subgenus
<taxonomicName id="88DA2489216624428B49FAF2DF8109E7" box="[521,674,1401,1425]" pageId="2" pageNumber="3" rank="subGenus" subGenus="Sahulotrigona">
<emphasis id="7DAE8318216624428B49FAF2DF8109E7" box="[521,674,1401,1425]" italics="true" pageId="2" pageNumber="3">Sahulotrigona</emphasis>
</taxonomicName>
is distinctive among species of
<taxonomicName id="88DA2489216624428D64FAF1DDF709C7" authorityName="Schwarz" authorityYear="1939" class="Insecta" family="Apidae" genus="Heterotrigona" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="7DAE8318216624428D64FAF1DDF709C7" italics="true" pageId="2" pageNumber="3">Heterotrigona</emphasis>
</taxonomicName>
for the combination of a basal propodeal area that is largely glabrous (
<figureCitation id="D7E1438F216624428D54FA12D97409C7" box="[1044,1111,1433,1457]" captionStart="Figures 4–6" captionStartId="4.[160,241,1306,1328]" captionTargetBox="[174,1152,248,1288]" captionTargetId="figure-175@4.[160,1168,224,1299]" captionTargetPageId="4" captionText="Figures 4–6. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 4. Dorsal view of head and mesosoma. 5. Detail of propodeum in oblique dorso-posterior view. 6. Lateral view of mesosoma, detailing mesoscutellar profile." figureDoi="http://doi.org/10.5281/zenodo.8057346" httpUri="https://zenodo.org/record/8057346/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
), the basal vein (1M) postfurcal (
<emphasis id="7DAE83182166244288A8FA31DF2309A7" box="[488,512,1466,1489]" italics="true" pageId="2" pageNumber="3">i.e</emphasis>
., distad 1cu-a) (
<figureCitation id="D7E1438F216624428B80FA32DE2709A7" box="[704,772,1465,1489]" captionStart="Figures 7–10" captionStartId="5.[160,241,1531,1553]" captionTargetBox="[177,1151,236,1507]" captionTargetId="figure-106@5.[160,1168,224,1525]" captionTargetPageId="5" captionText="Figures 7–10. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 7. Forewing. 8. Outer view of metatibia and metatarsus. 9. Inner view of metatibia and metatarsus with posterior glabrate (yellow), apical glabrate (blue), keirotrichiate (green), and sericeous (pink) areas colored (arrows indicate width of posterior glabrate and length of apical glabrate zones). 10. Unmodified view of inner surface of metatibia and metatarsus." figureDoi="http://doi.org/10.5281/zenodo.8057350" httpUri="https://zenodo.org/record/8057350/files/figure.png" pageId="2" pageNumber="3">Fig. 7</figureCitation>
), and the posterior glabrate zone of the metatibial inner surface narrower than the raised keirotrichiate zone (
<figureCitation id="D7E1438F216624428D7BFA52DD9D0A67" captionStart="Figures 7–10" captionStartId="5.[160,241,1531,1553]" captionTargetBox="[177,1151,236,1507]" captionTargetId="figure-106@5.[160,1168,224,1525]" captionTargetPageId="5" captionText="Figures 7–10. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 7. Forewing. 8. Outer view of metatibia and metatarsus. 9. Inner view of metatibia and metatarsus with posterior glabrate (yellow), apical glabrate (blue), keirotrichiate (green), and sericeous (pink) areas colored (arrows indicate width of posterior glabrate and length of apical glabrate zones). 10. Unmodified view of inner surface of metatibia and metatarsus." figureDoi="http://doi.org/10.5281/zenodo.8057350" httpUri="https://zenodo.org/record/8057350/files/figure.png" pageId="2" pageNumber="3">Figs. 9, 10</figureCitation>
). The subgenus includes the most easterly of the known species of
<taxonomicName id="88DA2489216624428AB9FA71D9B30A67" authorityName="Schwarz" authorityYear="1939" box="[1017,1168,1530,1553]" class="Insecta" family="Apidae" genus="Heterotrigona" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="7DAE8318216624428AB9FA71D9B30A67" box="[1017,1168,1530,1553]" italics="true" pageId="2" pageNumber="3">Heterotrigona</emphasis>
</taxonomicName>
and is most similar to the subgenera
<taxonomicName id="88DA2489216624428B0EF992DFF60A47" box="[590,725,1561,1585]" class="Insecta" family="Apidae" genus="Platytrigona" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="7DAE8318216624428B0EF992DFF60A47" box="[590,725,1561,1585]" italics="true" pageId="2" pageNumber="3">Platytrigona</emphasis>
</taxonomicName>
and
<taxonomicName id="88DA2489216624428A4FF992DE800A47" box="[783,931,1561,1585]" class="Insecta" family="Apidae" genus="Sundatrigona" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="7DAE8318216624428A4FF992DE800A47" box="[783,931,1561,1585]" italics="true" pageId="2" pageNumber="3">Sundatrigona</emphasis>
</taxonomicName>
, all three subgenera possessing a narrow glabrate to subglabrate zone on the posterior border of the inner surface of the metatibia. From the former,
<taxonomicName id="88DA2489216624428BE0F9D2DE1A0A07" box="[672,825,1625,1649]" pageId="2" pageNumber="13" rank="subGenus" subGenus="Sahulotrigona">
<emphasis id="7DAE8318216624428BE0F9D2DE1A0A07" box="[672,825,1625,1649]" italics="true" pageId="2" pageNumber="3">Sahulotrigona</emphasis>
</taxonomicName>
can be readily distinguished by the largely glabrous basal area of the propodeum, while both subgenera share a postfurcal basal vein (albeit in some
<taxonomicName id="88DA2489216624428B14F912DFF80AC7" box="[596,731,1689,1713]" class="Insecta" family="Apidae" genus="Platytrigona" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="7DAE8318216624428B14F912DFF80AC7" box="[596,731,1689,1713]" italics="true" pageId="2" pageNumber="3">Platytrigona</emphasis>
</taxonomicName>
the basal vein is confluent with 1cu- a). Species of
<taxonomicName id="88DA2489216624428810F932DCF40AA7" box="[336,471,1721,1745]" class="Insecta" family="Apidae" genus="Platytrigona" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="7DAE8318216624428810F932DCF40AA7" box="[336,471,1721,1745]" italics="true" pageId="2" pageNumber="3">Platytrigona</emphasis>
</taxonomicName>
are also larger than those of
<taxonomicName id="88DA2489216624428A79F932DEF30AA7" box="[825,976,1721,1745]" pageId="2" pageNumber="13" rank="subGenus" subGenus="Sahulotrigona">
<emphasis id="7DAE8318216624428A79F932DEF30AA7" box="[825,976,1721,1745]" italics="true" pageId="2" pageNumber="3">Sahulotrigona</emphasis>
</taxonomicName>
, the latter more closely approximate the smaller proportions of
<taxonomicName id="88DA2489216624428B82F952DE760A87" box="[706,853,1753,1777]" class="Insecta" family="Apidae" genus="Sundatrigona" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="7DAE8318216624428B82F952DE760A87" box="[706,853,1753,1777]" italics="true" pageId="2" pageNumber="3">Sundatrigona</emphasis>
</taxonomicName>
. From the latter subgenus,
<taxonomicName id="88DA24892167244389E0FA74DC1A0A61" box="[160,313,1535,1559]" pageId="3" pageNumber="13" rank="subGenus" subGenus="Sahulotrigona">
<emphasis id="7DAE83182167244389E0FA74DC1A0A61" box="[160,313,1535,1559]" italics="true" pageId="3" pageNumber="4">Sahulotrigona</emphasis>
</taxonomicName>
can be distinguished by the postfurcal basal vein (antefurcal in
<taxonomicName id="88DA2489216724438D67FA74DDF70A41" class="Insecta" family="Apidae" genus="Sundatrigona" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="7DAE8318216724438D67FA74DDF70A41" italics="true" pageId="3" pageNumber="4">Sundatrigona</emphasis>
</taxonomicName>
), longer scape which is about as long as the torulocellar distance (scape distinctly shorter than torulocellar distance in
<taxonomicName id="88DA2489216724438B08F9B4DFFF0A21" box="[584,732,1599,1623]" class="Insecta" family="Apidae" genus="Sundatrigona" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="7DAE8318216724438B08F9B4DFFF0A21" box="[584,732,1599,1623]" italics="true" pageId="3" pageNumber="4">Sundatrigona</emphasis>
</taxonomicName>
), and the gena about as wide as compound eye in profile (gena narrower than compound eye in profile in
<taxonomicName id="88DA2489216724438A94F9D4D94B0A01" box="[980,1128,1631,1655]" class="Insecta" family="Apidae" genus="Sundatrigona" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="7DAE8318216724438A94F9D4D94B0A01" box="[980,1128,1631,1655]" italics="true" pageId="3" pageNumber="4">Sundatrigona</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="1BA50F822167244389E0FA0EDC6C09A3" ID-DOI="http://doi.org/10.5281/zenodo.8057344" ID-Zenodo-Dep="8057344" httpUri="https://zenodo.org/record/8057344/files/figure.png" pageId="3" pageNumber="4" startId="3.[160,241,1413,1435]" targetBox="[164,1158,244,1397]" targetPageId="3" targetType="figure">
<paragraph id="4F655F0A2167244389E0FA0EDC6C09A3" blockId="3.[160,1168,1413,1493]" pageId="3" pageNumber="4">
<emphasis id="7DAE83182167244389E0FA0EDC0209EC" bold="true" box="[160,289,1413,1435]" pageId="3" pageNumber="4">Figures 1–3.</emphasis>
Worker of
<taxonomicName id="88DA24892167244388DDFA0DDE0D09EC" authority="Engel &amp; Rasmussen, 2017" authorityName="Engel &amp; Rasmussen" authorityYear="2017" box="[413,814,1413,1435]" class="Insecta" family="Apidae" genus="Heterotrigona" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="paradisaea" status="new subgenus and species" subGenus="Sahulotrigona">
<emphasis id="7DAE83182167244388DDFA0DDF0609ED" box="[413,549,1414,1435]" italics="true" pageId="3" pageNumber="4">Heterotrigona</emphasis>
(
<emphasis id="7DAE8318216724438B72FA0EDF9809EC" box="[562,699,1413,1434]" italics="true" pageId="3" pageNumber="4">Sahulotrigona</emphasis>
)
<emphasis id="7DAE8318216724438B89FA0EDE0D09EC" box="[713,814,1413,1434]" italics="true" pageId="3" pageNumber="4">paradisaea</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="669D3E63216724438A7AFA0DD97009ED" box="[826,1107,1413,1435]" pageId="3" pageNumber="4" rank="species">new subgenus and species</taxonomicNameLabel>
, from eastern Papua New Guinea.
<emphasis id="7DAE8318216724438897FA29DCC909C1" bold="true" box="[471,490,1442,1463]" pageId="3" pageNumber="4">1.</emphasis>
Lateral habitus.
<emphasis id="7DAE8318216724438BE3FA29DF9609C1" bold="true" box="[675,693,1442,1463]" pageId="3" pageNumber="4">2.</emphasis>
Facial view.
<emphasis id="7DAE8318216724438A08FA29DE7909C1" bold="true" box="[840,858,1442,1463]" pageId="3" pageNumber="4">3.</emphasis>
Outer view of mandible (P = preapical tooth).
</paragraph>
</caption>
<subSubSection id="07C00C81216724478990F9F4D9580D6F" lastPageId="7" lastPageNumber="8" pageId="3" pageNumber="4" type="description">
<paragraph id="4F655F0A216724448990F9F4DCE30A03" blockId="3.[160,1168,1535,1783]" lastBlockId="4.[160,1168,1437,1782]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
DESCRIPTION: Workers of small size, forewing length (including tegula) approximately 5.6–6.0 mm; integument shiny, smooth, with scattered microscopic punctures; integumental maculation absent (
<figureCitation id="D7E1438F216724438B6EF934DFBB0AA1" box="[558,664,1727,1751]" captionStart="Figures 1–3" captionStartId="3.[160,241,1413,1435]" captionTargetBox="[164,1158,244,1397]" captionTargetId="figure-144@3.[160,1168,224,1407]" captionTargetPageId="3" captionText="Figures 1–3. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 1. Lateral habitus. 2. Facial view. 3. Outer view of mandible (P = preapical tooth)." figureDoi="http://doi.org/10.5281/zenodo.8057344" httpUri="https://zenodo.org/record/8057344/files/figure.png" pageId="3" pageNumber="4">Figs. 1, 2</figureCitation>
,
<figureCitation id="D7E1438F216724438BE5F934DFF30AA1" box="[677,720,1727,1751]" captionStart="Figures 4–6" captionStartId="4.[160,241,1306,1328]" captionTargetBox="[174,1152,248,1288]" captionTargetId="figure-175@4.[160,1168,224,1299]" captionTargetPageId="4" captionText="Figures 4–6. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 4. Dorsal view of head and mesosoma. 5. Detail of propodeum in oblique dorso-posterior view. 6. Lateral view of mesosoma, detailing mesoscutellar profile." figureDoi="http://doi.org/10.5281/zenodo.8057346" httpUri="https://zenodo.org/record/8057346/files/figure.png" pageId="3" pageNumber="4">4–6</figureCitation>
); fine, minute, white to off-white plumose pubescence on face and clypeus and fine, plumose pubescence on lateral surfaces of mesosoma and somewhat obscuring integument; mesoscutum and mesoscutellum with evenly distributed, abundant, minute, erect, branched, fuscous to black setae (except pale in yellow morph of
<taxonomicName id="88DA24892160244488BFFA55DFA90983" baseAuthorityName="Smith" baseAuthorityYear="1865" box="[511,650,1502,1525]" class="Insecta" family="Apidae" genus="Trigona" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="atricornis">
<emphasis id="7DAE83182160244488BFFA55DF370983" box="[511,532,1502,1525]" italics="true" pageId="4" pageNumber="5">H</emphasis>
.
<emphasis id="7DAE8318216024448B62FA55DFA90983" box="[546,650,1502,1525]" italics="true" pageId="4" pageNumber="5">atricornis</emphasis>
</taxonomicName>
), intermingled with more scattered, longer, erect, simple setae, neither of which obscure integument but are more abundant ventrally on mesopleuron; metasomal terga with sparse, minute, simple setae except apical tergum with numerous, fuscous, erect, short, apically plumose setae; sterna with long, erect, simple setae.
</paragraph>
<caption id="1BA50F822160244489E0FA91D9A6091F" ID-DOI="http://doi.org/10.5281/zenodo.8057346" ID-Zenodo-Dep="8057346" httpUri="https://zenodo.org/record/8057346/files/figure.png" pageId="4" pageNumber="5" startId="4.[160,241,1306,1328]" targetBox="[174,1152,248,1288]" targetPageId="4" targetType="figure">
<paragraph id="4F655F0A2160244489E0FA91D9A6091F" blockId="4.[160,1168,1306,1385]" pageId="4" pageNumber="5">
<emphasis id="7DAE83182160244489E0FA91DC030959" bold="true" box="[160,288,1306,1328]" pageId="4" pageNumber="5">Figures 4–6.</emphasis>
Worker of
<taxonomicName id="88DA24892160244488DDFA91DE0D0959" authority="Engel &amp; Rasmussen, 2017" authorityName="Engel &amp; Rasmussen" authorityYear="2017" box="[413,814,1306,1327]" class="Insecta" family="Apidae" genus="Heterotrigona" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="paradisaea" status="new subgenus and species" subGenus="Sahulotrigona">
<emphasis id="7DAE83182160244488DDFA91DF060959" box="[413,549,1306,1327]" italics="true" pageId="4" pageNumber="5">Heterotrigona</emphasis>
(
<emphasis id="7DAE8318216024448B72FA91DF980959" box="[562,699,1306,1327]" italics="true" pageId="4" pageNumber="5">Sahulotrigona</emphasis>
)
<emphasis id="7DAE8318216024448B89FA91DE0D0959" box="[713,814,1306,1327]" italics="true" pageId="4" pageNumber="5">paradisaea</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="669D3E63216024448A7AFA91D9700946" box="[826,1107,1306,1328]" pageId="4" pageNumber="5" rank="species">new subgenus and species</taxonomicNameLabel>
, from eastern Papua New Guinea.
<emphasis id="7DAE831821602444889EFABDDCD3093D" bold="true" box="[478,496,1334,1355]" pageId="4" pageNumber="5">4.</emphasis>
Dorsal view of head and mesosoma.
<emphasis id="7DAE8318216024448AD3FABCDE86093A" bold="true" box="[915,933,1335,1356]" pageId="4" pageNumber="5">5.</emphasis>
Detail of propodeum in oblique dorso-posterior view.
<emphasis id="7DAE8318216024448B41FAD8DF30091E" bold="true" box="[513,531,1363,1384]" pageId="4" pageNumber="5">6.</emphasis>
Lateral view of mesosoma, detailing mesoscutellar profile.
</paragraph>
</caption>
<paragraph id="4F655F0A216024468990F9F6D9680D8F" blockId="4.[160,1168,1437,1782]" lastBlockId="6.[160,1175,225,1786]" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
Head as broad as mesosoma, with face slightly broader than compound eye length; vertex short, rounded; ocelloccipital distance slightly greater than one ocellar diameter; interocellar distance approximately 2.0 times ocellar diameter; ocellocular distance 2.0 times ocellar diameter; scape almost as long as torulocellar distance; first flagellomere longer than second, second and third flagellomeres equal in length; second through tenth flagellomeres each about as long as wide; intertorular distance a little more than one-half torulorbital distance; upper torular tangent near facial midlength; frontal line weakly impressed, not carinate, extending from median ocellus to near upper torular tangent; inner orbit of compound eye weakly concave in upper third; gena about as broad as compound eye in profile, posterior border rounded; malar area of moderate length, about as long as flagellar diameter (
<figureCitation id="D7E1438F216224468AC9FEEADEE80D0F" box="[905,971,353,377]" captionStart="Figures 1–3" captionStartId="3.[160,241,1413,1435]" captionTargetBox="[164,1158,244,1397]" captionTargetId="figure-144@3.[160,1168,224,1407]" captionTargetPageId="3" captionText="Figures 1–3. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 1. Lateral habitus. 2. Facial view. 3. Outer view of mandible (P = preapical tooth)." figureDoi="http://doi.org/10.5281/zenodo.8057344" httpUri="https://zenodo.org/record/8057344/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
); labrum weakly convex, short, much wider than long, apical margin medially blunt to weakly convex; mandible bidentate, teeth well defined and incised (as in other
<taxonomicName id="88DA2489216224468AC7FE29D93D0DCF" authorityName="Schwarz" authorityYear="1939" box="[903,1054,418,441]" class="Insecta" family="Apidae" genus="Heterotrigona" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="7DAE8318216224468AC7FE29D93D0DCF" box="[903,1054,418,441]" italics="true" pageId="6" pageNumber="7">Heterotrigona</emphasis>
</taxonomicName>
), incision between first and second preapical teeth slightly less than orthogonal (
<figureCitation id="D7E1438F216224468A98FE4AD93A0DAF" box="[984,1049,449,473]" captionStart="Figures 1–3" captionStartId="3.[160,241,1413,1435]" captionTargetBox="[164,1158,244,1397]" captionTargetId="figure-144@3.[160,1168,224,1407]" captionTargetPageId="3" captionText="Figures 1–3. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 1. Lateral habitus. 2. Facial view. 3. Outer view of mandible (P = preapical tooth)." figureDoi="http://doi.org/10.5281/zenodo.8057344" httpUri="https://zenodo.org/record/8057344/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
); first and second labial palpomeres with several elongate, apically wavy but simple setae.
</paragraph>
<caption id="1BA50F822161244589E0FA70DC390AD7" ID-DOI="http://doi.org/10.5281/zenodo.8057350" ID-Zenodo-Dep="8057350" httpUri="https://zenodo.org/record/8057350/files/figure.png" pageId="5" pageNumber="6" startId="5.[160,241,1531,1553]" targetBox="[177,1151,236,1507]" targetPageId="5" targetType="figure">
<paragraph id="4F655F0A2161244589E0FA70DC390AD7" blockId="5.[160,1169,1531,1697]" pageId="5" pageNumber="6">
<emphasis id="7DAE83182161244589E0FA70DC0F0A67" bold="true" box="[160,300,1531,1553]" pageId="5" pageNumber="6">Figures 7–10.</emphasis>
Worker of
<taxonomicName id="88DA24892161244588E6FA77DE100A67" authority="Engel &amp; Rasmussen, 2017" authorityName="Engel &amp; Rasmussen" authorityYear="2017" box="[422,819,1532,1553]" class="Insecta" family="Apidae" genus="Heterotrigona" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="paradisaea" status="new subgenus and species" subGenus="Sahulotrigona">
<emphasis id="7DAE83182161244588E6FA77DF0E0A67" box="[422,557,1532,1553]" italics="true" pageId="5" pageNumber="6">Heterotrigona</emphasis>
(
<emphasis id="7DAE8318216124458B7AFA77DFE10A67" box="[570,706,1532,1553]" italics="true" pageId="5" pageNumber="6">Sahulotrigona</emphasis>
)
<emphasis id="7DAE8318216124458B8FFA77DE100A67" box="[719,819,1532,1553]" italics="true" pageId="5" pageNumber="6">paradisaea</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="669D3E63216124458A7EFA77D9770A64" box="[830,1108,1532,1554]" pageId="5" pageNumber="6" rank="species">new subgenus and species</taxonomicNameLabel>
, from eastern Papua New Guinea.
<emphasis id="7DAE8318216124458897F992DCCA0A58" bold="true" box="[471,489,1561,1582]" pageId="5" pageNumber="6">7.</emphasis>
Forewing.
<emphasis id="7DAE8318216124458B29F993DF580A5B" bold="true" box="[617,635,1560,1581]" pageId="5" pageNumber="6">8.</emphasis>
Outer view of metatibia and metatarsus.
<emphasis id="7DAE8318216124458D7EF993D9730A5B" bold="true" box="[1086,1104,1560,1581]" pageId="5" pageNumber="6">9.</emphasis>
Inner view of metatibia and metatarsus with posterior glabrate (yellow), apical glabrate (blue), keirotrichiate (green), and sericeous (pink) areas colored (arrows indicate width of posterior glabrate and length of apical glabrate zones).
<emphasis id="7DAE8318216124458B7EF9E4DF7F0AF2" bold="true" box="[574,604,1647,1668]" pageId="5" pageNumber="6">10.</emphasis>
Unmodified view of inner surface of metatibia and metatarsus.
</paragraph>
</caption>
<paragraph id="4F655F0A216224468990FD89D9300F6F" blockId="6.[160,1175,225,1786]" pageId="6" pageNumber="7">
Mesoscutum with median sulcus weakly impressed; notauli scarcely evident; parapsidal lines short, slightly shorter than ocellar diameter, and weakly impressed. Mesoscutellum short, ending at profile of metanotum and not overhanging propodeum (
<figureCitation id="D7E1438F2162244689B7FDEADC1E0E0F" box="[247,317,609,633]" captionStart="Figures 4–6" captionStartId="4.[160,241,1306,1328]" captionTargetBox="[174,1152,248,1288]" captionTargetId="figure-175@4.[160,1168,224,1299]" captionTargetPageId="4" captionText="Figures 4–6. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 4. Dorsal view of head and mesosoma. 5. Detail of propodeum in oblique dorso-posterior view. 6. Lateral view of mesosoma, detailing mesoscutellar profile." figureDoi="http://doi.org/10.5281/zenodo.8057346" httpUri="https://zenodo.org/record/8057346/files/figure.png" pageId="6" pageNumber="7">Fig. 6</figureCitation>
), rounded (
<figureCitation id="D7E1438F21622446888DFDEADF370E0F" box="[461,532,609,633]" captionStart="Figures 4–6" captionStartId="4.[160,241,1306,1328]" captionTargetBox="[174,1152,248,1288]" captionTargetId="figure-175@4.[160,1168,224,1299]" captionTargetPageId="4" captionText="Figures 4–6. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 4. Dorsal view of head and mesosoma. 5. Detail of propodeum in oblique dorso-posterior view. 6. Lateral view of mesosoma, detailing mesoscutellar profile." figureDoi="http://doi.org/10.5281/zenodo.8057346" httpUri="https://zenodo.org/record/8057346/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
), slightly swollen in profile, with shining transverse depression along mesoscuto-mesoscutellar sulcus. Propodeum long and sloping, with distinct angle between basal area and posterior surface (
<figureCitation id="D7E1438F216224468A7FFD2ADEA10ECF" box="[831,898,673,697]" captionStart="Figures 4–6" captionStartId="4.[160,241,1306,1328]" captionTargetBox="[174,1152,248,1288]" captionTargetId="figure-175@4.[160,1168,224,1299]" captionTargetPageId="4" captionText="Figures 4–6. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 4. Dorsal view of head and mesosoma. 5. Detail of propodeum in oblique dorso-posterior view. 6. Lateral view of mesosoma, detailing mesoscutellar profile." figureDoi="http://doi.org/10.5281/zenodo.8057346" httpUri="https://zenodo.org/record/8057346/files/figure.png" pageId="6" pageNumber="7">Fig. 6</figureCitation>
), dorsal-facing surface about as long as posterior surface; basal area smooth, shining, largely glabrous except apicolateral patches of setae (but never largely pubescent as in
<taxonomicName id="88DA2489216224468AE6FD6AD9AB0E8F" authority="Moure" authorityName="Moure" box="[934,1160,737,761]" class="Insecta" family="Apidae" genus="Platytrigona" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="7DAE8318216224468AE6FD6AD90E0E8F" box="[934,1069,737,761]" italics="true" pageId="6" pageNumber="7">Platytrigona</emphasis>
Moure
</taxonomicName>
) (
<figureCitation id="D7E1438F2162244689E8FC8ADDC80F6F" box="[168,235,769,793]" captionStart="Figures 4–6" captionStartId="4.[160,241,1306,1328]" captionTargetBox="[174,1152,248,1288]" captionTargetId="figure-175@4.[160,1168,224,1299]" captionTargetPageId="4" captionText="Figures 4–6. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 4. Dorsal view of head and mesosoma. 5. Detail of propodeum in oblique dorso-posterior view. 6. Lateral view of mesosoma, detailing mesoscutellar profile." figureDoi="http://doi.org/10.5281/zenodo.8057346" httpUri="https://zenodo.org/record/8057346/files/figure.png" pageId="6" pageNumber="7">Fig. 5</figureCitation>
); propodeal spiracle elongate, approximately 4–4.5× as long as wide.
</paragraph>
<paragraph id="4F655F0A216224468990FCAADE46096F" blockId="6.[160,1175,225,1786]" pageId="6" pageNumber="7">
Forewing extending well beyond apex of metasoma (
<figureCitation id="D7E1438F216224468A04FCAADEA70F4F" box="[836,900,801,825]" captionStart="Figures 1–3" captionStartId="3.[160,241,1413,1435]" captionTargetBox="[164,1158,244,1397]" captionTargetId="figure-144@3.[160,1168,224,1407]" captionTargetPageId="3" captionText="Figures 1–3. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 1. Lateral habitus. 2. Facial view. 3. Outer view of mandible (P = preapical tooth)." figureDoi="http://doi.org/10.5281/zenodo.8057344" httpUri="https://zenodo.org/record/8057344/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
), with 2Rs, 1rs-m, 1m-cu, apical half 3M, 4M, apical third 1Cu, 2Cu, 3Cu, and 2cu-a indicated by brownish nebulous traces (
<figureCitation id="D7E1438F21622446882BFCEADC8C0F0F" box="[363,431,865,889]" captionStart="Figures 7–10" captionStartId="5.[160,241,1531,1553]" captionTargetBox="[177,1151,236,1507]" captionTargetId="figure-106@5.[160,1168,224,1525]" captionTargetPageId="5" captionText="Figures 7–10. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 7. Forewing. 8. Outer view of metatibia and metatarsus. 9. Inner view of metatibia and metatarsus with posterior glabrate (yellow), apical glabrate (blue), keirotrichiate (green), and sericeous (pink) areas colored (arrows indicate width of posterior glabrate and length of apical glabrate zones). 10. Unmodified view of inner surface of metatibia and metatarsus." figureDoi="http://doi.org/10.5281/zenodo.8057350" httpUri="https://zenodo.org/record/8057350/files/figure.png" pageId="6" pageNumber="7">Fig. 7</figureCitation>
); fenestrae demarcated by faint white spectral lines on otherwise infuscate wing membrane (lightly infuscate over most of membrane, darker in apical portion of radial cell and in and around marginal cell); membrane with dark brown microtrichia; prestigma short, about twice as long as anterior width of 1Rs; pterostigma slender; marginal cell approximately four times as long as maximum breadth, separated from wing apex by slightly more than its maximum width, with apex narrowly open, opening about one-fifth maximum marginal cell width, with nebulous, angled, appendiculate apex to 4Rs and nebulous 2r-rs (absent in
<taxonomicName id="88DA2489216224468AB5FBC9D9A3082F" baseAuthorityName="Smith" baseAuthorityYear="1865" box="[1013,1152,1090,1113]" class="Insecta" family="Apidae" genus="Trigona" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="atricornis">
<emphasis id="7DAE8318216224468AB5FBC9D929082F" box="[1013,1034,1090,1113]" italics="true" pageId="6" pageNumber="7">H</emphasis>
.
<emphasis id="7DAE8318216224468D58FBC9D9A3082F" box="[1048,1152,1090,1113]" italics="true" pageId="6" pageNumber="7">atricornis</emphasis>
</taxonomicName>
); 1M distad 1cu-a, thus minute M+Cuβ present, shorter than 1cu-a; submarginal angle (
<emphasis id="7DAE83182162244689E9FB09DDE208EF" box="[169,193,1154,1177]" italics="true" pageId="6" pageNumber="7">i.e</emphasis>
., anterior angle between 1Rs and Rs+M), nearly orthogonal; M obtusely angled at 1m-cu (
<emphasis id="7DAE831821622446885CFB29DC1708CF" box="[284,308,1186,1209]" italics="true" pageId="6" pageNumber="7">i.e</emphasis>
., angle between 2M and 3M); 3M tubular in basal half, then nebulous; 2Rs weakly nebulous, angulate; 1rs-m faintly nebulous, almost spectral, straight; r-rs slightly longer than 3Rs. Hind wing with 6–8 distal hamuli (
<quantity id="8822F2EF216224468A22FB6ADEAE088F" box="[866,909,1249,1273]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="6" pageNumber="7" unit="in" value="6.0">6 in</quantity>
<taxonomicName id="88DA2489216224468AD3FB69D93E088F" baseAuthorityName="Smith" baseAuthorityYear="1865" box="[915,1053,1250,1273]" class="Insecta" family="Apidae" genus="Trigona" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="atricornis">
<emphasis id="7DAE8318216224468AD3FB69DE8B088F" box="[915,936,1250,1273]" italics="true" pageId="6" pageNumber="7">H</emphasis>
.
<emphasis id="7DAE8318216224468AF5FB69D93E088F" box="[949,1053,1250,1273]" italics="true" pageId="6" pageNumber="7">atricornis</emphasis>
</taxonomicName>
,
<quantity id="8822F2EF216224468D69FB6AD94D088F" box="[1065,1134,1249,1273]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" metricValueMax="2.032" metricValueMin="1.524" pageId="6" pageNumber="7" unit="in" value="7.0" valueMax="8.0" valueMin="6.0">6–8 in</quantity>
<taxonomicName id="88DA2489216224468D35FB69DC32096F" authorityName="Engel &amp; Rasmussen" authorityYear="2017" class="Insecta" family="Apidae" genus="Heterotrigona" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="paradisaea">
<emphasis id="7DAE8318216224468D35FB69D9A9088F" box="[1141,1162,1250,1273]" italics="true" pageId="6" pageNumber="7">H</emphasis>
.
<emphasis id="7DAE83182162244689E0FA8ADC32096F" box="[160,273,1281,1305]" italics="true" pageId="6" pageNumber="7">paradisaea</emphasis>
</taxonomicName>
); radial and cubital cells closed by nebulous veins.
</paragraph>
<paragraph id="4F655F0A216224468990FAAADCFB0ACF" blockId="6.[160,1175,225,1786]" pageId="6" pageNumber="7">
Metatibia slightly less than three times as long as greatest width, elongate subtriangular; posterior margin gently recurved with subangulate distal angle (
<figureCitation id="D7E1438F216224468D4BFACAD9A7092F" box="[1035,1156,1345,1369]" captionStart="Figures 7–10" captionStartId="5.[160,241,1531,1553]" captionTargetBox="[177,1151,236,1507]" captionTargetId="figure-106@5.[160,1168,224,1525]" captionTargetPageId="5" captionText="Figures 7–10. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 7. Forewing. 8. Outer view of metatibia and metatarsus. 9. Inner view of metatibia and metatarsus with posterior glabrate (yellow), apical glabrate (blue), keirotrichiate (green), and sericeous (pink) areas colored (arrows indicate width of posterior glabrate and length of apical glabrate zones). 10. Unmodified view of inner surface of metatibia and metatarsus." figureDoi="http://doi.org/10.5281/zenodo.8057350" httpUri="https://zenodo.org/record/8057350/files/figure.png" pageId="6" pageNumber="7">Figs. 8–10</figureCitation>
), setae along posterior margin and upper outer surface mostly plumose (
<figureCitation id="D7E1438F216224468D49FAEAD9A7090F" box="[1033,1156,1377,1401]" captionStart="Figures 7–10" captionStartId="5.[160,241,1531,1553]" captionTargetBox="[177,1151,236,1507]" captionTargetId="figure-106@5.[160,1168,224,1525]" captionTargetPageId="5" captionText="Figures 7–10. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 7. Forewing. 8. Outer view of metatibia and metatarsus. 9. Inner view of metatibia and metatarsus with posterior glabrate (yellow), apical glabrate (blue), keirotrichiate (green), and sericeous (pink) areas colored (arrows indicate width of posterior glabrate and length of apical glabrate zones). 10. Unmodified view of inner surface of metatibia and metatarsus." figureDoi="http://doi.org/10.5281/zenodo.8057350" httpUri="https://zenodo.org/record/8057350/files/figure.png" pageId="6" pageNumber="7">Figs. 8–10</figureCitation>
); outer surface weakly concave apically, with corbicula occupying apical third (
<figureCitation id="D7E1438F216224468D7FFA0AD9A209EF" box="[1087,1153,1409,1433]" captionStart="Figures 7–10" captionStartId="5.[160,241,1531,1553]" captionTargetBox="[177,1151,236,1507]" captionTargetId="figure-106@5.[160,1168,224,1525]" captionTargetPageId="5" captionText="Figures 7–10. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 7. Forewing. 8. Outer view of metatibia and metatarsus. 9. Inner view of metatibia and metatarsus with posterior glabrate (yellow), apical glabrate (blue), keirotrichiate (green), and sericeous (pink) areas colored (arrows indicate width of posterior glabrate and length of apical glabrate zones). 10. Unmodified view of inner surface of metatibia and metatarsus." figureDoi="http://doi.org/10.5281/zenodo.8057350" httpUri="https://zenodo.org/record/8057350/files/figure.png" pageId="6" pageNumber="7">Fig. 8</figureCitation>
); apical margin transverse; inner surface with broad, elevated keirotrichiate zone and narrow subglabrous zone (
<figureCitation id="D7E1438F21622446889DFA4ADF7009AF" box="[477,595,1473,1497]" captionStart="Figures 7–10" captionStartId="5.[160,241,1531,1553]" captionTargetBox="[177,1151,236,1507]" captionTargetId="figure-106@5.[160,1168,224,1525]" captionTargetPageId="5" captionText="Figures 7–10. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 7. Forewing. 8. Outer view of metatibia and metatarsus. 9. Inner view of metatibia and metatarsus with posterior glabrate (yellow), apical glabrate (blue), keirotrichiate (green), and sericeous (pink) areas colored (arrows indicate width of posterior glabrate and length of apical glabrate zones). 10. Unmodified view of inner surface of metatibia and metatarsus." figureDoi="http://doi.org/10.5281/zenodo.8057350" httpUri="https://zenodo.org/record/8057350/files/figure.png" pageId="6" pageNumber="7">Figs. 9, 10</figureCitation>
), with abrupt clivulus (
<emphasis id="7DAE8318216224468A28FA49DE8609AF" box="[872,933,1474,1497]" italics="true" pageId="6" pageNumber="7">sensu</emphasis>
<taxonomicName id="88DA2489216224468AECFA49D91209AF" authority="Engel &amp; Rasmussen, 2017" authorityName="Engel &amp; Rasmussen" authorityYear="2017" box="[940,1073,1474,1497]" pageId="6" pageNumber="13" rank="subGenus" status="n. subgen." subGenus="Rasmussen">Rasmussen</taxonomicName>
<emphasis id="7DAE8318216224468D77FA49D94709AF" box="[1079,1124,1473,1497]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
., in press); keirotrichiate area broader than posterior glabrate zone, width of keirotrichiate zone in the subapical region (wider portion) slightly greater than length of apical glabrate zone and narrower than its width (
<figureCitation id="D7E1438F216224468BE2F9AADFC70A4F" box="[674,740,1569,1593]" captionStart="Figures 7–10" captionStartId="5.[160,241,1531,1553]" captionTargetBox="[177,1151,236,1507]" captionTargetId="figure-106@5.[160,1168,224,1525]" captionTargetPageId="5" captionText="Figures 7–10. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 7. Forewing. 8. Outer view of metatibia and metatarsus. 9. Inner view of metatibia and metatarsus with posterior glabrate (yellow), apical glabrate (blue), keirotrichiate (green), and sericeous (pink) areas colored (arrows indicate width of posterior glabrate and length of apical glabrate zones). 10. Unmodified view of inner surface of metatibia and metatarsus." figureDoi="http://doi.org/10.5281/zenodo.8057350" httpUri="https://zenodo.org/record/8057350/files/figure.png" pageId="6" pageNumber="7">Fig. 9</figureCitation>
); penicillus and rastellar comb present, each composed of stiff setae. Metabasitarsus weakly trapezoidal, with posterior margin gently arched, distal angle not projecting (
<figureCitation id="D7E1438F216224468BB6F9EADE4D0A0F" box="[758,878,1633,1657]" captionStart="Figures 7–10" captionStartId="5.[160,241,1531,1553]" captionTargetBox="[177,1151,236,1507]" captionTargetId="figure-106@5.[160,1168,224,1525]" captionTargetPageId="5" captionText="Figures 7–10. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 7. Forewing. 8. Outer view of metatibia and metatarsus. 9. Inner view of metatibia and metatarsus with posterior glabrate (yellow), apical glabrate (blue), keirotrichiate (green), and sericeous (pink) areas colored (arrows indicate width of posterior glabrate and length of apical glabrate zones). 10. Unmodified view of inner surface of metatibia and metatarsus." figureDoi="http://doi.org/10.5281/zenodo.8057350" httpUri="https://zenodo.org/record/8057350/files/figure.png" pageId="6" pageNumber="7">Figs. 8–10</figureCitation>
); outer surface with fine basal posterior fimbriate field bordering depression; inner surface with short basal sericeous area (
<figureCitation id="D7E1438F216224468815F92ADCEF0ACF" box="[341,460,1697,1721]" captionStart="Figures 7–10" captionStartId="5.[160,241,1531,1553]" captionTargetBox="[177,1151,236,1507]" captionTargetId="figure-106@5.[160,1168,224,1525]" captionTargetPageId="5" captionText="Figures 7–10. Worker of Heterotrigona (Sahulotrigona) paradisaea, new subgenus and species, from eastern Papua New Guinea. 7. Forewing. 8. Outer view of metatibia and metatarsus. 9. Inner view of metatibia and metatarsus with posterior glabrate (yellow), apical glabrate (blue), keirotrichiate (green), and sericeous (pink) areas colored (arrows indicate width of posterior glabrate and length of apical glabrate zones). 10. Unmodified view of inner surface of metatibia and metatarsus." figureDoi="http://doi.org/10.5281/zenodo.8057350" httpUri="https://zenodo.org/record/8057350/files/figure.png" pageId="6" pageNumber="7">Figs. 9, 10</figureCitation>
).
</paragraph>
<paragraph id="4F655F0A216224478990F949D9580D6F" blockId="6.[160,1175,225,1786]" lastBlockId="7.[160,1168,225,505]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">Metasoma narrow, with first metasomal tergum smooth and shining, second through fourth terga largely smooth and shining to faintly imbricate except exceedingly narrow apical marginal zones imbricate with appressed, minute, apically directed setae; postgradular surface of sixth tergum with short, erect, fuscous plumose setae.</paragraph>
</subSubSection>
<subSubSection id="07C00C81216324478990FEAADF360DEF" pageId="7" pageNumber="8" type="etymology">
<paragraph id="4F655F0A216324478990FEAADF360DEF" blockId="7.[160,1168,225,505]" pageId="7" pageNumber="8">
ETYMOLOGY: The subgeneric name is a combination of Sahul, a reference to the occurrence of the known species within a distinct zoogeographic region relative to those
<taxonomicName id="88DA24892163244789A6FEE9DC5E0D0F" authorityName="Schwarz" authorityYear="1939" box="[230,381,354,377]" class="Insecta" family="Apidae" genus="Heterotrigona" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="7DAE83182163244789A6FEE9DC5E0D0F" box="[230,381,354,377]" italics="true" pageId="7" pageNumber="8">Heterotrigona</emphasis>
</taxonomicName>
occurring in Sundaland, and the generic name
<taxonomicName id="88DA2489216324478AEBFEE9D9750D0F" authority="Jurine." authorityName="Jurine." box="[939,1110,353,377]" class="Insecta" family="Apidae" genus="Trigona" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="7DAE8318216324478AEBFEE9D9220D0F" box="[939,1025,354,377]" italics="true" pageId="7" pageNumber="8">Trigona</emphasis>
Jurine.
</taxonomicName>
The gender of the name is feminine.
</paragraph>
</subSubSection>
<subSubSection id="07C00C81216324478990FE2ADC4E0D8F" pageId="7" pageNumber="8" type="discussion">
<paragraph id="4F655F0A216324478990FE2ADC4E0D8F" blockId="7.[160,1168,225,505]" pageId="7" pageNumber="8">
INCLUDED SPECIES: The subgenus as currently constituted contains only two species, both from New
<collectingCountry id="37CD1F9A2163244788D6FE4ADCCC0DAF" box="[406,495,449,473]" name="Guinea-Bissau" pageId="7" pageNumber="8">Guinea</collectingCountry>
:
<taxonomicName id="88DA24892163244788B9FE49DE9A0DAF" authority="Engel &amp; Rasmussen, 2017" authorityName="Engel &amp; Rasmussen" authorityYear="2017" box="[505,953,449,473]" class="Insecta" family="Apidae" genus="Heterotrigona" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="paradisaea" status="n. sp." subGenus="Sahulotrigona">
<emphasis id="7DAE83182163244788B9FE49DFB30DAF" box="[505,656,450,473]" italics="true" pageId="7" pageNumber="8">Heterotrigona</emphasis>
(
<emphasis id="7DAE8318216324478BE0FE4ADE1A0DAF" box="[672,825,449,473]" italics="true" pageId="7" pageNumber="8">Sahulotrigona</emphasis>
)
<emphasis id="7DAE8318216324478A08FE4ADE9A0DAF" box="[840,953,449,473]" italics="true" pageId="7" pageNumber="8">paradisaea</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="669D3E63216324478A86FE49D9250DAF" box="[966,1030,450,473]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
, and
<emphasis id="7DAE8318216324478D07FE49D97F0DAF" box="[1095,1116,450,473]" italics="true" pageId="7" pageNumber="8">H</emphasis>
.
<taxonomicName id="88DA2489216324478D29FE4ADC4A0D8F" authority="(Smith)" baseAuthorityName="Smith" baseAuthorityYear="1865" class="Insecta" family="Apidae" genus="Trigona" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="atricornis" subGenus="Sahulotrigona">
(
<emphasis id="7DAE8318216324478D32FE49D9A20DAF" box="[1138,1153,450,473]" italics="true" pageId="7" pageNumber="8">S</emphasis>
.)
<emphasis id="7DAE83182163244789E0FE69DC2B0D8F" box="[160,264,482,505]" italics="true" pageId="7" pageNumber="8">atricornis</emphasis>
(Smith)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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