817 lines
105 KiB
XML
817 lines
105 KiB
XML
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<mods:title id="78AD3BE46741EB66725ECB5AE87070C8">New species, revision, and phylogeny of Ronzotherium Aymard, 1854 (Perissodactyla, Rhinocerotidae)</mods:title>
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<mods:namePart id="25C3A9494C77320F3D0AEE69D85E254F">Tissier, Jérémy</mods:namePart>
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<mods:affiliation id="A965B490CBE954C9576E5ECFD15C1ECC">Route de Fontenais 21, JURASSICA Museum, 2900 Porrentruy, Switzerland & Chemin du musée 4, Université de Fribourg, Department of Geosciences, 1700 Fribourg, Switzerland.</mods:affiliation>
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<mods:namePart id="507EF39FECAEE9E769BC508620A561F0">Antoine, Pierre-Olivier</mods:namePart>
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<mods:affiliation id="107CCBD34A0D652A866671E4EEE9765C">Place Eugène Bataillon, Institut des sciences de l’évolution de Montpellier-CNRS-IRD-RPHE, Université de Montpellier, 34095 Montpellier, France.</mods:affiliation>
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<mods:nameIdentifier id="F3BD7AE35680BC850F97D681F4AFD616" type="email">pierre-olivier.antoine@umontpellier.fr</mods:nameIdentifier>
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<mods:namePart id="F9F726BF966638AD04ABC4CD45D92151">Becker, Damien</mods:namePart>
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<mods:affiliation id="45F87BA91544677858997B44E3E58442">Route de Fontenais 21, JURASSICA Museum, 2900 Porrentruy, Switzerland & Chemin du musée 4, Université de Fribourg, Department of Geosciences, 1700 Fribourg, Switzerland.</mods:affiliation>
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<mods:nameIdentifier id="9A89506DAD8E86729892D89C1742E0CC" type="email">damien.becker@jurassica.ch</mods:nameIdentifier>
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<mods:title id="7EB0D279EAF12A00F2EA8EA2E90C7C55">European Journal of Taxonomy</mods:title>
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<mods:date id="6FD7DD1799060A4180E730E67CE4B033">2021</mods:date>
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<treatment id="C53FFD4DD73EFFE4FDF2D9221BC0A4AE" ID-DOI="http://doi.org/10.5281/zenodo.4959444" ID-GBIF-Taxon="182751316" ID-Zenodo-Dep="4959444" LSID="urn:lsid:zoobank.org:act:D1CFA5AE-6BC9-479B-AE43-F46EB9D86A49" httpUri="http://treatment.plazi.org/id/C53FFD4DD73EFFE4FDF2D9221BC0A4AE" lastPageId="67" lastPageNumber="68" pageId="58" pageNumber="59">
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<taxonomicName id="8A9637D8D73EFF9DFDF2D9221926A011" authority="Tissier & Antoine & Becker, 2021" authorityName="Tissier & Antoine & Becker" authorityYear="2021" box="[608,882,264,291]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="heissigi" status="sp. nov.">
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<emphasis id="7FE29049D73EFF9DFDF2D9221926A011" bold="true" box="[608,882,264,291]" italics="true" pageId="58" pageNumber="59">Ronzotherium heissigi</emphasis>
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<taxonomicNameLabel id="64D12D32D73EFF9DFCE8D9231980A011" box="[890,980,265,291]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="058C1FD0D73EFF9DFD4AD978190FA05E" box="[728,859,338,364]" pageId="58" pageNumber="59" type="description">
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<paragraph id="4D294C5BD73EFF9DFD4AD978190FA05E" blockId="58.[355,1232,264,364]" box="[728,859,338,364]" pageId="58" pageNumber="59">
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<figureCitation id="D5AD50DED73EFF9DFD4AD978190FA05E" box="[728,859,338,364]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="60.[189,232,1963,1989]" captionStartId-1="61.[189,232,1822,1848]" captionStartId-2="62.[189,232,1893,1919]" captionStartId-3="63.[189,232,1787,1813]" captionStartId-4="65.[188,231,1680,1706]" captionTargetBox-0="[194,1394,265,1940]" captionTargetBox-1="[252,1331,269,1782]" captionTargetBox-2="[227,1360,265,1868]" captionTargetId-0="figure-44@60.[194,1394,265,1940]" captionTargetId-1="figure-100@61.[239,1349,251,1797]" captionTargetId-2="figure-13@62.[227,1360,265,1868]" captionTargetId-3="figure-13@64.[189,1398,265,1957]" captionTargetId-4="figure-13@66.[189,1399,265,1975]" captionTargetPageId-0="60" captionTargetPageId-1="61" captionTargetPageId-2="62" captionTargetPageId-3="64" captionTargetPageId-4="66" captionText-0="Fig. 22. Ronzotherium heissigi sp. nov. from Vendèze (late early Oligocene, France). Skull MNHN.F.LIM181. A. Dorsal view. B. Left lateral view. C. Ventral view. Scale bar: 2 cm." captionText-1="Fig. 23. Ronzotherium heissigi sp. nov. from Vendèze (late early Oligocene, France). Drawing of the skull MNHN.F.LIM181. A. Dorsal view. B. Left lateral view. C. Ventral view. Abbreviations: AT = articular tubercle; BO = basioccipital; F = frontal; IOF = infraorbital foramen; J = jugal; M = maxilla; NT = nuchal tubercle; OC = occipital condyle; P = parietal; PGA = postglenoid apophysis; PMX = premaxilla; PoP = postorbital process; PP = paraoccipital process; SC = sagittal crest; SQ = squamosal. Scale bar: 2 cm." captionText-2="Fig. 24. Ronzotherium heissigi sp. nov. from Vendèze (late early Oligocene, France). A–D. Mandible MNHN.F.LIM181. A. Occlusal view. B. Lateral view. C. With left p2–p4 and m2–3 in labial view. View. D. Right p2–m3 in lingual view. – E. Skull MNHN.F.LIM181, belonging to the same individual as the mandible, close-up view of P2–M3 in occlusal view. Scale bars: 2 cm." captionText-3="Fig. 25 (next page). Ronzotherium heissigi sp. nov. from Bumbach, Bern Canton (early late Oligocene; Switzerland). – A–D. Mandible NMB-UM-6132. A. Occlusal view. B. Left lateral view. C. With right p3–m3 in labial view. D. With right p3–m3 in lingual view. – E–F. Right humerus NMB-UM-129a. E. Anterior view.F. Posterior view. –G–H. Left humerus NMB-UM-132. G. Anterior view.H. Posterior view. – I–J. Right radius NMB-UM-127a. I. Anterior view.J. Posterior view. – K–M. Right ulna NMB- UM-131b. K. Anterior view. L. Posterior view. M. Distal view. Abbreviation: im = insertion for the m. biceps brachii. Scale bars: 2 cm." captionText-4="Fig. 26 (next page). Ronzotherium heissigi sp. nov. from Bumbach, Bern Canton (early late Oligocene; Switzerland). – A–E. Left lunate NMBE-5035833. A. Anterior view. B. Proximal view. C. Distal view. D. Lateral view. E. Medial view. – F–J. Right trapezoid NMB-UM-6136b. F. Anterior view. G. Lateral view. H. Medial view. I. Proximal view. J. Distal view. – K–N. Right magnum NMB-UM- 6136c. K. Anterior view. L. Distal view. M. Lateral view. N. Medial view. – O–Q. Right McII NMB- UM-121. O. Anterior view. P. Lateral view. Q. Proximal view. – R–T. Right McII NMB-UM-6136a. R. Proximal view. S. Anterior view. T. Lateral view. – U–Y. Right cuboid NMBE-5035834. U. Anterior view. V. Proximal view. W. Distal view. X. Medial view. Y. Lateral view. Abbreviations: a = astragalus; ca = calcaneus; e = ectocuneiform; l = lunate; m = magnum; n = navicular; p = pyramidal; r = radius; s = scaphoid; td = trapezoid; tm = trapezium; un = unciform. Scale bar: 2 cm." figureDoi-0="http://doi.org/10.5281/zenodo.4945922" figureDoi-1="http://doi.org/10.5281/zenodo.4945926" figureDoi-2="http://doi.org/10.5281/zenodo.4945928" figureDoi-3="http://doi.org/10.5281/zenodo.4959442" figureDoi-4="http://doi.org/10.5281/zenodo.4959446" httpUri-0="https://zenodo.org/record/4945922/files/figure.png" httpUri-1="https://zenodo.org/record/4945926/files/figure.png" httpUri-2="https://zenodo.org/record/4945928/files/figure.png" httpUri-3="https://zenodo.org/record/4959442/files/figure.png" httpUri-4="https://zenodo.org/record/4959446/files/figure.png" pageId="58" pageNumber="59">Figs 22–26</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="058C1FD0D73EFF9DFF2FD9BE1E90A3E6" pageId="58" pageNumber="59" type="reference_group">
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<paragraph id="4D294C5BD73EFF9DFF2FD9BE1EFAA09D" blockId="58.[189,1398,404,724]" box="[189,1198,404,431]" pageId="58" pageNumber="59">
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<treatmentCitationGroup id="6D866B75D73EFF9DFF2FD9BE1EFAA09D" box="[189,1198,404,431]" pageId="58" pageNumber="59">
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<taxonomicName id="8A9637D8D73EFF9DFF2FD9BE1BB7A09C" baseAuthorityName="Pomel" baseAuthorityYear="1853" box="[189,483,404,430]" class="Mammalia" family="Rhinocerotidae" genus="Acerotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="lemanense">
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<emphasis id="7FE29049D73EFF9DFF2FD9BE1BB7A09C" box="[189,483,404,430]" italics="true" pageId="58" pageNumber="59">Acerotherium lemanense</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="CC376A4AD73EFF9DFD93D9BF1954A09C" author="Roman F." box="[513,768,404,431]" page="61" pageId="58" pageNumber="59" year="1912">
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<bibRefCitation id="290731AAD73EFF9DFD93D9BF1954A09C" author="Roman F." box="[513,768,404,431]" pageId="58" pageNumber="59" pagination="1 - 92" refId="ref42112" refString="Roman F. 1912 a. Les Rhinocerides de l'Oligocene d'Europe. Archives du Museum d'histoire naturelle de Lyon 11: 1 - 92." type="journal article" year="1912">Roman 1912a: 61–62</bibRefCitation>
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</treatmentCitation>
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, pl. VII (from Lamothe-Capdeville).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="4D294C5BD73EFF9DFF2FD993190EA0E6" blockId="58.[189,1398,404,724]" box="[189,858,441,468]" pageId="58" pageNumber="59">
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<treatmentCitationGroup id="6D866B75D73EFF9DFF2FD993190EA0E6" box="[189,858,441,468]" pageId="58" pageNumber="59">
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<taxonomicName id="8A9637D8D73EFF9DFF2FD9931BF8A0E1" authorityName="Osborn" authorityYear="1900" box="[189,428,441,467]" class="Mammalia" family="Rhinocerotidae" genus="Aceratherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="filholi">
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<emphasis id="7FE29049D73EFF9DFF2FD9931BF8A0E1" box="[189,428,441,467]" italics="true" pageId="58" pageNumber="59">Aceratherium filholi</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="CC376A4AD73EFF9DFE58D99318C9A0E1" author="Stehlin H. G." box="[458,669,441,468]" page="183" pageId="58" pageNumber="59" year="1914">
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||
<bibRefCitation id="290731AAD73EFF9DFE58D99318C9A0E1" author="Stehlin H. G." box="[458,669,441,468]" pageId="58" pageNumber="59" pagination="179 - 202" refId="ref42499" refString="Stehlin H. G. 1914. Ubersicht uber die Saugetiere der schweizerischen Molasseformation, ihre Fundorte und ihre stratigraphische Verbreitung. Verhandlungen der Naturforschenden Gesellschaft in Basel 25: 179 - 202." type="journal article" year="1914">Stehlin 1914: 183</bibRefCitation>
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</treatmentCitation>
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||
, 85 (Bumbach).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
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||
<paragraph id="4D294C5BD73EFF9DFF2FD9F4197CA32E" blockId="58.[189,1398,404,724]" pageId="58" pageNumber="59">
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<treatmentCitationGroup id="6D866B75D73EFF9DFF2FD9F4197CA32E" pageId="58" pageNumber="59">
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<taxonomicName id="8A9637D8D73EFF9DFF2FD9F41BE5A0CA" baseAuthorityName="Brunet" baseAuthorityYear="1979" box="[189,433,478,504]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="filholi">
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||
<emphasis id="7FE29049D73EFF9DFF2FD9F41BE5A0CA" box="[189,433,478,504]" italics="true" pageId="58" pageNumber="59">Ronzotherium filholi</emphasis>
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||
</taxonomicName>
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||
–
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<treatmentCitation id="CC376A4AD73EFF9DFE43D9F418E6A0CA" author="Lavocat R." box="[465,690,478,505]" page="116" pageId="58" pageNumber="59" year="1951">
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||
<bibRefCitation id="290731AAD73EFF9DFE43D9F418E6A0CA" author="Lavocat R." box="[465,690,478,505]" pageId="58" pageNumber="59" refId="ref41246" refString="Lavocat R. 1951. Revision de la Faune des Mammiferes oligocenes d'Auvergne et du Velay. Sciences et Avenir, Paris." type="book" year="1951">Lavocat 1951: 116</bibRefCitation>
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||
</treatmentCitation>
|
||
, pl. 19 fig. 3, pl. 26 fig. 1 (from Vendèze). —
|
||
<treatmentCitation id="CC376A4AD73EFF9DFB49D9F41B42A32F" author="Brunet M." page="105" pageId="58" pageNumber="59" year="1979">
|
||
<bibRefCitation id="290731AAD73EFF9DFB49D9F41B42A32F" author="Brunet M." pageId="58" pageNumber="59" refId="ref40079" refString="Brunet M. 1979. Les grands mammiferes chefs de file de l'immigration Oligocene et le probleme de la limite Eocene - Oligocene en Europe. Fondation Singer-Polignac, Paris." type="book" year="1979">Brunet 1979: 105</bibRefCitation>
|
||
</treatmentCitation>
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||
(from Bumbach). —
|
||
<treatmentCitation id="CC376A4AD73EFF9DFD87DA281970A32E" author="Becker D." box="[533,804,514,541]" page="213" pageId="58" pageNumber="59" year="2003">
|
||
<bibRefCitation id="290731AAD73EFF9DFD87DA281970A32E" author="Becker D." box="[533,804,514,541]" pageId="58" pageNumber="59" pagination="1 - 327" refId="ref39491" refString="Becker D. 2003. Paleoecologie et paleoclimat de la Molasse du Jura (Oligo - Miocene): apport des Rhinocerotoidea (Mammalia) et des mineraux argileux. GeoFocus 9: 1 - 327." type="journal article" year="2003">Becker 2003: 213–214</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73EFF9DFF2FDA0D1E6EA370" blockId="58.[189,1398,404,724]" box="[189,1082,551,578]" pageId="58" pageNumber="59">
|
||
<treatmentCitationGroup id="6D866B75D73EFF9DFF2FDA0D1E6EA370" box="[189,1082,551,578]" pageId="58" pageNumber="59">
|
||
<taxonomicName id="8A9637D8D73EFF9DFF2FDA0D1B8AA373" box="[189,478,551,577]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="velaunum">
|
||
<emphasis id="7FE29049D73EFF9DFF2FDA0D1B8AA373" box="[189,478,551,577]" italics="true" pageId="58" pageNumber="59">Ronzotherium velaunum</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="CC376A4AD73EFF9DFE6EDA0D18A5A373" author="Heissig K." box="[508,753,551,578]" page="20" pageId="58" pageNumber="59" year="1969">
|
||
<bibRefCitation id="290731AAD73EFF9DFE6EDA0D18A5A373" author="Heissig K." box="[508,753,551,578]" pageId="58" pageNumber="59" refId="ref40927" refString="Heissig K. 1969. Die Rhinocerotidae (Mammalia) aus der oberoligozanen Spaltenfullung von Gaimersheim bei Ingolstadt in Bayern und ihre phylogenetische Stellung. Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse 138. Bavarian Academy of Sciences, Munich." type="book" year="1969">Heissig 1969: 20–36</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 77, fig. 8b (from Vendèze).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73EFF9DFF2FDA611EA0A354" blockId="58.[189,1398,404,724]" box="[189,1268,587,614]" pageId="58" pageNumber="59">
|
||
<treatmentCitationGroup id="6D866B75D73EFF9DFF2FDA611EA0A354" box="[189,1268,587,614]" pageId="58" pageNumber="59">
|
||
<taxonomicName id="8A9637D8D73EFF9DFF2FDA611861A357" authorityName="Heissig" authorityYear="1969" box="[189,565,587,613]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="58" pageNumber="59" phylum="Chordata" rank="subSpecies" species="filholi" subSpecies="elongatum">
|
||
<emphasis id="7FE29049D73EFF9DFF2FDA611861A357" box="[189,565,587,613]" italics="true" pageId="58" pageNumber="59">Ronzotherium filholi elongatum</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="CC376A4AD73EFF9DFDC0DA61191CA354" author="Heissig K." box="[594,840,587,614]" page="46" pageId="58" pageNumber="59" year="1969">
|
||
<bibRefCitation id="290731AAD73EFF9DFDC0DA61191CA354" author="Heissig K." box="[594,840,587,614]" pageId="58" pageNumber="59" refId="ref40927" refString="Heissig K. 1969. Die Rhinocerotidae (Mammalia) aus der oberoligozanen Spaltenfullung von Gaimersheim bei Ingolstadt in Bayern und ihre phylogenetische Stellung. Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse 138. Bavarian Academy of Sciences, Munich." type="book" year="1969">Heissig 1969: 46–55</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 71, 75–77, 82–83 (from Bumbach).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73EFF9DFF2FDA5A1E11A3B9" blockId="58.[189,1398,404,724]" box="[189,1093,624,651]" pageId="58" pageNumber="59">
|
||
<treatmentCitationGroup id="6D866B75D73EFF9DFF2FDA5A1E11A3B9" box="[189,1093,624,651]" pageId="58" pageNumber="59">
|
||
<taxonomicName id="8A9637D8D73EFF9DFF2FDA5A1858A3B8" box="[189,524,624,650]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="58" pageNumber="59" phylum="Chordata" rank="subSpecies" species="filholi" subSpecies="romani">
|
||
<emphasis id="7FE29049D73EFF9DFF2FDA5A1858A3B8" box="[189,524,624,650]" italics="true" pageId="58" pageNumber="59">Ronzotherium filholi romani</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="CC376A4AD73EFF9DFDB8DA5A18A4A3B8" author="Heissig K." box="[554,752,624,651]" page="63" pageId="58" pageNumber="59" year="1969">
|
||
<bibRefCitation id="290731AAD73EFF9DFDB8DA5A18A4A3B8" author="Heissig K." box="[554,752,624,651]" pageId="58" pageNumber="59" refId="ref40927" refString="Heissig K. 1969. Die Rhinocerotidae (Mammalia) aus der oberoligozanen Spaltenfullung von Gaimersheim bei Ingolstadt in Bayern und ihre phylogenetische Stellung. Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse 138. Bavarian Academy of Sciences, Munich." type="book" year="1969">Heissig 1969: 63</bibRefCitation>
|
||
</treatmentCitation>
|
||
(from Lamothe-Capdeville).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73EFF9DFF2FDABF1E85A382" blockId="58.[189,1398,404,724]" box="[189,1233,661,688]" pageId="58" pageNumber="59">
|
||
<treatmentCitationGroup id="6D866B75D73EFF9DFF2FDABF1E85A382" box="[189,1233,661,688]" pageId="58" pageNumber="59">
|
||
<taxonomicName id="8A9637D8D73EFF9DFF2FDABF1BEBA39D" authorityName="Kretzoi" authorityYear="1940" box="[189,447,661,687]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="romani">
|
||
<emphasis id="7FE29049D73EFF9DFF2FDABF1BEBA39D" box="[189,447,661,687]" italics="true" pageId="58" pageNumber="59">Ronzotherium romani</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="CC376A4AD73EFF9DFE4FDABF18B3A382" author="Brunet M." box="[477,743,661,688]" page="135" pageId="58" pageNumber="59" year="1979">
|
||
<bibRefCitation id="290731AAD73EFF9DFE4FDABF18B3A382" author="Brunet M." box="[477,743,661,688]" pageId="58" pageNumber="59" refId="ref40079" refString="Brunet M. 1979. Les grands mammiferes chefs de file de l'immigration Oligocene et le probleme de la limite Eocene - Oligocene en Europe. Fondation Singer-Polignac, Paris." type="book" year="1979">Brunet 1979: 135–136</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 15, pls XVII–XVIII (from Vendèze).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73EFF9DFF2FDA931E90A3E6" blockId="58.[189,1398,404,724]" box="[189,1220,697,724]" pageId="58" pageNumber="59">
|
||
<treatmentCitationGroup id="6D866B75D73EFF9DFF2FDA931E90A3E6" box="[189,1220,697,724]" pageId="58" pageNumber="59">
|
||
<taxonomicName id="8A9637D8D73EFF9DFF2FDA931BABA3E1" baseAuthorityName="Pomel" baseAuthorityYear="1853" box="[189,511,697,723]" class="Mammalia" family="Rhinocerotidae" genus="Diaceratherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="lemanense">
|
||
<emphasis id="7FE29049D73EFF9DFF2FDA931BABA3E1" box="[189,511,697,723]" italics="true" pageId="58" pageNumber="59">Diaceratherium lemanense</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="CC376A4AD73EFF9DFD89DA93193BA3E6" author="Antoine P. - O. & Becker D." box="[539,879,697,724]" page="140" pageId="58" pageNumber="59" year="2013">
|
||
<bibRefCitation id="290731AAD73EFF9DFD89DA93193BA3E6" author="Antoine P. - O. & Becker D." box="[539,879,697,724]" pageId="58" pageNumber="59" pagination="135 - 146" refId="ref39152" refString="Antoine P. - O. & Becker D. 2013. A brief review of Agenian rhinocerotids in Western Europe. Swiss Journal of Geosciences 106 (2): 135 - 146. https: // doi. org / 10.1007 / s 00015 - 013 - 0126 - 8" type="journal article" year="2013">Antoine & Becker 2013: 140</bibRefCitation>
|
||
</treatmentCitation>
|
||
(from Lamothe-Capdeville).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="058C1FD0D73EFF9DFF2FDAD619ADA4D6" pageId="58" pageNumber="59" type="diagnosis">
|
||
<paragraph id="4D294C5BD73EFF9DFF2FDAD61B63A224" blockId="58.[189,311,764,790]" box="[189,311,764,790]" pageId="58" pageNumber="59">
|
||
<heading id="1661FB37D73EFF9DFF2FDAD61B63A224" bold="true" box="[189,311,764,790]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
|
||
<emphasis id="7FE29049D73EFF9DFF2FDAD61B63A224" bold="true" box="[189,311,764,790]" pageId="58" pageNumber="59">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73EFF9DFF2FDB061F22A2EB" blockId="58.[188,1400,812,985]" pageId="58" pageNumber="59">
|
||
Differs from
|
||
<taxonomicName id="8A9637D8D73EFF9DFEC6DB071B9CA274" authorityName="Kretzoi" authorityYear="1940" box="[340,456,813,838]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="romani">
|
||
<emphasis id="7FE29049D73EFF9DFEC6DB071B9CA274" box="[340,456,813,838]" italics="true" pageId="58" pageNumber="59">R. romani</emphasis>
|
||
</taxonomicName>
|
||
by the mandibular ramus inclined forward, the P1 sometimes absent and without anterolingual cingulum, the angular and V-shaped external groove of the lower cheek teeth, the lower premolars without lingual cingulum, the d/p1 always absent in the adult, the deep and wide gutter for the m. extensor carpi on the radius, the concave proximal border of the anterior side of the magnum, the salient insertion of the m. extensor carpalis of the metacarpals and the oval proximal side of the cuboid.
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73EFF9DFF2FDC2B1F05A572" blockId="58.[188,1398,1025,1089]" pageId="58" pageNumber="59">
|
||
Differs from
|
||
<taxonomicName id="8A9637D8D73EFF9DFEC4DC281BBDA529" box="[342,489,1025,1051]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="velaunum">
|
||
<emphasis id="7FE29049D73EFF9DFEC4DC281BBDA529" box="[342,489,1025,1051]" italics="true" pageId="58" pageNumber="59">R. velaunum</emphasis>
|
||
</taxonomicName>
|
||
by the presence of a lingual groove on the corpus mandibulae, the curved and not constricted paralophid on p2 and the deep median constriction of the distal humeral articulation.
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73EFF9DFF2FDC431BAAA5C3" blockId="58.[188,1398,1129,1265]" pageId="58" pageNumber="59">
|
||
Differs from
|
||
<taxonomicName id="8A9637D8D73EFF9DFEC6DC401BEDA5B1" baseAuthorityName="Brunet" baseAuthorityYear="1979" box="[340,441,1129,1155]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="filholi">
|
||
<emphasis id="7FE29049D73EFF9DFEC6DC401BEDA5B1" box="[340,441,1129,1155]" italics="true" pageId="58" pageNumber="59">R. filholi</emphasis>
|
||
</taxonomicName>
|
||
by a foramen infraorbitalis above P3, a zygomatic width/frontal width ratio above 1.5, a concave occipital crest, the reduced paraconid on p2, the high posterior expansion of the scaphoid facet on the radius, the open angle between the diaphysis of the ulna and the olecranon and the curved magnum facet on the McII.
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73EFF9DFF2FDD33184CA44F" blockId="58.[189,1398,1305,1405]" pageId="58" pageNumber="59">
|
||
Differs from
|
||
<taxonomicName id="8A9637D8D73EFF9DFECDDD301856A401" authorityName="Heissig" authorityYear="1969" box="[351,514,1305,1331]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="elongatum">
|
||
<emphasis id="7FE29049D73EFF9DFECDDD301856A401" box="[351,514,1305,1331]" italics="true" pageId="58" pageNumber="59">R. elongatum</emphasis>
|
||
</taxonomicName>
|
||
by the absence of processus lacrymalis, the reduction of the postorbital process on the zygomatic arch, its poorly developed processus posttympanicus and by the metaloph of P2 directed postero-lingually.
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73EFF9DFF2FDD8C19ADA4D6" blockId="58.[188,1399,1446,1509]" pageId="58" pageNumber="59">
|
||
Further differs from
|
||
<taxonomicName id="8A9637D8D73EFF9DFE39DD8C1845A4F2" baseAuthorityName="Brunet" baseAuthorityYear="1979" box="[427,529,1446,1472]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="filholi">
|
||
<emphasis id="7FE29049D73EFF9DFE39DD8C1845A4F2" box="[427,529,1446,1472]" italics="true" pageId="58" pageNumber="59">R. filholi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8A9637D8D73EFF9DFDDBDD8C18B2A4F2" authorityName="Heissig" authorityYear="1969" box="[585,742,1446,1472]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="58" pageNumber="59" phylum="Chordata" rank="species" species="elongatum">
|
||
<emphasis id="7FE29049D73EFF9DFDDBDD8C18B2A4F2" box="[585,742,1446,1472]" italics="true" pageId="58" pageNumber="59">R. elongatum</emphasis>
|
||
</taxonomicName>
|
||
by a convex processus postglenoidalis of the squamosal and by a narrow and V-shaped lingual opening of the lower premolars.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="058C1FD0D73EFF9DFF2FDE271B0DA793" pageId="58" pageNumber="59" type="etymology">
|
||
<paragraph id="4D294C5BD73EFF9DFF2FDE271B11A715" blockId="58.[189,325,1549,1575]" box="[189,325,1549,1575]" pageId="58" pageNumber="59">
|
||
<heading id="1661FB37D73EFF9DFF2FDE271B11A715" bold="true" box="[189,325,1549,1575]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
|
||
<emphasis id="7FE29049D73EFF9DFF2FDE271B11A715" bold="true" box="[189,325,1549,1575]" pageId="58" pageNumber="59">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73EFF9DFF2FDE171B0DA793" blockId="58.[189,1399,1597,1697]" pageId="58" pageNumber="59">
|
||
The specific epithet honours Prof. Dr Kurt Heissig for his major and imperishable contributions on the study of the
|
||
<taxonomicName id="8A9637D8D73EFF9DFEDDDE481851A74E" authorityName="Gray" authorityYear="1821" box="[335,517,1634,1660]" class="Mammalia" family="Rhinocerotidae" kingdom="Animalia" order="Perissodactyla" pageId="58" pageNumber="59" phylum="Chordata" rank="family">Rhinocerotidae</taxonomicName>
|
||
, and for providing the first systematic revision on
|
||
<taxonomicName id="8A9637D8D73EFF9DFBC1DE481EADA74E" baseAuthorityName="Menouret & Guerin" baseAuthorityYear="2009" box="[1107,1273,1634,1660]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="58" pageNumber="59" phylum="Chordata" rank="genus">
|
||
<emphasis id="7FE29049D73EFF9DFBC1DE481EADA74E" box="[1107,1273,1634,1660]" italics="true" pageId="58" pageNumber="59">Ronzotherium</emphasis>
|
||
</taxonomicName>
|
||
more than 40 years ago.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="058C1FD0D73EFF9CFF2FDEE31884A56D" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59" type="materials_examined">
|
||
<paragraph id="4D294C5BD73EFF9DFF2FDEE31B39A7D1" blockId="58.[189,365,1737,1763]" box="[189,365,1737,1763]" pageId="58" pageNumber="59">
|
||
<heading id="1661FB37D73EFF9DFF2FDEE31B39A7D1" bold="true" box="[189,365,1737,1763]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
|
||
<emphasis id="7FE29049D73EFF9DFF2FDEE31B39A7D1" bold="true" box="[189,365,1737,1763]" pageId="58" pageNumber="59">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73EFF9DFF78DED31B09A621" blockId="58.[189,1398,1785,1885]" box="[234,349,1785,1811]" pageId="58" pageNumber="59">
|
||
<emphasis id="7FE29049D73EFF9DFF78DED31B09A621" bold="true" box="[234,349,1785,1811]" pageId="58" pageNumber="59">
|
||
<typeStatus id="922DF2F9D73EFF9DFF78DED31B09A621" box="[234,349,1785,1811]" pageId="58" pageNumber="59" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73EFF9DFF2FDF34191EA66E" blockId="58.[189,1398,1785,1885]" pageId="58" pageNumber="59">
|
||
<materialsCitation id="FDFE4606D73EFF9DFF2FDF34191EA66E" ID-GBIF-Occurrence="3302594302" collectionCode="MNHN" country="France" county="Flour" latitude="45.043556" location="Cantal" longLatPrecision="1" longitude="3.103639" municipality="Vendeze" pageId="58" pageNumber="59" specimenCount="1" typeStatus="holotype">
|
||
<collectingCountry id="35810CCBD73EFF9DFF2FDF341B65A60A" box="[189,305,1822,1848]" name="France" pageId="58" pageNumber="59">FRANCE</collectingCountry>
|
||
• complete skull and associated mandible; Auvergne-Rhône-Alpes,
|
||
<location id="48491A80D73EFF9DFBC6DF341EF0A60A" LSID="urn:lsid:plazi:treatment:C53FFD4DD73EFFE4FDF2D9221BC0A4AE:48491A80D73EFF9DFBC6DF341EF0A60A" box="[1108,1188,1822,1848]" country="France" county="Flour" latitude="45.043556" longLatPrecision="1" longitude="3.103639" municipality="Vendeze" name="Cantal" pageId="58" pageNumber="59">Cantal</location>
|
||
,
|
||
<collectingMunicipality id="AD4DD621D73EFF9DFB3DDF341F46A60A" box="[1199,1298,1822,1848]" pageId="58" pageNumber="59">Vendèze</collectingMunicipality>
|
||
near St-
|
||
<collectingCounty id="A44834D7D73EFF9DFF2FDF681B54A66E" box="[189,256,1858,1884]" pageId="58" pageNumber="59">Flour</collectingCounty>
|
||
;
|
||
<geoCoordinate id="28A22A9CD73EFF9DFE9EDF6B1BECA66F" box="[268,440,1857,1885]" degrees="45" direction="north" minutes="02" orientation="latitude" pageId="58" pageNumber="59" precision="1" seconds="36.8" value="45.043556">45°02′36.8″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="28A22A9CD73EFF9DFE56DF6B180AA66F" box="[452,606,1857,1885]" degrees="3" direction="east" minutes="06" orientation="longitude" pageId="58" pageNumber="59" precision="1" seconds="13.1" value="3.103639">3°06′13.1″ E</geoCoordinate>
|
||
;
|
||
<collectionCode id="2B87D49ED73EFF9DFDF8DF68189FA66E" box="[618,715,1858,1884]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="58" pageNumber="59">MNHN</collectionCode>
|
||
.F.LIM181.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73EFF9DFF2FDFAF1BAAA6DB" blockId="58.[189,1399,1925,2025]" pageId="58" pageNumber="59">
|
||
According to the
|
||
<collectionCode id="2B87D49ED73EFF9DFE1DDFAC1BBDA692" box="[399,489,1926,1952]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="58" pageNumber="59">MNHN</collectionCode>
|
||
registry, it was discovered by M. Lauby (possibly Antoine Lauby), but sold to the
|
||
<collectionCode id="2B87D49ED73EFF9DFE9BDF801B37A6F6" box="[265,355,1962,1988]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="58" pageNumber="59">MNHN</collectionCode>
|
||
by M. Hugon, from St-Flour on the 19
|
||
<superScript id="BAE3E113D73EFF9DFCBDDF831968A685" attach="left" box="[815,828,1961,1975]" fontSize="6" pageId="58" pageNumber="59">th</superScript>
|
||
of
|
||
<date id="39286A9BD73EFF9DFCF7DF8019B4A6F6" box="[869,992,1962,1988]" pageId="58" pageNumber="59" value="1909-06">June 1909</date>
|
||
. It bears the old
|
||
<collectionCode id="2B87D49ED73EFF9DFB37DF801EABA6F6" box="[1189,1279,1962,1988]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="58" pageNumber="59">MNHN</collectionCode>
|
||
inventory number
|
||
<collectionCode id="2B87D49ED73EFF9DFE8CDFE41BD4A6DB" box="[286,384,1998,2025]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="58" pageNumber="59">MNHN</collectionCode>
|
||
.F.1909-25.
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73FFF9CFF78D9221BB0A010" blockId="59.[187,1400,264,718]" box="[234,484,264,290]" pageId="59" pageNumber="60">
|
||
<emphasis id="7FE29049D73FFF9CFF78D9221BB0A010" bold="true" box="[234,484,264,290]" pageId="59" pageNumber="60">Additional material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73FFF9CFF2FD9071854A3FC" blockId="59.[187,1400,264,718]" pageId="59" pageNumber="60">
|
||
<materialsCitation id="FDFE4606D73FFF9CFF2FD9071F33A075" ID-GBIF-Occurrence="3302594318" box="[189,1383,300,327]" country="Switzerland" location="Bumbach" pageId="59" pageNumber="60" specimenCode="NMB-UM-200" specimenCount="1">
|
||
<collectingCountry id="35810CCBD73FFF9CFF2FD9071BDAA075" box="[189,398,301,327]" name="Switzerland" pageId="59" pageNumber="60">SWITZERLAND</collectingCountry>
|
||
–
|
||
<emphasis id="7FE29049D73FFF9CFE34D90718D1A074" bold="true" box="[422,645,300,327]" pageId="59" pageNumber="60">
|
||
<location id="48491A80D73FFF9CFE34D907184BA075" LSID="urn:lsid:plazi:treatment:C53FFD4DD73EFFE4FDF2D9221BC0A4AE:48491A80D73FFF9CFE34D907184BA075" box="[422,543,301,327]" country="Switzerland" name="Bumbach" pageId="59" pageNumber="60">Bumbach</location>
|
||
(MP25)
|
||
</emphasis>
|
||
• 1 very poorly preserved and incomplete skull;
|
||
<specimenCode id="1D30E420D73FFF9CFB27D9071F33A075" box="[1205,1383,301,327]" pageId="59" pageNumber="60">NMB-UM-200</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFF29D97A19B1A059" ID-GBIF-Occurrence="3302594441" box="[187,997,336,363]" collectionCode="NMBE" pageId="59" pageNumber="60" specimenCode="NMBE-5035820" specimenCount="1">
|
||
1 fragment of parietal bone with occipital crest;
|
||
<specimenCode id="1D30E420D73FFF9CFCB3D97A19B1A059" box="[801,997,336,363]" pageId="59" pageNumber="60">NMBE-5035820</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFB99D97A1F59A058" ID-GBIF-Occurrence="3302594470" box="[1035,1293,336,362]" pageId="59" pageNumber="60" specimenCode="NMB-UM-463" specimenCount="1" specimenCount-pupa="1">
|
||
<specimenCount id="5B9087D2D73FFF9CFB99D97A1E69A058" box="[1035,1085,336,362]" pageId="59" pageNumber="60" type="pupa">1 P</specimenCount>
|
||
1;
|
||
<specimenCode id="1D30E420D73FFF9CFBC9D97A1F59A058" box="[1115,1293,336,362]" pageId="59" pageNumber="60">NMB-UM-463</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFAA0D97A1B2FA0BC" ID-GBIF-Occurrence="3302594437" pageId="59" pageNumber="60" specimenCode="NMB-UM-126a" specimenCount="1" specimenCount-pupa="1">
|
||
<specimenCount id="5B9087D2D73FFF9CFAA0D97A1F35A058" box="[1330,1377,336,362]" pageId="59" pageNumber="60" type="pupa">1 P</specimenCount>
|
||
2;
|
||
<specimenCode id="1D30E420D73FFF9CFF2FD95E1B2FA0BC" box="[189,379,372,398]" pageId="59" pageNumber="60">NMB-UM-126a</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFE37D95E18EBA0BD" ID-GBIF-Occurrence="3302594417" box="[421,703,372,399]" collectionCode="NMBE" pageId="59" pageNumber="60" specimenCode="NMBE-5014494" specimenCount="1" specimenCount-pupa="1">
|
||
<specimenCount id="5B9087D2D73FFF9CFE37D95E1B82A0BC" box="[421,470,372,398]" pageId="59" pageNumber="60" type="pupa">1 P</specimenCount>
|
||
2;
|
||
<specimenCode id="1D30E420D73FFF9CFE68D95E18EBA0BD" box="[506,703,372,399]" pageId="59" pageNumber="60">NMBE-5014494</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFD7BD95E1E57A0BD" ID-GBIF-Occurrence="3302594454" box="[745,1027,372,399]" collectionCode="NMBE" pageId="59" pageNumber="60" specimenCode="NMBE-5035822" specimenCount="1" specimenCount-pupa="1">
|
||
<specimenCount id="5B9087D2D73FFF9CFD7BD95E194FA0BC" box="[745,795,372,398]" pageId="59" pageNumber="60" type="pupa">1 P</specimenCount>
|
||
3;
|
||
<specimenCode id="1D30E420D73FFF9CFCACD95E1E57A0BD" box="[830,1027,372,399]" pageId="59" pageNumber="60">NMBE-5035822</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFBBFD95E1F53A0BC" ID-GBIF-Occurrence="3302594449" box="[1069,1287,372,398]" collectionCode="MGL" pageId="59" pageNumber="60" specimenCode="MGL-4264" specimenCount="1" specimenCount-pupa="1">
|
||
<specimenCount id="5B9087D2D73FFF9CFBBFD95E1E0BA0BC" box="[1069,1119,372,398]" pageId="59" pageNumber="60" type="pupa">1 P</specimenCount>
|
||
3;
|
||
<specimenCode id="1D30E420D73FFF9CFB10D95E1F53A0BC" box="[1154,1287,372,398]" pageId="59" pageNumber="60">MGL-4264</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFAA3D95E1B8DA081" ID-GBIF-Occurrence="3302594325" collectionCode="MGL" pageId="59" pageNumber="60" specimenCode="MGL-5265, MGL-5266" specimenCount="2" specimenCount-pupa="2">
|
||
<specimenCount id="5B9087D2D73FFF9CFAA3D95E1F35A0BC" box="[1329,1377,372,398]" pageId="59" pageNumber="60" type="pupa">2 P</specimenCount>
|
||
4;
|
||
<specimenCode id="1D30E420D73FFF9CFF2FD9B21B11A081" box="[189,325,408,435]" pageId="59" pageNumber="60">MGL-5265</specimenCode>
|
||
,
|
||
<specimenCode id="1D30E420D73FFF9CFEC1D9B21B8DA081" box="[339,473,408,435]" pageId="59" pageNumber="60">MGL-5266</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFE67D9B21958A081" ID-GBIF-Occurrence="3302594458" box="[501,780,408,435]" collectionCode="NMBE" pageId="59" pageNumber="60" specimenCode="NMBE-5014495" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="5B9087D2D73FFF9CFE67D9B2187EA080" box="[501,554,408,434]" pageId="59" pageNumber="60" type="male">1 M</specimenCount>
|
||
2;
|
||
<specimenCode id="1D30E420D73FFF9CFDD5D9B21958A081" box="[583,780,408,435]" pageId="59" pageNumber="60">NMBE-5014495</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFCBBD9B21F48A080" ID-GBIF-Occurrence="3302594390" box="[809,1308,408,434]" pageId="59" pageNumber="60" specimenCode="NMB-UM-6132" specimenCount="1">
|
||
1 subcomplete mandible;
|
||
<specimenCode id="1D30E420D73FFF9CFBCED9B21F48A080" box="[1116,1308,408,434]" pageId="59" pageNumber="60">NMB-UM-6132</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFAABD9B21B10A0E5" ID-GBIF-Occurrence="3302594353" collectionCode="MGL" pageId="59" pageNumber="60" specimenCode="MGL-5275" specimenCount="1">
|
||
1 d2;
|
||
<specimenCode id="1D30E420D73FFF9CFF2DD9961B10A0E5" box="[191,324,444,471]" pageId="59" pageNumber="60">MGL-5275</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFECCD9961810A0E4" ID-GBIF-Occurrence="3302594346" box="[350,580,444,470]" pageId="59" pageNumber="60" specimenCode="NMB-UM-13" specimenCount="1">
|
||
1 d4;
|
||
<specimenCode id="1D30E420D73FFF9CFE30D9961810A0E4" box="[418,580,444,470]" pageId="59" pageNumber="60">NMB-UM-13</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFDCCD9961932A0E5" ID-GBIF-Occurrence="3302594406" box="[606,870,444,471]" collectionCode="NMBE" pageId="59" pageNumber="60" specimenCode="NMBE-5035824" specimenCount="1" specimenCount-pupa="1">
|
||
<specimenCount id="5B9087D2D73FFF9CFDCCD99618D1A0E4" box="[606,645,444,470]" pageId="59" pageNumber="60" type="pupa">1 p</specimenCount>
|
||
2;
|
||
<specimenCode id="1D30E420D73FFF9CFD30D9961932A0E5" box="[674,870,444,471]" pageId="59" pageNumber="60">NMBE-5035824</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFC12D9961E1EA0E5" ID-GBIF-Occurrence="3302594412" box="[896,1098,444,471]" collectionCode="MGL" pageId="59" pageNumber="60" specimenCode="MGL-5274" specimenCount="1" specimenCount-pupa="1">
|
||
<specimenCount id="5B9087D2D73FFF9CFC12D99619FDA0E4" box="[896,937,444,470]" pageId="59" pageNumber="60" type="pupa">1 p</specimenCount>
|
||
3;
|
||
<specimenCode id="1D30E420D73FFF9CFC56D9961E1EA0E5" box="[964,1098,444,471]" pageId="59" pageNumber="60">MGL-5274</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFBF1D9961841A0C9" ID-GBIF-Occurrence="3302594339" collectionCode="NMBE" pageId="59" pageNumber="60" specimenCode="NMBE-5035825" specimenCount="1">
|
||
1 fragment of mandible with p3–4;
|
||
<specimenCode id="1D30E420D73FFF9CFEC2D9CA1841A0C9" box="[336,533,480,507]" pageId="59" pageNumber="60">NMBE-5035825</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFDA8D9CA1E7DA0C9" ID-GBIF-Occurrence="3302594350" box="[570,1065,480,507]" collectionCode="NMBE" pageId="59" pageNumber="60" specimenCode="NMBE-5035826, NMBE-5035827" specimenCount="2" specimenCount-pupa="2">
|
||
<specimenCount id="5B9087D2D73FFF9CFDA8D9CA1833A0C8" box="[570,615,480,506]" pageId="59" pageNumber="60" type="pupa">2 p</specimenCount>
|
||
4;
|
||
<specimenCode id="1D30E420D73FFF9CFD18D9CA1906A0C9" box="[650,850,480,507]" pageId="59" pageNumber="60">NMBE-5035826</specimenCode>
|
||
,
|
||
<specimenCode id="1D30E420D73FFF9CFCF6D9CA1E7DA0C9" box="[868,1065,480,507]" pageId="59" pageNumber="60">NMBE-5035827</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFBDDD9CA1F0BA0C8" ID-GBIF-Occurrence="3302594392" box="[1103,1375,480,506]" pageId="59" pageNumber="60" specimenCode="NMB-UM-6133" specimenCount="1" specimenCount-pupa="1">
|
||
<specimenCount id="5B9087D2D73FFF9CFBDDD9CA1E2FA0C8" box="[1103,1147,480,506]" pageId="59" pageNumber="60" type="pupa">1 p</specimenCount>
|
||
4;
|
||
<specimenCode id="1D30E420D73FFF9CFB0DD9CA1F0BA0C8" box="[1183,1375,480,506]" pageId="59" pageNumber="60">NMB-UM-6133</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFF2FDA2E1B9FA32C" ID-GBIF-Occurrence="3302594321" box="[189,459,516,542]" pageId="59" pageNumber="60" specimenCode="NMB-UM-806" specimenCount="1">
|
||
<quantity id="8A6EE1BED73FFF9CFF2FDA2E1AADA32C" box="[189,249,516,542]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="59" pageNumber="60" unit="m" value="1.0">
|
||
<specimenCount id="5B9087D2D73FFF9CFF2FDA2E1A98A32C" box="[189,204,516,542]" pageId="59" pageNumber="60">1</specimenCount>
|
||
m
|
||
</quantity>
|
||
1;
|
||
<specimenCode id="1D30E420D73FFF9CFE8BDA2E1B9FA32C" box="[281,459,516,542]" pageId="59" pageNumber="60">NMB-UM-806</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFE67DA2E1942A32D" ID-GBIF-Occurrence="3302594342" box="[501,790,516,543]" collectionCode="NMBE" pageId="59" pageNumber="60" specimenCode="NMBE-5035828" specimenCount="1">
|
||
<quantity id="8A6EE1BED73FFF9CFE67DA2E1864A32C" box="[501,560,516,542]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="59" pageNumber="60" unit="m" value="1.0">
|
||
<specimenCount id="5B9087D2D73FFF9CFE67DA2E1850A32C" box="[501,516,516,542]" pageId="59" pageNumber="60">1</specimenCount>
|
||
m
|
||
</quantity>
|
||
1;
|
||
<specimenCode id="1D30E420D73FFF9CFDC3DA2E1942A32D" box="[593,790,516,543]" pageId="59" pageNumber="60">NMBE-5035828</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFCD2DA2E1E70A32C" ID-GBIF-Occurrence="3302594462" box="[832,1060,516,542]" collectionCode="NMB" pageId="59" pageNumber="60" specimenCode="NMB-6278" specimenCount="1">
|
||
<quantity id="8A6EE1BED73FFF9CFCD2DA2E192DA32C" box="[832,889,516,542]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="59" pageNumber="60" unit="m" value="1.0">
|
||
<specimenCount id="5B9087D2D73FFF9CFCD2DA2E191BA32C" box="[832,847,516,542]" pageId="59" pageNumber="60">1</specimenCount>
|
||
m
|
||
</quantity>
|
||
2;
|
||
<specimenCode id="1D30E420D73FFF9CFC0EDA2E1E70A32C" box="[924,1060,516,542]" pageId="59" pageNumber="60">NMB-6278</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFBDCDA2E1803A371" ID-GBIF-Occurrence="3302594461" collectionCode="NMBE" pageId="59" pageNumber="60" specimenCode="NMBE-5035829, NMBE-5035830, NMBE-5035831" specimenCount="1">
|
||
<quantity id="8A6EE1BED73FFF9CFBDCDA2E1ED2A32C" box="[1102,1158,516,542]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="59" pageNumber="60" unit="m" value="3.0">3 m</quantity>
|
||
2;
|
||
<specimenCode id="1D30E420D73FFF9CFB38DA2E1F26A32D" box="[1194,1394,516,543]" pageId="59" pageNumber="60">NMBE-5035829</specimenCode>
|
||
,
|
||
<specimenCode id="1D30E420D73FFF9CFF2FDA021BD1A371" box="[189,389,552,579]" pageId="59" pageNumber="60">NMBE-5035830</specimenCode>
|
||
,
|
||
<specimenCode id="1D30E420D73FFF9CFE01DA021803A371" box="[403,599,552,579]" pageId="59" pageNumber="60">NMBE-5035831</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFDE6DA0219D3A371" ID-GBIF-Occurrence="3302594304" box="[628,903,552,579]" collectionCode="NMBE" pageId="59" pageNumber="60" specimenCode="NMBE-5035832" specimenCount="1">
|
||
<quantity id="8A6EE1BED73FFF9CFDE6DA0218F3A370" box="[628,679,552,578]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="59" pageNumber="60" unit="m" value="1.0">1 m</quantity>
|
||
3;
|
||
<specimenCode id="1D30E420D73FFF9CFD50DA0219D3A371" box="[706,903,552,579]" pageId="59" pageNumber="60">NMBE-5035832</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFC36DA021B2FA357" ID-GBIF-Occurrence="3302594323" pageId="59" pageNumber="60" specimenCode="NMB-UM-132, NMB-UM-129a" specimenCount="1">
|
||
2 incomplete humeri;
|
||
<specimenCode id="1D30E420D73FFF9CFB2FDA021F26A370" box="[1213,1394,552,578]" pageId="59" pageNumber="60">NMB-UM-132</specimenCode>
|
||
,
|
||
<specimenCode id="1D30E420D73FFF9CFF2FDA611B2FA357" box="[189,379,587,613]" pageId="59" pageNumber="60">NMB-UM-129a</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFE32DA611898A357" ID-GBIF-Occurrence="3302594347" box="[416,716,586,613]" collectionCode="NMB-UM" pageId="59" pageNumber="60" specimenCode="NMB-UM127a" specimenCount="1">
|
||
1 radius;
|
||
<specimenCode id="1D30E420D73FFF9CFD8ADA611898A357" box="[536,716,587,613]" collectionCode="NMB-UM" pageId="59" pageNumber="60">NMB-UM127a</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFD63DA611EB3A357" ID-GBIF-Occurrence="3302594435" box="[753,1255,586,613]" pageId="59" pageNumber="60" specimenCode="NMB-UM-131b, NMB-UM-131c" specimenCount="1">
|
||
1 ulna;
|
||
<specimenCode id="1D30E420D73FFF9CFCC6DA601E43A356" box="[852,1047,586,612]" pageId="59" pageNumber="60">NMB-UM-131b</specimenCode>
|
||
,
|
||
<specimenCode id="1D30E420D73FFF9CFBBADA611EB3A357" box="[1064,1255,587,613]" pageId="59" pageNumber="60">NMB-UM-131c</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFA99DA611BD6A3BA" ID-GBIF-Occurrence="3302594306" collectionCode="NMBE" pageId="59" pageNumber="60" specimenCode="NMBE-5035833" specimenCount="1">
|
||
1 lunate;
|
||
<specimenCode id="1D30E420D73FFF9CFF2FDA471BD6A3BA" box="[189,386,621,648]" pageId="59" pageNumber="60">NMBE-5035833</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFE3BDA47199AA3B5" ID-GBIF-Occurrence="3302594366" box="[425,974,621,647]" pageId="59" pageNumber="60" specimenCode="NMB-UM-6136b, NMB-UM-6" specimenCount="1">
|
||
2 trapezoids;
|
||
<specimenCode id="1D30E420D73FFF9CFDC4DA471973A3B5" box="[598,807,621,647]" pageId="59" pageNumber="60">NMB-UM-6136b</specimenCode>
|
||
,
|
||
<specimenCode id="1D30E420D73FFF9CFCA8DA47199AA3B5" box="[826,974,621,647]" pageId="59" pageNumber="60">NMB-UM-6</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFC64DA471F0AA3B5" ID-GBIF-Occurrence="3302594384" box="[1014,1374,621,648]" pageId="59" pageNumber="60" specimenCode="NMB-UM-6136c" specimenCount="1">
|
||
1 magnum;
|
||
<specimenCode id="1D30E420D73FFF9CFB02DA471F0AA3B5" box="[1168,1374,621,647]" pageId="59" pageNumber="60">NMB-UM-6136c</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFF2FDABA1BBFA398" ID-GBIF-Occurrence="3302594376" box="[189,491,656,683]" pageId="59" pageNumber="60" specimenCode="NMB-UM-6136a" specimenCount="1">
|
||
1 McII;
|
||
<specimenCode id="1D30E420D73FFF9CFE8CDABA1BBFA398" box="[286,491,656,682]" pageId="59" pageNumber="60">NMB-UM-6136a</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFD96DABA1943A398" ID-GBIF-Occurrence="3302594443" box="[516,791,656,683]" pageId="59" pageNumber="60" specimenCode="NMB-UM-121" specimenCount="1">
|
||
1 McII;
|
||
<specimenCode id="1D30E420D73FFF9CFDF7DABA1943A398" box="[613,791,656,682]" pageId="59" pageNumber="60">NMB-UM-121</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFCA2DABA1F31A399" ID-GBIF-Occurrence="3302594326" box="[816,1381,656,683]" collectionCode="NMBE" pageId="59" pageNumber="60" specimenCode="NMBE-5035835" specimenCount="1">
|
||
1 proximal fragment of femur;
|
||
<specimenCode id="1D30E420D73FFF9CFB0DDABA1F31A399" box="[1183,1381,656,683]" pageId="59" pageNumber="60">NMBE-5035835</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="FDFE4606D73FFF9CFF2FDA991BAFA3FC" ID-GBIF-Occurrence="3302594471" box="[189,507,691,718]" collectionCode="NMBE" pageId="59" pageNumber="60" specimenCode="NMBE-5035834" specimenCount="1">
|
||
1 cuboid;
|
||
<specimenCode id="1D30E420D73FFF9CFEA1DA991BAFA3FC" box="[307,507,691,718]" pageId="59" pageNumber="60">NMBE-5035834</specimenCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73FFF9CFF2FDADE1BAFA23F" blockId="59.[189,507,755,782]" box="[189,507,755,782]" pageId="59" pageNumber="60">
|
||
<heading id="1661FB37D73FFF9CFF2FDADE1BAFA23F" bold="true" box="[189,507,755,782]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
|
||
<emphasis id="7FE29049D73FFF9CFF2FDADE1BAFA23F" bold="true" box="[189,507,755,782]" pageId="59" pageNumber="60">Type horizon and locality</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73FFF9CFF2FDB081955A26D" blockId="59.[189,1398,802,863]" pageId="59" pageNumber="60">
|
||
Vendèze near St-Flour, Cantal, Auvergne-Rhône-Alpes,
|
||
<collectingCountry id="35810CCBD73FFF9CFCC6DB0819F0A20E" box="[852,932,802,828]" name="France" pageId="59" pageNumber="60">France</collectingCountry>
|
||
(
|
||
<collectionCode id="2B87D49ED73FFF9CFC26DB0819B6A20E" box="[948,994,802,828]" country="0" name="Mohonk Preserve, Inc." pageId="59" pageNumber="60">MP</collectionCode>
|
||
24, late early Oligocene), approximative coordinates:
|
||
<geoCoordinate id="28A22A9CD73FFF9CFE39DB691803A26D" box="[427,599,835,863]" degrees="45" direction="north" minutes="02" orientation="latitude" pageId="59" pageNumber="60" precision="1" seconds="36.8" value="45.043556">45°02′36.8″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="28A22A9CD73FFF9CFDF0DB6918A8A26D" box="[610,764,835,863]" degrees="3" direction="east" minutes="06" orientation="longitude" pageId="59" pageNumber="60" precision="1" seconds="13.1" value="3.103639">3°06′13.1″ E</geoCoordinate>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73FFF9CFF2FDBAF1843A2AD" blockId="59.[189,535,901,927]" box="[189,535,901,927]" pageId="59" pageNumber="60">
|
||
<heading id="1661FB37D73FFF9CFF2FDBAF1843A2AD" bold="true" box="[189,535,901,927]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
|
||
<emphasis id="7FE29049D73FFF9CFF2FDBAF1843A2AD" bold="true" box="[189,535,901,927]" pageId="59" pageNumber="60">Stratigraphical distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73FFF9CFF2FDB991B37A2FC" blockId="59.[189,355,947,974]" box="[189,355,947,974]" pageId="59" pageNumber="60">MP24–MP25.</paragraph>
|
||
<paragraph id="4D294C5BD73FFF9CFF2FDBDE1856A53C" blockId="59.[189,514,1012,1038]" box="[189,514,1012,1038]" pageId="59" pageNumber="60">
|
||
<heading id="1661FB37D73FFF9CFF2FDBDE1856A53C" bold="true" box="[189,514,1012,1038]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
|
||
<emphasis id="7FE29049D73FFF9CFF2FDBDE1856A53C" bold="true" box="[189,514,1012,1038]" pageId="59" pageNumber="60">Geographical distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73FFF9CFF2FDC091884A56D" blockId="59.[189,1398,1058,1119]" pageId="59" pageNumber="60">
|
||
<collectingCountry id="35810CCBD73FFF9CFF2FDC091B45A50F" box="[189,273,1059,1085]" name="France" pageId="59" pageNumber="60">France</collectingCountry>
|
||
: ‘Auvergne’ (without precision, which could possibly correspond to Vendèze), Lamothe- Capdeville, Vendèze.
|
||
<collectingCountry id="35810CCBD73FFF9CFE2EDC6F1819A56D" box="[444,589,1093,1119]" name="Switzerland" pageId="59" pageNumber="60">Switzerland</collectingCountry>
|
||
: Bumbach.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="058C1FD0D73FFFE4FF2FDCAC1BC0A4AE" lastPageId="67" lastPageNumber="68" pageId="59" pageNumber="60" type="description">
|
||
<paragraph id="4D294C5BD73FFF9CFF2FDCAC1B1AA592" blockId="59.[189,334,1158,1184]" box="[189,334,1158,1184]" pageId="59" pageNumber="60">
|
||
<heading id="1661FB37D73FFF9CFF2FDCAC1B1AA592" bold="true" box="[189,334,1158,1184]" fontSize="11" level="3" pageId="59" pageNumber="60" reason="3">
|
||
<emphasis id="7FE29049D73FFF9CFF2FDCAC1B1AA592" bold="true" box="[189,334,1158,1184]" pageId="59" pageNumber="60">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73FFF9CFF78DC9E1B08A5FC" blockId="59.[188,1400,1204,1890]" box="[234,348,1204,1230]" pageId="59" pageNumber="60">
|
||
<emphasis id="7FE29049D73FFF9CFF78DC9E1B08A5FC" bold="true" box="[234,348,1204,1230]" pageId="59" pageNumber="60">
|
||
<typeStatus id="922DF2F9D73FFF9CFF78DC9E1B08A5FC" box="[234,348,1204,1230]" pageId="59" pageNumber="60" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73FFF9CFF2FDCFD1F23A650" blockId="59.[188,1400,1204,1890]" pageId="59" pageNumber="60">
|
||
SKULL. The skull MNHN.F.LIM181 is quite well preserved but the nasals are broken (
|
||
<figureCitation id="D5AD50DED73FFF9CFB3EDCFD1F62A5C3" box="[1196,1334,1239,1265]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="60.[189,232,1963,1989]" captionStartId-1="61.[189,232,1822,1848]" captionTargetBox-0="[194,1394,265,1940]" captionTargetBox-1="[252,1331,269,1782]" captionTargetId-0="figure-44@60.[194,1394,265,1940]" captionTargetId-1="figure-100@61.[239,1349,251,1797]" captionTargetPageId-0="60" captionTargetPageId-1="61" captionText-0="Fig. 22. Ronzotherium heissigi sp. nov. from Vendèze (late early Oligocene, France). Skull MNHN.F.LIM181. A. Dorsal view. B. Left lateral view. C. Ventral view. Scale bar: 2 cm." captionText-1="Fig. 23. Ronzotherium heissigi sp. nov. from Vendèze (late early Oligocene, France). Drawing of the skull MNHN.F.LIM181. A. Dorsal view. B. Left lateral view. C. Ventral view. Abbreviations: AT = articular tubercle; BO = basioccipital; F = frontal; IOF = infraorbital foramen; J = jugal; M = maxilla; NT = nuchal tubercle; OC = occipital condyle; P = parietal; PGA = postglenoid apophysis; PMX = premaxilla; PoP = postorbital process; PP = paraoccipital process; SC = sagittal crest; SQ = squamosal. Scale bar: 2 cm." figureDoi-0="http://doi.org/10.5281/zenodo.4945922" figureDoi-1="http://doi.org/10.5281/zenodo.4945926" httpUri-0="https://zenodo.org/record/4945922/files/figure.png" httpUri-1="https://zenodo.org/record/4945926/files/figure.png" pageId="59" pageNumber="60">Figs 22–23</figureCitation>
|
||
). The premaxillae are long and contact each other only at their anterior extremity. The nasal notch extends up to P2 and the foramen infraorbitalis is located above P3. The nasal septum is not ossified. The suture between the nasals and the lacrimal is not visible and the lacrimal process is absent. The orbit is large and its anterior border is above the anterior side of M1. The processus postrobitalis of the frontal is large. The anterior base of the zygomatic process is high above the teeth neck. The zygomatic arch is high in lateral view, it is almost reaching the dorsal border of the skull. The postorbital process of the zygomatic arch is almost absent and very poorly distinguishable. The dorsal profile of the skull is overall concave in lateral view. The external auditory pseudomeatus is partially closed and the occipital side is inclined forward. The nuchal tubercle is developed. The back of the teeth row is in the posterior half of the skull in lateral view. In dorsal view, the skull is brachycephalic and it is hornless. The orbit is not laterally projected. The zygomatic width/frontal width ratio is above 1.5. A very thin sagittal crest is present, and the occipital crest is not preserved. In ventral view, the anterior tip of the zygomatic arch diverges progressively from the maxilla, without a sharp angle. The vomer and most of the basicranium are not preserved. The articular tubercle of the squamosal is rather smooth and its transverse profile is straight. The anterolateral sides of the processus postglenoidalis form a right dihedron. There is no posterior groove on the zygomatic process. The processus posttympanicus and paraoccipitalis are fused at their base. The processus post-tympanicus is poorly developed while the paraoccipitalis is developed. The foramen magnum is circular and there are no median ridges on the condyles, neither medial truncation.
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73FFF9CFF2FDFA31F22A6DA" blockId="59.[189,1400,1929,2025]" pageId="59" pageNumber="60">
|
||
MANDIBLE. The mandibular symphysis (
|
||
<figureCitation id="D5AD50DED73FFF9CFD06DFA31977A691" box="[660,803,1929,1955]" captionStart="Fig" captionStartId="62.[189,232,1893,1919]" captionTargetBox="[227,1360,265,1868]" captionTargetId="figure-13@62.[227,1360,265,1868]" captionTargetPageId="62" captionText="Fig. 24. Ronzotherium heissigi sp. nov. from Vendèze (late early Oligocene, France). A–D. Mandible MNHN.F.LIM181. A. Occlusal view. B. Lateral view. C. With left p2–p4 and m2–3 in labial view. View. D. Right p2–m3 in lingual view. – E. Skull MNHN.F.LIM181, belonging to the same individual as the mandible, close-up view of P2–M3 in occlusal view. Scale bars: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.4945928" httpUri="https://zenodo.org/record/4945928/files/figure.png" pageId="59" pageNumber="60">Fig. 24A–B</figureCitation>
|
||
) is slightly upraised and it is long and massive in dorsal view. Its posterior margin is at the level of p2. There are two mental foramen, one bellow p2 and one below the root of i2. The lingual groove of the sulcus mylohyoideus is slightly marked on the lingual
|
||
</paragraph>
|
||
<caption id="19E91CD3D738FF9BFF2FDF811E82A6DB" ID-DOI="http://doi.org/10.5281/zenodo.4945922" ID-Zenodo-Dep="4945922" httpUri="https://zenodo.org/record/4945922/files/figure.png" pageId="60" pageNumber="61" startId="60.[189,232,1963,1989]" targetBox="[194,1394,265,1940]" targetPageId="60">
|
||
<paragraph id="4D294C5BD738FF9BFF2FDF811E82A6DB" blockId="60.[189,1398,1963,2025]" pageId="60" pageNumber="61">
|
||
<emphasis id="7FE29049D738FF9BFF2FDF811B72A6F7" bold="true" box="[189,294,1963,1989]" pageId="60" pageNumber="61">Fig. 22.</emphasis>
|
||
<taxonomicName id="8A9637D8D738FF9BFEABDF81181AA6F7" authority="Tissier & Antoine & Becker, 2021" authorityName="Tissier & Antoine & Becker" authorityYear="2021" box="[313,590,1963,1989]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="heissigi" status="sp. nov.">
|
||
<emphasis id="7FE29049D738FF9BFEABDF81181AA6F7" box="[313,590,1963,1989]" italics="true" pageId="60" pageNumber="61">Ronzotherium heissigi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="64D12D32D738FF9BFDF3DF861893A6F4" box="[609,711,1964,1990]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from Vendèze (late early Oligocene, France). Skull MNHN.F.LIM181.
|
||
<emphasis id="7FE29049D738FF9BFE36DFE41BEDA6DA" bold="true" box="[420,441,1998,2024]" pageId="60" pageNumber="61">A</emphasis>
|
||
. Dorsal view.
|
||
<emphasis id="7FE29049D738FF9BFDF1DFE51823A6DB" bold="true" box="[611,631,1999,2025]" pageId="60" pageNumber="61">B</emphasis>
|
||
. Left lateral view.
|
||
<emphasis id="7FE29049D738FF9BFCC6DFE4193DA6DA" bold="true" box="[852,873,1998,2024]" pageId="60" pageNumber="61">C</emphasis>
|
||
. Ventral view. Scale bar: 2 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="19E91CD3D739FF9AFF2FDF341B2EA6DB" ID-DOI="http://doi.org/10.5281/zenodo.4945926" ID-Zenodo-Dep="4945926" httpUri="https://zenodo.org/record/4945926/files/figure.png" pageId="61" pageNumber="62" startId="61.[189,232,1822,1848]" targetBox="[252,1331,269,1782]" targetPageId="61">
|
||
<paragraph id="4D294C5BD739FF9AFF2FDF341B2EA6DB" blockId="61.[189,1400,1822,2025]" pageId="61" pageNumber="62">
|
||
<emphasis id="7FE29049D739FF9AFF2FDF341B48A60A" bold="true" box="[189,284,1822,1848]" pageId="61" pageNumber="62">Fig. 23.</emphasis>
|
||
<taxonomicName id="8A9637D8D739FF9AFEB7DF341864A60A" authority="Tissier & Antoine & Becker, 2021" authorityName="Tissier & Antoine & Becker" authorityYear="2021" box="[293,560,1822,1848]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="heissigi" status="sp. nov.">
|
||
<emphasis id="7FE29049D739FF9AFEB7DF341864A60A" box="[293,560,1822,1848]" italics="true" pageId="61" pageNumber="62">Ronzotherium heissigi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="64D12D32D739FF9AFDABDF3518C2A60B" box="[569,662,1823,1849]" pageId="61" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from Vendèze (late early Oligocene, France). Drawing of the skull MNHN.F.LIM181.
|
||
<emphasis id="7FE29049D739FF9AFE79DF6B1854A669" bold="true" box="[491,512,1857,1883]" pageId="61" pageNumber="62">A</emphasis>
|
||
. Dorsal view.
|
||
<emphasis id="7FE29049D739FF9AFD26DF6B189CA669" bold="true" box="[692,712,1857,1883]" pageId="61" pageNumber="62">B</emphasis>
|
||
. Left lateral view.
|
||
<emphasis id="7FE29049D739FF9AFC21DF6B199CA669" bold="true" box="[947,968,1857,1883]" pageId="61" pageNumber="62">C</emphasis>
|
||
. Ventral view. Abbreviations: AT = articular tubercle; BO = basioccipital; F = frontal; IOF = infraorbital foramen; J = jugal; M = maxilla; NT = nuchal tubercle; OC = occipital condyle; P = parietal; PGA = postglenoid apophysis; PMX = premaxilla; PoP = postorbital process; PP = paraoccipital process; SC = sagittal crest; SQ = squamosal. Scale bar: 2 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="19E91CD3D73AFF99FF2FDF4F19A1A6DB" ID-DOI="http://doi.org/10.5281/zenodo.4945928" ID-Zenodo-Dep="4945928" httpUri="https://zenodo.org/record/4945928/files/figure.png" pageId="62" pageNumber="63" startId="62.[189,232,1893,1919]" targetBox="[227,1360,265,1868]" targetPageId="62">
|
||
<paragraph id="4D294C5BD73AFF99FF2FDF4F19A1A6DB" blockId="62.[188,1399,1893,2025]" pageId="62" pageNumber="63">
|
||
<emphasis id="7FE29049D73AFF99FF2FDF4F1B48A64D" bold="true" box="[189,284,1893,1919]" pageId="62" pageNumber="63">Fig. 24.</emphasis>
|
||
<taxonomicName id="8A9637D8D73AFF99FEB7DF4F187AA64D" authority="Tissier & Antoine & Becker, 2021" authorityName="Tissier & Antoine & Becker" authorityYear="2021" box="[293,558,1893,1919]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="heissigi" status="sp. nov.">
|
||
<emphasis id="7FE29049D73AFF99FEB7DF4F1B9FA64D" box="[293,459,1893,1919]" italics="true" pageId="62" pageNumber="63">Ronzotherium</emphasis>
|
||
heissigi
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="64D12D32D73AFF99FDA5DF4C18C7A6B2" box="[567,659,1894,1920]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from Vendèze (late early Oligocene, France).
|
||
<emphasis id="7FE29049D73AFF99FB2CDF4F1EA1A64D" bold="true" box="[1214,1269,1893,1919]" pageId="62" pageNumber="63">A–D</emphasis>
|
||
. Mandible MNHN.F.LIM181.
|
||
<emphasis id="7FE29049D73AFF99FE30DFA21BE3A690" bold="true" box="[418,439,1928,1954]" pageId="62" pageNumber="63">A</emphasis>
|
||
. Occlusal view.
|
||
<emphasis id="7FE29049D73AFF99FDE7DFA318DDA691" bold="true" box="[629,649,1929,1955]" pageId="62" pageNumber="63">B</emphasis>
|
||
. Lateral view.
|
||
<emphasis id="7FE29049D73AFF99FCA1DFA2191CA690" bold="true" box="[819,840,1928,1954]" pageId="62" pageNumber="63">C</emphasis>
|
||
. With left p2–p4 and m2–3 in labial view. View.
|
||
<emphasis id="7FE29049D73AFF99FF2EDF861A85A6F4" bold="true" box="[188,209,1964,1990]" pageId="62" pageNumber="63">D</emphasis>
|
||
. Right p2–m3 in lingual view. –
|
||
<emphasis id="7FE29049D73AFF99FDCBDF861838A6F4" bold="true" box="[601,620,1964,1990]" pageId="62" pageNumber="63">E</emphasis>
|
||
. Skull MNHN.F.LIM181, belonging to the same individual as the mandible, close-up view of P2–M3 in occlusal view. Scale bars: 2 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4D294C5BD73BFF98FF2FD9231847A055" blockId="63.[188,1399,265,359]" pageId="63" pageNumber="64">border of the corpus. The ventral base of the corpus is completely straight. The ramus is inclined forward in lateral view, and the coronoid process is well developed. In medial view, the foramen mandibulare is located below the teeth neck.</paragraph>
|
||
<paragraph id="4D294C5BD73BFF98FF2FD9A41F58A0FB" blockId="63.[189,1398,397,458]" pageId="63" pageNumber="64">
|
||
DENTITION. The complete dental formula is I1–2, P2–M3 / i2, p2–m3. The Length(P3–4)/Length(M1–3) ratio is between 0.42 and 0.5. The cement is absent and the crowns are very low (
|
||
<figureCitation id="D5AD50DED73BFF98FBE4D9851F54A0FB" box="[1142,1280,431,457]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="60.[189,232,1963,1989]" captionStartId-1="61.[189,232,1822,1848]" captionStartId-2="62.[189,232,1893,1919]" captionTargetBox-0="[194,1394,265,1940]" captionTargetBox-1="[252,1331,269,1782]" captionTargetBox-2="[227,1360,265,1868]" captionTargetId-0="figure-44@60.[194,1394,265,1940]" captionTargetId-1="figure-100@61.[239,1349,251,1797]" captionTargetId-2="figure-13@62.[227,1360,265,1868]" captionTargetPageId-0="60" captionTargetPageId-1="61" captionTargetPageId-2="62" captionText-0="Fig. 22. Ronzotherium heissigi sp. nov. from Vendèze (late early Oligocene, France). Skull MNHN.F.LIM181. A. Dorsal view. B. Left lateral view. C. Ventral view. Scale bar: 2 cm." captionText-1="Fig. 23. Ronzotherium heissigi sp. nov. from Vendèze (late early Oligocene, France). Drawing of the skull MNHN.F.LIM181. A. Dorsal view. B. Left lateral view. C. Ventral view. Abbreviations: AT = articular tubercle; BO = basioccipital; F = frontal; IOF = infraorbital foramen; J = jugal; M = maxilla; NT = nuchal tubercle; OC = occipital condyle; P = parietal; PGA = postglenoid apophysis; PMX = premaxilla; PoP = postorbital process; PP = paraoccipital process; SC = sagittal crest; SQ = squamosal. Scale bar: 2 cm." captionText-2="Fig. 24. Ronzotherium heissigi sp. nov. from Vendèze (late early Oligocene, France). A–D. Mandible MNHN.F.LIM181. A. Occlusal view. B. Lateral view. C. With left p2–p4 and m2–3 in labial view. View. D. Right p2–m3 in lingual view. – E. Skull MNHN.F.LIM181, belonging to the same individual as the mandible, close-up view of P2–M3 in occlusal view. Scale bars: 2 cm." figureDoi-0="http://doi.org/10.5281/zenodo.4945922" figureDoi-1="http://doi.org/10.5281/zenodo.4945926" figureDoi-2="http://doi.org/10.5281/zenodo.4945928" httpUri-0="https://zenodo.org/record/4945922/files/figure.png" httpUri-1="https://zenodo.org/record/4945926/files/figure.png" httpUri-2="https://zenodo.org/record/4945928/files/figure.png" pageId="63" pageNumber="64">Figs 22–24</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD73BFF98FF2FD9DA1E7AA3A3" blockId="63.[189,1400,495,658]" pageId="63" pageNumber="64">I1 is oval in cross-section, pointed and not chisel-shaped. It is directed downwards, and it bears two crests, one anterior and one posterior. There was no contact with the lower incisor (no visible wear on any of them). It is separated from I2 by a very short diastema. I2 overall has the same shape as I1 but is smaller and less pointed. The diastema between I2 and P2 is very long. The i1 is absent, and the space between the two i2 is very short. The i2 are large, tusk-shaped and parallel.</paragraph>
|
||
<paragraph id="4D294C5BD73BFF98FF2FDA9D1E1DA52C" blockId="63.[188,1400,695,1055]" pageId="63" pageNumber="64">The labial cingulum of the upper premolars is always present, posteriorly, below the metastyle. The lingual one is always present, continuous and undulating in lingual view: it is high below the protocone, very low at the level of the median valley, and very high at the level of the hypocone. The crochet is always absent. The hypocone is connected to the ectoloph. The postfossette is narrow and the antecrochet is always absent. The P1 is absent. The protocone and hypocone of P2 are separated and the hypocone is posterior to the metacone. The metacone fold is strong. The protocone is same sized as the hypocone. The protoloph of P2 is joined to the ectoloph. The medifossette is always absent on the premolars and the protocone is never constricted. On P3–4, the protocone and hypocone are connected by a lingual wall. The metacone fold is strong. The hypocone is posterior to the metacone as on P2. The protoloph of P3 is joined to the ectoloph and the crista and pseudometaloph are absent. The antecrochet is always absent on P3–4. The metaloph of P4 is continuous.</paragraph>
|
||
<paragraph id="4D294C5BD73BFF98FF2FDC6E1977A45B" blockId="63.[187,1401,1092,1385]" pageId="63" pageNumber="64">The labial cingulum of the upper molars is always present, as on the premolars, below the metastyle. The antecrochet is present on M2, absent on M3 and not visible on M1. The crochet, crista, cristella and medifossette are always absent. The lingual cingulum is always absent. The protocone is never constricted. The paracone fold is strong and the metacone fold absent. The metastyle is long, but the parastyle is rather short. The metaloph is long. The posterior part of the ectoloph is straight. The posterior cingulum is continuous. There is no lingual groove on the protocone of M2 and the mesostyle is absent. The antecrochet and hypocone are well separated. The ectoloph and metaloph of M3 are fused into an ectometaloph, and it is quadrangular. The protocone is not constricted and the protoloph is transverse. The posterior groove of the ectometaloph is present.</paragraph>
|
||
<paragraph id="4D294C5BD73BFF98FF2FDDA41B8AA781" blockId="63.[188,1401,1422,1715]" pageId="63" pageNumber="64">The lower p2–3 do not bear vertical external rugosities. The external groove of the lower cheek teeth is angular and is developed until the neck. The trigonid is angular and forms an acute dihedron. The metaconid and entoconid are always joined to the hypolophid. The lingual opening of the lower premolars is V-shaped. The lingual cingulum is always absent on all lower cheek teeth and the labial one is basically absent, except anteriorly below the paralophid. The d/p1 is always absent. The paralophid of p2 is curved, without constriction and the paraconid is quite reduced. The posterior valley is lingually open. The lingual branch of the paralophid of p3 is long and developed. The anterolingual cingulum does not reach the metaconid. The hypolophid of the lower molar is transverse and there is no lingual groove of the entoconid.</paragraph>
|
||
<caption id="19E91CD3D73BFF98FF2FDED1186CA6DB" ID-DOI="http://doi.org/10.5281/zenodo.4959442" ID-Zenodo-Dep="4959442" httpUri="https://zenodo.org/record/4959442/files/figure.png" pageId="63" pageNumber="64" startId="63.[189,232,1787,1813]" subCaptionStartIDs="63.[940,1100,1963,1989]" subCaptionStarts="Abbr" targetBox="[189,1398,265,1957]" targetPageId="64">
|
||
<paragraph id="4D294C5BD73BFF98FF2FDED1186CA6DB" blockId="63.[187,1399,1787,2025]" pageId="63" pageNumber="64">
|
||
<emphasis id="7FE29049D73BFF98FF2FDED11B46A627" bold="true" box="[189,274,1787,1813]" pageId="63" pageNumber="64">Fig. 25</emphasis>
|
||
(next page).
|
||
<taxonomicName id="8A9637D8D73BFF98FE3CDED118E2A627" authority="Tissier & Antoine & Becker, 2021" authorityName="Tissier & Antoine & Becker" authorityYear="2021" box="[430,694,1787,1813]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="63" pageNumber="64" phylum="Chordata" rank="species" species="heissigi" status="sp. nov.">
|
||
<emphasis id="7FE29049D73BFF98FE3CDED118E2A627" box="[430,694,1787,1813]" italics="true" pageId="63" pageNumber="64">Ronzotherium heissigi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="64D12D32D73BFF98FD2FDED6194CA624" box="[701,792,1788,1814]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from Bumbach,
|
||
<collectingRegion id="8F5282B9D73BFF98FC4FDED61E42A624" box="[989,1046,1788,1814]" country="Switzerland" name="Bern" pageId="63" pageNumber="64">Bern</collectingRegion>
|
||
Canton (early late Oligocene;
|
||
<collectingCountry id="35810CCBD73BFF98FF2FDF341B1BA60B" box="[189,335,1822,1849]" name="Switzerland" pageId="63" pageNumber="64">Switzerland</collectingCountry>
|
||
). –
|
||
<emphasis id="7FE29049D73BFF98FEE8DF341BE7A60A" bold="true" box="[378,435,1822,1848]" pageId="63" pageNumber="64">A–D</emphasis>
|
||
. Mandible NMB-UM-6132.
|
||
<emphasis id="7FE29049D73BFF98FC99DF341974A60A" bold="true" box="[779,800,1822,1848]" pageId="63" pageNumber="64">A</emphasis>
|
||
. Occlusal view.
|
||
<emphasis id="7FE29049D73BFF98FC74DF3519ADA60B" bold="true" box="[998,1017,1823,1849]" pageId="63" pageNumber="64">B</emphasis>
|
||
. Left lateral view.
|
||
<emphasis id="7FE29049D73BFF98FB48DF341EBBA60A" bold="true" box="[1242,1263,1822,1848]" pageId="63" pageNumber="64">C</emphasis>
|
||
. With right p3–m
|
||
<quantity id="8A6EE1BED73BFF98FE90DF681B64A66E" box="[258,304,1858,1884]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="63" pageNumber="64" unit="in" value="3.0">3 in</quantity>
|
||
labial view.
|
||
<emphasis id="7FE29049D73BFF98FE5FDF681BB6A66E" bold="true" box="[461,482,1858,1884]" pageId="63" pageNumber="64">D</emphasis>
|
||
. With right p3–m
|
||
<quantity id="8A6EE1BED73BFF98FD2EDF6818BEA66E" box="[700,746,1858,1884]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="63" pageNumber="64" unit="in" value="3.0">3 in</quantity>
|
||
lingual view. –
|
||
<emphasis id="7FE29049D73BFF98FC22DF6819B1A66E" bold="true" box="[944,997,1858,1884]" pageId="63" pageNumber="64">E–F</emphasis>
|
||
. Right humerus NMB-UM-129a.
|
||
<emphasis id="7FE29049D73BFF98FF2FDF4F1A85A64D" bold="true" box="[189,209,1893,1919]" pageId="63" pageNumber="64">E</emphasis>
|
||
. Anterior view.
|
||
<emphasis id="7FE29049D73BFF98FE14DF4F1BCCA64D" bold="true" box="[390,408,1893,1919]" pageId="63" pageNumber="64">F</emphasis>
|
||
. Posterior view. –
|
||
<emphasis id="7FE29049D73BFF98FDFCDF4F18FFA64D" bold="true" box="[622,683,1893,1919]" pageId="63" pageNumber="64">G–H</emphasis>
|
||
. Left humerus NMB-UM-132.
|
||
<emphasis id="7FE29049D73BFF98FB8ADF4F1E7BA64D" bold="true" box="[1048,1071,1893,1919]" pageId="63" pageNumber="64">G</emphasis>
|
||
. Anterior view.
|
||
<emphasis id="7FE29049D73BFF98FB77DF4F1EA8A64D" bold="true" box="[1253,1276,1893,1919]" pageId="63" pageNumber="64">H</emphasis>
|
||
. Posterior view. –
|
||
<emphasis id="7FE29049D73BFF98FE87DFA21B6BA690" bold="true" box="[277,319,1928,1954]" pageId="63" pageNumber="64">I–J</emphasis>
|
||
. Right radius NMB-UM-127a.
|
||
<emphasis id="7FE29049D73BFF98FD3CDFA218EDA690" bold="true" box="[686,697,1928,1954]" pageId="63" pageNumber="64">I</emphasis>
|
||
. Anterior view.
|
||
<emphasis id="7FE29049D73BFF98FCE0DFA219D5A690" bold="true" box="[882,897,1928,1954]" pageId="63" pageNumber="64">J</emphasis>
|
||
. Posterior view. –
|
||
<emphasis id="7FE29049D73BFF98FBCBDFA21ECFA690" bold="true" box="[1113,1179,1928,1954]" pageId="63" pageNumber="64">K–M</emphasis>
|
||
. Right ulna NMB- UM-131b.
|
||
<emphasis id="7FE29049D73BFF98FED2DF811B03A6F7" bold="true" box="[320,343,1963,1989]" pageId="63" pageNumber="64">K</emphasis>
|
||
. Anterior view.
|
||
<emphasis id="7FE29049D73BFF98FD81DF811873A6F7" bold="true" box="[531,551,1963,1989]" pageId="63" pageNumber="64">L</emphasis>
|
||
. Posterior view.
|
||
<emphasis id="7FE29049D73BFF98FD7DDF81195FA6F7" bold="true" box="[751,779,1963,1989]" pageId="63" pageNumber="64">M</emphasis>
|
||
. Distal view. Abbreviation: im = insertion for the m. biceps brachii. Scale bars:
|
||
<quantity id="8A6EE1BED73BFF98FE65DFE51860A6DB" box="[503,564,1999,2025]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="63" pageNumber="64" unit="cm" value="2.0">2 cm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4D294C5BD745FFE6FF78D9221848A011" blockId="65.[188,1398,264,357]" box="[234,540,264,291]" pageId="65" pageNumber="66">
|
||
<emphasis id="7FE29049D745FFE6FF78D9221848A011" bold="true" box="[234,540,264,291]" pageId="65" pageNumber="66">Material from Bumbach</emphasis>
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD745FFE6FF2FD90018FAA057" blockId="65.[188,1398,264,357]" pageId="65" pageNumber="66">
|
||
Newly prepared specimens are referred to as
|
||
<taxonomicName id="8A9637D8D745FFE6FD5AD900199AA076" authority="Tissier & Antoine & Becker, 2021" authorityName="Tissier & Antoine & Becker" authorityYear="2021" box="[712,974,298,324]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="heissigi" status="sp. nov.">
|
||
<emphasis id="7FE29049D745FFE6FD5AD900199AA076" box="[712,974,298,324]" italics="true" pageId="65" pageNumber="66">Ronzotherium heissigi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="64D12D32D745FFE6FC46D9001E7AA076" box="[980,1070,298,324]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and they document the only known postcranial remains of this species.
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD745FFE6FF2FD9A11829A37F" blockId="65.[189,1400,395,590]" pageId="65" pageNumber="66">
|
||
HUMERUS. The two humeri from Bumbach are proximally broken (
|
||
<figureCitation id="D5AD50DED745FFE6FC46D9A11E30A094" box="[980,1124,395,422]" captionStart="Fig" captionStartId="63.[189,232,1787,1813]" captionTargetId="figure-13@64.[189,1398,265,1957]" captionTargetPageId="64" captionText="Fig. 25 (next page). Ronzotherium heissigi sp. nov. from Bumbach, Bern Canton (early late Oligocene; Switzerland). – A–D. Mandible NMB-UM-6132. A. Occlusal view. B. Left lateral view. C. With right p3–m3 in labial view. D. With right p3–m3 in lingual view. – E–F. Right humerus NMB-UM-129a. E. Anterior view.F. Posterior view. –G–H. Left humerus NMB-UM-132. G. Anterior view.H. Posterior view. – I–J. Right radius NMB-UM-127a. I. Anterior view.J. Posterior view. – K–M. Right ulna NMB- UM-131b. K. Anterior view. L. Posterior view. M. Distal view. Abbreviation: im = insertion for the m. biceps brachii. Scale bars: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.4959442" httpUri="https://zenodo.org/record/4959442/files/figure.png" pageId="65" pageNumber="66">Fig. 25E–H</figureCitation>
|
||
). In posterior view, the fossa olecrani is high on NMB-UM-132, but lower on the other humerus, NMB-UM-6132, possibly due to taphonomical deformation. The lateral epicondyle is very prominent, and distally elongated. The distal articulation is hourglass-shaped in anterior view, with a deep proximal incision between the two lips of the trochlea and the articulation is oblique. The epicondylar crest is high. The distal gutter on the epicondyle is strong in posterior view.
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD745FFE6FF2FDA5F18C9A229" blockId="65.[188,1399,628,795]" pageId="65" pageNumber="66">
|
||
RADIUS. The radius from Bumbach (NMB-UM-127a,
|
||
<figureCitation id="D5AD50DED745FFE6FCD9DA5F199CA3BD" box="[843,968,628,655]" captionStart="Fig" captionStartId="63.[189,232,1787,1813]" captionTargetId="figure-13@64.[189,1398,265,1957]" captionTargetPageId="64" captionText="Fig. 25 (next page). Ronzotherium heissigi sp. nov. from Bumbach, Bern Canton (early late Oligocene; Switzerland). – A–D. Mandible NMB-UM-6132. A. Occlusal view. B. Left lateral view. C. With right p3–m3 in labial view. D. With right p3–m3 in lingual view. – E–F. Right humerus NMB-UM-129a. E. Anterior view.F. Posterior view. –G–H. Left humerus NMB-UM-132. G. Anterior view.H. Posterior view. – I–J. Right radius NMB-UM-127a. I. Anterior view.J. Posterior view. – K–M. Right ulna NMB- UM-131b. K. Anterior view. L. Posterior view. M. Distal view. Abbreviation: im = insertion for the m. biceps brachii. Scale bars: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.4959442" httpUri="https://zenodo.org/record/4959442/files/figure.png" pageId="65" pageNumber="66">Fig. 25I–J</figureCitation>
|
||
) belongs to the same individual as the humerus NMB-UM-132. The anterior border of the proximal articulation is straight. The medial border of the diaphysis is straight. The insertion of the m. biceps brachii is deep and the gutter for the m. extensor carpi is deep and wide. The posterior expansion of the scaphoid facet is high and there is no secondary distal articulation for the ulna.
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD745FFE6FF2FDB691E66A2F5" blockId="65.[189,1399,835,967]" pageId="65" pageNumber="66">
|
||
ULNA. The ulna NMB-UM-131b-c (
|
||
<figureCitation id="D5AD50DED745FFE6FD16DB691974A26C" box="[644,800,835,862]" captionStart="Fig" captionStartId="63.[189,232,1787,1813]" captionTargetId="figure-13@64.[189,1398,265,1957]" captionTargetPageId="64" captionText="Fig. 25 (next page). Ronzotherium heissigi sp. nov. from Bumbach, Bern Canton (early late Oligocene; Switzerland). – A–D. Mandible NMB-UM-6132. A. Occlusal view. B. Left lateral view. C. With right p3–m3 in labial view. D. With right p3–m3 in lingual view. – E–F. Right humerus NMB-UM-129a. E. Anterior view.F. Posterior view. –G–H. Left humerus NMB-UM-132. G. Anterior view.H. Posterior view. – I–J. Right radius NMB-UM-127a. I. Anterior view.J. Posterior view. – K–M. Right ulna NMB- UM-131b. K. Anterior view. L. Posterior view. M. Distal view. Abbreviation: im = insertion for the m. biceps brachii. Scale bars: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.4959442" httpUri="https://zenodo.org/record/4959442/files/figure.png" pageId="65" pageNumber="66">Fig. 25K–M</figureCitation>
|
||
) belongs to the same individual as the radius (NMB-UM-127a) and the humerus (NMB-UM-132). It is very poorly preserved. The olecranon of the ulna is rather long and makes an open angle with the diaphysis. The distal end is large, the anterior tubercle and the lunate facet are absent, and the pyramidal facet is concave.
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD745FFE6FF2FDBC4181DA477" blockId="65.[188,1400,1006,1349]" pageId="65" pageNumber="66">
|
||
LUNATE. The lunate NMBE-5035833 is complete and very well preserved (
|
||
<figureCitation id="D5AD50DED745FFE6FBA1DBC41E97A53B" box="[1075,1219,1006,1033]" captionStart="Fig" captionStartId="65.[188,231,1680,1706]" captionTargetId="figure-13@66.[189,1399,265,1975]" captionTargetPageId="66" captionText="Fig. 26 (next page). Ronzotherium heissigi sp. nov. from Bumbach, Bern Canton (early late Oligocene; Switzerland). – A–E. Left lunate NMBE-5035833. A. Anterior view. B. Proximal view. C. Distal view. D. Lateral view. E. Medial view. – F–J. Right trapezoid NMB-UM-6136b. F. Anterior view. G. Lateral view. H. Medial view. I. Proximal view. J. Distal view. – K–N. Right magnum NMB-UM- 6136c. K. Anterior view. L. Distal view. M. Lateral view. N. Medial view. – O–Q. Right McII NMB- UM-121. O. Anterior view. P. Lateral view. Q. Proximal view. – R–T. Right McII NMB-UM-6136a. R. Proximal view. S. Anterior view. T. Lateral view. – U–Y. Right cuboid NMBE-5035834. U. Anterior view. V. Proximal view. W. Distal view. X. Medial view. Y. Lateral view. Abbreviations: a = astragalus; ca = calcaneus; e = ectocuneiform; l = lunate; m = magnum; n = navicular; p = pyramidal; r = radius; s = scaphoid; td = trapezoid; tm = trapezium; un = unciform. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.4959446" httpUri="https://zenodo.org/record/4959446/files/figure.png" pageId="65" pageNumber="66">Fig. 26A–E</figureCitation>
|
||
). The proximal articulation for the radius is very large and convex anteroposteriorly. There is no articulation with the ulna. In proximal view, there is a long drop-like posterior extension of the radius facet on the medial side. The anterior side is deeply keeled, and the distal border is acute in anterior view. Medially, the two proximal articulations for the scaphoid are fused in a single facet. It is separated from the distal scaphoid facet by a deep groove. The distal facet for the scaphoid is large. In lateral view, the proximal and distal articulations for the pyramidal are large and the proximal one is medially displaced. In distal view, there are three articulation facets: a large anterior one for the unciform, and two for the magnum, one of which is distal and very concave and the other thin, flat and elongated is anterior, and located between the scaphoid and unciform facets.
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD745FFE6FF2FDD471F22A721" blockId="65.[188,1400,1388,1556]" pageId="65" pageNumber="66">
|
||
TRAPEZOID. Two trapezoids are preserved from Bumbach (NMB-UM-6 and NMB-UM-6136b,
|
||
<figureCitation id="D5AD50DED745FFE6FF2FDDA51B12A498" box="[189,326,1423,1450]" captionStart="Fig" captionStartId="65.[188,231,1680,1706]" captionTargetId="figure-13@66.[189,1399,265,1975]" captionTargetPageId="66" captionText="Fig. 26 (next page). Ronzotherium heissigi sp. nov. from Bumbach, Bern Canton (early late Oligocene; Switzerland). – A–E. Left lunate NMBE-5035833. A. Anterior view. B. Proximal view. C. Distal view. D. Lateral view. E. Medial view. – F–J. Right trapezoid NMB-UM-6136b. F. Anterior view. G. Lateral view. H. Medial view. I. Proximal view. J. Distal view. – K–N. Right magnum NMB-UM- 6136c. K. Anterior view. L. Distal view. M. Lateral view. N. Medial view. – O–Q. Right McII NMB- UM-121. O. Anterior view. P. Lateral view. Q. Proximal view. – R–T. Right McII NMB-UM-6136a. R. Proximal view. S. Anterior view. T. Lateral view. – U–Y. Right cuboid NMBE-5035834. U. Anterior view. V. Proximal view. W. Distal view. X. Medial view. Y. Lateral view. Abbreviations: a = astragalus; ca = calcaneus; e = ectocuneiform; l = lunate; m = magnum; n = navicular; p = pyramidal; r = radius; s = scaphoid; td = trapezoid; tm = trapezium; un = unciform. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.4959446" httpUri="https://zenodo.org/record/4959446/files/figure.png" pageId="65" pageNumber="66">Fig. 26F–J</figureCitation>
|
||
). The latter is slightly different from the former, and it articulates with the magnum NMB-UM-6136c and the McII NMB-UM-6136a. In anterior view, they are both wider than high. The proximal border is sigmoid on NMB-UM-6136b, whereas it is symmetric on NMB-UM-6. The magnum facet occupies most of the lateral side, while the medial side is occupied by the medio-distal articulation
|
||
</paragraph>
|
||
<caption id="19E91CD3D745FFE6FF2EDEBA1E4AA6DA" ID-DOI="http://doi.org/10.5281/zenodo.4959446" ID-Zenodo-Dep="4959446" httpUri="https://zenodo.org/record/4959446/files/figure.png" pageId="65" pageNumber="66" startId="65.[188,231,1680,1706]" subCaptionStartIDs="65.[1050,1221,1927,1953]" subCaptionStarts="Abbr" targetBox="[189,1399,265,1975]" targetPageId="66">
|
||
<paragraph id="4D294C5BD745FFE6FF2EDEBA1E4AA6DA" blockId="65.[187,1399,1680,2024]" pageId="65" pageNumber="66">
|
||
<emphasis id="7FE29049D745FFE6FF2EDEBA1B45A798" bold="true" box="[188,273,1680,1706]" pageId="65" pageNumber="66">Fig. 26</emphasis>
|
||
(next page).
|
||
<taxonomicName id="8A9637D8D745FFE6FE3CDEBA18E2A798" authority="Tissier & Antoine & Becker, 2021" authorityName="Tissier & Antoine & Becker" authorityYear="2021" box="[430,694,1680,1706]" class="Mammalia" family="Rhinocerotidae" genus="Ronzotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perissodactyla" pageId="65" pageNumber="66" phylum="Chordata" rank="species" species="heissigi" status="sp. nov.">
|
||
<emphasis id="7FE29049D745FFE6FE3CDEBA18E2A798" box="[430,694,1680,1706]" italics="true" pageId="65" pageNumber="66">Ronzotherium heissigi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="64D12D32D745FFE6FD2FDEBB194CA799" box="[701,792,1681,1707]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from Bumbach,
|
||
<collectingRegion id="8F5282B9D745FFE6FC4FDEBB1E42A799" box="[989,1046,1681,1707]" country="Switzerland" name="Bern" pageId="65" pageNumber="66">Bern</collectingRegion>
|
||
Canton (early late Oligocene;
|
||
<collectingCountry id="35810CCBD745FFE6FF2FDE991B1BA7FC" box="[189,335,1715,1742]" name="Switzerland" pageId="65" pageNumber="66">Switzerland</collectingCountry>
|
||
). –
|
||
<emphasis id="7FE29049D745FFE6FE10DE991BEEA7FF" bold="true" box="[386,442,1715,1741]" pageId="65" pageNumber="66">A–E</emphasis>
|
||
. Left lunate NMBE-5035833.
|
||
<emphasis id="7FE29049D745FFE6FCABDE99191AA7FF" bold="true" box="[825,846,1715,1741]" pageId="65" pageNumber="66">A</emphasis>
|
||
. Anterior view.
|
||
<emphasis id="7FE29049D745FFE6FB8EDE9E1E64A7FC" bold="true" box="[1052,1072,1716,1742]" pageId="65" pageNumber="66">B</emphasis>
|
||
. Proximal view.
|
||
<emphasis id="7FE29049D745FFE6FA95DE991F48A7FF" bold="true" box="[1287,1308,1715,1741]" pageId="65" pageNumber="66">C</emphasis>
|
||
. Distal view.
|
||
<emphasis id="7FE29049D745FFE6FE95DEFD1B48A7C3" bold="true" box="[263,284,1751,1777]" pageId="65" pageNumber="66">D</emphasis>
|
||
. Lateral view.
|
||
<emphasis id="7FE29049D745FFE6FE44DEFD1BBEA7C3" bold="true" box="[470,490,1751,1777]" pageId="65" pageNumber="66">E</emphasis>
|
||
. Medial view. –
|
||
<emphasis id="7FE29049D745FFE6FD2CDEFD18BAA7C3" bold="true" box="[702,750,1751,1777]" pageId="65" pageNumber="66">F–J</emphasis>
|
||
. Right trapezoid NMB-UM-6136b.
|
||
<emphasis id="7FE29049D745FFE6FB34DEFD1EECA7C3" bold="true" box="[1190,1208,1751,1777]" pageId="65" pageNumber="66">F</emphasis>
|
||
. Anterior view.
|
||
<emphasis id="7FE29049D745FFE6FF2DDED01A80A626" bold="true" box="[191,212,1786,1812]" pageId="65" pageNumber="66">G</emphasis>
|
||
. Lateral view.
|
||
<emphasis id="7FE29049D745FFE6FE14DED01BC8A626" bold="true" box="[390,412,1786,1812]" pageId="65" pageNumber="66">H</emphasis>
|
||
. Medial view.
|
||
<emphasis id="7FE29049D745FFE6FDDDDED0180EA626" bold="true" box="[591,602,1786,1812]" pageId="65" pageNumber="66">I</emphasis>
|
||
. Proximal view.
|
||
<emphasis id="7FE29049D745FFE6FCB6DED01966A626" bold="true" box="[804,818,1786,1812]" pageId="65" pageNumber="66">J</emphasis>
|
||
. Distal view. –
|
||
<emphasis id="7FE29049D745FFE6FC7DDED01E7DA626" bold="true" box="[1007,1065,1786,1812]" pageId="65" pageNumber="66">K–N</emphasis>
|
||
. Right magnum NMB-UM- 6136c.
|
||
<emphasis id="7FE29049D745FFE6FE86DF371B7FA605" bold="true" box="[276,299,1821,1847]" pageId="65" pageNumber="66">K</emphasis>
|
||
. Anterior view.
|
||
<emphasis id="7FE29049D745FFE6FE79DF371BABA605" bold="true" box="[491,511,1821,1847]" pageId="65" pageNumber="66">L</emphasis>
|
||
. Distal view.
|
||
<emphasis id="7FE29049D745FFE6FD34DF371895A605" bold="true" box="[678,705,1821,1847]" pageId="65" pageNumber="66">M</emphasis>
|
||
. Lateral view.
|
||
<emphasis id="7FE29049D745FFE6FCE1DF3719D3A605" bold="true" box="[883,903,1821,1847]" pageId="65" pageNumber="66">N</emphasis>
|
||
. Medial view. –
|
||
<emphasis id="7FE29049D745FFE6FBC0DF371ED9A605" bold="true" box="[1106,1165,1821,1847]" pageId="65" pageNumber="66">O–Q</emphasis>
|
||
. Right McII NMB- UM-121.
|
||
<emphasis id="7FE29049D745FFE6FEA0DF6A1B1DA668" bold="true" box="[306,329,1856,1882]" pageId="65" pageNumber="66">O</emphasis>
|
||
. Anterior view.
|
||
<emphasis id="7FE29049D745FFE6FD9EDF6A184AA668" bold="true" box="[524,542,1856,1882]" pageId="65" pageNumber="66">P</emphasis>
|
||
. Lateral view.
|
||
<emphasis id="7FE29049D745FFE6FD41DF6A18BCA668" bold="true" box="[723,744,1856,1882]" pageId="65" pageNumber="66">Q</emphasis>
|
||
. Proximal view. –
|
||
<emphasis id="7FE29049D745FFE6FC5CDF6A1E50A668" bold="true" box="[974,1028,1856,1882]" pageId="65" pageNumber="66">R–T</emphasis>
|
||
. Right McII NMB-UM-6136a.
|
||
<emphasis id="7FE29049D745FFE6FF2DDF4E1A87A64C" bold="true" box="[191,211,1892,1918]" pageId="65" pageNumber="66">R</emphasis>
|
||
. Proximal view.
|
||
<emphasis id="7FE29049D745FFE6FE05DF491BF3A64F" bold="true" box="[407,423,1891,1917]" pageId="65" pageNumber="66">S</emphasis>
|
||
. Anterior view.
|
||
<emphasis id="7FE29049D745FFE6FDF3DF4E1821A64C" bold="true" box="[609,629,1892,1918]" pageId="65" pageNumber="66">T</emphasis>
|
||
. Lateral view. –
|
||
<emphasis id="7FE29049D745FFE6FCAADF4E1925A64C" bold="true" box="[824,881,1892,1918]" pageId="65" pageNumber="66">U–Y</emphasis>
|
||
. Right cuboid NMBE-5035834.
|
||
<emphasis id="7FE29049D745FFE6FB63DF4E1F52A64C" bold="true" box="[1265,1286,1892,1918]" pageId="65" pageNumber="66">U</emphasis>
|
||
. Anterior view.
|
||
<emphasis id="7FE29049D745FFE6FE90DFAD1B43A693" bold="true" box="[258,279,1927,1953]" pageId="65" pageNumber="66">V</emphasis>
|
||
. Proximal view.
|
||
<emphasis id="7FE29049D745FFE6FE49DFAD1BACA693" bold="true" box="[475,504,1927,1953]" pageId="65" pageNumber="66">W</emphasis>
|
||
. Distal view.
|
||
<emphasis id="7FE29049D745FFE6FD0ADFAD18F9A693" bold="true" box="[664,685,1927,1953]" pageId="65" pageNumber="66">X</emphasis>
|
||
. Medial view.
|
||
<emphasis id="7FE29049D745FFE6FCCBDFAD193AA693" bold="true" box="[857,878,1927,1953]" pageId="65" pageNumber="66">Y</emphasis>
|
||
. Lateral view. Abbreviations: a = astragalus; ca = calcaneus; e = ectocuneiform; l = lunate; m = magnum; n = navicular; p = pyramidal; r = radius; s = scaphoid; td = trapezoid; tm = trapezium; un = unciform. Scale bar:
|
||
<quantity id="8A6EE1BED745FFE6FC4EDFE71E4DA6DA" box="[988,1049,1997,2024]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="65" pageNumber="66" unit="cm" value="2.0">2 cm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4D294C5BD747FFE4FF2FD92319B2A075" blockId="67.[189,1399,265,327]" pageId="67" pageNumber="68">for the trapezium. The proximal side is fully occupied by the anteroposteriorly concave scaphoid facet. The distal articulation for the McII is also anteroposteriorly concave.</paragraph>
|
||
<paragraph id="4D294C5BD747FFE4FF2FD944192AA34D" blockId="67.[188,1399,366,640]" pageId="67" pageNumber="68">
|
||
MAGNUM. The magnum NMB-UM-6136c is complete (
|
||
<figureCitation id="D5AD50DED747FFE4FCC9D94419A6A0BB" box="[859,1010,366,393]" captionStart="Fig" captionStartId="65.[188,231,1680,1706]" captionTargetId="figure-13@66.[189,1399,265,1975]" captionTargetPageId="66" captionText="Fig. 26 (next page). Ronzotherium heissigi sp. nov. from Bumbach, Bern Canton (early late Oligocene; Switzerland). – A–E. Left lunate NMBE-5035833. A. Anterior view. B. Proximal view. C. Distal view. D. Lateral view. E. Medial view. – F–J. Right trapezoid NMB-UM-6136b. F. Anterior view. G. Lateral view. H. Medial view. I. Proximal view. J. Distal view. – K–N. Right magnum NMB-UM- 6136c. K. Anterior view. L. Distal view. M. Lateral view. N. Medial view. – O–Q. Right McII NMB- UM-121. O. Anterior view. P. Lateral view. Q. Proximal view. – R–T. Right McII NMB-UM-6136a. R. Proximal view. S. Anterior view. T. Lateral view. – U–Y. Right cuboid NMBE-5035834. U. Anterior view. V. Proximal view. W. Distal view. X. Medial view. Y. Lateral view. Abbreviations: a = astragalus; ca = calcaneus; e = ectocuneiform; l = lunate; m = magnum; n = navicular; p = pyramidal; r = radius; s = scaphoid; td = trapezoid; tm = trapezium; un = unciform. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.4959446" httpUri="https://zenodo.org/record/4959446/files/figure.png" pageId="67" pageNumber="68">Fig. 26K–N</figureCitation>
|
||
). In anterior view, the proximal border of the anterior side is concave and the anterior face is wider than high. The proximal apophysis is very high, very convex and narrow. The lunate facet on this apophysis is very long and contacts the small unciform facet. On the other side of the proximal apophysis, the facet for the scaphoid is also long and is very poorly distinguished from the medial facet for the trapezoid. This latter facet is longer than high. The medio-distal McII facet is flat and its distal border is very concave in medial view. On the distal side, the McIII facet is very large, quadrate, as long as wide and very concave anteroposteriorly. The posterior tuberosity of the magnum is short and curved.
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD747FFE4FF2FDA8D1909A27C" blockId="67.[188,1398,679,847]" pageId="67" pageNumber="68">
|
||
MCII. There are two McII preserved from Bumbach (NMB-UM-121 and McII NMB-6136a,
|
||
<figureCitation id="D5AD50DED747FFE4FA93DA8D1A86A3D7" captionStart="Fig" captionStartId="65.[188,231,1680,1706]" captionTargetId="figure-13@66.[189,1399,265,1975]" captionTargetPageId="66" captionText="Fig. 26 (next page). Ronzotherium heissigi sp. nov. from Bumbach, Bern Canton (early late Oligocene; Switzerland). – A–E. Left lunate NMBE-5035833. A. Anterior view. B. Proximal view. C. Distal view. D. Lateral view. E. Medial view. – F–J. Right trapezoid NMB-UM-6136b. F. Anterior view. G. Lateral view. H. Medial view. I. Proximal view. J. Distal view. – K–N. Right magnum NMB-UM- 6136c. K. Anterior view. L. Distal view. M. Lateral view. N. Medial view. – O–Q. Right McII NMB- UM-121. O. Anterior view. P. Lateral view. Q. Proximal view. – R–T. Right McII NMB-UM-6136a. R. Proximal view. S. Anterior view. T. Lateral view. – U–Y. Right cuboid NMBE-5035834. U. Anterior view. V. Proximal view. W. Distal view. X. Medial view. Y. Lateral view. Abbreviations: a = astragalus; ca = calcaneus; e = ectocuneiform; l = lunate; m = magnum; n = navicular; p = pyramidal; r = radius; s = scaphoid; td = trapezoid; tm = trapezium; un = unciform. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.4959446" httpUri="https://zenodo.org/record/4959446/files/figure.png" pageId="67" pageNumber="68">Fig. 26R– T</figureCitation>
|
||
). The outline of the proximal side is trapezoidal. The articulation for the trapezoid is concave lateromedially and slightly convex anteroposteriorly. On the lateral side, the magnum facet is curved and band-shaped. The anterior facet for the McIII is present but the posterior is absent. There is no trapezium facet. The insertion for the m. extensor carpalis is salient.
|
||
</paragraph>
|
||
<paragraph id="4D294C5BD747FFE4FF2FDB5C1F02A52F" blockId="67.[188,1400,885,1053]" pageId="67" pageNumber="68">FEMUR. Only a proximal fragment is preserved in Bumbach (NMBE-5035835). The head is hemispheric and the fovea capitis is high and narrow. The major trochanter is lower than the head and separated from it by a short neck. It is nearly triangular in proximal view and in lateral view the posterior part is higher than the anterior. The trochanteric fossa is deep on the posterior side, with a well-marked intertrochanteric crest. A fossa is also present on the anterior side, overhung by the major trochanter.</paragraph>
|
||
<paragraph id="4D294C5BD747FFE4FF2FDC6F1BC0A4AE" blockId="67.[189,1400,1092,1436]" pageId="67" pageNumber="68">
|
||
CUBOID. The cuboid NMBE-5035834 is complete (
|
||
<figureCitation id="D5AD50DED747FFE4FC85DC6E19FEA56D" box="[791,938,1092,1119]" captionStart="Fig" captionStartId="65.[188,231,1680,1706]" captionTargetId="figure-13@66.[189,1399,265,1975]" captionTargetPageId="66" captionText="Fig. 26 (next page). Ronzotherium heissigi sp. nov. from Bumbach, Bern Canton (early late Oligocene; Switzerland). – A–E. Left lunate NMBE-5035833. A. Anterior view. B. Proximal view. C. Distal view. D. Lateral view. E. Medial view. – F–J. Right trapezoid NMB-UM-6136b. F. Anterior view. G. Lateral view. H. Medial view. I. Proximal view. J. Distal view. – K–N. Right magnum NMB-UM- 6136c. K. Anterior view. L. Distal view. M. Lateral view. N. Medial view. – O–Q. Right McII NMB- UM-121. O. Anterior view. P. Lateral view. Q. Proximal view. – R–T. Right McII NMB-UM-6136a. R. Proximal view. S. Anterior view. T. Lateral view. – U–Y. Right cuboid NMBE-5035834. U. Anterior view. V. Proximal view. W. Distal view. X. Medial view. Y. Lateral view. Abbreviations: a = astragalus; ca = calcaneus; e = ectocuneiform; l = lunate; m = magnum; n = navicular; p = pyramidal; r = radius; s = scaphoid; td = trapezoid; tm = trapezium; un = unciform. Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.4959446" httpUri="https://zenodo.org/record/4959446/files/figure.png" pageId="67" pageNumber="68">Fig. 26U–Y</figureCitation>
|
||
). The anterior side is approximately as high as wide. In lateral view, the posterior side is higher than the anterior and the posterior apophysis of the cuboid is slightly lower than the distal articulation. In proximal view, the posterior apophysis is almost not visible. The proximal side is oval and the astragalus and calcaneal facets are almost equalsized. On the medial side, the postero-proximal and elongated facet for the navicular is concave and contacts the small, square and postero-mesial facet for the ectocuneiform. The navicular facet bears a thin extension up to the anterior border, bordering the astragalus facet. The small anterodistal facet for the ectocuneiform is separated from the posterior one by a wide groove. On the lateral side, the groove for the tendon of the m. fibularis longus is very large and deep. The distal articulation for the MtIV is flat and triangular.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |