260 lines
35 KiB
XML
260 lines
35 KiB
XML
<document ID-CLB-Dataset="21464" ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="C4C1340F83DE54568B677A4731BC6DFD" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole flavominuta Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="59199444-7C41-420D-8302-7EFBE5525DF0" docUuidSource="ZooBank" docVersion="6" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1732651982216" updateUser="ExternalLinkService">
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<mods:mods id="FBE354A2B95A12F6C674EF0D2D4D9CCC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="70F4633691D25F735B06FFC2890AD486">
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<mods:title id="179D16075DD2A90E149C533A9278A6E7">Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name id="45C06999FF7E9EE4A04ED78A913DA1D1" type="personal">
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<mods:roleTerm id="A0A81B130B6CF4E159185713D2C9C51D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5700C0A2BCCCB5B5D1EE19A71BA58AA5">Salata, Sebastian</mods:namePart>
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<mods:affiliation id="A57CBBF3D3CDB98A7521DEF705705483">Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier id="130D1FE5F6632382F599E6B2D1F35806" type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
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<mods:nameIdentifier id="AF060644B38890D6ABC8C2518899D277" type="email">sdsalata@gmail.com</mods:nameIdentifier>
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<mods:name id="E153BE9F75A46D0713A5C95A13AFE2A9" type="personal">
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<mods:roleTerm id="CADC4F77B6BA6CAF83E191FFE4213FBD">Author</mods:roleTerm>
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<mods:namePart id="53FC3BA2245175F0E3650AB13F029BEA">Fisher, Brian L.</mods:namePart>
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<mods:affiliation id="A7B94DB1863EEB9A8FE88DCBE2F7A0F8">Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier id="3DBBC9917087DE9A971AC5BF7285E5F6" type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
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<mods:typeOfResource id="D88D272B3E67D7156F13E7170A2AF521">text</mods:typeOfResource>
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<mods:titleInfo id="A8A02561F0EAF6B0603C311CEFD9EF77">
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<mods:title id="488573621A02ECC14F0631C295B622C2">ZooKeys</mods:title>
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<mods:part id="A58DDE75129793F3FE5777E07112F6F9">
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<mods:date id="8C236AE57D27482515CD06ADF81E6187">2020</mods:date>
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<mods:detail id="798237397DA6C4C32CE06D6CEA98BE96" type="volume">
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<mods:number id="D26083F82555DBDD2A6BC0DD0C701486">949</mods:number>
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<mods:start id="3C5DD9EAF943AC5E5FB1EFEE9B386396">1</mods:start>
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<mods:end id="35986FF9E86F2EFF0D284F2756454CE9">185</mods:end>
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<mods:location id="6187DADE376E6B021216381ADDF441EF">
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<mods:url id="954517B1CE1AA83791ECC208E6955F50">http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
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</mods:location>
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<mods:classification id="F536883EE13EA77E6FE976BE180E814C">journal article</mods:classification>
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<mods:identifier id="519AF9C9FC89CBC99CD2285ACEAE27CE" type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
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<mods:identifier id="616D08BD735B988673DD4CA19C404A86" type="Pensoft-Pub">1313-2970-949-1</mods:identifier>
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<treatment id="C4C1340F83DE54568B677A4731BC6DFD" ID-GBIF-Taxon="165090165" LSID="urn:lsid:zoobank.org:act:59199444-7C41-420D-8302-7EFBE5525DF0" httpUri="http://treatment.plazi.org/id/C4C1340F83DE54568B677A4731BC6DFD" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="1B895F18DCC7CB63F75DC99A35805510" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="AA882702DB48E54B5A56963DA3BB31E3" pageId="0" pageNumber="1">
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<taxonomicName id="B4EA1CF596E021B3CCEEC8F50819EBA0" ID-CoL="CML65" LSID="C4C1340F-83DE-5456-8B67-7A4731BC6DFD" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole flavominuta" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="flavominuta" status="sp. nov.">Pheidole flavominuta</taxonomicName>
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<taxonomicNameLabel id="3326B44EA94E5C4537FF86FB449A428E" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation id="AF4FC4EEDD10357EED01FBC77140EAEB" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Pheidole flavominuta sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0162186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923272)." figureDoi="10.3897/zookeys.949.51269.figure29" httpUri="https://binary.pensoft.net/fig/432198" pageId="0" pageNumber="1">Figs 29A-F</figureCitation>
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<figureCitation id="0004E77EA185F7549935AE2BA248A488" captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">, 63K</figureCitation>
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<figureCitation id="21EB97FFBAC204A1E6B145E26925AC0F" captionStart="Figure 65" captionStartId="F65" captionText="Figure 65. Distribution. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T)." figureDoi="10.3897/zookeys.949.51269.figure65" httpUri="https://binary.pensoft.net/fig/432234" pageId="0" pageNumber="1">, 65K</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="553B834776F9345524510075AAE1053C" pageId="0" pageNumber="1" type="type material">
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<paragraph id="8843A39397BBBE3130761CE150AD455B" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="F0CA077F8E40002E8E9AF09B5E920179" pageId="0" pageNumber="1">
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<emphasis id="F2EE0CC1A1FB3689EC726B01EAB36EDF" bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
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Madagascar. • 1 major worker; Toamasina;
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<normalizedToken id="232922EC4E488CC154C013DCFB1FE2A1" originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken id="0AB098166FAF6C5998C0BB75CF222222" originalValue="Spéciale">Speciale</normalizedToken>
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Ambatovaky, Sandrangato river;
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<geoCoordinate id="E64E1C394F6399B3D479DE867E799E34" degrees="16.81745" direction="south" orientation="latitude" precision="1" value="-16.81745">-16.81745</geoCoordinate>
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,
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<geoCoordinate id="4A00411EDA2022EA233A6CFD9B65B7A4" degrees="49.2925" direction="east" orientation="longitude" precision="5" value="49.2925">49.2925</geoCoordinate>
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; alt. 400 m; 26 Feb 2010; B. L. Fisher et al. leg.; rainforest, under root mat on ground; BLF24911; CASENT0923272 (CASC).
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<emphasis id="35781309CB46ABAA308C0AE54A386278" bold="true" italics="true" pageId="0" pageNumber="1">Paratype.</emphasis>
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• 1w.; same data as for holotype, CASENT0162186 (CASC).
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</paragraph>
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<caption id="98F426075DB532DE77630D87F838E6D6" doi="10.3897/zookeys.949.51269.figure29" httpUri="https://binary.pensoft.net/fig/432198" pageId="0" pageNumber="1" start="Figure 29" startId="F29">
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<paragraph id="B7132E0B615FAAEC8A1D31B64ECCEA49" pageId="0" pageNumber="1">
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<emphasis id="5EC690D11B580444E5D44D6AC53A9D03" bold="true" pageId="0" pageNumber="1">Figure 29.</emphasis>
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<taxonomicName id="7566F1177AF4186A0FCF9AF9948906C2" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole flavominuta" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="flavominuta">
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<emphasis id="E890147ECED83A78A1B072B28626F15C" italics="true" pageId="0" pageNumber="1">Pheidole flavominuta</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis id="CB05027EF39F4E645880A8A0EAB3096F" bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis id="E3580305F88644C2E1E2E5E7A0C53289" bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis id="AF0B16B92094260A8EB267D21DDD3103" bold="true" pageId="0" pageNumber="1">E</emphasis>
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) of paratype minor worker (CASENT0162186) and full-face view (
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<emphasis id="EC6DE713E5F658A2D50CCB7CA55BD65F" bold="true" pageId="0" pageNumber="1">B</emphasis>
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), profile (
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<emphasis id="755F71A95E4370DCDA72DDEBB5404D71" bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis id="162934A80AFA7F2DB9B08CD323D4D322" bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0923272).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="A3AAA51CB2E0E728E478E910209CE6F3" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="D609F82780427EEEF7FA7B91C3E1C379" pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph id="13879180507C42072630CAAF4C36EE70" pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis id="851A9AD5994895B827B6F32BE5A9571F" bold="true" pageId="0" pageNumber="1">Toamasina</emphasis>
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: • 1w., 1s.; Reserve Betampona, Camp Vohitsivalana, 37.1 km 338° Toamasina;
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<geoCoordinate id="FD731E4165ACA9592A2CC3CF5BDE84B1" degrees="17.88667" direction="south" orientation="latitude" precision="1" value="-17.88667">-17.88667</geoCoordinate>
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,
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<geoCoordinate id="ECCD44A5B8DDCF55B317605494860353" degrees="49.2025" direction="east" orientation="longitude" precision="5" value="49.2025">49.2025</geoCoordinate>
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; alt. 520 m; 1 Dec 2005; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF13262 (CASC). • 1w., 1s.;
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<normalizedToken id="426416831745E48AE9CEDCC1C16DA189" originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken id="C18425DD090C1D79C57255F375711C8D" originalValue="Spéciale">Speciale</normalizedToken>
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Ambatovaky, Sandrangato river;
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<geoCoordinate id="51928B1946FAA14E9EB446B407395399" degrees="16.77468" direction="south" orientation="latitude" precision="1" value="-16.77468">-16.77468</geoCoordinate>
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,
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<geoCoordinate id="780F59246139E8F4D6543E69264E20C8" degrees="49.26551" direction="east" orientation="longitude" precision="1" value="49.26551">49.26551</geoCoordinate>
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; alt. 355 m; 21 Feb 2010; B. L. Fisher et al. leg.; rainforest along river, under rootmat, litter on rock; BLF24505 (CASC). • 1w., 1s.;
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<normalizedToken id="FDA3F0D977C26AC5A8ECB01FDB2FFB9A" originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken id="2E754C103BA73B20052E4B274B8A4F41" originalValue="Spéciale">Speciale</normalizedToken>
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Ambatovaky, Sandrangato river;
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<geoCoordinate id="0611BC52C1914F5AF5935BEBDD032E4C" degrees="16.81745" direction="south" orientation="latitude" precision="1" value="-16.81745">-16.81745</geoCoordinate>
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,
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<geoCoordinate id="42FD94B703A15797EA39D7FA2D13FE8A" degrees="49.2925" direction="east" orientation="longitude" precision="5" value="49.2925">49.2925</geoCoordinate>
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; alt. 400 m; 26 Feb 2010; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF24913 (CASC). • 2w.;
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<normalizedToken id="ED8F29F7FAF2C469D6FCED0E6FA83C76" originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken id="6D1867CB8BB4ACB18C5A4423A64476A8" originalValue="Spéciale">Speciale</normalizedToken>
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Ambatovaky, Sandrangato river;
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<geoCoordinate id="1F7726CFA8DB3E7DAAB10A42BD9C80DA" degrees="16.81745" direction="south" orientation="latitude" precision="1" value="-16.81745">-16.81745</geoCoordinate>
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,
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<geoCoordinate id="812A4A081525F75F0DAF9A560BCA533E" degrees="49.2925" direction="east" orientation="longitude" precision="5" value="49.2925">49.2925</geoCoordinate>
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; alt. 400 m; 26 Feb 2010; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF24943 (CASC). • 1w., 1s.;
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<normalizedToken id="1EA2C08A3F4AD3DA229BA0BF46353BFB" originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken id="C9BC7C50F7B26CB0B2C0488C118EC734" originalValue="Spéciale">Speciale</normalizedToken>
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Ambatovaky, Sandrangato river;
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<geoCoordinate id="A5693C99899AD101BFEB9F34C9FC6B8F" degrees="16.81745" direction="south" orientation="latitude" precision="1" value="-16.81745">-16.81745</geoCoordinate>
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,
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<geoCoordinate id="43B0DE76832FC9294F1C72D91E53EB36" degrees="49.2925" direction="east" orientation="longitude" precision="5" value="49.2925">49.2925</geoCoordinate>
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; alt. 400 m; 26 Feb 2010; B. L. Fisher et al. leg.; rainforest, ex rotten log; BLF24958 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection id="8F4C64C3C760D724BD93CD802422154E" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="4A838D38D897C12D2697AC3D44075D42" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="AF6E76448992071791D7E926E42BCA09" pageId="0" pageNumber="1">
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Minute species.
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<emphasis id="5F6FBBC7A14B6A02635B3375179551E8" bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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HL <0.85 mm and WL <0.65 mm; head in full-face view elongate, not widening posteriorly with posterior sides slightly convex, in lateral view sub-oval and elongate with slightly convex dorsal and ventral sides; body yellow; sides of head with moderately dense, short, suberect to erect pilosity; entire head distinctly sculptured but with smooth interspaces; scape, when laid back, reaching the midlength of head; inner hypostomal teeth distinct, moderate, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, approximately as high and wide as inner teeth; inner and outer hypostomal teeth moderately closely spaced and not connected by concavity; base of first gastral tergite smooth.
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<emphasis id="800666E92DA562090D13641257DC2B2E" bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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HL <0.45 mm and WL <0.5 mm, scape, when laid back, reaching the posterior head margin, propodeal spines small and triangular; head relatively rectangular; body yellow; head predominantly smooth; mesosoma smooth.
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</paragraph>
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</subSubSection>
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<subSubSection id="BA492942925A1B259E45C063610F1A06" pageId="0" pageNumber="1" type="description">
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<paragraph id="94D4FEED91EB9F5B2AD9A9DCFC62E362" pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph id="DB3A551C9EC35CAE9DC590EC72719CDF" pageId="0" pageNumber="1">
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<emphasis id="EDB16986229E54D664EDF036730E0767" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis id="E6603E060AA8800C506AFADAA43C64EC" italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 5): HL: 0.76-0.83 (0.8); HW: 0.68-0.73 (0.71); SL: 0.39-0.41 (0.4); EL: 0.08-0.08 (0.08); WL: 0.61-0.66 (0.64); PSL: 0.1-0.13 (0.12); MTL: 0.33-0.36 (0.35); PNW: 0.36-0.4 (0.37); PTW: 0.09-0.1 (0.1); PPW: 0.17-0.19 (0.18); CI: 111.3-115.3 (113.0); SI: 53.7-58.7 (56.2); PSLI: 12.8-15.5 (14.5); PPI: 47.4-60.8 (54.6); PNI: 51.4-53.9 (52.6); MTI: 48.2-49.6 (49.0).
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</paragraph>
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<paragraph id="508A30B2A7EA7424534827DE6CA562CC" pageId="0" pageNumber="1">
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<emphasis id="B6705EBDEBCF53D08222E4145D85551A" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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In full-face view elongate, not widening posteriorly, with anterior and posterior sides slightly convex (Fig.
|
||
<figureCitation id="58161874EC4C2A36077831D9532E6FBF" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Pheidole flavominuta sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0162186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923272)." figureDoi="10.3897/zookeys.949.51269.figure29" httpUri="https://binary.pensoft.net/fig/432198" pageId="0" pageNumber="1">29B</figureCitation>
|
||
). In lateral view sub-oval and elongate; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with moderately dense, short, suberect to erect pilosity; whole head with dense, long, decumbent to erect pilosity. Medial part of frons with thick, longitudinal, and dense rugae and smooth to indistinctly rugulate interspaces, rugae directed slightly outward on posteromedial part; lateral sides with thick, dense, and irregular rugae, interspaces shiny and distinctly rugoreticulate. Occipital lobes with dense, irregular rugae and distinctly rugoreticulate interspaces. Area posterolateral from eyes with weaker sculpture, rugoreticulate. Gena with relatively sparse and thick longitudinal rugae and smooth interspaces. Centre of clypeus smooth and shiny, lateral sides with indistinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, reaching the midlength of head; pilosity subdecumbent to erect (Fig.
|
||
<figureCitation id="C95D204AC1D3B83AB50F3E951351913F" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Pheidole flavominuta sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0162186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923272)." figureDoi="10.3897/zookeys.949.51269.figure29" httpUri="https://binary.pensoft.net/fig/432198" pageId="0" pageNumber="1">29B, D</figureCitation>
|
||
). Inner hypostomal teeth distinct, moderate, closely spaced, triangular, with rounded apex directed upward; outer hypostomal teeth lobe-like, approximately as high and wide as inner teeth; inner and outer hypostomal teeth moderately closely spaced and not connected by concavity (Fig.
|
||
<figureCitation id="EFEA0867FA99D823D86C2EF258CDC7F7" captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">63K</figureCitation>
|
||
).
|
||
<emphasis id="93C547763B374AD4084BE8B6476FE6B7" bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
|
||
In lateral view, promesonotum short, angular, and moderately low, posterior mesonotum steep, mesonotal process very indistinct, bulge-like; promesonotal groove absent; metanotal groove absent; propodeal spines moderately long, with wide base and acute apex; humeral area laterally weakly produced (Fig.
|
||
<figureCitation id="6970B9CC06BADD071239A31856B0B1CB" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Pheidole flavominuta sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0162186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923272)." figureDoi="10.3897/zookeys.949.51269.figure29" httpUri="https://binary.pensoft.net/fig/432198" pageId="0" pageNumber="1">29D</figureCitation>
|
||
). Surface shiny; promesonotum predominantly smooth with very fine network of rugoreticulae; anepisternum, katepisternum, and propodeum with rugoreticulae denser and thicker. Pilosity moderately dense, moderately long, and erect (Fig.
|
||
<figureCitation id="05A47201CD71908609948045A7A47BA3" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Pheidole flavominuta sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0162186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923272)." figureDoi="10.3897/zookeys.949.51269.figure29" httpUri="https://binary.pensoft.net/fig/432198" pageId="0" pageNumber="1">29D, F</figureCitation>
|
||
).
|
||
<emphasis id="3078844BB565AB3F82C64ECEA7422B1C" bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
|
||
Shiny with dense foveolae; node smooth to finely foveolate, triangular, with rounded and thick apex, in rear view node dorsoventrally convex; pilosity moderately sparse and erect (Fig.
|
||
<figureCitation id="4E579394104C62459D529B0752A23CBB" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Pheidole flavominuta sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0162186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923272)." figureDoi="10.3897/zookeys.949.51269.figure29" httpUri="https://binary.pensoft.net/fig/432198" pageId="0" pageNumber="1">29D, F</figureCitation>
|
||
).
|
||
<emphasis id="3A0E49542ABCBF2B381F665CDFBE7632" bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
|
||
Shiny and foveolate; dorsum with reduced sculpture and smooth notch; in dorsal view oval, lateral margins medially with two dentate projections; pilosity long, moderately sparse, and erect (Fig.
|
||
<figureCitation id="57E6230E27E5250FCF8F2B01AFC9F928" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Pheidole flavominuta sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0162186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923272)." figureDoi="10.3897/zookeys.949.51269.figure29" httpUri="https://binary.pensoft.net/fig/432198" pageId="0" pageNumber="1">29D, F</figureCitation>
|
||
).
|
||
<emphasis id="489784A0CB1F04ED2F0A5E872C0BD318" bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
||
Shiny and smooth; pilosity moderately dense, moderately short, and erect (Fig.
|
||
<figureCitation id="7AF1705ECB77EEB2FF3599C688989D77" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Pheidole flavominuta sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0162186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923272)." figureDoi="10.3897/zookeys.949.51269.figure29" httpUri="https://binary.pensoft.net/fig/432198" pageId="0" pageNumber="1">29D, F</figureCitation>
|
||
).
|
||
<emphasis id="83E910B459B0865469434E280B1B7410" bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
||
Yellow (Fig.
|
||
<figureCitation id="FAA2405D37173AA57F96384C1D76A571" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Pheidole flavominuta sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0162186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923272)." figureDoi="10.3897/zookeys.949.51269.figure29" httpUri="https://binary.pensoft.net/fig/432198" pageId="0" pageNumber="1">29D, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F308CB9EF796DE2E43FE42EE82315982" pageId="0" pageNumber="1">
|
||
<emphasis id="9A353CE34A1AA57AA8460DF195AEC299" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
Measurements (
|
||
<emphasis id="19FBC4AA5E648CFA1897544113A98259" italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 5): HL: 0.42-0.44 (0.43); HW: 0.37-0.39 (0.38); SL: 0.34-0.36 (0.35); EL: 0.06-0.07 (0.06); WL: 0.46-0.49 (0.47); PSL: 0.05-0.07 (0.06); MTL: 0.25-0.28 (0.26); PNW: 0.23-0.26 (0.25); PTW: 0.05-0.07 (0.06); PPW: 0.09-0.11 (0.1); CI: 108.4-117.2 (114.8); SI: 88.2-95.7 (93.2); PSLI: 12.2-16.9 (14.6); PPI: 47.7-70.7 (63.8); PNI: 59.6-69.4 (66.4); MTI: 67.7-70.3 (69.0).
|
||
</paragraph>
|
||
<paragraph id="AAD8AE85D97BD8FEC15718A53C79B7D1" pageId="0" pageNumber="1">
|
||
<emphasis id="586093BE8230C3303049709EABDF64F5" bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
|
||
Cephalic margin indistinctly concave to straight (Fig.
|
||
<figureCitation id="CA7133E87F771AB3E1316B2855C9EEFC" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Pheidole flavominuta sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0162186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923272)." figureDoi="10.3897/zookeys.949.51269.figure29" httpUri="https://binary.pensoft.net/fig/432198" pageId="0" pageNumber="1">29A</figureCitation>
|
||
). Pilosity relatively dense, moderately long, decumbent to suberect. Sculpture shiny and smooth; lateral sides of frons with fine, sparse, longitudinal, and interrupted rugulae; vertex with indistinct, sparse, and transverse rugulae; antennal sockets with few thick, curved outward rugae and smooth interspaces. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, reaching the posterior head margin; pilosity dense, subdecumbent to erect (Fig.
|
||
<figureCitation id="4A5EDF80131D25F55EEAFE385BEFE309" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Pheidole flavominuta sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0162186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923272)." figureDoi="10.3897/zookeys.949.51269.figure29" httpUri="https://binary.pensoft.net/fig/432198" pageId="0" pageNumber="1">29A, C</figureCitation>
|
||
).
|
||
<emphasis id="90DA961DF9038A238E8C968F6AADC015" bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
|
||
In lateral view, promesonotum moderately high and short, arched; promesonotal groove absent; metanotal groove distinct; propodeal spines small and triangular (Fig.
|
||
<figureCitation id="20EB1A9D1AB31F4FCC37FDED6B1F50A3" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Pheidole flavominuta sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0162186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923272)." figureDoi="10.3897/zookeys.949.51269.figure29" httpUri="https://binary.pensoft.net/fig/432198" pageId="0" pageNumber="1">29C</figureCitation>
|
||
). Sculpture shiny and smooth. Pilosity moderately sparse, moderately long, and erect (Fig.
|
||
<figureCitation id="545B9C908359F0D884EAE592ACB84273" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Pheidole flavominuta sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0162186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923272)." figureDoi="10.3897/zookeys.949.51269.figure29" httpUri="https://binary.pensoft.net/fig/432198" pageId="0" pageNumber="1">29C, E</figureCitation>
|
||
).
|
||
<emphasis id="4E746D2F288C8AAAB21FD9C0649084B8" bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
||
With sparse, erect pilosity (Fig.
|
||
<figureCitation id="C0432129330FA0A6208A6DF20C5C203E" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Pheidole flavominuta sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0162186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923272)." figureDoi="10.3897/zookeys.949.51269.figure29" httpUri="https://binary.pensoft.net/fig/432198" pageId="0" pageNumber="1">29C, E</figureCitation>
|
||
).
|
||
<emphasis id="A1F9657B000964433FD4E12C5F34B508" bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
||
Yellow, head and gaster slightly darker (Fig.
|
||
<figureCitation id="85653849BFDD67972085C52EDD05831C" captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Pheidole flavominuta sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0162186) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923272)." figureDoi="10.3897/zookeys.949.51269.figure29" httpUri="https://binary.pensoft.net/fig/432198" pageId="0" pageNumber="1">29C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="98508FA7C557FC4779FAC3FCA4E6C5BC" pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph id="CE2B361E141A164F19E35EA938FF862C" pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph id="EC86F4BE8138429C76A018331C9900E4" pageId="0" pageNumber="1">Latin for bright and small in reference to small body size and bright colouration.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="33677E58C6D8D001157BF21DE3FFC1C7" pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph id="C469D7491F81C162CDE835298D47E696" pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph id="A269210E2F1E75B851F3910AEB9A4374" pageId="0" pageNumber="1">The species was collected between 355-520 m in elevation, in rainforest. Nests were located in rotten logs, and under rootmats on the ground.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="338B9D34AE5BD4237FBBC63210AEB041" pageId="0" pageNumber="1" type="comments">
|
||
<paragraph id="E597D48D8D11A56FDA1B7A47A46014D2" pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph id="D2E6D0AF48FBC7B668EBACEE669EC157" pageId="0" pageNumber="1">
|
||
<taxonomicName id="7BD027038CD2214C39C0AADE829B0FDB" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole flavominuta" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="flavominuta">
|
||
<emphasis id="0C763F2905DE14082B687FEEAC223F74" italics="true" pageId="0" pageNumber="1">Pheidole flavominuta</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. belongs to the group of species characterised by small body size (major workers: HL <1.05 mm, WL <0.9 mm and minor workers HL <0.5 mm, WL <0.6 mm), head in major workers elongate and not widening posteriorly, minor workers with predominantly smooth and relatively rectangular head, and yellow to brown body colouration. The group includes four species:
|
||
<taxonomicName id="86F345CA260F160C0B536FCE870702ED" lsidName="P. flavominuta" pageId="0" pageNumber="1" rank="species" species="flavominuta">
|
||
<emphasis id="1D207439A3E94CC2D7073BC88C024188" italics="true" pageId="0" pageNumber="1">P. flavominuta</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="EFB7D05D9427730C35BD8BC1F82C8B7E" lsidName="P. nitidobruna" pageId="0" pageNumber="1" rank="species" species="nitidobruna">
|
||
<emphasis id="90058889AE2CC98A74E1B71A12A1E29D" italics="true" pageId="0" pageNumber="1">P. nitidobruna</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="510A289615AAEA162A150E14B4C9036A" lsidName="P. mikros" pageId="0" pageNumber="1" rank="species" species="mikros">
|
||
<emphasis id="5C51177F567D6352AF30C4B1981916F9" italics="true" pageId="0" pageNumber="1">P. mikros</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., and
|
||
<taxonomicName id="160398367274B77A175DAF3D4015A26E" lsidName="P. beanka" pageId="0" pageNumber="1" rank="species" species="beanka">
|
||
<emphasis id="CE83D3398034E0806E3ABADCE0E6650C" italics="true" pageId="0" pageNumber="1">P. beanka</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Distribution of
|
||
<taxonomicName id="72E622BC317B387C31A7A64842AFD130" lsidName="P. flavominuta" pageId="0" pageNumber="1" rank="species" species="flavominuta">
|
||
<emphasis id="AB2C9D2495B7101B25343BA87A9A4931" italics="true" pageId="0" pageNumber="1">P. flavominuta</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. covers area north from Toamasina up to Antenina and
|
||
<normalizedToken id="BFFA23BADF43C7AFF4074A93690E5211" originalValue="doesn’t">doesn't</normalizedToken>
|
||
overlap with other members of this group.
|
||
<taxonomicName id="07244C401FA7FEBE3233C34F6C94C94B" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole flavominuta" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="flavominuta">
|
||
<emphasis id="DAB6F3958F0FC1F63820A218598B4680" italics="true" pageId="0" pageNumber="1">Pheidole flavominuta</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. strongly differs from other members of the group by having distinctly yellow body colouration in major and minor workers. Additionally, major workers have head, in lateral view, sub-oval and elongate with slightly convex dorsal and ventral sides, and first gastral tergite is entirely smooth. Minor workers of this species can be also separated by a short antennal scape which
|
||
<normalizedToken id="126B18205BB58CE096C2A89053D3922C" originalValue="doesn’t">doesn't</normalizedToken>
|
||
exceed the posterior head margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |