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<document id="4E0AA5B8144E27211F3CC514CDE125C0" ID-CLB-Dataset="21863" ID-DOI="10.5852/ejt.2016.241" ID-GBIF-Dataset="f694842b-3af3-4198-b01e-13d0b5466b5d" ID-ISSN="2118-9773" ID-Zenodo-Dep="3850299" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590443252253" checkinUser="valdenar" docAuthor="Frolov, Andrey V. &amp; Akhmetova, Lilia A." docDate="2016" docId="C4027A0914035552D358FEB2FA94B237" docLanguage="en" docName="ejt-2016-241.pdf.imf" docOrigin="European Journal of Taxonomy 241" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Orphnus (Phornus) renaudi Frolov &amp; Akhmetova 2016, sp. nov." docType="treatment" docUuid="0B5B7277-272A-43A7-8F95-2D8E5AE363F2" docUuidSource="ZooBank" docVersion="9" lastPageNumber="9" masterDocId="383B02711405555AD167FFB9FFF7B60D" masterDocTitle="Revision of the subgenus Orphnus (Phornus) (Coleoptera, Scarabaeidae, Orphninae)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="7" updateTime="1698841278433" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0A94382F70ECD04B0DEC41D483C5CCAC">Revision of the subgenus Orphnus (Phornus) (Coleoptera, Scarabaeidae, Orphninae)</mods:title>
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<mods:name id="629255A0E8E1D9D79EB19147A3055949" type="personal">
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<mods:namePart id="55BB80918B42201225C3569AD1265208">Frolov, Andrey V.</mods:namePart>
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<mods:affiliation id="8717203E2F7603B59480A4FFE5A3EBF6">Laboratory of Insect Systematics, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab., 1, St. - Petersburg, 199034 Russia. Federal University of Mato Grosso, Av. Fernando Corrêa da Costa, 2367, Boa Esperança, 78060 - 900 Cuiabá, MT, Brazil.</mods:affiliation>
<mods:nameIdentifier id="A3E2B32D852ACFAF6BBB55148E1DA816" type="email">frolov@scarabaeoidea.com</mods:nameIdentifier>
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<mods:namePart id="323140E7C9CDD8D34514844F2DED1B30">Akhmetova, Lilia A.</mods:namePart>
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<mods:affiliation id="5785AF74DAE3B688970B9D0A0D7F8D76">Laboratory of Insect Systematics, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab., 1, St. - Petersburg, 199034 Russia. Federal University of Mato Grosso, Av. Fernando Corrêa da Costa, 2367, Boa Esperança, 78060 - 900 Cuiabá, MT, Brazil.</mods:affiliation>
<mods:nameIdentifier id="A5918FB3D61C810E14F19D1AA9A681B0" type="email">akhmetova@scarabaeoidea.com</mods:nameIdentifier>
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<mods:title id="1DBD52FDA2887E9372908C15F9D41669">European Journal of Taxonomy</mods:title>
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<mods:date id="BBDBA50B00AA6D8FC7DF90935AB7B0BF">2016</mods:date>
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<treatment id="C4027A0914035552D358FEB2FA94B237" ID-DOI="http://doi.org/10.5281/zenodo.3854782" ID-GBIF-Taxon="164283140" ID-Zenodo-Dep="3854782" LSID="urn:lsid:zoobank.org:act:0B5B7277-272A-43A7-8F95-2D8E5AE363F2" httpUri="http://treatment.plazi.org/id/C4027A0914035552D358FEB2FA94B237" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection id="04B198941403555CD358FEB2FC02B72B" box="[575,1013,267,294]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="4C14CB1F1403555CD358FEB2FC02B72B" blockId="6.[370,1217,267,364]" box="[575,1013,267,294]" pageId="6" pageNumber="7">
<heading id="175C7C731403555CD358FEB2FC02B72B" box="[575,1013,267,294]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
<taxonomicName id="8BABB09C1403555CD358FEB2FC63B72B" authority="Frolov &amp; Akhmetova, 2016" authorityName="Frolov &amp; Akhmetova" authorityYear="2016" box="[575,916,267,294]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="renaudi" status="sp. nov." subGenus="Phornus">
<emphasis id="7EDF170D1403555CD358FEB2FC63B72B" bold="true" box="[575,916,267,294]" pageId="6" pageNumber="7">
<emphasis id="7EDF170D1403555CD358FEB2FD5CB72B" bold="true" box="[575,683,267,294]" italics="true" pageId="6" pageNumber="7">Orphnus</emphasis>
(
<emphasis id="7EDF170D1403555CD3DCFEB2FCD4B72B" bold="true" box="[699,803,267,294]" italics="true" pageId="6" pageNumber="7">Phornus</emphasis>
)
<emphasis id="7EDF170D1403555CD252FEB2FC63B72B" bold="true" box="[821,916,267,294]" italics="true" pageId="6" pageNumber="7">renaudi</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65ECAA761403555CD2FCFEB5FC02B72B" box="[923,1013,268,294]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="04B198941403555CD015FE97FC91B761" pageId="6" pageNumber="7" type="description">
<paragraph id="4C14CB1F1403555CD015FE97FB36B744" blockId="6.[370,1217,267,364]" box="[370,1217,302,329]" pageId="6" pageNumber="7">
<uri id="383AC71D1403555CD015FE97FB36B744" box="[370,1217,302,329]" pageId="6" pageNumber="7">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="4C14CB1F1403555CD3AAFEEBFC91B761" blockId="6.[370,1217,267,364]" box="[717,870,338,364]" pageId="6" pageNumber="7">
<figureCitation id="D490D79A1403555CD3AAFEEBFCDAB761" box="[717,813,338,364]" captionStart="Fig" captionStartId="4.[189,232,1860,1886]" captionTargetBox="[228,1350,266,1814]" captionTargetId="figure@4.[227,1357,265,1825]" captionTargetPageId="4" captionText="Fig. 1. — AM. Orphnus giganteus Paulian, 1948. AB. Habitus. CD. Head in dorsal and apical view. EG. Antennae. H. Aedeagus in lateral view. I. Parameres in dorsal view. J. Abdomen in dorsal view. KL. Stridulatory field. M. Habitus. — N. O. renaudi sp. nov., holotype, habitus. — O. O. ferrierei sp. nov., holotype, habitus. — P. O. valeriae sp. nov., holotype, habitus. — Q. O. parastrangulatus sp. nov., holotype, habitus. — R. O. strangulatus Paulian, 1948, habitus. MR = not to scale." figureDoi="http://doi.org/10.5281/zenodo.3850303" httpUri="https://zenodo.org/record/3850303/files/figure.png" pageId="6" pageNumber="7">Figs 1N</figureCitation>
,
<figureCitation id="D490D79A1403555CD25DFEEBFCBCB761" box="[826,843,338,364]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[203,1399,636,1876]" captionTargetId="figure@7.[189,1400,635,1888]" captionTargetPageId="7" captionText="Fig. 2. Orphnus renaudi sp. nov. AB. Habitus. CD. Head in dorsal and apical view. E. Aedeagus in lateral view. F. Parameres in dorsal view. G. Labels of holotype. H. Abdomen in dorsal view. I. Stridulatory field. A, CD, G. Holotype. B, EF, HI. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850307" httpUri="https://zenodo.org/record/3850307/files/figure.png" pageId="6" pageNumber="7">2</figureCitation>
,
<figureCitation id="D490D79A1403555CD230FEEBFC91B761" box="[855,870,338,364]" captionStart="Fig" captionStartId="16.[189,232,1966,1992]" captionTargetBox="[194,1393,1011,1923]" captionTargetId="figure@16.[189,1398,1009,1925]" captionTargetPageId="16" captionText="Fig. 6. Orphnus (Phornus) spp. Locality map. Orphnus parastrangulatus sp. nov. is not mapped, because there are no more or less precise locality data available for this species." figureDoi="http://doi.org/10.5281/zenodo.3850323" httpUri="https://zenodo.org/record/3850323/files/figure.png" pageId="6" pageNumber="7">6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="04B198941403555CD1DAFE2AFC51B42E" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="4C14CB1F1403555CD1DAFE2AFEC1B7A0" blockId="6.[189,310,403,429]" box="[189,310,403,429]" pageId="6" pageNumber="7">
<heading id="175C7C731403555CD1DAFE2AFEC1B7A0" bold="true" box="[189,310,403,429]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="7EDF170D1403555CD1DAFE2AFEC1B7A0" bold="true" box="[189,310,403,429]" pageId="6" pageNumber="7">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="4C14CB1F1403555CD1DAFE7BFC51B42E" blockId="6.[189,1399,450,547]" pageId="6" pageNumber="7">
<taxonomicName id="8BABB09C1403555CD1DAFE7BFE73B7D1" authority="Frolov &amp; Akhmetova, 2016" authorityName="Frolov &amp; Akhmetova" authorityYear="2016" box="[189,388,450,476]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="renaudi" status="sp. nov.">
<emphasis id="7EDF170D1403555CD1DAFE7BFE73B7D1" box="[189,388,450,476]" italics="true" pageId="6" pageNumber="7">Orphnus renaudi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65ECAA761403555CD0EBFE7BFE11B7D1" box="[396,486,450,476]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
differs from the other macropterous
<taxonomicName id="8BABB09C1403555CD2F4FE7BFC01B7D1" authorityName="Paulian" authorityYear="1948" box="[915,1014,450,476]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="18" phylum="Arthropoda" rank="subGenus" subGenus="Phornus">
<emphasis id="7EDF170D1403555CD2F4FE7BFC01B7D1" box="[915,1014,450,476]" italics="true" pageId="6" pageNumber="7">Phornus</emphasis>
</taxonomicName>
species (
<taxonomicName id="8BABB09C1403555CD504FE7BFB0FB7D1" authorityName="Paulian" authorityYear="1948" box="[1123,1272,450,476]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="giganteus">
<emphasis id="7EDF170D1403555CD504FE7BFB8FB7D1" box="[1123,1144,450,476]" italics="true" pageId="6" pageNumber="7">O</emphasis>
.
<emphasis id="7EDF170D1403555CD5E1FE7AFB0FB7D1" box="[1158,1272,451,476]" italics="true" pageId="6" pageNumber="7">giganteus</emphasis>
</taxonomicName>
) in having the pronotum relatively smaller and narrower than elytra, sparsely punctate disc of pronotum, narrower head with concave and sparsely punctate frons, and larger eyes.
</paragraph>
</subSubSection>
<subSubSection id="04B198941403555CD1DAFDF3FD34B49F" box="[189,707,586,658]" pageId="6" pageNumber="7" type="etymology">
<paragraph id="4C14CB1F1403555CD1DAFDF3FEB3B469" blockId="6.[189,324,586,612]" box="[189,324,586,612]" pageId="6" pageNumber="7">
<heading id="175C7C731403555CD1DAFDF3FEB3B469" bold="true" box="[189,324,586,612]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="7EDF170D1403555CD1DAFDF3FEB3B469" bold="true" box="[189,324,586,612]" pageId="6" pageNumber="7">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="4C14CB1F1403555CD1DAFDC1FD34B49F" blockId="6.[189,707,632,658]" box="[189,707,632,658]" pageId="6" pageNumber="7">This species is named after Renaud Paulian.</paragraph>
</subSubSection>
<subSubSection id="04B198941403555CD1DAFD00FD87B23D" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="4C14CB1F1403555CD1DAFD00FE5DB4DE" blockId="6.[189,426,697,723]" box="[189,426,697,723]" pageId="6" pageNumber="7">
<heading id="175C7C731403555CD1DAFD00FE5DB4DE" bold="true" box="[189,426,697,723]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="7EDF170D1403555CD1DAFD00FE5DB4DE" bold="true" box="[189,426,697,723]" pageId="6" pageNumber="7">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="FCC3C1421403555CD18DFD51FC9DB528" ID-GBIF-Occurrence="3761958365" collectionCode="MNHN" country="Nigeria" location="NIGERIA" pageId="6" pageNumber="7" specimenCount="1" typeStatus="holotype">
<paragraph id="4C14CB1F1403555CD18DFD51FEABB50F" blockId="6.[189,874,744,806]" box="[234,348,744,770]" pageId="6" pageNumber="7">
<emphasis id="7EDF170D1403555CD18DFD51FEABB50F" bold="true" box="[234,348,744,770]" pageId="6" pageNumber="7">
<typeStatus id="931075BD1403555CD18DFD51FEABB50F" box="[234,348,744,770]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="4C14CB1F1403555CD1DAFCB2FC9DB528" blockId="6.[189,874,744,806]" box="[189,874,779,806]" pageId="6" pageNumber="7">
<collectingCountry id="34BC8B8F1403555CD1DAFCB2FECDB528" box="[189,314,779,805]" name="Nigeria" pageId="6" pageNumber="7">NIGERIA</collectingCountry>
:
<specimenCount id="5AAD00961403555CD022FCB5FEABB52B" box="[325,348,780,806]" pageId="6" pageNumber="7"></specimenCount>
, “Old Calabar [Calabar, Nigeria]” (
<collectionCode id="2ABA53DA1403555CD39AFCB2FCA9B528" box="[765,862,779,805]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="7">MNHN</collectionCode>
).
</paragraph>
</materialsCitation>
<materialsCitation id="FCC3C1421403555CD18DFCF4FD74B5A0" ID-GBIF-Occurrence="3761958358" collectionCode="MNHN" country="Nigeria" location="Orphnus renaudi Frolov &amp; Akhmetova" pageId="6" pageNumber="7" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<paragraph id="4C14CB1F1403555CD18DFCF4FE90B56A" blockId="6.[189,1398,845,942]" box="[234,359,845,871]" pageId="6" pageNumber="7">
<emphasis id="7EDF170D1403555CD18DFCF4FE90B56A" bold="true" box="[234,359,845,871]" pageId="6" pageNumber="7">
<typeStatus id="931075BD1403555CD18DFCF4FE90B56A" box="[234,359,845,871]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="4C14CB1F1403555CD1DAFCC9FD74B5A0" blockId="6.[189,1398,845,942]" pageId="6" pageNumber="7">
<collectingCountry id="34BC8B8F1403555CD1DAFCC9FECDB587" box="[189,314,880,906]" name="Nigeria" pageId="6" pageNumber="7">NIGERIA</collectingCountry>
:
<specimenCount id="5AAD00961403555CD02EFCC9FE78B587" box="[329,399,880,906]" pageId="6" pageNumber="7" type="male">2 ♂♂</specimenCount>
with the same locality label as the
<typeStatus id="931075BD1403555CD22DFCC9FC46B587" box="[842,945,880,906]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
and “
<typeStatus id="931075BD1403555CD29DFCC9FB54B587" box="[1018,1187,880,906]" pageId="6" pageNumber="7" type="paratype">PARATYPUS</typeStatus>
<taxonomicName id="8BABB09C1403555CD5C9FCC9FE0BB5A3" authority="Frolov &amp; Akhmetova 2015" authorityName="Frolov &amp; Akhmetova" authorityYear="2015" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="renaudi">
<location id="49749DC41403555CD5C9FCC9FE4FB5A0" LSID="urn:lsid:plazi:treatment:C4027A0914035552D358FEB2FA94B237:49749DC41403555CD5C9FCC9FE4FB5A0" country="Nigeria" name="Orphnus renaudi Frolov &amp; Akhmetova" pageId="6" pageNumber="7">
Orphnus renaudi
<bibRefCitation id="283AB6EE1403555CD1DAFC2AFE4FB5A0" author="Frolov A. V. &amp; Akhmetova L. A." box="[189,440,915,941]" pageId="6" pageNumber="7" pagination="475 - 477" refId="ref8676" refString="Frolov A. V. &amp; Akhmetova L. A. 2015. A new brachypterous scarab species, Orphnus longicornis (Coleoptera: Scarabaeidae: Orphninae), from the East African Rift. Zootaxa 4039: 475 - 477. http: // dx. doi. org / 10.11646 / zootaxa. 4039.3.7" type="journal article" year="2015">Frolov &amp; Akhmetova</bibRefCitation>
</location>
2015
</taxonomicName>
” (
<collectionCode id="2ABA53DA1403555CD371FC2AFD80B5A0" box="[534,631,915,941]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="7">MNHN</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="4C14CB1F1403555CD1DAFC6CFB59B5E2" blockId="6.[189,1198,980,1007]" box="[189,1198,980,1007]" pageId="6" pageNumber="7">
<materialsCitation id="FCC3C1421403555CD1DAFC6CFB59B5E2" ID-GBIF-Occurrence="3761958314" box="[189,1198,980,1007]" collectionCode="MNHN" country="Cameroon" location="Mount Cameroon" municipality="Camaroons" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1">
<collectingCountry id="34BC8B8F1403555CD1DAFC6CFE9DB5E2" box="[189,362,981,1007]" name="Cameroon" pageId="6" pageNumber="7">CAMEROON</collectingCountry>
:
<specimenCount id="5AAD00961403555CD013FC6CFE56B5E2" box="[372,417,981,1007]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
, “
<collectingMunicipality id="AC7051651403555CD0DEFC6CFDB5B5E2" box="[441,578,981,1007]" pageId="6" pageNumber="7">Camaroons</collectingMunicipality>
[probably foothills of
<location id="49749DC41403555CD22CFC6CFBD6B5E2" LSID="urn:lsid:plazi:treatment:C4027A0914035552D358FEB2FA94B237:49749DC41403555CD22CFC6CFBD6B5E2" box="[843,1057,980,1007]" country="Cameroon" municipality="Camaroons" name="Mount Cameroon" pageId="6" pageNumber="7">Mount Cameroon</location>
]” (
<collectionCode id="2ABA53DA1403555CD526FC6CFB55B5E2" box="[1089,1186,981,1007]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="7">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="4C14CB1F1403555CD1DAFBAFFD87B23D" blockId="6.[189,624,1046,1073]" box="[189,624,1046,1073]" pageId="6" pageNumber="7">
<materialsCitation id="FCC3C1421403555CD1DAFBAFFD9CB23D" ID-GBIF-Occurrence="3761958318" box="[189,619,1046,1073]" collectionCode="BMNH" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1">
<specimenCount id="5AAD00961403555CD1DAFBAFFF1EB23C" box="[189,233,1046,1073]" pageId="6" pageNumber="7" type="male">1 ♂</specimenCount>
without locality labels (
<collectionCode id="2ABA53DA1403555CD362FBAFFD93B23D" box="[517,612,1046,1072]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="6" pageNumber="7">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="04B1989414035552D1DAFBEEFD8CB44D" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
<paragraph id="4C14CB1F1403555CD1DAFBEEFEB9B27C" blockId="6.[189,334,1111,1137]" box="[189,334,1111,1137]" pageId="6" pageNumber="7">
<heading id="175C7C731403555CD1DAFBEEFEB9B27C" bold="true" box="[189,334,1111,1137]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="7EDF170D1403555CD1DAFBEEFEB9B27C" bold="true" box="[189,334,1111,1137]" pageId="6" pageNumber="7">Description</emphasis>
</heading>
</paragraph>
<paragraph id="4C14CB1F1403555CD18DFB3FFE26B2AD" blockId="6.[189,793,1158,1220]" box="[234,465,1158,1184]" pageId="6" pageNumber="7">
<emphasis id="7EDF170D1403555CD18DFB3FFEABB2AD" bold="true" box="[234,348,1158,1184]" pageId="6" pageNumber="7">
<typeStatus id="931075BD1403555CD18DFB3FFEABB2AD" box="[234,348,1158,1184]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="D490D79A1403555CD00BFB3FFE3EB2AD" box="[364,457,1158,1184]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[203,1399,636,1876]" captionTargetId="figure@7.[189,1400,635,1888]" captionTargetPageId="7" captionText="Fig. 2. Orphnus renaudi sp. nov. AB. Habitus. CD. Head in dorsal and apical view. E. Aedeagus in lateral view. F. Parameres in dorsal view. G. Labels of holotype. H. Abdomen in dorsal view. I. Stridulatory field. A, CD, G. Holotype. B, EF, HI. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850307" httpUri="https://zenodo.org/record/3850307/files/figure.png" pageId="6" pageNumber="7">Fig. 2A</figureCitation>
)
</paragraph>
<paragraph id="4C14CB1F1403555CD1DAFB10FCEEB2CE" blockId="6.[189,793,1158,1220]" box="[189,793,1193,1220]" pageId="6" pageNumber="7">
Body length
<quantity id="8B5366FA1403555CD032FB10FE5DB2C9" box="[341,426,1193,1220]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="7" unit="mm" value="13.0">13 mm</quantity>
. Colour uniformly dark brown.
</paragraph>
<paragraph id="4C14CB1F1403555CD1DAFB52FE36B3B8" blockId="6.[189,1399,1259,1461]" pageId="6" pageNumber="7">
Anterior margin of frontoclypeus slightly convex in middle, slightly sinuate beside medial convexity, rounded laterally, with a narrow border (
<figureCitation id="D490D79A1403555CD3F0FAB7FD04B325" box="[663,755,1294,1320]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[203,1399,636,1876]" captionTargetId="figure@7.[189,1400,635,1888]" captionTargetPageId="7" captionText="Fig. 2. Orphnus renaudi sp. nov. AB. Habitus. CD. Head in dorsal and apical view. E. Aedeagus in lateral view. F. Parameres in dorsal view. G. Labels of holotype. H. Abdomen in dorsal view. I. Stridulatory field. A, CD, G. Holotype. B, EF, HI. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850307" httpUri="https://zenodo.org/record/3850307/files/figure.png" pageId="6" pageNumber="7">Fig. 2C</figureCitation>
). Frontoclypeus with a low transverse bimodal process medially approximately in middle of a line connecting anterior margins of eyes and anterior margin of frontoclypeus (
<figureCitation id="D490D79A1403555CD0FEFAEDFE0AB363" box="[409,509,1364,1390]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[203,1399,636,1876]" captionTargetId="figure@7.[189,1400,635,1888]" captionTargetPageId="7" captionText="Fig. 2. Orphnus renaudi sp. nov. AB. Habitus. CD. Head in dorsal and apical view. E. Aedeagus in lateral view. F. Parameres in dorsal view. G. Labels of holotype. H. Abdomen in dorsal view. I. Stridulatory field. A, CD, G. Holotype. B, EF, HI. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850307" httpUri="https://zenodo.org/record/3850307/files/figure.png" pageId="6" pageNumber="7">Fig. 2D</figureCitation>
). Frontoclypeus slightly concave behind process, punctate with rounded punctures separated by 13 puncture diameters in the anterior part and more sparsely behind the frontoclypeal process.
</paragraph>
<paragraph id="4C14CB1F1403555CD1DAFA64FDECB017" blockId="6.[189,1398,1500,1562]" pageId="6" pageNumber="7">Eyes relatively large: width about 1/5 distance between eyes in dorsal view. Antennae 10-segmented, without malformed segments.</paragraph>
<paragraph id="4C14CB1F1403555CD1DAF9F8FDE6B0EA" blockId="6.[189,1399,1601,1767]" pageId="6" pageNumber="7">
Pronotum widely rounded laterally, narrower than elytra (
<figureCitation id="D490D79A1403555CD200F9F8FC32B056" box="[871,965,1601,1627]" captionStart="Fig" captionStartId="4.[189,232,1860,1886]" captionTargetBox="[228,1350,266,1814]" captionTargetId="figure@4.[227,1357,265,1825]" captionTargetPageId="4" captionText="Fig. 1. — AM. Orphnus giganteus Paulian, 1948. AB. Habitus. CD. Head in dorsal and apical view. EG. Antennae. H. Aedeagus in lateral view. I. Parameres in dorsal view. J. Abdomen in dorsal view. KL. Stridulatory field. M. Habitus. — N. O. renaudi sp. nov., holotype, habitus. — O. O. ferrierei sp. nov., holotype, habitus. — P. O. valeriae sp. nov., holotype, habitus. — Q. O. parastrangulatus sp. nov., holotype, habitus. — R. O. strangulatus Paulian, 1948, habitus. MR = not to scale." figureDoi="http://doi.org/10.5281/zenodo.3850303" httpUri="https://zenodo.org/record/3850303/files/figure.png" pageId="6" pageNumber="7">Fig. 1N</figureCitation>
). Anterior border wide, with smooth posterior margin. Basal border narrow, keel-shaped, separated from pronotal disc by a deep groove having a row of coarse elongated punctures. Pronotal disc anteromedially slightly flattened, punctate. Sides with coarser punctures than disc, rounded to somewhat elongated. Disc almost smooth in basal half, with minute punctation.
</paragraph>
<paragraph id="4C14CB1F1403555CD1DAF8B6FBA4B124" blockId="6.[189,1107,1807,1833]" box="[189,1107,1807,1833]" pageId="6" pageNumber="7">Scutellum subtriangular, narrowly rounded apically, about 1/11 length of elytra.</paragraph>
<paragraph id="4C14CB1F1403555CD1DAF8E9FAC2B1BC" blockId="6.[189,1399,1872,1969]" pageId="6" pageNumber="7">Elytra 1.1 times longer than wide, with distinct humeral humps, widest in middle, with lateral margins almost parallel in basal half. First five striae feebly distinct as shallow densely punctured grooves. Laterad of fifth elytral stria, punctation is more uniform with relatively dense, rounded punctures.</paragraph>
<paragraph id="4C14CB1F1402555DD1DAFEB5FE9FB72B" blockId="7.[189,360,268,294]" box="[189,360,268,294]" pageId="7" pageNumber="8">Macropterous.</paragraph>
<paragraph id="4C14CB1F1402555DD1DAFEF4FB6AB76A" blockId="7.[189,1181,331,359]" box="[189,1181,331,359]" pageId="7" pageNumber="8">
Stridulatory field with carinae separated by 1/33 length of field in central ¼
<superScript id="BBDE66571402555DD579FEF2FBDBB754" attach="left" box="[1054,1068,331,345]" fontSize="6" pageId="7" pageNumber="8">rd</superScript>
(
<figureCitation id="D490D79A1402555DD55BFEF4FB78B76A" box="[1084,1167,333,359]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[203,1399,636,1876]" captionTargetId="figure@7.[189,1400,635,1888]" captionTargetPageId="7" captionText="Fig. 2. Orphnus renaudi sp. nov. AB. Habitus. CD. Head in dorsal and apical view. E. Aedeagus in lateral view. F. Parameres in dorsal view. G. Labels of holotype. H. Abdomen in dorsal view. I. Stridulatory field. A, CD, G. Holotype. B, EF, HI. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850307" httpUri="https://zenodo.org/record/3850307/files/figure.png" pageId="7" pageNumber="8">Fig. 2I</figureCitation>
).
</paragraph>
<paragraph id="4C14CB1F1402555DD1DAFE37FD71B41F" blockId="7.[189,1399,398,530]" pageId="7" pageNumber="8">
Abdominal sternite 8 medially longer than sternites 6 and 7 combined; sternite 6 slightly shorter than sternite 7. Pygidium almost not visible from above, with rounded apex. Plectrum trapezoidal, slightly longer than wide (
<figureCitation id="D490D79A1402555DD0E9FE6CFE1CB7E2" box="[398,491,469,495]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[203,1399,636,1876]" captionTargetId="figure@7.[189,1400,635,1888]" captionTargetPageId="7" captionText="Fig. 2. Orphnus renaudi sp. nov. AB. Habitus. CD. Head in dorsal and apical view. E. Aedeagus in lateral view. F. Parameres in dorsal view. G. Labels of holotype. H. Abdomen in dorsal view. I. Stridulatory field. A, CD, G. Holotype. B, EF, HI. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850307" httpUri="https://zenodo.org/record/3850307/files/figure.png" pageId="7" pageNumber="8">Fig. 2H</figureCitation>
). Lateral plate of second abdominal sternite well developed, sclerotized, about length of plectrum, with rounded apex.
</paragraph>
<caption id="18D49B971402555DD1DAF833FC3CB1E6" ID-DOI="http://doi.org/10.5281/zenodo.3850307" ID-Zenodo-Dep="3850307" httpUri="https://zenodo.org/record/3850307/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1930,1956]" targetBox="[203,1399,636,1876]" targetPageId="7">
<paragraph id="4C14CB1F1402555DD1DAF833FC3CB1E6" blockId="7.[189,1399,1930,2027]" pageId="7" pageNumber="8">
<emphasis id="7EDF170D1402555DD1DAF833FEF9B1A9" bold="true" box="[189,270,1930,1956]" pageId="7" pageNumber="8">Fig. 2.</emphasis>
<taxonomicName id="8BABB09C1402555DD07FF833FE14B1A9" authority="Frolov &amp; Akhmetova, 2016" authorityName="Frolov &amp; Akhmetova" authorityYear="2016" box="[280,483,1930,1956]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="renaudi" status="sp. nov.">
<emphasis id="7EDF170D1402555DD07FF833FE14B1A9" box="[280,483,1930,1956]" italics="true" pageId="7" pageNumber="8">Orphnus renaudi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65ECAA761402555DD08AF832FDBCB1A8" box="[493,587,1931,1957]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7EDF170D1402555DD332F833FD7BB1A9" bold="true" box="[597,652,1930,1956]" pageId="7" pageNumber="8">AB</emphasis>
. Habitus.
<emphasis id="7EDF170D1402555DD26CF833FCB3B1A9" bold="true" box="[779,836,1930,1956]" pageId="7" pageNumber="8">CD</emphasis>
. Head in dorsal and apical view.
<emphasis id="7EDF170D1402555DD5B8F832FB04B1A8" bold="true" box="[1247,1267,1931,1957]" pageId="7" pageNumber="8">E</emphasis>
. Aedeagus in lateral view.
<emphasis id="7EDF170D1402555DD0E6F817FE64B1C5" bold="true" box="[385,403,1966,1992]" pageId="7" pageNumber="8">F</emphasis>
. Parameres in dorsal view.
<emphasis id="7EDF170D1402555DD396F814FCFFB1CA" bold="true" box="[753,776,1965,1991]" pageId="7" pageNumber="8">G</emphasis>
. Labels of holotype.
<emphasis id="7EDF170D1402555DD570F817FBD9B1C5" bold="true" box="[1047,1070,1966,1992]" pageId="7" pageNumber="8">H</emphasis>
. Abdomen in dorsal view.
<emphasis id="7EDF170D1402555DD1DAF868FF3FB1E6" bold="true" box="[189,200,2001,2027]" pageId="7" pageNumber="8">I</emphasis>
. Stridulatory field. A, CD, G. Holotype. B, EF, HI. Paratypes.
</paragraph>
</caption>
<paragraph id="4C14CB1F140D5552D1DAFEB2FCBDB745" blockId="8.[189,1398,267,328]" pageId="8" pageNumber="9">
Aedeagus with relatively long parameres (0.65 length of phallobase,
<figureCitation id="D490D79A140D5552D56CFEB2FBA4B728" box="[1035,1107,267,293]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[203,1399,636,1876]" captionTargetId="figure@7.[189,1400,635,1888]" captionTargetPageId="7" captionText="Fig. 2. Orphnus renaudi sp. nov. AB. Habitus. CD. Head in dorsal and apical view. E. Aedeagus in lateral view. F. Parameres in dorsal view. G. Labels of holotype. H. Abdomen in dorsal view. I. Stridulatory field. A, CD, G. Holotype. B, EF, HI. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850307" httpUri="https://zenodo.org/record/3850307/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
EF), narrowly rounded apically in dorsal and lateral view, without excavations.
</paragraph>
<paragraph id="4C14CB1F140D5552D18DFEC9FEB2B787" blockId="8.[189,325,368,429]" box="[234,325,368,394]" pageId="8" pageNumber="9">
<emphasis id="7EDF170D140D5552D18DFEC9FEB2B787" bold="true" box="[234,325,368,394]" pageId="8" pageNumber="9">Female</emphasis>
</paragraph>
<paragraph id="4C14CB1F140D5552D1DAFE2AFECEB7A0" blockId="8.[189,325,368,429]" box="[189,313,403,429]" pageId="8" pageNumber="9">Unknown.</paragraph>
<paragraph id="4C14CB1F140D5552D1DAFE6DFEC4B7E3" blockId="8.[189,307,468,494]" box="[189,307,468,494]" pageId="8" pageNumber="9">
<heading id="175C7C73140D5552D1DAFE6DFEC4B7E3" bold="true" box="[189,307,468,494]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="7EDF170D140D5552D1DAFE6DFEC4B7E3" bold="true" box="[189,307,468,494]" pageId="8" pageNumber="9">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="4C14CB1F140D5552D1DAFDBAFD8CB44D" blockId="8.[189,1398,515,577]" pageId="8" pageNumber="9">
Body length of the
<typeStatus id="931075BD140D5552D0C9FDBDFDE9B413" box="[430,542,516,542]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
varies from 11.514.0 mm. In the smallest
<typeStatus id="931075BD140D5552D55CFDBDFB57B413" box="[1083,1184,516,542]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
the frontoclypeal process is feebly developed (
<figureCitation id="D490D79A140D5552D377FD9FFD9AB44D" box="[528,621,550,576]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[203,1399,636,1876]" captionTargetId="figure@7.[189,1400,635,1888]" captionTargetPageId="7" captionText="Fig. 2. Orphnus renaudi sp. nov. AB. Habitus. CD. Head in dorsal and apical view. E. Aedeagus in lateral view. F. Parameres in dorsal view. G. Labels of holotype. H. Abdomen in dorsal view. I. Stridulatory field. A, CD, G. Holotype. B, EF, HI. Paratypes." figureDoi="http://doi.org/10.5281/zenodo.3850307" httpUri="https://zenodo.org/record/3850307/files/figure.png" pageId="8" pageNumber="9">Fig. 2B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="04B19894140D5552D1DAFDD1FB7BB530" pageId="8" pageNumber="9" type="distribution">
<paragraph id="4C14CB1F140D5552D1DAFDD1FEA1B48F" blockId="8.[189,342,616,642]" box="[189,342,616,642]" pageId="8" pageNumber="9">
<heading id="175C7C73140D5552D1DAFDD1FEA1B48F" bold="true" box="[189,342,616,642]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="7EDF170D140D5552D1DAFDD1FEA1B48F" bold="true" box="[189,342,616,642]" pageId="8" pageNumber="9">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="4C14CB1F140D5552D1DAFD2FFB7BB530" blockId="8.[189,1399,662,830]" pageId="8" pageNumber="9">
The distribution and habitat of
<taxonomicName id="8BABB09C140D5552D34DFD2EFD5FB4BD" authorityName="Frolov &amp; Akhmetova" authorityYear="2016" box="[554,680,662,689]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="renaudi">
<emphasis id="7EDF170D140D5552D34DFD2EFD5FB4BD" box="[554,680,662,689]" italics="true" pageId="8" pageNumber="9">O. renaudi</emphasis>
</taxonomicName>
need further clarification. The only exact locality label, “Old Calabar” (modern town of Calabar,
<collectingCountry id="34BC8B8F140D5552D33CFD00FD42B4D9" box="[603,693,697,724]" name="Nigeria" pageId="8" pageNumber="9">Nigeria</collectingCountry>
) suggests that the
<typeStatus id="931075BD140D5552D2EFFD00FC18B4DE" box="[904,1007,697,723]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
and two
<typeStatus id="931075BD140D5552D530FD03FB30B4D9" box="[1111,1223,698,724]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
were collected in the coastal area near the
<collectingRegion id="8E6F05FD140D5552D377FD64FD69B4FA" box="[528,670,733,759]" country="Nigeria" name="Cross River" pageId="8" pageNumber="9">Cross River</collectingRegion>
estuary (
<figureCitation id="D490D79A140D5552D276FD64FCABB4F5" box="[785,860,733,760]" captionStart="Fig" captionStartId="16.[189,232,1966,1992]" captionTargetBox="[194,1393,1011,1923]" captionTargetId="figure@16.[189,1398,1009,1925]" captionTargetPageId="16" captionText="Fig. 6. Orphnus (Phornus) spp. Locality map. Orphnus parastrangulatus sp. nov. is not mapped, because there are no more or less precise locality data available for this species." figureDoi="http://doi.org/10.5281/zenodo.3850323" httpUri="https://zenodo.org/record/3850323/files/figure.png" pageId="8" pageNumber="9">Fig. 6</figureCitation>
). It is possible, however, that the specimens were collected inland of the town of Calabar, in the foothills of Western High
<collectingRegion id="8E6F05FD140D5552D529FCB9FB5FB517" box="[1102,1192,768,794]" country="Nigeria" name="Plateau" pageId="8" pageNumber="9">Plateau</collectingRegion>
. Another locality, “Camaroons”, may refer to foothills of Mount
<collectingCountry id="34BC8B8F140D5552D387FC9AFCADB530" box="[736,858,803,829]" name="Cameroon" pageId="8" pageNumber="9">Cameroon</collectingCountry>
or the state of
<collectingCountry id="34BC8B8F140D5552D56DFC9AFB7FB530" box="[1034,1160,803,829]" name="Cameroon" pageId="8" pageNumber="9">Cameroon</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="04B19894140D5552D1DAFCDCFA94B237" pageId="8" pageNumber="9" type="discussion">
<paragraph id="4C14CB1F140D5552D1DAFCDCFED8B572" blockId="8.[189,303,869,895]" box="[189,303,869,895]" pageId="8" pageNumber="9">
<heading id="175C7C73140D5552D1DAFCDCFED8B572" bold="true" box="[189,303,869,895]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="7EDF170D140D5552D1DAFCDCFED8B572" bold="true" box="[189,303,869,895]" pageId="8" pageNumber="9">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="4C14CB1F140D5552D1DAFC2AFA94B237" blockId="8.[189,1399,915,1082]" pageId="8" pageNumber="9">
One
<typeStatus id="931075BD140D5552D19DFC2DFEA8B5A3" box="[250,351,916,942]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
of
<taxonomicName id="8BABB09C140D5552D0F6FC2DFDE3B5A0" authority="Frolov &amp; Akhmetova, 2016" authorityName="Frolov &amp; Akhmetova" authorityYear="2016" box="[401,532,915,942]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="renaudi" status="sp. nov.">
<emphasis id="7EDF170D140D5552D0F6FC2DFDE3B5A0" box="[401,532,915,942]" italics="true" pageId="8" pageNumber="9">O. renaudi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65ECAA76140D5552D346FC2DFD76B5A3" box="[545,641,916,942]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
from Calabar bears the label “Brachyorphnus ferrierei
<taxonomicNameLabel id="65ECAA76140D5552D455FC2DFA81B5A3" box="[1330,1398,916,942]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
R. Paulian det. /
<typeStatus id="931075BD140D5552D0E5FC0FFDD7B5DD" box="[386,544,950,976]" pageId="8" pageNumber="9" type="allotype">ALLOTYPE</typeStatus>
”, and another specimen designated below as the
<typeStatus id="931075BD140D5552D505FC0FFB3EB5DD" box="[1122,1225,950,976]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
of
<taxonomicName id="8BABB09C140D5552D596FC0EFA80B5DC" authority="Frolov &amp; Akhmetova, 2016" authorityName="Frolov &amp; Akhmetova" authorityYear="2016" box="[1265,1399,951,977]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ferrierei" status="sp. nov.">
<emphasis id="7EDF170D140D5552D596FC0EFA80B5DC" box="[1265,1399,951,977]" italics="true" pageId="8" pageNumber="9">O. ferrierei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="65ECAA76140D5552D1DAFC63FEE9B5F9" box="[189,286,986,1012]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, bears the label “Brachyorphnus ferrierei
<taxonomicNameLabel id="65ECAA76140D5552D245FC63FCAEB5F9" box="[802,857,986,1012]" pageId="8" pageNumber="9" rank="species">n.sp.</taxonomicNameLabel>
R. Paulian det. /
<typeStatus id="931075BD140D5552D55EFC63FB2CB5F9" box="[1081,1243,986,1012]" pageId="8" pageNumber="9" type="holotype">HOLOTYPE</typeStatus>
”. Apparently Paulian considered the two specimens as conspecific and provisionally labelled them as a female and a male of a taxon he intended to describe, but his name “
<emphasis id="7EDF170D140D5552D231FB99FB83B237" box="[854,1140,1056,1082]" italics="true" pageId="8" pageNumber="9">
Brachyorphnus
<taxonomicName id="8BABB09C140D5552D574FB99FB83B237" authorityName="Frolov &amp; Akhmetova" authorityYear="2016" box="[1043,1140,1056,1082]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ferrierei">ferrierei</taxonomicName>
</emphasis>
” was not published.
</paragraph>
</subSubSection>
</treatment>
</document>