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<document id="8E97CD4C5040E9479EA5BD8B49FBB445" ID-CLB-Dataset="8469" ID-DOI="10.24349/acarologia/20184236" ID-GBIF-Dataset="3c0084e1-d3c9-4633-8e39-1513f2148596" ID-ISSN="2107-7207" ID-Zenodo-Dep="4487493" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612204954064" checkinUser="felipe" docAuthor="Pešić, Vladimir &amp; Smit, Harry" docDate="2018" docId="C37687EFEC082244AD86FEB9FD50103D" docLanguage="en" docName="Acarologia.58.1.165-185.pdf" docOrigin="Acarologia 58 (1)" docStyle="DocumentStyle:0387F17C4AF9F8D6336E46CC5E5664D0.6:Acarologia.2018-.journal_article" docStyleId="0387F17C4AF9F8D6336E46CC5E5664D0" docStyleName="Acarologia.2018-.journal_article" docStyleVersion="6" docTitle="Atractides (Polymegapus) grigorievka Pešić &amp; Smit 2018, n. sp." docType="treatment" docVersion="7" lastPageNumber="175" masterDocId="3F4FFF97EC01224FAC6DFF85FF96111D" masterDocTitle="A checklist of the water mites of Central Asia with description of six new species (Acari, Hydrachnidia) from Kyrgyzstan" masterLastPageNumber="185" masterPageNumber="165" pageNumber="173" updateTime="1698925910332" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C3715E9DF57B29B96936E78312EF3CC5">A checklist of the water mites of Central Asia with description of six new species (Acari, Hydrachnidia) from Kyrgyzstan</mods:title>
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<mods:namePart id="14BD836F6135DB05AB1C695DAD221142">Pešić, Vladimir</mods:namePart>
<mods:affiliation id="DF6AB670B1B7B210753F90C9A414A6D8">Department of Biology, University of Montenegro, Cetinjski put b. b., 81000 Podgorica, Montenegro.</mods:affiliation>
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<mods:namePart id="FEE786E8CDCF1FCFFB44667119403C39">Smit, Harry</mods:namePart>
<mods:affiliation id="6B8825E2044BAB307AE0DB68EC06D112">Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, The Netherlands.</mods:affiliation>
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<mods:title id="E14CCF6BAA7C9E10089C0AEB87CC5909">Acarologia</mods:title>
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<mods:date id="78F1C557905C6571E0ED0CB591B6B7B1">2018</mods:date>
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<emphasis id="79ABEAEBEC082246AD86FEB9FB4E1045" bold="true" box="[491,1240,316,344]" pageId="9" pageNumber="173">
<taxonomicName id="8CDF4D7AEC082246AD86FEB9FC641045" authority="Pešić &amp; Smit, 2018" authorityName="Pešić &amp; Smit" authorityYear="2018" box="[491,1010,316,344]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="173" phylum="Arthropoda" rank="species" species="grigorievka" status="sp. nov." subGenus="Polymegapus">
<emphasis id="79ABEAEBEC082246AD86FEB9FDED1045" bold="true" box="[491,635,316,344]" italics="true" pageId="9" pageNumber="173">Atractides</emphasis>
(
<emphasis id="79ABEAEBEC082246AEE0FEB9FCD31045" bold="true" box="[653,837,316,344]" italics="true" pageId="9" pageNumber="173">Polymegapus</emphasis>
)
<emphasis id="79ABEAEBEC082246AF3AFEB9FC641045" bold="true" box="[855,1010,316,344]" italics="true" pageId="9" pageNumber="173">grigorievka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="62985790EC082246AF97FEB9FBD11045" box="[1018,1095,316,344]" pageId="9" pageNumber="173" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="D3E42A7CEC082246A834FEB9FB5B1045" box="[1113,1229,316,344]" captionStart="Figure 3" captionStartId="10.[119,186,1863,1884]" captionTargetBox="[252,1333,301,1804]" captionTargetId="figure-2@10.[250,1348,297,1809]" captionTargetPageId="10" captionText="Figure 3 Atractides grigorievkan. sp., female holotype: A idiosoma, ventral view; B genital field; C palp, lateral view; D palp, medial view; E I-Leg; F I-L-5 and -6; G sword setae (S-1 and -2) at the ventrodistal margin of I-L-5. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4487499" httpUri="https://zenodo.org/record/4487499/files/figure.png" pageId="9" pageNumber="173">Figure 3</figureCitation>
)
</emphasis>
</heading>
</paragraph>
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Zoobank:
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</paragraph>
</subSubSection>
<subSubSection id="03C56572EC082244AE7EFE3DFD50103D" lastPageId="11" lastPageNumber="175" pageId="9" pageNumber="173" type="materials_examined">
<paragraph id="4B6036F9EC082246AE7EFE3DFB2B1344" blockId="9.[491,1509,439,1461]" pageId="9" pageNumber="173">
Type material —
<materialsCitation id="FBB73CA4EC082246AEBFFE32FCA81309" ID-GBIF-Occurrence="3029771301" collectingDate="2013-07-11" collectionCode="RMNH" collectorName="Pesic &amp; Smit" country="Kyrgyzstan" elevation="1960" latitude="42.776417" location="Grigorievka Gorge" longLatPrecision="1" longitude="77.47318" pageId="9" pageNumber="173" specimenCount="1" typeStatus="holotype">
<typeStatus id="9464885BEC082246AEBFFE32FCA610D2" box="[722,816,439,463]" pageId="9" pageNumber="173" type="holotype">Holotype</typeStatus>
female (
<collectionCode id="2DCEAE3CEC082246AFFEFE32FC7E10D2" box="[915,1000,439,463]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="9" pageNumber="173" type="Museum">RMNH</collectionCode>
),
<collectingCountry id="33C87669EC082246AF90FE3DFBE710CD" box="[1021,1137,440,464]" name="Kyrgyzstan" pageId="9" pageNumber="173">Kyrgyzstan</collectingCountry>
, KR10
<location id="4E006022EC082246A8A5FE32FA0010D2" LSID="urn:lsid:plazi:treatment:C37687EFEC082244AD86FEB9FD50103D:4E006022EC082246A8A5FE32FA0010D2" box="[1224,1430,439,463]" country="Kyrgyzstan" latitude="42.776417" longLatPrecision="1" longitude="77.47318" name="Grigorievka Gorge" pageId="9" pageNumber="173">Grigorievka Gorge</location>
, rheocrenic spring,
<geoCoordinate id="2EEB503EEC082246AED8FE5CFCAD10EC" box="[693,827,473,497]" degrees="42" direction="north" minutes="46.585" orientation="latitude" pageId="9" pageNumber="173" precision="1" value="42.776417">42°46.585N</geoCoordinate>
,
<geoCoordinate id="2EEB503EEC082246AF2AFE5CFC2710EC" box="[839,945,473,497]" degrees="77" direction="east" minutes="28.391" orientation="longitude" pageId="9" pageNumber="173" precision="1" value="77.47318">77°28.391E</geoCoordinate>
,
<elevation id="C0F2D1CAEC082246AFD1FE5CFBA510EC" box="[956,1075,473,497]" metricMagnitude="3" metricUnit="m" metricValue="1.96" pageId="9" pageNumber="173" unit="m" value="1960.0">
<quantity id="8C279B1CEC082246AFD1FE5CFB9F10EC" box="[956,1033,473,497]" metricMagnitude="3" metricUnit="m" metricValue="1.96" pageId="9" pageNumber="173" unit="m" value="1960.0">1960 m</quantity>
asl
</elevation>
.,
<date id="3F611039EC082246A829FE5CFB2310EC" box="[1092,1205,473,497]" pageId="9" pageNumber="173" value="2013-07-11">
<collectingDate id="2F25E9D1EC082246A829FE5CFB2310EC" box="[1092,1205,473,497]" pageId="9" pageNumber="173" value="2013-07-11">11.vii.2013</collectingDate>
</date>
, leg.
<collectorName id="E62A532FEC082246A89BFE5CFAB810EC" box="[1270,1326,473,497]" pageId="9" pageNumber="173">Pešić</collectorName>
&amp;
<collectorName id="E62A532FEC082246A93EFE5CFA1E10EC" box="[1363,1416,473,497]" pageId="9" pageNumber="173">Smit</collectorName>
, dissected and slide mounted
</materialsCitation>
.
<typeStatus id="9464885BEC082246AF3FFE79FC391309" box="[850,943,508,532]" pageId="9" pageNumber="173" type="paratype">Paratype</typeStatus>
: one female,
<collectingCountry id="33C87669EC082246A824FE79FB2B1309" box="[1097,1213,508,532]" name="Kyrgyzstan" pageId="9" pageNumber="173">Kyrgyzstan</collectingCountry>
, KR23 Karakal region, rheohelocrenic spring, Žeti-Egaz village,
<geoCoordinate id="2EEB503EEC082246AFC8FD9BFB8F132B" box="[933,1049,542,566]" degrees="42" direction="north" minutes="19.238" orientation="latitude" pageId="9" pageNumber="173" precision="1" value="42.320633">42°19.238N</geoCoordinate>
,
<geoCoordinate id="2EEB503EEC082246A849FD9BFB0A132B" box="[1060,1180,542,566]" degrees="78" direction="east" minutes="51.017" orientation="longitude" pageId="9" pageNumber="173" precision="1" value="78.85028">78°51.017E</geoCoordinate>
,
<quantity id="8C279B1CEC082246A8CAFD9BFB7A132B" box="[1191,1260,542,566]" metricMagnitude="3" metricUnit="m" metricValue="2.094" pageId="9" pageNumber="173" unit="m" value="2094.0">2094 m</quantity>
asl.,
<date id="3F611039EC082246A949FD9BFA04132B" box="[1316,1426,542,566]" pageId="9" pageNumber="173" value="2013-07-13">13.vii.2013</date>
, leg. Pešić &amp; Smit, left palp and I-L dissected and slide mounted.
</paragraph>
<paragraph id="4B6036F9EC082246AE7EFDE6FC0813FF" blockId="9.[491,1509,439,1461]" pageId="9" pageNumber="173">Diagnosis — Female (Male unknown). Integument striated; suture line Cx-III and IV in its medial part directed posteriorly; P-3 ventral margin slightly convex with one (occasionally two) ventral seta; P-5 short; excretory pore smooth; acetabula in a slightly curved line; I-L-5 with S-1 ending in a bluntly pointed tip.</paragraph>
<paragraph id="4B6036F9EC082246AE7EFD68FD1C1527" blockId="9.[491,1509,439,1461]" pageId="9" pageNumber="173">
Description — Female — Integument dorsally striated, muscle insertions unsclerotized. Posterior margin of Cx-I weakly curved, with a subcuticular protrusion; suture line Cx-III and -IV in its medial part directed posteriorly (
<figureCitation id="D3E42A7CEC082246AF9CFCB4FBC91254" box="[1009,1119,817,841]" captionStart="Figure 3" captionStartId="10.[119,186,1863,1884]" captionTargetBox="[252,1333,301,1804]" captionTargetId="figure-2@10.[250,1348,297,1809]" captionTargetPageId="10" captionText="Figure 3 Atractides grigorievkan. sp., female holotype: A idiosoma, ventral view; B genital field; C palp, lateral view; D palp, medial view; E I-Leg; F I-L-5 and -6; G sword setae (S-1 and -2) at the ventrodistal margin of I-L-5. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4487499" httpUri="https://zenodo.org/record/4487499/files/figure.png" pageId="9" pageNumber="173">Figure 3A</figureCitation>
). Genital plates with narrow sclerite border, slightly indented between Ac-1/2 and Ac-2/3, acetabula in a slightly curved line (
<figureCitation id="D3E42A7CEC082246A9F2FCD1FD9C1293" captionStart="Figure 3" captionStartId="10.[119,186,1863,1884]" captionTargetBox="[252,1333,301,1804]" captionTargetId="figure-2@10.[250,1348,297,1809]" captionTargetPageId="10" captionText="Figure 3 Atractides grigorievkan. sp., female holotype: A idiosoma, ventral view; B genital field; C palp, lateral view; D palp, medial view; E I-Leg; F I-L-5 and -6; G sword setae (S-1 and -2) at the ventrodistal margin of I-L-5. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4487499" httpUri="https://zenodo.org/record/4487499/files/figure.png" pageId="9" pageNumber="173">Figure 3B</figureCitation>
). Excretory pore smooth; Vgl-1 not fused to Vgl-2. P-3 ventral margin slightly convex, with one (occasionally two) ventral seta(e), inserted in the middle of segment; P-4 setae insertions dividing ventral margin into sectors 1:4:2, sword seta proximally from proximoventral seta; P-5 short (
<figureCitation id="D3E42A7CEC082246AE42FC58FD0612E8" box="[559,656,989,1013]" captionStart="Figure 3" captionStartId="10.[119,186,1863,1884]" captionTargetBox="[252,1333,301,1804]" captionTargetId="figure-2@10.[250,1348,297,1809]" captionTargetPageId="10" captionText="Figure 3 Atractides grigorievkan. sp., female holotype: A idiosoma, ventral view; B genital field; C palp, lateral view; D palp, medial view; E I-Leg; F I-L-5 and -6; G sword setae (S-1 and -2) at the ventrodistal margin of I-L-5. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4487499" httpUri="https://zenodo.org/record/4487499/files/figure.png" pageId="9" pageNumber="173">Figures 3</figureCitation>
C-D). I-L-5: S-1 with a blunt tip, more slender than S-2, S-2 pointed, strongly enlarged in the proximal part (
<figureCitation id="D3E42A7CEC082246AF55FB85FC061505" box="[824,912,1024,1048]" captionStart="Figure 3" captionStartId="10.[119,186,1863,1884]" captionTargetBox="[252,1333,301,1804]" captionTargetId="figure-2@10.[250,1348,297,1809]" captionTargetPageId="10" captionText="Figure 3 Atractides grigorievkan. sp., female holotype: A idiosoma, ventral view; B genital field; C palp, lateral view; D palp, medial view; E I-Leg; F I-L-5 and -6; G sword setae (S-1 and -2) at the ventrodistal margin of I-L-5. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4487499" httpUri="https://zenodo.org/record/4487499/files/figure.png" pageId="9" pageNumber="173">Fig. 3G</figureCitation>
); I-L-6 stout, very thick at the base, distally tapering (
<figureCitation id="D3E42A7CEC082246AD9EFBA7FDC21527" box="[499,596,1058,1082]" captionStart="Figure 3" captionStartId="10.[119,186,1863,1884]" captionTargetBox="[252,1333,301,1804]" captionTargetId="figure-2@10.[250,1348,297,1809]" captionTargetPageId="10" captionText="Figure 3 Atractides grigorievkan. sp., female holotype: A idiosoma, ventral view; B genital field; C palp, lateral view; D palp, medial view; E I-Leg; F I-L-5 and -6; G sword setae (S-1 and -2) at the ventrodistal margin of I-L-5. Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4487499" httpUri="https://zenodo.org/record/4487499/files/figure.png" pageId="9" pageNumber="173">Figures 3</figureCitation>
E-F).
</paragraph>
<paragraph id="4B6036F9EC082246AE7EFBC0FD2915D9" blockId="9.[491,1509,439,1461]" pageId="9" pageNumber="173">
Measurements — (
<typeStatus id="9464885BEC082246AEB1FBC0FCAC1540" box="[732,826,1093,1117]" pageId="9" pageNumber="173" type="holotype">holotype</typeStatus>
; in parentheses some measurements of
<typeStatus id="9464885BEC082246A8B0FBC0FAA01540" box="[1245,1334,1093,1117]" pageId="9" pageNumber="173" type="paratype">paratype</typeStatus>
) — Idiosoma L 938 (1113), W 688 (925); coxal field: L 434; Cx-III W 553; Cx-I+II mL 134, lL 355; genital field L/W 223/224, genital plates L 144-145, pregenital sclerite W 122, maximum diameter Ac 1-3: 44, 31-38, 47-51.
</paragraph>
<paragraph id="4B6036F9EC082246AE7EFB4BFB631436" blockId="9.[491,1509,439,1461]" pageId="9" pageNumber="173">Gnathosoma — palp total L 450; dL/H, dL/H ratio: P-1, 48/44, 1.1 (50/47, 1.06); P-2, 127/92, 1.38 (121/91, 1.34); P-3, 103/80, 1.29 (95/79, 1.2); P-4, 138/43, 3.2 (123/44, 2.82); P-5, 34/19, 1.83 (33/19, 1.75); L ratio P-2/P-4 0.92 (0.98). Chelicera total L 298.</paragraph>
<paragraph id="4B6036F9EC082246AE7EFAB3FD7314A8" blockId="9.[491,1509,439,1461]" pageId="9" pageNumber="173">Legs — I-L-5 dL 248 (232), vL 217 (186), dL/vL ratio 1.15 (1.25), maximum H 67 (72), dL/maximum H 3.7 (3.2), S-1 L 66, L/W ratio 5.3, S-2 L 70, L/W ratio 4.5, distance S-1-2 5.0, L ratio S-1/2 0.95; I-L-6 L 119 (108), maximum H 42 (41), L/maximum H ratio 2.8 (2.6); L ratio I-L-5/6, 2.1 (2.15).</paragraph>
<paragraph id="4B6036F9EC082246AD86FA64FB141669" blockId="9.[491,1511,1505,1909]" pageId="9" pageNumber="173">
Discussion — The new species is provisionally assigned to the subgenus
<taxonomicName id="8CDF4D7AEC082246A966FA64FDC91701" authority="K. Viets, 1926" authorityName="K. Viets" authorityYear="1926" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="173" phylum="Arthropoda" rank="subGenus" subGenus="Polymegapus">
<emphasis id="79ABEAEBEC082246A966FA64FA3514E4" box="[1291,1443,1505,1529]" italics="true" pageId="9" pageNumber="173">Polymegapus</emphasis>
K. Viets, 1926
</taxonomicName>
, which was considered by
<bibRefCitation id="2F4E4B08EC082246AFE6F981FBBF1701" author="Gerecke R." box="[907,1065,1540,1564]" pageId="9" pageNumber="173" refId="ref8910" refString="Gerecke R. 2003 - Water mites of the genus Atractides Koch, 1837 (Acari: Parasitengona: Hygrobatidae)" type="book" year="2003">Gerecke (2003)</bibRefCitation>
to be a monophyletic clade. The new species does not fit the diagnosis of the subgenus given by
<bibRefCitation id="2F4E4B08EC082246A8E8F9A3FAF11723" author="Gerecke R. &amp; Gledhill T. &amp; Pesic V. &amp; Smit H." box="[1157,1383,1574,1598]" pageId="9" pageNumber="173" pagination="1 - 429" refId="ref8973" refString="Gerecke R., Gledhill T., Pesic V., Smit H. 2016 - Chelicerata: Acari III - In: Gerecke R, ed. Susswasserfauna von Mitteleuropa, Bd. 7 / 2 - 3. Springer-Verlag Berlin, Heidelberg, pp. 1 - 429." type="book chapter" year="2016">
Gerecke
<emphasis id="79ABEAEBEC082246A8B1F9A2FA9B1723" box="[1244,1293,1574,1598]" italics="true" pageId="9" pageNumber="173">et al</emphasis>
. (2016)
</bibRefCitation>
in the following features: 1) suture line Cx-III and IV in its medial part directed posteriorly, 2) Cx-4 without triangular laterocaudal extensions of secondary sclerite, 3) I-L-5 with ventral margin shorter than dorsal margin, 4) P-5 short and less slender, and 5) excretory pore unsclerotized. The combination of the characters given under 1), 3) and 5) make the new species close to
<taxonomicName id="8CDF4D7AEC082246A9D7F935FD3217F7" authority="Wiles, 1991" authorityName="Wiles" authorityYear="1991" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="173" phylum="Arthropoda" rank="species" species="putihi">
<emphasis id="79ABEAEBEC082246A9D7F935FDBF17F7" italics="true" pageId="9" pageNumber="173">A. putihi</emphasis>
<bibRefCitation id="2F4E4B08EC082246AE5CF957FD3217F7" author="Wiles P. R." box="[561,676,1746,1770]" pageId="9" pageNumber="173" refId="ref9960" refString="Wiles P. R. 1991 - Rheophilic watermites (Acari: Hydrachnidia) from mainland Malaysia - Acarologia," type="book" year="1991">Wiles, 1991</bibRefCitation>
</taxonomicName>
from SE Asia (
<bibRefCitation id="2F4E4B08EC082246AF20F957FC2817F7" author="Wiles P. R." box="[845,958,1746,1770]" pageId="9" pageNumber="173" refId="ref9960" refString="Wiles P. R. 1991 - Rheophilic watermites (Acari: Hydrachnidia) from mainland Malaysia - Acarologia," type="book" year="1991">Wiles 1991</bibRefCitation>
,
<bibRefCitation id="2F4E4B08EC082246AFA4F957FB1D17F7" author="Pesic V. &amp; Smit H." box="[969,1163,1746,1770]" pageId="9" pageNumber="173" pagination="1 - 30" refId="ref9087" refString="Pesic V., Smit H. 2009 - New records of water mites of the genus Atractides Koch, 1837 (Acari: Hydrachnidia, Hygrobatidae) from Thailand, Malaysia and Sulawesi (Indonesia), with the description of four new species - Zootaxa, 2240: 1 - 30." type="journal article" year="2009">Pešić &amp; Smit 2009</bibRefCitation>
,
<bibRefCitation id="2F4E4B08EC082246A8FAF957FAF417F7" author="Pesic V. &amp; Smit H." box="[1175,1378,1746,1770]" pageId="9" pageNumber="173" pagination="1 - 72" refId="ref9141" refString="Pesic V., Smit H. 2014 - Torrenticolid water mites (Acari: Hydrachnidia: Torrenticolidae) from Malaysian Borneo - Zootaxa, 3840 (1): 1 - 72. doi: 10.11646 / zootaxa. 3840.1.1" type="journal article" year="2014">Smit &amp; Pešić 2014</bibRefCitation>
). The latter species differs in genital plates, restricted to the level of the postgenital platelet, with the acetabula in a triangular arrangement, P-3 ventral margin slightly concave, without setae and IV-L-5 with 2 swimming setae. Probably
<taxonomicName id="8CDF4D7AEC082246AFC6F8BCFBDA164C" authority="Pešić &amp; Smit, 2018" authorityName="Pešić &amp; Smit" authorityYear="2018" box="[939,1100,1849,1873]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="9" pageNumber="173" phylum="Arthropoda" rank="species" species="grigorievka" status="sp. nov.">
<emphasis id="79ABEAEBEC082246AFC6F8BCFBDA164C" box="[939,1100,1849,1873]" italics="true" pageId="9" pageNumber="173">A. grigorievka</emphasis>
</taxonomicName>
<emphasis id="79ABEAEBEC082246A821F8BFFB1A164F" bold="true" box="[1100,1164,1850,1874]" pageId="9" pageNumber="173">
<taxonomicNameLabel id="62985790EC082246A821F8BFFB1A164F" box="[1100,1164,1850,1874]" pageId="9" pageNumber="173" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
represents an early derivative member with the typical palp and I-L not yet fully developed.
</paragraph>
<paragraph id="4B6036F9EC082246AD86F824FD2116C6" blockId="9.[491,1461,1953,2011]" pageId="9" pageNumber="173">Etymology — Named after the canyon where the new species was found. The name is a noun in apposition.</paragraph>
<caption id="1FA06671EC0B2245AC1AF8C2FA99166E" ID-DOI="http://doi.org/10.5281/zenodo.4487499" ID-Zenodo-Dep="4487499" httpUri="https://zenodo.org/record/4487499/files/figure.png" pageId="10" pageNumber="174" startId="10.[119,186,1863,1884]" targetBox="[252,1333,301,1804]" targetPageId="10">
<paragraph id="4B6036F9EC0B2245AC1AF8C2FA99166E" blockId="10.[119,1489,1862,1907]" pageId="10" pageNumber="174">
<emphasis id="79ABEAEBEC0B2245AC1AF8C2FF5A1641" bold="true" box="[119,204,1863,1884]" pageId="10" pageNumber="174">Figure 3</emphasis>
<taxonomicName id="8CDF4D7AEC0B2245ACBBF8C3FE251641" authority="Pešić &amp; Smit, 2018" authorityName="Pešić &amp; Smit" authorityYear="2018" box="[214,435,1862,1884]" class="Arachnida" family="Hygrobatidae" genus="Atractides" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="174" phylum="Arthropoda" rank="species" species="grigorievka" status="sp. nov.">
<emphasis id="79ABEAEBEC0B2245ACBBF8C3FE251641" box="[214,435,1862,1884]" italics="true" pageId="10" pageNumber="174">Atractides grigorievka</emphasis>
</taxonomicName>
<emphasis id="79ABEAEBEC0B2245ADDEF8C2FE7D1641" bold="true" box="[435,491,1863,1884]" pageId="10" pageNumber="174">
<taxonomicNameLabel id="62985790EC0B2245ADDEF8C2FE7D1641" box="[435,491,1863,1884]" pageId="10" pageNumber="174" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female holotype: A idiosoma, ventral view; B genital field; C palp, lateral view; D palp, medial view; E I-Leg; F I-L-5 and -6; G sword setae (S-1 and -2) at the ventrodistal margin of I-L-5. Scale bars = 100 μm.
</paragraph>
</caption>
<paragraph id="4B6036F9EC0A2244AD86FF60FD50103D" blockId="11.[491,1489,229,288]" pageId="11" pageNumber="175">
Distribution —
<collectingCountry id="33C87669EC0A2244AEFBFF63FC9C11E3" box="[662,778,230,254]" name="Kyrgyzstan" pageId="11" pageNumber="175">Kyrgyzstan</collectingCountry>
, known from one rheocrenic (
<figureCitation id="D3E42A7CEC0A2244A827FF60FB5211E0" box="[1098,1220,229,253]" captionStart="Figure 10" captionStartId="20.[119,186,1853,1874]" captionTargetBox="[175,1421,220,1800]" captionTargetId="figure-2@20.[175,1421,220,1800]" captionTargetPageId="20" captionText="Figure 10 Photographs of sampling sites: A KR3, Ala Archa NP, spring outlet (locus typicusof Atractides alaarchaensis n. sp.); B KR32, stream near Son Kul lake (locus typicusof Atractides sonkulensisn. sp.); C KR10, spring in Grigorievka canyon (locus typicusof Atractides grigorievkan. sp.); D KR8, Chon-Kemin NP, upper part of stream near Ashu resort (locus typicusof Torrenticola kyrgyzican. sp.); E KR15, river on the road to May Saz Pass (locus typicusof Wandesia albertii n. sp.); F KR22, stream in Karakal region (locus typicusof Atractides manasi n. sp.); G KR23, rheohelocrenic spring in Karakal region (sampling site of Atractides grigorievkan. sp.); H KR31, river near Son Kol Lake (sampling site of Atractides sonkulensisn. sp.)." figureDoi="http://doi.org/10.5281/zenodo.4487513" httpUri="https://zenodo.org/record/4487513/files/figure.png" pageId="11" pageNumber="175">Figure 10C</figureCitation>
) and one rheohelocrenic spring (
<figureCitation id="D3E42A7CEC0A2244AE51FE8DFD2C103D" box="[572,698,264,288]" captionStart="Figure 10" captionStartId="20.[119,186,1853,1874]" captionTargetBox="[175,1421,220,1800]" captionTargetId="figure-2@20.[175,1421,220,1800]" captionTargetPageId="20" captionText="Figure 10 Photographs of sampling sites: A KR3, Ala Archa NP, spring outlet (locus typicusof Atractides alaarchaensis n. sp.); B KR32, stream near Son Kul lake (locus typicusof Atractides sonkulensisn. sp.); C KR10, spring in Grigorievka canyon (locus typicusof Atractides grigorievkan. sp.); D KR8, Chon-Kemin NP, upper part of stream near Ashu resort (locus typicusof Torrenticola kyrgyzican. sp.); E KR15, river on the road to May Saz Pass (locus typicusof Wandesia albertii n. sp.); F KR22, stream in Karakal region (locus typicusof Atractides manasi n. sp.); G KR23, rheohelocrenic spring in Karakal region (sampling site of Atractides grigorievkan. sp.); H KR31, river near Son Kol Lake (sampling site of Atractides sonkulensisn. sp.)." figureDoi="http://doi.org/10.5281/zenodo.4487513" httpUri="https://zenodo.org/record/4487513/files/figure.png" pageId="11" pageNumber="175">Figure 10G</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>