441 lines
57 KiB
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441 lines
57 KiB
XML
<document id="1C04D7FC561777DEDBE6C973D3F564A7" ID-DOI="10.1080/00222930701228132" ID-ISSN="1464-5262" ID-Zenodo-Dep="4582056" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614875044252" checkinUser="carolina" docAuthor="Peña Cantero, Alvaro L., Marques, Antonio C. & Migotto, Alvaro E." docDate="2007" docId="AC456037600F626DFE3CFC188E486AC2" docLanguage="en" docName="JNATHIST.41.5-8.229-291.pdf" docOrigin="Journal of Natural History 41 (5 - 8)" docSource="http://dx.doi.org/10.1080/00222930701228132" docStyle="DocumentStyle:7E99CF6C63B49250F2A87C3AEDC82D8C.2:JNatHist.2005-2007.journal_article.2cover" docStyleId="7E99CF6C63B49250F2A87C3AEDC82D8C" docStyleName="JNatHist.2005-2007.journal_article.2cover" docStyleVersion="2" docTitle="Acryptolaria operculata Stepanjants 1979" docType="treatment" docVersion="3" lastPageNumber="261" masterDocId="507C184F6010624FFFBDFFEE8D4B6862" masterDocTitle="Revision of the genus Acryptolaria Norman, 1875 (Cnidaria, Hydrozoa, Lafoeidae)" masterLastPageNumber="291" masterPageNumber="229" pageNumber="258" updateTime="1698880324352" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="8A934BE079F12E23FB67B3E610223FF5">Revision of the genus Acryptolaria Norman, 1875 (Cnidaria, Hydrozoa, Lafoeidae)</mods:title>
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<mods:namePart id="7682066600BCD11B85082B4F52972A84">Peña Cantero, Alvaro L.</mods:namePart>
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<mods:namePart id="41F2E176AF5A986325AB4B05393F3A22">Marques, Antonio C.</mods:namePart>
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<mods:namePart id="F15EBE0632FBF7A9705B92B768E75D46">Migotto, Alvaro E.</mods:namePart>
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<mods:title id="E47C6F855CB704B0197449A4FBE992C2">Journal of Natural History</mods:title>
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<mods:part id="F579F78ED0398BFAB5384C4182EE6ED1">
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<mods:date id="E72351204DD79F73AEF48444AA84B8B1">2007</mods:date>
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<mods:number id="C9C4699A69214FDE58D50681696E796F">2007-03-26</mods:number>
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<mods:number id="1EFA76E91CE56DD4A902D05991A2A0E6">5 - 8</mods:number>
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<treatment id="AC456037600F626DFE3CFC188E486AC2" LSID="urn:lsid:plazi:treatment:AC456037600F626DFE3CFC188E486AC2" httpUri="http://treatment.plazi.org/id/AC456037600F626DFE3CFC188E486AC2" lastPageId="34" lastPageNumber="261" pageId="31" pageNumber="258">
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<subSubSection id="6CF682AA600F6250FE3CFC188EDA6C6C" box="[385,913,1014,1038]" pageId="31" pageNumber="258" type="nomenclature">
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<paragraph id="2453D121600F6250FE3CFC188EDA6C6C" blockId="31.[125,1172,1014,1202]" box="[385,913,1014,1038]" pageId="31" pageNumber="258">
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<heading id="7F1B664D600F6250FE3CFC188EDA6C6C" box="[385,913,1014,1038]" centered="true" fontSize="10" level="2" pageId="31" pageNumber="258" reason="2">
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<taxonomicName id="E3ECAAA2600F6250FE3CFC188EDA6C6C" ID-CoL="9RXF" authority="Stepanjants, 1979" authorityName="Stepanjants" authorityYear="1979" box="[385,913,1014,1038]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="31" pageNumber="258" phylum="Cnidaria" rank="species" species="operculata">
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<emphasis id="16980D33600F6250FE3CFC188FFE6C6C" bold="true" box="[385,693,1014,1038]" italics="true" pageId="31" pageNumber="258">Acryptolaria operculata</emphasis>
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<bibRefCitation id="407DACD0600F6250FD02FC188EDA6C6C" author="Stepanjants SD" box="[703,913,1014,1038]" pageId="31" pageNumber="258" pagination="1 - 200" refId="ref22474" refString="Stepanjants SD. 1979. Hydroids of the antarctic and subantarctic waters. In: Biological results of the Soviet Antarctic Expeditions, 6. Issledovaniya Fauny Morei 20: 1 - 200. (Rus)." type="journal article" year="1979">Stepanjants, 1979</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="6CF682AA600F6250FE1FFBF98E336C4D" box="[418,888,1047,1071]" pageId="31" pageNumber="258" type="description">
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<paragraph id="2453D121600F6250FE1FFBF98E336C4D" blockId="31.[125,1172,1014,1202]" box="[418,888,1047,1071]" pageId="31" pageNumber="258">
|
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(
|
||
<figureCitation id="BCD7CDA4600F6250FE17FBF98F6C6C4D" box="[426,551,1047,1071]" captionStart="Figure 12" captionStartId="32.[138,198,1099,1118]" captionTargetBox="[314,1008,137,1059]" captionTargetId="figure-125@32.[310,1013,133,1063]" captionTargetPageId="32" captionText="Figure 12. Acryptolaria operculata Stepanjants, 1979. (A, B) Branch fragments showing shape and arrangement of hydrothecae. All drawings from the paratype. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4582092" httpUri="https://zenodo.org/record/4582092/files/figure.png" pageId="31" pageNumber="258">Figures 12</figureCitation>
|
||
,
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||
<figureCitation id="BCD7CDA4600F6250FD88FBF98F206C4D" box="[565,619,1047,1071]" captionStart="Figure 16" captionStartId="41.[125,185,1565,1584]" captionTargetBox="[9,1032,133,1529]" captionTargetId="figure-1@41.[266,1032,133,1529]" captionTargetPageId="41" captionText="Figure 16. Hydrothecal arrangement in Acryptolaria (continued). (A) A. longitheca; (B) A. minima; (C) A. minuta; (D) A. operculata; (E) A. pulchella; (F) A. tortugasensis. Scale bar: 500 Mm (A, D); 250 Mm (B, C, E, F)." figureDoi="http://doi.org/10.5281/zenodo.4582104" httpUri="https://zenodo.org/record/4582104/files/figure.png" pageId="31" pageNumber="258">16D</figureCitation>
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,
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<figureCitation id="BCD7CDA4600F6250FDC4FBF98FE46C4D" box="[633,687,1047,1071]" captionStart="Figure 18" captionStartId="43.[125,185,1625,1644]" captionTargetBox="[9,1010,135,1588]" captionTargetId="figure-1@43.[285,1012,135,1589]" captionTargetPageId="43" captionText="Figure 18. Shape of hydrotheca in Acryptolaria (continued). (A) A. gracilis; (B) A. longitheca; (C) A. minima; (D) A. operculata; (E) A. pulchella; (F) A. tortugasensis. Scale bar: 100 Mm (A, C, F); 250 Mm (B, D, E)." figureDoi="http://doi.org/10.5281/zenodo.4582110" httpUri="https://zenodo.org/record/4582110/files/figure.png" pageId="31" pageNumber="258">18D</figureCitation>
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,
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||
<figureCitation id="BCD7CDA4600F6250FD03FBF98FA46C4D" box="[702,751,1047,1071]" captionStart="Figure 19" captionStartId="44.[138,198,1482,1501]" captionTargetBox="[9,1044,133,1447]" captionTargetId="figure-1@44.[278,1044,133,1447]" captionTargetPageId="44" captionText="Figure 19. Shape of gonothecae: (A) Acryptolaria conferta; (B) A. minima. Detail of different structures: (C) bottleneck-shaped basal part of hydrotheca in A. gracilis; (D) opercular structure in A. minima; (E) opercular structure in A. operculata; (F) longitudinal striae in hydrotheca of A. flabellum; (G) intrathecal cusp of A. tortugasensis. Scale bar: 100 Mm (A, B, D, F, G); 50 Mm (C); 250 Mm (E)." figureDoi="http://doi.org/10.5281/zenodo.4582112" httpUri="https://zenodo.org/record/4582112/files/figure.png" pageId="31" pageNumber="258">19E</figureCitation>
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; Table XII)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="6CF682AA600F626FFFC0FBD68E3D6D52" lastPageId="32" lastPageNumber="259" pageId="31" pageNumber="258" type="reference_group">
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||
<paragraph id="2453D121600F6250FFC0FBD68ED86C32" blockId="31.[125,1172,1014,1202]" box="[125,915,1080,1104]" pageId="31" pageNumber="258">
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<taxonomicName id="E3ECAAA2600F6250FFC0FBD68FD86C32" ID-CoL="9RXF" authority="Stepanjants 1979, p 52" authorityName="Stepanjants" authorityPageNumber="52" authorityYear="1979" box="[125,659,1080,1104]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="31" pageNumber="258" phylum="Cnidaria" rank="species" species="operculata">
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<emphasis id="16980D33600F6250FFC0FBD68C306C32" box="[125,379,1080,1104]" italics="true" pageId="31" pageNumber="258">Acryptolaria operculata</emphasis>
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||
<bibRefCitation id="407DACD0600F6250FE38FBD68F196C32" author="Stepanjants SD" box="[389,594,1080,1104]" pageId="31" pageNumber="258" pagination="1 - 200" refId="ref22474" refString="Stepanjants SD. 1979. Hydroids of the antarctic and subantarctic waters. In: Biological results of the Soviet Antarctic Expeditions, 6. Issledovaniya Fauny Morei 20: 1 - 200. (Rus)." type="journal article" year="1979">Stepanjants 1979</bibRefCitation>
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, p 52
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</taxonomicName>
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, Plate 9
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<figureCitation id="BCD7CDA4600F6250FD4AFBD68EDB6C32" box="[759,912,1080,1104]" captionStart="Figure 5" captionStartId="17.[125,185,1450,1469]" captionTargetBox="[9,1028,140,1411]" captionTargetId="figure-21@17.[266,1032,135,1414]" captionTargetPageId="17" captionText="Figure 5. Acryptolaria corniformis Naumov and Stepanjants, 1962. (A) Branch fragment showing hydrothecal arrangement; (B, C) hydrothecae. All drawings from the holotype. Scale bar: 1 mm (A); 400 Mm (B, C)." figureDoi="http://doi.org/10.5281/zenodo.4582072" httpUri="https://zenodo.org/record/4582072/files/figure.png" pageId="31" pageNumber="258">Figure 5A, B</figureCitation>
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.
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</paragraph>
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||
<paragraph id="2453D121600F6250FFC0FBB789686C13" blockId="31.[125,1172,1014,1202]" box="[125,1059,1113,1137]" pageId="31" pageNumber="258">
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||
<taxonomicName id="E3ECAAA2600F6250FFC0FBB78FF26C13" ID-CoL="9RXG" authority="El Beshbeeshy 1991, p 67" authorityName="El Beshbeeshy" authorityPageNumber="67" authorityYear="1991" box="[125,697,1113,1137]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="31" pageNumber="258" phylum="Cnidaria" rank="species" species="patagonica">
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<emphasis id="16980D33600F6250FFC0FBB78CC86C13" box="[125,387,1113,1137]" italics="true" pageId="31" pageNumber="258">Acryptolaria patagonica</emphasis>
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||
<bibRefCitation id="407DACD0600F6250FE31FBB78F306C13" author="El Beshbeeshy M" box="[396,635,1113,1137]" pageId="31" pageNumber="258" refId="ref18937" refString="El Beshbeeshy M. 1991. Systematische, Morphologische und Zoogeographische Untersuchungen an den Thekaten Hydroiden des Patagonischen Schelfs [PhD thesis]. Hamburg: University of Hamburg. 390 p." type="book" year="1991">El Beshbeeshy 1991</bibRefCitation>
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, p 67
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</taxonomicName>
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–70,
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<figureCitation id="BCD7CDA4600F6250FD48FBB78E296C13" box="[757,866,1113,1137]" captionStart="Figure 14" captionStartId="39.[125,185,1589,1608]" captionTargetBox="[9,1091,137,1549]" captionTargetId="figure-0@39.[202,1096,133,1553]" captionTargetPageId="39" captionText="Figure 14. Acryptolaria tortugasensis Leloup, 1935. (A–F) Branch fragments showing shape and arrangement of hydrothecae. All drawings from the holotype. Scale bar: 1 mm (A); 400 Mm (B–F)." figureDoi="http://doi.org/10.5281/zenodo.4582098" httpUri="https://zenodo.org/record/4582098/files/figure.png" pageId="31" pageNumber="258">Figure 14</figureCitation>
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[
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<emphasis id="16980D33600F6250FCC8FBB7895A6C13" box="[885,1041,1113,1137]" italics="true" pageId="31" pageNumber="258">nomen nudum</emphasis>
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].
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</paragraph>
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<paragraph id="2453D121600F626FFFC0FB948C136CBD" blockId="31.[125,1172,1014,1202]" lastBlockId="32.[138,344,1223,1247]" lastPageId="32" lastPageNumber="259" pageId="31" pageNumber="258">
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||
<taxonomicName id="E3ECAAA2600F6250FFC0FB948F1F6CF0" ID-CoL="9RXG" authority="El Beshbeeshy" authorityName="El Beshbeeshy" authorityYear="1991" box="[125,596,1146,1170]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="31" pageNumber="258" phylum="Cnidaria" rank="species" species="patagonica">
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||
<emphasis id="16980D33600F6250FFC0FB948CC66CF0" box="[125,397,1146,1170]" italics="true" pageId="31" pageNumber="258">Acryptolaria patagonica</emphasis>
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||
El Beshbeeshy
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</taxonomicName>
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in
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<bibRefCitation id="407DACD0600F6250FD2EFB948E766CF0" author="Watson JW" box="[659,829,1146,1170]" pageId="31" pageNumber="258" pagination="151 - 180" refId="ref23225" refString="Watson JW. 2003. Deep-water hydroids (Hydrozoa: Leptolida) from Macquarie Island. Memoirs of Museum of Victoria 60: 151 - 180." type="journal article" year="2003">Watson 2003</bibRefCitation>
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, p 162–163,
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<figureCitation id="BCD7CDA4600F6250FC57FB94891E6CF0" box="[1002,1109,1146,1170]" captionStart="Figure 12" captionStartId="32.[138,198,1099,1118]" captionTargetBox="[314,1008,137,1059]" captionTargetId="figure-125@32.[310,1013,133,1063]" captionTargetPageId="32" captionText="Figure 12. Acryptolaria operculata Stepanjants, 1979. (A, B) Branch fragments showing shape and arrangement of hydrothecae. All drawings from the paratype. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4582092" httpUri="https://zenodo.org/record/4582092/files/figure.png" pageId="31" pageNumber="258">Figure 12</figureCitation>
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A–C;
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<bibRefCitation id="407DACD0600F6250FF25FB748C916CD0" author="Vervoort W & Watson JE" box="[152,474,1178,1202]" pageId="31" pageNumber="258" pagination="1 - 538" refId="ref23135" refString="Vervoort W, Watson JE. 2003. The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (Thecate Hydroids). NIWA Biodiversity Memoir 119: 1 - 538." type="journal article" year="2003">Vervoort and Watson 2003</bibRefCitation>
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, p 51–53,
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<figureCitation id="BCD7CDA4600F6250FDE9FB748FFA6CD0" box="[596,689,1178,1202]" captionStart="Figure 7" captionStartId="21.[125,185,1351,1370]" captionTargetBox="[9,1028,137,1310]" captionTargetId="figure-58@21.[266,1032,133,1315]" captionTargetPageId="21" captionText="Figure 7. Acryptolaria flabellum (Allman, 1888). (A, B) Branch fragments showing hydrothecal arrangement; (C, D) hydrothecae. All drawings from the holotype. Scale bar: 1 mm (A, B); 400 Mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.4582078" httpUri="https://zenodo.org/record/4582078/files/figure.png" pageId="31" pageNumber="258">Figure 7</figureCitation>
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A–G.
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<emphasis id="16980D336030626FFF37FB298C136CBD" box="[138,344,1223,1247]" italics="true" pageId="32" pageNumber="259">Material examined</emphasis>
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||
</paragraph>
|
||
<caption id="709381A9600F6250FFC0FAE58ED46D7C" box="[125,927,1289,1310]" pageId="31" pageNumber="258">
|
||
<paragraph id="2453D121600F6250FFC0FAE58ED46D7C" blockId="31.[125,927,1289,1310]" box="[125,927,1289,1310]" pageId="31" pageNumber="258">
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||
Table XII. Morphometric data of the type material of
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||
<taxonomicName id="E3ECAAA2600F6250FDC2FAE58E026D7C" ID-CoL="9RXF" authorityName="Stepanjants" authorityYear="1979" box="[639,841,1291,1310]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="31" pageNumber="258" phylum="Cnidaria" rank="species" species="operculata">
|
||
<emphasis id="16980D33600F6250FDC2FAE58E026D7C" box="[639,841,1291,1310]" italics="true" pageId="31" pageNumber="258">Acryptolaria operculata</emphasis>
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||
</taxonomicName>
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(in Mm).
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||
</paragraph>
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||
</caption>
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||
<paragraph id="2453D121600F6250FD11FAD0893F6EE1" pageId="31" pageNumber="258">
|
||
<table id="56EC2381600F9DB0FFC0FAD0893F6EE1" box="[125,1140,1342,1667]" colsContinueFrom="30.[138,1157,196,495]" colsContinueIn="34.[138,1154,1341,1666]" gridcols="4" gridrows="12" pageId="31" pageNumber="258">
|
||
<tr id="9ADCD363600F9DB0FFC0FAD0893F6D30" box="[125,1140,1342,1362]" gridrow="0" pageId="31" pageNumber="258" rowspan-0="1">
|
||
<th id="D90DBA1F600F9DB0FDC8FAD08E056D30" box="[629,846,1342,1362]" gridcol="1" gridrow="0" pageId="31" pageNumber="258">Mean¡SD</th>
|
||
<th id="D90DBA1F600F9DB0FC3BFAD0896D6D30" box="[902,1062,1342,1362]" gridcol="2" gridrow="0" pageId="31" pageNumber="258">Range</th>
|
||
<th id="D90DBA1F600F9DB0FBE1FAD0893F6D30" box="[1116,1140,1342,1362]" gridcol="3" gridrow="0" pageId="31" pageNumber="258">
|
||
<emphasis id="16980D33600F6250FBDFFAD089266D33" box="[1122,1133,1342,1361]" italics="true" pageId="31" pageNumber="258">n</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="9ADCD363600F9DB0FFC0FA88893F6D1B" box="[125,1140,1382,1401]" gridrow="1" pageId="31" pageNumber="258">
|
||
<th id="D90DBA1F600F9DB0FFC0FA88893F6D1B" box="[125,1140,1382,1401]" colspan="4" colspanRight="3" gridcol="0" gridrow="1" pageId="31" pageNumber="258">Hydrothecae</th>
|
||
</tr>
|
||
<tr id="9ADCD363600F9DB0FFC0FA6E893F6DF6" box="[125,1140,1408,1428]" gridrow="2" pageId="31" pageNumber="258">
|
||
<th id="D90DBA1F600F9DB0FFC0FA6E8CF56DF6" box="[125,446,1408,1428]" gridcol="0" gridrow="2" pageId="31" pageNumber="258">Length of abcauline wall</th>
|
||
<td id="D90DBA1F600F9DB0FDC8FA6E8E056DF6" box="[629,846,1408,1428]" gridcol="1" gridrow="2" pageId="31" pageNumber="258">2000.0¡141.4</td>
|
||
<td id="D90DBA1F600F9DB0FC3BFA6E896D6DF6" box="[902,1062,1408,1428]" gridcol="2" gridrow="2" pageId="31" pageNumber="258">1800–2100</td>
|
||
<td id="D90DBA1F600F9DB0FBE1FA6E893F6DF6" box="[1116,1140,1408,1428]" gridcol="3" gridrow="2" pageId="31" pageNumber="258">4</td>
|
||
</tr>
|
||
<tr id="9ADCD363600F9DB0FFC0FA75893F6DCD" box="[125,1140,1435,1455]" gridrow="3" pageId="31" pageNumber="258">
|
||
<th id="D90DBA1F600F9DB0FFC0FA758CF56DCD" box="[125,446,1435,1455]" gridcol="0" gridrow="3" pageId="31" pageNumber="258">Length of free adcauline wall</th>
|
||
<td id="D90DBA1F600F9DB0FDC8FA758E056DCD" box="[629,846,1435,1455]" gridcol="1" gridrow="3" pageId="31" pageNumber="258">987.5¡84.2</td>
|
||
<td id="D90DBA1F600F9DB0FC3BFA75896D6DCD" box="[902,1062,1435,1455]" gridcol="2" gridrow="3" pageId="31" pageNumber="258">910–1070</td>
|
||
<td id="D90DBA1F600F9DB0FBE1FA75893F6DCD" box="[1116,1140,1435,1455]" gridcol="3" gridrow="3" pageId="31" pageNumber="258">4</td>
|
||
</tr>
|
||
<tr id="9ADCD363600F9DB0FFC0FA5B893F6DAB" box="[125,1140,1461,1481]" gridrow="4" pageId="31" pageNumber="258">
|
||
<th id="D90DBA1F600F9DB0FFC0FA5B8CF56DAB" box="[125,446,1461,1481]" gridcol="0" gridrow="4" pageId="31" pageNumber="258">Length of adnate adcauline wall</th>
|
||
<td id="D90DBA1F600F9DB0FDC8FA5B8E056DAB" box="[629,846,1461,1481]" gridcol="1" gridrow="4" pageId="31" pageNumber="258">1525.0¡150.0</td>
|
||
<td id="D90DBA1F600F9DB0FC3BFA5B896D6DAB" box="[902,1062,1461,1481]" gridcol="2" gridrow="4" pageId="31" pageNumber="258">1400–1700</td>
|
||
<td id="D90DBA1F600F9DB0FBE1FA5B893F6DAB" box="[1116,1140,1461,1481]" gridcol="3" gridrow="4" pageId="31" pageNumber="258">4</td>
|
||
</tr>
|
||
<tr id="9ADCD363600F9DB0FFC0FA3E893F6D86" box="[125,1140,1488,1508]" gridrow="5" pageId="31" pageNumber="258">
|
||
<th id="D90DBA1F600F9DB0FFC0FA3E8CF56D86" box="[125,446,1488,1508]" gridcol="0" gridrow="5" pageId="31" pageNumber="258">Length of adcauline wall</th>
|
||
<td id="D90DBA1F600F9DB0FDC8FA3E8E056D86" box="[629,846,1488,1508]" gridcol="1" gridrow="5" pageId="31" pageNumber="258">2512.5¡158.4</td>
|
||
<td id="D90DBA1F600F9DB0FC3BFA3E896D6D86" box="[902,1062,1488,1508]" gridcol="2" gridrow="5" pageId="31" pageNumber="258">2320–2670</td>
|
||
<td id="D90DBA1F600F9DB0FBE1FA3E893F6D86" box="[1116,1140,1488,1508]" gridcol="3" gridrow="5" pageId="31" pageNumber="258">4</td>
|
||
</tr>
|
||
<tr id="9ADCD363600F9DB0FFC0FA05893F6D9C" box="[125,1140,1515,1534]" gridrow="6" pageId="31" pageNumber="258" rowspan-1="1" rowspan-3="1">
|
||
<th id="D90DBA1F600F9DB0FFC0FA058CF56D9C" box="[125,446,1515,1534]" gridcol="0" gridrow="6" pageId="31" pageNumber="258">Ratio adnate/free adcauline wall</th>
|
||
<td id="D90DBA1F600F9DB0FC3BFA05896D6D9C" box="[902,1062,1515,1534]" gridcol="2" gridrow="6" pageId="31" pageNumber="258">1.3–1.9</td>
|
||
</tr>
|
||
<tr id="9ADCD363600F9DB0FFC0F9EB893F6E7B" box="[125,1140,1541,1561]" gridrow="7" pageId="31" pageNumber="258">
|
||
<th id="D90DBA1F600F9DB0FFC0F9EB8CF56E7B" box="[125,446,1541,1561]" gridcol="0" gridrow="7" pageId="31" pageNumber="258">Diameter at aperture</th>
|
||
<td id="D90DBA1F600F9DB0FDC8F9EB8E056E7B" box="[629,846,1541,1561]" gridcol="1" gridrow="7" pageId="31" pageNumber="258">412.0¡21.7</td>
|
||
<td id="D90DBA1F600F9DB0FC3BF9EB896D6E7B" box="[902,1062,1541,1561]" gridcol="2" gridrow="7" pageId="31" pageNumber="258">400–450</td>
|
||
<td id="D90DBA1F600F9DB0FBE1F9EB893F6E7B" box="[1116,1140,1541,1561]" gridcol="3" gridrow="7" pageId="31" pageNumber="258">5</td>
|
||
</tr>
|
||
<tr id="9ADCD363600F9DB0FFC0F9CE893F6E56" box="[125,1140,1568,1588]" gridrow="8" pageId="31" pageNumber="258">
|
||
<th id="D90DBA1F600F9DB0FFC0F9CE8CF56E56" box="[125,446,1568,1588]" gridcol="0" gridrow="8" pageId="31" pageNumber="258">Diameter at base</th>
|
||
<td id="D90DBA1F600F9DB0FDC8F9CE8E056E56" box="[629,846,1568,1588]" gridcol="1" gridrow="8" pageId="31" pageNumber="258">206.7¡5.8</td>
|
||
<td id="D90DBA1F600F9DB0FC3BF9CE896D6E56" box="[902,1062,1568,1588]" gridcol="2" gridrow="8" pageId="31" pageNumber="258">200–210</td>
|
||
<td id="D90DBA1F600F9DB0FBE1F9CE893F6E56" box="[1116,1140,1568,1588]" gridcol="3" gridrow="8" pageId="31" pageNumber="258">3</td>
|
||
</tr>
|
||
<tr id="9ADCD363600F9DB0FFC0F9D5893F6E2C" box="[125,1140,1595,1614]" gridrow="9" pageId="31" pageNumber="258">
|
||
<th id="D90DBA1F600F9DB0FFC0F9D5893F6E2C" box="[125,1140,1595,1614]" colspan="4" colspanRight="3" gridcol="0" gridrow="9" pageId="31" pageNumber="258">Nematocysts</th>
|
||
</tr>
|
||
<tr id="9ADCD363600F9DB0FFC0F9BB893F6E0B" box="[125,1140,1621,1641]" gridrow="10" pageId="31" pageNumber="258">
|
||
<th id="D90DBA1F600F9DB0FFC0F9BB8CF56E0B" box="[125,446,1621,1641]" gridcol="0" gridrow="10" pageId="31" pageNumber="258">Larger group</th>
|
||
<td id="D90DBA1F600F9DB0FDC8F9BB8E056E0B" box="[629,846,1621,1641]" gridcol="1" gridrow="10" pageId="31" pageNumber="258">14.1¡0.5X5.0¡0.2</td>
|
||
<td id="D90DBA1F600F9DB0FC3BF9BB896D6E0B" box="[902,1062,1621,1641]" gridcol="2" gridrow="10" pageId="31" pageNumber="258">13.5–15X4.5–5</td>
|
||
<td id="D90DBA1F600F9DB0FBE1F9BB893F6E0B" box="[1116,1140,1621,1641]" gridcol="3" gridrow="10" pageId="31" pageNumber="258">10</td>
|
||
</tr>
|
||
<tr id="9ADCD363600F9DB0FFC0F981893F6EE1" box="[125,1140,1647,1667]" gridrow="11" pageId="31" pageNumber="258">
|
||
<th id="D90DBA1F600F9DB0FFC0F9818CF56EE1" box="[125,446,1647,1667]" gridcol="0" gridrow="11" pageId="31" pageNumber="258">Smaller group</th>
|
||
<td id="D90DBA1F600F9DB0FDC8F9818E056EE1" box="[629,846,1647,1667]" gridcol="1" gridrow="11" pageId="31" pageNumber="258">8.0¡0.4X3.1¡0.2</td>
|
||
<td id="D90DBA1F600F9DB0FC3BF981896D6EE1" box="[902,1062,1647,1667]" gridcol="2" gridrow="11" pageId="31" pageNumber="258">7.5–8.5X3–3.5</td>
|
||
<td id="D90DBA1F600F9DB0FBE1F981893F6EE1" box="[1116,1140,1647,1667]" gridcol="3" gridrow="11" pageId="31" pageNumber="258">5</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="709381A96030626FFF37FBA58F9E6C1B" ID-DOI="http://doi.org/10.5281/zenodo.4582092" ID-Zenodo-Dep="4582092" httpUri="https://zenodo.org/record/4582092/files/figure.png" pageId="32" pageNumber="259" startId="32.[138,198,1099,1118]" targetBox="[314,1008,137,1059]" targetPageId="32">
|
||
<paragraph id="2453D1216030626FFF37FBA58F9E6C1B" blockId="32.[138,1185,1099,1145]" pageId="32" pageNumber="259">
|
||
Figure 12.
|
||
<taxonomicName id="E3ECAAA26030626FFF48FBA58F266C3C" ID-CoL="9RXF" authority="Stepanjants, 1979" authorityName="Stepanjants" authorityYear="1979" box="[245,621,1099,1118]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="32" pageNumber="259" phylum="Cnidaria" rank="species" species="operculata">
|
||
<emphasis id="16980D336030626FFF48FBA58CF56C3C" box="[245,446,1099,1118]" italics="true" pageId="32" pageNumber="259">Acryptolaria operculata</emphasis>
|
||
<bibRefCitation id="407DACD06030626FFE79FBA58F266C3C" author="Stepanjants SD" box="[452,621,1099,1118]" pageId="32" pageNumber="259" pagination="1 - 200" refId="ref22474" refString="Stepanjants SD. 1979. Hydroids of the antarctic and subantarctic waters. In: Biological results of the Soviet Antarctic Expeditions, 6. Issledovaniya Fauny Morei 20: 1 - 200. (Rus)." type="journal article" year="1979">Stepanjants, 1979</bibRefCitation>
|
||
</taxonomicName>
|
||
. (A, B) Branch fragments showing shape and arrangement of hydrothecae. All drawings from the paratype. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2453D1216030626FFF37FB168E3D6D52" blockId="32.[138,1185,1272,1393]" pageId="32" pageNumber="259">
|
||
<materialsCitation id="9484DB7C6030626FFF37FB168E386D52" collectingDate="1974-05-27" collectionCode="ZIRAS" pageId="32" pageNumber="259" specimenCode="ZIRAS 2" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="FB576F836030626FFF37FB168DBF6D72" box="[138,244,1272,1296]" pageId="32" pageNumber="259" type="paratype">Paratype</typeStatus>
|
||
:
|
||
<specimenCode id="744A795A6030626FFEBCFB168C206D72" box="[257,363,1272,1296]" collectionCode="ZIRAS" pageId="32" pageNumber="259">ZIRAS 2</specimenCode>
|
||
/9463,
|
||
<emphasis id="16980D336030626FFE7DFB168CB16D72" box="[448,506,1272,1296]" italics="true" pageId="32" pageNumber="259">Zund</emphasis>
|
||
Sta. 194, 43
|
||
<emphasis id="16980D336030626FFD2BFB168FEB6D72" bold="true" box="[662,672,1272,1296]" italics="true" pageId="32" pageNumber="259">°</emphasis>
|
||
569 S, 59
|
||
<emphasis id="16980D336030626FFCB7FB168E5F6D72" bold="true" box="[778,788,1272,1296]" italics="true" pageId="32" pageNumber="259">°</emphasis>
|
||
089 W,
|
||
<date id="5052F7E16030626FFCD6FB1689446D72" box="[875,1039,1272,1296]" pageId="32" pageNumber="259" value="1974-05-27">
|
||
<collectingDate id="40160E096030626FFCD6FB1689446D72" box="[875,1039,1272,1296]" pageId="32" pageNumber="259" value="1974-05-27">
|
||
27 May
|
||
<quantity id="E3147CC46030626FFC6CFB1689446D72" box="[977,1039,1272,1296]" metricMagnitude="6" metricUnit="m" metricValue="1.97414" pageId="32" pageNumber="259" unit="m" value="1974140.0">1974</quantity>
|
||
</collectingDate>
|
||
</date>
|
||
, 140 m, one basally broken stem fragment ca
|
||
<quantity id="E3147CC46030626FFDB0FAF68F166D52" box="[525,605,1304,1328]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="32" pageNumber="259" unit="mm" value="40.0">40 mm</quantity>
|
||
long, without coppinia
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6CF682AA6030626DFF18FAD78C406992" lastPageId="34" lastPageNumber="261" pageId="32" pageNumber="259" type="description">
|
||
<paragraph id="2453D1216030626FFF18FAD78F226D13" blockId="32.[138,1185,1272,1393]" pageId="32" pageNumber="259">
|
||
<taxonomicName id="E3ECAAA26030626FFF18FAD78CE16D33" authorityName="El Beshbeeshy" authorityYear="1991" box="[165,426,1337,1361]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="32" pageNumber="259" phylum="Cnidaria" rank="species" species="patagonica">
|
||
<emphasis id="16980D336030626FFF18FAD78CE16D33" box="[165,426,1337,1361]" italics="true" pageId="32" pageNumber="259">Acryptolaria patagonica</emphasis>
|
||
</taxonomicName>
|
||
, one slide from Sta. 330, 40
|
||
<emphasis id="16980D336030626FFD4BFAD78E4B6D33" bold="true" box="[758,768,1337,1361]" italics="true" pageId="32" pageNumber="259">°</emphasis>
|
||
579 S, 56
|
||
<emphasis id="16980D336030626FFCD5FAD78E396D33" bold="true" box="[872,882,1337,1361]" italics="true" pageId="32" pageNumber="259">°</emphasis>
|
||
579 W,
|
||
<date id="5052F7E16030626FFC7AFAD789D76D33" box="[967,1180,1337,1361]" pageId="32" pageNumber="259" value="1971-02-23">
|
||
23 February
|
||
<quantity id="E3147CC46030626FFBE3FAD789D76D33" box="[1118,1180,1337,1361]" metricMagnitude="6" metricUnit="m" metricValue="1.9719799999999998" pageId="32" pageNumber="259" unit="m" value="1971980.0">1971</quantity>
|
||
</date>
|
||
, 980 m (Walter Herwig expedition 1971).
|
||
</paragraph>
|
||
<paragraph id="2453D1216030626FFF37FA448C486DA0" blockId="32.[138,259,1450,1474]" box="[138,259,1450,1474]" pageId="32" pageNumber="259">
|
||
<emphasis id="16980D336030626FFF37FA448C486DA0" box="[138,259,1450,1474]" italics="true" pageId="32" pageNumber="259">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="2453D1216030626FFF37FA358FC76E71" blockId="32.[138,1185,1499,1685]" pageId="32" pageNumber="259">
|
||
Stem fragment ca
|
||
<quantity id="E3147CC46030626FFECAFA358C8C6D91" box="[375,455,1499,1523]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="32" pageNumber="259" unit="mm" value="40.0">40 mm</quantity>
|
||
long, basally broken. No information concerning habit is available; other than its irregular branching.
|
||
</paragraph>
|
||
<paragraph id="2453D1216030626EFF18F9F28E6768A3" blockId="32.[138,1185,1499,1685]" lastBlockId="33.[125,1173,136,226]" lastPageId="33" lastPageNumber="260" pageId="32" pageNumber="259">
|
||
Hydrothecae alternate, in two planes, making an obtuse angle. Hydrotheca tubular, fusiform; maximum diameter approximately in the middle of hydrothecal length, decreasing distinctly towards basal part. Hydrotheca curved abcaulinarly, half to twothirds of adcauline wall adnate to internode. Adcauline wall convex; abcauline wall concave. Hydrothecal aperture circular, directed upwards, forming an angle of 55–75
|
||
<emphasis id="16980D336031626EFBECFF66891068C2" bold="true" box="[1105,1115,136,160]" italics="true" pageId="33" pageNumber="260">°</emphasis>
|
||
with long axis of branch. Rim even, with few short renovations.
|
||
</paragraph>
|
||
<paragraph id="2453D1216031626EFF25FF248CE06880" blockId="33.[125,1173,136,226]" box="[152,427,202,226]" pageId="33" pageNumber="260">Coppinia not observed.</paragraph>
|
||
<paragraph id="2453D1216031626EFFC0FEF58D976951" blockId="33.[125,220,283,307]" box="[125,220,283,307]" pageId="33" pageNumber="260">
|
||
<emphasis id="16980D336031626EFFC0FEF58D976951" box="[125,220,283,307]" italics="true" pageId="33" pageNumber="260">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="2453D1216031626EFFC0FEA58EE669A7" blockId="33.[125,1173,331,1685]" pageId="33" pageNumber="260">
|
||
<taxonomicName id="E3ECAAA26031626EFFC0FEA58C326901" authorityName="Stepanjants" authorityYear="1979" box="[125,377,331,355]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="33" pageNumber="260" phylum="Cnidaria" rank="species" species="operculata">
|
||
<emphasis id="16980D336031626EFFC0FEA58C326901" box="[125,377,331,355]" italics="true" pageId="33" pageNumber="260">Acryptolaria operculata</emphasis>
|
||
</taxonomicName>
|
||
is characterized by the size and arrangement of the hydrothecae. It is the species with the largest hydrothecae (cf. Table XV) and it is also characterized by the hydrothecal arrangement in two planes, making an obtuse angle, though this feature is also present in other species of the genus (e.g.
|
||
<taxonomicName id="E3ECAAA26031626EFDD3FE438E5169A7" authorityName="Leloup" authorityYear="1935" box="[622,794,429,453]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="33" pageNumber="260" phylum="Cnidaria" rank="species" species="tortugasensis">
|
||
<emphasis id="16980D336031626EFDD3FE438E5169A7" box="[622,794,429,453]" italics="true" pageId="33" pageNumber="260">A. tortugasensis</emphasis>
|
||
</taxonomicName>
|
||
, see below).
|
||
</paragraph>
|
||
<paragraph id="2453D1216031626EFF25FE208F3B6B6B" blockId="33.[125,1173,331,1685]" pageId="33" pageNumber="260">
|
||
<bibRefCitation id="407DACD06031626EFF25FE208C346984" author="Stepanjants SD" box="[152,383,462,486]" pageId="33" pageNumber="260" pagination="1 - 200" refId="ref22474" refString="Stepanjants SD. 1979. Hydroids of the antarctic and subantarctic waters. In: Biological results of the Soviet Antarctic Expeditions, 6. Issledovaniya Fauny Morei 20: 1 - 200. (Rus)." type="journal article" year="1979">Stepanjants (1979)</bibRefCitation>
|
||
also characterized this species by the presence of a one-flap operculum, pointing out the particular position of this species amongst the
|
||
<taxonomicName id="E3ECAAA26031626EFBA6FE0089DA6A64" authorityName="Hincks" authorityYear="1868" box="[1051,1169,494,518]" class="Hydrozoa" family="Lafoeidae" kingdom="Animalia" order="Leptothecata" pageId="33" pageNumber="260" phylum="Cnidaria" rank="family">Lafoeidae</taxonomicName>
|
||
. However, after studying many colonies of
|
||
<taxonomicName id="E3ECAAA26031626EFDDBFDE08FA66A44" authorityName="Norman" authorityYear="1875" box="[614,749,526,550]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="33" pageNumber="260" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="16980D336031626EFDDBFDE08FA66A44" box="[614,749,526,550]" italics="true" pageId="33" pageNumber="260">Acryptolaria</emphasis>
|
||
</taxonomicName>
|
||
, we have observed that the presence of that circular flap, in spite of being irregular, is quite common amongst species of
|
||
<taxonomicName id="E3ECAAA26031626EFFC0FDA18C486A05" authorityName="Norman" authorityYear="1875" box="[125,259,591,615]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="33" pageNumber="260" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="16980D336031626EFFC0FDA18C486A05" box="[125,259,591,615]" italics="true" pageId="33" pageNumber="260">Acryptolaria</emphasis>
|
||
</taxonomicName>
|
||
, being also common in species of another lafoeid genus,
|
||
<taxonomicName id="E3ECAAA26031626EFC31FDA189466A05" authorityName="Stimpson" authorityYear="1853" box="[908,1037,591,615]" class="Hydrozoa" family="Lafoeidae" genus="Grammaria" kingdom="Animalia" order="Leptothecata" pageId="33" pageNumber="260" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="16980D336031626EFC31FDA189466A05" box="[908,1037,591,615]" italics="true" pageId="33" pageNumber="260">Grammaria</emphasis>
|
||
</taxonomicName>
|
||
. That valve should not be considered as an opercular apparatus in the sense of that present in other families of hydroids (e.g.
|
||
<taxonomicName id="E3ECAAA26031626EFE02FD7E8F196ACA" authorityName="Lamouroux" authorityYear="1812" box="[447,594,656,680]" class="Hydrozoa" family="Sertulariidae" kingdom="Animalia" order="Leptothecata" pageId="33" pageNumber="260" phylum="Cnidaria" rank="family">Sertulariidae</taxonomicName>
|
||
or
|
||
<taxonomicName id="E3ECAAA26031626EFD35FD7E8E046ACA" authorityName="Hincks" authorityYear="1868" box="[648,847,656,680]" class="Hydrozoa" family="Campanulinidae" kingdom="Animalia" order="Leptothecata" pageId="33" pageNumber="260" phylum="Cnidaria" rank="family">Campanulinidae</taxonomicName>
|
||
). By contrast, it seems to constitute a sort of deciduous structure to temporally protect the polyps when they are in a state of inactivity or regeneration, as has already postulated for other groups such as
|
||
<taxonomicName id="E3ECAAA26031626EFFC0FD1F8DB56B6B" authorityName="Allman" authorityYear="1888" box="[125,254,753,777]" class="Hydrozoa" family="Sertulariidae" genus="Staurotheca" kingdom="Animalia" order="Leptothecata" pageId="33" pageNumber="260" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="16980D336031626EFFC0FD1F8DB56B6B" box="[125,254,753,777]" italics="true" pageId="33" pageNumber="260">Staurotheca</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<bibRefCitation id="407DACD06031626EFE8AFD1F8F286B6B" author="Pena Cantero AL & Svoboda A & Vervoort W" box="[311,611,753,777]" pageId="33" pageNumber="260" pagination="329 - 381" refId="ref21527" refString="Pena Cantero AL, Svoboda A, Vervoort W. 1997. Species of Staurotheca Allman, 1888 (Cnidaria: Hydrozoa) from recent antarctic expeditions with R. V. Polarstern, with the description of six new species. Journal of Natural History 31: 329 - 381." type="journal article" year="1997">Peña Cantero et al. 1997</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2453D1216031626EFF25FCFC895E6C0C" blockId="33.[125,1173,331,1685]" pageId="33" pageNumber="260">
|
||
In his PhD dissertation on hydrozoans from the Patagonian shelf,
|
||
<bibRefCitation id="407DACD06031626EFC2BFCFC89DF6B48" author="El Beshbeeshy M" box="[918,1172,786,810]" pageId="33" pageNumber="260" refId="ref18937" refString="El Beshbeeshy M. 1991. Systematische, Morphologische und Zoogeographische Untersuchungen an den Thekaten Hydroiden des Patagonischen Schelfs [PhD thesis]. Hamburg: University of Hamburg. 390 p." type="book" year="1991">El Beshbeeshy (1991)</bibRefCitation>
|
||
included the description of 22 new species, as well as many re-descriptions and some nomenclatural considerations. Many subsequent papers (e.g.
|
||
<bibRefCitation id="407DACD06031626EFCE1FCBD89DB6B09" author="Pena Cantero AL & Svoboda A & Vervoort W" box="[860,1168,851,875]" pageId="33" pageNumber="260" pagination="329 - 381" refId="ref21527" refString="Pena Cantero AL, Svoboda A, Vervoort W. 1997. Species of Staurotheca Allman, 1888 (Cnidaria: Hydrozoa) from recent antarctic expeditions with R. V. Polarstern, with the description of six new species. Journal of Natural History 31: 329 - 381." type="journal article" year="1997">Peña Cantero et al. 1997</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="407DACD06031626EFFC0FC9D8CB46BE9" author="Pena Cantero AL & Marques AC" box="[125,511,883,907]" pageId="33" pageNumber="260" pagination="83 - 93" refId="ref21491" refString="Pena Cantero AL, Marques AC. 1999. Phylogenetic analysis of the Antarctic genus Osaealdella Stechow, 1919 (Hydrozoa, Leptomedusae, Kirchenpaueriidae). Contributions to Zoology 68: 83 - 93." type="journal article" year="1999">Peña Cantero and Marques 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="407DACD06031626EFDB4FC9D8E0F6BE9" author="Vervoort W & Watson JE" box="[521,836,883,907]" pageId="33" pageNumber="260" pagination="1 - 538" refId="ref23135" refString="Vervoort W, Watson JE. 2003. The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (Thecate Hydroids). NIWA Biodiversity Memoir 119: 1 - 538." type="journal article" year="2003">Vervoort and Watson 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="407DACD06031626EFCF2FC9D8EA16BE9" author="Watson JW" box="[847,1002,883,907]" pageId="33" pageNumber="260" pagination="151 - 180" refId="ref23225" refString="Watson JW. 2003. Deep-water hydroids (Hydrozoa: Leptolida) from Macquarie Island. Memoirs of Museum of Victoria 60: 151 - 180." type="journal article" year="2003">Watson 2003</bibRefCitation>
|
||
) have used the names employed by
|
||
<bibRefCitation id="407DACD06031626EFED1FC7A8F256BCE" author="El Beshbeeshy M" box="[364,622,916,940]" pageId="33" pageNumber="260" refId="ref18937" refString="El Beshbeeshy M. 1991. Systematische, Morphologische und Zoogeographische Untersuchungen an den Thekaten Hydroiden des Patagonischen Schelfs [PhD thesis]. Hamburg: University of Hamburg. 390 p." type="book" year="1991">El Beshbeeshy (1991)</bibRefCitation>
|
||
. However, according to the International Code of
|
||
<bibRefCitation id="407DACD06031626EFF27FC5A8F526BAE" author="International Commission on Zoological Nomenclature" box="[154,537,948,972]" pageId="33" pageNumber="260" refId="ref19880" refString="International Commission on Zoological Nomenclature. 1999. International code of zoological nomenclature. 4 th ed. London: International Trust for Zoological Nomenclature. 306 p." type="book" year="1999">Zoological Nomenclature (1999)</bibRefCitation>
|
||
, a thesis is not to be taken as published for the purposes of zoological nomenclature, and therefore the specific names proposed by
|
||
<bibRefCitation id="407DACD06031626EFC56FC3A8D866C6F" author="El Beshbeeshy M" pageId="33" pageNumber="260" refId="ref18937" refString="El Beshbeeshy M. 1991. Systematische, Morphologische und Zoogeographische Untersuchungen an den Thekaten Hydroiden des Patagonischen Schelfs [PhD thesis]. Hamburg: University of Hamburg. 390 p." type="book" year="1991">El Beshbeeshy (1991)</bibRefCitation>
|
||
should be considered
|
||
<emphasis id="16980D336031626EFE6EFC1B8F2F6C6F" box="[467,612,1013,1037]" italics="true" pageId="33" pageNumber="260">nomina nuda</emphasis>
|
||
. According to the Code ‘‘A
|
||
<emphasis id="16980D336031626EFC0CFC1B89076C6F" box="[945,1100,1013,1037]" italics="true" pageId="33" pageNumber="260">nomen nudum</emphasis>
|
||
is not an available name, and therefore the same name may be made available later for the same or a different concept; in such a case it would take authorship and date [Arts. 50, 21] from that act of establishment, not from any earlier publication as a
|
||
<emphasis id="16980D336031626EFCDFFBB98EB56C0C" box="[866,1022,1110,1134]" italics="true" pageId="33" pageNumber="260">nomen nudum</emphasis>
|
||
’’.
|
||
</paragraph>
|
||
<paragraph id="2453D1216031626DFF25FB998FBC6882" blockId="33.[125,1173,331,1685]" lastBlockId="34.[138,1185,136,416]" lastPageId="34" lastPageNumber="261" pageId="33" pageNumber="260">
|
||
Indeed, many nominal species of El Beshbeeshy were subsequently described by other authors and, in this case, these should become the authors of the valid names. Nevertheless, Article 50.1.1. of the Code establishes that ‘‘however, if it is clear from the contents that some person other than an author of the work is alone responsible both for the name or act and for satisfying the criteria of availability other than actual publication, then that other person is the author of the name or act. If the identity of that other person is not explicit in the work itself, then the author is deemed to be the person who publishes the work’’.
|
||
<bibRefCitation id="407DACD06031626EFBC6FAD78C116D13" author="El Beshbeeshy M" pageId="33" pageNumber="260" refId="ref18937" refString="El Beshbeeshy M. 1991. Systematische, Morphologische und Zoogeographische Untersuchungen an den Thekaten Hydroiden des Patagonischen Schelfs [PhD thesis]. Hamburg: University of Hamburg. 390 p." type="book" year="1991">El Beshbeeshy (1991)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="E3ECAAA26031626EFE6BFAB78F256D13" authorityName="El Beshbeeshy" authorityYear="1991" box="[470,622,1369,1393]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="33" pageNumber="260" phylum="Cnidaria" rank="species" species="patagonica">
|
||
<emphasis id="16980D336031626EFE6BFAB78F256D13" box="[470,622,1369,1393]" italics="true" pageId="33" pageNumber="260">A. patagonica</emphasis>
|
||
</taxonomicName>
|
||
as a new species; the species was again reported by
|
||
<bibRefCitation id="407DACD06031626EFF18FA948C136DF0" author="Watson JW" box="[165,344,1402,1426]" pageId="33" pageNumber="260" pagination="151 - 180" refId="ref23225" refString="Watson JW. 2003. Deep-water hydroids (Hydrozoa: Leptolida) from Macquarie Island. Memoirs of Museum of Victoria 60: 151 - 180." type="journal article" year="2003">Watson (2003)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="407DACD06031626EFE26FA948FB16DF0" author="Vervoort W & Watson JE" box="[411,762,1402,1426]" pageId="33" pageNumber="260" pagination="1 - 538" refId="ref23135" refString="Vervoort W, Watson JE. 2003. The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (Thecate Hydroids). NIWA Biodiversity Memoir 119: 1 - 538." type="journal article" year="2003">Vervoort and Watson (2003)</bibRefCitation>
|
||
, and both papers clearly gave the authority of the species to El Beshbeeshy. Therefore, the species is herein assumed as ‘‘
|
||
<taxonomicName id="E3ECAAA26031626EFF30FA558F116DB1" authority="El Beshbeeshy" authorityName="El Beshbeeshy" authorityYear="1991" box="[141,602,1467,1491]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="33" pageNumber="260" phylum="Cnidaria" rank="species" species="patagonica">
|
||
<emphasis id="16980D336031626EFF30FA558CD16DB1" box="[141,410,1467,1491]" italics="true" pageId="33" pageNumber="260">Acryptolaria patagonica</emphasis>
|
||
El Beshbeeshy
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="407DACD06031626EFD2CFA558E0F6DB1" author="Watson JW" box="[657,836,1467,1491]" pageId="33" pageNumber="260" pagination="151 - 180" refId="ref23225" refString="Watson JW. 2003. Deep-water hydroids (Hydrozoa: Leptolida) from Macquarie Island. Memoirs of Museum of Victoria 60: 151 - 180." type="journal article" year="2003">Watson, 2003</bibRefCitation>
|
||
’’. In any case, the species nominally described as
|
||
<taxonomicName id="E3ECAAA26031626EFE2CFA358F606D91" authorityName="El Beshbeeshy" authorityYear="1991" box="[401,555,1499,1523]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="33" pageNumber="260" phylum="Cnidaria" rank="species" species="patagonica">
|
||
<emphasis id="16980D336031626EFE2CFA358F606D91" box="[401,555,1499,1523]" italics="true" pageId="33" pageNumber="260">A. patagonica</emphasis>
|
||
</taxonomicName>
|
||
would be characterized by its large hydrothecae and the arrangement of the hydrothecae in two planes, forming an obtuse angle. According to
|
||
<bibRefCitation id="407DACD06031626EFFC0F9F28CD96E56" author="El Beshbeeshy M" box="[125,402,1564,1588]" pageId="33" pageNumber="260" refId="ref18937" refString="El Beshbeeshy M. 1991. Systematische, Morphologische und Zoogeographische Untersuchungen an den Thekaten Hydroiden des Patagonischen Schelfs [PhD thesis]. Hamburg: University of Hamburg. 390 p." type="book" year="1991">El Beshbeeshy (1991)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="E3ECAAA26031626EFE14F9F28F076E56" authorityName="El Beshbeeshy" authorityYear="1991" box="[425,588,1564,1588]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="33" pageNumber="260" phylum="Cnidaria" rank="species" species="patagonica">
|
||
<emphasis id="16980D336031626EFE14F9F28F076E56" box="[425,588,1564,1588]" italics="true" pageId="33" pageNumber="260">A. patagonica</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="E3ECAAA26031626EFCBDF9F28ED76E56" authorityName="Stepanjants" authorityYear="1979" box="[768,924,1564,1588]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="33" pageNumber="260" phylum="Cnidaria" rank="species" species="operculata">
|
||
<emphasis id="16980D336031626EFCBDF9F28ED76E56" box="[768,924,1564,1588]" italics="true" pageId="33" pageNumber="260">A. operculata</emphasis>
|
||
</taxonomicName>
|
||
by the absence of operculum, the shorter free part of the adcauline wall (672–858 Mm), and the larger diameter at hydrothecal base (232–324 Mm). Nevertheless, we have re-examined some slides of the material described by El Beshbeeshy and there are no differences between it and the
|
||
<typeStatus id="FB576F836032626DFF41FF668C6668C2" box="[252,301,136,160]" pageId="34" pageNumber="261">type</typeStatus>
|
||
of
|
||
<taxonomicName id="E3ECAAA26032626DFEDAFF668F4968C2" authorityName="Stepanjants" authorityYear="1979" box="[359,514,136,160]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="34" pageNumber="261" phylum="Cnidaria" rank="species" species="operculata">
|
||
<emphasis id="16980D336032626DFEDAFF668F4968C2" box="[359,514,136,160]" italics="true" pageId="34" pageNumber="261">A. operculata</emphasis>
|
||
</taxonomicName>
|
||
. Additionally, both have similar dimensions of their nematocysts (13.5–14X4.5–5 Mm and 8X3–3.5 Mm in
|
||
<taxonomicName id="E3ECAAA26032626DFC8CFF468E9868A2" authorityName="El Beshbeeshy" authorityYear="1991" box="[817,979,168,192]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="34" pageNumber="261" phylum="Cnidaria" rank="species" species="patagonica">
|
||
<emphasis id="16980D336032626DFC8CFF468E9868A2" box="[817,979,168,192]" italics="true" pageId="34" pageNumber="261">A. patagonica</emphasis>
|
||
</taxonomicName>
|
||
). Therefore, we consider
|
||
<taxonomicName id="E3ECAAA26032626DFF48FF268CC46882" authorityName="El Beshbeeshy" authorityYear="1991" box="[245,399,200,224]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="34" pageNumber="261" phylum="Cnidaria" rank="species" species="patagonica">
|
||
<emphasis id="16980D336032626DFF48FF268CC46882" box="[245,399,200,224]" italics="true" pageId="34" pageNumber="261">A. patagonica</emphasis>
|
||
</taxonomicName>
|
||
conspecific with
|
||
<taxonomicName id="E3ECAAA26032626DFDE1FF268FA46882" authorityName="Stepanjants" authorityYear="1979" box="[604,751,200,224]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="34" pageNumber="261" phylum="Cnidaria" rank="species" species="operculata">
|
||
<emphasis id="16980D336032626DFDE1FF268FA46882" box="[604,751,200,224]" italics="true" pageId="34" pageNumber="261">A. operculata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="2453D1216032626DFF18FF068ED369C2" blockId="34.[138,1185,136,416]" pageId="34" pageNumber="261">
|
||
<bibRefCitation id="407DACD06032626DFF18FF068CBD6962" author="Vervoort W & Watson JE" box="[165,502,232,256]" pageId="34" pageNumber="261" pagination="1 - 538" refId="ref23135" refString="Vervoort W, Watson JE. 2003. The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (Thecate Hydroids). NIWA Biodiversity Memoir 119: 1 - 538." type="journal article" year="2003">Vervoort and Watson (2003)</bibRefCitation>
|
||
also assigned material from
|
||
<collectingCountry id="5CFB91B16032626DFCFEFF068EAB6962" box="[835,992,232,256]" name="New Zealand" pageId="34" pageNumber="261">New Zealand</collectingCountry>
|
||
to
|
||
<taxonomicName id="E3ECAAA26032626DFBBAFF0689EA6962" authorityName="El Beshbeeshy" authorityYear="1991" box="[1031,1185,232,256]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="34" pageNumber="261" phylum="Cnidaria" rank="species" species="patagonica">
|
||
<emphasis id="16980D336032626DFBBAFF0689EA6962" box="[1031,1185,232,256]" italics="true" pageId="34" pageNumber="261">A. patagonica</emphasis>
|
||
</taxonomicName>
|
||
and, again, the description fully agrees with the
|
||
<typeStatus id="FB576F836032626DFD12FEE68FAB6942" box="[687,736,264,288]" pageId="34" pageNumber="261">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="E3ECAAA26032626DFCD4FEE68EB16942" authorityName="Stepanjants" authorityYear="1979" box="[873,1018,264,288]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="34" pageNumber="261" phylum="Cnidaria" rank="species" species="operculata">
|
||
<emphasis id="16980D336032626DFCD4FEE68EB16942" box="[873,1018,264,288]" italics="true" pageId="34" pageNumber="261">A. operculata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="407DACD06032626DFBB4FEE68C776922" author="Vervoort W & Watson JE" pageId="34" pageNumber="261" pagination="1 - 538" refId="ref23135" refString="Vervoort W, Watson JE. 2003. The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (Thecate Hydroids). NIWA Biodiversity Memoir 119: 1 - 538." type="journal article" year="2003">Vervoort and Watson (2003)</bibRefCitation>
|
||
indicated that the most significant character in
|
||
<taxonomicName id="E3ECAAA26032626DFC3DFEC689576922" authorityName="El Beshbeeshy" authorityYear="1991" box="[896,1052,296,320]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="34" pageNumber="261" phylum="Cnidaria" rank="species" species="patagonica">
|
||
<emphasis id="16980D336032626DFC3DFEC689576922" box="[896,1052,296,320]" italics="true" pageId="34" pageNumber="261">A. patagonica</emphasis>
|
||
</taxonomicName>
|
||
is that the conspicuous hydrothecae are not in one plane, but point obliquely frontally. As we have demonstrated before, this feature is also found in the
|
||
<typeStatus id="FB576F836032626DFD50FE868E5569E2" box="[749,798,360,384]" pageId="34" pageNumber="261">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="E3ECAAA26032626DFC1AFE86897369E2" authorityName="Stepanjants" authorityYear="1979" box="[935,1080,360,384]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="34" pageNumber="261" phylum="Cnidaria" rank="species" species="operculata">
|
||
<emphasis id="16980D336032626DFC1AFE86897369E2" box="[935,1080,360,384]" italics="true" pageId="34" pageNumber="261">A. operculata</emphasis>
|
||
</taxonomicName>
|
||
, and it is clearly described by
|
||
<bibRefCitation id="407DACD06032626DFEC7FE668F1269C2" author="Stepanjants SD" box="[378,601,392,416]" pageId="34" pageNumber="261" pagination="1 - 200" refId="ref22474" refString="Stepanjants SD. 1979. Hydroids of the antarctic and subantarctic waters. In: Biological results of the Soviet Antarctic Expeditions, 6. Issledovaniya Fauny Morei 20: 1 - 200. (Rus)." type="journal article" year="1979">Stepanjants (1979)</bibRefCitation>
|
||
in her original description.
|
||
</paragraph>
|
||
<paragraph id="2453D1216032626DFF37FE368C406992" blockId="34.[138,267,472,496]" box="[138,267,472,496]" pageId="34" pageNumber="261">
|
||
<emphasis id="16980D336032626DFF37FE368C406992" box="[138,267,472,496]" italics="true" pageId="34" pageNumber="261">Distribution</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6CF682AA6032626DFF37FDE68E486AC2" pageId="34" pageNumber="261" type="reference_group">
|
||
<paragraph id="2453D1216032626DFF37FDE68E486AC2" blockId="34.[138,1185,520,672]" pageId="34" pageNumber="261">
|
||
<taxonomicName id="E3ECAAA26032626DFF37FDE68CCD6A42" ID-CoL="9RXF" authorityName="Stepanjants" authorityYear="1979" box="[138,390,520,544]" class="Hydrozoa" family="Lafoeidae" genus="Acryptolaria" kingdom="Animalia" order="Leptothecata" pageId="34" pageNumber="261" phylum="Cnidaria" rank="species" species="operculata">
|
||
<emphasis id="16980D336032626DFF37FDE68CCD6A42" box="[138,390,520,544]" italics="true" pageId="34" pageNumber="261">Acryptolaria operculata</emphasis>
|
||
</taxonomicName>
|
||
seems to be a shelf and bathyal species, having been found at depths between 98 (
|
||
<bibRefCitation id="407DACD06032626DFEA3FDC68F406A22" author="El Beshbeeshy M" box="[286,523,552,576]" pageId="34" pageNumber="261" refId="ref18937" refString="El Beshbeeshy M. 1991. Systematische, Morphologische und Zoogeographische Untersuchungen an den Thekaten Hydroiden des Patagonischen Schelfs [PhD thesis]. Hamburg: University of Hamburg. 390 p." type="book" year="1991">El Beshbeeshy 1991</bibRefCitation>
|
||
) and
|
||
<quantity id="E3147CC46032626DFDF3FDC68FEF6A22" box="[590,676,552,576]" metricMagnitude="3" metricUnit="m" metricValue="1.422" pageId="34" pageNumber="261" unit="m" value="1422.0">1422 m</quantity>
|
||
(
|
||
<bibRefCitation id="407DACD06032626DFD0EFDC68E1E6A22" author="Watson JW" box="[691,853,552,576]" pageId="34" pageNumber="261" pagination="151 - 180" refId="ref23225" refString="Watson JW. 2003. Deep-water hydroids (Hydrozoa: Leptolida) from Macquarie Island. Memoirs of Museum of Victoria 60: 151 - 180." type="journal article" year="2003">Watson 2003</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="407DACD06032626DFCD7FDC689536A22" author="Watson JW" box="[874,1048,552,576]" pageId="34" pageNumber="261" pagination="151 - 180" refId="ref23225" refString="Watson JW. 2003. Deep-water hydroids (Hydrozoa: Leptolida) from Macquarie Island. Memoirs of Museum of Victoria 60: 151 - 180." type="journal article" year="2003">Watson (2003)</bibRefCitation>
|
||
found it on primnoid gorgonians. It is known to be distributed in sub-Antarctic waters, from the Patagonian shelf (
|
||
<bibRefCitation id="407DACD06032626DFEE2FD868F796AE2" author="Stepanjants SD" box="[351,562,616,640]" pageId="34" pageNumber="261" pagination="1 - 200" refId="ref22474" refString="Stepanjants SD. 1979. Hydroids of the antarctic and subantarctic waters. In: Biological results of the Soviet Antarctic Expeditions, 6. Issledovaniya Fauny Morei 20: 1 - 200. (Rus)." type="journal article" year="1979">Stepanjants 1979</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="407DACD06032626DFDFCFD868E7C6AE2" author="El Beshbeeshy M" box="[577,823,616,640]" pageId="34" pageNumber="261" refId="ref18937" refString="El Beshbeeshy M. 1991. Systematische, Morphologische und Zoogeographische Untersuchungen an den Thekaten Hydroiden des Patagonischen Schelfs [PhD thesis]. Hamburg: University of Hamburg. 390 p." type="book" year="1991">El Beshbeeshy 1991</bibRefCitation>
|
||
),
|
||
<collectingCountry id="5CFB91B16032626DFCEDFD868EA56AE2" box="[848,1006,616,640]" name="New Zealand" pageId="34" pageNumber="261">New Zealand</collectingCountry>
|
||
(
|
||
<bibRefCitation id="407DACD06032626DFBBFFD868C606AC2" author="Vervoort W & Watson JE" pageId="34" pageNumber="261" pagination="1 - 538" refId="ref23135" refString="Vervoort W, Watson JE. 2003. The marine fauna of New Zealand: Leptothecata (Cnidaria: Hydrozoa) (Thecate Hydroids). NIWA Biodiversity Memoir 119: 1 - 538." type="journal article" year="2003">Vervoort and Watson 2003</bibRefCitation>
|
||
), and
|
||
<collectingCountry id="5CFB91B16032626DFEC8FD668F096AC2" box="[373,578,648,672]" name="Australia" pageId="34" pageNumber="261">Macquarie Island</collectingCountry>
|
||
(
|
||
<bibRefCitation id="407DACD06032626DFDEEFD668FBD6AC2" author="Watson JW" box="[595,758,648,672]" pageId="34" pageNumber="261" pagination="151 - 180" refId="ref23225" refString="Watson JW. 2003. Deep-water hydroids (Hydrozoa: Leptolida) from Macquarie Island. Memoirs of Museum of Victoria 60: 151 - 180." type="journal article" year="2003">Watson 2003</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |