216 lines
21 KiB
XML
216 lines
21 KiB
XML
<document id="9AE6B346C2C1419BD0D89466435F1253" ID-CLB-Dataset="306474" ID-DOI="10.11646/zootaxa.5542.1.1" ID-GBIF-Dataset="b349e73d-c49e-418c-81f7-c5d32e136362" ID-ISSN="1175-5334" ID-Zenodo-Dep="14385386" ID-ZooBank="2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1733917054948" checkinUser="plazi" docAuthor="Moghaddam, Masumeh & Watson, Gillian W." docDate="2024" docId="981B8798FFF6392292DA907030ADDF8B" docLanguage="en" docName="zootaxa.5542.1.1.pdf" docOrigin="Zootaxa 5542 (1)" docSource="https://doi.org/10.11646/zootaxa.5542.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Acanthococcus Signoret" docType="treatment" docVersion="3" lastPageNumber="110" masterDocId="6422FFE0FF9D394F924D97543531DD43" masterDocTitle="The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families" masterLastPageNumber="202" masterPageNumber="1" pageNumber="108" updateTime="1734101643649" updateUser="ExternalLinkService">
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<mods:titleInfo id="E772A37F441F88F65AB913C6C13B493D">
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<mods:title id="B5B346B52DF8010143190210CADAD51E">The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families</mods:title>
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<mods:name id="14E2C441BF55FEAD726DD903B3341BDB" type="personal">
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<mods:roleTerm id="60E6031B1664FF28C5CD0588BABD077D">Author</mods:roleTerm>
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<mods:namePart id="E4112A7F53687B9A48F9986D87708EFD">Moghaddam, Masumeh</mods:namePart>
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<mods:affiliation id="ACB4C88114B0E7C9F2E2632FEE224CFA">Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran.</mods:affiliation>
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<mods:name id="8E1C38DA70B32468ED9F26464E2FBAFA" type="personal">
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<mods:roleTerm id="FBA5DF973DF57050D483A1385561ED11">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="281D74E232DE07EE9F2C7F009F5DED85">Watson, Gillian W.</mods:namePart>
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<mods:nameIdentifier id="9B581801CE35B4035F3439D69B74733C" type="ORCID">0000-0001-9914-0094</mods:nameIdentifier>
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<mods:affiliation id="9E815080D50CA3BDD1F0EE659970275E">Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran.</mods:affiliation>
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<mods:nameIdentifier id="47ED676AB7B69BE3A49083C4B2B14AD3" type="email">gillian.watson@nhm.ac.uk</mods:nameIdentifier>
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<mods:title id="5D793957D3EEF4D3B71A4855A270024C">Zootaxa</mods:title>
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<mods:date id="060BBFA86B563DF8F221A31D7C5E46B7">2024</mods:date>
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<mods:number id="B7A0537391107236D006AB8A2CE08696">2024-11-29</mods:number>
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<mods:number id="1FDCA9C9789AFB78BF3C6C306CCFE5FF">5542</mods:number>
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<treatment id="981B8798FFF6392292DA907030ADDF8B" LSID="urn:lsid:plazi:treatment:981B8798FFF6392292DA907030ADDF8B" httpUri="http://treatment.plazi.org/id/981B8798FFF6392292DA907030ADDF8B" lastPageId="109" lastPageNumber="110" pageId="107" pageNumber="108">
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<subSubSection id="58A86505FFF6392492DA90703753DA7D" box="[151,610,1828,1854]" pageId="107" pageNumber="108" type="nomenclature">
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<paragraph id="100D368EFFF6392492DA90703753DA7D" blockId="107.[151,610,1828,1854]" box="[151,610,1828,1854]" pageId="107" pageNumber="108">
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<emphasis id="22C6EA9CFFF6392492DA90703753DA7D" bold="true" box="[151,610,1828,1854]" pageId="107" pageNumber="108">
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Genus
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<taxonomicName id="D7B24D0DFFF6392492A390703753DA7D" authority="Signoret" authorityName="Signoret" box="[238,610,1828,1854]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="107" pageNumber="108" phylum="Arthropoda" rank="genus">
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<emphasis id="22C6EA9CFFF6392492A3907034C0DA7D" bold="true" box="[238,497,1828,1854]" italics="true" pageId="107" pageNumber="108">ACANTHOCOCCUS</emphasis>
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Signoret
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</taxonomicName>
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</emphasis>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="58A86505FFF6392492DA903C316BDAC2" box="[151,1114,1896,1921]" pageId="107" pageNumber="108" type="reference_group">
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<paragraph id="100D368EFFF6392492DA903C316BDAC2" blockId="107.[151,1114,1896,1921]" box="[151,1114,1896,1921]" pageId="107" pageNumber="108">
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<treatmentCitationGroup id="30A211A0FFF6392492DA903C316BDAC2" box="[151,1114,1896,1921]" pageId="107" pageNumber="108">
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<treatmentCitation id="9113109FFFF6392492DA903C34C4DAC3" author="Signoret, V." box="[151,501,1896,1920]" page="16" pageId="107" pageNumber="108" year="1875">
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<taxonomicName id="D7B24D0DFFF6392492DA903C34C4DAC3" authority="Signoret, 1875: 16" authorityName="Signoret" authorityPageNumber="16" authorityYear="1875" box="[151,501,1896,1920]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="107" pageNumber="108" phylum="Arthropoda" rank="genus">
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<emphasis id="22C6EA9CFFF6392492DA903C3401DAC3" box="[151,304,1896,1920]" italics="true" pageId="107" pageNumber="108">Acanthococcus</emphasis>
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<bibRefCitation id="74234B7FFFF63924937A903C34C4DAC3" author="Signoret, V." box="[311,501,1896,1920]" pageId="107" pageNumber="108" pagination="15 - 40" refId="ref108733" refString="Signoret, V. (1875) Essai sur les cochenilles ou gallinsectes (Homopteres - Coccides), 14 e partie. Annales de la Societe entomologique de France, Serie 5, 5, 15 - 40." type="journal article" year="1875">Signoret, 1875: 16</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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.
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<typeStatus id="CF09882CFFF6392493B2903D3700DAC2" box="[511,561,1897,1921]" pageId="107" pageNumber="108">Type</typeStatus>
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species:
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<taxonomicName id="D7B24D0DFFF6392490C3903C36F9DAC3" authority="Signoret" authorityName="Signoret" box="[654,968,1896,1920]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="107" pageNumber="108" phylum="Arthropoda" rank="species" species="aceris">
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<emphasis id="22C6EA9CFFF6392490C3903C365DDAC3" box="[654,876,1896,1920]" italics="true" pageId="107" pageNumber="108">Acanthococcus aceris</emphasis>
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Signoret
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</taxonomicName>
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by monotypy.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="58A86505FFF6392292DA90FA30ACDC02" lastPageId="109" lastPageNumber="110" pageId="107" pageNumber="108" type="diagnosis">
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||
<paragraph id="100D368EFFF6392392DA90FA362ADC02" blockId="107.[151,1436,1966,2029]" lastBlockId="108.[151,1437,151,321]" lastPageId="108" lastPageNumber="109" pageId="107" pageNumber="108">
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<emphasis id="22C6EA9CFFF6392492DA90FA37AFDA8B" bold="true" box="[151,670,1966,1993]" pageId="107" pageNumber="108">Diagnostic characters of the adult female:</emphasis>
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Body of live adult female covered by waxy ovoid ovisac. Slide-mounted, body elongate oval, tapering posteriorly. Antennae each with 6 or 7 (rarely 8) segments. Frontal lobes or tubercles present. Eyes situated near margin. Legs well developed; mid- and hind coxae often with spinulae on anterior surfaces; translucent pores sometimes present on hind coxa and femur; claw usually with a denticle; tarsal and claw digitules longer than claw, all with apices knobbed. Anal lobes well developed and often sclerotized, frequently with tooth-like projections on inner margins. Anal ring well developed, sclerotized, with partly double rows of pores and bearing 8 setae. Cauda usually present.
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</paragraph>
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<caption id="44CD6606FFF1392392DA90DC37E4DAAB" ID-DOI="http://doi.org/10.5281/zenodo.14385554" ID-Zenodo-Dep="14385554" httpUri="https://zenodo.org/record/14385554/files/figure.png" pageId="108" pageNumber="109" startId="108.[151,250,1928,1952]" targetBox="[184,1404,352,1897]" targetPageId="108" targetType="figure">
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<paragraph id="100D368EFFF1392392DA90DC37E4DAAB" blockId="108.[151,1436,1928,2024]" pageId="108" pageNumber="109">
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<emphasis id="22C6EA9CFFF1392392DA90DC3411DAE3" bold="true" box="[151,288,1928,1952]" pageId="108" pageNumber="109">FIGURE 47.</emphasis>
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General morphology of
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<taxonomicName id="D7B24D0DFFF13923905490DC3791DAE3" box="[537,672,1928,1952]" class="Insecta" family="Eriococcidae" kingdom="Animalia" order="Hemiptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="family">Eriococcidae</taxonomicName>
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, adult female.
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<emphasis id="22C6EA9CFFF13923917990DC367DDAE3" bold="true" box="[820,844,1928,1952]" pageId="108" pageNumber="109">A,</emphasis>
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anal lobes;
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<emphasis id="22C6EA9CFFF13923918890DC36EDDAE3" bold="true" box="[965,988,1928,1952]" pageId="108" pageNumber="109">B,</emphasis>
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antenna;
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<emphasis id="22C6EA9CFFF13923967290DC3169DAE3" bold="true" box="[1087,1112,1928,1952]" pageId="108" pageNumber="109">C,</emphasis>
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||
frontal lobe;
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<emphasis id="22C6EA9CFFF1392396AD90DC31C9DAE3" bold="true" box="[1248,1272,1928,1952]" pageId="108" pageNumber="109">D,</emphasis>
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labium;
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<emphasis id="22C6EA9CFFF13923971F90DC3058DAE3" bold="true" box="[1362,1385,1928,1952]" pageId="108" pageNumber="109">E,</emphasis>
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hind coxa;
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<emphasis id="22C6EA9CFFF13923929B90F835D8DA87" bold="true" box="[214,233,1964,1988]" pageId="108" pageNumber="109">F,</emphasis>
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claw;
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<emphasis id="22C6EA9CFFF13923936290F83478DA87" bold="true" box="[303,329,1964,1988]" pageId="108" pageNumber="109">G,</emphasis>
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cruciform, trilocular, quinquelocular and multilocular pores;
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<emphasis id="22C6EA9CFFF13923918F90F836EDDA87" bold="true" box="[962,988,1964,1988]" pageId="108" pageNumber="109">H,</emphasis>
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microtubular duct;
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<emphasis id="22C6EA9CFFF1392396E590F83188DA87" bold="true" box="[1192,1209,1964,1988]" pageId="108" pageNumber="109">I,</emphasis>
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macrotubular duct;
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<emphasis id="22C6EA9CFFF1392397C490F830ADDA87" bold="true" box="[1417,1436,1964,1988]" pageId="108" pageNumber="109">J,</emphasis>
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enlarged conical setae;
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<emphasis id="22C6EA9CFFF1392393C9908434AFDAAB" bold="true" box="[388,414,2000,2024]" pageId="108" pageNumber="109">K,</emphasis>
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anal ring;
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<emphasis id="22C6EA9CFFF13923904690843713DAAB" bold="true" box="[523,546,2000,2024]" pageId="108" pageNumber="109">L,</emphasis>
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vulva;
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<emphasis id="22C6EA9CFFF139239023908437BDDAAB" bold="true" box="[622,652,2000,2024]" pageId="108" pageNumber="109">M,</emphasis>
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cauda.
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</paragraph>
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</caption>
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<paragraph id="100D368EFFF03922928A97C337E7DD96" blockId="109.[151,1437,151,393]" pageId="109" pageNumber="110">
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<emphasis id="22C6EA9CFFF03922928A97C33410DDF3" box="[199,289,151,176]" italics="true" pageId="109" pageNumber="110">Dorsum</emphasis>
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. Setae often enlarged and spinose. Macrotubular ducts all of 1 size, with macrotubular and microtubular ducts together forming transverse rows throughout.
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</paragraph>
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<paragraph id="100D368EFFF03922928A978B30ACDC02" blockId="109.[151,1437,151,393]" pageId="109" pageNumber="110">
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<emphasis id="22C6EA9CFFF03922928A978B343EDDBB" box="[199,271,223,248]" italics="true" pageId="109" pageNumber="110">Venter</emphasis>
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. Setae hair like with enlarged setae near body margin. Labium with basal segment sometimes weakly developed but bearing 2 unequal pairs of setae. Macrotubular ducts often of 2 or 3 sizes, present throughout. Disc-pores each with 3‒9 loculi, present only on venter; cruciform pores present on ventral prosoma in a marginal band.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="58A86505FFF03922928A961E36D7DCCA" pageId="109" pageNumber="110" type="discussion">
|
||
<paragraph id="100D368EFFF03922928A961E36D7DCCA" blockId="109.[151,1437,151,393]" pageId="109" pageNumber="110">
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||
<emphasis id="22C6EA9CFFF03922928A961E3463DC27" bold="true" box="[199,338,330,356]" pageId="109" pageNumber="110">Comments:</emphasis>
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Worldwide, 217 species are included in
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<taxonomicName id="D7B24D0DFFF039229168961F36FEDC26" box="[805,975,331,357]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="genus">
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||
<emphasis id="22C6EA9CFFF039229168961F36FEDC26" box="[805,975,331,357]" italics="true" pageId="109" pageNumber="110">Acanthococcus</emphasis>
|
||
</taxonomicName>
|
||
at present (
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||
<bibRefCitation id="74234B7FFFF03922961B961F30A1DC26" author="Garcia Morales M. & Denno B. D. & Miller D. R. & Miller G. L. & Ben-Dov Y. & Hardy N. B." box="[1110,1424,331,357]" pageId="109" pageNumber="110" refId="ref101399" refString="Garcia Morales M., Denno B. D., Miller D. R., Miller G. L., Ben-Dov Y. & Hardy N. B. (2016) ScaleNet: a literature-based model of scale insect biology and systematics. Database. Available from: http: // scalenet. info (accessed 31 January 2024) https: // doi. org / 10.1093 / database / bav 118" type="url" year="2016">
|
||
García Morales
|
||
<emphasis id="22C6EA9CFFF03922975D961F3072DC26" box="[1296,1347,331,357]" italics="true" pageId="109" pageNumber="110">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
); three species are known from
|
||
<collectingCountry id="68A5761EFFF0392293A5963B3724DCCA" box="[488,533,367,393]" name="Iran" pageId="109" pageNumber="110">Iran</collectingCountry>
|
||
(
|
||
<bibRefCitation id="74234B7FFFF03922906E963B3632DCCA" author="Rozedar, E. & Vahedi, H. & Mossadegh, M. S. & Samie, M. S." box="[547,771,367,393]" pageId="109" pageNumber="110" pagination="41 - 68" refId="ref108067" refString="Rozedar, E., Vahedi, H., Mossadegh, M. S. & Samie, M. S. (2013) The scale insects (Hem.: Coccoideae) of Behbahan county and their morphological characters. Plant Protection, Scientific Journal of Agriculture, 36 (2), 41 - 68." type="journal article" year="2013">
|
||
Rozedar
|
||
<emphasis id="22C6EA9CFFF0392290C7963B378ADCCA" box="[650,699,367,393]" italics="true" pageId="109" pageNumber="110">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="74234B7FFFF039229142963B36EBDCCA" author="Torabi, M. & Vahedi, H. A. & Hodgson, C. J." box="[783,986,367,393]" pageId="109" pageNumber="110" pagination="153 - 162" refId="ref109821" refString="Torabi, M., Vahedi, H. A. & Hodgson, C. J. (2010) Preliminary survey of the scale insects fauna in Kermanshah, western Iran. Entomologia Hellenica, 19, 153 - 162. https: // doi. org / 10.12681 / eh. 11584" type="journal article" year="2010">
|
||
Torabi
|
||
<emphasis id="22C6EA9CFFF039229113963B36A1DCCA" box="[862,912,367,393]" italics="true" pageId="109" pageNumber="110">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="58A86505FFF0392292DA968F30ADDF8B" pageId="109" pageNumber="110" type="key">
|
||
<paragraph id="100D368EFFF0392292DA968F3166DCB6" blockId="109.[151,1111,474,501]" box="[151,1111,474,501]" pageId="109" pageNumber="110">
|
||
<emphasis id="22C6EA9CFFF0392292DA968F37BCDCB6" bold="true" box="[151,653,474,501]" pageId="109" pageNumber="110">
|
||
Key to species of
|
||
<taxonomicName id="D7B24D0DFFF039229323968E3717DCB6" box="[366,550,474,501]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="22C6EA9CFFF039229323968E3717DCB6" bold="true" box="[366,550,474,501]" italics="true" pageId="109" pageNumber="110">Acanthococcus</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="68A5761EFFF039229003968F37B6DCB6" box="[590,647,475,501]" name="Iran" pageId="109" pageNumber="110">Iran</collectingCountry>
|
||
,
|
||
</emphasis>
|
||
based on slide-mounted adult females.
|
||
</paragraph>
|
||
<key id="642361BFFFF0392292DA957430ADDF8B" pageId="109" pageNumber="110">
|
||
<keyStep id="AB462E2BFFF0392292DA957430ADDFCE" pageId="109" pageNumber="110">
|
||
<paragraph id="100D368EFFF0392292DA957430ADDF10" blockId="109.[151,1436,544,712]" pageId="109" pageNumber="110">
|
||
<keyLead id="AB4395BBFFF0392292DA957430ADDF10" pageId="109" pageNumber="110">
|
||
1(0) Dorsum of abdominal segment VIII with enlarged setae. Many marginal setae more than 2x as long as a dorsal enlarged seta.........................................................................................
|
||
<taxonomicName id="D7B24D0DFFF039229747956930ADDF10" authority="Signoret" authorityName="Signoret" box="[1290,1436,573,595]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="aceris">
|
||
<emphasis id="22C6EA9CFFF03922974795693074DF11" bold="true" box="[1290,1349,573,594]" italics="true" pageId="109" pageNumber="110">aceris</emphasis>
|
||
Signoret
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="100D368EFFF0392292DA950F30ADDFCE" blockId="109.[151,1436,544,712]" pageId="109" pageNumber="110">
|
||
<keyLead id="AB4395BBFFF0392292DA950F30ADDFCE" pageId="109" pageNumber="110">- Dorsum of abdominal segment VIII without enlarged setae. Marginal setae all less than 2x as long as a dorsal enlarged seta..................................................................................................... 2</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="AB462E2BFFF0392292DA95C130ADDF8B" pageId="109" pageNumber="110">
|
||
<paragraph id="100D368EFFF0392292DA95C130ADDFE8" blockId="109.[151,1436,544,712]" box="[151,1436,661,683]" pageId="109" pageNumber="110">
|
||
<keyLead id="AB4395BBFFF0392292DA95C130ADDFE8" box="[151,1436,661,683]" pageId="109" pageNumber="110">
|
||
2(1) Antenna with 6 segments. Anal ring complete, bearing 8 setae.....................................
|
||
<taxonomicName id="D7B24D0DFFF03922974C95C130ADDFE8" ID-CoL="8V63" authority="Danzig" authorityName="Danzig" box="[1281,1436,661,683]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="costatus">
|
||
<emphasis id="22C6EA9CFFF03922974C95C13060DFE9" bold="true" box="[1281,1361,661,682]" italics="true" pageId="109" pageNumber="110">costatus</emphasis>
|
||
Danzig
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="100D368EFFF0392292DA95E730ADDF8B" blockId="109.[151,1436,544,712]" box="[151,1436,690,712]" pageId="109" pageNumber="110">
|
||
<keyLead id="AB4395BBFFF0392292DA95E730ADDF8B" box="[151,1436,690,712]" pageId="109" pageNumber="110">
|
||
- Antenna with 7 segments. Anal ring incomplete, open anteriorly, and bearing 6 setae................
|
||
<taxonomicName id="D7B24D0DFFF0392296AB95E630ADDF8B" ID-CoL="8V8J" authority="Danzig" authorityName="Danzig" box="[1254,1436,690,712]" class="Insecta" family="Eriococcidae" genus="Acanthococcus" kingdom="Animalia" order="Hemiptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="isacanthus">
|
||
<emphasis id="22C6EA9CFFF0392296AB95E63060DF8B" bold="true" box="[1254,1361,690,712]" italics="true" pageId="109" pageNumber="110">isacanthus</emphasis>
|
||
Danzig
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |