306 lines
38 KiB
XML
306 lines
38 KiB
XML
<document id="1552740A45E98FC7B234F1037254064F" ID-DOI="10.11646/zootaxa.5542.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14385386" ID-ZooBank="2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1733917054948" checkinUser="plazi" docAuthor="Moghaddam, Masumeh & Watson, Gillian W." docDate="2024" docId="981B8798FFD2391D92DA91A631A2DD96" docLanguage="en" docName="zootaxa.5542.1.1.pdf" docOrigin="Zootaxa 5542 (1)" docSource="https://doi.org/10.11646/zootaxa.5542.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Pulvinaria camelicola Signoret" docType="treatment" docVersion="4" lastPageNumber="82" masterDocId="6422FFE0FF9D394F924D97543531DD43" masterDocTitle="The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families" masterLastPageNumber="202" masterPageNumber="1" pageNumber="80" updateTime="1734093388678" updateUser="carolina">
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<mods:titleInfo id="70CEB89E6014F047398978880F21263B">
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<mods:title id="6E7E6CE27576A36392BE4D1B922E48CA">The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families</mods:title>
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<mods:name id="57A74C4A0380A42BB98EB6620CD0D2D4" type="personal">
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<mods:roleTerm id="A3C61B56E69D00820786E0D52E236D16">Author</mods:roleTerm>
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<mods:namePart id="77D430C6E1886998436B835B95A4DBFD">Moghaddam, Masumeh</mods:namePart>
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<mods:affiliation id="6C94C37C6FB34455D5CF4A511E7DC72E">Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran.</mods:affiliation>
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<mods:name id="52CE8B1E80701A702A2F4170BFE77B0A" type="personal">
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<mods:roleTerm id="8A1F55399E36D7433CD850C152A6A94F">Author</mods:roleTerm>
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<mods:namePart id="C92E56AE77A6C96CB6B3C4B0F0ECD1C7">Watson, Gillian W.</mods:namePart>
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<mods:nameIdentifier id="CE3B19598B784442B8B1F755BC02E510" type="ORCID">0000-0001-9914-0094</mods:nameIdentifier>
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<mods:affiliation id="43555DDB65941543B1479D7CC2B412A4">Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. & Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X & Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran.</mods:affiliation>
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<mods:nameIdentifier id="F2EB4C3C1D28C944DDB6CE8A74838229" type="email">gillian.watson@nhm.ac.uk</mods:nameIdentifier>
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<mods:title id="755E3D37A2902DAED1CB93118BE6322B">Zootaxa</mods:title>
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<mods:date id="2724C4FA44FE3478F369EAB588408F13">2024</mods:date>
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<mods:number id="B76B6F4A02AF801FE04236BAE27A2724">2024-11-29</mods:number>
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<treatment id="981B8798FFD2391D92DA91A631A2DD96" ID-DOI="http://doi.org/10.5281/zenodo.14385564" ID-Zenodo-Dep="14385564" LSID="urn:lsid:plazi:treatment:981B8798FFD2391D92DA91A631A2DD96" httpUri="http://treatment.plazi.org/id/981B8798FFD2391D92DA91A631A2DD96" lastPageId="82" lastPageNumber="82" pageId="79" pageNumber="80">
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<subSubSection id="58A86505FFD2390092DA91A63723DA4E" box="[151,530,1778,1805]" pageId="79" pageNumber="80" type="nomenclature">
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<paragraph id="100D368EFFD2390092DA91A63723DA4E" blockId="79.[151,530,1778,1841]" box="[151,530,1778,1805]" pageId="79" pageNumber="80">
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<heading id="4B4581E2FFD2390092DA91A63723DA4E" bold="true" box="[151,530,1778,1805]" fontSize="11" level="2" pageId="79" pageNumber="80" reason="4">
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<taxonomicName id="D7B24D0DFFD2390092DA91A63723DA4E" ID-CoL="4QH84" authority="Signoret" authorityName="Signoret" box="[151,530,1778,1805]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="camelicola">
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<emphasis id="22C6EA9CFFD2390092DA91A63723DA4E" bold="true" box="[151,530,1778,1805]" pageId="79" pageNumber="80">
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<emphasis id="22C6EA9CFFD2390092DA91A63490DA4E" bold="true" box="[151,417,1778,1805]" italics="true" pageId="79" pageNumber="80">Pulvinaria camelicola</emphasis>
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Signoret
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="58A86505FFD2390092DA904335C8DA72" box="[151,249,1815,1841]" pageId="79" pageNumber="80" type="description">
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||
<paragraph id="100D368EFFD2390092DA904335C8DA72" blockId="79.[151,530,1778,1841]" box="[151,249,1815,1841]" pageId="79" pageNumber="80">
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||
(
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<figureCitation id="88892A0BFFD2390092D2904335C0DA72" box="[159,241,1815,1841]" captionStart="FIGURE 33" captionStartId="80.[151,250,1943,1967]" captionTargetBox="[196,1397,188,1910]" captionTargetId="figure-22@80.[187,1401,181,1919]" captionTargetPageId="80" captionText="FIGURE 33. Adult female of Pulvinaria camelicola Signoret, from Tanaka & Kamitani (2022) page 441, fig. 1. © Magnolia Press, www.mapress.com/j/zt, reproduced with permission from the authors and the copyright holder. Distributions of tubular duct types (i), (ii) and (iii) shown; for details of their respective distributions, see also the diagnosis." figureDoi="http://doi.org/10.5281/zenodo.14385512" httpUri="https://zenodo.org/record/14385512/files/figure.png" pageId="79" pageNumber="80">Fig. 33</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="58A86505FFD2390092DA900F377EDA30" box="[151,591,1883,1907]" pageId="79" pageNumber="80" type="reference_group">
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<paragraph id="100D368EFFD2390092DA900F377EDA30" blockId="79.[151,591,1883,1907]" box="[151,591,1883,1907]" pageId="79" pageNumber="80">
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<treatmentCitationGroup id="30A211A0FFD2390092DA900F377EDA30" box="[151,591,1883,1907]" pageId="79" pageNumber="80">
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<treatmentCitation id="9113109FFFD2390092DA900F377ADA30" author="Signoret, V." box="[151,587,1883,1907]" page="32" pageId="79" pageNumber="80" year="1873">
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<taxonomicName id="D7B24D0DFFD2390092DA900F377ADA30" ID-CoL="4QH84" authority="Signoret, 1873 b: 32" authorityName="Signoret" authorityPageNumber="32" authorityYear="1873" box="[151,587,1883,1907]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="camelicola">
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<emphasis id="22C6EA9CFFD2390092DA900F3448DA30" box="[151,377,1883,1907]" italics="true" pageId="79" pageNumber="80">Pulvinaria camelicola</emphasis>
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<bibRefCitation id="74234B7FFFD239009332900F377ADA30" author="Signoret, V." box="[383,587,1883,1907]" pageId="79" pageNumber="80" pagination="27 - 48" refId="ref108692" refString="Signoret, V. (1873 b) Essai sur les cochenilles ou gallinsectes (Homopteres - Coccides), 10 e partie. Annales de la Societe entomologique de France, Serie 5, 3, 27 - 48." type="journal article" year="1873">Signoret, 1873b: 32</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="58A86505FFD2390092DA90F53004DA9C" pageId="79" pageNumber="80" type="multiple">
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<paragraph id="100D368EFFD2390092DA90F53004DA9C" blockId="79.[151,1436,1953,2016]" pageId="79" pageNumber="80">
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<emphasis id="22C6EA9CFFD2390092DA90F534A0DAF8" bold="true" box="[151,401,1953,1979]" pageId="79" pageNumber="80">Field characteristics:</emphasis>
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Live adult female elongate oval, slightly convex; body yellow with small reddish spots, quite similar to
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<taxonomicName id="D7B24D0DFFD239009344909237F9DAA3" ID-CoL="78PBJ" authority="(Tanaka & Kamitani 2022)" baseAuthorityName="Tanaka & Kamitani" baseAuthorityYear="2022" box="[265,712,1989,2016]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="floccifera">
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<emphasis id="22C6EA9CFFD239009344909234BEDA9C" box="[265,399,1989,2015]" italics="true" pageId="79" pageNumber="80">P. floccifera</emphasis>
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(
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<bibRefCitation id="74234B7FFFD2390093D3909137F1DAA3" author="Tanaka, H. & Kamitani, S." box="[414,704,1989,2016]" pageId="79" pageNumber="80" pagination="438 - 450" refId="ref109340" refString="Tanaka, H. & Kamitani, S. (2022) The identities of Pulvinaria camelicola Signoret 1873, stat. nov., and the related species, Pulvinaria floccifera (Westwood 1870) (Hemiptera: Coccidae) in Japan. Zootaxa, 5115 (3), 438 - 450. https: // doi. org / 10.11646 / zootaxa. 5115.3.8" type="journal article" year="2022">Tanaka & Kamitani 2022</bibRefCitation>
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||
)
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||
</taxonomicName>
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||
. Mature adult female produces a relatively long ovisac.
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||
</paragraph>
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||
</subSubSection>
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||
<caption id="44CD6606FFCD391F92DA90C331BCDAB4" ID-DOI="http://doi.org/10.5281/zenodo.14385512" ID-Zenodo-Dep="14385512" httpUri="https://zenodo.org/record/14385512/files/figure.png" pageId="80" pageNumber="81" startId="80.[151,250,1943,1967]" targetBox="[196,1397,188,1910]" targetPageId="80" targetType="figure">
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||
<paragraph id="100D368EFFCD391F92DA90C331BCDAB4" blockId="80.[151,1436,1943,2039]" pageId="80" pageNumber="81">
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<emphasis id="22C6EA9CFFCD391F92DA90C33410DAEC" bold="true" box="[151,289,1943,1967]" pageId="80" pageNumber="81">FIGURE 33.</emphasis>
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Adult female of
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<taxonomicName id="D7B24D0DFFCD391F939D90C33623DAEC" authority="Signoret" authorityName="Signoret" box="[464,786,1943,1967]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="camelicola">
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<emphasis id="22C6EA9CFFCD391F939D90C33782DAEC" box="[464,691,1943,1967]" italics="true" pageId="80" pageNumber="81">Pulvinaria camelicola</emphasis>
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Signoret
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</taxonomicName>
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, from
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<bibRefCitation id="74234B7FFFCD391F911B90C3315BDAEC" author="Tanaka, H. & Kamitani, S." box="[854,1130,1943,1967]" pageId="80" pageNumber="81" pagination="438 - 450" refId="ref109340" refString="Tanaka, H. & Kamitani, S. (2022) The identities of Pulvinaria camelicola Signoret 1873, stat. nov., and the related species, Pulvinaria floccifera (Westwood 1870) (Hemiptera: Coccidae) in Japan. Zootaxa, 5115 (3), 438 - 450. https: // doi. org / 10.11646 / zootaxa. 5115.3.8" type="journal article" year="2022">Tanaka & Kamitani (2022)</bibRefCitation>
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page 441, fig. 1. © Magnolia Press, www.mapress.com/j/zt, reproduced with permission from the authors and the copyright holder. Distributions of tubular duct types (i), (ii) and (iii) shown; for details of their respective distributions, see also the diagnosis.
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</paragraph>
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</caption>
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<subSubSection id="58A86505FFCC391E928A97C236A7D976" pageId="81" pageNumber="82" type="diagnosis">
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<paragraph id="100D368EFFCC391E928A97C23492DD96" blockId="81.[151,1437,150,2013]" pageId="81" pageNumber="82">
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<emphasis id="22C6EA9CFFCC391E928A97C234E9DDF3" bold="true" box="[199,472,150,176]" pageId="81" pageNumber="82">Microscopic diagnosis:</emphasis>
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Slide-mounted adult female body elongate oval. Stigmatic clefts rather shallow. Anal cleft of moderate depth.
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</paragraph>
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<paragraph id="100D368EFFCC391E928A978B31FFDC26" blockId="81.[151,1437,150,2013]" pageId="81" pageNumber="82">
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<emphasis id="22C6EA9CFFCC391E928A978B3410DDBB" box="[199,289,223,248]" italics="true" pageId="81" pageNumber="82">Dorsum</emphasis>
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. Derm entirely membranous; dermal areolations well developed. Setae spiniform, scattered throughout. Preopercular pores oval to circular, often difficult to see, present anterior to anal plates. Submarginal dorsal tubercles present, numbering 0−4 on each side. Tubular ducts frequent throughout, each situated within an areolation. Anal plates together quadrate, each plate with 3 or 4 apical setae. Anal ring usually bearing 5−7 setae.
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</paragraph>
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<paragraph id="100D368EFFCC391E928A963B34DCDCB6" blockId="81.[151,1437,150,2013]" pageId="81" pageNumber="82">
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<emphasis id="22C6EA9CFFCC391E928A963B342BDCCB" box="[199,282,367,392]" italics="true" pageId="81" pageNumber="82">Margin</emphasis>
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. Marginal setae with well-developed basal sockets and with tips fimbriate, frayed, spatulate, split, or simply pointed, those on either side of anal cleft tending to be longer; each side with 12–25 setae between anterior and posterior stigmatic clefts. Stigmatic clefts distinct but shallow, each containing 3 setae, median seta about 2−4 times longer than lateral setae.
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</paragraph>
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<paragraph id="100D368EFFCC391E928A96AB36A7D976" blockId="81.[151,1437,150,2013]" pageId="81" pageNumber="82">
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<emphasis id="22C6EA9CFFCC391E928A96AB343EDF5B" box="[199,271,511,536]" italics="true" pageId="81" pageNumber="82">Venter</emphasis>
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. Derm membranous. Pregenital disc-pores mostly each with 5−8 loculi, present around genital opening, and on medial areas of preceding 3 to 5 abdominal segments; a small group also present lateral to each metacoxa and occasionally to each mesocoxa also. Spiracular disc-pores each with 5 loculi, present in a narrow band between each spiracle and margin. Microducts scattered throughout venter. Tubular ducts of
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<specimenCount id="06B4FD07FFCC391E967E953F31B7DFC6" box="[1075,1158,619,645]" count="3" pageId="81" pageNumber="82" type="generic" typeStatus="types">3 types</specimenCount>
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:
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<typeStatus id="CF09882CFFCC391E96C2953F31F6DFC6" box="[1167,1223,619,645]" pageId="81" pageNumber="82">Type</typeStatus>
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I with a large outer ductule and a broad inner ductule ending in a well-developed flower-shaped terminal gland: present in posterior medial area of head, medial area of all thoracic segments and anterior abdominal segments, also in inner submarginal area extending from near anterior abdominal segments to prothoracic segments;
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<typeStatus id="CF09882CFFCC391E967695833142DFB2" box="[1083,1139,727,753]" pageId="81" pageNumber="82">Type</typeStatus>
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<collectionCode id="76A3AE4BFFCC391E9631958331BEDFB2" box="[1148,1167,727,753]" pageId="81" pageNumber="82">II</collectionCode>
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with a rather narrower outer ductule ending in a shallow cup-shaped invagination, and a long, slender inner ductule with a well-developed terminal gland, present medially on posterior abdominal segments; and submarginal band of tubular ducts but reaching only as far forward as posterior spiracular pore band; and
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<typeStatus id="CF09882CFFCC391E91C4941736F0DE1E" box="[905,961,835,861]" pageId="81" pageNumber="82">Type</typeStatus>
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<collectionCode id="76A3AE4BFFCC391E9185941736D5DE1E" box="[968,996,835,861]" pageId="81" pageNumber="82">III</collectionCode>
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with a short, filamentous inner ductule and minute terminal gland, present in a broad submarginal band on thoracic and abdominal segments, but mostly concentrated on posterior area, intermixed with
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<typeStatus id="CF09882CFFCC391E90FF94DF37DBDEE6" box="[690,746,907,933]" pageId="81" pageNumber="82">Type</typeStatus>
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I or
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<typeStatus id="CF09882CFFCC391E915294DF3666DEE6" box="[799,855,907,933]" pageId="81" pageNumber="82">Type</typeStatus>
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<collectionCode id="76A3AE4BFFCC391E911394DF3640DEE6" box="[862,881,907,933]" pageId="81" pageNumber="82">II</collectionCode>
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ducts. Posterior 3 abdominal segments each with 1 pair of long ventral setae on medial area. With 2–6 pairs of long setae present between antennal bases; other setae short and fine, distributed over entire venter. Legs well developed, each with an articulated tibio-tarsal joint and an articulatory sclerosis; claw without a denticle; both claw digitules rather broad and slightly shorter than thin tarsal digitules. Antennae each 6–8 segmented (
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<bibRefCitation id="74234B7FFFCC391E9025934F36BBD976" author="Tanaka, H. & Kamitani, S." box="[616,906,1051,1077]" pageId="81" pageNumber="82" pagination="438 - 450" refId="ref109340" refString="Tanaka, H. & Kamitani, S. (2022) The identities of Pulvinaria camelicola Signoret 1873, stat. nov., and the related species, Pulvinaria floccifera (Westwood 1870) (Hemiptera: Coccidae) in Japan. Zootaxa, 5115 (3), 438 - 450. https: // doi. org / 10.11646 / zootaxa. 5115.3.8" type="journal article" year="2022">Tanaka & Kamitani 2022</bibRefCitation>
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||
).
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</paragraph>
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</subSubSection>
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<subSubSection id="58A86505FFCC391E928A936A3058D93E" pageId="81" pageNumber="82" type="distribution">
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<paragraph id="100D368EFFCC391E928A936A3058D93E" blockId="81.[151,1437,150,2013]" pageId="81" pageNumber="82">
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<emphasis id="22C6EA9CFFCC391E928A936A3452D91B" bold="true" box="[199,355,1086,1112]" pageId="81" pageNumber="82">Distribution:</emphasis>
|
||
<taxonomicName id="D7B24D0DFFCC391E9325936B3751D91A" box="[360,608,1087,1113]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="camelicola">
|
||
<emphasis id="22C6EA9CFFCC391E9325936B3751D91A" box="[360,608,1087,1113]" italics="true" pageId="81" pageNumber="82">Pulvinaria camelicola</emphasis>
|
||
</taxonomicName>
|
||
has been reported from 54 countries in five zoogeograpical regions (
|
||
<bibRefCitation id="74234B7FFFCC391E971D936B3444D93E" author="Garcia Morales M. & Denno B. D. & Miller D. R. & Miller G. L. & Ben-Dov Y. & Hardy N. B." pageId="81" pageNumber="82" refId="ref101399" refString="Garcia Morales M., Denno B. D., Miller D. R., Miller G. L., Ben-Dov Y. & Hardy N. B. (2016) ScaleNet: a literature-based model of scale insect biology and systematics. Database. Available from: http: // scalenet. info (accessed 31 January 2024) https: // doi. org / 10.1093 / database / bav 118" type="url" year="2016">
|
||
García Morales
|
||
<emphasis id="22C6EA9CFFCC391E92B79337341AD93E" box="[250,299,1123,1149]" italics="true" pageId="81" pageNumber="82">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
), despite it not having been well described until very recently (
|
||
<bibRefCitation id="74234B7FFFCC391E96799337306CD93E" author="Tanaka, H. & Kamitani, S." box="[1076,1373,1123,1149]" pageId="81" pageNumber="82" pagination="438 - 450" refId="ref109340" refString="Tanaka, H. & Kamitani, S. (2022) The identities of Pulvinaria camelicola Signoret 1873, stat. nov., and the related species, Pulvinaria floccifera (Westwood 1870) (Hemiptera: Coccidae) in Japan. Zootaxa, 5115 (3), 438 - 450. https: // doi. org / 10.11646 / zootaxa. 5115.3.8" type="journal article" year="2022">Tanaka & Kamitani, 2022</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="58A86505FFCC391E928A93D331F5D986" pageId="81" pageNumber="82" type="biology_ecology">
|
||
<paragraph id="100D368EFFCC391E928A93D331F5D986" blockId="81.[151,1437,150,2013]" pageId="81" pageNumber="82">
|
||
<emphasis id="22C6EA9CFFCC391E928A93D3346AD9E2" bold="true" box="[199,347,1159,1185]" pageId="81" pageNumber="82">Host-plants:</emphasis>
|
||
<taxonomicName id="D7B24D0DFFCC391E932993D33751D9E2" box="[356,608,1159,1185]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="camelicola">
|
||
<emphasis id="22C6EA9CFFCC391E932993D33751D9E2" box="[356,608,1159,1185]" italics="true" pageId="81" pageNumber="82">Pulvinaria camelicola</emphasis>
|
||
</taxonomicName>
|
||
has been recorded on host-plants in 62 genera belonging to 44 families (
|
||
<bibRefCitation id="74234B7FFFCC391E92D293FF34E0D986" author="Garcia Morales M. & Denno B. D. & Miller D. R. & Miller G. L. & Ben-Dov Y. & Hardy N. B." box="[159,465,1195,1221]" pageId="81" pageNumber="82" refId="ref101399" refString="Garcia Morales M., Denno B. D., Miller D. R., Miller G. L., Ben-Dov Y. & Hardy N. B. (2016) ScaleNet: a literature-based model of scale insect biology and systematics. Database. Available from: http: // scalenet. info (accessed 31 January 2024) https: // doi. org / 10.1093 / database / bav 118" type="url" year="2016">
|
||
García Morales
|
||
<emphasis id="22C6EA9CFFCC391E931893FF34B7D986" box="[341,390,1195,1221]" italics="true" pageId="81" pageNumber="82">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
). The host-plant in
|
||
<collectingCountry id="68A5761EFFCC391E90E493FF37E7D986" box="[681,726,1195,1221]" name="Iran" pageId="81" pageNumber="82">Iran</collectingCountry>
|
||
was not recorded by
|
||
<bibRefCitation id="74234B7FFFCC391E918B93FF31F1D986" author="Bodenheimer, F. S." box="[966,1216,1195,1221]" pageId="81" pageNumber="82" pagination="85 - 100" refId="ref98321" refString="Bodenheimer, F. S. (1944 b) Note on the Coccoidea of Iran, with description of new species. Bulletin de la Societe Fouad 1 er d'Entomologie, 28, 85 - 100." type="journal article" year="1944">Bodenheimer (1944b)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="58A86505FFCC391E928A939A37A0D84E" pageId="81" pageNumber="82" type="multiple">
|
||
<paragraph id="100D368EFFCC391E928A939A37F8D9AA" blockId="81.[151,1437,150,2013]" box="[199,713,1230,1257]" pageId="81" pageNumber="82">
|
||
<emphasis id="22C6EA9CFFCC391E928A939A34E6D9AB" bold="true" box="[199,471,1230,1256]" pageId="81" pageNumber="82">Economic importance:</emphasis>
|
||
Not recorded in
|
||
<collectingCountry id="68A5761EFFCC391E90D9939B37F4D9AA" box="[660,709,1231,1257]" name="Iran" pageId="81" pageNumber="82">Iran</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="100D368EFFCC391E928A93A737A0D84E" blockId="81.[151,1437,150,2013]" box="[199,657,1266,1293]" pageId="81" pageNumber="82">
|
||
<emphasis id="22C6EA9CFFCC391E928A93A734A5D84F" bold="true" box="[199,404,1266,1293]" pageId="81" pageNumber="82">Natural enemies:</emphasis>
|
||
None recorded in
|
||
<collectingCountry id="68A5761EFFCC391E902993A737A0D84E" box="[612,657,1267,1293]" name="Iran" pageId="81" pageNumber="82">Iran</collectingCountry>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="58A86505FFCC391D928A924231A2DD96" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82" type="discussion">
|
||
<paragraph id="100D368EFFCC391E928A92423060D83A" blockId="81.[151,1437,150,2013]" pageId="81" pageNumber="82">
|
||
<emphasis id="22C6EA9CFFCC391E928A92423463D873" bold="true" box="[199,338,1302,1328]" pageId="81" pageNumber="82">Comments:</emphasis>
|
||
<bibRefCitation id="74234B7FFFCC391E931692433764D872" author="Bodenheimer, F. S." box="[347,597,1303,1329]" pageId="81" pageNumber="82" pagination="85 - 100" refId="ref98321" refString="Bodenheimer, F. S. (1944 b) Note on the Coccoidea of Iran, with description of new species. Bulletin de la Societe Fouad 1 er d'Entomologie, 28, 85 - 100." type="journal article" year="1944">Bodenheimer (1944b)</bibRefCitation>
|
||
recorded
|
||
<taxonomicName id="D7B24D0DFFCC391E908792433655D872" box="[714,868,1303,1329]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="camelicola">
|
||
<emphasis id="22C6EA9CFFCC391E908792433655D872" box="[714,868,1303,1329]" italics="true" pageId="81" pageNumber="82">P. camelicola</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="68A5761EFFCC391E91E3924336EFD872" box="[942,990,1303,1329]" name="Iran" pageId="81" pageNumber="82">Iran</collectingCountry>
|
||
, but neither his material nor any other specimens from
|
||
<collectingCountry id="68A5761EFFCC391E931F926F344ED816" box="[338,383,1339,1365]" name="Iran" pageId="81" pageNumber="82">Iran</collectingCountry>
|
||
have been seen in the present study so it has not been possible to confirm its presence in the country. It is possible that
|
||
<bibRefCitation id="74234B7FFFCC391E93F3920B37F8D83A" author="Bodenheimer, F. S." box="[446,713,1375,1401]" pageId="81" pageNumber="82" pagination="85 - 100" refId="ref98321" refString="Bodenheimer, F. S. (1944 b) Note on the Coccoidea of Iran, with description of new species. Bulletin de la Societe Fouad 1 er d'Entomologie, 28, 85 - 100." type="journal article" year="1944">Bodenheimer’s (1944b)</bibRefCitation>
|
||
record may have been a misidentification of
|
||
<taxonomicName id="D7B24D0DFFCC391E9689920B307BD83A" baseAuthorityName="Tanaka & Kamitani" baseAuthorityYear="2022" box="[1220,1354,1375,1401]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="floccifera">
|
||
<emphasis id="22C6EA9CFFCC391E9689920B307BD83A" box="[1220,1354,1375,1401]" italics="true" pageId="81" pageNumber="82">P. floccifera</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="100D368EFFCC391E928A92D73677DB6E" blockId="81.[151,1437,150,2013]" pageId="81" pageNumber="82">
|
||
In the past, several little-studied species (including
|
||
<taxonomicName id="D7B24D0DFFCC391E914392D73697D8DE" box="[782,934,1411,1437]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="camelicola">
|
||
<emphasis id="22C6EA9CFFCC391E914392D73697D8DE" box="[782,934,1411,1437]" italics="true" pageId="81" pageNumber="82">P. camelicola</emphasis>
|
||
</taxonomicName>
|
||
) have been misidentified as the well-known polyphagous pest species,
|
||
<taxonomicName id="D7B24D0DFFCC391E938892F3377CD882" baseAuthorityName="Tanaka & Kamitani" baseAuthorityYear="2022" box="[453,589,1447,1473]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="floccifera">
|
||
<emphasis id="22C6EA9CFFCC391E938892F3377CD882" box="[453,589,1447,1473]" italics="true" pageId="81" pageNumber="82">P. floccifera</emphasis>
|
||
</taxonomicName>
|
||
, because its original description is too brief and subsequent redescriptions were based on misidentified material. This situation subjectively invalidates the identity of
|
||
<taxonomicName id="D7B24D0DFFCC391E96FC929F300BD8A6" baseAuthorityName="Tanaka & Kamitani" baseAuthorityYear="2022" box="[1201,1338,1483,1509]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="floccifera">
|
||
<emphasis id="22C6EA9CFFCC391E96FC929F300BD8A6" box="[1201,1338,1483,1509]" italics="true" pageId="81" pageNumber="82">P. floccifera</emphasis>
|
||
</taxonomicName>
|
||
because several species have been studied under the same (inappropriate) scientific name in the absence of
|
||
<typeStatus id="CF09882CFFCC391E974892BB3004DB4A" box="[1285,1333,1519,1545]" pageId="81" pageNumber="82">type</typeStatus>
|
||
fixation, leading to taxonomic confusion (
|
||
<bibRefCitation id="74234B7FFFCC391E90459147360BDB6E" author="Garcia Morales M. & Denno B. D. & Miller D. R. & Miller G. L. & Ben-Dov Y. & Hardy N. B." box="[520,826,1555,1581]" pageId="81" pageNumber="82" refId="ref101399" refString="Garcia Morales M., Denno B. D., Miller D. R., Miller G. L., Ben-Dov Y. & Hardy N. B. (2016) ScaleNet: a literature-based model of scale insect biology and systematics. Database. Available from: http: // scalenet. info (accessed 31 January 2024) https: // doi. org / 10.1093 / database / bav 118" type="url" year="2016">
|
||
García Morales
|
||
<emphasis id="22C6EA9CFFCC391E90F3914737C7DB6E" box="[702,758,1555,1581]" italics="true" pageId="81" pageNumber="82">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="100D368EFFCC391E928A9163318CDA0E" blockId="81.[151,1437,150,2013]" pageId="81" pageNumber="82">
|
||
<bibRefCitation id="74234B7FFFCC391E928A916334DADB12" author="Tanaka, H. & Amano, H." box="[199,491,1591,1617]" pageId="81" pageNumber="82" pagination="211 - 215" refId="ref109295" refString="Tanaka, H. & Amano, H. (2007) Redescription of Pulvinaria floccifera (Insecta: Hemiptera: Coccidae). Species Diversity, 12, 211 - 215. https: // doi. org / 10.12782 / specdiv. 12.211" type="journal article" year="2007">
|
||
Tanaka &
|
||
<collectingRegion id="D276F86CFFCC391E930F916334A5DB12" box="[322,404,1591,1617]" country="Jordan" name="'Amman" pageId="81" pageNumber="82">Amano</collectingRegion>
|
||
(2007)
|
||
</bibRefCitation>
|
||
found that the morphology of
|
||
<specimenCount id="06B4FD07FFCC391E9116916336FBDB12" box="[859,970,1591,1617]" count="5" pageId="81" pageNumber="82" type="adult">five adult</specimenCount>
|
||
female
|
||
<typeStatus id="CF09882CFFCC391E9661916331BFDB12" box="[1068,1166,1591,1617]" pageId="81" pageNumber="82" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="D7B24D0DFFCC391E96F591633073DB12" baseAuthorityName="Tanaka & Kamitani" baseAuthorityYear="2022" box="[1208,1346,1591,1617]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="floccifera">
|
||
<emphasis id="22C6EA9CFFCC391E96F591633073DB12" box="[1208,1346,1591,1617]" italics="true" pageId="81" pageNumber="82">P. floccifera</emphasis>
|
||
</taxonomicName>
|
||
did not correspond with the descriptions of several of the species that were considered synonyms. To confer taxonomic stability to
|
||
<taxonomicName id="D7B24D0DFFCC391E935E912B34A8DBDA" baseAuthorityName="Tanaka & Kamitani" baseAuthorityYear="2022" box="[275,409,1663,1689]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="floccifera">
|
||
<emphasis id="22C6EA9CFFCC391E935E912B34A8DBDA" box="[275,409,1663,1689]" italics="true" pageId="81" pageNumber="82">P. floccifera</emphasis>
|
||
</taxonomicName>
|
||
, in accordance with the provisions of Article 23.3.5 of the Fourth edition of the International Code of Zoological Nomenclature (
|
||
<bibRefCitation id="74234B7FFFCC391E906C91F73797DBFE" author="ICZN" box="[545,678,1699,1725]" pageId="81" pageNumber="82" refId="ref103090" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. International Commission on Zoological Nomenclature. Available from: https: // code. iczn. org /? frame = 1 (accessed 19 May 2024)." type="url" year="1999">ICZN 1999</bibRefCitation>
|
||
), its oldest synonym (
|
||
<taxonomicName id="D7B24D0DFFCC391E91D491F7311EDBFE" box="[921,1071,1699,1725]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="camelicola">
|
||
<emphasis id="22C6EA9CFFCC391E91D491F7311EDBFE" box="[921,1071,1699,1725]" italics="true" pageId="81" pageNumber="82">P. camelicola</emphasis>
|
||
</taxonomicName>
|
||
) was revived and redescribed by Tanaka & Kanitani (2022) based on Japanese specimens of a distinct species hitherto misidentified as
|
||
<taxonomicName id="D7B24D0DFFCC391E9743919330A4DBA2" baseAuthorityName="Tanaka & Kamitani" baseAuthorityYear="2022" box="[1294,1429,1735,1761]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="floccifera">
|
||
<emphasis id="22C6EA9CFFCC391E9743919330A4DBA2" box="[1294,1429,1735,1761]" italics="true" pageId="81" pageNumber="82">P. floccifera</emphasis>
|
||
</taxonomicName>
|
||
. However, they could not determine which other species had been misidentified as
|
||
<taxonomicName id="D7B24D0DFFCC391E961291BF31D8DA46" baseAuthorityName="Tanaka & Kamitani" baseAuthorityYear="2022" box="[1119,1257,1771,1797]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="floccifera">
|
||
<emphasis id="22C6EA9CFFCC391E961291BF31D8DA46" box="[1119,1257,1771,1797]" italics="true" pageId="81" pageNumber="82">P. floccifera</emphasis>
|
||
</taxonomicName>
|
||
because of the difficulty of obtaining the relevant type specimens for study. Aspects of this taxonomic confusion therefore have not yet been resolved and molecular studies are needed to further elucidate the species boundaries.
|
||
</paragraph>
|
||
<paragraph id="100D368EFFCC391D928A900331A2DD96" blockId="81.[151,1437,150,2013]" lastBlockId="82.[151,1437,151,213]" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82">
|
||
As presently understood,
|
||
<taxonomicName id="D7B24D0DFFCC391E93A3900337B6DA32" box="[494,647,1879,1905]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="camelicola">
|
||
<emphasis id="22C6EA9CFFCC391E93A3900337B6DA32" box="[494,647,1879,1905]" italics="true" pageId="81" pageNumber="82">P. camelicola</emphasis>
|
||
</taxonomicName>
|
||
is close to
|
||
<taxonomicName id="D7B24D0DFFCC391E915C900336ABDA32" baseAuthorityName="Tanaka & Kamitani" baseAuthorityYear="2022" box="[785,922,1879,1905]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="floccifera">
|
||
<emphasis id="22C6EA9CFFCC391E915C900336ABDA32" box="[785,922,1879,1905]" italics="true" pageId="81" pageNumber="82">P. floccifera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D7B24D0DFFCC391E919B90033165DA32" box="[982,1108,1879,1905]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="urbicola">
|
||
<emphasis id="22C6EA9CFFCC391E919B90033165DA32" box="[982,1108,1879,1905]" italics="true" pageId="81" pageNumber="82">P. urbicola</emphasis>
|
||
</taxonomicName>
|
||
in having multilocular disc-pores each with 6‒8 loculi and similarly distributed ventral tubular ducts, each with a filamentous inner ductule and lacking a terminal gland (
|
||
<bibRefCitation id="74234B7FFFCC391E93F090CB37D4DAFA" author="Tanaka, H. & Kamitani, S." box="[445,741,1951,1977]" pageId="81" pageNumber="82" pagination="438 - 450" refId="ref109340" refString="Tanaka, H. & Kamitani, S. (2022) The identities of Pulvinaria camelicola Signoret 1873, stat. nov., and the related species, Pulvinaria floccifera (Westwood 1870) (Hemiptera: Coccidae) in Japan. Zootaxa, 5115 (3), 438 - 450. https: // doi. org / 10.11646 / zootaxa. 5115.3.8" type="journal article" year="2022">Tanaka & Kamitani 2022</bibRefCitation>
|
||
).
|
||
<taxonomicName id="D7B24D0DFFCC391E90B490CB36C4DAFA" box="[761,1013,1951,1977]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="camelicola">
|
||
<emphasis id="22C6EA9CFFCC391E90B490CB36C4DAFA" box="[761,1013,1951,1977]" italics="true" pageId="81" pageNumber="82">Pulvinaria camelicola</emphasis>
|
||
</taxonomicName>
|
||
differs from these species by having (contrasting character states of
|
||
<taxonomicName id="D7B24D0DFFCC391E93B59097374EDA9E" baseAuthorityName="Tanaka & Kamitani" baseAuthorityYear="2022" box="[504,639,1987,2013]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="floccifera">
|
||
<emphasis id="22C6EA9CFFCC391E93B59097374EDA9E" box="[504,639,1987,2013]" italics="true" pageId="81" pageNumber="82">P. floccifera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D7B24D0DFFCC391E90F590973602DA9E" box="[696,819,1987,2013]" class="Insecta" family="Coccidae" genus="Pulvinaria" kingdom="Animalia" order="Hemiptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="urbicola">
|
||
<emphasis id="22C6EA9CFFCC391E90F590973602DA9E" box="[696,819,1987,2013]" italics="true" pageId="81" pageNumber="82">P. urbicola</emphasis>
|
||
</taxonomicName>
|
||
given in parenthesis): (1) large, well-developed dorsal areolations (areolations rarely present, small and restricted to submarginal areas); and (2) dorsum with a small number of tubular ducts (dorsum mostly lacking tubular ducts) (
|
||
<bibRefCitation id="74234B7FFFCF391D912897EF31B6DD96" author="Tanaka, H. & Kamitani, S." box="[869,1159,187,213]" pageId="82" pageNumber="83" pagination="438 - 450" refId="ref109340" refString="Tanaka, H. & Kamitani, S. (2022) The identities of Pulvinaria camelicola Signoret 1873, stat. nov., and the related species, Pulvinaria floccifera (Westwood 1870) (Hemiptera: Coccidae) in Japan. Zootaxa, 5115 (3), 438 - 450. https: // doi. org / 10.11646 / zootaxa. 5115.3.8" type="journal article" year="2022">Tanaka & Kamitani 2022</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |