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<document id="97822106C77DE2E5EADFF75333E973F6" ID-CLB-Dataset="306474" ID-DOI="10.11646/zootaxa.5542.1.1" ID-GBIF-Dataset="b349e73d-c49e-418c-81f7-c5d32e136362" ID-ISSN="1175-5334" ID-Zenodo-Dep="14385386" ID-ZooBank="2DB3A5B7-4292-4CD9-B6D8-FA97EB48DD16" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1733917054948" checkinUser="plazi" docAuthor="Moghaddam, Masumeh &amp; Watson, Gillian W." docDate="2024" docId="981B8798FF1139C292DA92FB30ADDC09" docLanguage="en" docName="zootaxa.5542.1.1.pdf" docOrigin="Zootaxa 5542 (1)" docSource="https://doi.org/10.11646/zootaxa.5542.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Kermes Boitard" docType="treatment" docVersion="3" lastPageNumber="142" masterDocId="6422FFE0FF9D394F924D97543531DD43" masterDocTitle="The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families" masterLastPageNumber="202" masterPageNumber="1" pageNumber="141" updateTime="1734101643649" updateUser="ExternalLinkService">
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<mods:title id="741D616E81559AC97754A16E0B162309">The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 3 The Soft Scales (Coccidae) And Other Families</mods:title>
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<mods:namePart id="72B10AB72FE199FD211761A027ED965B">Moghaddam, Masumeh</mods:namePart>
<mods:affiliation id="270D14EDA4998F4B615B8B5B6200DC00">Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran.</mods:affiliation>
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<mods:namePart id="7472207868762DF453F9F527AF4D463D">Watson, Gillian W.</mods:namePart>
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<mods:affiliation id="CBF0C686209DD7AF6E70078C3BD66FA1">Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran. &amp; Science: Research, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. gillian. watson @ nhm. ac. uk; https: // orcid. org / 0000 - 0001 - 9914 - 0094 * Corresponding author. moghadam @ iripp. ir; https: // orcid. org / 0000 - 0003 - 0908 - 838 X &amp; Department of Insect Taxonomy Research, Iranian Research Institute of Plant Protection, Agricultural Research Education and Extension Organisation, Tehran, Iran.</mods:affiliation>
<mods:nameIdentifier id="60ABE804FCDE76B9DB7012C8E46CA0EA" type="email">gillian.watson@nhm.ac.uk</mods:nameIdentifier>
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<treatment id="981B8798FF1139C292DA92FB30ADDC09" LSID="urn:lsid:plazi:treatment:981B8798FF1139C292DA92FB30ADDC09" httpUri="http://treatment.plazi.org/id/981B8798FF1139C292DA92FB30ADDC09" lastPageId="141" lastPageNumber="142" pageId="140" pageNumber="141">
<subSubSection id="58A86505FF1139C392DA92FB34FFD88A" box="[151,462,1455,1481]" pageId="140" pageNumber="141" type="nomenclature">
<paragraph id="100D368EFF1139C392DA92FB34FFD88A" blockId="140.[151,462,1455,1481]" box="[151,462,1455,1481]" pageId="140" pageNumber="141">
<emphasis id="22C6EA9CFF1139C392DA92FB34FFD88A" bold="true" box="[151,462,1455,1481]" pageId="140" pageNumber="141">
Genus
<taxonomicName id="D7B24D0DFF1139C392A392FB34FFD88A" ID-CoL="637WL" ID-ENA="535997" authority="Boitard" authorityName="Boitard" box="[238,462,1455,1481]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="140" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="22C6EA9CFF1139C392A392FB3456D88A" bold="true" box="[238,359,1455,1481]" italics="true" pageId="140" pageNumber="141">KERMES</emphasis>
Boitard
</taxonomicName>
</emphasis>
</paragraph>
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<subSubSection id="58A86505FF1139C392DA92A134B2DB68" pageId="140" pageNumber="141" type="reference_group">
<paragraph id="100D368EFF1139C392DA92A134B2DB68" blockId="140.[151,1436,1524,1579]" pageId="140" pageNumber="141">
<treatmentCitationGroup id="30A211A0FF1139C392DA92A134B2DB68" pageId="140" pageNumber="141">
<treatmentCitation id="9113109FFF1139C392DA92A1349ADB4F" author="Boitard, P." box="[151,427,1524,1548]" page="171" pageId="140" pageNumber="141" year="1828">
<taxonomicName id="D7B24D0DFF1139C392DA92A1349ADB4F" ID-CoL="637WL" authority="Boitard, 1828: 171" authorityName="Boitard" authorityPageNumber="171" authorityYear="1828" box="[151,427,1524,1548]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="140" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="22C6EA9CFF1139C392DA92A135D5DB4F" box="[151,228,1525,1548]" italics="true" pageId="140" pageNumber="141">Kermes</emphasis>
<bibRefCitation id="74234B7FFF1139C392A792A0349ADB4F" author="Boitard, P." box="[234,427,1524,1548]" pageId="140" pageNumber="141" refId="ref98424" refString="Boitard, P. (1828) Les gallinsectes. Manuel d'entomologie ou Histoire naturelle des Insectes. Roret, Paris, 435 pp. https: // doi. org / 10.5962 / bhl. title. 122860" type="book" year="1828">Boitard, 1828: 171</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="CF09882CFF1139C393FE92A034D4DB4F" box="[435,485,1524,1548]" pageId="140" pageNumber="141">Type</typeStatus>
species:
<taxonomicName id="D7B24D0DFF1139C3900C92A03660DB4F" authority="Gmelin" authorityName="Gmelin" authorityYear="1790" box="[577,849,1524,1548]" class="Insecta" family="Coccidae" genus="Coccus" kingdom="Animalia" order="Hemiptera" pageId="140" pageNumber="141" phylum="Arthropoda" rank="species" species="variegatus">
<emphasis id="22C6EA9CFF1139C3900C92A037CCDB4F" box="[577,765,1524,1548]" italics="true" pageId="140" pageNumber="141">Coccus variegatus</emphasis>
Gmelin
</taxonomicName>
, by subsequent designation of
<treatmentCitation id="9113109FFF1139C396C892A03005DB4F" author="Fernald, M. E." box="[1157,1332,1524,1548]" page="60" pageId="140" pageNumber="141" year="1903">
<bibRefCitation id="74234B7FFF1139C396C892A03005DB4F" author="Fernald, M. E." box="[1157,1332,1524,1548]" pageId="140" pageNumber="141" pagination="1 - 360" refId="ref101196" refString="Fernald, M. E. (1903) A catalogue of the Coccidae of the world. Bulletin of the Hatch Experiment Station of the Massachusetts Agricultural College, 88, 1 - 360. https: // doi. org / 10.5962 / bhl. title. 8533" type="journal article" year="1903">Fernald 1903: 60</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="9113109FFF1139C3977092A0344EDB68" author="von Heyden, C." page="90" pageId="140" pageNumber="141" year="1860">
<taxonomicName id="D7B24D0DFF1139C3977092A0344EDB68" authority="von Heyden, 1860: 90" authorityName="von Heyden" authorityPageNumber="90" authorityYear="1860" class="Insecta" family="Kermesidae" genus="Talla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="140" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="22C6EA9CFF1139C3977092A03041DB4F" box="[1341,1392,1524,1548]" italics="true" pageId="140" pageNumber="141">Talla</emphasis>
<bibRefCitation id="74234B7FFF1139C3973B92A0344EDB68" author="von Heyden, C." pageId="140" pageNumber="141" pagination="89 - 90" refId="ref110338" refString="von Heyden, C. (1860) [From my entomological diaries. (1). On the natural history of coccids]. Korrespondenzblatt fur Sammler von Insekten, 1, 89 - 90. [in German]" type="journal article" year="1860">von Heyden, 1860: 90</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
</subSubSection>
<subSubSection id="58A86505FF1139C392DA910D30A0DA64" pageId="140" pageNumber="141" type="diagnosis">
<paragraph id="100D368EFF1139C392DA910D30A0DA64" blockId="140.[151,1437,1625,1903]" pageId="140" pageNumber="141">
<emphasis id="22C6EA9CFF1139C392DA910D37A6DB30" bold="true" box="[151,663,1625,1651]" pageId="140" pageNumber="141">Diagnostic characters of the adult female:</emphasis>
Live post-reproductive adult female
<taxonomicName id="D7B24D0DFF1139C39609910E31ABDB30" ID-CoL="637WL" box="[1092,1178,1626,1651]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="140" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="22C6EA9CFF1139C39609910E31ABDB30" box="[1092,1178,1626,1651]" italics="true" pageId="140" pageNumber="141">Kermes</emphasis>
</taxonomicName>
are heavy sclerotized, spherical or shaped like a heart, bud or gall; they are light brown to black, and do not produce an ovisac. Slide-mounted adult female with legs each with 3‒5 segments; dorsum with some conical setae and tubular ducts; anal ring may contain setae but never pores. venter with broad submarginal band of tubular ducts, with some ducts wrench shaped (
<figureCitation id="88892A0BFF1139C3931991BD348ADA40" box="[340,443,1769,1795]" captionStart="FIGURE 66" captionStartId="141.[151,250,1875,1899]" captionTargetBox="[153,1436,364,1849]" captionTargetId="figure-187@141.[151,1436,355,1852]" captionTargetPageId="141" captionText="FIGURE 66. Adult female of Kermes quercus (Linnaeus), modified from Podsiadło (2014), page 265, fig. 3, published in Polish Journal of Entomology. A, antenna; B, base of hind leg; C, anterior thoracic spiracle; D, Disc-pores each with 4‒12 loculi; E, anal ring, situated on venter; F, macroducts, varying in size, some with inner ends wrench shaped in longitudinal section." figureDoi="http://doi.org/10.5281/zenodo.14385625" httpUri="https://zenodo.org/record/14385625/files/figure.png" pageId="140" pageNumber="141">Fig. 66F</figureCitation>
); without submarginal rows of quinquelocular or bilocular pores. Irregular sclerotized spots often present on both surfaces of abdomen. Anterior spiracles usually much smaller than posterior spiracles.
</paragraph>
</subSubSection>
<subSubSection id="58A86505FF1139C3928A90653109DA2C" pageId="140" pageNumber="141" type="discussion">
<paragraph id="100D368EFF1139C3928A90653109DA2C" blockId="140.[151,1437,1625,1903]" pageId="140" pageNumber="141">
<emphasis id="22C6EA9CFF1139C3928A90653463DA08" bold="true" box="[199,338,1841,1867]" pageId="140" pageNumber="141">Comments:</emphasis>
Worldwide, 64 species are included in
<taxonomicName id="D7B24D0DFF1139C3914E906631AADA08" authority="(Garcia Morales et al. 2016)" baseAuthorityName="Garcia Morales" baseAuthorityYear="2016" box="[771,1179,1841,1867]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="140" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="22C6EA9CFF1139C3914E90663668DA08" box="[771,857,1842,1867]" italics="true" pageId="140" pageNumber="141">Kermes</emphasis>
(
<bibRefCitation id="74234B7FFF1139C3912B906531A5DA08" author="Garcia Morales M. &amp; Denno B. D. &amp; Miller D. R. &amp; Miller G. L. &amp; Ben-Dov Y. &amp; Hardy N. B." box="[870,1172,1841,1867]" pageId="140" pageNumber="141" refId="ref101399" refString="Garcia Morales M., Denno B. D., Miller D. R., Miller G. L., Ben-Dov Y. &amp; Hardy N. B. (2016) ScaleNet: a literature-based model of scale insect biology and systematics. Database. Available from: http: // scalenet. info (accessed 31 January 2024) https: // doi. org / 10.1093 / database / bav 118" type="url" year="2016">
García Morales
<emphasis id="22C6EA9CFF1139C396579066317BDA08" box="[1050,1098,1841,1867]" italics="true" pageId="140" pageNumber="141">et al</emphasis>
. 2016
</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="D7B24D0DFF1139C396EB906630ACDA08" authority="(Linnaeus)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1190,1437,1841,1867]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="140" pageNumber="141" phylum="Arthropoda" rank="species" species="quercus">
<emphasis id="22C6EA9CFF1139C396EB9066302CDA08" box="[1190,1309,1842,1867]" italics="true" pageId="140" pageNumber="141">K. quercus</emphasis>
(Linnaeus)
</taxonomicName>
and
<taxonomicName id="D7B24D0DFF1139C3928A900234C9DA2C" authority="Balachowsky" authorityName="Balachowsky" box="[199,504,1877,1903]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="140" pageNumber="141" phylum="Arthropoda" rank="species" species="spatulatus">
<emphasis id="22C6EA9CFF1139C3928A9002346BDA2C" box="[199,346,1877,1903]" italics="true" pageId="140" pageNumber="141">K. spatulatus</emphasis>
Balachowsky
</taxonomicName>
have been recorded from
<collectingCountry id="68A5761EFF1139C391509001367BDA2C" box="[797,842,1877,1903]" name="Iran" pageId="140" pageNumber="141">Iran</collectingCountry>
(
<bibRefCitation id="74234B7FFF1139C391149001311DDA2C" author="Moghaddam, M." box="[857,1068,1877,1903]" pageId="140" pageNumber="141" pagination="1 - 45" refId="ref105743" refString="Moghaddam, M. (2009) Insects of Iran: the list of Coccoidea in the Hayk Mirzayans Insect Museum of Iranian Research Institute of Plant Protection (2). Hemiptera: Coccoidea. Publications, Plant Pests &amp; Diseases Research Institute, Insect Taxonomy Research Department, No. 16, 1 - 45." type="book chapter" year="2009">Moghaddam 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="58A86505FF1039C292DA97CC30ADDC09" pageId="141" pageNumber="142" type="key">
<paragraph id="100D368EFF1039C292DA97CC36CBDDF2" blockId="141.[151,1018,151,178]" box="[151,1018,151,178]" pageId="141" pageNumber="142">
<emphasis id="22C6EA9CFF1039C292DA97CC3701DDF1" bold="true" box="[151,560,151,178]" pageId="141" pageNumber="142">
Key to species of
<taxonomicName id="D7B24D0DFF1039C2932397CC34F8DDF1" ID-CoL="637WL" box="[366,457,152,178]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="22C6EA9CFF1039C2932397CC34F8DDF1" bold="true" box="[366,457,152,178]" italics="true" pageId="141" pageNumber="142">Kermes</emphasis>
</taxonomicName>
in
<collectingCountry id="68A5761EFF1039C293BC97CC371BDDF1" box="[497,554,152,178]" name="Iran" pageId="141" pageNumber="142">Iran</collectingCountry>
,
</emphasis>
based on slide-mounted adult females.
</paragraph>
<keyStep id="AB462E2BFF1039C292DA978930ADDC09" pageId="141" pageNumber="142">
<key id="642361BFFF1039C292DA978930ADDC09" pageId="141" pageNumber="142">
<paragraph id="100D368EFF1039C292DA978930ACDC53" blockId="141.[151,1437,220,330]" pageId="141" pageNumber="142">
<keyLead id="AB4395BBFF1039C292DA978930ACDC53" pageId="141" pageNumber="142">
1(0) Antennae each with 2 or 3 segments. Legs reduced, each with 3 segments. Anal ring situated near abdominal apex; margin of ring complete. Venter with tubular ducts of 1 size............................................
<taxonomicName id="D7B24D0DFF1039C296AD97AE30ACDC53" ID-CoL="3R66V" authority="(Linnaeus)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1248,1437,250,272]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="quercus">
<emphasis id="22C6EA9CFF1039C296AD97AE301EDC4C" bold="true" box="[1248,1327,250,271]" italics="true" pageId="141" pageNumber="142">quercus</emphasis>
(Linnaeus)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="100D368EFF1039C292DA964C30ADDC09" blockId="141.[151,1437,220,330]" pageId="141" pageNumber="142">
<keyLead id="AB4395BBFF1039C292DA964C30ADDC09" pageId="141" pageNumber="142">
- Antennae each with 5 or 6 segments. Legs well developed, each with 4 or 5 segments. Anal ring dorsal; anterior margin of ring open. Venter with tubular ducts of 2 sizes...............................................
<taxonomicName id="D7B24D0DFF1039C296FD966030ADDC09" ID-CoL="3R67B" authority="Balachowsky" authorityName="Balachowsky" box="[1200,1436,308,330]" class="Insecta" family="Kermesidae" genus="Kermes" kingdom="Animalia" order="Hemiptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="spatulatus">
<emphasis id="22C6EA9CFF1039C296FD96603024DC09" bold="true" box="[1200,1301,308,330]" italics="true" pageId="141" pageNumber="142">spatulatus</emphasis>
Balachowsky
</taxonomicName>
</keyLead>
</paragraph>
</key>
</keyStep>
</subSubSection>
</treatment>
</document>
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