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<document id="80FC4E12DECEB2E7821087C276EB7D3D" ID-DOI="10.5281/zenodo.5181756" ID-ISSN="1942-1354" ID-Zenodo-Dep="5181756" ID-ZooBank="367B2C10-0F57-46E6-AAB5-EDF240370778" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628297108097" checkinUser="felipe" docAuthor="Larochelle, André, Zeperyphodes, Broun &amp; Larivière, Marie-Claude" docDate="2015" docId="980DC26FE33BFFD4FF6DFAD093D2FAD1" docLanguage="en" docName="InsectaMundi.0415.1-78.pdf" docOrigin="Insecta Mundi 2015 (415)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Bembidion (Zecillenus) puponga Larochelle &amp; Zeperyphodes &amp; Broun &amp; Larivière 2015, new species" docType="treatment" docVersion="4" lastPageNumber="16" masterDocId="6434BA17E32BFFC5FFADFFF1926CFFAB" masterDocTitle="Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini)" masterLastPageNumber="78" masterPageNumber="1" pageNumber="15" updateTime="1699041462693" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="D6F70FBAB864731421F83C1399E839B9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1F6A1DFE50FB9201F7CA3B00F1573435">
<mods:title id="FCB5792472E4877B9F48EFD1433D3E18">Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini)</mods:title>
</mods:titleInfo>
<mods:name id="4B0BEC6D3776BA9479A3C1B32BB92CEC" type="personal">
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<mods:namePart id="76BB01D15C16FE26655D50B27A9DACCE">Larochelle, André</mods:namePart>
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<mods:namePart id="4A4EA95EDF8070B1BA8233ECC57D5964">Zeperyphodes</mods:namePart>
</mods:name>
<mods:name id="F81E3F3C2AD98D5ADCBC3515058052DA" type="personal">
<mods:role id="9C1B21AB1F6CB1AA47BB4CC964481DAA">
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<mods:namePart id="79A464A0D66392DFB8CE9908989E9AA0">Broun</mods:namePart>
</mods:name>
<mods:name id="329B8BC56897A24E6EDCDAE3D37F6BF3" type="personal">
<mods:role id="02ACC14ADBA537BEFBBF8D32F18F7382">
<mods:roleTerm id="4BB46AE42EB3FA615AD0D950BD5032E9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8DFBE286703AE05CA22A184C4F95363A">Larivière, Marie-Claude</mods:namePart>
</mods:name>
<mods:typeOfResource id="2E905305A56D2C5D8263614B8CDF79A8">text</mods:typeOfResource>
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<mods:title id="666C1417E796798C91B5A8D8153DF8D3">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="050C22CA9118A9CAE61046F80727A91F">
<mods:date id="2E57C04E650101213017B8F3263DCAAB">2015</mods:date>
<mods:detail id="B7F8C1F3F4E4115A14CF07967CDEA9D7" type="pubDate">
<mods:number id="33ABDD78ABF6D1662A301FCD91372EE5">2015-04-03</mods:number>
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<mods:number id="D50FDC7B9ED8780D36CA3A923F84896C">2015</mods:number>
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<mods:number id="20D76FEC809FAC3AC16CE86BDA617EB4">415</mods:number>
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<mods:start id="1AB82D1EEA2AE9E82D893467474556DB">1</mods:start>
<mods:end id="4FD1531B9F75B5B235824B4D0EDE97FF">78</mods:end>
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<mods:classification id="2941CCD5107BEF8D88AD75362E51A344">journal article</mods:classification>
<mods:identifier id="969D0951863A6D4E88D73EDF2ADAEF57" type="DOI">10.5281/zenodo.5181756</mods:identifier>
<mods:identifier id="FFFA3C9C38B4370D508C23D437D1CFC3" type="ISSN">1942-1354</mods:identifier>
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<treatment id="980DC26FE33BFFD4FF6DFAD093D2FAD1" ID-DOI="http://doi.org/10.5281/zenodo.5190987" ID-Zenodo-Dep="5190987" LSID="urn:lsid:plazi:treatment:980DC26FE33BFFD4FF6DFAD093D2FAD1" httpUri="http://treatment.plazi.org/id/980DC26FE33BFFD4FF6DFAD093D2FAD1" lastPageId="17" lastPageNumber="16" pageId="16" pageNumber="15">
<subSubSection id="58BE20F2E33BFFD5FF6DFAD09146FA92" box="[192,810,1313,1338]" pageId="16" pageNumber="15" type="nomenclature">
<paragraph id="101B7379E33BFFD5FF6DFAD09146FA92" blockId="16.[192,810,1313,1370]" box="[192,810,1313,1338]" pageId="16" pageNumber="15">
<heading id="4B53C415E33BFFD5FF6DFAD09146FA92" bold="true" box="[192,810,1313,1338]" fontSize="10" level="2" pageId="16" pageNumber="15" reason="3">
<emphasis id="22D0AF6BE33BFFD5FF6DFAD09146FA92" bold="true" box="[192,810,1313,1338]" pageId="16" pageNumber="15">
<taxonomicName id="D7A408FAE33BFFD5FF6DFAD09011FA92" ID-CoL="5WHHQ" authority="Larochelle &amp; Zeperyphodes &amp; Broun &amp; Larivière, 2015" authorityName="Larochelle &amp; Zeperyphodes &amp; Broun &amp; Larivière" authorityYear="2015" box="[192,637,1313,1338]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="puponga" status="sp. nov." subGenus="Zecillenus">
<emphasis id="22D0AF6BE33BFFD5FF6DFAD09338FA91" bold="true" box="[192,340,1313,1338]" italics="true" pageId="16" pageNumber="15">Bembidion</emphasis>
(
<emphasis id="22D0AF6BE33BFFD5FECBFAD0939FFA91" bold="true" box="[358,499,1313,1338]" italics="true" pageId="16" pageNumber="15">Zecillenus</emphasis>
)
<emphasis id="22D0AF6BE33BFFD5FDA8FAD39011FA92" bold="true" box="[517,637,1314,1337]" italics="true" pageId="16" pageNumber="15">puponga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="39E31210E33BFFD5FD28FAD39146FA92" box="[645,810,1313,1338]" pageId="16" pageNumber="15" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="58BE20F2E33BFFD4FF6DFAB390AAFE71" lastPageId="17" lastPageNumber="16" pageId="16" pageNumber="15" type="description">
<paragraph id="101B7379E33BFFD5FF6DFAB393F6FAF1" blockId="16.[192,810,1313,1370]" box="[192,410,1345,1370]" pageId="16" pageNumber="15">
<figureCitation id="889F6FFCE33BFFD5FF6DFAB39364FAF2" box="[192,264,1345,1370]" captionStart="Figures 5–8" captionStartId="55.[192,288,1875,1897]" captionTargetBox="[223,1390,255,1848]" captionTargetId="figure-43@55.[192,1440,192,1856]" captionTargetPageId="55" captionText="Figures 5–8. Dorsal habitus. 5) Bembidion waimarama new species. 6) B. puponga new species. 7) B. tillyardi. 8) B. chalmeri. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5181760" httpUri="https://zenodo.org/record/5181760/files/figure.png" pageId="16" pageNumber="15">Fig. 6</figureCitation>
,
<figureCitation id="889F6FFCE33BFFD5FEB8FAB39359FAF1" box="[277,309,1346,1370]" captionStart="Figures 37–44" captionStartId="63.[192,288,1875,1897]" captionTargetBox="[251,1397,255,1850]" captionTargetId="figure-56@63.[192,1441,192,1857]" captionTargetPageId="63" captionText="Figures 37–44. Aedeagi, lateral. 37) Bembidion karikari new species. 38) B. albescens. 39) B. alacre. 40) B. tepaki new species. 41) B. waimarama new species. 42) B. puponga new species. 43) B. tillyardi. 44) B. chalmeri." figureDoi="http://doi.org/10.5281/zenodo.5181776" httpUri="https://zenodo.org/record/5181776/files/figure.png" pageId="16" pageNumber="15">42</figureCitation>
,
<figureCitation id="889F6FFCE33BFFD5FEECFAB3930DFAF1" box="[321,353,1346,1370]" captionStart="Figures 89–94" captionStartId="71.[192,287,1875,1897]" captionTargetBox="[192,1440,192,1857]" captionTargetId="figure-47@71.[192,1440,192,1857]" captionTargetPageId="71" captionText="Figures 89–94. Species distribution maps. 89) Bembidion maorinum maorinum. 90) B. musae. 91) B. nesophilum. 92) B. orbiferum. 93) B. parviceps. 94) B. puponga new species." figureDoi="http://doi.org/10.5281/zenodo.5181792" httpUri="https://zenodo.org/record/5181792/files/figure.png" pageId="16" pageNumber="15">94</figureCitation>
,
<figureCitation id="889F6FFCE33BFFD5FEC3FAB393F6FAF1" box="[366,410,1346,1370]" captionStart="Figures 111–112" captionStartId="76.[192,288,1875,1897]" captionTargetBox="[342,1303,255,1800]" captionTargetId="figure-48@76.[192,1440,192,1872]" captionTargetPageId="76" captionText="Figures 111–112. Species habitats, Bembidion (Zecillenus). 111) Hawkes Bay (HB), Waimarama, B. waimarama new species. 112) Nelson (NN), Puponga Farm Park, B. puponga new species." figureDoi="http://doi.org/10.5281/zenodo.5181802" httpUri="https://zenodo.org/record/5181802/files/figure.png" pageId="16" pageNumber="15">112</figureCitation>
</paragraph>
<paragraph id="101B7379E33BFFD5FF6DFA7390F2F991" blockId="16.[192,1440,1410,1594]" pageId="16" pageNumber="15">
<taxonomicName id="D7A408FAE33BFFD5FF6DFA73916CFA31" authority="Larochelle and Lariviere" authorityName="Larochelle and Lariviere" authorityYear="2015" box="[192,768,1410,1434]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="puponga" status="sp. nov.">
<emphasis id="22D0AF6BE33BFFD5FF6DFA7393D6FA32" box="[192,442,1410,1434]" italics="true" pageId="16" pageNumber="15">Bembidion puponga</emphasis>
<materialsCitation id="A0CC7924E33BFFD5FE68FA73916CFA31" accessionNumber="S 17239" box="[453,768,1410,1434]" collectingDate="2003-03-04" collectionCode="NZAC, NN, CMNZ" country="New Zealand" county="Green Hills Stream" elevation="2" location="Puponga Forest" municipality="Park" pageId="16" pageNumber="15" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">Larochelle and Larivière</materialsCitation>
</taxonomicName>
,
<taxonomicNameLabel id="39E31210E33BFFD5FCBDFA7391C4FA31" box="[784,936,1410,1434]" pageId="16" pageNumber="15" rank="species">new species</taxonomicNameLabel>
.
<typeStatus id="CF1FCDDBE33BFFD5FC1BFA739645FA31" box="[950,1065,1410,1434]" pageId="16" pageNumber="15" type="holotype">Holotype</typeStatus>
: male (NZAC) labeled: “
<collectingCountry id="68B333E9E33BFFD5FAF0FA73932AFA11" name="New Zealand" pageId="16" pageNumber="15">NEW ZEALAND</collectingCountry>
NN
<location id="157B25A2E33BFFD5FE29FA539024FA12" LSID="urn:lsid:plazi:treatment:980DC26FE33BFFD4FF6DFAD093D2FAD1:157B25A2E33BFFD5FE29FA539024FA12" box="[388,584,1442,1466]" country="New Zealand" county="Green Hills Stream" municipality="Park" name="Puponga Forest" pageId="16" pageNumber="15">Puponga Forest</location>
[=
<location id="157B25A2E33BFFD5FDC4FA5390DCFA12" LSID="urn:lsid:plazi:treatment:980DC26FE33BFFD4FF6DFAD093D2FAD1:157B25A2E33BFFD5FDC4FA5390DCFA12" box="[617,688,1442,1465]" country="New Zealand" county="Green Hills Stream" municipality="Park" name="Farm" pageId="16" pageNumber="15">Farm</location>
]
<collectingMunicipality id="F07FE903E33BFFD5FD6DFA539093FA11" box="[704,767,1442,1466]" pageId="16" pageNumber="15">Park</collectingMunicipality>
,
<collectingCounty id="F97A0BF5E33BFFD5FCA1FA53966CFA11" box="[780,1024,1442,1466]" pageId="16" pageNumber="15">Green Hills Stream</collectingCounty>
mouth 4030
<accessionNumber id="0FF7EE9AE33BFFD5FB0DFA53976EFA11" box="[1184,1282,1442,1466]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S17239" pageId="16" pageNumber="15" type="EnaNcbi">S 17239</accessionNumber>
E
<date id="641A55B9E33BFFD5FA8BFA5397CCFA11" box="[1318,1440,1442,1466]" pageId="16" pageNumber="15" value="2003-03-04">
<collectingDate id="745EAC51E33BFFD5FA8BFA5397CCFA11" box="[1318,1440,1442,1466]" pageId="16" pageNumber="15" value="2003-03-04">4.III.2003</collectingDate>
</date>
Larochelle &amp; Larivière (typed)/ Estuary stream and lagoon: wet, bare, white-silver sand flat near dune;
<quantity id="D75CDE9CE33BFFD5FF6DFA139368FA52" box="[192,260,1506,1530]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="16" pageNumber="15" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">
<elevation id="9B89944AE33BFFD5FF6DFA139368FA52" box="[192,260,1506,1530]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="16" pageNumber="15" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1-2 m</elevation>
</quantity>
from water (typed)/
<typeStatus id="CF1FCDDBE33BFFD5FE54FA1390F8FA51" box="[505,660,1506,1530]" pageId="16" pageNumber="15" type="holotype">HOLOTYPE</typeStatus>
[male symbol]
<taxonomicName id="D7A408FAE33BFFD5FCE4FA1397CCFA52" authority="Larochelle &amp; Lariviere, 2015" authorityName="Larochelle &amp; Lariviere" authorityYear="2015" box="[841,1440,1505,1530]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="puponga">
<emphasis id="22D0AF6BE33BFFD5FCE4FA139652FA52" box="[841,1086,1506,1530]" italics="true" pageId="16" pageNumber="15">Bembidion puponga</emphasis>
Larochelle &amp; Larivière, 2015
</taxonomicName>
(red label; typed).”
<typeStatus id="CF1FCDDBE33BFFD5FE04F9F39046F9B1" box="[425,554,1538,1562]" pageId="16" pageNumber="15" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="06A2B8F0E33BFFD5FD98F9F390CFF9B1" box="[565,675,1538,1562]" pageId="16" pageNumber="15" type="male">one male</specimenCount>
(CMNZ) and
<specimenCount id="06A2B8F0E33BFFD5FCE1F9F391BCF9B1" box="[844,976,1538,1562]" pageId="16" pageNumber="15" type="female">one female</specimenCount>
(NZAC) from the same locality as the holotype, bearing blue
<typeStatus id="CF1FCDDBE33BFFD5FE74F9D3902BF991" box="[473,583,1570,1594]" pageId="16" pageNumber="15" type="paratype">paratype</typeStatus>
labels.
</paragraph>
<paragraph id="101B7379E33BFFD4FF6DF99090AAFE71" blockId="16.[192,1441,1633,1914]" lastBlockId="17.[192,1441,194,474]" lastPageId="17" lastPageNumber="16" pageId="16" pageNumber="15">
<emphasis id="22D0AF6BE33BFFD5FF6DF990930FF9D2" bold="true" box="[192,355,1633,1657]" pageId="16" pageNumber="15">Description</emphasis>
. Body length 3.0–
<quantity id="D75CDE9CE33BFFD5FDE8F99090C9F9D2" box="[581,677,1633,1657]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="16" pageNumber="15" unit="mm" value="4.5">4.5 mm</quantity>
. Head and pronotum black or piceous; elytra pale testaceous, with a reduced brownish irregular discal marking (reaching interval 6) and interval 2 infuscated behind middle; antennae with segments 1–4 pale testaceous and segments 5–11 infuscated; legs pale testaceous, with apex of femora and base of tibiae slightly infuscated; abdomen piceous. Microsculpture strong and moderately transverse on disc of head; obsolete, with traces of transverse meshes (microlines) on disc of pronotum; strong on elytra, isodiametric on disc, granulate elsewhere. Head, pronotum, and disc of elytra shiny, remainder of elytra dull; metallic lustre present on head and pronotum (aeneous).
<emphasis id="22D0AF6BE33BFFD5FF6DF8B39348F8F1" bold="true" box="[192,292,1858,1882]" pageId="16" pageNumber="15">Thorax</emphasis>
. Pronotum widest before middle; sides moderately rounded anteriorly, not sinuate posteriorly; posterolateral angles rectangular; laterobasal foveae shallow, ill-defined (rounded), short, not reaching basal margin.
<emphasis id="22D0AF6BE33AFFD4FEDFFF3393A5FF71" bold="true" box="[370,457,194,218]" pageId="17" pageNumber="16">Elytra</emphasis>
. Shorter than in other
<taxonomicName id="D7A408FAE33AFFD4FD46FF3391AAFF71" box="[747,966,194,218]" class="Insecta" family="Carabidae" genus="Zecillenus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="22D0AF6BE33AFFD4FD46FF33910BFF71" box="[747,871,194,218]" italics="true" pageId="17" pageNumber="16">Zecillenus</emphasis>
species
</taxonomicName>
(except
<taxonomicName id="D7A408FAE33AFFD4FB83FF3396DAFF71" baseAuthorityName="Brookes" baseAuthorityYear="1927" box="[1070,1206,194,218]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="tillyardi">
<emphasis id="22D0AF6BE33AFFD4FB83FF3396DAFF71" box="[1070,1206,194,218]" italics="true" pageId="17" pageNumber="16">B. tillyardi</emphasis>
</taxonomicName>
). Elliptical, widest about middle. Shoulders prominent. Sides moderately rounded. Striae incomplete, barely suggested and impunctate laterally and apically; striae 2–5 poorly impressed, shallow and finely punctate on disc; striae 6–7 obsolete. Intervals subdepressed on disc. Lateral margins slightly widened subapically; subapical tooth obtuse. Subapical sinuations moderately strong. Sutural apices obtusely rounded. Apex strongly rounded.
<emphasis id="22D0AF6BE33AFFD4FE9AFE9093D0FED2" bold="true" box="[311,444,353,377]" pageId="17" pageNumber="16">Aedeagus</emphasis>
. Lateral view (
<figureCitation id="889F6FFCE33AFFD4FDD1FE9390BAFED1" box="[636,726,354,378]" captionStart="Figures 37–44" captionStartId="63.[192,288,1875,1897]" captionTargetBox="[251,1397,255,1850]" captionTargetId="figure-56@63.[192,1441,192,1857]" captionTargetPageId="63" captionText="Figures 37–44. Aedeagi, lateral. 37) Bembidion karikari new species. 38) B. albescens. 39) B. alacre. 40) B. tepaki new species. 41) B. waimarama new species. 42) B. puponga new species. 43) B. tillyardi. 44) B. chalmeri." figureDoi="http://doi.org/10.5281/zenodo.5181776" httpUri="https://zenodo.org/record/5181776/files/figure.png" pageId="17" pageNumber="16">Fig. 42</figureCitation>
): moderately strongly arcuate, slightly widened in apical half; base moderately convex dorsally; middle moderately convex dorsally, slightly concave towards its apex, almost straight ventrally towards its apex; apex subtriangular, moderately concave dorsally and ventrally, with extreme tip wide and long.
</paragraph>
</subSubSection>
<subSubSection id="58BE20F2E33AFFD4FF6DFDF0934DFD11" pageId="17" pageNumber="16" type="materials_examined">
<paragraph id="101B7379E33AFFD4FF6DFDF0966FFDB1" blockId="17.[192,1027,513,538]" box="[192,1027,513,538]" pageId="17" pageNumber="16">
<emphasis id="22D0AF6BE33AFFD4FF6DFDF093A9FDB2" bold="true" box="[192,453,513,537]" pageId="17" pageNumber="16">Material examined</emphasis>
.
<specimenCount id="06A2B8F0E33AFFD4FE79FDF39014FDB1" box="[468,632,514,538]" pageId="17" pageNumber="16" type="generic">93 specimens</specimenCount>
(
<collectionCode id="76B5EBBCE33AFFD4FD2AFDF390B2FDB1" box="[647,734,514,538]" country="New Zealand" httpUri="http://grbio.org/cool/n6z3-qvv0" name="Canterbury Museum" pageId="17" pageNumber="16">CMNZ</collectionCode>
,
<collectionCode id="76B5EBBCE33AFFD4FD44FDF3915BFDB2" box="[745,823,514,537]" pageId="17" pageNumber="16">JNNZ</collectionCode>
,
<collectionCode id="76B5EBBCE33AFFD4FCEEFDF391F6FDB1" box="[835,922,514,538]" country="New Zealand" httpUri="http://grbio.org/cool/15kb-kde6" name="Museum of New Zealand Te Papa Tongarewa - Entomology" pageId="17" pageNumber="16" type="Museum">MONZ</collectionCode>
,
<collectionCode id="76B5EBBCE33AFFD4FC0BFDF39194FDB1" box="[934,1016,514,538]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="17" pageNumber="16" type="Other institutional type">NZAC</collectionCode>
).
</paragraph>
<paragraph id="101B7379E33AFFD4FF6DFDB0934DFD11" blockId="17.[192,1440,577,698]" pageId="17" pageNumber="16">
<emphasis id="22D0AF6BE33AFFD4FF6DFDB09063FDF2" bold="true" box="[192,527,577,601]" pageId="17" pageNumber="16">Geographic distribution</emphasis>
(
<figureCitation id="889F6FFCE33AFFD4FDB2FDB3901BFDF1" box="[543,631,578,602]" captionStart="Figures 89–94" captionStartId="71.[192,287,1875,1897]" captionTargetBox="[192,1440,192,1857]" captionTargetId="figure-47@71.[192,1440,192,1857]" captionTargetPageId="71" captionText="Figures 89–94. Species distribution maps. 89) Bembidion maorinum maorinum. 90) B. musae. 91) B. nesophilum. 92) B. orbiferum. 93) B. parviceps. 94) B. puponga new species." figureDoi="http://doi.org/10.5281/zenodo.5181792" httpUri="https://zenodo.org/record/5181792/files/figure.png" pageId="17" pageNumber="16">Fig. 94</figureCitation>
). North Island: WI–Santoft Forest (Koitiata Stream mouth).
<collectionCode id="76B5EBBCE33AFFD4FACCFDB397FDFDF2" box="[1377,1425,578,601]" pageId="17" pageNumber="16">WN</collectionCode>
– Kuku Beach (near Levin) [=Kuku Beach Road end beach]. Lake Waiorongomai Stream mouth. Titahi Bay. South Island:
<collectionCode id="76B5EBBCE33AFFD4FE06FD7393BBFD31" box="[427,471,642,666]" pageId="17" pageNumber="16">NN</collectionCode>
–Puponga Forest [=Farm] Park (Green Hills Stream mouth; Wharariki Stream mouth).
</paragraph>
</subSubSection>
<subSubSection id="58BE20F2E33AFFD4FF6DFD139690FB91" pageId="17" pageNumber="16" type="biology_ecology">
<paragraph id="101B7379E33AFFD4FF6DFD13918EFCF1" blockId="17.[192,1440,738,858]" pageId="17" pageNumber="16">
<emphasis id="22D0AF6BE33AFFD4FF6DFD139342FD51" bold="true" box="[192,302,738,762]" pageId="17" pageNumber="16">Ecology</emphasis>
. Coastal lowland. Fossorial. Banks of meandering estuarine streams (
<quantity id="D75CDE9CE33AFFD4FB31FD139684FD52" box="[1180,1256,738,762]" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="17" pageNumber="16" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">1–2 m</quantity>
wide) running through sand dunes (
<figureCitation id="889F6FFCE33AFFD4FE60FCF3905AFCB1" box="[461,566,770,794]" captionStart="Figures 111–112" captionStartId="76.[192,288,1875,1897]" captionTargetBox="[342,1303,255,1800]" captionTargetId="figure-48@76.[192,1440,192,1872]" captionTargetPageId="76" captionText="Figures 111–112. Species habitats, Bembidion (Zecillenus). 111) Hawkes Bay (HB), Waimarama, B. waimarama new species. 112) Nelson (NN), Puponga Farm Park, B. puponga new species." figureDoi="http://doi.org/10.5281/zenodo.5181802" httpUri="https://zenodo.org/record/5181802/files/figure.png" pageId="17" pageNumber="16">Fig. 112</figureCitation>
), just above the highest tidal line, at a certain distance (
<quantity id="D75CDE9CE33AFFD4FAA8FCF39738FCB1" box="[1285,1364,770,794]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="17" pageNumber="16" unit="m" value="2.5" valueMax="3.0" valueMin="2.0">2–3 m</quantity>
) from water. Open ground; wet, sandy (white, black), bare soil. Nocturnal; hides during the day in the sand. Gregarious. Occurs in association with staphylinids (
<taxonomicName id="D7A408FAE33AFFD4FCE1FCB391BBFCF1" authorityName="Linnaeus" authorityYear="1758" box="[844,983,834,858]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
).
</paragraph>
<paragraph id="101B7379E33AFFD4FF6DFC70973FFC11" blockId="17.[192,1440,897,954]" pageId="17" pageNumber="16">
<emphasis id="22D0AF6BE33AFFD4FF6DFC709344FC32" bold="true" box="[192,296,897,921]" pageId="17" pageNumber="16">Biology</emphasis>
. Seasonality: September, December, March. Tenerals: December, March. Occasionally infested with fungi (Laboulbeniales). Defence mechanism: when alarmed, the adult escapes by running.
</paragraph>
<paragraph id="101B7379E33AFFD4FF6DFC1097CDFC51" blockId="17.[192,1441,993,1018]" box="[192,1441,993,1018]" pageId="17" pageNumber="16">
<emphasis id="22D0AF6BE33AFFD4FF6DFC1093F1FC51" bold="true" box="[192,413,993,1018]" pageId="17" pageNumber="16">Dispersal power</emphasis>
. Brachypterous (incapable of flight). Fast runner. Vagility limited by flight incapacity.
</paragraph>
<paragraph id="101B7379E33AFFD4FF6DFBD09690FB91" blockId="17.[192,1276,1057,1082]" box="[192,1276,1057,1082]" pageId="17" pageNumber="16">
<emphasis id="22D0AF6BE33AFFD4FF6DFBD09381FB92" bold="true" box="[192,493,1057,1081]" pageId="17" pageNumber="16">Collecting techniques</emphasis>
. Pouring water over the ground; treading the soil with the feet.
</paragraph>
</subSubSection>
<subSubSection id="58BE20F2E33AFFD4FF6DFB9393D2FAD1" pageId="17" pageNumber="16" type="discussion">
<paragraph id="101B7379E33AFFD4FF6DFB9393D2FAD1" blockId="17.[192,1440,1122,1402]" pageId="17" pageNumber="16">
<emphasis id="22D0AF6BE33AFFD4FF6DFB939350FBD1" bold="true" box="[192,316,1122,1146]" pageId="17" pageNumber="16">Remarks</emphasis>
. This species is named after its
<typeStatus id="CF1FCDDBE33AFFD4FD4FFB93917BFBD1" box="[738,791,1122,1146]" pageId="17" pageNumber="16">type</typeStatus>
locality, Puponga Farm Park.
<taxonomicName id="D7A408FAE33AFFD4FB08FB9397CCFBD2" authorityName="Larochelle &amp; Lariviere" authorityYear="2015" box="[1189,1440,1122,1146]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="puponga">
<emphasis id="22D0AF6BE33AFFD4FB08FB9397CCFBD2" box="[1189,1440,1122,1146]" italics="true" pageId="17" pageNumber="16">Bembidion puponga</emphasis>
</taxonomicName>
is morphologically close to
<taxonomicName id="D7A408FAE33AFFD4FDBCFB7390F7FB31" baseAuthorityName="Brookes" baseAuthorityYear="1927" box="[529,667,1154,1178]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="tillyardi">
<emphasis id="22D0AF6BE33AFFD4FDBCFB7390F7FB31" box="[529,667,1154,1178]" italics="true" pageId="17" pageNumber="16">B. tillyardi</emphasis>
</taxonomicName>
. In addition to diagnostic characters of the male genitalia,
<taxonomicName id="D7A408FAE33AFFD4FA28FB739346FB12" authorityName="Larochelle &amp; Lariviere" authorityYear="2015" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="puponga">
<emphasis id="22D0AF6BE33AFFD4FA28FB739346FB12" italics="true" pageId="17" pageNumber="16">B. puponga</emphasis>
</taxonomicName>
has the following distinguishing features: head and pronotum black or piceous; elytra pale testaceous with a reduced brownish irregular discal marking and interval 2 infuscated behind middle; legs pale testaceous with apex of femora and base of tibiae slightly infuscated; abdomen piceous; elytra with intervals subdepressed on disc and subapical sinuations moderately strong.
<taxonomicName id="D7A408FAE33AFFD4FB04FAF397CCFAB2" authorityName="Larochelle &amp; Lariviere" authorityYear="2015" box="[1193,1440,1282,1306]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="puponga">
<emphasis id="22D0AF6BE33AFFD4FB04FAF397CCFAB2" box="[1193,1440,1282,1306]" italics="true" pageId="17" pageNumber="16">Bembidion puponga</emphasis>
</taxonomicName>
occurs in the southwestern North Island (WI, WN) and northwestern South Island (NN, Puponga Farm Park) while
<taxonomicName id="D7A408FAE33AFFD4FE32FAB3904AFAF1" baseAuthorityName="Brookes" baseAuthorityYear="1927" box="[415,550,1346,1370]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="tillyardi">
<emphasis id="22D0AF6BE33AFFD4FE32FAB3904AFAF1" box="[415,550,1346,1370]" italics="true" pageId="17" pageNumber="16">B. tillyardi</emphasis>
</taxonomicName>
is restricted to a single locality from the northwestern South Island (NN,
<collectingRegion id="D260BD9BE33AFFD4FF6DFA939374FAD1" box="[192,280,1378,1402]" country="New Zealand" name="Nelson" pageId="17" pageNumber="16">Nelson</collectingRegion>
, Tahunanui).
</paragraph>
</subSubSection>
</treatment>
</document>
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