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<document id="8EED37B64F933A0CEC9F19F7D402C9ED" ID-DOI="10.5281/zenodo.5181756" ID-ISSN="1942-1354" ID-Zenodo-Dep="5181756" ID-ZooBank="367B2C10-0F57-46E6-AAB5-EDF240370778" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628297108097" checkinUser="felipe" docAuthor="Larochelle, André, Zeperyphodes, Broun &amp; Larivière, Marie-Claude" docDate="2015" docId="980DC26FE305FFEAFF6DFA10939CFB31" docLanguage="en" docName="InsectaMundi.0415.1-78.pdf" docOrigin="Insecta Mundi 2015 (415)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Bembidion (Zemetallina) chalceipes Bates 1878" docType="treatment" docVersion="4" lastPageNumber="46" masterDocId="6434BA17E32BFFC5FFADFFF1926CFFAB" masterDocTitle="Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini)" masterLastPageNumber="78" masterPageNumber="1" pageNumber="45" updateTime="1699041462693" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="3F742266B156B09700E0942625DE33F7">Synopsis of the genus Bembidion Latreille in New Zealand (Coleoptera: Carabidae: Bembidiini)</mods:title>
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<mods:namePart id="2FEDC2DBFD6B7FFE0A76D5904022E747">Larochelle, André</mods:namePart>
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<mods:namePart id="013087EC4C12FD32383F3B552D8BAC1A">Larivière, Marie-Claude</mods:namePart>
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<paragraph id="101B7379E305FFEBFF6DFA109120FA52" blockId="46.[192,844,1505,1562]" box="[192,844,1505,1530]" pageId="46" pageNumber="45">
<heading id="4B53C415E305FFEBFF6DFA109120FA52" bold="true" box="[192,844,1505,1530]" fontSize="10" level="2" pageId="46" pageNumber="45" reason="3">
<taxonomicName id="D7A408FAE305FFEBFF6DFA109120FA52" ID-CoL="5WHJ7" authority="Bates, 1878" authorityName="Bates" authorityYear="1878" box="[192,844,1505,1530]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="45" phylum="Arthropoda" rank="species" species="chalceipes" subGenus="Zemetallina">
<emphasis id="22D0AF6BE305FFEBFF6DFA109338FA51" bold="true" box="[192,340,1505,1530]" italics="true" pageId="46" pageNumber="45">Bembidion</emphasis>
(
<emphasis id="22D0AF6BE305FFEBFEC9FA109067FA51" bold="true" box="[356,523,1505,1530]" italics="true" pageId="46" pageNumber="45">Zemetallina</emphasis>
)
<emphasis id="22D0AF6BE305FFEBFDB1FA109120FA52" bold="true" box="[540,844,1505,1530]" pageId="46" pageNumber="45">
<emphasis id="22D0AF6BE305FFEBFDB1FA1090C7FA51" bold="true" box="[540,683,1505,1530]" italics="true" pageId="46" pageNumber="45">chalceipes</emphasis>
Bates, 1878
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="58BE20F2E305FFEBFF6DF9F39302F9B1" box="[192,366,1537,1562]" pageId="46" pageNumber="45" type="description">
<paragraph id="101B7379E305FFEBFF6DF9F39302F9B1" blockId="46.[192,844,1505,1562]" box="[192,366,1537,1562]" pageId="46" pageNumber="45">
<figureCitation id="889F6FFCE305FFEBFF6DF9F3937BF9B2" box="[192,279,1537,1562]" captionStart="Figures 3336" captionStartId="62.[192,288,1875,1897]" captionTargetBox="[238,1427,233,1800]" captionTargetId="figure-47@62.[192,1440,213,1867]" captionTargetPageId="62" captionText="Figures 3336. Dorsal habitus. 33) Bembidion bullerense new species. 34) B. hokitikense. 35) B. chalceipes. 36) B. wanakense. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5181774" httpUri="https://zenodo.org/record/5181774/files/figure.png" pageId="46" pageNumber="45">Fig. 35</figureCitation>
,
<figureCitation id="889F6FFCE305FFEBFE89F9F09328F9B2" box="[292,324,1537,1561]" captionStart="Figures 6169" captionStartId="66.[192,288,1846,1868]" captionTargetBox="[249,1403,200,1826]" captionTargetId="figure-64@66.[192,1424,192,1834]" captionTargetPageId="66" captionText="Figures 6169. Aedeagi, lateral. 61) Bembidion parviceps. 62) B. waiho new species. 63) B. urewerense. 64) B. mangamuka new species. 65) B. solitarium. 66) B. bullerense new species. 67) B. hokitikense. 68) B. chalceipes. 69) B. wanakense." figureDoi="http://doi.org/10.5281/zenodo.5181782" httpUri="https://zenodo.org/record/5181782/files/figure.png" pageId="46" pageNumber="45">68</figureCitation>
,
<figureCitation id="889F6FFCE305FFEBFEFDF9F39302F9B1" box="[336,366,1538,1562]" captionStart="Figures 7782" captionStartId="69.[192,288,1875,1897]" captionTargetBox="[192,1440,192,1857]" captionTargetId="figure-45@69.[192,1440,192,1857]" captionTargetPageId="69" captionText="Figures 7782. Species distribution maps. 77) Bembidion callipeplum. 78) B. chalceipes. 79) B. chalmeri. 80) B. charile. 81) B. dehiscens. 82) B. embersoni." figureDoi="http://doi.org/10.5281/zenodo.5181788" httpUri="https://zenodo.org/record/5181788/files/figure.png" pageId="46" pageNumber="45">78</figureCitation>
</paragraph>
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<subSubSection id="58BE20F2E305FFEBFF6DF9B391D1F9D1" pageId="46" pageNumber="45" type="reference_group">
<paragraph id="101B7379E305FFEBFF6DF9B396CEF9F1" blockId="46.[192,1186,1602,1658]" box="[192,1186,1602,1626]" pageId="46" pageNumber="45">
<treatmentCitationGroup id="30B45457E305FFEBFF6DF9B396CEF9F1" box="[192,1186,1602,1626]" pageId="46" pageNumber="45">
<treatmentCitation id="91055568E305FFEBFF6DF9B390C5F9F1" author="Bates, H. W." box="[192,681,1602,1626]" page="24" pageId="46" pageNumber="45" year="1878">
<taxonomicName id="D7A408FAE305FFEBFF6DF9B390C5F9F1" authority="Bates, 1878 b: 24" authorityName="Bates" authorityPageNumber="24" authorityYear="1878" box="[192,681,1602,1626]" class="Insecta" family="Carabidae" genus="Bembidium" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="45" phylum="Arthropoda" rank="species" species="chalceipes">
<emphasis id="22D0AF6BE305FFEBFF6DF9B393BFF9F1" box="[192,467,1602,1626]" italics="true" pageId="46" pageNumber="45">Bembidium chalceipes</emphasis>
<bibRefCitation id="74350E88E305FFEBFE77F9B390C5F9F1" author="Bates, H. W." box="[474,681,1602,1626]" pageId="46" pageNumber="45" pagination="57 - 58" refId="ref32928" refString="Bates, H. W. 1878 b. New species of geodephagous Coleoptera from New Zealand. Entomologist's Monthly Magazine 15: 22 - 28, 57 - 58." type="journal article" year="1878">Bates, 1878b: 24</bibRefCitation>
</taxonomicName>
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.
<typeStatus id="CF1FCDDBE305FFEBFD18F9B3909EF9F1" box="[693,754,1602,1626]" pageId="46" pageNumber="45">Type</typeStatus>
locality:
<collectingRegion id="D260BD9BE305FFEBFCCFF9B3919CF9F1" box="[866,1008,1602,1626]" country="New Zealand" name="West Coast" pageId="46" pageNumber="45">West Coast</collectingRegion>
, South Island.
</treatmentCitationGroup>
</paragraph>
<paragraph id="101B7379E305FFEBFF6DF99391D1F9D1" blockId="46.[192,1186,1602,1658]" box="[192,957,1634,1658]" pageId="46" pageNumber="45">
Other synonymy as in
<bibRefCitation id="74350E88E305FFEBFE7AF99391D5F9D1" author="Larochelle, A. &amp; M. - C. Lariviere" box="[471,953,1634,1658]" pageId="46" pageNumber="45" pagination="1 - 281" refId="ref33655" refString="Larochelle, A., and M. - C. Lariviere. 2001. Carabidae (Insecta: Coleoptera): catalogue. Fauna of New Zealand 43: 1 - 281." type="journal article" year="2001">Larochelle and Larivière (2001: 8182)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="58BE20F2E305FFEAFF6DF9509644FE71" lastPageId="47" lastPageNumber="46" pageId="46" pageNumber="45" type="description">
<paragraph id="101B7379E305FFEAFF6DF9509644FE71" blockId="46.[192,1441,1697,1914]" lastBlockId="47.[192,1440,194,474]" lastPageId="47" lastPageNumber="46" pageId="46" pageNumber="45">
<emphasis id="22D0AF6BE305FFEBFF6DF950930EF912" bold="true" box="[192,354,1697,1721]" pageId="46" pageNumber="45">Description</emphasis>
. Body length
<quantity id="D75CDE9CE305FFEBFDABF95090FBF912" box="[518,663,1697,1721]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="5.2" metricValueMin="4.4" pageId="46" pageNumber="45" unit="mm" value="4.8" valueMax="5.2" valueMin="4.4">4.45.2 mm</quantity>
. Black; antennae black (segment 1 rufous underneath, segments 23 rarely rufous underneath); legs black to piceous (tibiae rarely rufous). Microsculpture very strong, isodiametric. Shiny, with strong metallic lustre (aeneous, rarely greenish); tibiae sometimes with slight metallic lustre. Forebody moderately wide in comparison to elytra.
<emphasis id="22D0AF6BE305FFEBFC4DF8F39644F8B1" bold="true" box="[992,1064,1794,1818]" pageId="46" pageNumber="45">Head</emphasis>
. Antennae filiform, moderately long (reaching about elytral shoulders).
<emphasis id="22D0AF6BE305FFEBFD01F8D3917CF891" bold="true" box="[684,784,1826,1850]" pageId="46" pageNumber="45">Thorax</emphasis>
. Pronotum strongly convex, very wide; sides strongly rounded anteriorly, moderately sinuate posteriorly; posterolateral angles rectangular, vaguely obtuse at tip; laterobasal foveae moderately deep, linear, moderately long, reaching or almost reaching basal margin. Epipleura (in dorsal view) exposed in front of posterolateral angles.
<emphasis id="22D0AF6BE304FFEAFBFAFF3396C2FF71" bold="true" box="[1111,1198,194,218]" pageId="47" pageNumber="46">Elytra</emphasis>
. Moderately convex (somewhat less than in
<taxonomicName id="D7A408FAE304FFEAFE4EFF1390DCFF51" authorityName="Bates" authorityYear="1878" box="[483,688,226,250]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="46" phylum="Arthropoda" rank="species" species="anchonoderus">
<emphasis id="22D0AF6BE304FFEAFE4EFF1390DCFF51" box="[483,688,226,250]" italics="true" pageId="47" pageNumber="46">B. anchonoderus</emphasis>
</taxonomicName>
), subovate, moderately wide, widest about middle. Shoulders moderately rounded. Sides moderately rounded. Scutellar striole consisting of a row of punctures. Striae 23 complete, 46 incomplete; striae shallow, finely punctate; stria 7 obsolete, incomplete, at most with rudimentary punctures. Intervals depressed. Apical striole shallow, connected to stria 5; preapical setiferous puncture not isolated.
<emphasis id="22D0AF6BE304FFEAFD9FFE9090D9FED2" bold="true" box="[562,693,353,377]" pageId="47" pageNumber="46">Abdomen</emphasis>
. Last visible sternum (sternum VII) of female with four long ambulatory setae only.
<emphasis id="22D0AF6BE304FFEAFE4FFE70900BFE32" bold="true" box="[482,615,385,409]" pageId="47" pageNumber="46">Aedeagus</emphasis>
. Lateral view (
<figureCitation id="889F6FFCE304FFEAFC8EFE739110FE32" box="[803,892,385,410]" captionStart="Figures 6169" captionStartId="66.[192,288,1846,1868]" captionTargetBox="[249,1403,200,1826]" captionTargetId="figure-64@66.[192,1424,192,1834]" captionTargetPageId="66" captionText="Figures 6169. Aedeagi, lateral. 61) Bembidion parviceps. 62) B. waiho new species. 63) B. urewerense. 64) B. mangamuka new species. 65) B. solitarium. 66) B. bullerense new species. 67) B. hokitikense. 68) B. chalceipes. 69) B. wanakense." figureDoi="http://doi.org/10.5281/zenodo.5181782" httpUri="https://zenodo.org/record/5181782/files/figure.png" pageId="47" pageNumber="46">Fig. 68</figureCitation>
): strongly arcuate, gradually narrowed from base to apex; base moderately concave dorsally; middle strongly convex dorsally, moderately concave ventrally; apex triangular, with extreme tip moderately wide and long.
</paragraph>
</subSubSection>
<subSubSection id="58BE20F2E304FFEAFF6DFDF093FCFDD2" pageId="47" pageNumber="46" type="materials_examined">
<paragraph id="101B7379E304FFEAFF6DFDF091DDFDB1" blockId="47.[192,945,513,538]" box="[192,945,513,538]" pageId="47" pageNumber="46">
<emphasis id="22D0AF6BE304FFEAFF6DFDF093A9FDB2" bold="true" box="[192,453,513,537]" pageId="47" pageNumber="46">Material examined</emphasis>
.
<specimenCount id="06A2B8F0E304FFEAFE79FDF390EBFDB1" box="[468,647,514,538]" pageId="47" pageNumber="46" type="generic">301 specimens</specimenCount>
(
<collectionCode id="76B5EBBCE304FFEAFD3BFDF39088FDB1" box="[662,740,514,538]" pageId="47" pageNumber="46">JNNZ</collectionCode>
,
<collectionCode id="76B5EBBCE304FFEAFD5DFDF3912BFDB1" box="[752,839,514,538]" country="New Zealand" httpUri="http://grbio.org/cool/15kb-kde6" name="Museum of New Zealand Te Papa Tongarewa - Entomology" pageId="47" pageNumber="46" type="Museum">MONZ</collectionCode>
,
<collectionCode id="76B5EBBCE304FFEAFCFEFDF391C9FDB1" box="[851,933,514,538]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="47" pageNumber="46" type="Other institutional type">NZAC</collectionCode>
).
</paragraph>
<paragraph id="101B7379E304FFEAFF6DFDB093FCFDD2" blockId="47.[192,1440,577,634]" pageId="47" pageNumber="46">
<emphasis id="22D0AF6BE304FFEAFF6DFDB0907CFDF2" bold="true" box="[192,528,577,601]" pageId="47" pageNumber="46">Geographic distribution</emphasis>
(
<figureCitation id="889F6FFCE304FFEAFD8DFDB39015FDF1" box="[544,633,578,602]" captionStart="Figures 7782" captionStartId="69.[192,288,1875,1897]" captionTargetBox="[192,1440,192,1857]" captionTargetId="figure-45@69.[192,1440,192,1857]" captionTargetPageId="69" captionText="Figures 7782. Species distribution maps. 77) Bembidion callipeplum. 78) B. chalceipes. 79) B. chalmeri. 80) B. charile. 81) B. dehiscens. 82) B. embersoni." figureDoi="http://doi.org/10.5281/zenodo.5181788" httpUri="https://zenodo.org/record/5181788/files/figure.png" pageId="47" pageNumber="46">Fig. 78</figureCitation>
). South Island:
<collectionCode id="76B5EBBCE304FFEAFC96FDB39108FDF2" box="[827,868,578,601]" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="47" pageNumber="46">BR</collectionCode>
, CO,
<collectionCode id="76B5EBBCE304FFEAFC0AFDB391B8FDF2" box="[935,980,578,601]" pageId="47" pageNumber="46">DN</collectionCode>
,
<collectionCode id="76B5EBBCE304FFEAFC4DFDB39665FDF2" box="[992,1033,578,601]" pageId="47" pageNumber="46">FD</collectionCode>
, KA,
<collectionCode id="76B5EBBCE304FFEAFBE0FDB39610FDF2" box="[1101,1148,578,601]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="47" pageNumber="46">MB</collectionCode>
,
<collectionCode id="76B5EBBCE304FFEAFB25FDB396DBFDF1" box="[1160,1207,578,602]" country="0" name="Museo de Cipolleti" pageId="47" pageNumber="46">MC</collectionCode>
,
<collectionCode id="76B5EBBCE304FFEAFB6EFDB39698FDF2" box="[1219,1268,578,601]" country="Kenya" name="National Museum of Kenya" pageId="47" pageNumber="46">MK</collectionCode>
,
<collectionCode id="76B5EBBCE304FFEAFAADFDB39740FDF1" box="[1280,1324,578,602]" pageId="47" pageNumber="46">NC</collectionCode>
,
<collectionCode id="76B5EBBCE304FFEAFA95FDB3970AFDF2" box="[1336,1382,578,601]" pageId="47" pageNumber="46">NN</collectionCode>
, OL, SC, SD, SL,
<collectionCode id="76B5EBBCE304FFEAFEF4FD9393E0FDD2" box="[345,396,610,633]" pageId="47" pageNumber="46">WD</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="58BE20F2E304FFEAFF6DFD53939CFB31" pageId="47" pageNumber="46" type="biology_ecology">
<paragraph id="101B7379E304FFEAFF6DFD53933DFCB1" blockId="47.[192,1441,674,794]" pageId="47" pageNumber="46">
<emphasis id="22D0AF6BE304FFEAFF6DFD539342FD11" bold="true" box="[192,302,674,698]" pageId="47" pageNumber="46">Ecology</emphasis>
. Lowland, montane, subalpine. Fossorial, epigean. Banks and beds of rivers and brooks, at a certain distance (
<quantity id="D75CDE9CE304FFEAFE03FD339397FD71" box="[430,507,706,730]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="47" pageNumber="46" unit="m" value="2.5" valueMax="4.0" valueMin="1.0">14 m</quantity>
) from water. Open ground; moist or wet, sandy, bare or sparsely vegetated soil, sometimes mixed with silt. Nocturnal; hides during the day in the sand and under small stones. Gregarious.
</paragraph>
<paragraph id="101B7379E304FFEAFF6DFCB09778FCD1" blockId="47.[192,1440,833,890]" pageId="47" pageNumber="46">
<emphasis id="22D0AF6BE304FFEAFF6DFCB09345FCF2" bold="true" box="[192,297,833,857]" pageId="47" pageNumber="46">Biology</emphasis>
. Seasonality: September, NovemberApril. Tenerals: FebruaryMarch. Often infested with fungi (Laboulbeniales). Defence mechanism: when alarmed, the adult escapes by running.
</paragraph>
<paragraph id="101B7379E304FFEAFF6DFC50931BFC71" blockId="47.[192,1440,929,986]" pageId="47" pageNumber="46">
<emphasis id="22D0AF6BE304FFEAFF6DFC5093CFFC11" bold="true" box="[192,419,929,954]" pageId="47" pageNumber="46">Dispersal power</emphasis>
. Macropterous, probably capable of flight. Fast runner. Vagility likely favoured by flight capacity.
</paragraph>
<paragraph id="101B7379E304FFEAFF6DFBF0937AFB92" blockId="47.[192,1440,1025,1081]" pageId="47" pageNumber="46">
<emphasis id="22D0AF6BE304FFEAFF6DFBF09381FBB2" bold="true" box="[192,493,1025,1049]" pageId="47" pageNumber="46">Collecting techniques</emphasis>
. Pouring water over the ground; treading the soil with the feet; turning small stones.
</paragraph>
<paragraph id="101B7379E304FFEAFF6DFB90939CFB31" blockId="47.[192,1440,1121,1178]" pageId="47" pageNumber="46">
<emphasis id="22D0AF6BE304FFEAFF6DFB909320FBD2" bold="true" box="[192,332,1121,1145]" pageId="47" pageNumber="46">Reference</emphasis>
.
<bibRefCitation id="74350E88E304FFEAFEF4FB93909EFBD1" author="Larochelle, A. &amp; M. - C. Lariviere" box="[345,754,1122,1146]" pageId="47" pageNumber="46" pagination="1 - 281" refId="ref33655" refString="Larochelle, A., and M. - C. Lariviere. 2001. Carabidae (Insecta: Coleoptera): catalogue. Fauna of New Zealand 43: 1 - 281." type="journal article" year="2001">Larochelle and Larivière 2001: 81</bibRefCitation>
(catalogue; biology, dispersal power, ecology, geographic distribution, references).
</paragraph>
</subSubSection>
</treatment>
</document>