96 lines
14 KiB
XML
96 lines
14 KiB
XML
<document id="8420210329018A2E1207FEACFE2FF22E" ENCODING="UTF-8" ID-GBIF-Dataset="d42acefb-3c32-4b96-8400-e1575aa17bc6" ModsDocID="z01578p041" checkinTime="1247322647604" checkinUser="thomas" docAuthor="Paul H. Skelton" docDate="2007" docId="939D8420C5DAF0F09BA873CF2C24587F" docLanguage="en" docName="2007_Skelton_gg1.xml" docOrigin="Zootaxa 1578" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:91D85038-3B5D-4493-AE8C-6D2ECA205CAA" docTitle="Paramphilius Pellegrin" docType="treatment" docVersion="11" lastPageNumber="56" masterDocId="69594900A1F947C1E217DCD0031C8796" masterDocTitle="New species of the amphiliid catfish genera Amphilius, Doumea and Phractura and the taxonomy of Paramphilius from West Central Africa (Siluriformes, Amphiliidae)." masterLastPageNumber="68" masterPageNumber="41" pageNumber="54" updateTime="1701314704579" updateUser="plazi">
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<mods:title id="40463C929ED3C505D40B87F4BA4F331B">New species of the amphiliid catfish genera Amphilius, Doumea and Phractura and the taxonomy of Paramphilius from West Central Africa (Siluriformes, Amphiliidae).</mods:title>
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<mods:namePart id="FEDC852D9C70CC76D2A2325810E896CA">Paul H. Skelton</mods:namePart>
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<mods:title id="118C84135F4EBA4C141975F6189F98F0">Zootaxa</mods:title>
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<mods:date id="0D6F529C75E07B3D34B4FE4AAED4DBA6">2007</mods:date>
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<mods:number id="8EAA4E67C0D6408137DF6BA191EE5D65">1578</mods:number>
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<mods:classification id="C79EA1C40A020D1724A74C0B3629C518">journal article</mods:classification>
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<mods:identifier id="629018E2384C5F81344963B7521F4C98" type="Plazi-Custom">z01578p041</mods:identifier>
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<treatment id="939D8420C5DAF0F09BA873CF2C24587F" ID-DOI="http://doi.org/10.5281/zenodo.6236494" ID-GBIF-Taxon="100121966" ID-Zenodo-Dep="6236494" LSID="urn:lsid:plazi:treatment:939D8420C5DAF0F09BA873CF2C24587F" httpUri="http://treatment.plazi.org/id/939D8420C5DAF0F09BA873CF2C24587F" lastPageNumber="56" pageNumber="54">
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<subSubSection id="588E5668FB010D5A0B2B9022B5C68BA2" type="nomenclature">
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<paragraph id="3C6B8BF77EE373B0BBAC8C1286F19DF9" pageNumber="54">
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Genus
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<taxonomicName id="44974B0DE69EC2E20C0FE18D51AA89BF" ID-CoL="6FNN" ID-ENA="1233696" LSID-ZBK="urn:lsid:zoobank.org:act:EB1AFD4D-9B6D-4D03-9A67-B53999EAAC78" class="Actinopterygii" family="Amphiliidae" genus="Paramphilius" kingdom="Animalia" lsidName-ZBK="Paramphilius Pellegrin 1907:23" order="Siluriformes" pageNumber="54" phylum="Chordata" rank="genus">Paramphilius Pellegrin</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="1AED7D17EC513F68ACA7763B8FD84A31" type="diagnosis">
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<paragraph id="BF1B3D76F61AAE33A00E675197519A21" lastPageNumber="55" pageNumber="54">
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Skelton (1989) redefined the genus
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<taxonomicName id="8C33D9AB8073022A62AFBD410BDAC39D" LSID-ZBK="urn:lsid:zoobank.org:act:EB1AFD4D-9B6D-4D03-9A67-B53999EAAC78" class="Actinopterygii" family="Amphiliidae" genus="Paramphilius" kingdom="Animalia" lsidName-ZBK="Paramphilius Pellegrin 1907:23" order="Siluriformes" pageNumber="54" phylum="Chordata" rank="genus">Paramphilius</taxonomicName>
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emphasizing a highly modified head structure as the most obvious derived feature separating these fishes from other amphiliids. In
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<taxonomicName id="07775FC63116A9414E09A439141FEDB2" LSID-ZBK="urn:lsid:zoobank.org:act:EB1AFD4D-9B6D-4D03-9A67-B53999EAAC78" class="Actinopterygii" family="Amphiliidae" genus="Paramphilius" kingdom="Animalia" lsidName-ZBK="Paramphilius Pellegrin 1907:23" order="Siluriformes" pageNumber="54" phylum="Chordata" rank="genus">Paramphilius</taxonomicName>
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the tiny eyes are displaced anteriorly on the head by the extreme development of the adductor mandibulae muscles which extend behind the orbits to insert onto the dorsal aspect of the neurocranium. In larger males of
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<taxonomicName id="2D4DF8369746861E8CA3067AF698953A" LSID-ZBK="urn:lsid:zoobank.org:act:EB1AFD4D-9B6D-4D03-9A67-B53999EAAC78" class="Actinopterygii" family="Amphiliidae" genus="Paramphilius" kingdom="Animalia" lsidName-ZBK="Paramphilius Pellegrin 1907:23" order="Siluriformes" pageNumber="54" phylum="Chordata" rank="genus">Paramphilus</taxonomicName>
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the adductor mandibulae are enlarged (hypertrophied) and cause the post-orbit region to bulge noticeably.
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<taxonomicName id="317287E3DE96E35333B3A581036C1DC2" LSID-ZBK="urn:lsid:zoobank.org:act:EB1AFD4D-9B6D-4D03-9A67-B53999EAAC78" class="Actinopterygii" family="Amphiliidae" genus="Paramphilius" kingdom="Animalia" lsidName-ZBK="Paramphilius Pellegrin 1907:23" order="Siluriformes" pageNumber="54" phylum="Chordata" rank="genus">Paramphilius</taxonomicName>
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are soft bodied and cylindrical in shape, the fins are short, rounded and relatively weak compared to other amphiliids. The predorsal length is extended so that the dorsal fin is displaced relatively far back from the head and is positioned above the pelvic fins. A low number of gill rakers (<5 on leading arch) is characteristic. The genus was originally defined by Pellegrin (1907) on having a long, low adipose fin confluent with the dorsal fold of the caudal fin. This character is however unreliable at the generic level as there are both
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<taxonomicName id="E75857A6F91EED35577C0EFEF03DA788" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" order="Siluriformes" pageNumber="54" phylum="Chordata" rank="genus">Amphilius</taxonomicName>
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species with such a condition, and secondly that it is variable within a species (see Skelton, 1984). Within
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<taxonomicName id="47C2BE0D06287F372F5C0AAF0AF09782" LSID-ZBK="urn:lsid:zoobank.org:act:EB1AFD4D-9B6D-4D03-9A67-B53999EAAC78" class="Actinopterygii" family="Amphiliidae" genus="Paramphilius" kingdom="Animalia" lsidName-ZBK="Paramphilius Pellegrin 1907:23" order="Siluriformes" pageNumber="54" phylum="Chordata" rank="genus">Paramphilius</taxonomicName>
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the character appears to be consistent within the species, but it is not within the genus (Skelton, 1989). The largest species in the genus is
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<taxonomicName id="C8890875FA4F51E89E925DFAE8667E86" class="Actinopterygii" family="Amphiliidae" genus="Paramphilius" kingdom="Animalia" order="Siluriformes" pageNumber="54" phylum="Chordata" rank="species" species="baudoni">Paramphilius baudoni</taxonomicName>
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(see below) and only this species has a short notched adipose fin.
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</paragraph>
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</subSubSection>
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<subSubSection id="7B9A207CCE13D67DB6B7026E74294C7B" type="discussion">
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<paragraph id="2A30AAC0B7DC4CF108E82C0C860FFCA7" pageNumber="56">
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This review indicates that a single somewhat variable species occurs in West Central Africa and is represented in collections mostly by a species formerly known as
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<taxonomicName id="E1319BF8367CE1B6DAA8BC6962CB280B" LSID-ZBK="urn:lsid:zoobank.org:act:CFF782B6-14A0-4B44-87B2-53E7BF96D8B0" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" lsidName-ZBK="Amphilius baudoni Pellegrin 1928:453" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="baudoni">Amphilius baudoni</taxonomicName>
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but also by the species
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<taxonomicName id="A3CD815FF119E0039374FFA0CC7C2AD2" LSID-ZBK="urn:lsid:zoobank.org:act:1CE23B87-879F-44A8-89F0-A74DB11E01C2" class="Actinopterygii" family="Amphiliidae" genus="Paramphilius" kingdom="Animalia" lsidName-ZBK="Paramphilius goodi Harry 1953:203" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="goodi">Paramphilius goodi</taxonomicName>
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. A detailed comparison of the type specimens of
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<taxonomicName id="394BF973B19345A8F80BD0208825C9DC" LSID-ZBK="urn:lsid:zoobank.org:act:CFF782B6-14A0-4B44-87B2-53E7BF96D8B0" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" lsidName-ZBK="Amphilius baudoni Pellegrin 1928:453" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="baudoni">A. baudoni</taxonomicName>
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with the holotype and one paratype of
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<taxonomicName id="4B0E1A04FE5A4586086B7E457D228555" LSID-ZBK="urn:lsid:zoobank.org:act:1CE23B87-879F-44A8-89F0-A74DB11E01C2" class="Actinopterygii" family="Amphiliidae" genus="Paramphilius" kingdom="Animalia" lsidName-ZBK="Paramphilius goodi Harry 1953:203" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="goodi">P.goodi</taxonomicName>
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(Table 5) indicates the close physical similarity of the specimens on which the decision to synonymize the nominal species rests. The main difference between the types is in the Principal Caudal fin rays of
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<taxonomicName id="47C8C0A118BC858B958118CAD38C0562" LSID-ZBK="urn:lsid:zoobank.org:act:1CE23B87-879F-44A8-89F0-A74DB11E01C2" class="Actinopterygii" family="Amphiliidae" genus="Paramphilius" kingdom="Animalia" lsidName-ZBK="Paramphilius goodi Harry 1953:203" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="goodi">P. goodi</taxonomicName>
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(7+8 vs. 8+9 of
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<taxonomicName id="C0E50A4E649F2550E38DF30CB6924DE6" LSID-ZBK="urn:lsid:zoobank.org:act:CFF782B6-14A0-4B44-87B2-53E7BF96D8B0" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" lsidName-ZBK="Amphilius baudoni Pellegrin 1928:453" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="baudoni">A.baudoni</taxonomicName>
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). Although this character might indicate a taxonomic difference with only the type specimens of
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<taxonomicName id="37296F977EBADA42EE3986279FC7A653" LSID-ZBK="urn:lsid:zoobank.org:act:1CE23B87-879F-44A8-89F0-A74DB11E01C2" class="Actinopterygii" family="Amphiliidae" genus="Paramphilius" kingdom="Animalia" lsidName-ZBK="Paramphilius goodi Harry 1953:203" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="goodi">P. goodi</taxonomicName>
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available for examination, there is insufficient reason to avoid the synonymy at present.
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</paragraph>
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<paragraph id="8B8641EC8639AE60236809DE50636298" pageNumber="56">
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There are three
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<taxonomicName id="891D87605ED82E95A9B7625FDD77FA1E" LSID-ZBK="urn:lsid:zoobank.org:act:EB1AFD4D-9B6D-4D03-9A67-B53999EAAC78" class="Actinopterygii" family="Amphiliidae" genus="Paramphilius" kingdom="Animalia" lsidName-ZBK="Paramphilius Pellegrin 1907:23" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="genus">Paramphilius</taxonomicName>
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species in the Upper Guinean region of West Africa,
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<taxonomicName id="3D7B71AFF075159787AB3FD7741A19D8" LSID-ZBK="urn:lsid:zoobank.org:act:E6909ED7-3DA5-4CC1-8EEF-980082A208B6" class="Actinopterygii" family="Amphiliidae" genus="Paramphilius" kingdom="Animalia" lsidName-ZBK="Paramphilius firestonei Schultz 1942:(328) 333" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="firestonei">Paramphilius firestonei Schultz, 1941</taxonomicName>
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,
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<taxonomicName id="1409ED9BA912C06B40A28CDBBA829E6A" LSID-ZBK="urn:lsid:zoobank.org:act:EB476FA1-6CE4-4816-BB11-E209425BFEBA" class="Actinopterygii" family="Amphiliidae" genus="Paramphilius" kingdom="Animalia" lsidName-ZBK="Paramphilius teugelsi Skelton 1989:2" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="teugelsi">Paramphilius teugelsi Skelton, 1989</taxonomicName>
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, and
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<taxonomicName id="E9A907A6C64AF971BC6888A3F318A365" LSID-ZBK="urn:lsid:zoobank.org:act:135C3E32-14C6-431D-82C3-40AE0ADEF753" class="Actinopterygii" family="Amphiliidae" genus="Paramphilius" kingdom="Animalia" lsidName-ZBK="Paramphilius trichomycteroides Pellegrin 1907:24" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="trichomycteroides">Paramphilius trichomycteroides Pellegrin, 1907</taxonomicName>
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(Skelton et al., 2003). These species are all smaller sized species and differ in body proportions, especially depth, fin ray counts and the form of the adipose fin (short and notched posteriorly in
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<taxonomicName id="8D9323122B41E7078E8C2A1D467D148F" LSID-ZBK="urn:lsid:zoobank.org:act:CFF782B6-14A0-4B44-87B2-53E7BF96D8B0" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" lsidName-ZBK="Amphilius baudoni Pellegrin 1928:453" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="species" species="baudoni">A. baudoni</taxonomicName>
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vs.long, low and confluent with dorsal caudal fin fold in West African
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<taxonomicName id="645A403DC7FF2AC3F263A8D3D9816E4B" LSID-ZBK="urn:lsid:zoobank.org:act:EB1AFD4D-9B6D-4D03-9A67-B53999EAAC78" class="Actinopterygii" family="Amphiliidae" genus="Paramphilius" kingdom="Animalia" lsidName-ZBK="Paramphilius Pellegrin 1907:23" order="Siluriformes" pageNumber="56" phylum="Chordata" rank="genus">Paramphilius</taxonomicName>
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species).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |