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<document id="97A8CF0035910EC1B1946F5A0367A552" ID-CLB-Dataset="54975" ID-DOI="10.5852/ejt.2022.808.1721" ID-GBIF-Dataset="3eba97d9-c3b4-4437-8922-d01358c276bb" ID-ISSN="2118-9773" ID-Zenodo-Dep="6419368" ID-ZooBank="7898E29D-1F57-4DC2-AB70-8532234CA118" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649272993249" checkinUser="felipe" docAuthor="Hegg, Danilo, Morgan-Richards, Mary &amp; Trewick, Steven A." docDate="2022" docId="91320800FFE3FFA1FD832E91FCC3FD6D" docLanguage="en" docName="ejt-808_1-58.pdf" docOrigin="European Journal of Taxonomy 808 (1)" docSource="http://dx.doi.org/10.5852/ejt.2022.808.1721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pharmacus cochleatus subsp. nauclerus Hegg &amp; Morgan-Richards &amp; Trewick 2022, subsp. nov." docType="treatment" docUuid="D540A190-8DEE-4CDD-AA28-EA92A31550BF" docUuidSource="ZooBank" docVersion="10" lastPageNumber="31" masterDocId="6D0B7078FFFEFFBFFF882D00FF8BFFE9" masterDocTitle="High alpine sorcerers: revision of the cave wētā genus Pharmacus Pictet &amp; de Saussure (Orthoptera: Rhaphidophoridae: Macropathinae), with the description of six new species and three new subspecies" masterLastPageNumber="58" masterPageNumber="1" pageNumber="30" updateTime="1699319494823" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6263B2A52304814891345A7C332EA402">High alpine sorcerers: revision of the cave wētā genus Pharmacus Pictet &amp; de Saussure (Orthoptera: Rhaphidophoridae: Macropathinae), with the description of six new species and three new subspecies</mods:title>
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<mods:namePart id="17A179DD6E310795C4AB56CB91E17A7C">Hegg, Danilo</mods:namePart>
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<mods:affiliation id="1A8043BAE0E49D2FE547461FB5F633F6">135 Blacks Road, Opoho, Dunedin 9010, New Zealand.</mods:affiliation>
<mods:nameIdentifier id="F68608CAAF00FB8CBE7506B2E2011E2C" type="email">danilo_hegg@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="F291CC1A1C6D88BB37F29D57A2BE5A58">Morgan-Richards, Mary</mods:namePart>
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<mods:affiliation id="9EA7C453DC199E933A0CFFDE54A63123">Wildlife &amp; Ecology Group, Massey University, Private Bag 11 - 222, Palmerston North 4442, New Zealand.</mods:affiliation>
<mods:nameIdentifier id="E1EFE4D27467939437F0452A089A9800" type="email">M.Morgan-Richards@massey.ac.nz</mods:nameIdentifier>
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<mods:namePart id="D14DE42F3E19871E75FB7CB38AD074B6">Trewick, Steven A.</mods:namePart>
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<mods:affiliation id="FCB021229498F2D7DC93F01BC0B55887">Wildlife &amp; Ecology Group, Massey University, Private Bag 11 - 222, Palmerston North 4442, New Zealand.</mods:affiliation>
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<mods:title id="85DDF3E4F68355A305BB0D994A20DB55">European Journal of Taxonomy</mods:title>
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<treatment id="91320800FFE3FFA1FD832E91FCC3FD6D" ID-DOI="http://doi.org/10.5281/zenodo.6425129" ID-GBIF-Taxon="194518708" ID-Zenodo-Dep="6425129" LSID="urn:lsid:zoobank.org:act:D540A190-8DEE-4CDD-AA28-EA92A31550BF" httpUri="http://treatment.plazi.org/id/91320800FFE3FFA1FD832E91FCC3FD6D" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<subSubSection id="5181EA9DFFE3FFA2FD832E91FBA2FC45" box="[523,1065,913,940]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="1924B916FFE3FFA2FD832E91FBA2FC45" blockId="29.[353,1235,913,1010]" box="[523,1065,913,940]" pageId="29" pageNumber="30">
<heading id="426C0E7AFFE3FFA2FD832E91FBA2FC45" box="[523,1065,913,940]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="30" reason="2">
<taxonomicName id="DE9BC295FFE3FFA2FD832E91FC15FC45" authority="Hegg &amp; Morgan-Richards &amp; Trewick, 2022" authorityName="Hegg &amp; Morgan-Richards &amp; Trewick" authorityYear="2022" box="[523,926,913,940]" class="Insecta" family="Rhaphidophoridae" genus="Pharmacus" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="cochleatus" status="subsp. nov." subSpecies="nauclerus">
<emphasis id="2BEF6504FFE3FFA2FD832E91FC15FC45" bold="true" box="[523,926,913,940]" italics="true" pageId="29" pageNumber="30">Pharmacus cochleatus nauclerus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30DCD87FFFE3FFA2FC2D2E92FBA2FC45" box="[933,1065,914,940]" pageId="29" pageNumber="30" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
</heading>
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<subSubSection id="5181EA9DFFE3FFA2FEE92EB5FC17FC1B" pageId="29" pageNumber="30" type="description">
<paragraph id="1924B916FFE3FFA2FEE92EB5FB58FC39" blockId="29.[353,1235,913,1010]" box="[353,1235,949,976]" pageId="29" pageNumber="30">
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urn:lsid:zoobank.org:act:
<uuid id="6D3D83C3FFE3FFA2FDF32EB5FB58FC39" box="[635,1235,949,976]" pageId="29" pageNumber="30">D540A190-8DEE-4CDD-AA28-EA92A31550BF</uuid>
</uri>
</paragraph>
<paragraph id="1924B916FFE3FFA2FD102ED8FC17FC1B" blockId="29.[353,1235,913,1010]" box="[664,924,984,1010]" pageId="29" pageNumber="30">
<figureCitation id="81A0A593FFE3FFA2FD102ED8FC8BFC1B" box="[664,768,984,1010]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="4.[189,232,1815,1841]" captionStartId-1="6.[189,232,1958,1984]" captionTargetBox-0="[241,1345,267,1776]" captionTargetBox-1="[267,1371,262,1917]" captionTargetId-0="figure-17@4.[241,1346,265,1776]" captionTargetId-1="figure-17@6.[214,1371,262,1917]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionText-0="Fig. 1. A. Genetic relationships of cave wētā support monophyly of the genus Pharmacus Pictet &amp; de Saussure, 1893. Gene tree from mtDNA sequences (~1500 bp of COI) using Maximum Likelihood with 1000 bootstraps and Macropathus filifer Walker, 1869 to root the tree. B. Map of South Island of New Zealand, showing locations of Pharmacus specimens used in phylogenetic analyses (Figs 1A, 2). Colours correspond to different species. Two letter codes indicate the New Zealand entomological regions (Crosby et al. 1998). These codes are reported in the Material examined section for each species." captionText-1="Fig. 2. Gene tree for the eight morphologically identified species of Pharmacus Pictet &amp; de Saussure, 1893 using Maximum Likelihood analysis of ~850 bp of mtDNA (COI) from 60 specimens ofPharmacus." figureDoi-0="http://doi.org/10.5281/zenodo.6419370" figureDoi-1="http://doi.org/10.5281/zenodo.6419372" httpUri-0="https://zenodo.org/record/6419370/files/figure.png" httpUri-1="https://zenodo.org/record/6419372/files/figure.png" pageId="29" pageNumber="30">Figs 1–2</figureCitation>
,
<figureCitation id="81A0A593FFE3FFA2FC842ED8FC96FC1B" box="[780,797,984,1010]" captionStart="Fig" captionStartId="12.[189,232,1945,1971]" captionTargetBox="[189,1398,273,1856]" captionTargetId="figure-17@12.[189,1398,273,1893]" captionTargetPageId="12" captionText="Fig. 6. Known distribution of cave wētā in the genus Pharmacus Pictet &amp; de Saussure, 1893 in the south of the South Island, New Zealand." figureDoi="http://doi.org/10.5281/zenodo.6419380" httpUri="https://zenodo.org/record/6419380/files/figure.png" pageId="29" pageNumber="30">6</figureCitation>
,
<figureCitation id="81A0A593FFE3FFA2FCA22ED8FCD4FC1B" box="[810,863,984,1010]" captionStart="Fig" captionStartId="40.[189,232,1671,1697]" captionTargetBox="[189,1398,824,1633]" captionTargetId="figure-150@40.[189,1398,824,1633]" captionTargetPageId="40" captionText="Fig. 17. Habitat of Pharmacus Pictet &amp; de Saussure, 1893. A. Mount Dick, Eyre Mountains. a. Rock bluffs at 1550 m a.s.l., collection locality of Pharmacus concinnus sp. nov. (MPN CW4472–4475, CW4501–4504, CW4592, CW4593). B. Dunstan, Dunstan Mountains. b. Rock tors at 1670 m a.s.l. collection locality of P. senex sp. nov. (MPN CW4381–4385, CW4402–4405, CW4506–4509). C. Spence Peak, Takitimu Mountains. c. North ridge, 1450 m a.s.l., collection locality of P. perfidus sp. nov. (NMNZ AI.052300, AI.052301; MPN CW4487, CW4489, CW4490, CW4494–4496, CW4498, CW4499, CW4553–4555). D. Skippers Range High Point, South Westland. d–e. Scree and rock bluffs around lake, 1600 m a.s.l., collection locality of P. cochleatus nauclerus subsp. nov. (NMNZ AI.052289, MPN CW4363) and of P. cristatus sp. nov. (NMNZ AI.052292, AI.052293; MPN CW4365, CW4376, CW4484, CW4485, CW4562)." figureDoi="http://doi.org/10.5281/zenodo.6419402" httpUri="https://zenodo.org/record/6419402/files/figure.png" pageId="29" pageNumber="30">17D</figureCitation>
,
<figureCitation id="81A0A593FFE3FFA2FCE32ED8FC17FC1B" box="[875,924,984,1010]" captionStart="Fig" captionStartId="46.[189,232,1810,1836]" captionTargetBox="[189,1398,802,1769]" captionTargetId="figure-184@46.[189,1398,797,1769]" captionTargetPageId="46" captionText="Fig. 18. Map of South Island, New Zealand, showing the known distribution of cave wētā in the genus Pharmacus Pictet &amp; de Saussure, 1893. A–B. Solid shapes indicate material collected as part of this study; empty circles indicate additional locations of material examined by Richards (1972). B. Shapes represent different subspecies: dark blue circle = Pharmacus cochleatus cochleatus; light blue circle = Pharmacus cochleatus nauclerus subsp. nov.; diamond = Pharmacus cochleatus fiordensis subsp. nov.; triangle = Pharmacus cochleatus rawhiti subsp. nov." figureDoi="http://doi.org/10.5281/zenodo.6419404" httpUri="https://zenodo.org/record/6419404/files/figure.png" pageId="29" pageNumber="30">18B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="5181EA9DFFE3FFA2FF352919FEACFA9E" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="1924B916FFE3FFA2FF352919FEBDFBDA" blockId="29.[189,310,1049,1075]" box="[189,310,1049,1075]" pageId="29" pageNumber="30">
<heading id="426C0E7AFFE3FFA2FF352919FEBDFBDA" bold="true" box="[189,310,1049,1075]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="2BEF6504FFE3FFA2FF352919FEBDFBDA" bold="true" box="[189,310,1049,1075]" pageId="29" pageNumber="30">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="1924B916FFE3FFA2FF352948FE28FB40" blockId="29.[189,1398,1096,1193]" pageId="29" pageNumber="30">
A mid-sized cave wētā endemic to the Skippers Range in South Westland. Morphologically indistinguishable from
<taxonomicName id="DE9BC295FFE3FFA2FE45296BFCD7FB6C" authority="cochleatus" authorityName="cochleatus" box="[461,860,1131,1157]" class="Insecta" family="Rhaphidophoridae" genus="Pharmacus" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="cochleatus" status="comb. nov." subSpecies="cochleatus">
<emphasis id="2BEF6504FFE3FFA2FE45296BFCD7FB6C" box="[461,860,1131,1157]" italics="true" pageId="29" pageNumber="30">Pharmacus cochleatus cochleatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30DCD87FFFE3FFA2FCEC296BFC63FB6F" box="[868,1000,1131,1158]" pageId="29" pageNumber="30" rank="species">comb. nov.</taxonomicNameLabel>
, but geographically separated and genetically distinct.
</paragraph>
<paragraph id="1924B916FFE3FFA2FF3529D0FEACFA9E" blockId="29.[189,1399,1232,1399]" pageId="29" pageNumber="30">
<taxonomicName id="DE9BC295FFE3FFA2FF3529D0FDC9FB03" authorityName="Hegg &amp; Morgan-Richards &amp; Trewick" authorityYear="2022" box="[189,578,1232,1258]" class="Insecta" family="Rhaphidophoridae" genus="Pharmacus" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="subSpecies" species="cochleatus" subSpecies="nauclerus">
<emphasis id="2BEF6504FFE3FFA2FF3529D0FDC9FB03" box="[189,578,1232,1258]" italics="true" pageId="29" pageNumber="30">Pharmacus cochleatus nauclerus</emphasis>
</taxonomicName>
is sympatric with
<taxonomicName id="DE9BC295FFE3FFA2FC9F29D0FB82FB03" authority="Hegg &amp; Morgan-Richards &amp; Trewick, 2022" authorityName="Hegg &amp; Morgan-Richards &amp; Trewick" authorityYear="2022" box="[791,1033,1232,1258]" class="Insecta" family="Rhaphidophoridae" genus="Pharmacus" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cristatus" status="sp. nov.">
<emphasis id="2BEF6504FFE3FFA2FC9F29D0FB82FB03" box="[791,1033,1232,1258]" italics="true" pageId="29" pageNumber="30">Pharmacus cristatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30DCD87FFFE3FFA2FB9829D1FBE1FB02" box="[1040,1130,1233,1259]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
and with
<taxonomicName id="DE9BC295FFE3FFA2FB5429D0FEDDFAE4" authorityName="Hegg &amp; Morgan-Richards &amp; Trewick" authorityYear="2022" baseAuthorityName="Richards" baseAuthorityYear="1972" class="Insecta" family="Rhaphidophoridae" genus="Notoplectron" kingdom="Animalia" order="Orthoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="brewsterense" status="comb. nov.">
<emphasis id="2BEF6504FFE3FFA2FB5429D0FEDDFAE4" italics="true" pageId="29" pageNumber="30">Notoplectron brewsterense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30DCD87FFFE3FFA2FED629F3FE6AFAE7" box="[350,481,1267,1294]" pageId="29" pageNumber="30" rank="species">comb. nov.</taxonomicNameLabel>
It is easily differentiated from the former by its darker coloration and lack of prolateral apical spines on the fore and mid femora and from the latter by its darker colour, longer legs with more numerous spines on the hind tibiae and the presence of dorsal spines on the first hind tarsus segment.
</paragraph>
</subSubSection>
<subSubSection id="5181EA9DFFE3FFA2FF35289EFBD1FA0E" box="[189,1114,1438,1511]" pageId="29" pageNumber="30" type="etymology">
<paragraph id="1924B916FFE3FFA2FF35289EFECFFA51" blockId="29.[189,324,1438,1464]" box="[189,324,1438,1464]" pageId="29" pageNumber="30">
<heading id="426C0E7AFFE3FFA2FF35289EFECFFA51" bold="true" box="[189,324,1438,1464]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="2BEF6504FFE3FFA2FF35289EFECFFA51" bold="true" box="[189,324,1438,1464]" pageId="29" pageNumber="30">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="1924B916FFE3FFA2FF3528CDFBD1FA0E" blockId="29.[189,1114,1485,1511]" box="[189,1114,1485,1511]" pageId="29" pageNumber="30">
‘
<emphasis id="2BEF6504FFE3FFA2FF4F28CDFECBFA0E" box="[199,320,1485,1511]" italics="true" pageId="29" pageNumber="30">Nauclērus</emphasis>
’ is Latin for ‘skipper’, after the Skippers Range in South Westland.
</paragraph>
</subSubSection>
<subSubSection id="5181EA9DFFE3FFA1FF352B0EFBF2FE59" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="1924B916FFE3FFA2FF352B0EFCAEF9C1" blockId="29.[189,805,1550,1577]" box="[189,805,1550,1577]" pageId="29" pageNumber="30">
<emphasis id="2BEF6504FFE3FFA2FF352B0EFE21F9C1" bold="true" box="[189,426,1550,1576]" pageId="29" pageNumber="30">Material examined</emphasis>
(see also Supp. file 1: Table S3)
</paragraph>
<paragraph id="1924B916FFE3FFA2FF622B3DFAB6F974" blockId="29.[189,1398,1597,1694]" pageId="29" pageNumber="30">
<materialsCitation id="A9F3B34BFFE3FFA2FF622B3DFAB6F974" ID-GBIF-Occurrence="3748656324" collectedFrom="on rock bluffs" collectingDate="2019-02-07" collectingMethod="night search" collectionCode="NMNZ" collectorName="D. Hegg" country="New Zealand" elevation="1400" latitude="-44.50459" location="Westland" longLatPrecision="1" longitude="168.12086" municipality="Skippers Range" pageId="29" pageNumber="30" specimenCode="AI.052288" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="2BEF6504FFE3FFA2FF622B3DFED7F9BE" bold="true" box="[234,348,1597,1623]" pageId="29" pageNumber="30">
<typeStatus id="C62007B4FFE3FFA2FF622B3DFED7F9BE" box="[234,348,1597,1623]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="618CF986FFE3FFA2FF352B61FE05F993" box="[189,398,1632,1659]" name="New Zealand" pageId="29" pageNumber="30">NEW ZEALAND</collectingCountry>
•
<specimenCount id="0F9D729FFFE3FFA2FE2B2B61FE30F992" box="[419,443,1633,1659]" pageId="29" pageNumber="30" type="male">♂</specimenCount>
, adult;
<location id="1C44EFCDFFE3FFA2FD852B60FDF2F993" LSID="urn:lsid:plazi:treatment:91320800FFE3FFA1FD832E91FCC3FD6D:1C44EFCDFFE3FFA2FD852B60FDF2F993" box="[525,633,1632,1658]" country="New Zealand" latitude="-44.50459" longLatPrecision="1" longitude="168.12086" municipality="Skippers Range" name="Westland" pageId="29" pageNumber="30">Westland</location>
(WD),
<collectingMunicipality id="F940236CFFE3FFA2FD582B60FC01F992" box="[720,906,1632,1659]" pageId="29" pageNumber="30">Skippers Range</collectingMunicipality>
;
<geoCoordinate id="7CAFDFD1FFE3FFA2FC1C2B60FBAFF993" box="[916,1060,1632,1659]" degrees="44.50459" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="1" value="-44.50459">44.50459° S</geoCoordinate>
,
<geoCoordinate id="7CAFDFD1FFE3FFA2FBA72B60FB59F992" box="[1071,1234,1632,1659]" degrees="168.12086" direction="east" orientation="longitude" pageId="29" pageNumber="30" precision="1" value="168.12086">168.12086° E</geoCoordinate>
;
<elevation id="92B65E25FFE3FFA2FB542B60FAFBF993" box="[1244,1392,1632,1659]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="29" pageNumber="30" unit="m" value="1400.0">
<quantity id="DE6314F3FFE3FFA2FB542B60FAB8F992" box="[1244,1331,1632,1659]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="29" pageNumber="30" unit="m" value="1400.0">1400 m</quantity>
a.s.l.
</elevation>
;
<date id="6D259FD6FFE3FFA2FF352B84FEC7F974" box="[189,332,1667,1694]" pageId="29" pageNumber="30" value="2019-02-07">
<collectingDate id="7D61663EFFE3FFA2FF352B84FEC7F974" box="[189,332,1667,1694]" pageId="29" pageNumber="30" value="2019-02-07">7 Feb. 2019</collectingDate>
</date>
;
<collectorName id="B46EDCC0FFE3FFA2FED02B84FE37F977" box="[344,444,1668,1694]" pageId="29" pageNumber="30">D. Hegg</collectorName>
leg.;
<collectedFrom id="AA87FAEBFFE3FFA2FE752B84FD2EF974" box="[509,677,1667,1694]" pageId="29" pageNumber="30">on rock bluffs</collectedFrom>
;
<collectingMethod id="C0DAC101FFE3FFA2FD392B83FCC8F974" box="[689,835,1667,1693]" pageId="29" pageNumber="30">night search</collectingMethod>
;
<collectionCode id="7F8A21D3FFE3FFA2FCC72B84FC2EF977" box="[847,933,1668,1694]" country="New Zealand" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="29" pageNumber="30" type="Museum">NMNZ</collectionCode>
<specimenCode id="493D116DFFE3FFA2FC232B83FBA2F977" box="[939,1065,1667,1694]" pageId="29" pageNumber="30">AI.052288</specimenCode>
(prev. MPN CW4412).
</materialsCitation>
</paragraph>
<paragraph id="1924B916FFE3FFA2FF622BC5FEF5F8A0" blockId="29.[188,1399,1733,1865]" pageId="29" pageNumber="30">
<materialsCitation id="A9F3B34BFFE3FFA2FF622BC5FEF1F8A0" ID-GBIF-Occurrence="3748656398" collectedFrom="on scree and rock bluffs" collectingDate="2019-02-03" collectingMethod="night search" collectionCode="NMNZ" collectorName="D. Hegg" country="New Zealand" elevation="1600" latitude="-44.43914" location="Westland" longLatPrecision="1" longitude="168.16515" municipality="Skippers Range High Point" pageId="29" pageNumber="30" specimenCode="AI.052289" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="2BEF6504FFE3FFA2FF622BC5FED7F936" bold="true" box="[234,348,1733,1759]" pageId="29" pageNumber="30">
<typeStatus id="C62007B4FFE3FFA2FF622BC5FED7F936" box="[234,348,1733,1759]" pageId="29" pageNumber="30" type="paratype">Paratype</typeStatus>
</emphasis>
<collectingCountry id="618CF986FFE3FFA2FF352BE8FE04F8EB" box="[189,399,1768,1794]" name="New Zealand" pageId="29" pageNumber="30">NEW ZEALAND</collectingCountry>
•
<specimenCount id="0F9D729FFFE3FFA2FE2D2BE8FE5BF8EB" box="[421,464,1768,1794]" pageId="29" pageNumber="30" type="female">1 ♀</specimenCount>
, adult;
<location id="1C44EFCDFFE3FFA2FDAD2BE8FD1AF8EB" LSID="urn:lsid:plazi:treatment:91320800FFE3FFA1FD832E91FCC3FD6D:1C44EFCDFFE3FFA2FDAD2BE8FD1AF8EB" box="[549,657,1768,1794]" country="New Zealand" latitude="-44.43914" longLatPrecision="1" longitude="168.16515" municipality="Skippers Range High Point" name="Westland" pageId="29" pageNumber="30">Westland</location>
(WD),
<collectingMunicipality id="F940236CFFE3FFA2FD602BE8FBA3F8EB" box="[744,1064,1768,1794]" pageId="29" pageNumber="30">Skippers Range High Point</collectingMunicipality>
;
<geoCoordinate id="7CAFDFD1FFE3FFA2FBB92BE8FB48F8EB" box="[1073,1219,1768,1794]" degrees="44.43914" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="1" value="-44.43914">44.43914° S</geoCoordinate>
,
<geoCoordinate id="7CAFDFD1FFE3FFA2FB462BE8FAFAF8EB" box="[1230,1393,1768,1795]" degrees="168.16515" direction="east" orientation="longitude" pageId="29" pageNumber="30" precision="1" value="168.16515">168.16515° E</geoCoordinate>
;
<elevation id="92B65E25FFE3FFA2FF352A0BFED8F8CC" box="[189,339,1803,1830]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="29" pageNumber="30" unit="m" value="1600.0">
<quantity id="DE6314F3FFE3FFA2FF352A0BFE9EF8CF" box="[189,277,1803,1830]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="29" pageNumber="30" unit="m" value="1600.0">1600 m</quantity>
a.s.l.
</elevation>
;
<date id="6D259FD6FFE3FFA2FED72A0BFE60F8CC" box="[351,491,1803,1829]" pageId="29" pageNumber="30" value="2019-02-03">
<collectingDate id="7D61663EFFE3FFA2FED72A0BFE60F8CC" box="[351,491,1803,1829]" pageId="29" pageNumber="30" value="2019-02-03">3 Feb. 2019</collectingDate>
</date>
;
<collectorName id="B46EDCC0FFE3FFA2FE7E2A0CFDD3F8CF" box="[502,600,1804,1830]" pageId="29" pageNumber="30">D. Hegg</collectorName>
leg.;
<collectedFrom id="AA87FAEBFFE3FFA2FD1F2A0CFC24F8CC" box="[663,943,1803,1830]" pageId="29" pageNumber="30">on scree and rock bluffs</collectedFrom>
;
<collectingMethod id="C0DAC101FFE3FFA2FC312A0BFBC1F8CC" box="[953,1098,1803,1829]" pageId="29" pageNumber="30">night search</collectingMethod>
;
<collectionCode id="7F8A21D3FFE3FFA2FBDC2A0CFB21F8CF" box="[1108,1194,1804,1830]" country="New Zealand" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="29" pageNumber="30" type="Museum">NMNZ</collectionCode>
<specimenCode id="493D116DFFE3FFA2FB272A0BFAA6F8CF" box="[1199,1325,1803,1830]" pageId="29" pageNumber="30">AI.052289</specimenCode>
(prev. MPN CW4376)
</materialsCitation>
.
</paragraph>
<paragraph id="1924B916FFE3FFA2FF622A6FFE2DF860" blockId="29.[189,1398,1903,2001]" box="[234,422,1903,1929]" pageId="29" pageNumber="30">
<emphasis id="2BEF6504FFE3FFA2FF622A6FFE2DF860" bold="true" box="[234,422,1903,1929]" pageId="29" pageNumber="30">Other material</emphasis>
</paragraph>
<paragraph id="1924B916FFE3FFA2FF352A93FAFDF839" blockId="29.[189,1398,1903,2001]" pageId="29" pageNumber="30">
<materialsCitation id="A9F3B34BFFE3FFA2FF352A93FAFDF839" ID-GBIF-Occurrence="3748656349" collectedFrom="on rock bluffs" collectingDate="2019-02-02" collectingMethod="night search" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" elevation="1360" latitude="-44.42005" location="Skippers Range" longLatPrecision="1" longitude="168.15399" pageId="29" pageNumber="30" specimenCode="CW4366, CW4378, CW4413" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Westland">
<collectingCountry id="618CF986FFE3FFA2FF352A93FE04F844" box="[189,399,1939,1965]" name="New Zealand" pageId="29" pageNumber="30">NEW ZEALAND</collectingCountry>
–
<emphasis id="2BEF6504FFE3FFA2FE222A93FDC4F844" bold="true" box="[426,591,1939,1965]" pageId="29" pageNumber="30">
<collectingRegion id="DB5F77F4FFE3FFA2FE222A93FD9AF844" box="[426,529,1939,1965]" pageId="29" pageNumber="30">Westland</collectingRegion>
(WD)
</emphasis>
•
<specimenCount id="0F9D729FFFE3FFA2FD0C2A93FD3AF844" box="[644,689,1939,1965]" pageId="29" pageNumber="30" type="male">1 ♂</specimenCount>
,
<specimenCount id="0F9D729FFFE3FFA2FD342A93FD6CF844" box="[700,743,1939,1965]" pageId="29" pageNumber="30" type="female">1 ♀</specimenCount>
, 1 nymph;
<location id="1C44EFCDFFE3FFA2FCE22A93FBAEF844" LSID="urn:lsid:plazi:treatment:91320800FFE3FFA1FD832E91FCC3FD6D:1C44EFCDFFE3FFA2FCE22A93FBAEF844" box="[874,1061,1939,1965]" country="New Zealand" latitude="-44.42005" longLatPrecision="1" longitude="168.15399" name="Skippers Range" pageId="29" pageNumber="30" stateProvince="Westland">Skippers Range</location>
;
<geoCoordinate id="7CAFDFD1FFE3FFA2FBB82A93FB49F844" box="[1072,1218,1939,1966]" degrees="44.42005" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="1" value="-44.42005">44.42005° S</geoCoordinate>
,
<geoCoordinate id="7CAFDFD1FFE3FFA2FB462A93FAFAF844" box="[1230,1393,1939,1966]" degrees="168.15399" direction="east" orientation="longitude" pageId="29" pageNumber="30" precision="1" value="168.15399">168.15399° E</geoCoordinate>
;
<quantity id="DE6314F3FFE3FFA2FF352AB6FE9FF838" box="[189,276,1974,2001]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="29" pageNumber="30" unit="m" value="1360.0">
<elevation id="92B65E25FFE3FFA2FF352AB6FE9FF838" box="[189,276,1974,2001]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="29" pageNumber="30" unit="m" value="1360.0">1360 m</elevation>
</quantity>
a.s.l;
<date id="6D259FD6FFE3FFA2FEDF2AB6FE68F839" box="[343,483,1974,2000]" pageId="29" pageNumber="30" value="2019-02-02">
<collectingDate id="7D61663EFFE3FFA2FEDF2AB6FE68F839" box="[343,483,1974,2000]" pageId="29" pageNumber="30" value="2019-02-02">2 Feb. 2019</collectingDate>
</date>
;
<collectorName id="B46EDCC0FFE3FFA2FE662AB7FDDBF838" box="[494,592,1975,2001]" pageId="29" pageNumber="30">D. Hegg</collectorName>
leg.;
<collectedFrom id="AA87FAEBFFE3FFA2FD072AB7FCBFF839" box="[655,820,1974,2001]" pageId="29" pageNumber="30">on rock bluffs</collectedFrom>
;
<collectingMethod id="C0DAC101FFE3FFA2FCB62AB6FC44F839" box="[830,975,1974,2000]" pageId="29" pageNumber="30">night search</collectingMethod>
;
<collectionCode id="7F8A21D3FFE3FFA2FC512AB7FB92F838" box="[985,1049,1975,2001]" pageId="29" pageNumber="30">MPN</collectionCode>
<specimenCode id="493D116DFFE3FFA2FB962AB6FB00F839" box="[1054,1163,1974,2000]" collectionCode="MPN" pageId="29" pageNumber="30">CW4366</specimenCode>
,
<specimenCode id="493D116DFFE3FFA2FB1D2AB6FA89F839" box="[1173,1282,1974,2000]" collectionCode="MPN" pageId="29" pageNumber="30">CW4378</specimenCode>
,
<specimenCode id="493D116DFFE3FFA2FA852AB6FAFDF839" box="[1293,1398,1974,2000]" collectionCode="MPN" pageId="29" pageNumber="30">CW4413</specimenCode>
</materialsCitation>
</paragraph>
<paragraph id="1924B916FFE0FFA1FF352C09FBF2FE59" blockId="30.[189,1399,265,432]" pageId="30" pageNumber="31">
•
<materialsCitation id="A9F3B34BFFE0FFA1FF462C09FB37FECA" ID-GBIF-Occurrence="3748656357" accessionNumber="OM293711" box="[206,1212,265,291]" collectedFrom="on scree and rock bluffs" collectingDate="2019-02-03" collectingMethod="night search" collectionCode="MPN" collectorName="GenBank" country="New Zealand" elevation="1600" latitude="-44.43914" location="Westland" longLatPrecision="1" longitude="168.16515" municipality="Skippers Range High Point" pageId="30" pageNumber="31" specimenCode="CW4363" specimenCount="1">
<specimenCount id="0F9D729FFFE0FFA1FF462C09FEB2FECA" box="[206,313,265,291]" pageId="30" pageNumber="31">1 nymph</specimenCount>
; same collection data as for paratype; GenBank:
<accessionNumber id="06C824F5FFE0FFA1FCFE2C09FB8BFECA" box="[886,1024,265,291]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OM293711" pageId="30" pageNumber="31">OM293711</accessionNumber>
;
<collectionCode id="7F8A21D3FFE0FFA1FB842C09FBC7FECA" box="[1036,1100,265,291]" pageId="30" pageNumber="31">MPN</collectionCode>
<specimenCode id="493D116DFFE0FFA1FBDA2C09FB37FECA" box="[1106,1212,265,291]" collectionCode="MPN" pageId="30" pageNumber="31">CW4363</specimenCode>
</materialsCitation>
•
<materialsCitation id="A9F3B34BFFE0FFA1FB5D2C09FD51FE80" ID-GBIF-Occurrence="3748656426" accessionNumber="OM293713" collectedFrom="on rock bluffs" collectingDate="2019-02-06" collectingMethod="night search" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" elevation="1200" latitude="-44.46821" location="Skippers Range" longLatPrecision="1" longitude="168.10095" pageId="30" pageNumber="31" specimenCode="CW4374" specimenCount="1" specimenCount-female="1">
<specimenCount id="0F9D729FFFE0FFA1FB5D2C09FA8AFECA" box="[1237,1281,265,291]" pageId="30" pageNumber="31" type="female">1 ♀</specimenCount>
;
<location id="1C44EFCDFFE0FFA1FA982C09FE87FEAE" LSID="urn:lsid:plazi:treatment:91320800FFE3FFA1FD832E91FCC3FD6D:1C44EFCDFFE0FFA1FA982C09FE87FEAE" country="New Zealand" latitude="-44.46821" longLatPrecision="1" longitude="168.10095" name="Skippers Range" pageId="30" pageNumber="31">Skippers Range</location>
;
<geoCoordinate id="7CAFDFD1FFE0FFA1FE932C2CFE3AFEAF" box="[283,433,300,326]" degrees="44.46821" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="1" value="-44.46821">44.46821° S</geoCoordinate>
,
<geoCoordinate id="7CAFDFD1FFE0FFA1FE492C2CFDE2FEAE" box="[449,617,300,327]" degrees="168.10095" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="1" value="168.10095">168.10095° E</geoCoordinate>
;
<elevation id="92B65E25FFE0FFA1FDF12C2CFC93FEAF" box="[633,792,300,327]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="30" pageNumber="31" unit="m" value="1200.0">
<quantity id="DE6314F3FFE0FFA1FDF12C2CFD5EFEAE" box="[633,725,300,327]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="30" pageNumber="31" unit="m" value="1200.0">1200 m</quantity>
a.s.l.
</elevation>
;
<date id="6D259FD6FFE0FFA1FCA12C2CFC35FEAF" box="[809,958,300,326]" pageId="30" pageNumber="31" value="2019-02-06">
<collectingDate id="7D61663EFFE0FFA1FCA12C2CFC35FEAF" box="[809,958,300,326]" pageId="30" pageNumber="31" value="2019-02-06">6 Feb. 2019</collectingDate>
</date>
;
<collectorName id="B46EDCC0FFE0FFA1FC462C2DFBBEFEAE" box="[974,1077,301,327]" pageId="30" pageNumber="31">D. Hegg</collectorName>
leg.;
<collectedFrom id="AA87FAEBFFE0FFA1FBF52C2DFAA0FEAF" box="[1149,1323,300,327]" pageId="30" pageNumber="31">on rock bluffs</collectedFrom>
;
<collectingMethod id="C0DAC101FFE0FFA1FAB22C2CFE87FE80" pageId="30" pageNumber="31">night search</collectingMethod>
; GenBank:
<accessionNumber id="06C824F5FFE0FFA1FE1C2C4FFD94FE80" box="[404,543,335,361]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OM293713" pageId="30" pageNumber="31">OM293713</accessionNumber>
;
<collectionCode id="7F8A21D3FFE0FFA1FDA22C50FDE1FE83" box="[554,618,336,362]" pageId="30" pageNumber="31">MPN</collectionCode>
<specimenCode id="493D116DFFE0FFA1FDF82C4FFD51FE80" box="[624,730,335,361]" collectionCode="MPN" pageId="30" pageNumber="31">CW4374</specimenCode>
</materialsCitation>
•
<materialsCitation id="A9F3B34BFFE0FFA1FD792C4FFD92FE64" ID-GBIF-Occurrence="3748656313" accessionNumber="OM293713" collectedFrom="on rock bluffs" collectingDate="2019-02-06" collectingMethod="night search" collectionCode="MPN" collectorName="D. Hegg &amp; GenBank" country="New Zealand" elevation="1200" latitude="-44.46821" location="Skippers Range" longLatPrecision="1" longitude="168.10095" pageId="30" pageNumber="31" specimenCode="CW4377, CW4410, CW4411" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="0F9D729FFFE0FFA1FD792C4FFCBFFE83" box="[753,820,335,362]" pageId="30" pageNumber="31" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="0F9D729FFFE0FFA1FCB72C4FFCE7FE83" box="[831,876,335,362]" pageId="30" pageNumber="31" type="female">1 ♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="7F8A21D3FFE0FFA1FABF2C50FAFCFE83" box="[1335,1399,336,362]" pageId="30" pageNumber="31">MPN</collectionCode>
<specimenCode id="493D116DFFE0FFA1FF352C73FEA1FE64" box="[189,298,371,397]" collectionCode="MPN" pageId="30" pageNumber="31">CW4377</specimenCode>
,
<specimenCode id="493D116DFFE0FFA1FEBE2C73FE2FFE64" box="[310,420,371,397]" collectionCode="MPN" pageId="30" pageNumber="31">CW4410</specimenCode>
,
<specimenCode id="493D116DFFE0FFA1FE382C73FD92FE64" box="[432,537,371,397]" collectionCode="MPN" pageId="30" pageNumber="31">CW4411</specimenCode>
</materialsCitation>
•
<materialsCitation id="A9F3B34BFFE0FFA1FDBB2C73FEE5FE59" ID-GBIF-Occurrence="3748656384" accessionNumber="OM293710" collectedFrom="on rock bluffs" collectingDate="2019-02-07" collectingMethod="night search" collectionCode="MPN" collectorName="GenBank" country="New Zealand" elevation="1400" latitude="-44.50459" location="Westland" longLatPrecision="1" longitude="168.12086" municipality="Skippers Range" pageId="30" pageNumber="31" specimenCode="CW4361" specimenCount="1">
<specimenCount id="0F9D729FFFE0FFA1FDBB2C73FD14FE65" box="[563,671,370,397]" pageId="30" pageNumber="31">1 nymph</specimenCount>
; same collection data as for holotype; GenBank:
<accessionNumber id="06C824F5FFE0FFA1FB6F2C73FAF9FE64" box="[1255,1394,371,397]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OM293710" pageId="30" pageNumber="31">OM293710</accessionNumber>
;
<collectionCode id="7F8A21D3FFE0FFA1FF352C96FF76FE59" box="[189,253,406,432]" pageId="30" pageNumber="31">MPN</collectionCode>
<specimenCode id="493D116DFFE0FFA1FE8C2C96FEE5FE59" box="[260,366,406,432]" collectionCode="MPN" pageId="30" pageNumber="31">CW4361</specimenCode>
</materialsCitation>
•
<materialsCitation id="A9F3B34BFFE0FFA1FE0F2C96FBFEFE59" ID-GBIF-Occurrence="3748656434" box="[391,1141,406,432]" collectedFrom="on rock bluffs" collectingDate="2019-02-07" collectingMethod="night search" collectionCode="NMNZ, MPN" collectorName="D. Hegg" country="New Zealand" elevation="1400" latitude="-44.50459" location="Westland" longLatPrecision="1" longitude="168.12086" municipality="Skippers Range" pageId="30" pageNumber="31" specimenCode="CW4379, CW4380" specimenCount="2" specimenCount-female="2">
<specimenCount id="0F9D729FFFE0FFA1FE0F2C96FE41FE59" box="[391,458,406,432]" pageId="30" pageNumber="31" type="female">2 ♀♀</specimenCount>
; same collection data as for holotype;
<specimenCode id="493D116DFFE0FFA1FC072C96FC70FE59" box="[911,1019,406,432]" pageId="30" pageNumber="31">CW4379</specimenCode>
,
<specimenCode id="493D116DFFE0FFA1FB8F2C96FBFEFE59" box="[1031,1141,406,432]" pageId="30" pageNumber="31">CW4380</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5181EA9DFFE0FFA1FF352CD7FCC3FD6D" pageId="30" pageNumber="31" type="description">
<paragraph id="1924B916FFE0FFA1FF352CD7FEC5FE18" blockId="30.[189,334,471,497]" box="[189,334,471,497]" pageId="30" pageNumber="31">
<heading id="426C0E7AFFE0FFA1FF352CD7FEC5FE18" bold="true" box="[189,334,471,497]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="2BEF6504FFE0FFA1FF352CD7FEC5FE18" bold="true" box="[189,334,471,497]" pageId="30" pageNumber="31">Description</emphasis>
</heading>
</paragraph>
<paragraph id="1924B916FFE0FFA1FF352F06FD87FDC9" blockId="30.[189,524,518,544]" box="[189,524,518,544]" pageId="30" pageNumber="31">
MEASUREMENTS. See
<tableCitation id="54198CADFFE0FFA1FE262F06FD8DFDC9" box="[430,518,518,544]" captionStart="Table 1" captionStartId="14.[189,259,265,291]" captionTargetPageId="14" captionText="Table 1 (continued on next three pages). Dimensions and spine count of species of Pharmacus Pictet &amp; de Saussure, 1893." httpUri="http://table.plazi.org/id/4DE4E99EFFF0FFB1FF352C09FE13FEAF" pageId="30" pageNumber="31" tableUuid="4DE4E99EFFF0FFB1FF352C09FE13FEAF">Table 1</tableCitation>
.
</paragraph>
<paragraph id="1924B916FFE0FFA1FF352F47FCC3FD6D" blockId="30.[189,1398,583,645]" pageId="30" pageNumber="31">
<taxonomicName id="DE9BC295FFE0FFA1FF352F47FDC0FD88" authority="Hegg &amp; Morgan-Richards &amp; Trewick, 2022" authorityName="Hegg &amp; Morgan-Richards &amp; Trewick" authorityYear="2022" box="[189,587,583,609]" class="Insecta" family="Rhaphidophoridae" genus="Pharmacus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="cochleatus" status="subsp. nov." subSpecies="nauclerus">
<emphasis id="2BEF6504FFE0FFA1FF352F47FDC0FD88" box="[189,587,583,609]" italics="true" pageId="30" pageNumber="31">Pharmacus cochleatus nauclerus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30DCD87FFFE0FFA1FDDE2F47FD55FD8B" box="[598,734,583,610]" pageId="30" pageNumber="31" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
is morphologically identical to
<taxonomicName id="DE9BC295FFE0FFA1FBE02F47FEB1FD6D" authority="cochleatus" authorityName="cochleatus" class="Insecta" family="Rhaphidophoridae" genus="Pharmacus" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="cochleatus" status="comb. nov." subSpecies="cochleatus">
<emphasis id="2BEF6504FFE0FFA1FBE02F47FEB1FD6D" italics="true" pageId="30" pageNumber="31">Pharmacus cochleatus cochleatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30DCD87FFFE0FFA1FECA2F6AFE48FD6C" box="[322,451,618,645]" pageId="30" pageNumber="31" rank="species">comb. nov.</taxonomicNameLabel>
See the description for the latter.
</paragraph>
</subSubSection>
</treatment>
</document>
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