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<mods:title id="93E83F95DE744F96A208A760E9939D11">High alpine sorcerers: revision of the cave wētā genus Pharmacus Pictet &amp; de Saussure (Orthoptera: Rhaphidophoridae: Macropathinae), with the description of six new species and three new subspecies</mods:title>
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<mods:namePart id="0E7E02B2249D3AA5A7896032A198C6F4">Hegg, Danilo</mods:namePart>
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<mods:affiliation id="2A09C2696CEDFE92D37DCAC48B2A6CDB">Wildlife &amp; Ecology Group, Massey University, Private Bag 11 - 222, Palmerston North 4442, New Zealand.</mods:affiliation>
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<emphasis id="2BEF6504FFD8FF99FDE42C08FCEDFECA" bold="true" box="[620,870,264,291]" italics="true" pageId="38" pageNumber="39">Pharmacus notabilis</emphasis>
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<figureCitation id="81A0A593FFD8FF99FE662C4FFDDCFE80" box="[494,599,335,361]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="4.[189,232,1815,1841]" captionStartId-1="6.[189,232,1958,1984]" captionTargetBox-0="[241,1345,267,1776]" captionTargetBox-1="[267,1371,262,1917]" captionTargetId-0="figure-17@4.[241,1346,265,1776]" captionTargetId-1="figure-17@6.[214,1371,262,1917]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionText-0="Fig. 1. A. Genetic relationships of cave wētā support monophyly of the genus Pharmacus Pictet &amp; de Saussure, 1893. Gene tree from mtDNA sequences (~1500 bp of COI) using Maximum Likelihood with 1000 bootstraps and Macropathus filifer Walker, 1869 to root the tree. B. Map of South Island of New Zealand, showing locations of Pharmacus specimens used in phylogenetic analyses (Figs 1A, 2). Colours correspond to different species. Two letter codes indicate the New Zealand entomological regions (Crosby et al. 1998). These codes are reported in the Material examined section for each species." captionText-1="Fig. 2. Gene tree for the eight morphologically identified species of Pharmacus Pictet &amp; de Saussure, 1893 using Maximum Likelihood analysis of ~850 bp of mtDNA (COI) from 60 specimens ofPharmacus." figureDoi-0="http://doi.org/10.5281/zenodo.6419370" figureDoi-1="http://doi.org/10.5281/zenodo.6419372" httpUri-0="https://zenodo.org/record/6419370/files/figure.png" httpUri-1="https://zenodo.org/record/6419372/files/figure.png" pageId="38" pageNumber="39">Figs 12</figureCitation>
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<figureCitation id="81A0A593FFD8FF99FDEB2C4FFDFFFE80" box="[611,628,335,361]" captionStart="Fig" captionStartId="12.[189,232,1945,1971]" captionTargetBox="[189,1398,273,1856]" captionTargetId="figure-17@12.[189,1398,273,1893]" captionTargetPageId="12" captionText="Fig. 6. Known distribution of cave wētā in the genus Pharmacus Pictet &amp; de Saussure, 1893 in the south of the South Island, New Zealand." figureDoi="http://doi.org/10.5281/zenodo.6419380" httpUri="https://zenodo.org/record/6419380/files/figure.png" pageId="38" pageNumber="39">6</figureCitation>
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<figureCitation id="81A0A593FFD8FF99FD082C4FFD2DFE80" box="[640,678,335,361]" captionStart="Fig" captionStartId="22.[189,232,1642,1668]" captionTargetBox="[347,1238,265,1605]" captionTargetId="figure-17@22.[347,1240,265,1605]" captionTargetPageId="22" captionText="Fig. 8. Dorsal views of adult ♂ cave wētā in the genus Pharmacus Pictet &amp; de Saussure, 1893. Notice loss of natural pigmentation of pale body parts due to preservation in ethanol. AB. Pharmacus montanus Pictet &amp; de Saussure, 1893 A. Mt Annette, Sealy Range, Mt Cook (MPN CW3302). B. Lake Anna, Mt Franklin, Arthurs Pass (MPN CW3989). C. Pharmacus senex sp. nov., Old Woman Range, Central Otago (MPN CW4387). D. Pharmacus cochleatus cochleatus (Karny, 1935) comb. nov., Topheavy, Mt Brewster, Haast Pass (MPN CW3324). E. Pharmacus cochleatus rawhiti subsp. nov., Mt Tūwhakarōria, Hector Mountains (NMNZ AI.052290). F. Pharmacus cristatus sp. nov., Skippers Range, South Westland (MPN CW4562). G. Pharmacus vallestris sp. nov., Matukituki River West Branch (MPN CW3700). H. Pharmacus notabilis sp. nov., Remarkables ski-field access road (NMNZ AI.052296). I. Pharmacus concinnus sp. nov., Eyre Peak, Eyre Mountains (MPN CW4482). J. Pharmacus perfidus sp. nov., Spence Peak, Takitimu Mountains (MPN CW4494). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6419384" httpUri="https://zenodo.org/record/6419384/files/figure.png" pageId="38" pageNumber="39">8H</figureCitation>
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<figureCitation id="81A0A593FFD8FF99FD3B2C4FFD5EFE80" box="[691,725,335,361]" captionStart="Fig" captionStartId="23.[189,232,1673,1699]" captionTargetBox="[213,1375,290,1608]" captionTargetId="figure-11@23.[213,1375,265,1629]" captionTargetPageId="23" captionText="Fig. 9. Left hind tibia of adult ♂ cave wētā in the genus Pharmacus Pictet &amp; de Saussure, 1893, dorsal view. A. Pharmacus montanus Pictet &amp; de Saussure, 1893, Lake Anna, Mt Franklin, Arthurs Pass (MPN CW3989). B. Pharmacus cochleatus cochleatus (Karny, 1935) comb. nov., Gertrude Saddle, Darran Mountains (MPN CW3413). C. Pharmacus cochleatus rawhiti subsp. nov., Mt Tūwhakarōria, Hector Mountains (MPN CW4431). D. Pharmacus cristatus sp. nov., Skippers Range, South Westland (NMNZ AI.052292). E. Pharmacus notabilis sp. nov., Remarkables ski-field access road (NMNZ AI.052296). F. Pharmacus senex sp. nov., Obelisk, Old Man Range (NMNZ AI.052294). G. Pharmacus concinnus sp. nov., Eyre Peak, Eyre Mountains (MPN CW4480). H. Pharmacus perfidus sp. nov., Spence Peak, Takitimu Mountains (NMNZ AI.052300). I. Pharmacus vallestris sp. nov., Matukituki River West Branch (MPN CW3700). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6419386" httpUri="https://zenodo.org/record/6419386/files/figure.png" pageId="38" pageNumber="39">9E</figureCitation>
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<figureCitation id="81A0A593FFD8FF99FD6A2C4FFCA5FE80" box="[738,814,335,361]" captionStart="Fig" captionStartId="26.[189,232,1782,1808]" captionTargetBox="[226,1375,266,1741]" captionTargetId="figure-18@26.[202,1384,263,1741]" captionTargetPageId="26" captionText="Fig. 10. Adult male terminalia of cave wētā in the genus Pharmacus Pictet &amp; de Saussure, 1893. Left column: dorsal view; central column: ventral view (subgenital plate); right column: lateral view. AC. Pharmacus montanus Pictet &amp; de Saussure, 1893, Mueller Hut Track, Sealy Range, Mt Cook (MPN CW3532). DF. Pharmacus cochleatus (Karny, 1935) comb. nov. DE. Humboldt Mountains (MPN CW3698). F. Blue Lake Creek, Garvie Mountains (MPN CW4462). GI. Pharmacus cristatus sp. nov., Skippers Range High Point, South Westland (NMNZ AI.052292). JL. Pharmacus notabilis sp. nov., Remarkables Ski-field access road (NMNZ AI.052296). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6419388" httpUri="https://zenodo.org/record/6419388/files/figure.png" pageId="38" pageNumber="39">10JL</figureCitation>
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<figureCitation id="81A0A593FFD8FF99FCB22C4FFC0DFE80" box="[826,902,335,361]" captionStart="Fig" captionStartId="31.[189,232,1776,1802]" captionTargetBox="[189,1378,569,1736]" captionTargetId="figure-110@31.[189,1398,545,1736]" captionTargetPageId="31" captionText="Fig. 12. Adult female terminalia of cave wētā in the genus Pharmacus Pictet &amp; de Saussure, 1893. Left column: subgenital plate; central and right columns: ovipositor. AC. Pharmacus montanus Pictet &amp; de Saussure, 1893. A. Mt Annette, Sealy Range, Mt Cook (MPN CW3303). BC. Mt Wakefield, Mount Cook Range (MPN CW3362). DF. Pharmacus cochleatus (Karny, 1935) comb. nov., Lochnagar, Richardson Mountains (MPN CW4590). GI. Pharmacus cristatus sp. nov., Skippers Range High Point, South Westland (NMNZ AI.052293). JL. Pharmacus notabilis sp. nov., Two Mile Hut, Hector Mountains (NMNZ AI.052297). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6419392" httpUri="https://zenodo.org/record/6419392/files/figure.png" pageId="38" pageNumber="39">12JL</figureCitation>
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<figureCitation id="81A0A593FFD8FF99FC1A2C4FFC4EFE83" box="[914,965,335,362]" captionStart="Fig" captionStartId="36.[189,232,1542,1568]" captionTargetBox="[189,1398,265,1482]" captionTargetId="figure-17@36.[189,1398,265,1482]" captionTargetPageId="36" captionText="Fig. 15. Live Pharmacus Pictet &amp; de Saussure, 1893 in their natural environments. AB. Pharmacus senex sp. nov. A. Adult ♂, Dunstan, Dunstan Mountains, 1670 m a.s.l. B. Adult ♂ and ♀. The Obelisk, Old Man Range, 1680 m a.s.l. C. Pharmacus notabilis sp. nov. Adult ♂, Remarkables Ski-field Access Road, Wakatipu, 1200 m a.s.l. D. Pharmacus concinnus sp. nov. Adult ♂, Symmetry Peaks, Eyre Mountains, 1500 m a.s.l. E. Pharmacus perfidus sp. nov. Adult ♀ and ♂, Spence Peak, Takitimu Mountains, 1450 m a.s.l. F. Pharmacus vallestris sp. nov. Adult ♀, French Ridge, Matukituki River West Branch, 1700 m a.s.l. (MPN CW5145)." figureDoi="http://doi.org/10.5281/zenodo.6419398" httpUri="https://zenodo.org/record/6419398/files/figure.png" pageId="38" pageNumber="39">15C</figureCitation>
,
<figureCitation id="81A0A593FFD8FF99FC592C4FFB8DFE80" box="[977,1030,335,361]" captionStart="Fig" captionStartId="39.[189,232,1535,1561]" captionTargetBox="[189,1398,687,1496]" captionTargetId="figure-105@39.[189,1398,687,1496]" captionTargetPageId="39" captionText="Fig. 16. Habitat of Pharmacus Pictet &amp; de Saussure, 1893.A. Sealy Range, Mount Cook.ab. Collection localities of Pharmacus montanus Pictet &amp; de Saussure, 1893. a. Mueller Hut Track, 1500 m a.s.l. (MPN CW3299, CW3300, CW35323535). b. Mount Annette, 2200 m a.s.l. (MPN CW3302, CW3303, CW3323). c. Mount Edgar Thomson, Ben Ohau Range, 2350 m a.s.l., collection locality of P. cochleatus cochleatus (Karny, 1935) comb. nov. (MPN CW3278). B. Lochnagar, Richardson Mountains. d. Rock bluffs at 2080 m a.s.l., collection locality of P. cochleatus cochleatus (Karny, 1935) comb. nov. (MPN CW4590). C. Matukituki River West Branch. ef. Collection localities of P. vallestris sp. nov. e. Valley floor, dry creek bed at 1000 m a.s.l. (MPN CW3430, CW3700). f. French Ridge, 1650 m a.s.l. (MPN CW5145, CW5146, CW52135215). gh. Collection localities of P. cochleatus cochleatus (Karny, 1935) comb. nov. g. French Ridge, 1800 m a.s.l. (MPN CW5207, CW5208). h. Bevan Col, 1850 m a.s.l. (OMNZ IV7927 and MPN CW5178 to 5180). D. The Remarkables, Queenstown. ij. Collection localities of Pharmacus notabilis sp. nov. i. Remarkables Ski-field Access Road, 1220 m a.s.l. (NMNZ AI.052296). j. The Remarkables North ridge, 1600 m a.s.l. (MPN CW2588 2594). k. Lake Alta, 1820 m a.s.l., collection locality of Pharmacus cochleatus rawhiti subsp. nov. (MPN CW5219 5221)." figureDoi="http://doi.org/10.5281/zenodo.6419400" httpUri="https://zenodo.org/record/6419400/files/figure.png" pageId="38" pageNumber="39">16D</figureCitation>
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<figureCitation id="81A0A593FFD8FF99FB9B2C4FFBCEFE80" box="[1043,1093,335,361]" captionStart="Fig" captionStartId="46.[189,232,1810,1836]" captionTargetBox="[189,1398,802,1769]" captionTargetId="figure-184@46.[189,1398,797,1769]" captionTargetPageId="46" captionText="Fig. 18. Map of South Island, New Zealand, showing the known distribution of cave wētā in the genus Pharmacus Pictet &amp; de Saussure, 1893. AB. Solid shapes indicate material collected as part of this study; empty circles indicate additional locations of material examined by Richards (1972). B. Shapes represent different subspecies: dark blue circle = Pharmacus cochleatus cochleatus; light blue circle = Pharmacus cochleatus nauclerus subsp. nov.; diamond = Pharmacus cochleatus fiordensis subsp. nov.; triangle = Pharmacus cochleatus rawhiti subsp. nov." figureDoi="http://doi.org/10.5281/zenodo.6419404" httpUri="https://zenodo.org/record/6419404/files/figure.png" pageId="38" pageNumber="39">18D</figureCitation>
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<emphasis id="2BEF6504FFD8FF99FF352C90FEBDFE43" bold="true" box="[189,310,400,426]" pageId="38" pageNumber="39">Diagnosis</emphasis>
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<paragraph id="1924B916FFD8FF99FF352CC0FE53FDAA" blockId="38.[189,1399,447,579]" pageId="38" pageNumber="39">
A mid-sized cave wētā found in the low alpine regions of The Remarkables and Hector Mountains, east of Lake Wakatipu, at elevations between
<quantity id="DE6314F3FFD8FF99FD522CE3FCBFFE14" box="[730,820,483,509]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="38" pageNumber="39" unit="m" value="1200.0">1200 m</quantity>
and
<quantity id="DE6314F3FFD8FF99FCE62CE2FC40FE14" box="[878,971,482,509]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="38" pageNumber="39" unit="m" value="1600.0">1600 m</quantity>
. Body colour is chequered turquois/ grey and eye colour is blue/grey. A prolateral apical spine on the fore femur is always present in males, occasionally in females.
</paragraph>
<paragraph id="1924B916FFD8FF99FF352F6AFA82FCFB" blockId="38.[186,1399,618,786]" pageId="38" pageNumber="39">
The sympatric
<taxonomicName id="DE9BC295FFD8FF99FEEC2F6AFD4EFD6D" authority="Hegg &amp; Morgan-Richards &amp; Trewick, 2022" authorityName="Hegg &amp; Morgan-Richards &amp; Trewick" authorityYear="2022" box="[356,709,618,644]" class="Insecta" family="Rhaphidophoridae" genus="Pharmacus" kingdom="Animalia" order="Orthoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="subSpecies" species="cochleatus" status="subsp. nov." subSpecies="rawhiti">
<emphasis id="2BEF6504FFD8FF99FEEC2F6AFD4EFD6D" box="[356,709,618,644]" italics="true" pageId="38" pageNumber="39">Pharmacus cochleatus rawhiti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30DCD87FFFD8FF99FD442F6AFCC7FD6C" box="[716,844,618,645]" pageId="38" pageNumber="39" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
is mainly found at higher elevations and is easily differentiated by its brown tergites and reddish legs and antennae.Two other species of
<taxonomicName id="DE9BC295FFD8FF99FB132F8DFAFFFD4E" authorityName="Walker" authorityYear="1869" box="[1179,1396,653,679]" class="Insecta" family="Rhaphidophoridae" kingdom="Animalia" order="Orthoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="family">Rhaphidophoridae</taxonomicName>
share the habitat with
<taxonomicName id="DE9BC295FFD8FF99FE422FB2FDDBFD22" authority="Hegg &amp; Morgan-Richards &amp; Trewick, 2022" authorityName="Hegg &amp; Morgan-Richards &amp; Trewick" authorityYear="2022" box="[458,592,689,715]" class="Insecta" family="Rhaphidophoridae" genus="Pharmacus" kingdom="Animalia" order="Orthoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="notabilis" status="sp. nov.">
<emphasis id="2BEF6504FFD8FF99FE422FB2FDDBFD22" box="[458,592,689,715]" italics="true" pageId="38" pageNumber="39">P. notabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30DCD87FFFD8FF99FDD22FB1FD31FD22" box="[602,698,689,715]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
, these are
<taxonomicName id="DE9BC295FFD8FF99FCB42FB1FBA1FD22" baseAuthorityName="Karny" baseAuthorityYear="1937" box="[828,1066,689,715]" class="Insecta" family="Rhaphidophoridae" genus="Talitropsis" kingdom="Animalia" order="Orthoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="chopardi">
<emphasis id="2BEF6504FFD8FF99FCB42FB1FBA1FD22" box="[828,1066,689,715]" italics="true" pageId="38" pageNumber="39">Talitropsis chopardi</emphasis>
</taxonomicName>
and
<taxonomicName id="DE9BC295FFD8FF99FBE02FB1FAA5FD22" box="[1128,1326,689,715]" class="Insecta" family="Rhaphidophoridae" genus="Macropathus" kingdom="Animalia" order="Orthoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="2BEF6504FFD8FF99FBE02FB1FA88FD22" box="[1128,1283,689,715]" italics="true" pageId="38" pageNumber="39">Macropathus</emphasis>
sp.
</taxonomicName>
. The former has shorter legs and fewer, larger spines on the hind tibiae, and unarmed hind tarsi; the latter has uniform brown colour and much longer legs, which give the insect a very slender appearance.
</paragraph>
</subSubSection>
<subSubSection id="5181EA9DFFD8FF99FF352E39FB07FC4D" pageId="38" pageNumber="39" type="etymology">
<paragraph id="1924B916FFD8FF99FF352E39FECFFCBA" blockId="38.[189,324,825,851]" box="[189,324,825,851]" pageId="38" pageNumber="39">
<heading id="426C0E7AFFD8FF99FF352E39FECFFCBA" bold="true" box="[189,324,825,851]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="2BEF6504FFD8FF99FF352E39FECFFCBA" bold="true" box="[189,324,825,851]" pageId="38" pageNumber="39">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="1924B916FFD8FF99FF352E67FB07FC4D" blockId="38.[189,1399,871,933]" pageId="38" pageNumber="39">
<taxonomicName id="DE9BC295FFD8FF99FF4F2E67FEB8FC68" authorityName="Hegg &amp; Morgan-Richards &amp; Trewick" authorityYear="2022" box="[199,307,871,897]" class="Insecta" family="Rhaphidophoridae" genus="Pharmacus" kingdom="Animalia" order="Orthoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="notabilis">
<emphasis id="2BEF6504FFD8FF99FF4F2E67FEB8FC68" box="[199,307,871,897]" italics="true" pageId="38" pageNumber="39">Nŏtābĭlis</emphasis>
</taxonomicName>
is Latin for remarkable, after the location where the species was first discovered, The Remarkables near Queenstown.
<taxonomicName id="DE9BC295FFD8FF99FDB12E8AFCA7FC4D" authorityName="Hegg &amp; Morgan-Richards &amp; Trewick" authorityYear="2022" box="[569,812,906,932]" class="Insecta" family="Rhaphidophoridae" genus="Pharmacus" kingdom="Animalia" order="Orthoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="notabilis">
<emphasis id="2BEF6504FFD8FF99FDB12E8AFCA7FC4D" box="[569,812,906,932]" italics="true" pageId="38" pageNumber="39">Pharmacus notabilis</emphasis>
</taxonomicName>
means remarkable sorcerer.
</paragraph>
</subSubSection>
<subSubSection id="5181EA9DFFD8FF99FF352ECBFB7EF98B" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="1924B916FFD8FF99FF352ECBFCAEFC0F" blockId="38.[189,805,971,998]" box="[189,805,971,998]" pageId="38" pageNumber="39">
<emphasis id="2BEF6504FFD8FF99FF352ECBFE21FC0C" bold="true" box="[189,426,971,997]" pageId="38" pageNumber="39">Material examined</emphasis>
(see also Supp. file 1: Table S7)
</paragraph>
<paragraph id="1924B916FFD8FF99FF622EFAFC1EFB96" blockId="38.[189,1399,1018,1151]" pageId="38" pageNumber="39">
<materialsCitation id="A9F3B34BFFD8FF99FF622EFAFC1EFB96" ID-GBIF-Occurrence="3748656370" collectedFrom="under boulders in road escarpment" collectingDate="2017-05-12" collectingMethod="night search + insect net" collectionCode="NMNZ" collectorName="D. Hegg" country="New Zealand" elevation="1220" latitude="-45.02651" location="Central Otago" longLatPrecision="1" longitude="168.79749" municipality="Central" pageId="38" pageNumber="39" specimenCode="AI.052296" specimenCount="1" specimenCount-male="1" stateProvince="Otago" typeStatus="holotype">
<emphasis id="2BEF6504FFD8FF99FF622EFAFED7FBFD" bold="true" box="[234,348,1018,1044]" pageId="38" pageNumber="39">
<typeStatus id="C62007B4FFD8FF99FF622EFAFED7FBFD" box="[234,348,1018,1044]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="618CF986FFD8FF99FF35291EFE18FBD1" box="[189,403,1054,1080]" name="New Zealand" pageId="38" pageNumber="39">NEW ZEALAND</collectingCountry>
<specimenCount id="0F9D729FFFD8FF99FE38291EFE43FBD1" box="[432,456,1054,1080]" pageId="38" pageNumber="39" type="male"></specimenCount>
, adult;
<location id="1C44EFCDFFD8FF99FDAB291DFD40FBD1" LSID="urn:lsid:plazi:treatment:91320800FFD8FF97FDE42C08FA80FE80:1C44EFCDFFD8FF99FDAB291DFD40FBD1" box="[547,715,1053,1080]" country="New Zealand" latitude="-45.02651" longLatPrecision="1" longitude="168.79749" municipality="Central" name="Central Otago" pageId="38" pageNumber="39" stateProvince="Otago">Central Otago</location>
(CO),
<location id="1C44EFCDFFD8FF99FCA9291DFB45FBDE" LSID="urn:lsid:plazi:treatment:91320800FFD8FF97FDE42C08FA80FE80:1C44EFCDFFD8FF99FCA9291DFB45FBDE" box="[801,1230,1053,1080]" country="New Zealand" latitude="-45.02651" longLatPrecision="1" longitude="168.79749" municipality="Central" name="Remarkables Ski-field Access Road" pageId="38" pageNumber="39" stateProvince="Otago">Remarkables Ski-field Access Road</location>
;
<geoCoordinate id="7CAFDFD1FFD8FF99FB54291DFAFAFBD1" box="[1244,1393,1053,1080]" degrees="45.02651" direction="south" orientation="latitude" pageId="38" pageNumber="39" precision="1" value="-45.02651">45.02651° S</geoCoordinate>
,
<geoCoordinate id="7CAFDFD1FFD8FF99FF352941FEE9FBB2" box="[189,354,1089,1115]" degrees="168.79748" direction="east" orientation="longitude" pageId="38" pageNumber="39" precision="1" value="168.79749">168.79748° E</geoCoordinate>
;
<elevation id="92B65E25FFD8FF99FEFC2941FD98FBB2" box="[372,531,1089,1115]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="38" pageNumber="39" unit="m" value="1220.0">
<quantity id="DE6314F3FFD8FF99FEFC2941FE5BFBB2" box="[372,464,1089,1115]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="38" pageNumber="39" unit="m" value="1220.0">1220 m</quantity>
a.s.l.
</elevation>
;
<date id="6D259FD6FFD8FF99FDAB2941FD42FBB2" box="[547,713,1089,1115]" pageId="38" pageNumber="39" value="2017-05-12">
<collectingDate id="7D61663EFFD8FF99FDAB2941FD42FBB2" box="[547,713,1089,1115]" pageId="38" pageNumber="39" value="2017-05-12">12 May 2017</collectingDate>
</date>
;
<collectorName id="B46EDCC0FFD8FF99FD502941FCB4FBB2" box="[728,831,1089,1115]" pageId="38" pageNumber="39">D. Hegg</collectorName>
leg.;
<collectedFrom id="AA87FAEBFFD8FF99FC0E2941FAA0FBB2" box="[902,1323,1089,1115]" pageId="38" pageNumber="39">under boulders in road escarpment</collectedFrom>
;
<collectingMethod id="C0DAC101FFD8FF99FAB22941FE11FB97" pageId="38" pageNumber="39">night search + insect net</collectingMethod>
;
<collectionCode id="7F8A21D3FFD8FF99FE2E2964FE77FB97" box="[422,508,1124,1150]" country="New Zealand" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="38" pageNumber="39" type="Museum">NMNZ</collectionCode>
<specimenCode id="493D116DFFD8FF99FD8A2964FD0BFB96" box="[514,640,1124,1151]" pageId="38" pageNumber="39">AI.052296</specimenCode>
(prev. MPN CW3528).
</materialsCitation>
</paragraph>
<paragraph id="1924B916FFD8FF99FF6229A5FD26FAC0" blockId="38.[189,1399,1189,1322]" pageId="38" pageNumber="39">
<materialsCitation id="A9F3B34BFFD8FF99FF6229A5FD22FAC0" ID-GBIF-Occurrence="3748656390" collectedFrom="on large boulders" collectingDate="2019-04-26" collectingMethod="night search + insect net" collectionCode="NMNZ" collectorName="D. Hegg" country="New Zealand" county="Central" elevation="1460" latitude="-45.17392" location="Central Otago" longLatPrecision="1" longitude="168.8027" municipality="Two Mile Hut" pageId="38" pageNumber="39" specimenCode="AI.052297" specimenCount="1" specimenCount-female="1" stateProvince="Otago" typeStatus="paratype">
<emphasis id="2BEF6504FFD8FF99FF6229A5FED7FB56" bold="true" box="[234,348,1189,1215]" pageId="38" pageNumber="39">
<typeStatus id="C62007B4FFD8FF99FF6229A5FED7FB56" box="[234,348,1189,1215]" pageId="38" pageNumber="39" type="paratype">Paratype</typeStatus>
</emphasis>
<collectingCountry id="618CF986FFD8FF99FF3529C9FE18FB0A" box="[189,403,1225,1251]" name="New Zealand" pageId="38" pageNumber="39">NEW ZEALAND</collectingCountry>
<specimenCount id="0F9D729FFFD8FF99FE3829C9FE55FB0A" box="[432,478,1225,1251]" pageId="38" pageNumber="39" type="female">1 ♀</specimenCount>
, adult;
<location id="1C44EFCDFFD8FF99FDB329C8FD68FB0A" LSID="urn:lsid:plazi:treatment:91320800FFD8FF97FDE42C08FA80FE80:1C44EFCDFFD8FF99FDB329C8FD68FB0A" box="[571,739,1224,1251]" country="New Zealand" county="Central" latitude="-45.17392" longLatPrecision="1" longitude="168.8027" municipality="Two Mile Hut" name="Central Otago" pageId="38" pageNumber="39" stateProvince="Otago">Central Otago</location>
(CO),
<collectingMunicipality id="F940236CFFD8FF99FCB129C9FC6DFB0A" box="[825,998,1224,1251]" pageId="38" pageNumber="39">Two Mile Hut</collectingMunicipality>
,
<location id="1C44EFCDFFD8FF99FC7B29C9FB45FB0B" LSID="urn:lsid:plazi:treatment:91320800FFD8FF97FDE42C08FA80FE80:1C44EFCDFFD8FF99FC7B29C9FB45FB0B" box="[1011,1230,1224,1251]" country="New Zealand" county="Central" latitude="-45.17392" longLatPrecision="1" longitude="168.8027" municipality="Two Mile Hut" name="Hector Mountains" pageId="38" pageNumber="39" stateProvince="Otago">Hector Mountains</location>
;
<geoCoordinate id="7CAFDFD1FFD8FF99FB5429C8FAFAFB0A" box="[1244,1393,1224,1251]" degrees="45.17392" direction="south" orientation="latitude" pageId="38" pageNumber="39" precision="1" value="-45.17392">45.17392° S</geoCoordinate>
,
<geoCoordinate id="7CAFDFD1FFD8FF99FF3529ECFEEBFAEF" box="[189,352,1260,1286]" degrees="168.80270" direction="east" orientation="longitude" pageId="38" pageNumber="39" precision="1" value="168.8027">168.80270° E</geoCoordinate>
;
<elevation id="92B65E25FFD8FF99FEF829ECFD80FAEF" box="[368,523,1260,1286]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="38" pageNumber="39" unit="m" value="1460.0">
<quantity id="DE6314F3FFD8FF99FEF829ECFE40FAEF" box="[368,459,1260,1286]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="38" pageNumber="39" unit="m" value="1460.0">1460 m</quantity>
a.s.l.
</elevation>
;
<date id="6D259FD6FFD8FF99FD9229ECFD33FAEF" box="[538,696,1260,1286]" pageId="38" pageNumber="39" value="2019-04-26">
<collectingDate id="7D61663EFFD8FF99FD9229ECFD33FAEF" box="[538,696,1260,1286]" pageId="38" pageNumber="39" value="2019-04-26">26 Apr. 2019</collectingDate>
</date>
;
<collectorName id="B46EDCC0FFD8FF99FD4D29ECFCA1FAEF" box="[709,810,1260,1286]" pageId="38" pageNumber="39">D. Hegg</collectorName>
leg.;
<collectedFrom id="AA87FAEBFFD8FF99FCE629ECFBCBFAEF" box="[878,1088,1260,1286]" pageId="38" pageNumber="39">on large boulders</collectedFrom>
;
<collectingMethod id="C0DAC101FFD8FF99FBC429ECFAFAFAEF" box="[1100,1393,1260,1286]" pageId="38" pageNumber="39">night search + insect net</collectingMethod>
;
<collectionCode id="7F8A21D3FFD8FF99FF36280FFE9FFAC0" box="[190,276,1295,1321]" country="New Zealand" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="38" pageNumber="39" type="Museum">NMNZ</collectionCode>
<specimenCode id="493D116DFFD8FF99FE92280FFE13FAC3" box="[282,408,1295,1322]" pageId="38" pageNumber="39">AI.052297</specimenCode>
(prev. MPN CW4394)
</materialsCitation>
.
</paragraph>
<paragraph id="1924B916FFD8FF99FF622850FE2DFA83" blockId="38.[189,1399,1360,1634]" box="[234,422,1360,1386]" pageId="38" pageNumber="39">
<emphasis id="2BEF6504FFD8FF99FF622850FE2DFA83" bold="true" box="[234,422,1360,1386]" pageId="38" pageNumber="39">Other material</emphasis>
</paragraph>
<paragraph id="1924B916FFD8FF99FF352874FB7EF98B" blockId="38.[189,1399,1360,1634]" pageId="38" pageNumber="39">
<materialsCitation id="A9F3B34BFFD8FF99FF352874FE6DFA3C" ID-GBIF-Occurrence="3748656423" accessionNumber="OM293682, OM293683" collectedFrom="in cracks in rock bluffs" collectingDate="2013-01" collectionCode="MPN" collectorName="T. Jewell" country="New Zealand" elevation="1600" latitude="-45.0413" location="Remarkables North ridge" longLatPrecision="6" longitude="168.7977" pageId="38" pageNumber="39" specimenCode="CW2587, CW2589" specimenCount="2" specimenCount-male="2" stateProvince="Otago">
<collectingCountry id="618CF986FFD8FF99FF352874FE1BFA67" box="[189,400,1396,1422]" name="New Zealand" pageId="38" pageNumber="39">NEW ZEALAND</collectingCountry>
<emphasis id="2BEF6504FFD8FF99FE262873FD22FA64" bold="true" box="[430,681,1395,1421]" pageId="38" pageNumber="39">
Central
<collectingRegion id="DB5F77F4FFD8FF99FD9D2873FDE9FA64" box="[533,610,1395,1421]" country="New Zealand" name="Otago" pageId="38" pageNumber="39">Otago</collectingRegion>
(CO)
</emphasis>
<specimenCount id="0F9D729FFFD8FF99FD492874FC8DFA67" box="[705,774,1396,1422]" pageId="38" pageNumber="39" type="male">2 ♂♂</specimenCount>
;
<location id="1C44EFCDFFD8FF99FC9B2873FBB4FA64" LSID="urn:lsid:plazi:treatment:91320800FFD8FF97FDE42C08FA80FE80:1C44EFCDFFD8FF99FC9B2873FBB4FA64" box="[787,1087,1395,1421]" country="New Zealand" latitude="-45.0413" longLatPrecision="6" longitude="168.7977" name="Remarkables North ridge" pageId="38" pageNumber="39" stateProvince="Otago">Remarkables North ridge</location>
;
<geoCoordinate id="7CAFDFD1FFD8FF99FBC22873FB44FA67" box="[1098,1231,1395,1422]" degrees="45.0413" direction="south" orientation="latitude" pageId="38" pageNumber="39" precision="5" value="-45.0413">45.0413° S</geoCoordinate>
,
<geoCoordinate id="7CAFDFD1FFD8FF99FB532873FAFAFA67" box="[1243,1393,1395,1422]" degrees="168.7977" direction="east" orientation="longitude" pageId="38" pageNumber="39" precision="5" value="168.7977">168.7977° E</geoCoordinate>
;
<elevation id="92B65E25FFD8FF99FF352897FED0FA58" box="[189,347,1431,1457]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="38" pageNumber="39" unit="m" value="1600.0">
<quantity id="DE6314F3FFD8FF99FF352897FE92FA58" box="[189,281,1431,1457]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="38" pageNumber="39" unit="m" value="1600.0">1600 m</quantity>
a.s.l.
</elevation>
;
<date id="6D259FD6FFD8FF99FEE32897FE69FA58" box="[363,482,1431,1457]" pageId="38" pageNumber="39" value="2013-01">
<collectingDate id="7D61663EFFD8FF99FEE32897FE69FA58" box="[363,482,1431,1457]" pageId="38" pageNumber="39" value="2013-01">Jan. 2013</collectingDate>
</date>
;
<collectorName id="B46EDCC0FFD8FF99FE782897FDD7FA58" box="[496,604,1431,1457]" pageId="38" pageNumber="39">T. Jewell</collectorName>
leg.;
<collectedFrom id="AA87FAEBFFD8FF99FD2B2897FC4BFA58" box="[675,960,1431,1457]" pageId="38" pageNumber="39">in cracks in rock bluffs</collectedFrom>
; GenBank:
<accessionNumber id="06C824F5FFD8FF99FBC72897FB53FA58" box="[1103,1240,1431,1457]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OM293682" pageId="38" pageNumber="39">OM293682</accessionNumber>
,
<accessionNumber id="06C824F5FFD8FF99FB6F2897FAF8FA58" box="[1255,1395,1431,1457]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OM293683" pageId="38" pageNumber="39">OM293683</accessionNumber>
;
<collectionCode id="7F8A21D3FFD8FF99FF3528BBFF76FA3C" box="[189,253,1467,1493]" pageId="38" pageNumber="39">MPN</collectionCode>
<specimenCode id="493D116DFFD8FF99FE8C28BAFEFAFA3C" box="[260,369,1466,1493]" collectionCode="MPN" pageId="38" pageNumber="39">CW2587</specimenCode>
,
<specimenCode id="493D116DFFD8FF99FEF428BAFE6DFA3C" box="[380,486,1466,1493]" collectionCode="MPN" pageId="38" pageNumber="39">CW2589</specimenCode>
</materialsCitation>
<materialsCitation id="A9F3B34BFFD8FF99FE7728BAFECEFA11" ID-GBIF-Occurrence="3748656339" accessionNumber="OM293682, OM293683" collectedFrom="in cracks in rock bluffs" collectingDate="2013-01" collectionCode="MPN" collectorName="T. Jewell &amp; GenBank" country="New Zealand" elevation="1600" latitude="-45.0413" location="Remarkables North" longLatPrecision="6" longitude="168.7977" pageId="38" pageNumber="39" specimenCode="CW2588, CW2590, CW2594" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="Central">
<specimenCount id="0F9D729FFFD8FF99FE7728BAFDA6FA3C" box="[511,557,1466,1493]" pageId="38" pageNumber="39" type="male">1 ♂</specimenCount>
,
<specimenCount id="0F9D729FFFD8FF99FDB128BAFDF7FA3C" box="[569,636,1466,1493]" pageId="38" pageNumber="39" type="female">5 ♀♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="7F8A21D3FFD8FF99FBC528BBFB06FA3C" box="[1101,1165,1467,1493]" pageId="38" pageNumber="39">MPN</collectionCode>
<specimenCode id="493D116DFFD8FF99FB1C28BAFA8AFA3C" box="[1172,1281,1466,1493]" collectionCode="MPN" pageId="38" pageNumber="39">CW2588</specimenCode>
,
<specimenCode id="493D116DFFD8FF99FA8528BAFAFCFA3C" box="[1293,1399,1466,1493]" collectionCode="MPN" pageId="38" pageNumber="39">CW2590</specimenCode>
to
<specimenCode id="493D116DFFD8FF99FF5328DDFECEFA11" box="[219,325,1501,1528]" collectionCode="MPN" pageId="38" pageNumber="39">CW2594</specimenCode>
</materialsCitation>
<materialsCitation id="A9F3B34BFFD8FF99FED628DDFA9FFA11" ID-GBIF-Occurrence="3748656414" accessionNumber="OM293701" box="[350,1300,1501,1528]" collectedFrom="under boulders in road escarpment" collectingDate="2017-05-12" collectingMethod="night search + insect net" collectionCode="MPN" collectorName="GenBank" country="New Zealand" elevation="1220" latitude="-45.02651" location="Remarkables Ski-field Access Road" longLatPrecision="1" longitude="168.79749" municipality="Central" pageId="38" pageNumber="39" specimenCode="CW3509" specimenCount="1" specimenCount-female="1" stateProvince="Central">
<specimenCount id="0F9D729FFFD8FF99FED628DDFE00FA11" box="[350,395,1501,1528]" pageId="38" pageNumber="39" type="female">1 ♀</specimenCount>
; same collection data as for holotype; GenBank:
<accessionNumber id="06C824F5FFD8FF99FC4528DDFBD3FA1E" box="[973,1112,1501,1527]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OM293701" pageId="38" pageNumber="39">OM293701</accessionNumber>
;
<collectionCode id="7F8A21D3FFD8FF99FBEC28DEFB2FFA11" box="[1124,1188,1502,1528]" pageId="38" pageNumber="39">MPN</collectionCode>
<specimenCode id="493D116DFFD8FF99FB2228DDFA9FFA11" box="[1194,1300,1501,1528]" collectionCode="MPN" pageId="38" pageNumber="39">CW3509</specimenCode>
</materialsCitation>
<materialsCitation id="A9F3B34BFFD8FF99FAA528DDFE3FF9D7" ID-GBIF-Occurrence="3748656430" collectedFrom="under boulders in road escarpment" collectingDate="2017-05-12" collectingMethod="night search + insect net" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" elevation="1220" latitude="-45.02651" location="Remarkables Ski-field Access Road" longLatPrecision="1" longitude="168.79749" municipality="Central" pageId="38" pageNumber="39" specimenCode="CW3510, CW3513, CW3524, CW3527, CW3529, CW3531" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Central">
<specimenCount id="0F9D729FFFD8FF99FAA528DDFAFAFA11" box="[1325,1393,1501,1528]" pageId="38" pageNumber="39" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="0F9D729FFFD8FF99FF352B01FF75F9F2" box="[189,254,1537,1563]" pageId="38" pageNumber="39" type="female">3 ♀♀</specimenCount>
, 5 nymphs; same collection data as for holotype;
<collectionCode id="7F8A21D3FFD8FF99FCBB2B01FCF8F9F2" box="[819,883,1537,1563]" pageId="38" pageNumber="39">MPN</collectionCode>
<specimenCode id="493D116DFFD8FF99FCF02B00FC6AF9F2" box="[888,993,1536,1563]" collectionCode="MPN" pageId="38" pageNumber="39">CW3510</specimenCode>
to
<specimenCode id="493D116DFFD8FF99FB8B2B00FBFBF9F2" box="[1027,1136,1536,1563]" collectionCode="MPN" pageId="38" pageNumber="39">CW3513</specimenCode>
,
<specimenCode id="493D116DFFD8FF99FBF22B00FB68F9F2" box="[1146,1251,1536,1563]" collectionCode="MPN" pageId="38" pageNumber="39">CW3524</specimenCode>
to
<specimenCode id="493D116DFFD8FF99FA8D2B00FAF9F9F2" box="[1285,1394,1536,1563]" collectionCode="MPN" pageId="38" pageNumber="39">CW3527</specimenCode>
,
<specimenCode id="493D116DFFD8FF99FF352B23FEACF9D7" box="[189,295,1571,1598]" collectionCode="MPN" pageId="38" pageNumber="39">CW3529</specimenCode>
to
<specimenCode id="493D116DFFD8FF99FEC22B23FE3FF9D7" box="[330,436,1571,1598]" collectionCode="MPN" pageId="38" pageNumber="39">CW3531</specimenCode>
</materialsCitation>
<materialsCitation id="A9F3B34BFFD8FF99FE432B24FAFCF9D7" ID-GBIF-Occurrence="3748656335" accessionNumber="OM293716" box="[459,1399,1571,1598]" collectedFrom="on large boulders" collectingDate="2019-04-26" collectingMethod="night search + insect net" collectionCode="MPN" collectorName="GenBank" country="New Zealand" county="Central" elevation="1460" latitude="-45.17392" location="Hector Mountains" longLatPrecision="1" longitude="168.8027" municipality="Two Mile Hut" pageId="38" pageNumber="39" specimenCode="CW4391" specimenCount="1" specimenCount-male="1" stateProvince="Central">
<specimenCount id="0F9D729FFFD8FF99FE432B24FE72F9D7" box="[459,505,1572,1598]" pageId="38" pageNumber="39" type="male">1 ♂</specimenCount>
; same collection data as for paratype; GenBank:
<accessionNumber id="06C824F5FFD8FF99FBB92B23FB37F9D4" box="[1073,1212,1571,1597]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/OM293716" pageId="38" pageNumber="39">OM293716</accessionNumber>
;
<collectionCode id="7F8A21D3FFD8FF99FB4F2B24FA8CF9D7" box="[1223,1287,1572,1598]" pageId="38" pageNumber="39">MPN</collectionCode>
<specimenCode id="493D116DFFD8FF99FA852B24FAFCF9D7" box="[1293,1399,1572,1598]" collectionCode="MPN" pageId="38" pageNumber="39">CW4391</specimenCode>
</materialsCitation>
<materialsCitation id="A9F3B34BFFD8FF99FF472B47FB7AF98B" ID-GBIF-Occurrence="3748656401" box="[207,1265,1607,1634]" collectedFrom="on large boulders" collectingDate="2019-04-26" collectingMethod="night search + insect net" collectionCode="MPN" collectorName="D. Hegg" country="New Zealand" county="Central" elevation="1460" latitude="-45.17392" location="Hector Mountains" longLatPrecision="1" longitude="168.8027" municipality="Two Mile Hut" pageId="38" pageNumber="39" specimenCode="CW4392, CW4393, CW4500" specimenCount="2" specimenCount-male="2" stateProvince="Central">
<specimenCount id="0F9D729FFFD8FF99FF472B47FE98F988" box="[207,275,1607,1633]" pageId="38" pageNumber="39" type="male">2 ♂♂</specimenCount>
, 1 nymph; same collection data as for paratype;
<collectionCode id="7F8A21D3FFD8FF99FCC32B47FC00F988" box="[843,907,1607,1633]" pageId="38" pageNumber="39">MPN</collectionCode>
<specimenCode id="493D116DFFD8FF99FC1A2B47FC75F988" box="[914,1022,1607,1633]" collectionCode="MPN" pageId="38" pageNumber="39">CW4392</specimenCode>
,
<specimenCode id="493D116DFFD8FF99FB822B47FBFCF988" box="[1034,1143,1607,1633]" collectionCode="MPN" pageId="38" pageNumber="39">CW4393</specimenCode>
,
<specimenCode id="493D116DFFD8FF99FB0B2B47FB7AF98B" box="[1155,1265,1607,1634]" collectionCode="MPN" pageId="38" pageNumber="39">CW4500</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5181EA9DFFD8FF97FF352B88FA80FE80" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39" type="description">
<paragraph id="1924B916FFD8FF99FF352B88FEC5F94B" blockId="38.[189,334,1672,1698]" box="[189,334,1672,1698]" pageId="38" pageNumber="39">
<heading id="426C0E7AFFD8FF99FF352B88FEC5F94B" bold="true" box="[189,334,1672,1698]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="2BEF6504FFD8FF99FF352B88FEC5F94B" bold="true" box="[189,334,1672,1698]" pageId="38" pageNumber="39">Description</emphasis>
</heading>
</paragraph>
<paragraph id="1924B916FFD8FF99FF352BB7FD87F938" blockId="38.[189,524,1719,1745]" box="[189,524,1719,1745]" pageId="38" pageNumber="39">
MEASUREMENTS. See
<tableCitation id="54198CADFFD8FF99FE262BB7FD87F938" box="[430,524,1719,1745]" captionStart="Table 1" captionStartId="14.[189,259,265,291]" captionTargetPageId="14" captionText="Table 1 (continued on next three pages). Dimensions and spine count of species of Pharmacus Pictet &amp; de Saussure, 1893." httpUri="http://table.plazi.org/id/4DE4E99EFFF0FFB1FF352C09FE13FEAF" pageId="38" pageNumber="39" tableUuid="4DE4E99EFFF0FFB1FF352C09FE13FEAF">Table 1.</tableCitation>
</paragraph>
<paragraph id="1924B916FFD8FF99FF352BF9FC5FF8FB" blockId="38.[189,980,1784,1811]" box="[189,980,1784,1811]" pageId="38" pageNumber="39">HEAD. As per generic description. Eyes grey/blue. Antennae brown.</paragraph>
<paragraph id="1924B916FFD8FF99FF352A3AFDE8F8BD" blockId="38.[189,611,1850,1876]" box="[189,611,1850,1876]" pageId="38" pageNumber="39">THORAX. As per generic description.</paragraph>
<paragraph id="1924B916FFD8FF98FF352A7CFC07FEAF" blockId="38.[189,1399,1915,2012]" lastBlockId="39.[189,1398,265,327]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">LEGS. The hind tibiae are approx. 10% Shorter than the body in females, equally long as body in males. Fore femora always armed at the apex with one prolateral spine in males, occasionally also in females. Mid femora armed with one retrolateral spine at the apex; prolateral apical spine absent. The first hind tarsus segment is armed with a variable number of linear spines above (up to eight); the second hind tarsus segment is only rarely armed with dorsal linear spines.</paragraph>
<paragraph id="1924B916FFD9FF98FF352C6EFE7EFE26" blockId="39.[189,1399,366,463]" pageId="39" pageNumber="40">
ABDOMEN. Tergites glabrous or sparsely tomentose (
<figureCitation id="81A0A593FFD9FF98FCA62C6EFC04FE61" box="[814,911,366,392]" captionStart="Fig" captionStartId="22.[189,232,1642,1668]" captionTargetBox="[347,1238,265,1605]" captionTargetId="figure-17@22.[347,1240,265,1605]" captionTargetPageId="22" captionText="Fig. 8. Dorsal views of adult ♂ cave wētā in the genus Pharmacus Pictet &amp; de Saussure, 1893. Notice loss of natural pigmentation of pale body parts due to preservation in ethanol. AB. Pharmacus montanus Pictet &amp; de Saussure, 1893 A. Mt Annette, Sealy Range, Mt Cook (MPN CW3302). B. Lake Anna, Mt Franklin, Arthurs Pass (MPN CW3989). C. Pharmacus senex sp. nov., Old Woman Range, Central Otago (MPN CW4387). D. Pharmacus cochleatus cochleatus (Karny, 1935) comb. nov., Topheavy, Mt Brewster, Haast Pass (MPN CW3324). E. Pharmacus cochleatus rawhiti subsp. nov., Mt Tūwhakarōria, Hector Mountains (NMNZ AI.052290). F. Pharmacus cristatus sp. nov., Skippers Range, South Westland (MPN CW4562). G. Pharmacus vallestris sp. nov., Matukituki River West Branch (MPN CW3700). H. Pharmacus notabilis sp. nov., Remarkables ski-field access road (NMNZ AI.052296). I. Pharmacus concinnus sp. nov., Eyre Peak, Eyre Mountains (MPN CW4482). J. Pharmacus perfidus sp. nov., Spence Peak, Takitimu Mountains (MPN CW4494). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.6419384" httpUri="https://zenodo.org/record/6419384/files/figure.png" pageId="39" pageNumber="40">Fig. 8H</figureCitation>
), coloured chequered brown in nymphs, chequered turquois/grey in adults (
<figureCitation id="81A0A593FFD9FF98FDE82C91FD45FE45" box="[608,718,401,428]" captionStart="Fig" captionStartId="36.[189,232,1542,1568]" captionTargetBox="[189,1398,265,1482]" captionTargetId="figure-17@36.[189,1398,265,1482]" captionTargetPageId="36" captionText="Fig. 15. Live Pharmacus Pictet &amp; de Saussure, 1893 in their natural environments. AB. Pharmacus senex sp. nov. A. Adult ♂, Dunstan, Dunstan Mountains, 1670 m a.s.l. B. Adult ♂ and ♀. The Obelisk, Old Man Range, 1680 m a.s.l. C. Pharmacus notabilis sp. nov. Adult ♂, Remarkables Ski-field Access Road, Wakatipu, 1200 m a.s.l. D. Pharmacus concinnus sp. nov. Adult ♂, Symmetry Peaks, Eyre Mountains, 1500 m a.s.l. E. Pharmacus perfidus sp. nov. Adult ♀ and ♂, Spence Peak, Takitimu Mountains, 1450 m a.s.l. F. Pharmacus vallestris sp. nov. Adult ♀, French Ridge, Matukituki River West Branch, 1700 m a.s.l. (MPN CW5145)." figureDoi="http://doi.org/10.5281/zenodo.6419398" httpUri="https://zenodo.org/record/6419398/files/figure.png" pageId="39" pageNumber="40">Fig. 15C</figureCitation>
). A thin, pale median line along the whole length of the insect is generally present.
</paragraph>
<paragraph id="1924B916FFD9FF98FF352CF6FDF1FDDA" blockId="39.[187,1398,501,563]" pageId="39" pageNumber="40">
MALE TERMINALIA. Subgenital plate unmistakeable, elongated, the vertex of the spoon acuminate, shallow when seen from the side (
<figureCitation id="81A0A593FFD9FF98FE602F19FDE5FDDA" box="[488,622,537,563]" captionStart="Fig" captionStartId="26.[189,232,1782,1808]" captionTargetBox="[226,1375,266,1741]" captionTargetId="figure-18@26.[202,1384,263,1741]" captionTargetPageId="26" captionText="Fig. 10. Adult male terminalia of cave wētā in the genus Pharmacus Pictet &amp; de Saussure, 1893. Left column: dorsal view; central column: ventral view (subgenital plate); right column: lateral view. AC. Pharmacus montanus Pictet &amp; de Saussure, 1893, Mueller Hut Track, Sealy Range, Mt Cook (MPN CW3532). DF. Pharmacus cochleatus (Karny, 1935) comb. nov. DE. Humboldt Mountains (MPN CW3698). F. Blue Lake Creek, Garvie Mountains (MPN CW4462). GI. Pharmacus cristatus sp. nov., Skippers Range High Point, South Westland (NMNZ AI.052292). JL. Pharmacus notabilis sp. nov., Remarkables Ski-field access road (NMNZ AI.052296). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6419388" httpUri="https://zenodo.org/record/6419388/files/figure.png" pageId="39" pageNumber="40">Fig. 10JL</figureCitation>
).
</paragraph>
<caption id="4DE4E99EFFD9FF98FF3528FFFAFCF80D" ID-DOI="http://doi.org/10.5281/zenodo.6419400" ID-Zenodo-Dep="6419400" httpUri="https://zenodo.org/record/6419400/files/figure.png" pageId="39" pageNumber="40" startId="39.[189,232,1535,1561]" targetBox="[189,1398,687,1496]" targetPageId="39">
<paragraph id="1924B916FFD9FF98FF3528FFFAFCF80D" blockId="39.[187,1399,1535,2020]" pageId="39" pageNumber="40">
<emphasis id="2BEF6504FFD9FF98FF3528FFFE93F9F0" bold="true" box="[189,280,1535,1561]" pageId="39" pageNumber="40">Fig. 16.</emphasis>
Habitat of
<taxonomicName id="DE9BC295FFD9FF98FE1D2B00FCD2F9F3" authority="Pictet &amp; de Saussure, 1893" authorityName="Pictet &amp; de Saussure" authorityYear="1893" box="[405,857,1536,1562]" class="Insecta" family="Rhaphidophoridae" genus="Pharmacus" kingdom="Animalia" order="Orthoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="2BEF6504FFD9FF98FE1D2B00FD90F9F3" box="[405,539,1536,1562]" italics="true" pageId="39" pageNumber="40">Pharmacus</emphasis>
<bibRefCitation id="7D0AC4E7FFD9FF98FDAA2B00FCD2F9F3" author="Pictet A. &amp; de Saussure H." box="[546,857,1536,1562]" pageId="39" pageNumber="40" pagination="293 - 318" refId="ref32748" refString="Pictet A. &amp; de Saussure H. 1893. De quelques orthopteres nouveaux. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 8: 293 - 318." type="journal article" year="1893">Pictet &amp; de Saussure, 1893</bibRefCitation>
</taxonomicName>
.
<emphasis id="2BEF6504FFD9FF98FCE928FFFCFDF9F0" bold="true" box="[865,886,1535,1561]" pageId="39" pageNumber="40">A</emphasis>
. Sealy Range, Mount Cook.
<emphasis id="2BEF6504FFD9FF98FB482B00FB65F9F3" bold="true" box="[1216,1262,1536,1562]" pageId="39" pageNumber="40">ab</emphasis>
. Collection localities of
<taxonomicName id="DE9BC295FFD9FF98FEDE2B23FC38F9D4" authority="Pictet &amp; de Saussure, 1893" authorityName="Pictet &amp; de Saussure" authorityYear="1893" box="[342,947,1571,1597]" class="Insecta" family="Rhaphidophoridae" genus="Pharmacus" kingdom="Animalia" order="Orthoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="2BEF6504FFD9FF98FEDE2B23FDD3F9D4" box="[342,600,1571,1597]" italics="true" pageId="39" pageNumber="40">Pharmacus montanus</emphasis>
<bibRefCitation id="7D0AC4E7FFD9FF98FDEC2B23FC38F9D4" author="Pictet A. &amp; de Saussure H." box="[612,947,1571,1597]" pageId="39" pageNumber="40" pagination="293 - 318" refId="ref32748" refString="Pictet A. &amp; de Saussure H. 1893. De quelques orthopteres nouveaux. Mitteilungen der Schweizerischen Entomologischen Gesellschaft 8: 293 - 318." type="journal article" year="1893">Pictet &amp; de Saussure, 1893</bibRefCitation>
</taxonomicName>
.
<emphasis id="2BEF6504FFD9FF98FC4A2B23FC5AF9D4" bold="true" box="[962,977,1571,1597]" pageId="39" pageNumber="40">a</emphasis>
. Mueller Hut Track, 1500 m a.s.l. (MPN CW3299, CW3300, CW35323535).
<emphasis id="2BEF6504FFD9FF98FD4E2B46FD5DF989" bold="true" box="[710,726,1606,1632]" pageId="39" pageNumber="40">b</emphasis>
. Mount Annette, 2200 m a.s.l. (MPN CW3302, CW3303, CW3323).
<emphasis id="2BEF6504FFD9FF98FEB62B69FEC0F96A" bold="true" box="[318,331,1641,1667]" pageId="39" pageNumber="40">c</emphasis>
. Mount Edgar Thomson, Ben Ohau Range, 2350 m a.s.l., collection locality of
<taxonomicName id="DE9BC295FFD9FF98FB542B6AFE6CF94E" authority="cochleatus (Karny, 1935)" authorityName="cochleatus (Karny" authorityYear="1935" baseAuthorityName="cochleatus (Karny" baseAuthorityYear="1935" class="Insecta" family="Rhaphidophoridae" genus="Pharmacus" kingdom="Animalia" order="Orthoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="subSpecies" species="cochleatus" status="comb. nov." subSpecies="cochleatus">
<emphasis id="2BEF6504FFD9FF98FB542B6AFEB1F94F" italics="true" pageId="39" pageNumber="40">P. cochleatus cochleatus</emphasis>
(
<bibRefCitation id="7D0AC4E7FFD9FF98FEC32B8DFE54F94E" author="Karny H." box="[331,479,1676,1703]" pageId="39" pageNumber="40" pagination="383 - 385" refId="ref31566" refString="Karny H. 1935. Die Gryllacrididen des Pariser Museum und der Collection L. Chopard. Eos 10 (3 - 4): 383 - 385." type="journal article" year="1935">Karny, 1935</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="30DCD87FFFD9FF98FE782B8CFDF9F94E" box="[496,626,1676,1703]" pageId="39" pageNumber="40" rank="species">comb. nov.</taxonomicNameLabel>
(MPN CW3278).
<emphasis id="2BEF6504FFD9FF98FCD82B8DFCE8F94E" bold="true" box="[848,867,1677,1703]" pageId="39" pageNumber="40">B</emphasis>
. Lochnagar, Richardson Mountains.
<emphasis id="2BEF6504FFD9FF98FA922B8DFAA2F94E" bold="true" box="[1306,1321,1677,1703]" pageId="39" pageNumber="40">d</emphasis>
. Rock bluffs at 2080 m a.s.l., collection locality of
<taxonomicName id="DE9BC295FFD9FF98FD452BB0FB11F923" authority="cochleatus (Karny, 1935)" authorityName="cochleatus (Karny" authorityYear="1935" baseAuthorityName="cochleatus (Karny" baseAuthorityYear="1935" box="[717,1178,1711,1738]" class="Insecta" family="Rhaphidophoridae" genus="Pharmacus" kingdom="Animalia" order="Orthoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="subSpecies" species="cochleatus" status="comb. nov." subSpecies="cochleatus">
<emphasis id="2BEF6504FFD9FF98FD452BB0FC67F923" box="[717,1004,1712,1738]" italics="true" pageId="39" pageNumber="40">P. cochleatus cochleatus</emphasis>
(
<bibRefCitation id="7D0AC4E7FFD9FF98FC752BB0FB19F923" author="Karny H." box="[1021,1170,1711,1738]" pageId="39" pageNumber="40" pagination="383 - 385" refId="ref31566" refString="Karny H. 1935. Die Gryllacrididen des Pariser Museum und der Collection L. Chopard. Eos 10 (3 - 4): 383 - 385." type="journal article" year="1935">Karny, 1935</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="30DCD87FFFD9FF98FB2B2BB0FAAEF923" box="[1187,1317,1712,1738]" pageId="39" pageNumber="40" rank="species">comb. nov.</taxonomicNameLabel>
(MPN CW4590).
<emphasis id="2BEF6504FFD9FF98FEB72BD3FEDFF904" bold="true" box="[319,340,1747,1773]" pageId="39" pageNumber="40">C</emphasis>
. Matukituki River West Branch.
<emphasis id="2BEF6504FFD9FF98FD512BD3FD75F904" bold="true" box="[729,766,1747,1773]" pageId="39" pageNumber="40">ef</emphasis>
. Collection localities of
<taxonomicName id="DE9BC295FFD9FF98FB972BD4FB23F904" authority="Hegg &amp; Morgan-Richards &amp; Trewick, 2022" authorityName="Hegg &amp; Morgan-Richards &amp; Trewick" authorityYear="2022" box="[1055,1192,1747,1773]" class="Insecta" family="Rhaphidophoridae" genus="Pharmacus" kingdom="Animalia" order="Orthoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="vallestris" status="sp. nov.">
<emphasis id="2BEF6504FFD9FF98FB972BD4FB23F904" box="[1055,1192,1747,1773]" italics="true" pageId="39" pageNumber="40">P. vallestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30DCD87FFFD9FF98FB272BD3FA82F904" box="[1199,1289,1747,1773]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="2BEF6504FFD9FF98FA982BD3FA96F904" bold="true" box="[1296,1309,1747,1773]" pageId="39" pageNumber="40">e</emphasis>
. Valley floor, dry creek bed at 1000 m a.s.l. (MPN CW3430, CW3700).
<emphasis id="2BEF6504FFD9FF98FC482BF6FC41F8F9" bold="true" box="[960,970,1782,1808]" pageId="39" pageNumber="40">f</emphasis>
. French Ridge, 1650 m a.s.l. (MPN CW5145, CW5146, CW52135215).
<emphasis id="2BEF6504FFD9FF98FD002A19FD3DF8DA" bold="true" box="[648,694,1817,1843]" pageId="39" pageNumber="40">gh</emphasis>
. Collection localities of
<taxonomicName id="DE9BC295FFD9FF98FC622A1AFE8AF8BE" authority="cochleatus (Karny, 1935)" authorityName="cochleatus (Karny" authorityYear="1935" baseAuthorityName="cochleatus (Karny" baseAuthorityYear="1935" class="Insecta" family="Rhaphidophoridae" genus="Pharmacus" kingdom="Animalia" order="Orthoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="subSpecies" species="cochleatus" status="comb. nov." subSpecies="cochleatus">
<emphasis id="2BEF6504FFD9FF98FC622A1AFA9AF8DA" box="[1002,1297,1817,1843]" italics="true" pageId="39" pageNumber="40">P. cochleatus cochleatus</emphasis>
(
<bibRefCitation id="7D0AC4E7FFD9FF98FAAE2A19FF72F8BE" author="Karny H." pageId="39" pageNumber="40" pagination="383 - 385" refId="ref31566" refString="Karny H. 1935. Die Gryllacrididen des Pariser Museum und der Collection L. Chopard. Eos 10 (3 - 4): 383 - 385." type="journal article" year="1935">Karny, 1935</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="30DCD87FFFD9FF98FE842A3CFE1BF8BE" box="[268,400,1852,1879]" pageId="39" pageNumber="40" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="2BEF6504FFD9FF98FE122A3DFE22F8BE" bold="true" box="[410,425,1853,1879]" pageId="39" pageNumber="40">g</emphasis>
. French Ridge, 1800 m a.s.l. (MPN CW5207, CW5208).
<emphasis id="2BEF6504FFD9FF98FBE02A3DFBF3F8BE" bold="true" box="[1128,1144,1853,1879]" pageId="39" pageNumber="40">h</emphasis>
. Bevan Col, 1850 m a.s.l. (OMNZ IV7927 and MPN CW5178 to 5180).
<emphasis id="2BEF6504FFD9FF98FCAE2A60FCB0F893" bold="true" box="[806,827,1888,1914]" pageId="39" pageNumber="40">D</emphasis>
. The Remarkables, Queenstown.
<emphasis id="2BEF6504FFD9FF98FB442A5FFB66F890" bold="true" box="[1228,1261,1887,1913]" pageId="39" pageNumber="40">ij</emphasis>
. Collection localities of
<taxonomicName id="DE9BC295FFD9FF98FEC72A83FDC8F874" authority="Hegg &amp; Morgan-Richards &amp; Trewick, 2022" authorityName="Hegg &amp; Morgan-Richards &amp; Trewick" authorityYear="2022" box="[335,579,1923,1949]" class="Insecta" family="Rhaphidophoridae" genus="Pharmacus" kingdom="Animalia" order="Orthoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="notabilis" status="sp. nov.">
<emphasis id="2BEF6504FFD9FF98FEC72A83FDC8F874" box="[335,579,1923,1949]" italics="true" pageId="39" pageNumber="40">Pharmacus notabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30DCD87FFFD9FF98FDC22A83FD2EF874" box="[586,677,1923,1949]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="2BEF6504FFD9FF98FD242A83FD3FF874" bold="true" box="[684,692,1923,1949]" pageId="39" pageNumber="40">i</emphasis>
. Remarkables Ski-field Access Road, 1220 m a.s.l. (NMNZ AI.052296).
<emphasis id="2BEF6504FFD9FF98FED02AA6FEE9F829" bold="true" box="[344,354,1958,1984]" pageId="39" pageNumber="40">j</emphasis>
. The Remarkables North ridge, 1600 m a.s.l. (MPN CW2588 2594).
<emphasis id="2BEF6504FFD9FF98FB5D2AA6FB6EF829" bold="true" box="[1237,1253,1958,1984]" pageId="39" pageNumber="40">k</emphasis>
. Lake Alta, 1820 m a.s.l., collection locality of
<taxonomicName id="DE9BC295FFD9FF98FDD42AC9FC49F80A" authority="Hegg &amp; Morgan-Richards &amp; Trewick, 2022" authorityName="Hegg &amp; Morgan-Richards &amp; Trewick" authorityYear="2022" box="[604,962,1993,2019]" class="Insecta" family="Rhaphidophoridae" genus="Pharmacus" kingdom="Animalia" order="Orthoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="subSpecies" species="cochleatus" status="subsp. nov." subSpecies="rawhiti">
<emphasis id="2BEF6504FFD9FF98FDD42AC9FC49F80A" box="[604,962,1993,2019]" italics="true" pageId="39" pageNumber="40">Pharmacus cochleatus rawhiti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="30DCD87FFFD9FF98FC412AC9FBC7F80D" box="[969,1100,1993,2020]" pageId="39" pageNumber="40" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
(MPN CW5219 5221).
</paragraph>
</caption>
<paragraph id="1924B916FFD6FF97FF352C09FA80FE80" blockId="40.[189,1399,265,362]" pageId="40" pageNumber="41">
FEMALE TERMINALIA. Subgenital plate small, narrow at the base and widening at the vertex, bilobed, the two rounded, narrow lobes very hairy (
<figureCitation id="81A0A593FFD6FF97FD072C2CFD7FFEAF" box="[655,756,300,326]" captionStart="Fig" captionStartId="31.[189,232,1776,1802]" captionTargetBox="[189,1378,569,1736]" captionTargetId="figure-110@31.[189,1398,545,1736]" captionTargetPageId="31" captionText="Fig. 12. Adult female terminalia of cave wētā in the genus Pharmacus Pictet &amp; de Saussure, 1893. Left column: subgenital plate; central and right columns: ovipositor. AC. Pharmacus montanus Pictet &amp; de Saussure, 1893. A. Mt Annette, Sealy Range, Mt Cook (MPN CW3303). BC. Mt Wakefield, Mount Cook Range (MPN CW3362). DF. Pharmacus cochleatus (Karny, 1935) comb. nov., Lochnagar, Richardson Mountains (MPN CW4590). GI. Pharmacus cristatus sp. nov., Skippers Range High Point, South Westland (NMNZ AI.052293). JL. Pharmacus notabilis sp. nov., Two Mile Hut, Hector Mountains (NMNZ AI.052297). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6419392" httpUri="https://zenodo.org/record/6419392/files/figure.png" pageId="40" pageNumber="41">Fig. 12J</figureCitation>
). Ovipositor relatively short, on average 60% of body length, gently curving upwards; lower valve with 5 to 7 teeth below at the apex (
<figureCitation id="81A0A593FFD6FF97FBF82C4FFB74FE80" box="[1136,1279,335,361]" captionStart="Fig" captionStartId="31.[189,232,1776,1802]" captionTargetBox="[189,1378,569,1736]" captionTargetId="figure-110@31.[189,1398,545,1736]" captionTargetPageId="31" captionText="Fig. 12. Adult female terminalia of cave wētā in the genus Pharmacus Pictet &amp; de Saussure, 1893. Left column: subgenital plate; central and right columns: ovipositor. AC. Pharmacus montanus Pictet &amp; de Saussure, 1893. A. Mt Annette, Sealy Range, Mt Cook (MPN CW3303). BC. Mt Wakefield, Mount Cook Range (MPN CW3362). DF. Pharmacus cochleatus (Karny, 1935) comb. nov., Lochnagar, Richardson Mountains (MPN CW4590). GI. Pharmacus cristatus sp. nov., Skippers Range High Point, South Westland (NMNZ AI.052293). JL. Pharmacus notabilis sp. nov., Two Mile Hut, Hector Mountains (NMNZ AI.052297). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6419392" httpUri="https://zenodo.org/record/6419392/files/figure.png" pageId="40" pageNumber="41">Fig. 12KL</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>