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<mods:title id="8B10C8C852390BF1742648136FB17B45">Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)</mods:title>
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<mods:namePart id="5DB2DCD93FBFC075E4D367B6C7B80D68">Humberto Y. Yamaguti</mods:namePart>
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<taxonomicName id="C0CC27C2FF9DB10338A8CCC8B1436E89" ID-CoL="6PYV" authority="Mello-Leitao, 1934" authorityName="Mello-Leitao" authorityYear="1934" box="[200,543,228,252]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="87" phylum="Arthropoda" rank="genus">
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<bibRefCitation id="635D21B0FF9DB1033932CCC8B1436E89" author="Mello-Leitao" box="[338,543,228,252]" firstAuthor="Mello-Leitao" journalOrPublisher="Octava Reunion de la Sociedad Argentina, Santiago del Estero" pageId="87" pageNumber="87" pagination="1 - 97" part="1933" refId="ref69252" refString="Mello-Leitao, C. de. 1934 a. (&quot; 1933 &quot;). Estudo monografico dos escorpioes da Republica Argentina. Octava Reunion de la Sociedad Argentina, Santiago del Estero 1933: 1 - 97." title="Estudo monografico dos escorpioes da Republica Argentina" type="journal article" year="1934">Mello-Leitão, 1934</bibRefCitation>
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<figureCitation id="9FF740C4FF9DB1033815CD3AB3B66F5B" box="[117,234,278,302]" captionStart="FIG" captionStartId="16.[107,147,1517,1538]" captionTargetBox="[188,1135,224,1487]" captionTargetId="figure@16.[142,1179,224,1487]" captionTargetPageId="16" captionText="FIG. 1. Representative species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971, habitus in life. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂, Isla Mona, Puerto Rico. B. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂, Guanambí, Bahía, Brazil. C. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀, Parque Zoobotânico de Teresina, Piauí, Brazil. D. Jaguajir rochae (Borelli, 1910), comb. nov., ♂, Jeremoabo, Bahía, Brazil. E. Physoctonus debilis, ♀, Guanambí, Bahía, Brazil. F. Rhopalurus laticauda Thorell, 1876., ♀, Alter do Chão, Santarem, Pará, Brazil. G. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀, Lapa do Bode, Itaeté, Bahía, Brazil. H. Troglorhopalurus translucidus Lourenço et al., 2004, ♀, Gruta Rio dos Pombos, Igatu, Andaraí, Bahía, Brazil." httpUri="https://zenodo.org/record/851384/files/figure.png" pageId="87" pageNumber="87">
<figureCitation id="9FF740C4FF9DB1033815CD3AB38A6F5B" box="[117,214,278,302]" captionStart="FIG. 1" captionStartId="16.[107,147,1517,1538]" captionTargetBox="[188,1135,224,1487]" captionTargetId="figure@16.[142,1179,224,1487]" captionTargetPageId="16" captionText="FIG. 1. Representative species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971, habitus in life. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂, Isla Mona, Puerto Rico. B. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂, Guanambí, Bahía, Brazil. C. Jaguajir agamemnon (C. L. Koch, 1839), comb. nov., ♀, Parque Zoobotânico de Teresina, Piauí, Brazil. D. Jaguajir rochae (Borelli, 1910), comb. nov., ♂, Jeremoabo, Bahía, Brazil. E. Physoctonus debilis, ♀, Guanambí, Bahía, Brazil. F. Rhopalurus laticauda Thorell, 1876., ♀, Alter do Chão, Santarem, Pará, Brazil. G. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀, Lapa do Bode, Itaeté, Bahía, Brazil. H. Troglorhopalurus translucidus Lourenço et al., 2004, ♀, Gruta Rio dos Pombos, Igatu, Andaraí, Bahía, Brazil." httpUri="https://zenodo.org/record/851384/files/figure.png" pageId="87" pageNumber="87">Figures 1</figureCitation>
E
</figureCitation>
, 
<figureCitation id="9FF740C4FF9DB1033896CD3AB2446F5B" box="[246,280,278,302]" captionStart="FIG" captionStartId="17.[111,153,1486,1507]" captionTargetBox="[155,1166,224,1455]" captionTargetId="figure@17.[111,1214,224,1455]" captionTargetPageId="17" captionText="FIG. 2. Representative habitats of species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971. A. Isla Mona, Puerto Rico, habitat of Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov. B. Sierra de Cubitas, Camagüey, Cuba, habitat of Heteroctenus junceus (Herbst, 1800), comb. nov. C. Guanambí, Bahía, Brazil, habitat of Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., and Jaguajir rochae (Borelli, 1910), comb. nov. D. Exu, Pernambuco, Brazil, habitat of J. rochae and Physoctonus debilis (C.L. Koch, 1840). E. Santarém, Pará, Brazil, habitat of Rhopalurus laticauda Thorell, 1876. F. Normandia, Roraima, Brazil, habitat of R. laticauda. G. Lapa do Bode, Itaeté, Bahía, Brazil, habitat of Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov. H. Gruta do Lapão, Lençóis, Bahía, Brazil, habitat of Troglorhopalurus translucidus Lourenço et al., 2004." httpUri="https://zenodo.org/record/851386/files/figure.png" pageId="87" pageNumber="87">
<figureCitation id="9FF740C4FF9DB1033896CD3AB2586F5B" box="[246,260,278,302]" captionStart="FIG. 2" captionStartId="17.[111,153,1486,1507]" captionTargetBox="[155,1166,224,1455]" captionTargetId="figure@17.[111,1214,224,1455]" captionTargetPageId="17" captionText="FIG. 2. Representative habitats of species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971. A. Isla Mona, Puerto Rico, habitat of Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov. B. Sierra de Cubitas, Camagüey, Cuba, habitat of Heteroctenus junceus (Herbst, 1800), comb. nov. C. Guanambí, Bahía, Brazil, habitat of Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., and Jaguajir rochae (Borelli, 1910), comb. nov. D. Exu, Pernambuco, Brazil, habitat of J. rochae and Physoctonus debilis (C. L. Koch, 1840). E. Santarém, Pará, Brazil, habitat of Rhopalurus laticauda Thorell, 1876. F. Normandia, Roraima, Brazil, habitat of R. laticauda. G. Lapa do Bode, Itaeté, Bahía, Brazil, habitat of Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov. H. Gruta do Lapão, Lençóis, Bahía, Brazil, habitat of Troglorhopalurus translucidus Lourenço et al., 2004." httpUri="https://zenodo.org/record/851386/files/figure.png" pageId="87" pageNumber="87">2</figureCitation>
D
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, 
<figureCitation id="9FF740C4FF9DB1033944CD3AB26F6F5B" box="[292,307,278,302]" captionStart="FIG" captionStartId="24.[107,149,1416,1437]" captionTargetBox="[138,1207,234,1374]" captionTargetId="figure@24.[107,1211,224,1383]" captionTargetPageId="24" captionText="FIG. 9. Map of northeastern South America, plotting known locality records of six species of Rhopalurusinae Bücherl, 1971: A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov. (star), Ischnotelson peruassu, sp. nov. (triangle), Physoctonus striatus, sp. nov. (circle), Troglorhopalurus lacrau (Lourenço and Pinto-da- Rocha, 1997), comb. nov (diamonds). B. Physoctonus debilis (C.L. Koch, 1840) (squares), Troglorhopalurus translucidus Lourenço et al., 2004 (crosses)." httpUri="https://zenodo.org/record/851400/files/figure.png" pageId="87" pageNumber="87">9</figureCitation>
, 
<figureCitation id="9FF740C4FF9DB103395ECD3AB2316F5B" box="[318,365,278,302]" captionStart="FIG" captionStartId="26.[109,151,1396,1417]" captionTargetBox="[109,1213,224,1368]" captionTargetId="figure@26.[109,1213,224,1368]" captionTargetPageId="26" captionText="FIG. 10. Rhopalurusinae Bücherl, 1971, sternites III (A–C, E, F), V and VI (D), ventral aspect. A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). B. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀ (MZSP). C, D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). E. Physoctonus debilis (C.L. Koch, 1840), ♂ (MZSP). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 1 mm (A, E, F), 2 mm (B, C, D)." httpUri="https://zenodo.org/record/851402/files/figure.png" pageId="87" pageNumber="87">10D</figureCitation>
, 
<figureCitation id="9FF740C4FF9DB1033919CD3AB2F46F5B" box="[377,424,278,302]" captionStart="FIG" captionStartId="28.[108,150,1167,1188]" captionTargetBox="[108,1213,224,1136]" captionTargetId="figure@28.[108,1213,224,1136]" captionTargetPageId="28" captionText="FIG. 11. Rhopalurusinae Bücherl, 1971, dextral pecten, ventral aspect (A, C, E, G) and pectinal teeth, dorsal aspect (B, D, F). A. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♀ (AMNH). C. Rhopalurus laticauda Thorell, 1876, ♂ (MZSP) D. Jaguajir rochae (Borelli, 1910), comb. nov., (AMNH). E. Rhopalurus laticauda Thorell, 1876, ♂ (AMCC [LP 2845]). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha 1997), comb. nov., ♀ (AMCC [LP 3260]). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (MZSP)." httpUri="https://zenodo.org/record/851404/files/figure.png" pageId="87" pageNumber="87">11G</figureCitation>
, 
<figureCitation id="9FF740C4FF9DB10339D4CD3AB2806F5B" box="[436,476,278,302]" captionStart="FIG" captionStartId="29.[107,149,984,1005]" captionTargetBox="[107,1211,224,956]" captionTargetId="figure@29.[107,1211,224,956]" captionTargetPageId="29" captionText="FIG. 12. Rhopalurusinae Bücherl, 1971, pectinal teeth, dorsal aspect, fine structure. A. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., (MZSP). B. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). C. Rhopalurus laticauda Thorell, 1876, (MZSP). D. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha 1997), comb. nov., ♀ (MZSP). E. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). F. Physoctonus debilis (C.L. Koch, 1840), ♀ (MZSP)." httpUri="https://zenodo.org/record/851406/files/figure.png" pageId="87" pageNumber="87">12F</figureCitation>
, 
<figureCitation id="9FF740C4FF9DB1033987CD3AB1686F5B" box="[487,564,278,302]" captionStart="FIG" captionStartId="34.[109,151,1628,1649]" captionTargetBox="[208,1116,224,1599]" captionTargetId="figure@34.[109,1212,224,1599]" captionTargetPageId="34" captionText="FIG. 16. Rhopalurusinae Bücherl, 1971, carapace, dorsal aspect. A. Physoctonus debilis (C.L. Koch, 1840), ♀ (MZSP). B. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). C. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). D. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. E. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Guatire, Venezuela. F. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). Scale bars = 1 mm (A, B), 2 mm (C–F)." httpUri="https://zenodo.org/record/851414/files/figure.png" pageId="87" pageNumber="87">16A, B</figureCitation>
, 
<figureCitation id="9FF740C4FF9DB1033A5FCD3AB1326F5B" box="[575,622,278,302]" captionStart="FIG" captionStartId="36.[106,149,1153,1174]" captionTargetBox="[106,634,224,1125]" captionTargetId="figure@36.[106,634,224,1125]" captionTargetPageId="36" captionText="FIG. 17. Rhopalurusinae Bücherl, 1971, pedipalp chela movable finger, dorsal aspect, illustrating dentition (denticle rows). A. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). B. Ischnotelson peruassu, sp. nov., paratype ♀ (MZSP). C. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). D. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). E. Rhopalurus ochoai, sp. nov., paratype ♀ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851416/files/figure.png" pageId="87" pageNumber="87">17D</figureCitation>
, 
<figureCitation id="9FF740C4FF9DB10338A1CD14B2516F3A" box="[193,269,312,336]" captionStart="FIG" captionStartId="40.[108,151,1490,1511]" captionTargetBox="[108,1212,224,1461]" captionTargetId="figure@40.[108,1212,224,1462]" captionTargetPageId="40" captionText="FIG. 20. Rhopalurusinae Bücherl, 1971, sternum, pectines and sternite III, ventral aspect. A. Physoctonus debilis (C.L. Koch, 1840), ♂ (MZSP). B. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). C. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). D. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. E. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Puerto Colombia, Venezuela. F. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851422/files/figure.png" pageId="87" pageNumber="87">20A, B</figureCitation>
, 
<figureCitation id="9FF740C4FF9DB1033978CD14B2146F25" box="[280,328,312,336]" captionStart="FIG" captionStartId="44.[107,149,1352,1373]" captionTargetBox="[108,1212,224,1323]" captionTargetId="figure@44.[108,1212,224,1323]" captionTargetPageId="44" captionText="FIG. 21. Rhopalurusinae Bücherl, 1971, telson, lateral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). C. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). D. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀ (AMNH). E. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♀ (AMNH). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (AMNH). H. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). I. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. J. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). K. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). L. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851424/files/figure.png" pageId="87" pageNumber="87">21G</figureCitation>
, 
<figureCitation id="9FF740C4FF9DB1033934CD14B2DF6F25" box="[340,387,312,336]" captionStart="FIG" captionStartId="45.[108,147,1513,1534]" captionTargetBox="[176,1145,224,1484]" captionTargetId="figure@45.[108,1212,224,1484]" captionTargetPageId="45" captionText="FIG. 22. Rhopalurusinae Bücherl, 1971, telson, ventral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). C. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). D. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀ (AMNH). E. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♀ (AMNH). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (AMNH). H. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). I. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH). J. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). K. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (AMNH). L. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851426/files/figure.png" pageId="87" pageNumber="87">22G</figureCitation>
, 
<figureCitation id="9FF740C4FF9DB10339EFCD14B2856F25" box="[399,473,312,336]" captionStart="FIG" captionStartId="47.[107,149,1521,1542]" captionTargetBox="[107,1211,228,1487]" captionTargetId="figure@47.[107,1211,228,1493]" captionTargetPageId="47" captionText="FIG. 24. Rhopalurusinae Bücherl, 1971, hemispermatophores, dorsal aspect (E), dorsal aspect, detail (B, H, M, P), ectal aspect (F, I, N, S), ectal aspect, detail (D, J, R, T, U), ental aspect (A, G, K, O), and ental aspect, detail (C, L, Q). A–D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). E–H. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). I–M. Rhopalurus laticauda Thorell, 1876, ♂ (MZSP). N–R. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (MZSP). S–U. Physoctonus debilis (C.L. Koch, 1840), ♂ (MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/851430/files/figure.png" pageId="87" pageNumber="87">24S–U</figureCitation>
, 
<figureCitation id="9FF740C4FF9DB1033983CD14B1796F25" box="[483,549,312,336]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStart-4="FIG" captionStartId-0="88.[108,150,1167,1188]" captionStartId-1="90.[107,149,1293,1314]" captionStartId-2="91.[107,149,1276,1297]" captionStartId-3="92.[107,149,1467,1488]" captionStartId-4="96.[107,149,1652,1673]" captionTargetBox-0="[108,1212,224,1137]" captionTargetBox-1="[120,1209,224,1263]" captionTargetBox-2="[115,1204,224,1246]" captionTargetBox-3="[201,1115,224,1429]" captionTargetBox-4="[108,1212,224,1623]" captionTargetId-0="figure@88.[108,1212,224,1137]" captionTargetId-1="figure@90.[120,1209,224,1263]" captionTargetId-2="figure@91.[115,1204,224,1246]" captionTargetId-3="figure@92.[201,1115,224,1429]" captionTargetId-4="figure@96.[108,1212,224,1623]" captionTargetPageId-0="88" captionTargetPageId-1="90" captionTargetPageId-2="91" captionTargetPageId-3="92" captionTargetPageId-4="96" captionText-0="FIG. 48. Physoctonus Mello-Leitão, 1934, pedipalp chela, retrolateral aspect. A. P. debilis (C.L. Koch, 1840), ♂ (MZSP). B. P. debilis (C.L. Koch, 1840), ♀ (MZSP). C. P. striatus, sp. nov., ♂ (MZSP). Scale bars = 1 mm." captionText-1="FIG. 49. Physoctonus debilis (C.L. Koch, 1840), metasoma and telson, dorsal (A–C) and ventral (D–F) aspects. A, C, D, F. 2 ♂ (MZSP). B, E. ♀ (AMNH). Scale bars = 2.5 mm." captionText-2="FIG. 50. Physoctonus debilis (C.L. Koch, 1840), metasoma and telson, lateral aspect. A, C. 2♂ (MZSP). B. ♀ (AMNH). Scale bars = 2.5 mm." captionText-3="FIG. 51. Physoctonus debilis (C.L. Koch, 1840), 2 ♀ (AMNH), habitus, dorsal (A, C) and ventral (B, D) aspects. Scale bars = 1 mm." captionText-4="FIG. 52. Rhopalurus Thorell, 1876, pedipalp chela, retrolateral aspect. A. R. caribensis Teruel and Roncallo, 2008, ♂ (AMNH). B, C. R. laticauda Thorell, 1876. B. ♂ (AMNH), Laguna Canaima, Venezuela. C. ♂ (AMNH), Guatire, Venezuela. D. R. ochoai, sp. nov., holotype ♂ (AMNH). Scale bars = 2 mm." httpUri-0="https://zenodo.org/record/851478/files/figure.png" httpUri-1="https://zenodo.org/record/851480/files/figure.png" httpUri-2="https://zenodo.org/record/851482/files/figure.png" httpUri-3="https://zenodo.org/record/851484/files/figure.png" httpUri-4="https://zenodo.org/record/851486/files/figure.png" pageId="87" pageNumber="87">48–52</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFF9DB103380BCD43B10F6A9F" pageId="87" pageNumber="87" type="reference_group">
<paragraph id="07735C41FF9DB103380BCD43B2C16FB9" blockId="87.[107,635,366,1258]" pageId="87" pageNumber="87">
<taxonomicName id="C0CC27C2FF9DB103380BCD43B2E36FF3" authority="C. L. Koch, 1840" authorityName="C. L. Koch" authorityYear="1840" box="[107,447,366,390]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="87" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="35B88053FF9DB103380BCD43B2576FF3" box="[107,267,366,390]" italics="true" pageId="87" pageNumber="87">Vaejovis debilis</emphasis>
C.L. 
<bibRefCitation id="635D21B0FF9DB1033926CD42B2E36FF3" author="Koch" box="[326,447,366,390]" firstAuthor="Koch" journalOrPublisher="Nurnberg: C. H. Zeh'sche Buchhandlung" pageId="87" pageNumber="87" pagination="1 - 114" part="8" refId="ref66739" refString="Koch, C. L. 1840. Die Arachniden. Nurnberg: C. H. Zeh'sche Buchhandlung 8: 1 - 114 (pts. 1, 2, 1840; pts. 2, 3, 1841)." title="Die Arachniden" type="journal article" year="1840">Koch, 1840</bibRefCitation>
</taxonomicName>
[= 
<taxonomicName id="C0CC27C2FF9DB1033986CD42B2F46FDC" ID-CoL="4HHR8" authority="(C. L. Koch, 1840)" authorityName="C.L. Koch" authorityYear="1840" baseAuthorityName="C. L. Koch" baseAuthorityYear="1840" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="87" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="35B88053FF9DB1033986CD42B3816FDC" italics="true" pageId="87" pageNumber="87">Physoctonus debilis</emphasis>
(C.L. 
<bibRefCitation id="635D21B0FF9DB1033941CDBDB2FC6FDC" author="Koch" box="[289,416,401,425]" firstAuthor="Koch" journalOrPublisher="Nurnberg: C. H. Zeh'sche Buchhandlung" pageId="87" pageNumber="87" pagination="1 - 114" part="8" refId="ref66739" refString="Koch, C. L. 1840. Die Arachniden. Nurnberg: C. H. Zeh'sche Buchhandlung 8: 1 - 114 (pts. 1, 2, 1840; pts. 2, 3, 1841)." title="Die Arachniden" type="journal article" year="1840">Koch, 1840</bibRefCitation>
)
</taxonomicName>
], type species, by subsequent designation.
</paragraph>
<paragraph id="07735C41FF9DB103380BCDFBB17B6C64" blockId="87.[107,635,366,1258]" pageId="87" pageNumber="87">
<taxonomicName id="C0CC27C2FF9DB103380BCDFBB39E6F9B" ID-CoL="85MN" authorityName="C. L. Koch" authorityYear="1836" box="[107,194,471,494]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF9DB103380BCDFBB39E6F9B" box="[107,194,471,494]" italics="true" pageId="87" pageNumber="87">Vaejovis</emphasis>
</taxonomicName>
(part): C.L. 
<bibRefCitation id="635D21B0FF9DB1033928CDFAB2B56F9A" author="Koch" box="[328,489,470,495]" firstAuthor="Koch" journalOrPublisher="Nurnberg: C. H. Zeh'sche Buchhandlung" pageId="87" pageNumber="87" pagination="1 - 114" part="8" refId="ref66739" refString="Koch, C. L. 1840. Die Arachniden. Nurnberg: C. H. Zeh'sche Buchhandlung 8: 1 - 114 (pts. 1, 2, 1840; pts. 2, 3, 1841)." title="Die Arachniden" type="journal article" year="1840">Koch, 1840: 21</bibRefCitation>
, 22, pl. CCLIX, fig. 605; 
<bibRefCitation id="635D21B0FF9DB1033930CDD5B17F6C64" author="Kraepelin" bookContentInfo="265 pp." box="[336,547,505,529]" editor="F. Dahl" firstAuthor="Kraepelin" journalOrPublisher="Friedlander and Sohn Verlag" pageId="87" pageNumber="87" refId="ref66921" refString="Kraepelin, K. 1899. Scorpiones und Pedipalpi. In F. Dahl (editor), Das Tierreich. Berlin: R. Friedlander and Sohn Verlag. 265 pp." title="Scorpiones und Pedipalpi" type="book" volumeTitle="Das Tierreich. Berlin: R" year="1899">Kraepelin, 1899: 96</bibRefCitation>
.
</paragraph>
<paragraph id="07735C41FF9DB103380BCE31B1506C41" blockId="87.[107,635,366,1258]" box="[107,524,540,564]" pageId="87" pageNumber="87">
<emphasis id="35B88053FF9DB103380BCE31B3956C41" box="[107,201,541,564]" italics="true" pageId="87" pageNumber="87">Waejovis</emphasis>
(lapsus): 
<bibRefCitation id="635D21B0FF9DB1033952CE30B29B6C41" author="Gervais" box="[306,455,540,564]" firstAuthor="Gervais" journalOrPublisher="Archives du Museum d'Histoire Naturelle, Paris" pageId="87" pageNumber="87" pagination="201 - 240" part="4" refId="ref66060" refString="Gervais, P. M. 1844 a. Remarques sur la famille des scorpions et description des plusieurs especes nouvelles de la collection du Museum. Archives du Museum d'Histoire Naturelle, Paris 4: 201 - 240." title="Remarques sur la famille des scorpions et description des plusieurs especes nouvelles de la collection du Museum" type="journal article" year="1844">Gervais, 1844</bibRefCitation>
b: 458.
</paragraph>
<paragraph id="07735C41FF9DB103380BCE13B2726D82" blockId="87.[107,635,366,1258]" pageId="87" pageNumber="87">
<taxonomicName id="C0CC27C2FF9DB103380BCE13B3B86C22" ID-CoL="63QFB" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" box="[107,228,575,599]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF9DB103380BCE13B3B86C22" box="[107,228,575,599]" italics="true" pageId="87" pageNumber="87">Rhopalurus</emphasis>
</taxonomicName>
(part): 
<bibRefCitation id="635D21B0FF9DB1033956CE13B2A86C23" author="Borelli" box="[310,500,574,599]" firstAuthor="Borelli" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia Comparata della Realle Universita di Torino" pageId="87" pageNumber="87" pagination="1 - 8" part="25" refId="ref64441" refString="Borelli, A. 1910. Scorpioni nuovi e poco noti del Brasile. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Realle Universita di Torino 25 (629): 1 - 8." title="Scorpioni nuovi e poco noti del Brasile" type="journal article" year="1910">Borelli, 1910: 5–8</bibRefCitation>
, fig. 1; 
<bibRefCitation id="635D21B0FF9DB10338FBCE4DB2216C0C" author="Mello-Campos" box="[155,381,609,633]" firstAuthor="Mello-Campos" journalOrPublisher="Memorias do Instituto Oswaldo Cruz" pageId="87" pageNumber="87" pagination="237 - 369" part="17" refId="ref69164" refString="Mello-Campos, O. de. 1924 a. Os escorpioes brasileiros. Memorias do Instituto Oswaldo Cruz 17 (2): 237 - 369." title="Os escorpioes brasileiros" type="journal article" year="1924">Mello-Campos, 1924</bibRefCitation>
a: 252, 275–277; Mello- Leitão, 1932: 14, 30; 
<bibRefCitation id="635D21B0FF9DB1033918CEA8B17C6CE9" author="Meise" box="[376,544,644,668]" firstAuthor="Meise" journalOrPublisher="Nyt Magazin for Naturvidenskaberne" pageId="87" pageNumber="87" pagination="25 - 43" part="72" refId="ref69146" refString="Meise, W. 1934. Scorpiones. Nyt Magazin for Naturvidenskaberne 72: 25 - 43." title="Scorpiones" type="journal article" year="1934">Meise, 1934: 42</bibRefCitation>
; 
<bibRefCitation id="635D21B0FF9DB1033A4BCEA8B3AB6CCA" author="Prado" firstAuthor="Prado" journalOrPublisher="Sinopse das especies de Rhopalurus. Memorias do Instituto Butantan" pageId="87" pageNumber="87" pagination="25 - 36" part="13" refId="ref70234" refString="Prado, A. 1940. Contribuicao ao conhecimento dos escorpioes sul-americanos. Sinopse das especies de Rhopalurus. Memorias do Instituto Butantan 13: 25 - 36." title="Contribuicao ao conhecimento dos escorpioes sul-americanos" type="journal article" year="1940">Prado, 1940: 26</bibRefCitation>
, 29, 30; 
<bibRefCitation id="635D21B0FF9DB1033930CE8BB1136CCA" author="Mello-Leitao" box="[336,591,679,703]" firstAuthor="Mello-Leitao" journalOrPublisher="Arquivos do Museu Nacional" pageId="87" pageNumber="87" pagination="1 - 468" part="40" refId="ref69384" refString="Mello-Leitao, C. F. de. 1945. Escorpioes sul americanos. Arquivos do Museu Nacional 40: 1 - 468." title="Escorpioes sul americanos" type="journal article" year="1945">Mello-Leitão, 1945: 266</bibRefCitation>
, 272, 273; 
<bibRefCitation id="635D21B0FF9DB1033962CEE5B2906C94" author="Bucherl" box="[258,460,713,737]" firstAuthor="Bucherl" journalOrPublisher="Catalogo da colecao escorpionica do Instituto Butantan. Memorias do Instituto Butantan" pageId="87" pageNumber="87" pagination="255 - 275" part="29" refId="ref64652" refString="Bucherl, W. 1959. Escorpioes e escorpionismo no Brasil. X. Catalogo da colecao escorpionica do Instituto Butantan. Memorias do Instituto Butantan 29: 255 - 275." title="Escorpioes e escorpionismo no Brasil. X" type="journal article" year="1959">Bücherl, 1959: 268</bibRefCitation>
; 
<bibRefCitation id="635D21B0FF9DB10339B6CEE6B1636C97" author="Bucherl" box="[470,575,714,738]" editor="W. Bucherl &amp; E. R. Buckley" firstAuthor="Bucherl" journalOrPublisher="New York: Academic Press" pageId="87" pageNumber="87" pagination="317 - 348" part="3" refId="ref64808" refString="Bucherl, W. 1971. Classification, biology and venom extraction of scorpions. In W. Bucherl and E. R. Buckley (editors), Venomous animals and their venoms 3: 317 - 348. New York: Academic Press." title="Classification, biology and venom extraction of scorpions" type="journal article" volumeTitle="Venomous animals and their venoms" year="1971">1971: 327</bibRefCitation>
; 
<bibRefCitation id="635D21B0FF9DB10338FBCEC0B26F6D71" author="Francke" box="[155,307,748,772]" firstAuthor="Francke" journalOrPublisher="Journal of Arachnology" pageId="87" pageNumber="87" pagination="125 - 135" part="4" refId="ref65579" refString="Francke, O. F. 1977 a. Two emendations to Stahnke's (1974) Vaejovidae revision (Scorpionida, Vaejovidae). Journal of Arachnology 4 (2): 125 - 135." title="Two emendations to Stahnke's (1974) Vaejovidae revision (Scorpionida, Vaejovidae)" type="journal article" year="1977">Francke, 1977</bibRefCitation>
a: 125, 127–134, figs. 1–15; 
<bibRefCitation id="635D21B0FF9DB10338FBCF23B21F6D52" author="Lourenco" box="[155,323,783,807]" firstAuthor="Lourenco" journalOrPublisher="Revue Arachnologique" pageId="87" pageNumber="87" pagination="107 - 141" part="4" refId="ref67639" refString="Lourenco, W. R. 1982 a. Revision du genre Rhopalurus Thorell, 1876 (Scorpiones, Buthidae). Revue Arachnologique 4: 107 - 141." title="Revision du genre Rhopalurus Thorell, 1876 (Scorpiones, Buthidae)" type="journal article" year="1982">Lourenço, 1982</bibRefCitation>
a: 108, 133, 135–137, fig. 78; 
<bibRefCitation id="635D21B0FF9DB10338FBCF1EB2626D3F" author="Armas" box="[155,318,818,842]" firstAuthor="Armas" journalOrPublisher="Poeyana" pageId="87" pageNumber="87" pagination="1 - 11" part="284" refId="ref63754" refString="Armas, L. F. de. 1984. Tipos de Arachnida depositados en el Instituto de Zoologia de la Academia de Ciencias de Cuba. 1. Amblypygi, Opiliones, Ricinulei, Scorpiones, Schizomida e Uropygi. Poeyana 284: 1 - 11." title="Tipos de Arachnida depositados en el Instituto de Zoologia de la Academia de Ciencias de Cuba. 1. Amblypygi, Opiliones, Ricinulei, Scorpiones, Schizomida e Uropygi" type="journal article" year="1984">Armas, 1984: 8</bibRefCitation>
; 
<bibRefCitation id="635D21B0FF9DB103392ACF1EB2AF6D3F" author="Lourenco" box="[330,499,818,842]" firstAuthor="Lourenco" journalOrPublisher="Memoires de la Societe Royale Belge d'Entomologie" pageId="87" pageNumber="87" pagination="129 - 137" part="33" refId="ref67782" refString="Lourenco, W. R. 1986 a. Biogeographie et phylogenie des scorpions du genre Rhopalurus Thorell, 1876 (Scorpiones, Buthidae). Memoires de la Societe Royale Belge d'Entomologie 33: 129 - 137." title="Biogeographie et phylogenie des scorpions du genre Rhopalurus Thorell, 1876 (Scorpiones, Buthidae)" type="journal article" year="1986">Lourenço, 1986</bibRefCitation>
a: 133, 135, figs. 12, 16; 1986b: 165, fig. 7; 1990: 161; 1992: 55; 
<bibRefCitation id="635D21B0FF9DB1033962CF5BB2916DFA" author="Kovarik" bookContentInfo="175 pp." box="[258,461,887,911]" firstAuthor="Kovarik" journalOrPublisher="Madagaskar: Jihlava" pageId="87" pageNumber="87" refId="ref66846" refString="Kovarik, F. 1998. Stiri [Scorpions]. Madagaskar: Jihlava. 175 pp. [in Czech]" title="Stiri [Scorpions]" type="book" year="1998">Kovařík, 1998: 118</bibRefCitation>
; 
<bibRefCitation id="635D21B0FF9DB10339B9CF54B25F6DC7" author="Lourenco" bookContentInfo="306 pp." firstAuthor="Lourenco" journalOrPublisher="Paris: Les Editions de l'If" pageId="87" pageNumber="87" refId="ref68189" refString="Lourenco, W. R. 2002. Scorpions of Brazil. Paris: Les Editions de l'If. 306 pp." title="Scorpions of Brazil" type="book" year="2002">Lourenço, 2002: 101</bibRefCitation>
, 111, figs. 225, 226; 
<bibRefCitation id="635D21B0FF9DB10339BCCFB6B2756DA0" author="Lira-da-Silva" etAl="et al." firstAuthor="Lira-da-Silva" journalOrPublisher="Biota Neotropica" pageId="87" pageNumber="87" pagination="1 - 3" part="5" refId="ref67409" refString="Lira-da-Silva, R. M., G. M. Jordao, T. F. Silva, D. M. Candido, and T. K. Brazil. 2005. Ocorrencia de Rhopalurus debilis (C. L. Koch, 1840) (Scorpiones, Buthidae) no estado da Bahia, Brasil. Biota Neotropica 5 (1 A): 1 - 3." title="Ocorrencia de Rhopalurus debilis (C. L. Koch, 1840) (Scorpiones, Buthidae) no estado da Bahia, Brasil" type="journal article" year="2005">Lira-da-Silva et al., 2005: 1</bibRefCitation>
, 2; 
<bibRefCitation id="635D21B0FF9DB1033930CF91B2A06DA1" author="Teruel" box="[336,508,956,981]" firstAuthor="Teruel" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="87" pageNumber="87" pagination="43 - 56" part="38" refId="ref71569" refString="Teruel, R. 2006. Apuntes sobre la taxonomia y biogeografia del genero Rhopalurus Thorell 1876 (Scorpiones: Buthidae), con la descripcion de dos nuevas especies de Cuba. Boletin de la Sociedad Entomologica Aragonesa 38: 43 - 56." title="Apuntes sobre la taxonomia y biogeografia del genero Rhopalurus Thorell 1876 (Scorpiones: Buthidae), con la descripcion de dos nuevas especies de Cuba" type="journal article" year="2006">Teruel, 2006: 51</bibRefCitation>
; 
<bibRefCitation id="635D21B0FF9DB1033A68CF91B2766D82" author="Ubinski" etAl="et al." firstAuthor="Ubinski" journalOrPublisher="Cytogenetic and Genome Research" pageId="87" pageNumber="87" pagination="83 - 155" part="148" refId="ref72292" refString="Ubinski, C. V., L. S. Carvalho, and M. C. Schneider. 2016. Chromosome evolution in Brazilian scorpions of the genus Rhopalurus and related genera (Scorpiones: Buthidae). Abstracts of the 21 st International Chromosome Conference, July 10 - 13, Foz do Iguacu, Brazil. Cytogenetic and Genome Research 148: 83 - 155. [doi: 10.1159 / 000446523]" title="Chromosome evolution in Brazilian scorpions of the genus Rhopalurus and related genera (Scorpiones: Buthidae). Abstracts of the 21 st International Chromosome Conference, July 10 - 13, Foz do Iguacu, Brazil" type="journal article" year="2016">Ubinski et al., 2016: 122</bibRefCitation>
.
</paragraph>
<paragraph id="07735C41FF9DB103380BC82EB10F6A9F" blockId="87.[107,635,366,1258]" pageId="87" pageNumber="87">
<taxonomicName id="C0CC27C2FF9DB103380BC82EB2AE6A6F" ID-CoL="6PYV" authority="Mello-Leitao, 1934 b: 76" authorityName="Mello-Leitao" authorityPageNumber="76" authorityYear="1934" box="[107,498,1026,1050]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF9DB103380BC82EB3B76A6F" box="[107,235,1026,1050]" italics="true" pageId="87" pageNumber="87">Physoctonus</emphasis>
<bibRefCitation id="635D21B0FF9DB1033892C82EB2E26A6F" author="Mello-Leitao" box="[242,446,1026,1050]" firstAuthor="Mello-Leitao" journalOrPublisher="Octava Reunion de la Sociedad Argentina, Santiago del Estero" pageId="87" pageNumber="87" pagination="1 - 97" part="1933" refId="ref69252" refString="Mello-Leitao, C. de. 1934 a. (&quot; 1933 &quot;). Estudo monografico dos escorpioes da Republica Argentina. Octava Reunion de la Sociedad Argentina, Santiago del Estero 1933: 1 - 97." title="Estudo monografico dos escorpioes da Republica Argentina" type="journal article" year="1934">Mello-Leitão, 1934</bibRefCitation>
b: 76
</taxonomicName>
, 77, figs. 1–7; 1942: 129; 1945: 129–132, figs. 40, 41; 
<bibRefCitation id="635D21B0FF9DB10338FBC86BB23F6A2A" author="Bucherl" box="[155,355,1095,1119]" editor="H. Lent" firstAuthor="Bucherl" journalOrPublisher="Zoologia" pageId="87" pageNumber="87" pagination="111 - 125" part="5" refId="ref64708" refString="Bucherl, W. 1967. Escorpioes, aranhas e escolopendromorfos da Amazonia. In H. Lent (editor), Atas do Simposio sobre a Biota Amazonica 5 (Zoologia): 111 - 125." title="Escorpioes, aranhas e escolopendromorfos da Amazonia" type="journal article" volumeTitle="Atas do Simposio sobre a Biota Amazonica" year="1967">Bücherl, 1967: 115</bibRefCitation>
; 
<bibRefCitation id="635D21B0FF9DB103390FC86BB2846A2A" author="Bucherl" box="[367,472,1095,1119]" editor="E. J. Fittkau &amp; J. Illies &amp; H. King &amp; G. H. Schwabe &amp; H. Sioli" firstAuthor="Bucherl" journalOrPublisher="2" pageId="87" pageNumber="87" pagination="764 - 793" part="19" refId="ref64746" refString="Bucherl, W. 1969. Giftige Arthropoden. In E. J. Fittkau, J. Illies, H. King, G. H. Schwabe, and H. Sioli (editors), Biogeography and ecology in South America. Monographiae Biologicae 19 (2): 764 - 793. Dordrecht: W. Junk." title="Giftige Arthropoden" type="journal article" volumeTitle="Biogeography and ecology in South America" year="1969">1969: 768</bibRefCitation>
; 
<bibRefCitation id="635D21B0FF9DB1033983C86BB39F6AF7" author="Vachon" firstAuthor="Vachon" journalOrPublisher="Bulletin du Museum National d'Histoire Naturelle, Paris" pageId="87" pageNumber="87" pagination="161 - 166" part="2" refId="ref72387" refString="Vachon, M. 1963. De l'utilite, en systematique, d'une nomenclature des dents de cheliceres chez les Scorpions. Bulletin du Museum National d'Histoire Naturelle, Paris (2) 35 (2): 161 - 166." title="De l'utilite, en systematique, d'une nomenclature des dents de cheliceres chez les Scorpions" type="journal article" year="1963">Vachon, 1963: 161</bibRefCitation>
, 165; 
<bibRefCitation id="635D21B0FF9DB1033962C846B2926AF7" author="Stahnke" box="[258,462,1130,1154]" firstAuthor="Stahnke" journalOrPublisher="Journal of Arachnology" pageId="87" pageNumber="87" pagination="107 - 141" part="1" refId="ref71310" refString="Stahnke, H. L. 1974. Revision and keys to the higher categories of Vejovidae (Scorpionida). Journal of Arachnology 1 (2): 107 - 141." title="Revision and keys to the higher categories of Vejovidae (Scorpionida)" type="journal article" year="1974">Stahnke, 1974: 129</bibRefCitation>
; 
<bibRefCitation id="635D21B0FF9DB10339B9C846B38D6AD0" author="Francke" firstAuthor="Francke" journalOrPublisher="Journal of Arachnology" pageId="87" pageNumber="87" pagination="125 - 135" part="4" refId="ref65579" refString="Francke, O. F. 1977 a. Two emendations to Stahnke's (1974) Vaejovidae revision (Scorpionida, Vaejovidae). Journal of Arachnology 4 (2): 125 - 135." title="Two emendations to Stahnke's (1974) Vaejovidae revision (Scorpionida, Vaejovidae)" type="journal article" year="1977">Francke, 1977</bibRefCitation>
a: 127; 
<bibRefCitation id="635D21B0FF9DB103397BC8A1B1776AD0" author="Lourenco" box="[283,555,1165,1189]" firstAuthor="Lourenco" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="87" pageNumber="87" pagination="359 - 365" part="40" refId="ref68211" refString="Lourenco, W. R. 2007. New considerations on the taxonomic status of the genus Physoctonus Mello-Leitao, 1934 (Scorpiones, Buthidae). Boletin de la Sociedad Entomologica Aragonesa 40: 359 - 365." title="New considerations on the taxonomic status of the genus Physoctonus Mello-Leitao, 1934 (Scorpiones, Buthidae)" type="journal article" year="2007">Lourenço, 2007: 359–364</bibRefCitation>
; 
<bibRefCitation id="635D21B0FF9DB1033A56C8A1B2906ABD" author="Outeda-Jorge" etAl="et al." firstAuthor="Outeda-Jorge" journalOrPublisher="Zoologia" pageId="87" pageNumber="87" pagination="43 - 53" part="26" refId="ref69741" refString="Outeda-Jorge, S., T. Mello, and R. Pinto-da-Rocha. 2009. Litter size, effects of maternal body size, and date of birth in South American scorpions (Arachnida: Scorpiones). Zoologia 26 (1): 43 - 53." title="Litter size, effects of maternal body size, and date of birth in South American scorpions (Arachnida: Scorpiones)" type="journal article" year="2009">Outeda-Jorge et al., 2009: 43–46</bibRefCitation>
; 
<bibRefCitation id="635D21B0FF9DB10339B7C89CB2586A9F" author="Prendini" etAl="et al." firstAuthor="Prendini" journalOrPublisher="Journal of Arachnology" pageId="87" pageNumber="87" pagination="206 - 224" part="37" refId="ref70522" refString="Prendini, L., L. A. Esposito, J. Huff, and E. S. Volschenk. 2009. Redescription of Rhopalurus abudi (Scorpiones, Buthidae), with first description of the male and first record from mainland Hispaniola. Journal of Arachnology 37: 206 - 224." title="Redescription of Rhopalurus abudi (Scorpiones, Buthidae), with first description of the male and first record from mainland Hispaniola" type="journal article" year="2009">Prendini et al., 2009: 222</bibRefCitation>
; 
<bibRefCitation id="635D21B0FF9DB103396FC8FEB1746A9F" author="Brazil and Porto" bookContentInfo="84 pp." box="[271,552,1234,1258]" firstAuthor="Brazil" journalOrPublisher="Salvador, BA, Brazil: EDUFBA" pageId="87" pageNumber="87" refId="ref64569" refString="Brazil, T. K., and Porto, T. J. 2010. Os escorpioes. Salvador, BA, Brazil: EDUFBA. 84 pp." title="Os escorpioes" type="book" year="2010">Brazil and Porto, 2010: 50</bibRefCitation>
, 57.
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFF9DB10338EBC8D3B1AB6CFC" pageId="87" pageNumber="87" type="diagnosis">
<paragraph id="07735C41FF9DB10338EBC8D3B1AB6CFC" blockId="87.[107,637,1279,1741]" lastBlockId="87.[684,1214,228,1740]" pageId="87" pageNumber="87">
DIAGNOSIS: 
<taxonomicName id="C0CC27C2FF9DB1033978C8D3B2C26B62" authorityName="Mello-Leitao" authorityYear="1934" box="[280,414,1279,1303]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF9DB1033978C8D3B2C26B62" box="[280,414,1279,1303]" italics="true" pageId="87" pageNumber="87">Physoctonus</emphasis>
</taxonomicName>
differs from other rhopalurusine genera by the obsolete carapacial carinae, the asetose proximal dorsal fulcra of the pectines, the simple (nonbifurcate) prolateral pedal spur of leg I, and the oblique subrows of primary denticles on the pedipalp chela fingers flanked by small, widely spaced prolateral accessory (supernumerary) denticles and sparse retrolateral accessory denticles. It differs further from 
<taxonomicName id="C0CC27C2FF9DB103380BCA03B2CD6833" authority="Ischnotelson" authorityName=", Ischnotelson, Jaguajir" authorityYear="2017" box="[107,401,1582,1606]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="87" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="35B88053FF9DB103380BCA03B3A56833" box="[107,249,1583,1606]" italics="true" pageId="87" pageNumber="87">Heteroctenus</emphasis>
, 
<emphasis id="35B88053FF9DB103396CCA02B2CD6833" box="[268,401,1582,1606]" italics="true" pageId="87" pageNumber="87">Ischnotelson</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="2E8B3D28FF9DB10339C4CA02B1536830" box="[420,527,1582,1606]" pageId="87" pageNumber="87" rank="genus">gen. nov.</taxonomicNameLabel>
, 
<taxonomicName id="C0CC27C2FF9DB1033A40CA03B12A6833" authority="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; CLOSE Lorenzo Prendini Scorpion Systematics Research Group, 2017" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" box="[544,630,1583,1606]" class="Arachnida" family="Buthidae" genus="Jaguajir" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="87" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="35B88053FF9DB1033A40CA03B12A6833" box="[544,630,1583,1606]" italics="true" pageId="87" pageNumber="87">Jaguajir</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="2E8B3D28FF9DB103380BCA7CB38E6812" box="[107,210,1616,1640]" pageId="87" pageNumber="87" rank="genus">gen. nov.</taxonomicNameLabel>
, 
<taxonomicName id="C0CC27C2FF9DB10338BFCA7CB207681D" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" box="[223,347,1616,1640]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF9DB10338BFCA7CB207681D" box="[223,347,1616,1640]" italics="true" pageId="87" pageNumber="87">Rhopalurus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="C0CC27C2FF9DB10339FFCA7CB105681D" box="[415,601,1616,1640]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF9DB10339FFCA7CB105681D" box="[415,601,1616,1640]" italics="true" pageId="87" pageNumber="87">Troglorhopalurus</emphasis>
</taxonomicName>
by the small size (total length, 20–25 mm); from 
<taxonomicName id="C0CC27C2FF9DB103380BCAB8B2BB68DE" authority=", Ischnotelson, Jaguajir" authorityName=", Ischnotelson, Jaguajir" authorityYear="2017" box="[107,487,1683,1707]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF9DB103380BCAB8B3A968DE" box="[107,245,1684,1707]" italics="true" pageId="87" pageNumber="87">Heteroctenus</emphasis>
, 
<emphasis id="35B88053FF9DB1033964CABFB2D968DE" box="[260,389,1683,1707]" italics="true" pageId="87" pageNumber="87">Ischnotelson</emphasis>
, 
<emphasis id="35B88053FF9DB10339F4CAB8B2BB68DE" box="[404,487,1684,1707]" italics="true" pageId="87" pageNumber="87">Jaguajir</emphasis>
</taxonomicName>
, and 
<taxonomicName id="C0CC27C2FF9DB1033A47CABFB3C768B9" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF9DB1033A47CABFB3C768B9" italics="true" pageId="87" pageNumber="87">Rhopalurus</emphasis>
</taxonomicName>
by the slender pedipalp chela manus of the adult male; from 
<taxonomicName id="C0CC27C2FF9DB1033B06CCC9B7126E89" authority=", Jaguajir" authorityName=", Ischnotelson, Jaguajir" authorityYear="2017" box="[870,1102,229,252]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF9DB1033B06CCC9B0B36E89" box="[870,1007,229,252]" italics="true" pageId="87" pageNumber="87">Heteroctenus</emphasis>
, 
<emphasis id="35B88053FF9DB1033B9BCCC9B7126E89" box="[1019,1102,229,252]" italics="true" pageId="87" pageNumber="87">Jaguajir</emphasis>
</taxonomicName>
, and 
<taxonomicName id="C0CC27C2FF9DB1033CEACCC8B1A16F68" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF9DB1033CEACCC8B1A16F68" italics="true" pageId="87" pageNumber="87">Rhopalurus</emphasis>
</taxonomicName>
by the absence of a pecten-sternite stridulatory organ; from 
<taxonomicName id="C0CC27C2FF9DB1033BEBCD0BB7496F4B" authorityName=", Ischnotelson, Jaguajir" authorityYear="2017" box="[907,1045,295,318]" class="Arachnida" family="Buthidae" genus="Heteroctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF9DB1033BEBCD0BB7496F4B" box="[907,1045,295,318]" italics="true" pageId="87" pageNumber="87">Heteroctenus</emphasis>
</taxonomicName>
by the absence of depressions in the male pectinal plate and the presence of a subaculear tubercle on the telson; from 
<taxonomicName id="C0CC27C2FF9DB1033A8BCDA5B0306FD4" authority="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; CLOSE Lorenzo Prendini Scorpion Systematics Research Group, 2017" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" box="[747,876,393,417]" class="Arachnida" family="Buthidae" genus="Ischnotelson" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="116" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="35B88053FF9DB1033A8BCDA5B0306FD4" box="[747,876,393,417]" italics="true" pageId="87" pageNumber="87">Ischnotelson</emphasis>
</taxonomicName>
by the separate (unfused) lateral ocular and central lateral carinae of the carapace and the telson vesicle not laterally compressed; and from 
<taxonomicName id="C0CC27C2FF9DB1033BD2CDC1B7306C70" box="[946,1132,493,517]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF9DB1033BD2CDC1B7306C70" box="[946,1132,493,517]" italics="true" pageId="87" pageNumber="87">Troglorhopalurus</emphasis>
</taxonomicName>
by the proximal dentate margin of the chela fixed and movable fingers of the adult male emarginate, with a distinct gap evident between them, when closed.
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFF9DB10D3AACCEBEB04B6DE4" lastPageId="89" lastPageNumber="89" pageId="87" pageNumber="87" type="description">
<paragraph id="07735C41FF9DB1033AACCEBEB7FF6C99" blockId="87.[684,1214,228,1740]" pageId="87" pageNumber="87">
DESCRIPTION: The following general description outlines characters common to both species of 
<taxonomicName id="C0CC27C2FF9DB1033AA8CEF8B0146C99" authorityName="Mello-Leitao" authorityYear="1934" box="[712,840,724,748]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="87" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF9DB1033AA8CEF8B0146C99" box="[712,840,724,748]" italics="true" pageId="87" pageNumber="87">Physoctonus</emphasis>
</taxonomicName>
(for measurements, see 
<tableCitation id="4A4E69FAFF9DB1033C2ECEF8B7CA6C99" box="[1102,1174,724,748]" captionStart="TABLE 3" captionStartId="69.[610,690,228,252]" captionTargetBox="[131,1171,439,1728]" captionText="TABLE 3 Meristic data for type material of Ischnotelson peruassu, sp. n., Physoctonus striatus, sp. n., and Physoctonus debilis (C. L. Koch, 1840). Measurements follow Prendini (2000, 2003, 2004). 1 Sum of carapace, tergites I – VII, metasomal segments I – V, and telson; 2 sum of metasomal segments I – V and telson; 3 measured from base of condyle to tip of fixed finger." httpUri="http://table.plazi.org/id/53B30CC9FF8FB1113A02CCC8B0CF6FEA" pageId="87" pageNumber="87" tableUuid="53B30CC9FF8FB1113A02CCC8B0CF6FEA">table 3</tableCitation>
).
</paragraph>
<paragraph id="07735C41FF9DB1033AACCED9B0D96D5B" blockId="87.[684,1214,228,1740]" pageId="87" pageNumber="87">
<emphasis id="35B88053FF9DB1033AACCED9B0116D78" box="[716,845,757,781]" italics="true" pageId="87" pageNumber="87">Total length</emphasis>
: Relatively small scorpions (total length, 20–25 mm).
</paragraph>
<paragraph id="07735C41FF9DB1033AACCF1BB7406BD7" blockId="87.[684,1214,228,1740]" pageId="87" pageNumber="87">
<emphasis id="35B88053FF9DB1033AACCF1BB04D6D3A" box="[716,785,823,847]" italics="true" pageId="87" pageNumber="87">Color:</emphasis>
Base color pale to dark yellow (fig. 1E). Carapace immaculate except interocular surface infuscate, forming dark triangle. Tergites immaculate except for dorsomedian band of infuscation, forming longitudinal stripe on mesosoma. Coxosternal region, pectines, and sternites immaculate, pale to dark yellow. Metasomal segments I–III, dorsal surfaces immaculate, yellow, similar color as carapace and tergites, segments IV and V, dorsal surfaces darker than preceding segments; I–III, ventral surfaces slightly darker than dorsal surfaces, IV and V noticeably darker than I–III, V darker than IV; I–IV each with dark ventromedian band of infuscation. Telson vesicle yellow, similar to metasomal segment V, aculeus almost black. Chelicerae, pedipalps, and legs base color yellow, similar to tergites, with reticulate infuscation; chela fingers dark brown.
</paragraph>
<paragraph id="07735C41FF9DB1033AACC980B7D86B90" blockId="87.[684,1214,228,1740]" pageId="87" pageNumber="87">
<emphasis id="35B88053FF9DB1033AACC980B0676BB1" box="[716,827,1452,1476]" italics="true" pageId="87" pageNumber="87">Chelicerae</emphasis>
: Base, dorsal surface with medial transverse row of well-developed tubercles.
</paragraph>
<paragraph id="07735C41FF9DB10C3AACC9C2B3F76BF4" blockId="87.[684,1214,228,1740]" lastBlockId="88.[107,637,1286,1707]" lastPageId="88" lastPageNumber="88" pageId="87" pageNumber="87">
<emphasis id="35B88053FF9DB1033AACC9C2B06D6870" box="[716,817,1518,1541]" italics="true" pageId="87" pageNumber="87">Carapace</emphasis>
: Median ocular tubercle low (fig. 16A, B); two median ocelli; three pairs of lateral macroocelli; one pair of lateral microocelli. Anteromedian, median ocular, and posteromedian sulci present, forming single, almost continuous, longitudinal sulcus. Carinae obsolete, finely granular, and barely distinguishable from surface granulation; lateral ocular and anterior central submedian carinae separate (unfused); central lateral and posterior central submedian carinae fused.
</paragraph>
<caption id="53B30CC9FF92B10C380CC8A3B7F86AB7" httpUri="https://zenodo.org/record/851478/files/figure.png" pageId="88" pageNumber="88" targetBox="[108,1212,224,1137]" targetPageId="88">
<paragraph id="07735C41FF92B10C380CC8A3B7F86AB7" blockId="88.[108,1212,1163,1219]" pageId="88" pageNumber="88">
FIG. 48. 
<taxonomicName id="C0CC27C2FF92B10C38A3C8A1B1586AD3" authority="Mello-Leitao, 1934" authorityName="Mello-Leitao" authorityYear="1934" box="[195,516,1165,1190]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="88" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF92B10C38A3C8A1B2616AD0" box="[195,317,1165,1189]" italics="true" pageId="88" pageNumber="88">Physoctonus</emphasis>
<bibRefCitation id="635D21B0FF92B10C3923C8A1B1586AD3" author="Mello-Leitao" box="[323,516,1165,1190]" firstAuthor="Mello-Leitao" journalOrPublisher="Octava Reunion de la Sociedad Argentina, Santiago del Estero" pageId="88" pageNumber="88" pagination="1 - 97" part="1933" refId="ref69252" refString="Mello-Leitao, C. de. 1934 a. (&quot; 1933 &quot;). Estudo monografico dos escorpioes da Republica Argentina. Octava Reunion de la Sociedad Argentina, Santiago del Estero 1933: 1 - 97." title="Estudo monografico dos escorpioes da Republica Argentina" type="journal article" year="1934">Mello-Leitão, 1934</bibRefCitation>
</taxonomicName>
, pedipalp chela, retrolateral aspect. 
<emphasis id="35B88053FF92B10C3B0DC8A1B0DA6AD0" bold="true" box="[877,902,1165,1189]" pageId="88" pageNumber="88">A.</emphasis>
<taxonomicName id="C0CC27C2FF92B10C3BECC8A2B7C16AD3" authority="(C. L. Koch, 1840)" authorityName="C.L. Koch" authorityYear="1840" baseAuthorityName="C. L. Koch" baseAuthorityYear="1840" box="[908,1181,1165,1190]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="88" pageNumber="88" phylum="Arthropoda" rank="species" species="debilis">
P. 
<emphasis id="35B88053FF92B10C3BC2C8A1B0BD6AD0" box="[930,993,1165,1189]" italics="true" pageId="88" pageNumber="88">debilis</emphasis>
(C.L. 
<bibRefCitation id="635D21B0FF92B10C3C40C8A1B7CA6AD3" author="Koch" box="[1056,1174,1165,1190]" firstAuthor="Koch" journalOrPublisher="Nurnberg: C. H. Zeh'sche Buchhandlung" pageId="88" pageNumber="88" pagination="1 - 114" part="8" refId="ref66739" refString="Koch, C. L. 1840. Die Arachniden. Nurnberg: C. H. Zeh'sche Buchhandlung 8: 1 - 114 (pts. 1, 2, 1840; pts. 2, 3, 1841)." title="Die Arachniden" type="journal article" year="1840">Koch, 1840</bibRefCitation>
)
</taxonomicName>
, ♂ (MZSP). 
<emphasis id="35B88053FF92B10C38AAC887B3BC6AB6" bold="true" box="[202,224,1195,1219]" pageId="88" pageNumber="88">B.</emphasis>
<taxonomicName id="C0CC27C2FF92B10C3887C880B2A76AB6" authority="(C. L. Koch, 1840)" authorityName="C.L. Koch" authorityYear="1840" baseAuthorityName="C. L. Koch" baseAuthorityYear="1840" box="[231,507,1195,1219]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="88" pageNumber="88" phylum="Arthropoda" rank="species" species="debilis">
P. 
<emphasis id="35B88053FF92B10C389EC887B2616AB6" box="[254,317,1195,1219]" italics="true" pageId="88" pageNumber="88">debilis</emphasis>
(C.L. 
<bibRefCitation id="635D21B0FF92B10C391EC887B2A86AB6" author="Koch" box="[382,500,1195,1219]" firstAuthor="Koch" journalOrPublisher="Nurnberg: C. H. Zeh'sche Buchhandlung" pageId="88" pageNumber="88" pagination="1 - 114" part="8" refId="ref66739" refString="Koch, C. L. 1840. Die Arachniden. Nurnberg: C. H. Zeh'sche Buchhandlung 8: 1 - 114 (pts. 1, 2, 1840; pts. 2, 3, 1841)." title="Die Arachniden" type="journal article" year="1840">Koch, 1840</bibRefCitation>
)
</taxonomicName>
, ♀ (MZSP). 
<emphasis id="35B88053FF92B10C3AE0C887B1C46AB6" bold="true" box="[640,664,1195,1219]" pageId="88" pageNumber="88">C.</emphasis>
<taxonomicName id="C0CC27C2FF92B10C3AFEC880B1A36AB6" authority="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; CLOSE Lorenzo Prendini Scorpion Systematics Research Group, 2017" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" box="[670,767,1196,1219]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="88" pageNumber="88" phylum="Arthropoda" rank="species" species="striatus" status="sp. nov.">
<emphasis id="35B88053FF92B10C3AFEC880B1A36AB6" box="[670,767,1196,1219]" italics="true" pageId="88" pageNumber="88">P. striatus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="2E8B3D28FF92B10C3B6CC887B0076AB7" box="[780,859,1195,1218]" pageId="88" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ (MZSP). Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="07735C41FF92B10D38EBC9A6B2666FF5" blockId="88.[107,637,1286,1707]" lastBlockId="89.[107,637,228,1707]" lastPageId="89" lastPageNumber="89" pageId="88" pageNumber="88">
<emphasis id="35B88053FF92B10C38EBC9A6B3AE6BD7" box="[139,242,1418,1442]" italics="true" pageId="88" pageNumber="88">Pedipalps</emphasis>
: Pedipalp femur retrolateral accessory carinae present. Pedipalp chela manus of adult male slender, proximal dentate margins of fixed and movable fingers slightly curved proximally (fixed finger curved dorsally, movable finger curved ventrally), such that proximal dentate margin emarginate, slight gap present between fingers proximally, when closed (fig. 48A, C), manus of female not incrassate, fixed and movable fingers not curved proximally, such that proximal dentate margin sublinear, little or no gap present between them proximally, when closed (fig. 48B); manus, proventral carina absent, promedian carina present; fixed and movable fingers, median denticle rows each comprising eight oblique subrows of primary denticles flanked by small, widely spaced prolateral accessory (supernumerary) denticles and sparse retrolateral accessory denticles; movable finger without proximal lobe (
<figureCitation id="9FF740C4FF92B10C3C6ACA7CB708681D" box="[1034,1108,1616,1640]" captionStart="FIG. 17" captionStartId="36.[106,149,1153,1174]" captionTargetBox="[106,634,224,1125]" captionTargetId="figure@36.[106,634,224,1125]" captionTargetPageId="36" captionText="FIG. 17. Rhopalurusinae Bücherl, 1971, pedipalp chela movable finger, dorsal aspect, illustrating dentition (denticle rows). A. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). B. Ischnotelson peruassu, sp. nov., paratype ♀ (MZSP). C. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). D. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). E. Rhopalurus ochoai, sp. nov., paratype ♀ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851416/files/figure.png" pageId="88" pageNumber="88">fig. 17</figureCitation>
D). Pedipalps orthobothriotaxic Type A, α configuration; femur with five dorsal trichobothria, trichobothrium 
<emphasis id="35B88053FF93B10D3947CCC8B2696E89" box="[295,309,228,252]" italics="true" pageId="89" pageNumber="89">d</emphasis>
<subScript id="9B485E04FF93B10D3955CCC3B2626F75" attach="left" box="[309,318,239,256]" fontSize="7" pageId="89" pageNumber="89">2</subScript>
situated on prolateral surface; patella trichobothrium 
<emphasis id="35B88053FF93B10D39C6CD29B2E86F68" box="[422,436,261,285]" italics="true" pageId="89" pageNumber="89">d</emphasis>
<subScript id="9B485E04FF93B10D39D4CD3CB2E16F54" attach="left" box="[436,445,272,289]" fontSize="7" pageId="89" pageNumber="89">3</subScript>
situated retrolateral to dorsomedian carina; chela fixed finger trichobothrium 
<emphasis id="35B88053FF93B10D3941CD6BB2606F2A" box="[289,316,327,351]" italics="true" pageId="89" pageNumber="89">db</emphasis>
proximal to or aligned with trichobothrium 
<emphasis id="35B88053FF93B10D3940CD45B26F6FF5" box="[288,307,361,384]" italics="true" pageId="89" pageNumber="89">et</emphasis>
.
</paragraph>
<paragraph id="07735C41FF93B10D38EBCDA6B2B76C71" blockId="89.[107,637,228,1707]" pageId="89" pageNumber="89">
<emphasis id="35B88053FF93B10D38EBCDA6B3E46FD4" box="[139,184,394,417]" italics="true" pageId="89" pageNumber="89">Legs</emphasis>
: Legs III and IV, tibial spurs absent; I and II, basitarsi each with simple prolateral pedal spur; telotarsi each with distinct pro- and retroventral rows of fine, acuminate macrosetae.
</paragraph>
<paragraph id="07735C41FF93B10D38EBCE22B1206D78" blockId="89.[107,637,228,1707]" pageId="89" pageNumber="89">
<emphasis id="35B88053FF93B10D38EBCE22B3BC6C50" box="[139,224,526,549]" italics="true" pageId="89" pageNumber="89">Pectines</emphasis>
: Pectinal plate without depressions (male), anterior margin with sulcus (fig. 20A, B). Pectines not proximally expanded; proximal dorsal fulcra asetose; pectinal teeth almost straight, slightly curved laterally, proximal teeth, dorsal surfaces without striations but covered with small denticles, dorsobasal surfaces without macrosetae; pectinal sensillae short and blunt (figs. 11G, 12F).
</paragraph>
<paragraph id="07735C41FF93B10D38EBCF3BB2F26AEF" blockId="89.[107,637,228,1707]" pageId="89" pageNumber="89">
<emphasis id="35B88053FF93B10D38EBCF3BB3A76D5B" box="[139,251,791,814]" italics="true" pageId="89" pageNumber="89">Mesosoma</emphasis>
: Tergites V–VII slightly wider than than I–IV; I–VI unicarinate, dorsosubmedian carinae absent, dorsomedian carina reduced to posterior half on I–VI, complete on V and VI. Tergite VII pentacarinate, dorsomedian carina complete (fig. 51). Sternites smooth, carinae absent or obsolete; sternite III, lateral margins not forming smooth, raised carina, ventromedian carina not elevated anteriorly, ventrosubmedian surfaces not forming paired depressions, smooth; respiratory spiracles (stigmata) width less than 5× length (fig. 10E).
</paragraph>
<paragraph id="07735C41FF93B10D38EBC888B0896FF5" blockId="89.[107,637,228,1707]" lastBlockId="89.[684,1214,228,1707]" pageId="89" pageNumber="89">
<emphasis id="35B88053FF93B10D38EBC888B3A26ACE" box="[139,254,1188,1211]" italics="true" pageId="89" pageNumber="89">Metasoma</emphasis>
: Metasoma slender, increasing slightly in width posteriorly, segment V only slightly wider than I in adult male, I and V similar width in adult female (figs. 49, 50). Segments I–III each with 10 distinct, granular carinae, IV with eight distinct, granular carinae, V with five distinct, granular carinae; dorsosubmedian carinae obsolete, reduced to rows of granules on dorsal surfaces of segments I–IV, more pronounced on segment I; dorsolateral carinae complete on segments I–IV, and terminating in slightly larger, subspiniform granules posteriorly on II–IV, absent on V; lateral supramedian carinae complete on segments I–V; lateral inframedian carinae complete on segments I–III, complete but obsolete on IV, and absent on V; ventrosubmedian carinae complete on segments I–IV, absent on V; ventromedian carina absent on segments I–IV, complete on V. Intercarinal surfaces finely and densely granular on lateral and ventral surfaces of segments I–V and dorsal surfaces of I–III.
</paragraph>
<paragraph id="07735C41FF93B10D3AACCDA5B72C6C53" blockId="89.[684,1214,228,1707]" pageId="89" pageNumber="89">
<emphasis id="35B88053FF93B10D3AACCDA5B0516FD4" box="[716,781,393,417]" italics="true" pageId="89" pageNumber="89">Telson</emphasis>
: Vesicle oval, not laterally compressed, narrower than metasoma V; anterodorsal lateral lobes prominent; lateral and ventral surfaces granular, pentacarinate with distinct ventromedian carina; subaculear tubercle vestigial.
</paragraph>
<paragraph id="07735C41FF93B10D3AACCE03B0A06D5B" blockId="89.[684,1214,228,1707]" pageId="89" pageNumber="89">
<emphasis id="35B88053FF93B10D3AACCE03B0F96C32" box="[716,933,559,583]" italics="true" pageId="89" pageNumber="89">Hemispermatophore</emphasis>
: Flagelliform; flagellum, elongate and narrow (fig. 24S–U); trunk concave; three lobules, ental (LI), ectal (LE), and basal (LB); LI inclined to ental side of trunk and continuous to flagellar base; flagellar base wide, ca. two thirds width of trunk; LE length ca. half that of LI, spiniform with curved tip; LB base wide and slightly elongate, apex thin and curved.
</paragraph>
<paragraph id="07735C41FF93B10D3AACCF14B04B6DE4" blockId="89.[684,1214,228,1707]" pageId="89" pageNumber="89">
<emphasis id="35B88053FF93B10D3AACCF14B00C6D3A" box="[716,848,824,847]" italics="true" pageId="89" pageNumber="89">Cytogenetics</emphasis>
: The diploid chromosome number of 
<taxonomicName id="C0CC27C2FF93B10D3A98CF75B00E6D05" baseAuthorityName="C. L. Koch" baseAuthorityYear="1840" box="[760,850,856,880]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="89" pageNumber="89" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="35B88053FF93B10D3A98CF75B00E6D05" box="[760,850,856,880]" italics="true" pageId="89" pageNumber="89">P. debilis</emphasis>
</taxonomicName>
(
<tableCitation id="4A4E69FAFF93B10D3B06CF74B0E86D05" box="[870,948,856,880]" captionStart="TABLE 2" captionStartId="7.[610,690,228,252]" captionTargetBox="[235,1058,354,747]" captionText="TABLE 2 Diploid chromosome numbers (2 n) for Brazilian species of Rhopalurusinae from Ubinski et al., (2016) with corresponding genera indicated." httpUri="http://table.plazi.org/id/53B30CC9FFCDB1533A02CCC8B0866F32" pageId="89" pageNumber="89" tableUuid="53B30CC9FFCDB1533A02CCC8B0866F32">table 2</tableCitation>
) is 2n = 26 (
<bibRefCitation id="635D21B0FF93B10D3C28CF74B0506DE4" author="Ubinski" etAl="et al." firstAuthor="Ubinski" journalOrPublisher="Cytogenetic and Genome Research" pageId="89" pageNumber="89" pagination="83 - 155" part="148" refId="ref72292" refString="Ubinski, C. V., L. S. Carvalho, and M. C. Schneider. 2016. Chromosome evolution in Brazilian scorpions of the genus Rhopalurus and related genera (Scorpiones: Buthidae). Abstracts of the 21 st International Chromosome Conference, July 10 - 13, Foz do Iguacu, Brazil. Cytogenetic and Genome Research 148: 83 - 155. [doi: 10.1159 / 000446523]" title="Chromosome evolution in Brazilian scorpions of the genus Rhopalurus and related genera (Scorpiones: Buthidae). Abstracts of the 21 st International Chromosome Conference, July 10 - 13, Foz do Iguacu, Brazil" type="journal article" year="2016">Ubinski et al., 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFF93B10D3AACCFB7B72A6DA6" pageId="89" pageNumber="89" type="discussion">
<paragraph id="07735C41FF93B10D3AACCFB7B72A6DA6" blockId="89.[684,1214,228,1707]" pageId="89" pageNumber="89">
INCLUDED SPECIES: 
<taxonomicName id="C0CC27C2FF93B10D3BCCCFB7B06F6DA1" authority="(C. L. Koch, 1840)" authorityName="C.L. Koch" authorityYear="1840" baseAuthorityName="C. L. Koch" baseAuthorityYear="1840" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="89" pageNumber="89" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="35B88053FF93B10D3BCCCFB7B7216DC6" box="[940,1149,923,947]" italics="true" pageId="89" pageNumber="89">Physoctonus debilis</emphasis>
(C.L. 
<bibRefCitation id="635D21B0FF93B10D3ACCCF90B0776DA1" author="Koch" box="[684,811,956,980]" firstAuthor="Koch" journalOrPublisher="Nurnberg: C. H. Zeh'sche Buchhandlung" pageId="89" pageNumber="89" pagination="1 - 114" part="8" refId="ref66739" refString="Koch, C. L. 1840. Die Arachniden. Nurnberg: C. H. Zeh'sche Buchhandlung 8: 1 - 114 (pts. 1, 2, 1840; pts. 2, 3, 1841)." title="Die Arachniden" type="journal article" year="1840">Koch, 1840</bibRefCitation>
)
</taxonomicName>
; 
<taxonomicName id="C0CC27C2FF93B10D3B5ECF90B7446DA6" authority="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; CLOSE Lorenzo Prendini Scorpion Systematics Research Group, 2017" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" box="[830,1048,956,980]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="89" pageNumber="89" phylum="Arthropoda" rank="species" species="striatus" status="sp. nov.">
<emphasis id="35B88053FF93B10D3B5ECF90B7446DA6" box="[830,1048,956,980]" italics="true" pageId="89" pageNumber="89">Physoctonus striatus</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="2E8B3D28FF93B10D3C46CF90B72A6DA6" box="[1062,1142,956,979]" pageId="89" pageNumber="89" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFF93B10D3AACCFF1B0366A2D" pageId="89" pageNumber="89" type="distribution">
<paragraph id="07735C41FF93B10D3AACCFF1B0366A2D" blockId="89.[684,1214,228,1707]" pageId="89" pageNumber="89">
DISTRIBUTION: 
<taxonomicName id="C0CC27C2FF93B10D3BE4CFF1B7576D80" authorityName="Mello-Leitao" authorityYear="1934" box="[900,1035,989,1013]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="89" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF93B10D3BE4CFF1B7576D80" box="[900,1035,989,1013]" italics="true" pageId="89" pageNumber="89">Physoctonus</emphasis>
</taxonomicName>
is endemic to northeastern Brazilian, and has been recorded in the states of Bahía, Ceará, Paraíba, Pernambuco, and Piauí (fig. 9).
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFF93B10D3AACC84DB0806A88" pageId="89" pageNumber="89" type="biology_ecology">
<paragraph id="07735C41FF93B10D3AACC84DB0806A88" blockId="89.[684,1214,228,1707]" pageId="89" pageNumber="89">
ECOLOGY: The two species of 
<taxonomicName id="C0CC27C2FF93B10D3C57C84DB7E26A0C" authorityName="Mello-Leitao" authorityYear="1934" box="[1079,1214,1121,1145]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="89" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF93B10D3C57C84DB7E26A0C" box="[1079,1214,1121,1145]" italics="true" pageId="89" pageNumber="89">Physoctonus</emphasis>
</taxonomicName>
inhabit the semiarid Brazilian caatinga and cerrado (fig. 2D). These small, lapidicolous scorpions have been collected under stones and with UV light detection at night.
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFF93B10D3AACC92AB08368DE" pageId="89" pageNumber="89" type="discussion">
<paragraph id="07735C41FF93B10D3AACC92AB08368DE" blockId="89.[684,1214,228,1707]" pageId="89" pageNumber="89">
REMARKS: 
<taxonomicName id="C0CC27C2FF93B10D3B23C92AB0946B6B" authorityName="Mello-Leitao" authorityYear="1934" box="[835,968,1286,1310]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="89" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF93B10D3B23C92AB0946B6B" box="[835,968,1286,1310]" italics="true" pageId="89" pageNumber="89">Physoctonus</emphasis>
</taxonomicName>
, created to accommodate 
<taxonomicName id="C0CC27C2FF93B10D3A85C90BB7E56B4A" authority="Mello-Leitao, 1934" authorityName="Mello-Leitao" authorityYear="1934" box="[741,1209,1319,1343]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="89" pageNumber="89" phylum="Arthropoda" rank="species" species="physurus">
<emphasis id="35B88053FF93B10D3A85C90BB08B6B4A" box="[741,983,1319,1343]" italics="true" pageId="89" pageNumber="89">Physoctonus physurus</emphasis>
<bibRefCitation id="635D21B0FF93B10D3B81C90BB7E56B4A" author="Mello-Leitao" box="[993,1209,1319,1343]" firstAuthor="Mello-Leitao" journalOrPublisher="Octava Reunion de la Sociedad Argentina, Santiago del Estero" pageId="89" pageNumber="89" pagination="1 - 97" part="1933" refId="ref69252" refString="Mello-Leitao, C. de. 1934 a. (&quot; 1933 &quot;). Estudo monografico dos escorpioes da Republica Argentina. Octava Reunion de la Sociedad Argentina, Santiago del Estero 1933: 1 - 97." title="Estudo monografico dos escorpioes da Republica Argentina" type="journal article" year="1934">Mello-Leitão, 1934</bibRefCitation>
</taxonomicName>
, was synonymized with 
<taxonomicName id="C0CC27C2FF93B10D3B85C964B7376B15" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" box="[997,1131,1352,1376]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="89" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF93B10D3B85C964B7376B15" box="[997,1131,1352,1376]" italics="true" pageId="89" pageNumber="89">Rhopalurus</emphasis>
</taxonomicName>
when 
<bibRefCitation id="635D21B0FF93B10D3ACCC945B0126BF7" author="Francke" box="[684,846,1385,1410]" firstAuthor="Francke" journalOrPublisher="Journal of Arachnology" pageId="89" pageNumber="89" pagination="125 - 135" part="4" refId="ref65579" refString="Francke, O. F. 1977 a. Two emendations to Stahnke's (1974) Vaejovidae revision (Scorpionida, Vaejovidae). Journal of Arachnology 4 (2): 125 - 135." title="Two emendations to Stahnke's (1974) Vaejovidae revision (Scorpionida, Vaejovidae)" type="journal article" year="1977">Francke (1977</bibRefCitation>
a) synonymized 
<taxonomicName id="C0CC27C2FF93B10D3C67C946B7DF6BF4" authorityName="Mello-Leitao" authorityYear="1934" box="[1031,1155,1385,1409]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="89" pageNumber="89" phylum="Arthropoda" rank="species" species="physurus">
<emphasis id="35B88053FF93B10D3C67C946B7DF6BF4" box="[1031,1155,1385,1409]" italics="true" pageId="89" pageNumber="89">P. physurus</emphasis>
</taxonomicName>
with 
<taxonomicName id="C0CC27C2FF93B10D3ACCC9A7B0296BD6" authorityName=": Borelli" authorityYear="1910" box="[684,885,1419,1443]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="89" pageNumber="89" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="35B88053FF93B10D3ACCC9A7B0296BD6" box="[684,885,1419,1443]" italics="true" pageId="89" pageNumber="89">Rhopalurus debilis</emphasis>
</taxonomicName>
. 
<taxonomicName id="C0CC27C2FF93B10D3BE2C9A7B75B6BD6" authorityName="Mello-Leitao" authorityYear="1934" box="[898,1031,1419,1443]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="89" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF93B10D3BE2C9A7B75B6BD6" box="[898,1031,1419,1443]" italics="true" pageId="89" pageNumber="89">Physoctonus</emphasis>
</taxonomicName>
was later revalidated by 
<bibRefCitation id="635D21B0FF93B10D3B45C980B0AC6BB1" author="Lourenco" bookContentInfo="306 pp." box="[805,1008,1452,1476]" firstAuthor="Lourenco" journalOrPublisher="Paris: Les Editions de l'If" pageId="89" pageNumber="89" refId="ref68189" refString="Lourenco, W. R. 2002. Scorpions of Brazil. Paris: Les Editions de l'If. 306 pp." title="Scorpions of Brazil" type="book" year="2002">Lourenço (2002)</bibRefCitation>
. Its validity was upheld by the analyses of Esposito et al. (in review), which consistently recovered the monophyly of its two species as distinct from the species of 
<taxonomicName id="C0CC27C2FF93B10D3B25CA1CB09E683D" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" box="[837,962,1584,1608]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="89" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="35B88053FF93B10D3B25CA1CB09E683D" box="[837,962,1584,1608]" italics="true" pageId="89" pageNumber="89">Rhopalurus</emphasis>
</taxonomicName>
(fig. 13), and the cytogenetic study of 
<bibRefCitation id="635D21B0FF93B10D3B15CA7DB731681C" author="Ubinski" box="[885,1133,1617,1641]" etAl="et al." firstAuthor="Ubinski" journalOrPublisher="Cytogenetic and Genome Research" pageId="89" pageNumber="89" pagination="83 - 155" part="148" refId="ref72292" refString="Ubinski, C. V., L. S. Carvalho, and M. C. Schneider. 2016. Chromosome evolution in Brazilian scorpions of the genus Rhopalurus and related genera (Scorpiones: Buthidae). Abstracts of the 21 st International Chromosome Conference, July 10 - 13, Foz do Iguacu, Brazil. Cytogenetic and Genome Research 148: 83 - 155. [doi: 10.1159 / 000446523]" title="Chromosome evolution in Brazilian scorpions of the genus Rhopalurus and related genera (Scorpiones: Buthidae). Abstracts of the 21 st International Chromosome Conference, July 10 - 13, Foz do Iguacu, Brazil" type="journal article" year="2016">Ubinski et al. (2016)</bibRefCitation>
which identified a diploid chromosome number of 2n = 26 for 
<taxonomicName id="C0CC27C2FF93B10D3B6ECAB8B02E68DE" authorityName=": Borelli" authorityYear="1910" box="[782,882,1683,1707]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="89" pageNumber="89" phylum="Arthropoda" rank="species" species="debilis">
<emphasis id="35B88053FF93B10D3B6ECAB8B02E68DE" box="[782,882,1683,1707]" italics="true" pageId="89" pageNumber="89">R. debilis</emphasis>
</taxonomicName>
(
<tableCitation id="4A4E69FAFF93B10D3BE2CABFB08D68DE" box="[898,977,1683,1707]" captionStart="TABLE 2" captionStartId="7.[610,690,228,252]" captionTargetBox="[235,1058,354,747]" captionText="TABLE 2 Diploid chromosome numbers (2 n) for Brazilian species of Rhopalurusinae from Ubinski et al., (2016) with corresponding genera indicated." httpUri="http://table.plazi.org/id/53B30CC9FFCDB1533A02CCC8B0866F32" pageId="89" pageNumber="89" tableUuid="53B30CC9FFCDB1533A02CCC8B0866F32">table 2</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>