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<document id="2B890C26C9ED93635B012E46B4E18411" ID-CLB-Dataset="29930" ID-DOI="10.11646/zootaxa.4433.1.11" ID-GBIF-Dataset="63a66d44-a2f0-478a-9e74-9c439bf873d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="1287073" ID-ZooBank="DB01DC36-A123-41C3-B870-C73DD35BF6C2" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1528732591500" checkinUser="plazi" docAuthor="Bartsch, Daniel" docDate="2018" docId="847D3258EB72181962ADFC5EFD45BD51" docLanguage="en" docName="zootaxa.4433.1.11.pdf" docOrigin="Zootaxa 4433 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cinctosesia Bartsch 2018, gen. nov." docType="treatment" docVersion="7" lastPageNumber="180" masterDocId="78444A20EB74181E623AFFABFF91BF6D" masterDocTitle="Taxonomic changes in Synanthedonini from Madagascar, with description of two new genera and species (Lepidoptera: Sesiidae)" masterLastPageNumber="186" masterPageNumber="174" pageNumber="180" updateTime="1698548243284" updateUser="ExternalLinkService">
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<mods:title id="72C5BF27CF217FF31063EE85A9DE33CA">Taxonomic changes in Synanthedonini from Madagascar, with description of two new genera and species (Lepidoptera: Sesiidae)</mods:title>
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<treatment id="847D3258EB72181962ADFC5EFD45BD51" ID-DOI="http://doi.org/10.5281/zenodo.5953581" ID-GBIF-Taxon="145939518" ID-Zenodo-Dep="5953581" LSID="urn:lsid:plazi:treatment:847D3258EB72181962ADFC5EFD45BD51" httpUri="http://treatment.plazi.org/id/847D3258EB72181962ADFC5EFD45BD51" lastPageId="7" lastPageNumber="180" pageId="6" pageNumber="180">
<subSubSection id="44CED0C5EB72181862ADFC5EFE07BB62" box="[151,406,1013,1039]" pageId="6" pageNumber="180" type="nomenclature">
<paragraph id="0C6B834EEB72181862ADFC5EFE07BB62" blockId="6.[151,406,1013,1039]" box="[151,406,1013,1039]" pageId="6" pageNumber="180">
<heading id="57233422EB72181862ADFC5EFE07BB62" bold="true" box="[151,406,1013,1039]" fontSize="11" level="1" pageId="6" pageNumber="180" reason="1">
<emphasis id="3EA05F5CEB72181862ADFC5EFE07BB62" bold="true" box="[151,406,1013,1039]" pageId="6" pageNumber="180">
<taxonomicName id="CBD4F8CDEB72181862ADFC5EFEB3BB62" ID-CoL="B8XND" authority="Bartsch, 2018" authorityName="Bartsch" authorityYear="2018" box="[151,290,1013,1039]" class="Insecta" family="Sesiidae" genus="Cinctosesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="3EA05F5CEB72181862ADFC5EFEB3BB62" bold="true" box="[151,290,1013,1039]" italics="true" pageId="6" pageNumber="180">Cinctosesia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2593E227EB7218186312FC5EFE07BB62" box="[296,406,1013,1039]" pageId="6" pageNumber="180" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="44CED0C5EB72181862ADFB96FCBABB3F" box="[151,811,1084,1106]" pageId="6" pageNumber="180" type="materials_examined">
<paragraph id="0C6B834EEB72181862ADFB96FCBABB3F" blockId="6.[151,811,1084,1106]" box="[151,811,1084,1106]" pageId="6" pageNumber="180">
Type species:
<taxonomicName id="CBD4F8CDEB721818631CFB97FDB3BB3C" authority="Bartsch, 2018" authorityName="Bartsch" authorityYear="2018" box="[294,546,1084,1105]" class="Insecta" family="Sesiidae" genus="Cinctosesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="species" species="polistiformis" status="sp. nov.">
<emphasis id="3EA05F5CEB721818631CFB97FDB3BB3C" box="[294,546,1084,1105]" italics="true" pageId="6" pageNumber="180">Cinctosesia polistiformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2593E227EB7218186013FB97FDE8BB3F" box="[553,633,1084,1106]" pageId="6" pageNumber="180" rank="species">
<emphasis id="3EA05F5CEB7218186013FB97FDE2BB3C" bold="true" box="[553,627,1084,1105]" pageId="6" pageNumber="180">sp. nov</emphasis>
.
</taxonomicNameLabel>
, designated here.
</paragraph>
</subSubSection>
<subSubSection id="44CED0C5EB72181862ADFBD5FC77BBFA" box="[151,998,1150,1175]" pageId="6" pageNumber="180" type="etymology">
<paragraph id="0C6B834EEB72181862ADFBD5FC77BBFA" blockId="6.[151,1437,1150,2003]" box="[151,998,1150,1175]" pageId="6" pageNumber="180">
<emphasis id="3EA05F5CEB72181862ADFBD5FE8FBBFA" bold="true" box="[151,286,1150,1175]" pageId="6" pageNumber="180">Etymology.</emphasis>
Latin cinctus (= waist, belt),
<taxonomicName id="CBD4F8CDEB721818605DFBD4FD7BBBFB" authority="Bartsch, 2018" authorityName="Bartsch" authorityYear="2018" box="[615,746,1151,1174]" class="Insecta" family="Sesiidae" genus="Cinctosesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="3EA05F5CEB721818605DFBD4FD7BBBFB" box="[615,746,1151,1174]" italics="true" pageId="6" pageNumber="180">Cinctosesia</emphasis>
</taxonomicName>
is of feminine gender.
</paragraph>
</subSubSection>
<subSubSection id="44CED0C5EB72181862FDFB09FD88B873" pageId="6" pageNumber="180" type="description">
<paragraph id="0C6B834EEB72181862FDFB09FC51B92A" blockId="6.[151,1437,1150,2003]" pageId="6" pageNumber="180">
<emphasis id="3EA05F5CEB72181862FDFB09FEC8BBD6" bold="true" box="[199,345,1186,1211]" pageId="6" pageNumber="180">Description.</emphasis>
Medium sized (wingspan
<quantity id="CB2C2EABEB72181860B9FB08FD6FBBD6" box="[643,766,1187,1211]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="3.1" metricValueMin="2.3" pageId="6" pageNumber="180" unit="mm" value="27.0" valueMax="31.0" valueMin="23.0">2331 mm</quantity>
) with distinct paper-wasp-like appearance. Head with welldeveloped and possibly functional proboscis; labial palpi large, strongly upturned; first and third palpomeres nearly of equal length, about two thirds of second palpomere; basal two palpomeres with long, rough scales ventrally; frons with somewhat rough, vertex with smooth scales; antennae short, nearly half the length of forewing, clavate, ciliate in male. Thorax, legs and abdomen smoothly scaled, without long, hair-like scales; coxa of foreleg long and narrow; hindleg nearly as long as abdomen; spurs of all legs with outer spur less than half as long as inner one. Abdomen long, strongly wasp-waisted due to the lateral and ventral constriction of segments 24; analtuft developed, but scales pressed against abdomen, which therefore appears pointed. Wings long and narrow with large transparent areas; wing venation of forewing with veins R1R3 closely approximated, parallel; R4 and R5 with long common stalk, well separated from R3; CuA1 and CuA2 arise close together, parallel; hindwing with M2 from costad 2/5 of cross vein; M3 and CuA1 from common point or minutely stalked; discal spot of forewing narrow; discal spot of hindwing very small, triangular, extending to M2.
</paragraph>
<paragraph id="0C6B834EEB72181862FDF9F9FD88B873" blockId="6.[151,1437,1150,2003]" pageId="6" pageNumber="180">
<emphasis id="3EA05F5CEB72181862FDF9F9FEECB906" bold="true" box="[199,381,1618,1643]" pageId="6" pageNumber="180">Male genitalia.</emphasis>
Tegumen-uncus-complex long and narrow; scopula androconialis well-developed; gnathos well-developed, forming an ovoid, distally horizontally truncated plate, medial crista absent; valva somewhat rectangular shaped; inner surface nearly completely covert with bifurcate setae; crista sacculi indistinct, flat and broad, forming a double arched fold toward apex of valva, not separated from setaceous area, very densely covert with bifurcate setae; juxta strongly protruded; vinculum with short saccus; phallus (aedeagus) long and slender; vesica with single, small cornutus.
</paragraph>
</subSubSection>
<subSubSection id="44CED0C5EB72181962FDF881FD45BD51" lastPageId="7" lastPageNumber="181" pageId="6" pageNumber="180" type="diagnosis">
<paragraph id="0C6B834EEB72181962FDF881FD45BD51" blockId="6.[151,1437,1150,2003]" lastBlockId="7.[151,1436,151,573]" lastPageId="7" lastPageNumber="181" pageId="6" pageNumber="180">
<emphasis id="3EA05F5CEB72181862FDF881FED0B82E" bold="true" box="[199,321,1834,1859]" pageId="6" pageNumber="180">Diagnosis.</emphasis>
<taxonomicName id="CBD4F8CDEB721818636AF880FE42B82F" authority="Bartsch, 2018" authorityName="Bartsch" authorityYear="2018" box="[336,467,1835,1858]" class="Insecta" family="Sesiidae" genus="Cinctosesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="3EA05F5CEB721818636AF880FE42B82F" box="[336,467,1835,1858]" italics="true" pageId="6" pageNumber="180">Cinctosesia</emphasis>
</taxonomicName>
cannot be confused with any other genus within African
<taxonomicName id="CBD4F8CDEB721818669FF881FAC8B82E" authorityName="Niculescu" authorityYear="1964" box="[1189,1369,1834,1859]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
. Two characters are here considered generic autapomorphies: 1) remarkable paper-wasp-like mimicry; 2) strongly waspwaisted abdomen with constricted segments 24 (more pronounced in older specimens). Other taxa with waisted abdomen (e.g.
<emphasis id="3EA05F5CEB721818637CF83DFE34B8C2" box="[326,421,1942,1967]" italics="true" pageId="6" pageNumber="180">Lolibaja</emphasis>
<bibRefCitation id="6845FEBFEB721818638AF83DFD71B8C2" box="[432,736,1942,1967]" pageId="6" pageNumber="180" refId="ref6556" refString="Gorbunov, O. G. &amp; Gurko, V. O. (2017) A new genus and species of clearwing moths (Lepidoptera: Sesiidae) from South Sudan. Zootaxa, 4276 (2), 270 - 276. https: // doi. org / 10.11646 / zootaxa. 4276.2.8" type="journal article">Gorbunov &amp; Gurko, 2017</bibRefCitation>
and
<taxonomicName id="CBD4F8CDEB7218186125F83DFBC4B8C2" authority="Holland, 1893" authorityName="Holland" authorityYear="1893" box="[799,1109,1942,1967]" class="Insecta" family="Sesiidae" genus="Tipulamima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="genus">
<emphasis id="3EA05F5CEB7218186125F83DFC34B8C2" box="[799,933,1942,1967]" italics="true" pageId="6" pageNumber="180">Tipulamima</emphasis>
<bibRefCitation id="6845FEBFEB721818618AF83DFBC4B8C2" box="[944,1109,1942,1967]" pageId="6" pageNumber="180" refId="ref6718" refString="Holland, W. J. (1893) Four new genera and species of West African Sesiidae. Journal of the New York Entomological Society, 1, 181 - 184." type="journal article">Holland, 1893</bibRefCitation>
</taxonomicName>
partim) have the waist less pronounced and only segments 12 narrowed. Additionally, the latter taxa differ from
<taxonomicName id="CBD4F8CDEB721818665BF810FB75B8BF" authority="Bartsch, 2018" authorityName="Bartsch" authorityYear="2018" box="[1121,1252,1979,2002]" class="Insecta" family="Sesiidae" genus="Cinctosesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="180" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="3EA05F5CEB721818665BF810FB75B8BF" box="[1121,1252,1979,2002]" italics="true" pageId="6" pageNumber="180">Cinctosesia</emphasis>
</taxonomicName>
by the length of their hindlegs, which are considerably longer than the abdomen (shorter than abdomen in
<taxonomicName id="CBD4F8CDEB73181966ACFF33FA88BFC2" authority="Bartsch, 2018" authorityName="Bartsch" authorityYear="2018" box="[1174,1305,152,175]" class="Insecta" family="Sesiidae" genus="Cinctosesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="180" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="3EA05F5CEB73181966ACFF33FA88BFC2" box="[1174,1305,152,175]" italics="true" pageId="7" pageNumber="181">Cinctosesia</emphasis>
</taxonomicName>
) and in the genitalia structure: tegumen shorter; uncus stronger with scopula androconialis shorter; crista sacculi with distal portion down curved toward ventral margin and base of valva (distal portion toward apex of valva in
<taxonomicName id="CBD4F8CDEB7318196732FF4BFA1ABF9A" authority="Bartsch, 2018" authorityName="Bartsch" authorityYear="2018" box="[1288,1419,224,247]" class="Insecta" family="Sesiidae" genus="Cinctosesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="180" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="3EA05F5CEB7318196732FF4BFA1ABF9A" box="[1288,1419,224,247]" italics="true" pageId="7" pageNumber="181">Cinctosesia</emphasis>
</taxonomicName>
); phallus shorter and broader, basally not bulbous-like enlarged;
<taxonomicName id="CBD4F8CDEB7318196161FEAFFC70BE70" authorityName="Holland" authorityYear="1894" box="[859,993,260,285]" class="Insecta" family="Sesiidae" genus="Tipulamima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="3EA05F5CEB7318196161FEAFFC70BE70" box="[859,993,260,285]" italics="true" pageId="7" pageNumber="181">Tipulamima</emphasis>
</taxonomicName>
differs further by an apical hook of the valva. The most distinct external characteristics of
<taxonomicName id="CBD4F8CDEB73181960D6FE83FCFEBE52" authority="Bartsch, 2018" authorityName="Bartsch" authorityYear="2018" box="[748,879,296,319]" class="Insecta" family="Sesiidae" genus="Cinctosesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="180" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="3EA05F5CEB73181960D6FE83FCFEBE52" box="[748,879,296,319]" italics="true" pageId="7" pageNumber="181">Cinctosesia</emphasis>
</taxonomicName>
are: antennae short and strong, around half the length of forewing (clearly longer, two thirds to three-fourths of forewing in most
<taxonomicName id="CBD4F8CDEB731819665DFEE7FA8DBE08" authorityName="Niculescu" authorityYear="1964" box="[1127,1308,332,357]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="181" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
); smoothly scaled body (thorax, legs and first abdominal tergites with hair-like scales in most
<taxonomicName id="CBD4F8CDEB731819660EFEC4FB78BEE5" authorityName="Niculescu" authorityYear="1964" box="[1076,1257,367,392]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="181" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
); narrow wings, veins R1 and R2 not fused distally (distally fused or rarely completely coincident in many
<taxonomicName id="CBD4F8CDEB7318196681FE3FFAE1BEC0" authorityName="Niculescu" authorityYear="1964" box="[1211,1392,404,429]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="181" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
). In male genitalia, the valva is distinct with crista sacculi and setaceous area fused, apex barely tapering (more or less pointed in most
<taxonomicName id="CBD4F8CDEB7318196370FE77FE6FBE98" authorityName="Niculescu" authorityYear="1964" box="[330,510,476,501]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="181" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
), inner surface nearly completely covered with sensory hairs (with bald area dorsodistally of crista sacculi in most
<taxonomicName id="CBD4F8CDEB731819603EFE54FD29BD75" authorityName="Niculescu" authorityYear="1964" box="[516,696,511,536]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="181" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
), and the vesica with only a single cornutus (with numerous short spines in many
<taxonomicName id="CBD4F8CDEB731819637CFD8FFE68BD50" authorityName="Niculescu" authorityYear="1964" box="[326,505,548,573]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="181" phylum="Arthropoda" rank="tribe" tribe="Synanthedonini">Synanthedonini</taxonomicName>
, e.g.
<taxonomicName id="CBD4F8CDEB7318196009FD8FFD55BD50" box="[563,708,548,573]" class="Insecta" family="Sesiidae" genus="Synanthedon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="181" phylum="Arthropoda" rank="genus">
<emphasis id="3EA05F5CEB7318196009FD8FFD55BD50" box="[563,708,548,573]" italics="true" pageId="7" pageNumber="181">Synanthedon</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
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