treatments-xml/data/79/7C/87/797C87A3FFC36224FF25A1F9FAF6FD35.xml

<document id="1BB1A2AFFD7CC4B253CC434B9C1D9CB7" ID-DOI="10.5281/zenodo.196825" ID-GBIF-Dataset="94931097-3db5-4a60-99e3-9457eaf91a55" ID-ISSN="1175-5326" ID-Zenodo-Dep="196825" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460253684862" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2010" docId="797C87A3FFC36224FF25A1F9FAF6FD35" docLanguage="en" docName="finalversionAnkerpaper.imf" docOrigin="Zootaxa 2552" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Metabetaeus mcphersonae Anker, 2010, n. sp." docType="treatment" docVersion="9" lastPageNumber="48" masterDocId="8545FFDBFFC0622FFFB2A123FFA7FF89" masterDocTitle="Metabetaeus Borradaile, 1899 revisited, with description of a new marine species from French Polynesia (Crustacea: Decapoda: Alpheidae)" masterLastPageNumber="54" masterPageNumber="37" originalUpdateDomain="Boston" originalUpdateTime="1545145707360" originalUpdateUser="gglab" pageNumber="40" updateTime="1698243125732" updateUser="plazi">
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<mods:title id="E569B70E4B5CC27DCB4C06A2B2014CD3">Metabetaeus Borradaile, 1899 revisited, with description of a new marine species from French Polynesia (Crustacea: Decapoda: Alpheidae)</mods:title>
</mods:titleInfo>
<mods:name id="8296C9AD10A5FE8FBD5D3907596C6334" type="personal">
<mods:role id="7E93CAF0321CAEB8DBD4EBEA8174A69D">
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<mods:namePart id="D7BD6FBDCC3EB57E689C187384A6A8FA">Anker, Arthur</mods:namePart>
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<mods:title id="54C0BB18D1CC459858D45820CD39D6BD">Zootaxa</mods:title>
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<mods:part id="554DF994D31258A7A76F0DC5DD27D64A">
<mods:date id="E828E41233AA8B61C7387F125EC4BE95">2010</mods:date>
<mods:detail id="595CCF669038CCB1F8C654B82F48C625" type="volume">
<mods:number id="B7356F24CA83B070BF4939B1659DCE73">2552</mods:number>
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<mods:classification id="D42FD7F7F07FC596060ED94510A5C89C">journal article</mods:classification>
<mods:identifier id="E89EDDEEF7DBE856AAFB8AD588EDB69B" type="DOI">10.5281/zenodo.196825</mods:identifier>
<mods:identifier id="7D910146F90821CC4F3A297A324060A8" type="GBIF-Dataset">94931097-3db5-4a60-99e3-9457eaf91a55</mods:identifier>
<mods:identifier id="2A39ADF2F7D248E47356BF365472F841" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E8AB3E47C1A9CC6FCB2C18C40ADE5FE9" type="Zenodo-Dep">196825</mods:identifier>
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<treatment id="797C87A3FFC36224FF25A1F9FAF6FD35" ID-DOI="http://doi.org/10.5281/zenodo.6206640" ID-GBIF-Taxon="119392128" ID-Zenodo-Dep="6206640" LSID="urn:lsid:plazi:treatment:797C87A3FFC36224FF25A1F9FAF6FD35" httpUri="http://treatment.plazi.org/id/797C87A3FFC36224FF25A1F9FAF6FD35" lastPageId="11" lastPageNumber="48" pageId="3" pageNumber="40">
<subSubSection id="B9CF653EFFC3622CFF25A1F9FEA3FE9E" pageId="3" pageNumber="40" type="nomenclature">
<paragraph id="F16A36B5FFC3622CFF25A1F9FD86FF7D" blockId="3.[151,545,218,279]" box="[151,545,218,244]" pageId="3" pageNumber="40">
<heading id="AA2281D9FFC3622CFF25A1F9FD86FF7D" bold="true" box="[151,545,218,244]" fontSize="11" level="1" pageId="3" pageNumber="40" reason="1">
<emphasis id="C3A1EAA7FFC3622CFF25A1F9FD86FF7D" bold="true" box="[151,545,218,244]" pageId="3" pageNumber="40">
<taxonomicName id="36D54D36FFC3622CFF25A1F9FE70FF7D" ID-CoL="85V8X" box="[151,471,218,244]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="mcphersonae" status="sp. nov.">
<emphasis id="C3A1EAA7FFC3622CFF25A1F9FE70FF7D" bold="true" box="[151,471,218,244]" italics="true" pageId="3" pageNumber="40">Metabetaeus mcphersonae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D89257DCFFC3622CFE6DA1F9FD86FF7D" box="[479,545,218,244]" pageId="3" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F16A36B5FFC3622CFF25A1DEFEA3FE9E" blockId="3.[151,545,218,279]" box="[151,260,253,279]" pageId="3" pageNumber="40">
<figureCitation id="69EE2A30FFC3622CFF25A1DEFEA3FE9E" box="[151,260,253,279]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="3.[151,258,1713,1737]" captionStartId-1="5.[151,258,1844,1868]" captionStartId-2="6.[151,258,1517,1541]" captionStartId-3="7.[151,258,1508,1532]" captionStartId-4="8.[151,258,1948,1972]" captionStartId-5="9.[151,258,1942,1966]" captionStartId-6="10.[151,258,1613,1637]" captionTargetBox-0="[162,1422,654,1677]" captionTargetBox-1="[187,1399,194,1843]" captionTargetBox-2="[164,1430,187,1516]" captionTargetBox-3="[151,1424,188,1480]" captionTargetBox-4="[151,1436,194,1926]" captionTargetBox-5="[151,1436,194,1919]" captionTargetBox-6="[151,1436,194,1590]" captionTargetId-0="figure@3.[151,1436,634,1689]" captionTargetId-1="figure@5.[151,1436,163,1857]" captionTargetId-2="figure@6.[151,1436,172,1534]" captionTargetId-3="figure@7.[151,1436,178,1487]" captionTargetId-4="figure@8.[151,1436,194,1926]" captionTargetId-5="figure@9.[151,1436,194,1921]" captionTargetId-6="figure@10.[151,1436,194,1592]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionText-0="FIGURE 1. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, frontal region, dorsal; B, same, lateral; C, tooth on ventromesial carina of first segment of antennular peduncle, lateral; D, first pleopod with two embryos attached, lateral; E, second pleopod, lateral; F, uropod, dorsal; G, telson, dorsal." captionText-1="FIGURE 2. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, mandible, mesial; B, same, incisor process and palp, lateral; C, maxillule, lateral; D, maxilla, lateral; E, first maxilliped, lateral; F, second maxilliped, lateral (epipod missing, detached during dissection); G, third maxilliped (right), lateral; H, third maxilliped (left), ultimate segment, mesial." captionText-2="FIGURE 3. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, cheliped (right), lateral; B, same, row of spiniform setae on ventromesial margin of merus, mesial; C, same, chela, mesial; D, second pereiopod, lateral; E, third pereiopod, lateral; F, same, propodus and dactylus, lateral; G, fourth pereiopod, lateral; H, fifth pereiopod, lateral." captionText-3="FIGURE 4. Metabetaeus mcphersonae n. sp., holotype, ovigerous female from Moorea (FLMNH UF Arthropoda 23247): A, frontal region, dorsal; B, major cheliped (right), lateral; C, same, chela, mesial; D, same, chela fingers; E, minor cheliped (left), chela, mesial; paratype, male from Moorea (FLMNH UF Arthropoda 23249): F, cheliped (right), chela, mesial; G, second pleopod, lateral." captionText-4="FIGURE 5. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, habitus, lateral; B, same, dorsal." captionText-5="FIGURE 6. Metabetaeus mcphersonae n. sp., holotype, ovigerous female from Moorea (FLMNH UF Arthropoda 23247): A, habitus, lateral; B, same, dorsal." captionText-6="FIGURE 7. Metabetaeus mcphersonae n. sp., holotype, ovigerous female from Moorea (FLMNH UF Arthropoda 23247): A, frontal region, dorsal; B, major cheliped (right), chela, mesial (= dorsal); paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): C, anterior region, lateral; D, chelipeds, lateral (= ventral); E, uropod, dorsal; paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23248): F, embryos attached to pleopods; G, zoea." httpUri-0="https://zenodo.org/record/196826/files/figure.png" httpUri-1="https://zenodo.org/record/196827/files/figure.png" httpUri-2="https://zenodo.org/record/196828/files/figure.png" httpUri-3="https://zenodo.org/record/196829/files/figure.png" httpUri-4="https://zenodo.org/record/196830/files/figure.png" httpUri-5="https://zenodo.org/record/196831/files/figure.png" httpUri-6="https://zenodo.org/record/196832/files/figure.png" pageId="3" pageNumber="40">Figs. 1–7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="B9CF653EFFC3622CFF25A064FC03FDCB" pageId="3" pageNumber="40" type="materials_examined">
<paragraph id="F16A36B5FFC3622CFF25A064FC03FDCB" blockId="3.[151,1436,327,578]" pageId="3" pageNumber="40">
<emphasis id="C3A1EAA7FFC3622CFF25A064FE2DFEE8" bold="true" box="[151,394,327,353]" pageId="3" pageNumber="40">Material examined</emphasis>
. 
<materialsCitation id="41BD3CE8FFC3622CFE28A06BFCA3FE5B" ID-GBIF-Occurrence="1948792858" collectingDate="2009-12-05" collectionCode="FLMNH" collectorName="S. McPherson" country="French Polynesia" elevation="-15" latitude="-17.47564" location="Moorea, in front of Hilton Hotel" longitude="-149.84247" pageId="3" pageNumber="40" specimenCode="FLMNH UF Arthropoda 23247" specimenCount="1" specimenCount-female="1" stateProvince="Society Islands" typeStatus="holotype">
<typeStatus id="2E6E8817FFC3622CFE28A06BFDA8FEEB" box="[410,527,328,354]" pageId="3" pageNumber="40" type="holotype">Holotype</typeStatus>
: ovigerous 
<specimenCount id="E7D3FD3CFFC3622CFD2BA06BFD4CFEEB" box="[665,747,328,354]" pageId="3" pageNumber="40" type="female">female</specimenCount>
(cl 3.0 mm, tl 
<quantity id="362D9B50FFC3622CFC13A06BFBA5FEEB" box="[929,1026,328,354]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="3" pageNumber="40" unit="mm" value="8.1">8.1 mm</quantity>
), 
<collectionCode id="97C4AE70FFC3622CFBAAA06BFB27FEEB" box="[1048,1152,328,354]" collectionName="FLMNH" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="3" pageNumber="40">FLMNH</collectionCode>
UF Arthropoda 
<specimenCode id="A1739ECEFFC3622CFAF8A06BFA3FFEEB" box="[1354,1432,328,354]" pageId="3" pageNumber="40">23247</specimenCode>
, 
<collectingCountry id="89C27625FFC3622CFF25A04EFEC2FE0E" box="[151,357,365,391]" name="French Polynesia" pageId="3" pageNumber="40" value="French Polynesia">French Polynesia</collectingCountry>
, 
<collectingRegion id="3311F857FFC3622CFEC0A04EFD8FFE0E" box="[370,552,365,391]" pageId="3" pageNumber="40">Society Islands</collectingRegion>
, 
<location id="F40A606EFFC3622CFD86A04EFC11FE0E" LSID="urn:lsid:plazi:treatment:797C87A3FFC36224FF25A1F9FAF6FD35:F40A606EFFC3622CFD86A04EFC11FE0E" box="[564,950,365,391]" country="French Polynesia" latitude="-17.47564" longLatPrecision="1" longitude="-149.84247" name="Moorea, in front of Hilton Hotel" pageId="3" pageNumber="40" stateProvince="Society Islands">Moorea, in front of Hilton Hotel</location>
, 
<geoCoordinate id="94E15072FFC3622CFC71A04EFBC3FE0E" box="[963,1124,365,391]" direction="west" orientation="longitude" pageId="3" pageNumber="40" precision="1" value="-149.84247">149.84247 W</geoCoordinate>
, 
<geoCoordinate id="94E15072FFC3622CFBC2A04EFB5FFE0E" box="[1136,1272,365,391]" direction="south" orientation="latitude" pageId="3" pageNumber="40" precision="1" value="-17.47564">17.47564 S</geoCoordinate>
, depth: 
<elevation id="7AF8D186FFC3622CFAE5A04EFF16FE25" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="3" pageNumber="40" unit="m" value="15.0" valueMax="17.0" valueMin="13.0">
<quantity id="362D9B50FFC3622CFAE5A04EFF16FE25" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="3" pageNumber="40" unit="m" value="15.0" valueMax="17.0" valueMin="13.0">13-17 m</quantity>
</elevation>
, fore-reef, digging and fanning of coarse sand and small-sized rubble, collector 
<collectorName id="5C205363FFC3622CFBDCA0B1FAB2FE25" box="[1134,1301,402,428]" pageId="3" pageNumber="40">S. McPherson</collectorName>
(processed by A. Anker), 
<collectingDate id="952FE99DFFC3622CFEF6A09BFDBFFE5B" box="[324,536,440,466]" pageId="3" pageNumber="40" value="2009-12-05">
<date id="856B1075FFC3622CFEF6A09BFDBFFE5B" box="[324,536,440,466]" pageId="3" pageNumber="40" value="2009-12-05">5 December 2009</date>
</collectingDate>
(fcn BMOO 9789).
</materialsCitation>
<materialsCitation id="41BD3CE8FFC3622CFCBEA09BFDC0FE7E" ID-GBIF-Occurrence="1948792750" collectingDate="2009-12-05" collectionCode="FLMNH" collectorName="S. McPherson" country="French Polynesia" elevation="-15" latitude="-17.47564" location="Moorea, in front of Hilton Hotel" longitude="-149.84247" pageId="3" pageNumber="40" specimenCode="FLMNH UF Arthropoda 23249" specimenCount="1" specimenCount-male="1" stateProvince="Society Islands" typeStatus="paratype">
<typeStatus id="2E6E8817FFC3622CFCBEA09BFC24FE5B" box="[780,899,440,466]" pageId="3" pageNumber="40" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="E7D3FD3CFFC3622CFC3DA09BFC78FE5B" box="[911,991,440,466]" pageId="3" pageNumber="40" type="male">1 male</specimenCount>
(cl 
<quantity id="362D9B50FFC3622CFBBDA09BFBCAFE5B" box="[1039,1133,440,466]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="3" pageNumber="40" unit="mm" value="2.1">2.1 mm</quantity>
, tl 
<quantity id="362D9B50FFC3622CFB20A09BFB56FE5B" box="[1170,1265,440,466]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="3" pageNumber="40" unit="mm" value="6.3">6.3 mm</quantity>
), 
<collectionCode id="97C4AE70FFC3622CFAB4A09BFACAFE5B" box="[1286,1389,440,466]" collectionName="FLMNH" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="3" pageNumber="40">FLMNH</collectionCode>
UF Arthropoda 
<specimenCode id="A1739ECEFFC3622CFE98A0FEFED2FE7E" box="[298,373,477,503]" pageId="3" pageNumber="40">23249</specimenCode>
(fcn BMOO 9791);
</materialsCitation>
<materialsCitation id="41BD3CE8FFC3622CFDDDA0FEFDC7FD95" ID-GBIF-Occurrence="1948792859" collectingDate="2009-12-05" collectionCode="FLMNH" collectorName="S. McPherson" country="French Polynesia" elevation="-15" latitude="-17.47564" location="Moorea, in front of Hilton Hotel" longitude="-149.84247" pageId="3" pageNumber="40" specimenCode="FLMNH UF Arthropoda 23250" specimenCount="1" specimenCount-female="1" stateProvince="Society Islands" typeStatus="paratype">
<specimenCount id="E7D3FD3CFFC3622CFDDDA0FEFDD9FE7E" box="[623,638,477,503]" pageId="3" pageNumber="40">1</specimenCount>
ovigerous 
<specimenCount id="E7D3FD3CFFC3622CFCB4A0FEFCFFFE7E" box="[774,856,477,503]" pageId="3" pageNumber="40" type="female">female</specimenCount>
(cl 
<quantity id="362D9B50FFC3622CFC3AA0FEFC4EFE7E" box="[904,1001,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="40" unit="mm" value="2.8">2.8 mm</quantity>
, tl 
<quantity id="362D9B50FFC3622CFBBDA0FEFBC8FE7E" box="[1039,1135,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="3" pageNumber="40" unit="mm" value="8.3">8.3 mm</quantity>
, dissected), 
<collectionCode id="97C4AE70FFC3622CFAB7A0FEFAC9FE7E" box="[1285,1390,477,503]" collectionName="FLMNH" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="3" pageNumber="40">FLMNH</collectionCode>
UF Arthropoda 
<specimenCode id="A1739ECEFFC3622CFE9AA321FED5FD95" box="[296,370,514,540]" pageId="3" pageNumber="40">23250</specimenCode>
(fcn BMOO 9788);
</materialsCitation>
<materialsCitation id="41BD3CE8FFC3622CFDDAA321FC03FDCB" ID-GBIF-Occurrence="1948792751" collectingDate="2009-12-05" collectionCode="FLMNH" collectorName="S. McPherson" country="French Polynesia" elevation="-15" latitude="-17.47564" location="Moorea, in front of Hilton Hotel" longitude="-149.84247" pageId="3" pageNumber="40" specimenCode="FLNMH UF Arthropoda 23248" specimenCount="1" specimenCount-female="1" stateProvince="Society Islands" typeStatus="paratype">
<specimenCount id="E7D3FD3CFFC3622CFDDAA321FDD0FD95" box="[616,631,514,540]" pageId="3" pageNumber="40">1</specimenCount>
ovigerous 
<specimenCount id="E7D3FD3CFFC3622CFD4EA321FCEAFD95" box="[764,845,514,540]" pageId="3" pageNumber="40" type="female">female</specimenCount>
(cl 
<quantity id="362D9B50FFC3622CFCCFA321FC7CFD95" box="[893,987,514,540]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="40" unit="mm" value="2.4">2.4 mm</quantity>
, tl 
<quantity id="362D9B50FFC3622CFBB2A321FBC7FD95" box="[1024,1120,514,540]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="3" pageNumber="40" unit="mm" value="7.1">7.1 mm</quantity>
), 
<collectionCode id="97C4AE70FFC3622CFBC7A321FB7BFD95" box="[1141,1244,514,540]" collectionName="FLMNH" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="3" pageNumber="40">FLMNH</collectionCode>
UF Arthropoda 
<specimenCode id="A1739ECEFFC3622CFF25A30BFF47FDCB" box="[151,224,552,578]" pageId="3" pageNumber="40">23248</specimenCode>
(fcn BMOO 9790); all same collection data as for holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="A5AA663DFFC3622CFF25A792FE8BF8A1" httpUri="https://zenodo.org/record/196826/files/figure.png" pageId="3" pageNumber="40" targetBox="[162,1422,654,1677]" targetPageId="3">
<paragraph id="F16A36B5FFC3622CFF25A792FE8BF8A1" blockId="3.[151,1437,1713,1832]" pageId="3" pageNumber="40">
<emphasis id="C3A1EAA7FFC3622CFF25A792FEBFF940" bold="true" box="[151,280,1713,1737]" pageId="3" pageNumber="40">FIGURE 1</emphasis>
. 
<taxonomicName id="36D54D36FFC3622CFE95A792FDF6F941" box="[295,593,1713,1736]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="mcphersonae" status="sp. nov.">
<emphasis id="C3A1EAA7FFC3622CFE95A792FDF6F941" box="[295,593,1713,1736]" italics="true" pageId="3" pageNumber="40">Metabetaeus mcphersonae</emphasis>
</taxonomicName>
<emphasis id="C3A1EAA7FFC3622CFDE8A792FD3EF940" bold="true" box="[602,665,1713,1737]" pageId="3" pageNumber="40">
<taxonomicNameLabel id="D89257DCFFC3622CFDE8A792FD3EF940" box="[602,665,1713,1737]" pageId="3" pageNumber="40" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, frontal region, dorsal; B, same, lateral; C, tooth on ventromesial carina of first segment of antennular peduncle, lateral; D, first pleopod with two embryos attached, lateral; E, second pleopod, lateral; F, uropod, dorsal; G, telson, dorsal.
</paragraph>
</caption>
<subSubSection id="B9CF653EFFC36229FF74A675FB05F934" lastPageId="6" lastPageNumber="43" pageId="3" pageNumber="40" type="description">
<paragraph id="F16A36B5FFC3622BFF74A675FD65FF7E" blockId="3.[151,1437,1878,2008]" lastBlockId="4.[151,1438,152,2015]" lastPageId="4" lastPageNumber="41" pageId="3" pageNumber="40">
<emphasis id="C3A1EAA7FFC3622CFF74A675FEF0F8F9" bold="true" box="[198,343,1878,1904]" pageId="3" pageNumber="40">Description</emphasis>
. Carapace smooth, not setose. Frontal margin with rostrum and extra-corneal teeth; rostrum variable in shape, triangular or subtriangular, as broad as long or longer than broad, reaching to mid-length of first segment of antennular peduncle; orbital teeth well developed, acute distally, directed more or less mesially (
<figureCitation id="69EE2A30FFC3622CFEB8A69DFEF0F851" box="[266,343,1982,2008]" captionStart="FIGURE 1" captionStartId="3.[151,258,1713,1737]" captionTargetBox="[162,1422,654,1677]" captionTargetId="figure@3.[151,1436,634,1689]" captionTargetPageId="3" captionText="FIGURE 1. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, frontal region, dorsal; B, same, lateral; C, tooth on ventromesial carina of first segment of antennular peduncle, lateral; D, first pleopod with two embryos attached, lateral; E, second pleopod, lateral; F, uropod, dorsal; G, telson, dorsal." httpUri="https://zenodo.org/record/196826/files/figure.png" pageId="3" pageNumber="40">Figs. 1</figureCitation>
A, 4A). Pterygostomial angle rounded, not protruding anteriorly. Posterior margin with welldeveloped cardiac notch. Epistomial sclerites not projecting ventrally. Ocellar beak obscure, not visible in lateral view. Eyestalks completely concealed in dorsal and lateral views; cornea well developed, ovalelongate, in lateral position (
<figureCitation id="69EE2A30FFC4622BFE54A1FEFD94FF7E" box="[486,563,221,247]" captionStart="FIGURE 1" captionStartId="3.[151,258,1713,1737]" captionTargetBox="[162,1422,654,1677]" captionTargetId="figure@3.[151,1436,634,1689]" captionTargetPageId="3" captionText="FIGURE 1. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, frontal region, dorsal; B, same, lateral; C, tooth on ventromesial carina of first segment of antennular peduncle, lateral; D, first pleopod with two embryos attached, lateral; E, second pleopod, lateral; F, uropod, dorsal; G, telson, dorsal." httpUri="https://zenodo.org/record/196826/files/figure.png" pageId="4" pageNumber="41">Figs. 1</figureCitation>
A, 4A, 7A).
</paragraph>
<paragraph id="F16A36B5FFC4622BFF74A023FF6BFE63" blockId="4.[151,1438,152,2015]" pageId="4" pageNumber="41">
Pleura of second to fourth abdominal somites rounded; fifth pleuron with posteroventral angle produced into small subacute tooth; sixth somite without distinct articulated plate at posteroventral angle (indistinct suture visible in living specimens); preanal plate rounded. Telson subrectangular, slightly tapering posteriorly; dorsal surface with two pairs of strong spiniform setae inserted first at mid-length and second at 0.7–0.8 length of telson, at some distance from edges; posterior margin rounded medially, each angle with two pairs of strong spiniform setae, mesial distinctly longer than lateral; central area with three long plumose setae (
<figureCitation id="69EE2A30FFC4622BFADEA08EFF05FE63" captionStart="FIGURE 1" captionStartId="3.[151,258,1713,1737]" captionTargetBox="[162,1422,654,1677]" captionTargetId="figure@3.[151,1436,634,1689]" captionTargetPageId="3" captionText="FIGURE 1. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, frontal region, dorsal; B, same, lateral; C, tooth on ventromesial carina of first segment of antennular peduncle, lateral; D, first pleopod with two embryos attached, lateral; E, second pleopod, lateral; F, uropod, dorsal; G, telson, dorsal." httpUri="https://zenodo.org/record/196826/files/figure.png" pageId="4" pageNumber="41">Fig. 1</figureCitation>
G).
</paragraph>
<paragraph id="F16A36B5FFC4622BFF74A0D1FA9EFD33" blockId="4.[151,1438,152,2015]" pageId="4" pageNumber="41">
Antennular peduncle reaching to end of antennal carpocerite, greatly overreachiung scaphocerite; first segment with stylocerite reaching to 0.3 length of second segment, with acute tip; mesioventral carina with strong, anteriorly directed tooth (
<figureCitation id="69EE2A30FFC4622BFD84A31BFDDAFDDB" box="[566,637,568,594]" captionStart="FIGURE 1" captionStartId="3.[151,258,1713,1737]" captionTargetBox="[162,1422,654,1677]" captionTargetId="figure@3.[151,1436,634,1689]" captionTargetPageId="3" captionText="FIGURE 1. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, frontal region, dorsal; B, same, lateral; C, tooth on ventromesial carina of first segment of antennular peduncle, lateral; D, first pleopod with two embryos attached, lateral; E, second pleopod, lateral; F, uropod, dorsal; G, telson, dorsal." httpUri="https://zenodo.org/record/196826/files/figure.png" pageId="4" pageNumber="41">Fig. 1</figureCitation>
C); second segment shorter than dorsally visible portion of first segment, somewhat longer than wide (
<figureCitation id="69EE2A30FFC4622BFDE5A379FD3FFDFD" box="[599,664,602,628]" captionStart="FIGURE 1" captionStartId="3.[151,258,1713,1737]" captionTargetBox="[162,1422,654,1677]" captionTargetId="figure@3.[151,1436,634,1689]" captionTargetPageId="3" captionText="FIGURE 1. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, frontal region, dorsal; B, same, lateral; C, tooth on ventromesial carina of first segment of antennular peduncle, lateral; D, first pleopod with two embryos attached, lateral; E, second pleopod, lateral; F, uropod, dorsal; G, telson, dorsal." httpUri="https://zenodo.org/record/196826/files/figure.png" pageId="4" pageNumber="41">Fig. 1</figureCitation>
A, 4A). Antenna with basicerite bearing strong, sharp distoventral tooth (
<figureCitation id="69EE2A30FFC4622BFF54A35EFE8CFD1E" box="[230,299,637,663]" captionStart="FIGURE 1" captionStartId="3.[151,258,1713,1737]" captionTargetBox="[162,1422,654,1677]" captionTargetId="figure@3.[151,1436,634,1689]" captionTargetPageId="3" captionText="FIGURE 1. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, frontal region, dorsal; B, same, lateral; C, tooth on ventromesial carina of first segment of antennular peduncle, lateral; D, first pleopod with two embryos attached, lateral; E, second pleopod, lateral; F, uropod, dorsal; G, telson, dorsal." httpUri="https://zenodo.org/record/196826/files/figure.png" pageId="4" pageNumber="41">Fig. 1</figureCitation>
B); carpocerite significantly overreaching scaphocerite; scaphocerite ovate, with strong, sharp distolateral tooth and convex distal margin, latter slightly exceeding distolateral tooth (
<figureCitation id="69EE2A30FFC4622BFB3DA383FB7AFD33" box="[1167,1245,672,698]" captionStart="FIGURE 1" captionStartId="3.[151,258,1713,1737]" captionTargetBox="[162,1422,654,1677]" captionTargetId="figure@3.[151,1436,634,1689]" captionTargetPageId="3" captionText="FIGURE 1. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, frontal region, dorsal; B, same, lateral; C, tooth on ventromesial carina of first segment of antennular peduncle, lateral; D, first pleopod with two embryos attached, lateral; E, second pleopod, lateral; F, uropod, dorsal; G, telson, dorsal." httpUri="https://zenodo.org/record/196826/files/figure.png" pageId="4" pageNumber="41">Figs. 1</figureCitation>
A, 4A).
</paragraph>
<paragraph id="F16A36B5FFC4622BFF74A3E1FC92FB9D" blockId="4.[151,1438,152,2015]" pageId="4" pageNumber="41">
Mouthparts typical to genus. Mandible with two-segmented palp, well-developed molar process and incisor process bearing at least six teeth distally and large dark-brown spot proximally (
<figureCitation id="69EE2A30FFC4622BFB27A3C6FB7CFD76" box="[1173,1243,741,767]" captionStart="FIGURE 2" captionStartId="5.[151,258,1844,1868]" captionTargetBox="[187,1399,194,1843]" captionTargetId="figure@5.[151,1436,163,1857]" captionTargetPageId="5" captionText="FIGURE 2. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, mandible, mesial; B, same, incisor process and palp, lateral; C, maxillule, lateral; D, maxilla, lateral; E, first maxilliped, lateral; F, second maxilliped, lateral (epipod missing, detached during dissection); G, third maxilliped (right), lateral; H, third maxilliped (left), ultimate segment, mesial." httpUri="https://zenodo.org/record/196827/files/figure.png" pageId="4" pageNumber="41">Fig. 2</figureCitation>
A, B). Maxillule with bilobed palp, each furnished with one seta; basal endite with particularly strong spine-like setae (
<figureCitation id="69EE2A30FFC4622BFAD9A22BFF00FCCD" captionStart="FIGURE 2" captionStartId="5.[151,258,1844,1868]" captionTargetBox="[187,1399,194,1843]" captionTargetId="figure@5.[151,1436,163,1857]" captionTargetPageId="5" captionText="FIGURE 2. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, mandible, mesial; B, same, incisor process and palp, lateral; C, maxillule, lateral; D, maxilla, lateral; E, first maxilliped, lateral; F, second maxilliped, lateral (epipod missing, detached during dissection); G, third maxilliped (right), lateral; H, third maxilliped (left), ultimate segment, mesial." httpUri="https://zenodo.org/record/196827/files/figure.png" pageId="4" pageNumber="41">Fig. 2</figureCitation>
C). Maxilla with simple endopod; basal endite rounded and notched distally (
<figureCitation id="69EE2A30FFC4622BFB80A209FBDEFCCD" box="[1074,1145,810,836]" captionStart="FIGURE 2" captionStartId="5.[151,258,1844,1868]" captionTargetBox="[187,1399,194,1843]" captionTargetId="figure@5.[151,1436,163,1857]" captionTargetPageId="5" captionText="FIGURE 2. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, mandible, mesial; B, same, incisor process and palp, lateral; C, maxillule, lateral; D, maxilla, lateral; E, first maxilliped, lateral; F, second maxilliped, lateral (epipod missing, detached during dissection); G, third maxilliped (right), lateral; H, third maxilliped (left), ultimate segment, mesial." httpUri="https://zenodo.org/record/196827/files/figure.png" pageId="4" pageNumber="41">Fig. 2</figureCitation>
D). First maxilliped with non-segmented endopod; caridean lobe of exopod moderately developed (
<figureCitation id="69EE2A30FFC4622BFB8BA26EFB21FCEE" box="[1081,1158,845,871]" captionStart="FIGURE 2" captionStartId="5.[151,258,1844,1868]" captionTargetBox="[187,1399,194,1843]" captionTargetId="figure@5.[151,1436,163,1857]" captionTargetPageId="5" captionText="FIGURE 2. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, mandible, mesial; B, same, incisor process and palp, lateral; C, maxillule, lateral; D, maxilla, lateral; E, first maxilliped, lateral; F, second maxilliped, lateral (epipod missing, detached during dissection); G, third maxilliped (right), lateral; H, third maxilliped (left), ultimate segment, mesial." httpUri="https://zenodo.org/record/196827/files/figure.png" pageId="4" pageNumber="41">Fig. 2</figureCitation>
E). Second maxilliped without specific features, as illustrated (
<figureCitation id="69EE2A30FFC4622BFD1DA253FCA6FC03" box="[687,769,880,906]" captionStart="FIGURE 2" captionStartId="5.[151,258,1844,1868]" captionTargetBox="[187,1399,194,1843]" captionTargetId="figure@5.[151,1436,163,1857]" captionTargetPageId="5" captionText="FIGURE 2. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, mandible, mesial; B, same, incisor process and palp, lateral; C, maxillule, lateral; D, maxilla, lateral; E, first maxilliped, lateral; F, second maxilliped, lateral (epipod missing, detached during dissection); G, third maxilliped (right), lateral; H, third maxilliped (left), ultimate segment, mesial." httpUri="https://zenodo.org/record/196827/files/figure.png" pageId="4" pageNumber="41">Fig. 2</figureCitation>
F). Third maxilliped with blunt lateral plate above mastigobranch; antepenultimate segment long, flattened lateroventrally, ventromesial margin setose; penultimate segment about twice as long as wide; ultimate segment tapering distally, with minute spine-like seta near blunt tip (
<figureCitation id="69EE2A30FFC4622BFE39A2FBFE71FC7B" box="[395,470,984,1010]" captionStart="FIGURE 2" captionStartId="5.[151,258,1844,1868]" captionTargetBox="[187,1399,194,1843]" captionTargetId="figure@5.[151,1436,163,1857]" captionTargetPageId="5" captionText="FIGURE 2. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, mandible, mesial; B, same, incisor process and palp, lateral; C, maxillule, lateral; D, maxilla, lateral; E, first maxilliped, lateral; F, second maxilliped, lateral (epipod missing, detached during dissection); G, third maxilliped (right), lateral; H, third maxilliped (left), ultimate segment, mesial." httpUri="https://zenodo.org/record/196827/files/figure.png" pageId="4" pageNumber="41">Fig. 2</figureCitation>
G, H), with transverse rows of finely serrulate setae dorsomesially (
<figureCitation id="69EE2A30FFC4622BFA9AA2FBFAD2FC7B" box="[1320,1397,984,1010]" captionStart="FIGURE 2" captionStartId="5.[151,258,1844,1868]" captionTargetBox="[187,1399,194,1843]" captionTargetId="figure@5.[151,1436,163,1857]" captionTargetPageId="5" captionText="FIGURE 2. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, mandible, mesial; B, same, incisor process and palp, lateral; C, maxillule, lateral; D, maxilla, lateral; E, first maxilliped, lateral; F, second maxilliped, lateral (epipod missing, detached during dissection); G, third maxilliped (right), lateral; H, third maxilliped (left), ultimate segment, mesial." httpUri="https://zenodo.org/record/196827/files/figure.png" pageId="4" pageNumber="41">Fig. 2</figureCitation>
H); arthrobranch present, small, composed of a few lamellae.
</paragraph>
<paragraph id="F16A36B5FFC4622BFF74A53EFB20FAE6" blockId="4.[151,1438,152,2015]" pageId="4" pageNumber="41">
First pereiopods (= chelipeds) somewhat polymorphic, especially in the development of chelae; ischium slender, with usually four slender spiniform setae on dorsal margin (
<figureCitation id="69EE2A30FFC4622BFC0AA563FBAFFBD3" box="[952,1032,1088,1114]" captionStart="FIGURE 3" captionStartId="6.[151,258,1517,1541]" captionTargetBox="[164,1430,187,1516]" captionTargetId="figure@6.[151,1436,172,1534]" captionTargetPageId="6" captionText="FIGURE 3. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, cheliped (right), lateral; B, same, row of spiniform setae on ventromesial margin of merus, mesial; C, same, chela, mesial; D, second pereiopod, lateral; E, third pereiopod, lateral; F, same, propodus and dactylus, lateral; G, fourth pereiopod, lateral; H, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/196828/files/figure.png" pageId="4" pageNumber="41">Figs. 3</figureCitation>
A, 4B); merus flattened on ventral side, ending bluntly distoventrally and distodorsally, ventromesial margin with row of five to ten slender spiniform setae (
<figureCitation id="69EE2A30FFC4622BFEEDA5A6FE16FB16" box="[351,433,1157,1183]" captionStart="FIGURE 3" captionStartId="6.[151,258,1517,1541]" captionTargetBox="[164,1430,187,1516]" captionTargetId="figure@6.[151,1436,172,1534]" captionTargetPageId="6" captionText="FIGURE 3. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, cheliped (right), lateral; B, same, row of spiniform setae on ventromesial margin of merus, mesial; C, same, chela, mesial; D, second pereiopod, lateral; E, third pereiopod, lateral; F, same, propodus and dactylus, lateral; G, fourth pereiopod, lateral; H, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/196828/files/figure.png" pageId="4" pageNumber="41">Figs. 3</figureCitation>
A, B, 4B); carpus variable in shape from subcylindrical, more than twice as long as wide (
<figureCitation id="69EE2A30FFC4622BFF50A58BFE8EFB4B" box="[226,297,1192,1218]" captionStart="FIGURE 3" captionStartId="6.[151,258,1517,1541]" captionTargetBox="[164,1430,187,1516]" captionTargetId="figure@6.[151,1436,172,1534]" captionTargetPageId="6" captionText="FIGURE 3. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, cheliped (right), lateral; B, same, row of spiniform setae on ventromesial margin of merus, mesial; C, same, chela, mesial; D, second pereiopod, lateral; E, third pereiopod, lateral; F, same, propodus and dactylus, lateral; G, fourth pereiopod, lateral; H, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/196828/files/figure.png" pageId="4" pageNumber="41">Fig. 3</figureCitation>
A), to vase-shaped, widening distally (
<figureCitation id="69EE2A30FFC4622BFD4EA58BFCE2FB4B" box="[764,837,1192,1218]" captionStart="FIGURE 4" captionStartId="7.[151,258,1508,1532]" captionTargetBox="[151,1424,188,1480]" captionTargetId="figure@7.[151,1436,178,1487]" captionTargetPageId="7" captionText="FIGURE 4. Metabetaeus mcphersonae n. sp., holotype, ovigerous female from Moorea (FLMNH UF Arthropoda 23247): A, frontal region, dorsal; B, major cheliped (right), lateral; C, same, chela, mesial; D, same, chela fingers; E, minor cheliped (left), chela, mesial; paratype, male from Moorea (FLMNH UF Arthropoda 23249): F, cheliped (right), chela, mesial; G, second pleopod, lateral." httpUri="https://zenodo.org/record/196829/files/figure.png" pageId="4" pageNumber="41">Fig. 4</figureCitation>
B); chela varying in proportions from slender, not swollen, subcylindrical, with palm about three times as long as high (
<figureCitation id="69EE2A30FFC4622BFC71A5E9FBAFFB6D" box="[963,1032,1226,1252]" captionStart="FIGURE 3" captionStartId="6.[151,258,1517,1541]" captionTargetBox="[164,1430,187,1516]" captionTargetId="figure@6.[151,1436,172,1534]" captionTargetPageId="6" captionText="FIGURE 3. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, cheliped (right), lateral; B, same, row of spiniform setae on ventromesial margin of merus, mesial; C, same, chela, mesial; D, second pereiopod, lateral; E, third pereiopod, lateral; F, same, propodus and dactylus, lateral; G, fourth pereiopod, lateral; H, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/196828/files/figure.png" pageId="4" pageNumber="41">Fig. 3</figureCitation>
A, C), to somewhat swollen, more ovoid, with palm twice or slightly more as long as high (
<figureCitation id="69EE2A30FFC4622BFC82A5CEFCDFFA8E" box="[816,888,1261,1287]" captionStart="FIGURE 4" captionStartId="7.[151,258,1508,1532]" captionTargetBox="[151,1424,188,1480]" captionTargetId="figure@7.[151,1436,178,1487]" captionTargetPageId="7" captionText="FIGURE 4. Metabetaeus mcphersonae n. sp., holotype, ovigerous female from Moorea (FLMNH UF Arthropoda 23247): A, frontal region, dorsal; B, major cheliped (right), lateral; C, same, chela, mesial; D, same, chela fingers; E, minor cheliped (left), chela, mesial; paratype, male from Moorea (FLMNH UF Arthropoda 23249): F, cheliped (right), chela, mesial; G, second pleopod, lateral." httpUri="https://zenodo.org/record/196829/files/figure.png" pageId="4" pageNumber="41">Fig. 4</figureCitation>
C, E, F); palm smooth, setose along dorsal and ventral margins; length ratio fingers to palm variable approximately from 
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non-enlarged chelae to 
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enlarged chelae; fingers unarmed in non-enlarged chelae (
<figureCitation id="69EE2A30FFC4622BFC67A411FBBBFAC5" box="[981,1052,1330,1356]" captionStart="FIGURE 3" captionStartId="6.[151,258,1517,1541]" captionTargetBox="[164,1430,187,1516]" captionTargetId="figure@6.[151,1436,172,1534]" captionTargetPageId="6" captionText="FIGURE 3. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, cheliped (right), lateral; B, same, row of spiniform setae on ventromesial margin of merus, mesial; C, same, chela, mesial; D, second pereiopod, lateral; E, third pereiopod, lateral; F, same, propodus and dactylus, lateral; G, fourth pereiopod, lateral; H, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/196828/files/figure.png" pageId="4" pageNumber="41">Fig. 3</figureCitation>
C) or armed with blunt teeth on proximal half of cutting edge in enlarged chelae (
<figureCitation id="69EE2A30FFC4622BFD6AA476FCB9FAE6" box="[728,798,1365,1391]" captionStart="FIGURE 4" captionStartId="7.[151,258,1508,1532]" captionTargetBox="[151,1424,188,1480]" captionTargetId="figure@7.[151,1436,178,1487]" captionTargetPageId="7" captionText="FIGURE 4. Metabetaeus mcphersonae n. sp., holotype, ovigerous female from Moorea (FLMNH UF Arthropoda 23247): A, frontal region, dorsal; B, major cheliped (right), lateral; C, same, chela, mesial; D, same, chela fingers; E, minor cheliped (left), chela, mesial; paratype, male from Moorea (FLMNH UF Arthropoda 23249): F, cheliped (right), chela, mesial; G, second pleopod, lateral." httpUri="https://zenodo.org/record/196829/files/figure.png" pageId="4" pageNumber="41">Fig. 4</figureCitation>
C–F), with elongate stiff setae.
</paragraph>
<paragraph id="F16A36B5FFC4622BFF74A45BFAE0F92E" blockId="4.[151,1438,152,2015]" pageId="4" pageNumber="41">
Second pereiopod long, slender, ischium and merus subequal in length; carpus five-segmented, with ratio of segments being approximately equal to 2: 1: 1.5: 0.7: 1.2; chela simple, with palm longer than fingers (
<figureCitation id="69EE2A30FFC4622BFF2DA49EFF41FA5E" box="[159,230,1469,1495]" captionStart="FIGURE 3" captionStartId="6.[151,258,1517,1541]" captionTargetBox="[164,1430,187,1516]" captionTargetId="figure@6.[151,1436,172,1534]" captionTargetPageId="6" captionText="FIGURE 3. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, cheliped (right), lateral; B, same, row of spiniform setae on ventromesial margin of merus, mesial; C, same, chela, mesial; D, second pereiopod, lateral; E, third pereiopod, lateral; F, same, propodus and dactylus, lateral; G, fourth pereiopod, lateral; H, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/196828/files/figure.png" pageId="4" pageNumber="41">Fig. 3</figureCitation>
D). Third pereiopod slender; ischium with two spiniform setae on ventrolateral surface; merus slightly more than six times as long as wide, with three spiniform setae on ventrolateral surface; carpus more slender than merus, unarmed; propodus more slender than carpus, with three to five spiniform setae on ventral margin; dactylus simple, conical, faintly curved, about 0.25 length of propodus, with slightly thickened setae inserted at about 0.7 of dactylar length. Fourth pereiopod very similar to third except for being more slender; merus with one or two spiniform setae on ventrolateral surface (
<figureCitation id="69EE2A30FFC4622BFC31A749FC60F90D" box="[899,967,1642,1668]" captionStart="FIGURE 3" captionStartId="6.[151,258,1517,1541]" captionTargetBox="[164,1430,187,1516]" captionTargetId="figure@6.[151,1436,172,1534]" captionTargetPageId="6" captionText="FIGURE 3. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, cheliped (right), lateral; B, same, row of spiniform setae on ventromesial margin of merus, mesial; C, same, chela, mesial; D, second pereiopod, lateral; E, third pereiopod, lateral; F, same, propodus and dactylus, lateral; G, fourth pereiopod, lateral; H, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/196828/files/figure.png" pageId="4" pageNumber="41">Fig. 3</figureCitation>
G). Fifth pereiopod as slender as fourth; ischium and merus unarmed; propodus with spiniform setae and some rows of setae distally (
<figureCitation id="69EE2A30FFC4622BFB69A7AEFA87F92E" box="[1243,1312,1677,1703]" captionStart="FIGURE 3" captionStartId="6.[151,258,1517,1541]" captionTargetBox="[164,1430,187,1516]" captionTargetId="figure@6.[151,1436,172,1534]" captionTargetPageId="6" captionText="FIGURE 3. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, cheliped (right), lateral; B, same, row of spiniform setae on ventromesial margin of merus, mesial; C, same, chela, mesial; D, second pereiopod, lateral; E, third pereiopod, lateral; F, same, propodus and dactylus, lateral; G, fourth pereiopod, lateral; H, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/196828/files/figure.png" pageId="4" pageNumber="41">Fig. 3</figureCitation>
H).
</paragraph>
<paragraph id="F16A36B5FFC4622BFF74A793FE4EF8FE" blockId="4.[151,1438,152,2015]" pageId="4" pageNumber="41">
First pleopod with endopod less than half-length of exopod (
<figureCitation id="69EE2A30FFC4622BFC73A793FBAEF943" box="[961,1033,1712,1738]" captionStart="FIGURE 1" captionStartId="3.[151,258,1713,1737]" captionTargetBox="[162,1422,654,1677]" captionTargetId="figure@3.[151,1436,634,1689]" captionTargetPageId="3" captionText="FIGURE 1. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, frontal region, dorsal; B, same, lateral; C, tooth on ventromesial carina of first segment of antennular peduncle, lateral; D, first pleopod with two embryos attached, lateral; E, second pleopod, lateral; F, uropod, dorsal; G, telson, dorsal." httpUri="https://zenodo.org/record/196826/files/figure.png" pageId="4" pageNumber="41">Fig. 1</figureCitation>
D); female second pleopod with appendix interna (
<figureCitation id="69EE2A30FFC4622BFEC8A7F1FE66F965" box="[378,449,1746,1772]" captionStart="FIGURE 1" captionStartId="3.[151,258,1713,1737]" captionTargetBox="[162,1422,654,1677]" captionTargetId="figure@3.[151,1436,634,1689]" captionTargetPageId="3" captionText="FIGURE 1. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, frontal region, dorsal; B, same, lateral; C, tooth on ventromesial carina of first segment of antennular peduncle, lateral; D, first pleopod with two embryos attached, lateral; E, second pleopod, lateral; F, uropod, dorsal; G, telson, dorsal." httpUri="https://zenodo.org/record/196826/files/figure.png" pageId="4" pageNumber="41">Fig. 1</figureCitation>
E); male second pleopod with appendix interna and appendix masculina, latter elongate, reaching beyond endopod, furnished with at least three strong setae (
<figureCitation id="69EE2A30FFC4622BFB8CA7D6FB21F886" box="[1086,1158,1781,1807]" captionStart="FIGURE 4" captionStartId="7.[151,258,1508,1532]" captionTargetBox="[151,1424,188,1480]" captionTargetId="figure@7.[151,1436,178,1487]" captionTargetPageId="7" captionText="FIGURE 4. Metabetaeus mcphersonae n. sp., holotype, ovigerous female from Moorea (FLMNH UF Arthropoda 23247): A, frontal region, dorsal; B, major cheliped (right), lateral; C, same, chela, mesial; D, same, chela fingers; E, minor cheliped (left), chela, mesial; paratype, male from Moorea (FLMNH UF Arthropoda 23249): F, cheliped (right), chela, mesial; G, second pleopod, lateral." httpUri="https://zenodo.org/record/196829/files/figure.png" pageId="4" pageNumber="41">Fig. 4</figureCitation>
G). Uropod with lateral lobe of protopod obscurely bifid, main ramus somewhat projecting, acute; exopod with diaeresis bearing large subtriangular tooth adjacent to strong, long distolateral spiniform seta; endopod with conspicuous tooth on distolateral margin (
<figureCitation id="69EE2A30FFC4622BFE31A67EFE62F8FE" box="[387,453,1885,1911]" captionStart="FIGURE 1" captionStartId="3.[151,258,1713,1737]" captionTargetBox="[162,1422,654,1677]" captionTargetId="figure@3.[151,1436,634,1689]" captionTargetPageId="3" captionText="FIGURE 1. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, frontal region, dorsal; B, same, lateral; C, tooth on ventromesial carina of first segment of antennular peduncle, lateral; D, first pleopod with two embryos attached, lateral; E, second pleopod, lateral; F, uropod, dorsal; G, telson, dorsal." httpUri="https://zenodo.org/record/196826/files/figure.png" pageId="4" pageNumber="41">Fig. 1</figureCitation>
F).
</paragraph>
<paragraph id="F16A36B5FFC4622BFF74A65CFBE6F856" blockId="4.[151,1438,152,2015]" pageId="4" pageNumber="41">
<emphasis id="C3A1EAA7FFC4622BFF74A65CFE23F810" bold="true" box="[198,388,1919,1945]" pageId="4" pageNumber="41">Colour pattern</emphasis>
. Semitransparent with reddish chromatophores forming more or less distinct bands, two on the carapace and six on the abdomen; scattered red chromatophores also on antennular peduncles, chelipeds, walking legs and tail fan; eggs / embryos dark brown-red (
<figureCitation id="69EE2A30FFC4622BFC73A6E6FB94F856" box="[961,1075,1989,2015]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="8.[151,258,1948,1972]" captionStartId-1="9.[151,258,1942,1966]" captionStartId-2="10.[151,258,1613,1637]" captionTargetBox-0="[151,1436,194,1926]" captionTargetBox-1="[151,1436,194,1919]" captionTargetBox-2="[151,1436,194,1590]" captionTargetId-0="figure@8.[151,1436,194,1926]" captionTargetId-1="figure@9.[151,1436,194,1921]" captionTargetId-2="figure@10.[151,1436,194,1592]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURE 5. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, habitus, lateral; B, same, dorsal." captionText-1="FIGURE 6. Metabetaeus mcphersonae n. sp., holotype, ovigerous female from Moorea (FLMNH UF Arthropoda 23247): A, habitus, lateral; B, same, dorsal." captionText-2="FIGURE 7. Metabetaeus mcphersonae n. sp., holotype, ovigerous female from Moorea (FLMNH UF Arthropoda 23247): A, frontal region, dorsal; B, major cheliped (right), chela, mesial (= dorsal); paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): C, anterior region, lateral; D, chelipeds, lateral (= ventral); E, uropod, dorsal; paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23248): F, embryos attached to pleopods; G, zoea." httpUri-0="https://zenodo.org/record/196830/files/figure.png" httpUri-1="https://zenodo.org/record/196831/files/figure.png" httpUri-2="https://zenodo.org/record/196832/files/figure.png" pageId="4" pageNumber="41">Figs. 5–7</figureCitation>
).
</paragraph>
<caption id="A5AA663DFFC5622AFF25A617FCA9F822" httpUri="https://zenodo.org/record/196827/files/figure.png" pageId="5" pageNumber="42" targetBox="[187,1399,194,1843]" targetPageId="5">
<paragraph id="F16A36B5FFC5622AFF25A617FCA9F822" blockId="5.[151,1437,1844,1963]" pageId="5" pageNumber="42">
<emphasis id="C3A1EAA7FFC5622AFF25A617FEBFF8C5" bold="true" box="[151,280,1844,1868]" pageId="5" pageNumber="42">FIGURE 2</emphasis>
. 
<taxonomicName id="36D54D36FFC5622AFE95A616FDF6F8C5" box="[295,593,1845,1868]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="mcphersonae" status="sp. nov.">
<emphasis id="C3A1EAA7FFC5622AFE95A616FDF6F8C5" box="[295,593,1845,1868]" italics="true" pageId="5" pageNumber="42">Metabetaeus mcphersonae</emphasis>
</taxonomicName>
<emphasis id="C3A1EAA7FFC5622AFDE8A617FD3EF8C5" bold="true" box="[602,665,1844,1868]" pageId="5" pageNumber="42">
<taxonomicNameLabel id="D89257DCFFC5622AFDE8A617FD3EF8C5" box="[602,665,1844,1868]" pageId="5" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, mandible, mesial; B, same, incisor process and palp, lateral; C, maxillule, lateral; D, maxilla, lateral; E, first maxilliped, lateral; F, second maxilliped, lateral (epipod missing, detached during dissection); G, third maxilliped (right), lateral; H, third maxilliped (left), ultimate segment, mesial.
</paragraph>
</caption>
<caption id="A5AA663DFFC66229FF25A4CEFDD0F9ED" httpUri="https://zenodo.org/record/196828/files/figure.png" pageId="6" pageNumber="43" targetBox="[164,1430,187,1516]" targetPageId="6">
<paragraph id="F16A36B5FFC66229FF25A4CEFDD0F9ED" blockId="6.[151,1437,1517,1636]" pageId="6" pageNumber="43">
<emphasis id="C3A1EAA7FFC66229FF25A4CEFEBFF98C" bold="true" box="[151,280,1517,1541]" pageId="6" pageNumber="43">FIGURE 3</emphasis>
. 
<taxonomicName id="36D54D36FFC66229FE95A4CDFDF6F98C" box="[295,593,1518,1541]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="mcphersonae" status="sp. nov.">
<emphasis id="C3A1EAA7FFC66229FE95A4CDFDF6F98C" box="[295,593,1518,1541]" italics="true" pageId="6" pageNumber="43">Metabetaeus mcphersonae</emphasis>
</taxonomicName>
<emphasis id="C3A1EAA7FFC66229FDE8A4CEFD3EF98C" bold="true" box="[602,665,1517,1541]" pageId="6" pageNumber="43">
<taxonomicNameLabel id="D89257DCFFC66229FDE8A4CEFD3EF98C" box="[602,665,1517,1541]" pageId="6" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, cheliped (right), lateral; B, same, row of spiniform setae on ventromesial margin of merus, mesial; C, same, chela, mesial; D, second pereiopod, lateral; E, third pereiopod, lateral; F, same, propodus and dactylus, lateral; G, fourth pereiopod, lateral; H, fifth pereiopod, lateral.
</paragraph>
</caption>
<paragraph id="F16A36B5FFC66229FF74A781FB05F934" blockId="6.[151,1436,1698,2037]" box="[198,1186,1698,1725]" pageId="6" pageNumber="43">
<emphasis id="C3A1EAA7FFC66229FF74A781FEEFF935" bold="true" box="[198,328,1698,1724]" pageId="6" pageNumber="43">Size range</emphasis>
. Females at cl 2.4–3.0 mm, tl 
<quantity id="362D9B50FFC66229FD19A780FC9CF934" box="[683,827,1699,1725]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" metricValueMax="8.3" metricValueMin="7.1" pageId="6" pageNumber="43" unit="mm" value="7.7" valueMax="8.3" valueMin="7.1">7.1–8.3 mm</quantity>
; male at cl 
<quantity id="362D9B50FFC66229FC72A780FBBAF934" box="[960,1053,1699,1725]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="43" unit="mm" value="2.1">2.1 mm</quantity>
, tl 
<quantity id="362D9B50FFC66229FBF2A780FB39F934" box="[1088,1182,1699,1725]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="6" pageNumber="43" unit="mm" value="6.3">6.3 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="B9CF653EFFC66229FF74A7E6FE6DF8AC" pageId="6" pageNumber="43" type="etymology">
<paragraph id="F16A36B5FFC66229FF74A7E6FA22F969" blockId="6.[151,1436,1698,2037]" box="[198,1413,1733,1760]" pageId="6" pageNumber="43">
<emphasis id="C3A1EAA7FFC66229FF74A7E6FEECF956" bold="true" box="[198,331,1733,1759]" pageId="6" pageNumber="43">Etymology</emphasis>
. This very interesting new species is named after its collector, Sarah McPherson (FLMNH).
</paragraph>
<paragraph id="F16A36B5FFC66229FF74A7CBFE6DF8AC" blockId="6.[151,1436,1698,2037]" pageId="6" pageNumber="43">
<emphasis id="C3A1EAA7FFC66229FF74A7CBFE8FF88B" bold="true" box="[198,296,1768,1794]" pageId="6" pageNumber="43">Ecology</emphasis>
. All specimens were collected by digging and fanning of coarse sand and rubble on a fore-reef, at a depth range of 
<quantity id="362D9B50FFC66229FEE9A628FE61F8AC" box="[347,454,1803,1829]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="6" pageNumber="43" unit="m" value="15.0" valueMax="17.0" valueMin="13.0">
13– 
<specimenCount id="E7D3FD3CFFC66229FE3AA628FE61F8AC" box="[392,454,1803,1829]" pageId="6" pageNumber="43" type="male">17 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="B9CF653EFFC66229FF74A60EFD32F839" pageId="6" pageNumber="43" type="description">
<paragraph id="F16A36B5FFC66229FF74A60EFD32F839" blockId="6.[151,1436,1698,2037]" pageId="6" pageNumber="43">
<emphasis id="C3A1EAA7FFC66229FF74A60EFE2AF8CE" bold="true" box="[198,397,1837,1863]" pageId="6" pageNumber="43">Biological notes</emphasis>
. All three ovigerous females of 
<taxonomicName id="36D54D36FFC66229FCA9A60DFC46F8CE" box="[795,993,1838,1863]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="43" phylum="Arthropoda" rank="species" species="mcphersonae" status="sp. nov.">
<emphasis id="C3A1EAA7FFC66229FCA9A60DFC46F8CE" box="[795,993,1838,1863]" italics="true" pageId="6" pageNumber="43">M. mcphersonae</emphasis>
</taxonomicName>
<emphasis id="C3A1EAA7FFC66229FC5EA60EFB95F8CE" bold="true" box="[1004,1074,1837,1863]" pageId="6" pageNumber="43">
<taxonomicNameLabel id="D89257DCFFC66229FC5EA60EFB95F8CE" box="[1004,1074,1837,1863]" pageId="6" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<typeStatus id="2E6E8817FFC66229FBF6A60DFB0AF8C1" box="[1092,1197,1838,1864]" pageId="6" pageNumber="43" type="holotype">holotype</typeStatus>
and two 
<typeStatus id="2E6E8817FFC66229FA90A60DFA33F8C1" box="[1314,1428,1838,1864]" pageId="6" pageNumber="43" type="paratype">paratypes</typeStatus>
) were carrying a low number (4–18) of large embryos (relative to body size, see 
<figureCitation id="69EE2A30FFC66229FBD1A673FB11F8E3" box="[1123,1206,1872,1898]" captionStart="FIGURE 5" captionStartId="8.[151,258,1948,1972]" captionTargetBox="[151,1436,194,1926]" captionTargetId="figure@8.[151,1436,194,1926]" captionTargetPageId="8" captionText="FIGURE 5. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, habitus, lateral; B, same, dorsal." httpUri="https://zenodo.org/record/196830/files/figure.png" pageId="6" pageNumber="43">Figs. 5</figureCitation>
A, 6A; diameter of embryos ~ 
<quantity id="362D9B50FFC66229FE91A650FE1CF804" box="[291,443,1907,1933]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="6" pageNumber="43" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.6–0.8 mm</quantity>
, see 
<figureCitation id="69EE2A30FFC66229FE4AA650FD98F804" box="[504,575,1907,1933]" captionStart="FIGURE 1" captionStartId="3.[151,258,1713,1737]" captionTargetBox="[162,1422,654,1677]" captionTargetId="figure@3.[151,1436,634,1689]" captionTargetPageId="3" captionText="FIGURE 1. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, frontal region, dorsal; B, same, lateral; C, tooth on ventromesial carina of first segment of antennular peduncle, lateral; D, first pleopod with two embryos attached, lateral; E, second pleopod, lateral; F, uropod, dorsal; G, telson, dorsal." httpUri="https://zenodo.org/record/196826/files/figure.png" pageId="6" pageNumber="43">Fig. 1</figureCitation>
D). Some embryos hatched into relatively advanced zoeae (
<figureCitation id="69EE2A30FFC66229FA9BA650FAD2F804" box="[1321,1397,1907,1933]" captionStart="FIGURE 7" captionStartId="10.[151,258,1613,1637]" captionTargetBox="[151,1436,194,1590]" captionTargetId="figure@10.[151,1436,194,1592]" captionTargetPageId="10" captionText="FIGURE 7. Metabetaeus mcphersonae n. sp., holotype, ovigerous female from Moorea (FLMNH UF Arthropoda 23247): A, frontal region, dorsal; B, major cheliped (right), chela, mesial (= dorsal); paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): C, anterior region, lateral; D, chelipeds, lateral (= ventral); E, uropod, dorsal; paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23248): F, embryos attached to pleopods; G, zoea." httpUri="https://zenodo.org/record/196832/files/figure.png" pageId="6" pageNumber="43">Fig. 7</figureCitation>
G), suggesting abbreviated larval development.
</paragraph>
</subSubSection>
<subSubSection id="B9CF653EFFC66229FF74A69BFCD7F85B" box="[198,880,1976,2002]" pageId="6" pageNumber="43" type="materials_examined">
<paragraph id="F16A36B5FFC66229FF74A69BFCD7F85B" blockId="6.[151,1436,1698,2037]" box="[198,880,1976,2002]" pageId="6" pageNumber="43">
<emphasis id="C3A1EAA7FFC66229FF74A69BFEC3F85B" bold="true" box="[198,356,1976,2002]" pageId="6" pageNumber="43">
<typeStatus id="2E6E8817FFC66229FF74A69BFEA3F85B" box="[198,260,1976,2002]" pageId="6" pageNumber="43">Type</typeStatus>
locality
</emphasis>
. Moorea, Society Islands, French Polynesia.
</paragraph>
</subSubSection>
<subSubSection id="B9CF653EFFC66229FF74A6F9FAA5F87C" box="[198,1282,2010,2037]" pageId="6" pageNumber="43" type="distribution">
<paragraph id="F16A36B5FFC66229FF74A6F9FAA5F87C" blockId="6.[151,1436,1698,2037]" box="[198,1282,2010,2037]" pageId="6" pageNumber="43">
<emphasis id="C3A1EAA7FFC66229FF74A6F9FEF8F87D" bold="true" box="[198,351,2010,2036]" pageId="6" pageNumber="43">Distribution</emphasis>
. Presently known only from the 
<typeStatus id="2E6E8817FFC66229FD52A6F8FCB5F87C" box="[736,786,2011,2037]" pageId="6" pageNumber="43">type</typeStatus>
locality in French Polynesia (see 
<figureCitation id="69EE2A30FFC66229FB2DA6F8FB53F87C" box="[1183,1268,2011,2037]" captionStart="FIGURE 11" captionStartId="15.[151,255,1405,1429]" captionTargetBox="[151,1434,194,1383]" captionTargetId="figure@15.[151,1436,194,1384]" captionTargetPageId="15" captionText="FIGURE 11. Metabetaeus minutus (Whitelegge, 1897), male from Runaway Cave, Christmas Island (ZRC 2010.0013): A, habitus, lateral. Metabetaeus lohena Banner &amp; Banner, 1960 from Hawai’i (not deposited): B, C, two individuals from anchialine pond at Kona, Kaloko-Honokohau National Park, Hawai’i, showing different state of contraction of chromatophores (photographs: Peter K. L. Ng [A]; Karl Magnacca [B, C])." httpUri="https://zenodo.org/record/196836/files/figure.png" pageId="6" pageNumber="43">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="A5AA663DFFC76228FF25A4C7FDECF9D5" httpUri="https://zenodo.org/record/196829/files/figure.png" pageId="7" pageNumber="44" targetBox="[151,1424,188,1480]" targetPageId="7">
<paragraph id="F16A36B5FFC76228FF25A4C7FDECF9D5" blockId="7.[151,1437,1508,1628]" pageId="7" pageNumber="44">
<emphasis id="C3A1EAA7FFC76228FF25A4C7FEBFFA75" bold="true" box="[151,280,1508,1532]" pageId="7" pageNumber="44">FIGURE 4</emphasis>
. 
<taxonomicName id="36D54D36FFC76228FE95A4C6FDF6FA75" box="[295,593,1509,1532]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="mcphersonae" status="sp. nov.">
<emphasis id="C3A1EAA7FFC76228FE95A4C6FDF6FA75" box="[295,593,1509,1532]" italics="true" pageId="7" pageNumber="44">Metabetaeus mcphersonae</emphasis>
</taxonomicName>
<emphasis id="C3A1EAA7FFC76228FDEBA4C7FD3EFA75" bold="true" box="[601,665,1508,1532]" pageId="7" pageNumber="44">
<taxonomicNameLabel id="D89257DCFFC76228FDEBA4C7FD3EFA75" box="[601,665,1508,1532]" pageId="7" pageNumber="44" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, ovigerous female from Moorea (FLMNH UF Arthropoda 23247): A, frontal region, dorsal; B, major cheliped (right), lateral; C, same, chela, mesial; D, same, chela fingers; E, minor cheliped (left), chela, mesial; paratype, male from Moorea (FLMNH UF Arthropoda 23249): F, cheliped (right), chela, mesial; G, second pleopod, lateral.
</paragraph>
</caption>
<subSubSection id="B9CF653EFFC76224FF74A7A9FAF6FD35" lastPageId="11" lastPageNumber="48" pageId="7" pageNumber="44" type="discussion">
<paragraph id="F16A36B5FFC76225FF74A7A9FB59F864" blockId="7.[151,1437,1674,1999]" lastBlockId="10.[151,1437,1779,2029]" lastPageId="10" lastPageNumber="47" pageId="7" pageNumber="44">
<emphasis id="C3A1EAA7FFC76228FF74A7A9FE9FF92D" bold="true" box="[198,312,1674,1700]" pageId="7" pageNumber="44">Remarks</emphasis>
. 
<taxonomicName id="36D54D36FFC76228FEF8A7A9FD21F92A" box="[330,646,1674,1699]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="mcphersonae" status="sp. nov.">
<emphasis id="C3A1EAA7FFC76228FEF8A7A9FD21F92A" box="[330,646,1674,1699]" italics="true" pageId="7" pageNumber="44">Metabetaeus mcphersonae</emphasis>
</taxonomicName>
<emphasis id="C3A1EAA7FFC76228FD23A7A9FD71F92D" bold="true" box="[657,726,1674,1700]" pageId="7" pageNumber="44">
<taxonomicNameLabel id="D89257DCFFC76228FD23A7A9FD71F92D" box="[657,726,1674,1700]" pageId="7" pageNumber="44" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from both 
<taxonomicName id="36D54D36FFC76228FC0BA7A9FBE7F92A" box="[953,1088,1674,1699]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="C3A1EAA7FFC76228FC0BA7A9FBE7F92A" box="[953,1088,1674,1699]" italics="true" pageId="7" pageNumber="44">M. minutus</emphasis>
</taxonomicName>
and 
<taxonomicName id="36D54D36FFC76228FBCDA7A9FB5EF92A" box="[1151,1273,1674,1699]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="lohena">
<emphasis id="C3A1EAA7FFC76228FBCDA7A9FB5EF92A" box="[1151,1273,1674,1699]" italics="true" pageId="7" pageNumber="44">M. lohena</emphasis>
</taxonomicName>
by the better developed cornea (compare 
<figureCitation id="69EE2A30FFC76228FE4BA793FDEAF943" box="[505,589,1712,1738]" captionStart="FIGURE 1" captionStartId="3.[151,258,1713,1737]" captionTargetBox="[162,1422,654,1677]" captionTargetId="figure@3.[151,1436,634,1689]" captionTargetPageId="3" captionText="FIGURE 1. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, frontal region, dorsal; B, same, lateral; C, tooth on ventromesial carina of first segment of antennular peduncle, lateral; D, first pleopod with two embryos attached, lateral; E, second pleopod, lateral; F, uropod, dorsal; G, telson, dorsal." httpUri="https://zenodo.org/record/196826/files/figure.png" pageId="7" pageNumber="44">Figs. 1</figureCitation>
A, 4A, 8A; see also 
<bibRefCitation id="95444B44FFC76228FCE3A793FBDFF943" author="Banner" box="[849,1144,1712,1738]" pageId="7" pageNumber="54" refString="Banner, A. H. &amp; Banner, D. M. (1960) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean, part VII: On Metabetaeus Borradaile, with a new species from Hawaii. Pacific Science, 15, 299 - 303." type="journal article" year="1960">Banner &amp; Banner 1960</bibRefCitation>
, figs. 1a, 2a); the more conspicuous distolateral tooth on the uropodal endopod (compare 
<figureCitation id="69EE2A30FFC76228FC12A7F6FC4AF966" box="[928,1005,1749,1775]" captionStart="FIGURE 1" captionStartId="3.[151,258,1713,1737]" captionTargetBox="[162,1422,654,1677]" captionTargetId="figure@3.[151,1436,634,1689]" captionTargetPageId="3" captionText="FIGURE 1. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, frontal region, dorsal; B, same, lateral; C, tooth on ventromesial carina of first segment of antennular peduncle, lateral; D, first pleopod with two embryos attached, lateral; E, second pleopod, lateral; F, uropod, dorsal; G, telson, dorsal." httpUri="https://zenodo.org/record/196826/files/figure.png" pageId="7" pageNumber="44">Figs. 1</figureCitation>
F, 8F); the propodus of the third and fourth pereiopods armed with only a few spiniform setae (vs. numerous spiniform setae in 
<taxonomicName id="36D54D36FFC76228FB0DA7D9FAE4F89A" box="[1215,1347,1786,1811]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="C3A1EAA7FFC76228FB0DA7D9FAE4F89A" box="[1215,1347,1786,1811]" italics="true" pageId="7" pageNumber="44">M. minutus</emphasis>
</taxonomicName>
and 
<taxonomicName id="36D54D36FFC76228FACEA7D9FF40F8B0" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="lohena">
<emphasis id="C3A1EAA7FFC76228FACEA7D9FF40F8B0" italics="true" pageId="7" pageNumber="44">M. lohena</emphasis>
</taxonomicName>
; compare 
<figureCitation id="69EE2A30FFC76228FED0A603FE16F8B3" box="[354,433,1824,1850]" captionStart="FIGURE 3" captionStartId="6.[151,258,1517,1541]" captionTargetBox="[164,1430,187,1516]" captionTargetId="figure@6.[151,1436,172,1534]" captionTargetPageId="6" captionText="FIGURE 3. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, cheliped (right), lateral; B, same, row of spiniform setae on ventromesial margin of merus, mesial; C, same, chela, mesial; D, second pereiopod, lateral; E, third pereiopod, lateral; F, same, propodus and dactylus, lateral; G, fourth pereiopod, lateral; H, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/196828/files/figure.png" pageId="7" pageNumber="44">Figs. 3</figureCitation>
E, G, 8D); and the presence of a small arthrobranch at the base of the third maxilliped (absent in 
<taxonomicName id="36D54D36FFC76228FEA4A666FE3AF8D7" box="[278,413,1861,1886]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="C3A1EAA7FFC76228FEA4A666FE3AF8D7" box="[278,413,1861,1886]" italics="true" pageId="7" pageNumber="44">M. minutus</emphasis>
</taxonomicName>
and 
<taxonomicName id="36D54D36FFC76228FE6BA666FDF4F8D7" box="[473,595,1861,1886]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="lohena">
<emphasis id="C3A1EAA7FFC76228FE6BA666FDF4F8D7" box="[473,595,1861,1886]" italics="true" pageId="7" pageNumber="44">M. lohena</emphasis>
</taxonomicName>
). In addition, in most specimens of 
<taxonomicName id="36D54D36FFC76228FBB9A666FB74F8D7" box="[1035,1235,1861,1886]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="mcphersonae" status="sp. nov.">
<emphasis id="C3A1EAA7FFC76228FBB9A666FB74F8D7" box="[1035,1235,1861,1886]" italics="true" pageId="7" pageNumber="44">M. mcphersonae</emphasis>
</taxonomicName>
<emphasis id="C3A1EAA7FFC76228FB69A666FA87F8D6" bold="true" box="[1243,1312,1861,1887]" pageId="7" pageNumber="44">
<taxonomicNameLabel id="D89257DCFFC76228FB69A666FA87F8D6" box="[1243,1312,1861,1887]" pageId="7" pageNumber="44" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, the chela palm is stouter and more swollen, while the cutting edges of finger have a fairly strong proximal dentition (
<figureCitation id="69EE2A30FFC76228FF2DA6B3FF41F823" box="[159,230,1936,1962]" captionStart="FIGURE 4" captionStartId="7.[151,258,1508,1532]" captionTargetBox="[151,1424,188,1480]" captionTargetId="figure@7.[151,1436,178,1487]" captionTargetPageId="7" captionText="FIGURE 4. Metabetaeus mcphersonae n. sp., holotype, ovigerous female from Moorea (FLMNH UF Arthropoda 23247): A, frontal region, dorsal; B, major cheliped (right), lateral; C, same, chela, mesial; D, same, chela fingers; E, minor cheliped (left), chela, mesial; paratype, male from Moorea (FLMNH UF Arthropoda 23249): F, cheliped (right), chela, mesial; G, second pleopod, lateral." httpUri="https://zenodo.org/record/196829/files/figure.png" pageId="7" pageNumber="44">Fig. 4</figureCitation>
D); in 
<taxonomicName id="36D54D36FFC76228FE82A6B3FE14F820" box="[304,435,1936,1961]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="C3A1EAA7FFC76228FE82A6B3FE14F820" box="[304,435,1936,1961]" italics="true" pageId="7" pageNumber="44">M. minutus</emphasis>
</taxonomicName>
and 
<taxonomicName id="36D54D36FFC76228FE5EA6B3FDC5F820" box="[492,610,1936,1961]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="lohena">
<emphasis id="C3A1EAA7FFC76228FE5EA6B3FDC5F820" box="[492,610,1936,1961]" italics="true" pageId="7" pageNumber="44">M. lohena</emphasis>
</taxonomicName>
, the palm is more slender, while the cutting edges are either straight or bearing minute teeth proximally (
<figureCitation id="69EE2A30FFC76228FDBEA696FDFFF846" box="[524,600,1973,1999]" captionStart="FIGURE 9" captionStartId="13.[151,255,1896,1920]" captionTargetBox="[215,1390,417,1862]" captionTargetId="figure@13.[206,1401,369,1873]" captionTargetPageId="13" captionText="FIGURE 9. Metabetaeus minutus (Whitelegge, 1897), male from Runaway Cave, Christmas Island (ZRC 2010.0013): A, frontal margin of carapace, dorsal; B, frontal region of carapace and frontal appendages, lateral; C, left cheliped, lateral; D, same, chela, mesial; E, second pereiopod, lateral; F, third or fourth pereiopod (detached), lateral; G, fifth pereiopod, lateral." httpUri="https://zenodo.org/record/196834/files/figure.png" pageId="7" pageNumber="44">Figs. 9</figureCitation>
D, 10B, D). Furthermore, in 
<taxonomicName id="36D54D36FFC76228FC19A696FBBBF847" box="[939,1052,1973,1998]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="lohena">
<emphasis id="C3A1EAA7FFC76228FC19A696FBBBF847" box="[939,1052,1973,1998]" italics="true" pageId="7" pageNumber="44">M. lohena</emphasis>
</taxonomicName>
, the fingers are more slender and elongate, usually longer than the palm, and somewhat twisted (
<figureCitation id="69EE2A30FFCA6225FC18A7D0FBA0F884" box="[938,1031,1779,1805]" captionStart="FIGURE 10" captionStartId="14.[151,255,1946,1970]" captionTargetBox="[175,1397,806,1929]" captionTargetId="figure@14.[175,1411,805,1933]" captionTargetPageId="14" captionText="FIGURE 10. Metabetaeus lohena Banner &amp; Banner, 1960, female from Rapa Nui (OUMNH. ZC- 2010 - 20 - 009): A, right cheliped, lateral; B, same, chela, mesial; paratype from Hawaii: C, left cheliped, lateral; D, same, chela, mesial (C, D after Banner &amp; Banner 1960)." httpUri="https://zenodo.org/record/196835/files/figure.png" pageId="10" pageNumber="47">Fig. 10</figureCitation>
). The length of the stylocerite is apparently variable in both 
<taxonomicName id="36D54D36FFCA6225FE69A63AFDC7F8BB" box="[475,608,1817,1842]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="C3A1EAA7FFCA6225FE69A63AFDC7F8BB" box="[475,608,1817,1842]" italics="true" pageId="10" pageNumber="47">M. minutus</emphasis>
</taxonomicName>
and 
<taxonomicName id="36D54D36FFCA6225FD2BA63AFBE6F8BB" authority="Banner &amp; Banner 1983" authorityName="Banner &amp; Banner" authorityYear="1983" box="[665,1089,1816,1842]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="lohena">
<emphasis id="C3A1EAA7FFCA6225FD2BA63AFCB6F8BB" box="[665,785,1817,1842]" italics="true" pageId="10" pageNumber="47">M. lohena</emphasis>
(
<bibRefCitation id="95444B44FFCA6225FC92A63BFB9EF8BB" author="Banner" box="[800,1081,1816,1842]" pageId="10" pageNumber="54" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner 1983</bibRefCitation>
)
</taxonomicName>
; nevertheless, in all available specimens of 
<taxonomicName id="36D54D36FFCA6225FE88A61DFE58F8DE" box="[314,511,1854,1879]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="mcphersonae" status="sp. nov.">
<emphasis id="C3A1EAA7FFCA6225FE88A61DFE58F8DE" box="[314,511,1854,1879]" italics="true" pageId="10" pageNumber="47">M. mcphersonae</emphasis>
</taxonomicName>
<emphasis id="C3A1EAA7FFCA6225FDB5A61EFDEDF8DE" bold="true" box="[519,586,1853,1879]" pageId="10" pageNumber="47">
<taxonomicNameLabel id="D89257DCFFCA6225FDB5A61EFDEDF8DE" box="[519,586,1853,1879]" pageId="10" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, the stylocerite always exceeds the distal margin of the first segment of the antennular peduncle, reaching to at least 0.2 of the second segment (
<figureCitation id="69EE2A30FFCA6225FC47A640FB9FF8F4" box="[1013,1080,1891,1917]" captionStart="FIGURE 1" captionStartId="3.[151,258,1713,1737]" captionTargetBox="[162,1422,654,1677]" captionTargetId="figure@3.[151,1436,634,1689]" captionTargetPageId="3" captionText="FIGURE 1. Metabetaeus mcphersonae n. sp., paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, frontal region, dorsal; B, same, lateral; C, tooth on ventromesial carina of first segment of antennular peduncle, lateral; D, first pleopod with two embryos attached, lateral; E, second pleopod, lateral; F, uropod, dorsal; G, telson, dorsal." httpUri="https://zenodo.org/record/196826/files/figure.png" pageId="10" pageNumber="47">Fig. 1</figureCitation>
A, 4A). The new species also differs from 
<taxonomicName id="36D54D36FFCA6225FE9EA6AAFE16F82B" box="[300,433,1929,1954]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="C3A1EAA7FFCA6225FE9EA6AAFE16F82B" box="[300,433,1929,1954]" italics="true" pageId="10" pageNumber="47">M. minutus</emphasis>
</taxonomicName>
and 
<taxonomicName id="36D54D36FFCA6225FE5EA6AAFDC4F82B" box="[492,611,1929,1954]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="lohena">
<emphasis id="C3A1EAA7FFCA6225FE5EA6AAFDC4F82B" box="[492,611,1929,1954]" italics="true" pageId="10" pageNumber="47">M. lohena</emphasis>
</taxonomicName>
by its much smaller size (maximum cl 
<quantity id="362D9B50FFCA6225FB8AA6ABFBDAF82B" box="[1080,1149,1928,1954]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="47" unit="mm" value="3.0">3 mm</quantity>
vs. cl 
<quantity id="362D9B50FFCA6225FB7EA6ABFAA8F82B" box="[1228,1295,1928,1954]" metricMagnitude="-1" metricUnit="m" metricValue="1.4731999999999998" pageId="10" pageNumber="47" unit="in" value="5.8">5.8 in</quantity>
<taxonomicName id="36D54D36FFCA6225FAA5A6AAFA3BF82B" box="[1303,1436,1929,1954]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="C3A1EAA7FFCA6225FAA5A6AAFA3BF82B" box="[1303,1436,1929,1954]" italics="true" pageId="10" pageNumber="47">M. minutus</emphasis>
</taxonomicName>
and cl ~6.0 in 
<taxonomicName id="36D54D36FFCA6225FEF0A68DFE1EF84E" box="[322,441,1966,1991]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="lohena">
<emphasis id="C3A1EAA7FFCA6225FEF0A68DFE1EF84E" box="[322,441,1966,1991]" italics="true" pageId="10" pageNumber="47">M. lohena</emphasis>
</taxonomicName>
), and ecologically by its association with marine coral reef habitats rather than with land-locked anchialine pools and caves, which are typical habitats of 
<taxonomicName id="36D54D36FFCA6225FC76A6F0FBEFF865" box="[964,1096,2003,2028]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="C3A1EAA7FFCA6225FC76A6F0FBEFF865" box="[964,1096,2003,2028]" italics="true" pageId="10" pageNumber="47">M. minutus</emphasis>
</taxonomicName>
and 
<taxonomicName id="36D54D36FFCA6225FB32A6F0FB50F865" box="[1152,1271,2003,2028]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="lohena">
<emphasis id="C3A1EAA7FFCA6225FB32A6F0FB50F865" box="[1152,1271,2003,2028]" italics="true" pageId="10" pageNumber="47">M. lohena</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="A5AA663DFFC86227FF25A6BFFDCFF85A" httpUri="https://zenodo.org/record/196830/files/figure.png" pageId="8" pageNumber="45" targetBox="[151,1436,194,1926]" targetPageId="8">
<paragraph id="F16A36B5FFC86227FF25A6BFFDCFF85A" blockId="8.[151,1437,1948,2003]" pageId="8" pageNumber="45">
<emphasis id="C3A1EAA7FFC86227FF25A6BFFEBFF83D" bold="true" box="[151,280,1948,1972]" pageId="8" pageNumber="45">FIGURE 5</emphasis>
. 
<taxonomicName id="36D54D36FFC86227FE95A6BFFDF6F83A" box="[295,593,1948,1971]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="45" phylum="Arthropoda" rank="species" species="mcphersonae" status="sp. nov.">
<emphasis id="C3A1EAA7FFC86227FE95A6BFFDF6F83A" box="[295,593,1948,1971]" italics="true" pageId="8" pageNumber="45">Metabetaeus mcphersonae</emphasis>
</taxonomicName>
<emphasis id="C3A1EAA7FFC86227FDE8A6BFFD3EF83D" bold="true" box="[602,665,1948,1972]" pageId="8" pageNumber="45">
<taxonomicNameLabel id="D89257DCFFC86227FDE8A6BFFD3EF83D" box="[602,665,1948,1972]" pageId="8" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): A, habitus, lateral; B, same, dorsal.
</paragraph>
</caption>
<caption id="A5AA663DFFC96226FF25A6B5FDCFF844" httpUri="https://zenodo.org/record/196831/files/figure.png" pageId="9" pageNumber="46" targetBox="[151,1436,194,1919]" targetPageId="9">
<paragraph id="F16A36B5FFC96226FF25A6B5FDCFF844" blockId="9.[151,1437,1942,1997]" pageId="9" pageNumber="46">
<emphasis id="C3A1EAA7FFC96226FF25A6B5FEBFF827" bold="true" box="[151,280,1942,1966]" pageId="9" pageNumber="46">FIGURE 6</emphasis>
. 
<taxonomicName id="36D54D36FFC96226FE95A6B4FDF6F827" box="[295,593,1943,1966]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="46" phylum="Arthropoda" rank="species" species="mcphersonae" status="sp. nov.">
<emphasis id="C3A1EAA7FFC96226FE95A6B4FDF6F827" box="[295,593,1943,1966]" italics="true" pageId="9" pageNumber="46">Metabetaeus mcphersonae</emphasis>
</taxonomicName>
<emphasis id="C3A1EAA7FFC96226FDEBA6B5FD3EF827" bold="true" box="[601,665,1942,1966]" pageId="9" pageNumber="46">
<taxonomicNameLabel id="D89257DCFFC96226FDEBA6B5FD3EF827" box="[601,665,1942,1966]" pageId="9" pageNumber="46" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, ovigerous female from Moorea (FLMNH UF Arthropoda 23247): A, habitus, lateral; B, same, dorsal.
</paragraph>
</caption>
<caption id="A5AA663DFFCA6225FF25A76EFAD8F94D" httpUri="https://zenodo.org/record/196832/files/figure.png" pageId="10" pageNumber="47" targetBox="[151,1436,194,1590]" targetPageId="10">
<paragraph id="F16A36B5FFCA6225FF25A76EFAD8F94D" blockId="10.[151,1437,1613,1732]" pageId="10" pageNumber="47">
<emphasis id="C3A1EAA7FFCA6225FF25A76EFEBFF9EC" bold="true" box="[151,280,1613,1637]" pageId="10" pageNumber="47">FIGURE 7</emphasis>
. 
<taxonomicName id="36D54D36FFCA6225FE95A76DFDF6F9EC" box="[295,593,1614,1637]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="47" phylum="Arthropoda" rank="species" species="mcphersonae" status="sp. nov.">
<emphasis id="C3A1EAA7FFCA6225FE95A76DFDF6F9EC" box="[295,593,1614,1637]" italics="true" pageId="10" pageNumber="47">Metabetaeus mcphersonae</emphasis>
</taxonomicName>
<emphasis id="C3A1EAA7FFCA6225FDEBA76EFD3EF9EC" bold="true" box="[601,665,1613,1637]" pageId="10" pageNumber="47">
<taxonomicNameLabel id="D89257DCFFCA6225FDEBA76EFD3EF9EC" box="[601,665,1613,1637]" pageId="10" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, ovigerous female from Moorea (FLMNH UF Arthropoda 23247): A, frontal region, dorsal; B, major cheliped (right), chela, mesial (= dorsal); paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23250): C, anterior region, lateral; D, chelipeds, lateral (= ventral); E, uropod, dorsal; paratype, ovigerous female from Moorea (FLMNH UF Arthropoda 23248): F, embryos attached to pleopods; G, zoea.
</paragraph>
</caption>
<paragraph id="F16A36B5FFCB6224FF74A1BBFAF6FD35" blockId="11.[151,1437,152,700]" pageId="11" pageNumber="48">
<bibRefCitation id="95444B44FFCB6224FF74A1BBFE58FF3B" author="Banner" box="[198,511,152,178]" pageId="11" pageNumber="54" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner (1983)</bibRefCitation>
reported as 
<taxonomicName id="36D54D36FFCB6224FD10A1BBFC9FFF38" box="[674,824,152,177]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="C3A1EAA7FFCB6224FD10A1BBFC9FFF38" box="[674,824,152,177]" italics="true" pageId="11" pageNumber="48">Metabetaeus</emphasis>
</taxonomicName>
sp. five incomplete specimens (all missing their chelipeds) from Tuléar, southwestern Madagascar. These specimens were collected by an airlift suction sampler on the outer reef front (&quot;gravely sedimentary accumulation at the foot of a vertical cliff along the edges of an outer creed [= outer re-entrant] of the outer reef front”), at a depth of about 
<specimenCount id="E7D3FD3CFFCB6224FB25A02BFB6BFEAB" box="[1175,1228,264,290]" pageId="11" pageNumber="48" type="generic">
<quantity id="362D9B50FFCB6224FB25A02BFB60FEAB" box="[1175,1223,264,290]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="48" unit="m" value="7.0">7 m</quantity>
.
</specimenCount>
The whereabouts of these specimens are unknown and their taxonomic identity remains unclear. 
<bibRefCitation id="95444B44FFCB6224FB8BA00EFAF8FECE" author="Banner" box="[1081,1375,301,327]" pageId="11" pageNumber="54" refString="Banner, A. H. &amp; Banner, D. M. (1983) An annotated checklist of the alpheid shrimp from the western Indian Ocean. Travaux et Documents de l'ORSTOM, 158, 1 - 164." type="journal article" year="1983">Banner &amp; Banner (1983)</bibRefCitation>
tried to provide a plausible explanation for the presence of 
<taxonomicName id="36D54D36FFCB6224FCA0A071FC0FFEE2" box="[786,936,338,363]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="C3A1EAA7FFCB6224FCA0A071FC0FFEE2" box="[786,936,338,363]" italics="true" pageId="11" pageNumber="48">Metabetaeus</emphasis>
</taxonomicName>
in rubble substrate of a coral reef, e.g., by the possible proximity of karstic systems (according to B. Thomassin) or the tolerance of saltwater by hypogeal euryhaline shrimps (according to J. Maciolek). However, the finding of 
<taxonomicName id="36D54D36FFCB6224FBE4A0BEFABEFE3F" box="[1110,1305,413,438]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="mcphersonae" status="sp. nov.">
<emphasis id="C3A1EAA7FFCB6224FBE4A0BEFABEFE3F" box="[1110,1305,413,438]" italics="true" pageId="11" pageNumber="48">M. mcphersonae</emphasis>
</taxonomicName>
<emphasis id="C3A1EAA7FFCB6224FA93A0BEFAC4FE3E" bold="true" box="[1313,1379,413,439]" pageId="11" pageNumber="48">
<taxonomicNameLabel id="D89257DCFFCB6224FA93A0BEFAC4FE3E" box="[1313,1379,413,439]" pageId="11" pageNumber="48" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
on a fore-reef in Moorea, where no karstic systems are known, suggests that the ancestors of the three recent species of 
<taxonomicName id="36D54D36FFCB6224FEA7A0CBFE0BFD88" box="[277,428,488,513]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="C3A1EAA7FFCB6224FEA7A0CBFE0BFD88" box="[277,428,488,513]" italics="true" pageId="11" pageNumber="48">Metabetaeus</emphasis>
</taxonomicName>
were probably truly marine shrimps associated with rubble and/or deep reef matrix. Following this hypothetical scenario, 
<taxonomicName id="36D54D36FFCB6224FDE3A32EFCB3FDAF" box="[593,788,525,550]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="mcphersonae" status="sp. nov.">
<emphasis id="C3A1EAA7FFCB6224FDE3A32EFCB3FDAF" box="[593,788,525,550]" italics="true" pageId="11" pageNumber="48">M. mcphersonae</emphasis>
</taxonomicName>
<emphasis id="C3A1EAA7FFCB6224FCAEA32EFCF8FDAE" bold="true" box="[796,863,525,551]" pageId="11" pageNumber="48">
<taxonomicNameLabel id="D89257DCFFCB6224FCAEA32EFCF8FDAE" box="[796,863,525,551]" pageId="11" pageNumber="48" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
remained a truly marine shrimp, while ancestors of 
<taxonomicName id="36D54D36FFCB6224FF0AA311FE99FDC2" box="[184,318,562,587]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="C3A1EAA7FFCB6224FF0AA311FE99FDC2" box="[184,318,562,587]" italics="true" pageId="11" pageNumber="48">M. minutus</emphasis>
</taxonomicName>
and 
<taxonomicName id="36D54D36FFCB6224FECBA311FE55FDC2" box="[377,498,562,587]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="lohena">
<emphasis id="C3A1EAA7FFCB6224FECBA311FE55FDC2" box="[377,498,562,587]" italics="true" pageId="11" pageNumber="48">M. lohena</emphasis>
</taxonomicName>
invaded anchialine habitats. A similar evolutionary scenario was proposed for another anchialine cave alpheid, 
<taxonomicName id="36D54D36FFCB6224FD9EA37BFB83FDFB" authority="Anker &amp; Iliffe, 2000" authorityName="Anker &amp; Iliffe" authorityYear="2000" box="[556,1060,600,626]" class="Malacostraca" family="Alpheidae" genus="Bermudacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="harti">
<emphasis id="C3A1EAA7FFCB6224FD9EA37BFCBAFDF8" box="[556,797,600,625]" italics="true" pageId="11" pageNumber="48">Bermudacaris harti</emphasis>
<bibRefCitation id="95444B44FFCB6224FC97A37BFB83FDFB" author="Anker" box="[805,1060,600,626]" pageId="11" pageNumber="54" refString="Anker, A. &amp; Iliffe, T. M. (2000) Description of Bermudacaris harti, a new genus, and species (Crustacea: Decapoda: Alpheidae) from anchialine caves of Bermuda. Proceedings of the Biological Society of Washington, 113, 761 - 775." type="journal article" year="2000">Anker &amp; Iliffe, 2000</bibRefCitation>
</taxonomicName>
(see 
<bibRefCitation id="95444B44FFCB6224FBD4A37BFAC5FDFB" author="Anker" box="[1126,1378,600,626]" pageId="11" pageNumber="54" refString="Anker, A. &amp; Iliffe, T. M. (2000) Description of Bermudacaris harti, a new genus, and species (Crustacea: Decapoda: Alpheidae) from anchialine caves of Bermuda. Proceedings of the Biological Society of Washington, 113, 761 - 775." type="journal article" year="2000">Anker &amp; Iliffe 2000</bibRefCitation>
). An alternative scenario, i.e. 
<taxonomicName id="36D54D36FFCB6224FE7AA35EFD32FD1F" box="[456,661,637,662]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="48" phylum="Arthropoda" rank="species" species="mcphersonae" status="sp. nov.">
<emphasis id="C3A1EAA7FFCB6224FE7AA35EFD32FD1F" box="[456,661,637,662]" italics="true" pageId="11" pageNumber="48">M. mcphersonae</emphasis>
</taxonomicName>
<emphasis id="C3A1EAA7FFCB6224FD2FA35EFD43FD1E" bold="true" box="[669,740,637,663]" pageId="11" pageNumber="48">
<taxonomicNameLabel id="D89257DCFFCB6224FD2FA35EFD43FD1E" box="[669,740,637,663]" pageId="11" pageNumber="48" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
having evolved from anchialine ancestors, is less likely considering the marine ancestry of the vast majority of alpheid shrimps, but cannot be totally excluded.
</paragraph>
</subSubSection>
</treatment>
</document>