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<mods:title id="66C98712092A1BF31700F7BCA0E915B3">Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina)</mods:title>
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<mods:namePart id="1356F7BF78A23BD2D0D09A323BD13AC9">Hutchinson, Paul M.</mods:namePart>
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<mods:namePart id="465A9C0982666DCA75CF2EDE6869A21B">Allsopp, Peter G.</mods:namePart>
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Key to
<taxonomicName id="368F4F92FFDFFFB7CB83D53FFECEFA95" authority="Arrow, 1908" authorityName="Arrow" authorityYear="1908" box="[241,327,1307,1333]" class="Insecta" family="Dynastidae" genus="Anomalomorpha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="457" phylum="Arthropoda" rank="genus">species</taxonomicName>
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<paragraph id="F1303411FFDFFFB7CBE5D545FA14FA11" blockId="18.[151,1437,1377,1809]" pageId="18" pageNumber="457">
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1 Pygidium with apical ridge glabrous (
<figureCitation id="69B42894FFDFFFB7C92FD545FD38FAD6" box="[605,689,1377,1399]" captionStart="FIGURES 3134" captionStartId="13.[151,264,1982,2006]" captionTargetBox="[173,1414,921,1943]" captionTargetId="figure-330@13.[173,1414,921,1958]" captionTargetPageId="13" captionText="FIGURES 3134. Anomalomorpha pilbara Hutchinson &amp; Allsopp new species, holotype male. 31, Head dorsal view; 32, head lateral view; 33, pygidium; 34, labels and mounted aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5749023" httpUri="https://zenodo.org/record/5749023/files/figure.png" pageId="18" pageNumber="457">Figs. 33</figureCitation>
,
<figureCitation id="69B42894FFDFFFB7C9CCD545FD50FAD7" box="[702,729,1377,1399]" captionStart="FIGURES 3840" captionStartId="15.[151,264,1821,1845]" captionTargetBox="[151,1435,741,1798]" captionTargetId="figure-261@15.[151,1435,741,1798]" captionTargetPageId="15" captionText="FIGURES 3840. Anomalomorpha pilbara Hutchinson &amp; Allsopp new species, allotype female. 38, Head dorsal view; 39, head lateral view; 40, pygidium." figureDoi="http://doi.org/10.5281/zenodo.5749027" httpUri="https://zenodo.org/record/5749027/files/figure.png" pageId="18" pageNumber="457">40</figureCitation>
); metacoxa with posterior margin angulate at midlength then narrowly arcuate at posterolateral angle (
<figureCitation id="69B42894FFDFFFB7C97FD55AFDDDFA34" box="[525,596,1406,1428]" captionStart="FIGURES 4143" captionStartId="16.[151,264,637,661]" captionTargetBox="[151,1435,181,613]" captionTargetId="figure-26@16.[151,1435,181,613]" captionTargetPageId="16" captionText="FIGURES 4143. Anomalomorpha metacoxae. 41, Anomalomorpha anthracina Arrow, 1908; 42, A. pilbara Hutchinson &amp; Allsopp new species; 43, A. monteithi Allsopp, 2000." figureDoi="http://doi.org/10.5281/zenodo.5749029" httpUri="https://zenodo.org/record/5749029/files/figure.png" pageId="18" pageNumber="457">Fig. 42</figureCitation>
); parameres with lateral tooth obtusely rounded, tapering to acute apices that are curved inwards (
<figureCitation id="69B42894FFDFFFB7CA49D5BFFE38FA11" box="[315,433,1435,1457]" captionStart="FIGURES 4449" captionStartId="16.[151,264,1680,1704]" captionTargetBox="[175,1435,744,1639]" captionTargetId="figure-53@16.[151,1435,722,1656]" captionTargetPageId="16" captionText="FIGURES 4449. Anomalomorpha male aedeagi. 4445, Anomalomorpha anthracina Arrow, 1908. 44, parameres dorsal; 45, aedeagus lateral; 4647, A. pilbara Hutchinson &amp; Allsopp new species. 46, parameres dorsal; 47, aedeagus lateral; 4849, A. monteithi Allsopp, 2000. 48, parameres dorsal; 49, aedeagus lateral. Arrows show regions of major differences." figureDoi="http://doi.org/10.5281/zenodo.5749031" httpUri="https://zenodo.org/record/5749031/files/figure.png" pageId="18" pageNumber="457">Figs. 4647</figureCitation>
);
<collectingRegion id="334BFAF3FFDFFFB7CAB3D5B8FDE5FA11" box="[449,620,1435,1457]" country="Australia" name="Western Australia" pageId="18" pageNumber="457">Western Australia</collectingRegion>
.................
<taxonomicName id="368F4F92FFDFFFB7C83CD5BFFA95FA11" authority="Hutchinson &amp; Allsopp" authorityName="Hutchinson &amp; Allsopp" authorityYear="2021" box="[846,1308,1435,1457]" class="Insecta" family="Dynastidae" genus="Anomalomorpha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="457" phylum="Arthropoda" rank="species" species="pilbara" status="sp. nov.">
<emphasis id="C3FBE803FFDFFFB7C83CD5BFFBB4FA10" box="[846,1085,1435,1456]" italics="true" pageId="18" pageNumber="457">Anomalomorpha pilbara</emphasis>
Hutchinson &amp; Allsopp
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,
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<taxonomicNameLabel id="D8C85578FFDFFFB7CE54D5BFFA14FA11" box="[1318,1437,1435,1457]" pageId="18" pageNumber="457" rank="species">new species</taxonomicNameLabel>
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<paragraph id="F1303411FFDFFFB7CBE5D59DFA15F986" blockId="18.[151,1437,1377,1809]" pageId="18" pageNumber="457">
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- Pygidium with apical ridge setose (
<figureCitation id="69B42894FFDFFFB7C944D59DFD0FFA6E" box="[566,646,1465,1487]" captionStart="FIGURES 1518" captionStartId="8.[151,264,1923,1947]" captionTargetBox="[151,1433,396,1900]" captionTargetId="figure-108@8.[151,1435,381,1900]" captionTargetPageId="8" captionText="FIGURES 1518. Anomalomorpha anthracina Arrow, 1908, male. 15, Dorsal view; 16, ventral view; 17, head dorso-lateral view; 18, pygidium." figureDoi="http://doi.org/10.5281/zenodo.5749013" httpUri="https://zenodo.org/record/5749013/files/figure.png" pageId="18" pageNumber="457">Figs. 18</figureCitation>
,
<figureCitation id="69B42894FFDFFFB7C9E2D59DFD23FA6E" box="[656,682,1465,1486]" captionStart="FIGURES 1922" captionStartId="9.[151,264,1764,1788]" captionTargetBox="[151,1435,181,1741]" captionTargetId="figure-19@9.[151,1435,181,1741]" captionTargetPageId="9" captionText="FIGURES 1922. Anomalomorpha anthracina Arrow, 1908, female. 19, Dorsal view; 20, ventral view; 21, head dorso-lateral view; 22, pygidium." figureDoi="http://doi.org/10.5281/zenodo.5749015" httpUri="https://zenodo.org/record/5749015/files/figure.png" pageId="18" pageNumber="457">22</figureCitation>
,
<figureCitation id="69B42894FFDFFFB7C9C7D59DFD59FA6F" box="[693,720,1465,1487]" captionStart="FIGURES 2327" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[272,1345,200,1919]" captionTargetId="figure-26@10.[241,1345,181,1919]" captionTargetPageId="10" captionText="FIGURES 2327. Anomalomorpha monteithi Allsopp, 2000, male. 23, Dorsal view; 24, ventral view; 25, lateral view; 26, head dorso-lateral view; 27, pygidium." figureDoi="http://doi.org/10.5281/zenodo.5749019" httpUri="https://zenodo.org/record/5749019/files/figure.png" pageId="18" pageNumber="457">27</figureCitation>
); metacoxa with posterior margin broadly arcuate from midlength through posterolateral angle or sublinear then arcuate through posterolateral angle (
<figureCitation id="69B42894FFDFFFB7C8C3D5F2FBAEFA4C" box="[945,1063,1494,1516]" captionStart="FIGURES 4143" captionStartId="16.[151,264,637,661]" captionTargetBox="[151,1435,181,613]" captionTargetId="figure-26@16.[151,1435,181,613]" captionTargetPageId="16" captionText="FIGURES 4143. Anomalomorpha metacoxae. 41, Anomalomorpha anthracina Arrow, 1908; 42, A. pilbara Hutchinson &amp; Allsopp new species; 43, A. monteithi Allsopp, 2000." figureDoi="http://doi.org/10.5281/zenodo.5749029" httpUri="https://zenodo.org/record/5749029/files/figure.png" pageId="18" pageNumber="457">Figs. 41, 43</figureCitation>
); parameres with lateral tooth angulate and prominent, tapering to truncate or acute apices, apices curved slightly outwards (
<figureCitation id="69B42894FFDFFFB7CF63D5D7FB46F9A9" box="[1041,1231,1523,1545]" captionStart="FIGURES 4449" captionStartId="16.[151,264,1680,1704]" captionTargetBox="[175,1435,744,1639]" captionTargetId="figure-53@16.[151,1435,722,1656]" captionTargetPageId="16" captionText="FIGURES 4449. Anomalomorpha male aedeagi. 4445, Anomalomorpha anthracina Arrow, 1908. 44, parameres dorsal; 45, aedeagus lateral; 4647, A. pilbara Hutchinson &amp; Allsopp new species. 46, parameres dorsal; 47, aedeagus lateral; 4849, A. monteithi Allsopp, 2000. 48, parameres dorsal; 49, aedeagus lateral. Arrows show regions of major differences." figureDoi="http://doi.org/10.5281/zenodo.5749031" httpUri="https://zenodo.org/record/5749031/files/figure.png" pageId="18" pageNumber="457">Figs. 4445, 4849</figureCitation>
); eastern
<collectingCountry id="89987481FFDFFFB7CE5AD5D7FA09F9A9" box="[1320,1408,1523,1545]" name="Australia" pageId="18" pageNumber="457">Australia</collectingCountry>
. ................................................................................................... 2
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<keyStep id="4A7B2CB4FFDFFFB7CBE5D60AFA15F8B0" pageId="18" pageNumber="457">
<paragraph id="F1303411FFDFFFB7CBE5D60AFA15F93B" blockId="18.[151,1437,1377,1809]" pageId="18" pageNumber="457">
<keyLead id="4A7E9724FFDFFFB7CBE5D60AFA15F93B" pageId="18" pageNumber="457">
2 Dorsal surface shiny (
<figureCitation id="69B42894FFDFFFB7CAC7D60AFD8FF9E4" box="[437,518,1582,1604]" captionStart="FIGURES 1518" captionStartId="8.[151,264,1923,1947]" captionTargetBox="[151,1433,396,1900]" captionTargetId="figure-108@8.[151,1435,381,1900]" captionTargetPageId="8" captionText="FIGURES 1518. Anomalomorpha anthracina Arrow, 1908, male. 15, Dorsal view; 16, ventral view; 17, head dorso-lateral view; 18, pygidium." figureDoi="http://doi.org/10.5281/zenodo.5749013" httpUri="https://zenodo.org/record/5749013/files/figure.png" pageId="18" pageNumber="457">Figs. 15</figureCitation>
,
<figureCitation id="69B42894FFDFFFB7C962D60AFDA3F9E4" box="[528,554,1582,1604]" captionStart="FIGURES 1922" captionStartId="9.[151,264,1764,1788]" captionTargetBox="[151,1435,181,1741]" captionTargetId="figure-19@9.[151,1435,181,1741]" captionTargetPageId="9" captionText="FIGURES 1922. Anomalomorpha anthracina Arrow, 1908, female. 19, Dorsal view; 20, ventral view; 21, head dorso-lateral view; 22, pygidium." figureDoi="http://doi.org/10.5281/zenodo.5749015" httpUri="https://zenodo.org/record/5749015/files/figure.png" pageId="18" pageNumber="457">19</figureCitation>
); distance between outer edges of canthus 1.81.9 times interocular distance (
<figureCitation id="69B42894FFDFFFB7CE68D60AFAE2F9E3" box="[1306,1387,1582,1604]" captionStart="FIGURES 1518" captionStartId="8.[151,264,1923,1947]" captionTargetBox="[151,1433,396,1900]" captionTargetId="figure-108@8.[151,1435,381,1900]" captionTargetPageId="8" captionText="FIGURES 1518. Anomalomorpha anthracina Arrow, 1908, male. 15, Dorsal view; 16, ventral view; 17, head dorso-lateral view; 18, pygidium." figureDoi="http://doi.org/10.5281/zenodo.5749013" httpUri="https://zenodo.org/record/5749013/files/figure.png" pageId="18" pageNumber="457">Figs. 17</figureCitation>
,
<figureCitation id="69B42894FFDFFFB7CE07D60AFA19F9E4" box="[1397,1424,1582,1604]" captionStart="FIGURES 1922" captionStartId="9.[151,264,1764,1788]" captionTargetBox="[151,1435,181,1741]" captionTargetId="figure-19@9.[151,1435,181,1741]" captionTargetPageId="9" captionText="FIGURES 1922. Anomalomorpha anthracina Arrow, 1908, female. 19, Dorsal view; 20, ventral view; 21, head dorso-lateral view; 22, pygidium." figureDoi="http://doi.org/10.5281/zenodo.5749015" httpUri="https://zenodo.org/record/5749015/files/figure.png" pageId="18" pageNumber="457">21</figureCitation>
); pygidium with few long setae at sides in anterior third (
<figureCitation id="69B42894FFDFFFB7C983D66FFCC8F9C0" box="[753,833,1611,1633]" captionStart="FIGURES 1518" captionStartId="8.[151,264,1923,1947]" captionTargetBox="[151,1433,396,1900]" captionTargetId="figure-108@8.[151,1435,381,1900]" captionTargetPageId="8" captionText="FIGURES 1518. Anomalomorpha anthracina Arrow, 1908, male. 15, Dorsal view; 16, ventral view; 17, head dorso-lateral view; 18, pygidium." figureDoi="http://doi.org/10.5281/zenodo.5749013" httpUri="https://zenodo.org/record/5749013/files/figure.png" pageId="18" pageNumber="457">Figs. 18</figureCitation>
,
<figureCitation id="69B42894FFDFFFB7C838D66FFCECF9C1" box="[842,869,1611,1633]" captionStart="FIGURES 1922" captionStartId="9.[151,264,1764,1788]" captionTargetBox="[151,1435,181,1741]" captionTargetId="figure-19@9.[151,1435,181,1741]" captionTargetPageId="9" captionText="FIGURES 1922. Anomalomorpha anthracina Arrow, 1908, female. 19, Dorsal view; 20, ventral view; 21, head dorso-lateral view; 22, pygidium." figureDoi="http://doi.org/10.5281/zenodo.5749015" httpUri="https://zenodo.org/record/5749015/files/figure.png" pageId="18" pageNumber="457">22</figureCitation>
); parameres tapering to truncate apices (
<figureCitation id="69B42894FFDFFFB7CF94D66FFAD5F9C1" box="[1254,1372,1611,1633]" captionStart="FIGURES 4449" captionStartId="16.[151,264,1680,1704]" captionTargetBox="[175,1435,744,1639]" captionTargetId="figure-53@16.[151,1435,722,1656]" captionTargetPageId="16" captionText="FIGURES 4449. Anomalomorpha male aedeagi. 4445, Anomalomorpha anthracina Arrow, 1908. 44, parameres dorsal; 45, aedeagus lateral; 4647, A. pilbara Hutchinson &amp; Allsopp new species. 46, parameres dorsal; 47, aedeagus lateral; 4849, A. monteithi Allsopp, 2000. 48, parameres dorsal; 49, aedeagus lateral. Arrows show regions of major differences." figureDoi="http://doi.org/10.5281/zenodo.5749031" httpUri="https://zenodo.org/record/5749031/files/figure.png" pageId="18" pageNumber="457">Figs. 4445</figureCitation>
); body 11.317.0 mm long; eastern
<collectingCountry id="89987481FFDFFFB7CA80D64DFDC3F9DF" box="[498,586,1641,1663]" name="Australia" pageId="18" pageNumber="457">Australia</collectingCountry>
from northeastern
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to inland
<collectingRegion id="334BFAF3FFDFFFB7C8A1D64DFBA9F9DF" box="[979,1056,1641,1663]" country="Australia" name="Victoria" pageId="18" pageNumber="457">Victoria</collectingRegion>
.................................................................................................
<taxonomicName id="368F4F92FFDFFFB7CF7CD6A2FA15F93B" authority="Arrow, 1908" authorityName="Arrow" authorityYear="1908" box="[1038,1436,1670,1691]" class="Insecta" family="Dynastidae" genus="Anomalomorpha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="457" phylum="Arthropoda" rank="species" species="anthracina">
<emphasis id="C3FBE803FFDFFFB7CF7CD6A2FA97F93B" box="[1038,1310,1670,1691]" italics="true" pageId="18" pageNumber="457">Anomalomorpha anthracina</emphasis>
<bibRefCitation id="951E49E0FFDFFFB7CE51D6A2FA15F93B" author="Arrow, G. J." box="[1315,1436,1670,1691]" pageId="18" pageNumber="457" pagination="321 - 358" refId="ref10294" refString="Arrow, G. J. (1908) A contribution to the classification of the coleopterous family Dynastidae. Transactions of the Entomological Society of London, 1908, 321 - 358. https: // doi. org / 10.1111 / j. 1365 - 2311.1908. tb 02151. x" type="journal article" year="1908">Arrow, 1908</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="F1303411FFDFFFB7CBE5D680FA15F8B0" blockId="18.[151,1437,1377,1809]" pageId="18" pageNumber="457">
<keyLead id="4A7E9724FFDFFFB7CBE5D680FA15F8B0" pageId="18" pageNumber="457">
- Dorsal surface sericeous (
<figureCitation id="69B42894FFDFFFB7CAAFD687FDAFF919" box="[477,550,1699,1721]" captionStart="FIGURES 2327" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[272,1345,200,1919]" captionTargetId="figure-26@10.[241,1345,181,1919]" captionTargetPageId="10" captionText="FIGURES 2327. Anomalomorpha monteithi Allsopp, 2000, male. 23, Dorsal view; 24, ventral view; 25, lateral view; 26, head dorso-lateral view; 27, pygidium." figureDoi="http://doi.org/10.5281/zenodo.5749019" httpUri="https://zenodo.org/record/5749019/files/figure.png" pageId="18" pageNumber="457">Fig. 23</figureCitation>
); distance between outer edges of canthi 1.6 times interocular distance (
<figureCitation id="69B42894FFDFFFB7CF96D687FAA4F919" box="[1252,1325,1699,1721]" captionStart="FIGURES 2327" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[272,1345,200,1919]" captionTargetId="figure-26@10.[241,1345,181,1919]" captionTargetPageId="10" captionText="FIGURES 2327. Anomalomorpha monteithi Allsopp, 2000, male. 23, Dorsal view; 24, ventral view; 25, lateral view; 26, head dorso-lateral view; 27, pygidium." figureDoi="http://doi.org/10.5281/zenodo.5749019" httpUri="https://zenodo.org/record/5749019/files/figure.png" pageId="18" pageNumber="457">Fig. 26</figureCitation>
); pygidium with line of very long setae across posterior third (
<figureCitation id="69B42894FFDFFFB7C9B8D6E5FC9BF977" box="[714,786,1729,1751]" captionStart="FIGURES 2327" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[272,1345,200,1919]" captionTargetId="figure-26@10.[241,1345,181,1919]" captionTargetPageId="10" captionText="FIGURES 2327. Anomalomorpha monteithi Allsopp, 2000, male. 23, Dorsal view; 24, ventral view; 25, lateral view; 26, head dorso-lateral view; 27, pygidium." figureDoi="http://doi.org/10.5281/zenodo.5749019" httpUri="https://zenodo.org/record/5749019/files/figure.png" pageId="18" pageNumber="457">Fig. 27</figureCitation>
); parameres tapering to acute apices (
<figureCitation id="69B42894FFDFFFB7CF0CD6E5FB7DF977" box="[1150,1268,1729,1751]" captionStart="FIGURES 4449" captionStartId="16.[151,264,1680,1704]" captionTargetBox="[175,1435,744,1639]" captionTargetId="figure-53@16.[151,1435,722,1656]" captionTargetPageId="16" captionText="FIGURES 4449. Anomalomorpha male aedeagi. 4445, Anomalomorpha anthracina Arrow, 1908. 44, parameres dorsal; 45, aedeagus lateral; 4647, A. pilbara Hutchinson &amp; Allsopp new species. 46, parameres dorsal; 47, aedeagus lateral; 4849, A. monteithi Allsopp, 2000. 48, parameres dorsal; 49, aedeagus lateral. Arrows show regions of major differences." figureDoi="http://doi.org/10.5281/zenodo.5749031" httpUri="https://zenodo.org/record/5749031/files/figure.png" pageId="18" pageNumber="457">Figs. 4849</figureCitation>
); body 19.220.0 mm long; wet sclerophyll forest to the west of tropical rainforest in northeastern
<collectingRegion id="334BFAF3FFDFFFB7C89BD6FAFBD3F954" box="[1001,1114,1758,1780]" country="Australia" name="Queensland" pageId="18" pageNumber="457">Queensland</collectingRegion>
............................................................................................
<taxonomicName id="368F4F92FFDFFFB7CF7CD6DFFA15F8B0" authority="Allsopp, 2000" authorityName="Allsopp" authorityYear="2000" box="[1038,1436,1787,1809]" class="Insecta" family="Dynastidae" genus="Anomalomorpha" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="457" phylum="Arthropoda" rank="species" species="monteithi" status="gen. nov.">
<emphasis id="C3FBE803FFDFFFB7CF7CD6DFFA99F8B0" box="[1038,1296,1787,1808]" italics="true" pageId="18" pageNumber="457">Anomalomorpha monteithi</emphasis>
<bibRefCitation id="951E49E0FFDFFFB7CE66D6DFFA15F8B0" author="Allsopp, P. G." box="[1300,1436,1787,1809]" pageId="18" pageNumber="457" pagination="1 - 7" refId="ref10108" refString="Allsopp, P. G. (2000) Revision of the Australian genus Anomalomorpha Arrow (Coleoptera: Scarabaeidae: Dynastinae) with a new species from the wet tropics of Queensland. Memoirs of the Queensland Museum, 46, 1 - 7." type="journal article" year="2000">Allsopp, 2000</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</subSection>
</document>