treatments-xml/data/79/26/85/79268507FFCFFFA1CBE5D1DBFCE0FE0D.xml
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<document id="3B0E606B415B22DC8B6A5B5090690CDB" ID-CLB-Dataset="3142" ID-DOI="10.11646/zootaxa.5072.5.2" ID-GBIF-Dataset="cdde4106-d3d5-4294-9f6c-1e82afd8d580" ID-ISSN="1175-5326" ID-Zenodo-Dep="5748997" ID-ZooBank="B1D30BB4-8CAB-419E-98DE-02D97726137C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638425477373" checkinUser="plazi" docAuthor="Hutchinson, Paul M. &amp; Allsopp, Peter G." docDate="2021" docId="79268507FFCFFFA1CBE5D1DBFCE0FE0D" docLanguage="en" docName="zootaxa.5072.5.2.pdf" docOrigin="Zootaxa 5072 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enracius longipes Dechambre 1999" docType="treatment" docVersion="7" lastPageNumber="443" masterDocId="851FFD7FFFCDFFA5CB72D024FF89FFA0" masterDocTitle="Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina)" masterLastPageNumber="462" masterPageNumber="439" pageNumber="441" updateTime="1699471722501" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="79DE73E4EC8DC88B8F3F0992D2ADDEBD">Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina)</mods:title>
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<mods:namePart id="7A5451FA3CBAD8CC847FDE45C449FC01">Hutchinson, Paul M.</mods:namePart>
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<mods:namePart id="9D78C730378A38844192E63856C848B9">Allsopp, Peter G.</mods:namePart>
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<mods:date id="4F9FD49CDAE5C56783AC9B45D37C0DF3">2021</mods:date>
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<treatment id="79268507FFCFFFA1CBE5D1DBFCE0FE0D" ID-DOI="http://doi.org/10.5281/zenodo.5750342" ID-GBIF-Taxon="190874937" ID-Zenodo-Dep="5750342" LSID="urn:lsid:plazi:treatment:79268507FFCFFFA1CBE5D1DBFCE0FE0D" httpUri="http://treatment.plazi.org/id/79268507FFCFFFA1CBE5D1DBFCE0FE0D" lastPageId="4" lastPageNumber="443" pageId="2" pageNumber="441">
<subSubSection id="B995679AFFCFFFA7CBE5D1DBFDDDFDB8" box="[151,596,510,537]" pageId="2" pageNumber="441" type="nomenclature">
<paragraph id="F1303411FFCFFFA7CBE5D1DBFDDDFDB8" blockId="2.[151,596,510,537]" box="[151,596,510,537]" pageId="2" pageNumber="441">
<heading id="AA78837DFFCFFFA7CBE5D1DBFDDDFDB8" bold="true" box="[151,596,510,537]" fontSize="11" level="1" pageId="2" pageNumber="441" reason="1">
<taxonomicName id="368F4F92FFCFFFA7CBE5D1DBFDDDFDB8" authority="Dechambre, 1999" authorityName="Dechambre" authorityYear="1999" box="[151,596,510,537]" class="Insecta" family="Dynastidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="441" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="C3FBE803FFCFFFA7CBE5D1DBFDDDFDB8" bold="true" box="[151,596,510,537]" pageId="2" pageNumber="441">
<emphasis id="C3FBE803FFCFFFA7CBE5D1DBFEFBFDB9" bold="true" box="[151,370,510,537]" italics="true" pageId="2" pageNumber="441">Enracius longipes</emphasis>
<bibRefCitation id="951E49E0FFCFFFA7CA08D1DBFDDDFDB8" author="Dechambre, R. - P." box="[378,596,510,537]" pageId="2" pageNumber="441" pagination="37 - 38" refId="ref10492" refString="Dechambre, R. - P. (1999) Enracius longipes n. gen., n. sp., nouveau Dynastidae d'Australie (Col. Scarabaeoidea Dynastidae). Bulletin de la Societe Entomologique de France, 104, 37 - 38. https: // doi. org / 10.3406 / bsef. 1999.16541" type="journal article" year="1999">Dechambre, 1999</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="F1303411FFCFFFA7CBE5D260FC90FDFB" blockId="2.[151,793,579,603]" box="[151,793,579,603]" pageId="2" pageNumber="441">
<treatmentCitationGroup id="D19F133FFFCFFFA7CBE5D260FC90FDFB" box="[151,793,579,603]" pageId="2" pageNumber="441">
<treatmentCitation id="702E1200FFCFFFA7CBE5D260FDBAFDFB" author="Dechambre, R. - P." box="[151,563,579,603]" page="38" pageId="2" pageNumber="441" year="1999">
<taxonomicName id="368F4F92FFCFFFA7CBE5D260FDBAFDFB" authority="Dechambre, 1999: 38" authorityName="Dechambre" authorityPageNumber="38" authorityYear="1999" box="[151,563,579,603]" class="Insecta" family="Dynastidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="441" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="C3FBE803FFCFFFA7CBE5D260FEC6FDFB" box="[151,335,579,603]" italics="true" pageId="2" pageNumber="441">Enracius longipes</emphasis>
<bibRefCitation id="951E49E0FFCFFFA7CA27D267FDBAFDFB" author="Dechambre, R. - P." box="[341,563,579,603]" pageId="2" pageNumber="441" pagination="37 - 38" refId="ref10492" refString="Dechambre, R. - P. (1999) Enracius longipes n. gen., n. sp., nouveau Dynastidae d'Australie (Col. Scarabaeoidea Dynastidae). Bulletin de la Societe Entomologique de France, 104, 37 - 38. https: // doi. org / 10.3406 / bsef. 1999.16541" type="journal article" year="1999">Dechambre, 1999: 38</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="702E1200FFCFFFA7C94FD267FC9DFDFB" author="Dechambre, R. - P." box="[573,788,579,603]" page="53" pageId="2" pageNumber="441" year="2005">
<bibRefCitation id="951E49E0FFCFFFA7C94FD267FC9DFDFB" author="Dechambre, R. - P." box="[573,788,579,603]" pageId="2" pageNumber="441" refId="ref10613" refString="Dechambre, R. - P. (2005) Dynastidae Australiens et Oceaniens. Les Coleopteres du Monde 30. Hillside Books, Canterbury, 132 pp." type="book" year="2005">Dechambre 2005: 53</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="B995679AFFCFFFA7CBE5D2A7FEE1FB89" pageId="2" pageNumber="441" type="materials_examined">
<paragraph id="F1303411FFCFFFA7CBE5D2A7FABEFD61" blockId="2.[151,1437,642,2037]" pageId="2" pageNumber="441">
<emphasis id="C3FBE803FFCFFFA7CBE5D2A7FEA8FD3C" bold="true" box="[151,289,642,669]" pageId="2" pageNumber="441">Type series.</emphasis>
<materialsCitation id="41E73E4CFFCFFFA7CA54D2A7FABEFD61" ID-GBIF-Occurrence="3413046318" collectingDate="1978-09-30" collectorName="S. Gaudaire-Thore" latitude="-32.333332" location="Caiguna" longLatPrecision="1211" longitude="125.083336" pageId="2" pageNumber="441" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="C3FBE803FFCFFFA7CA54D2A7FC89FD3C" bold="true" box="[294,768,642,669]" pageId="2" pageNumber="441">
<typeStatus id="2E348AB3FFCFFFA7CA54D2A7FE1AFD3D" box="[294,403,643,669]" pageId="2" pageNumber="441" type="holotype">Holotype</typeStatus>
male (by original designation):
</emphasis>
<collectingRegion id="334BFAF3FFCFFFA7C87CD2A7FC50FD3D" box="[782,985,643,669]" country="Australia" name="Western Australia" pageId="2" pageNumber="441">Western Australia</collectingRegion>
,
<quantity id="367799F4FFCFFFA7C890D2A7FBAEFD3D" box="[994,1063,643,669]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="441" unit="km" value="40.0">
<locationDeviation id="E0822D33FFCFFFA7C890D2A7FBAEFD3D" box="[994,1063,643,669]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="441" unit="km" value="40.0">40 km</locationDeviation>
</quantity>
W of
<location id="F45062CAFFCFFFA7CF14D2A7FB4AFD3D" LSID="urn:lsid:plazi:treatment:79268507FFCFFFA1CBE5D1DBFCE0FE0D:F45062CAFFCFFFA7CF14D2A7FB4AFD3D" box="[1126,1219,643,669]" latitude="-32.333332" longLatPrecision="1211" longitude="125.083336" name="Caiguna" pageId="2" pageNumber="441" stateProvince="Western Australia">Caiguna</location>
<geoCoordinate id="94BB52D6FFCFFFA7CFBBD2A6FABFFD3D" box="[1225,1334,642,669]" degrees="125" direction="east" minutes="05" orientation="longitude" pageId="2" pageNumber="441" precision="925" value="125.083336">125°05´E</geoCoordinate>
,
<geoCoordinate id="94BB52D6FFCFFFA7CE32D2A7FA14FD3D" box="[1344,1437,643,669]" degrees="32" direction="south" minutes="20" orientation="latitude" pageId="2" pageNumber="441" precision="925" value="-32.333332">32°20´S</geoCoordinate>
[32.33°S, 125.08°E],
<date id="853112D1FFCFFFA7CAFBD283FD87FD61" box="[393,526,679,705]" pageId="2" pageNumber="441" value="1978-09-30">
<collectingDate id="9575EB39FFCFFFA7CAFBD283FD87FD61" box="[393,526,679,705]" pageId="2" pageNumber="441" value="1978-09-30">30.IX.1978</collectingDate>
</date>
”; in “coll.
<collectorName id="5C7A51C7FFCFFFA7C9F7D283FCD1FD61" box="[645,856,679,705]" pageId="2" pageNumber="441">S. Gaudaire-Thore</collectorName>
” [label data taken from
<bibRefCitation id="951E49E0FFCFFFA7CF14D283FAA5FD61" author="Dechambre, R. - P." box="[1126,1324,678,705]" pageId="2" pageNumber="441" pagination="37 - 38" refId="ref10492" refString="Dechambre, R. - P. (1999) Enracius longipes n. gen., n. sp., nouveau Dynastidae d'Australie (Col. Scarabaeoidea Dynastidae). Bulletin de la Societe Entomologique de France, 104, 37 - 38. https: // doi. org / 10.3406 / bsef. 1999.16541" type="journal article" year="1999">Dechambre 1999</bibRefCitation>
].
</materialsCitation>
</paragraph>
<paragraph id="F1303411FFCFFFA7CBB5D2EFFEFCFCF2" blockId="2.[151,1437,642,2037]" pageId="2" pageNumber="441">
<materialsCitation id="41E73E4CFFCFFFA7CBB5D2EFFEF8FCF2" ID-GBIF-Occurrence="3413046306" collectingDate="1978-09-30" collectionCode="FD" collectorName="M. S. &amp; B. J. Moulds" determinerName="P. B. Carne &amp; R. - P. Dechambre" latitude="-32.333332" location="Caiguna" longLatPrecision="1211" longitude="125.083336" pageId="2" pageNumber="441" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<emphasis id="C3FBE803FFCFFFA7CBB5D2EFFE37FD44" bold="true" box="[199,446,714,741]" pageId="2" pageNumber="441">
<typeStatus id="2E348AB3FFCFFFA7CBB5D2EFFEBDFD45" box="[199,308,715,741]" pageId="2" pageNumber="441" type="paratype">Paratype</typeStatus>
(
<figureCitation id="69B42894FFCFFFA7CA30D2EEFE26FD44" box="[322,431,714,740]" captionStart="FIGURES 14" captionStartId="3.[151,264,1304,1328]" captionTargetBox="[196,1432,179,1277]" captionTargetId="figure-19@3.[156,1432,179,1277]" captionTargetPageId="3" captionText="FIGURES 14. Enracius longipes Dechambre, 1999, paratype male. 1, Dorsal view; 2, pygidium; 3, phallobase and parameres, lateral view; 4, labels (photograph by Fabien Dupuis)." figureDoi="http://doi.org/10.5281/zenodo.5749001" httpUri="https://zenodo.org/record/5749001/files/figure.png" pageId="2" pageNumber="441">Figs. 14</figureCitation>
).
</emphasis>
<specimenCount id="E789FF98FFCFFFA7CAB6D2EFFE61FD45" box="[452,488,715,741]" pageId="2" pageNumber="441" type="male">1♂</specimenCount>
, “
<quantity id="367799F4FFCFFFA7CA8CD2EFFDCEFD45" box="[510,583,715,741]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="441" unit="km" value="40.0">40 km</quantity>
W of
<location id="F45062CAFFCFFFA7C9F8D2EFFD63FD45" LSID="urn:lsid:plazi:treatment:79268507FFCFFFA1CBE5D1DBFCE0FE0D:F45062CAFFCFFFA7C9F8D2EFFD63FD45" box="[650,746,715,741]" latitude="-32.333332" longLatPrecision="1211" longitude="125.083336" name="Caiguna" pageId="2" pageNumber="441">Caiguna</location>
, W.A.
<geoCoordinate id="94BB52D6FFCFFFA7C847D2EEFC2AFD45" box="[821,931,714,741]" degrees="125" direction="east" minutes="05" orientation="longitude" pageId="2" pageNumber="441" precision="925" value="125.083336">125°05´E</geoCoordinate>
,
<geoCoordinate id="94BB52D6FFCFFFA7C8DCD2EFFB85FD45" box="[942,1036,715,741]" degrees="32" direction="south" minutes="20" orientation="latitude" pageId="2" pageNumber="441" precision="925" value="-32.333332">32°20´S</geoCoordinate>
,
<date id="853112D1FFCFFFA7CF65D2EFFB20FD45" box="[1047,1193,715,741]" pageId="2" pageNumber="441" value="1978-09-30">
<collectingDate id="9575EB39FFCFFFA7CF65D2EFFB20FD45" box="[1047,1193,715,741]" pageId="2" pageNumber="441" value="1978-09-30">30 Sep. 1978</collectingDate>
</date>
<collectorName id="5C7A51C7FFCFFFA7CFDDD2EFFA1BFD45" box="[1199,1426,715,741]" pageId="2" pageNumber="441">M.S. &amp; B.J. Moulds</collectorName>
” (white label) | “new genus,
<determinerName id="04ECD8D5FFCFFFA7CAB6D2CBFDB0FCA9" box="[452,569,751,777]" pageId="2" pageNumber="441">P.B. Carne</determinerName>
det. 1983” (white label) | “
<taxonomicName id="368F4F92FFCFFFA7C82DD2CBFC4DFCA9" authority="Hutchinson &amp; Allsopp, 2021" authorityName="Hutchinson &amp; Allsopp" authorityYear="2021" box="[863,964,751,777]" class="Insecta" family="Dynastidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="441" phylum="Arthropoda" rank="genus" status="gen. nov.">Enracius</taxonomicName>
<taxonomicNameLabel id="D8C85578FFCFFFA7C8BBD2CBFB9DFCA9" box="[969,1044,751,777]" pageId="2" pageNumber="441" rank="genus">n. gen.</taxonomicNameLabel>
<specimenCount id="E789FF98FFCFFFA7CF6BD2CBFBA7FCA9" box="[1049,1070,751,777]" pageId="2" pageNumber="441" type="male"></specimenCount>
longipes
<taxonomicNameLabel id="D8C85578FFCFFFA7CFEAD2CBFB5AFCA9" box="[1176,1235,751,777]" pageId="2" pageNumber="441" rank="species">n. sp.</taxonomicNameLabel>
mihi
<typeStatus id="2E348AB3FFCFFFA7CE63D2CBFA15FCA9" box="[1297,1436,751,777]" pageId="2" pageNumber="441" type="paratype">PARATYPE</typeStatus>
<determinerName id="04ECD8D5FFCFFFA7CBE5D337FEDEFC8D" box="[151,343,787,813]" pageId="2" pageNumber="441">R.-P. Dechambre</determinerName>
det. 1999” (white label,
<typeStatus id="2E348AB3FFCFFFA7C903D337FD75FC8D" box="[625,764,787,813]" pageId="2" pageNumber="441" type="paratype">PARATYPE</typeStatus>
written in red) | mouthparts glued to a piece of card; in
<collectionCode id="979EACD4FFCFFFA7CE0AD337FA15FC8D" box="[1400,1436,787,813]" pageId="2" pageNumber="441">FD</collectionCode>
(images examined)
</materialsCitation>
.
</paragraph>
<paragraph id="F1303411FFCFFFA7CBB5D37EFEE1FB89" blockId="2.[151,1437,642,2037]" pageId="2" pageNumber="441">
<materialsCitation id="41E73E4CFFCFFFA7CBB5D37EFAEEFCD5" ID-GBIF-Occurrence="3413046317" box="[199,1383,858,885]" collectingDate="1978-09-30" collectionCode="ANIC" collectorName="M. S. &amp; B. J. Moulds" country="Australia" determinerName="P. B. Carne &amp; R. - P. Dechambre" latitude="-32.333332" location="Caiguna" longLatPrecision="1211" longitude="125.083336" pageId="2" pageNumber="441" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia">
<emphasis id="C3FBE803FFCFFFA7CBB5D37EFE72FCD4" bold="true" box="[199,507,858,884]" pageId="2" pageNumber="441">Other material examined.</emphasis>
<collectingCountry id="89987481FFCFFFA7C972D37FFD2BFCD5" box="[512,674,859,885]" name="Australia" pageId="2" pageNumber="441">AUSTRALIA</collectingCountry>
,
<collectingRegion id="334BFAF3FFCFFFA7C9D9D37FFC51FCD5" box="[683,984,859,885]" country="Australia" name="Western Australia" pageId="2" pageNumber="441">WESTERN AUSTRALIA</collectingRegion>
:
<specimenCount id="E789FF98FFCFFFA7C890D37FFB8FFCD5" box="[994,1030,859,885]" pageId="2" pageNumber="441" type="male">1♂</specimenCount>
, same data as paratype (
<collectionCode id="979EACD4FFCFFFA7CE67D37FFAE9FCD5" box="[1301,1376,859,885]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="2" pageNumber="441">ANIC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="41E73E4CFFCFFFA7CE00D37FFD7DFC1D" ID-GBIF-Occurrence="3413046319" collectingDate="2011-02-18" collectionCode="PMH" country="Australia" latitude="-31.64" location="Western Australia" longLatPrecision="729" longitude="129.0" pageId="2" pageNumber="441" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount id="E789FF98FFCFFFA7CE00D37FFA1CFCD5" box="[1394,1429,859,885]" pageId="2" pageNumber="441" type="female">1♀</specimenCount>
, “
<quantity id="367799F4FFCFFFA7CBD0D35BFF67FC39" box="[162,238,895,921]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="441" unit="km" value="12.0">12 km</quantity>
e[ast] of Eucla West. Aust. [
<geoCoordinate id="94BB52D6FFCFFFA7C931D35BFD2AFC39" box="[579,675,895,921]" degrees="31.64" direction="south" orientation="latitude" pageId="2" pageNumber="441" precision="555" value="-31.64">31.64°S</geoCoordinate>
,
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<collectingRegion id="334BFAF3FFCFFFA7C857D35BFC78FC39" box="[805,1009,895,921]" country="Australia" name="Western Australia" pageId="2" pageNumber="441">Western Australia</collectingRegion>
/
<collectingRegion id="334BFAF3FFCFFFA7C88BD35BFB23FC39" box="[1017,1194,895,921]" country="Australia" name="South Australia" pageId="2" pageNumber="441">South Australia</collectingRegion>
border]
<date id="853112D1FFCFFFA7CE63D35BFA15FC39" box="[1297,1436,895,921]" pageId="2" pageNumber="441" value="2011-02-18">
<collectingDate id="9575EB39FFCFFFA7CE63D35BFA15FC39" box="[1297,1436,895,921]" pageId="2" pageNumber="441" value="2011-02-18">18.Feb.2011</collectingDate>
</date>
|
<taxonomicName id="368F4F92FFCFFFA7CBD5D387FEF8FC1D" authorityName="Dechambre" authorityYear="1999" box="[167,369,931,957]" class="Insecta" family="Dynastidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="441" phylum="Arthropoda" rank="species" species="longipes">Enracius longipes</taxonomicName>
|
<collectionCode id="979EACD4FFCFFFA7CAF9D387FE41FC1D" box="[395,456,931,957]" pageId="2" pageNumber="441">PMH</collectionCode>
Coll# Dyn 0422” (
<collectionCode id="979EACD4FFCFFFA7C9D8D387FD64FC1D" box="[682,749,931,957]" pageId="2" pageNumber="441">PMH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="41E73E4CFFCFFFA7C871D387FD74FBA5" ID-GBIF-Occurrence="3413046303" collectingDate="1970-03-18" collectingDateMax="1970-03-28" collectingDateMin="1970-03-18" collectionCode="WAM" collectorName="W. A. M Reg. No &amp; M. Archer" country="Australia" county="Possibly" latitude="-31.68" location="Madura" longLatPrecision="729" longitude="127.43" municipality="Horseshoe Cave" pageId="2" pageNumber="441" specimenCount="3" specimenCount-female="3" stateProvince="Western Australia">
<specimenCount id="E789FF98FFCFFFA7C871D387FCAEFC1D" box="[771,807,931,957]" pageId="2" pageNumber="441" type="female">2♀</specimenCount>
, “
<collectingCounty id="18514C9DFFCFFFA7C832D387FC2BFC1D" box="[832,930,931,957]" pageId="2" pageNumber="441">Possibly</collectingCounty>
<specimenCount id="E789FF98FFCFFFA7C8DED387FC48FC1D" box="[940,961,931,957]" pageId="2" pageNumber="441" type="female"></specimenCount>
of
<taxonomicName id="368F4F92FFCFFFA7C89ED387FA15FC1D" authority="P. B. Carne" authorityName="P. B. Carne" box="[1004,1436,931,957]" class="Insecta" family="Dynastidae" genus="Orthocavonus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="441" phylum="Arthropoda" rank="species" species="occidentalis">Orthocavonus occidentalis P.B. Carne</taxonomicName>
det. 1981 |
<collectorName id="5C7A51C7FFCFFFA7CA67D3E3FE44FC41" box="[277,461,967,993]" pageId="2" pageNumber="441">W.A.M Reg. No</collectorName>
78/68 or 78/69 | outside N59 or
<collectingMunicipality id="1154AE6BFFCFFFA7C832D3E3FC71FC41" box="[832,1016,967,993]" pageId="2" pageNumber="441">Horseshoe Cave</collectingMunicipality>
N.E. of
<location id="F45062CAFFCFFFA7CF2BD3E3FB39FC41" LSID="urn:lsid:plazi:treatment:79268507FFCFFFA1CBE5D1DBFCE0FE0D:F45062CAFFCFFFA7CF2BD3E3FB39FC41" box="[1113,1200,967,993]" country="Australia" county="Possibly" latitude="-31.68" longLatPrecision="729" longitude="127.43" municipality="Horseshoe Cave" name="Madura" pageId="2" pageNumber="441" stateProvince="Western Australia">Madura</location>
[
<geoCoordinate id="94BB52D6FFCFFFA7CFCDD3E3FA97FC41" box="[1215,1310,967,993]" degrees="31.68" direction="south" orientation="latitude" pageId="2" pageNumber="441" precision="555" value="-31.68">31.68°S</geoCoordinate>
,
<geoCoordinate id="94BB52D6FFCFFFA7CE58D3E2FA1CFC41" box="[1322,1429,966,993]" degrees="127.43" direction="east" orientation="longitude" pageId="2" pageNumber="441" precision="555" value="127.43">127.43°E</geoCoordinate>
] W.
<collectingCountry id="89987481FFCFFFA7CBCFD3CFFEADFBA5" box="[189,292,1003,1029]" name="Australia" pageId="2" pageNumber="441">Australia</collectingCountry>
<date id="853112D1FFCFFFA7CA5ED3CFFE46FBA5" box="[300,463,1003,1029]" pageId="2" pageNumber="441" value="1970-03-18" valueMax="1970-03-28" valueMin="1970-03-18">
<collectingDate id="9575EB39FFCFFFA7CA5ED3CFFE46FBA5" box="[300,463,1003,1029]" pageId="2" pageNumber="441" value="1970-03-18" valueMax="1970-03-28" valueMin="1970-03-18">1828.iii.1970</collectingDate>
</date>
<collectorName id="5C7A51C7FFCFFFA7CAAAD3CFFDC4FBA5" box="[472,589,1003,1029]" pageId="2" pageNumber="441">M. Archer</collectorName>
et al.” (
<collectionCode id="979EACD4FFCFFFA7C9DBD3CFFD7DFBA5" box="[681,756,1003,1029]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="2" pageNumber="441" type="Museum">WAM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="41E73E4CFFCFFFA7C87BD3CFFEEDFB89" ID-GBIF-Occurrence="3413046309" collectingDate="1969-01" collectionCode="ANIC" collectorName="J. Bywater" country="Australia" latitude="-31.0" location="Rawlinna" longLatPrecision="731" longitude="125.33" pageId="2" pageNumber="441" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount id="E789FF98FFCFFFA7C87BD3CFFCA5FBA5" box="[777,812,1003,1029]" pageId="2" pageNumber="441" type="female">1♀</specimenCount>
, “10 N
<location id="F45062CAFFCFFFA7C8FBD3CFFC7DFBA5" LSID="urn:lsid:plazi:treatment:79268507FFCFFFA1CBE5D1DBFCE0FE0D:F45062CAFFCFFFA7C8FBD3CFFC7DFBA5" box="[905,1012,1003,1029]" country="Australia" latitude="-31.0" longLatPrecision="731" longitude="125.33" name="Rawlinna" pageId="2" pageNumber="441" stateProvince="Western Australia">Rawlinna</location>
[
<geoCoordinate id="94BB52D6FFCFFFA7CF71D3CFFBEAFBA5" box="[1027,1123,1003,1029]" degrees="31.00" direction="south" orientation="latitude" pageId="2" pageNumber="441" precision="555" value="-31.0">31.00°S</geoCoordinate>
,
<geoCoordinate id="94BB52D6FFCFFFA7CF1DD3CEFB52FBA5" box="[1135,1243,1002,1029]" degrees="125.33" direction="east" orientation="longitude" pageId="2" pageNumber="441" precision="555" value="125.33">125.33°E</geoCoordinate>
] WA
<date id="853112D1FFCFFFA7CE6ED3CFFA0BFBA5" box="[1308,1410,1003,1029]" pageId="2" pageNumber="441" value="1969-01">
<collectingDate id="9575EB39FFCFFFA7CE6ED3CFFA0BFBA5" box="[1308,1410,1003,1029]" pageId="2" pageNumber="441" value="1969-01">Jan 1969</collectingDate>
</date>
<collectorName id="5C7A51C7FFCFFFA7CEF8D3CFFF70FB89" pageId="2" pageNumber="441">J. Bywater</collectorName>
” (
<collectionCode id="979EACD4FFCFFFA7CA63D42BFED5FB89" box="[273,348,1039,1065]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="2" pageNumber="441">ANIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B995679AFFCFFFA6CBB5D416FC11F9A9" lastPageId="3" lastPageNumber="442" pageId="2" pageNumber="441" type="description">
<paragraph id="F1303411FFCFFFA7CBB5D416FDDFF855" blockId="2.[151,1437,642,2037]" pageId="2" pageNumber="441">
<emphasis id="C3FBE803FFCFFFA7CBB5D416FDD5FBEC" bold="true" box="[199,604,1074,1100]" pageId="2" pageNumber="441">
Description of female (
<figureCitation id="69B42894FFCFFFA7CAACD416FDC6FBEC" box="[478,591,1074,1100]" captionStart="FIGURES 57" captionStartId="4.[151,264,2000,2024]" captionTargetBox="[174,1423,460,1977]" captionTargetId="figure-154@4.[162,1426,453,1977]" captionTargetPageId="4" captionText="FIGURES 57. Enracius longipes Dechambre, 1999, female. 5, Dorsal view; 6, ventral view; 7, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5749003" httpUri="https://zenodo.org/record/5749003/files/figure.png" pageId="2" pageNumber="441">Figs. 57</figureCitation>
).
</emphasis>
Body
<quantity id="367799F4FFCFFFA7C9DFD416FCDAFBED" box="[685,851,1074,1101]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" metricValueMax="1.75" metricValueMin="1.3699999999999999" pageId="2" pageNumber="441" unit="mm" value="15.6" valueMax="17.5" valueMin="13.7">13.717.5 mm</quantity>
long, pronotum
<quantity id="367799F4FFCFFFA7CF64D417FB29FBED" box="[1046,1184,1075,1101]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="7.2" metricValueMin="6.1" pageId="2" pageNumber="441" unit="mm" value="6.65" valueMax="7.2" valueMin="6.1">6.17.2 mm</quantity>
wide at widest point, elytra
<quantity id="367799F4FFCFFFA7CB93D473FEE5FBD1" box="[225,364,1111,1137]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" metricValueMax="9.6" metricValueMin="7.8" pageId="2" pageNumber="441" unit="mm" value="8.7" valueMax="9.6" valueMin="7.8">7.89.6 mm</quantity>
wide (
<emphasis id="C3FBE803FFCFFFA7CAB2D473FE47FBD0" box="[448,462,1111,1136]" italics="true" pageId="2" pageNumber="441">n</emphasis>
= 3); dorsal surface shiny, head, pronotum, scutellum and pygidium brown, ventral surfaces and legs brown with black suffusions, elytra dark brown to black with suture brown, clypeus with pair of black paramedian maculae. Maxillary palpomeres 13 subequal in length, palpomere 4 longest and subequal to palpomeres 2 and 3 combined, tear-drop shaped and 2.5 times longer than greatest width and bearing shallow, oval sensorium on basal half; mandibles with lateral margin bearing stout setae; mentum narrow, bearing long setae over entire surface. Antenna with 9 antennomeres, club with antennomeres 8 and 9 1.5 times width of antennomere 7 and longer than antennomeres 26 combined, scape and pedicel bearing long setae, inner margin of club setose. Clypeus transverse almost trapezoidal with anterior angles rounded, anterior margin emarginate, anterior face truncate, vertical and glabrous, surface rugose, setose; clypeofrontal ridge linear, slightly raised. Frons rugose; ocular canthi angulate to clypeus, dorsal surface rugulose, anterior margin bearing stout setae. Pronotum widest basally, subparallel on basal half, angulate medially, linear and tapering to acute anterolateral angles, broadly recurved linear, transverse anterior margin, anterior margin membranous, lateral and anterior ridges complete, basal ridge obsolete across medial two-thirds, disc evenly convex, bearing fine punctures, becoming rugulose along lateral and anterior margins, anterior margin weakly raised medially, preceding small flat area and narrow impunctate midline. Scutellum sparsely micropunctate on disc. Elytral striae weakly impressed, linear-punctate, intervals abundantly micropunctate to rugulose, laterally and apical declivity coarsely rugulose, suture punctate-striate, epipleura nonemarginate, continuous and bearing lateral setae to apical calli; pygidium almost flat, glabrous, disc punctate, becoming denser and becoming finely rugulose at basolateral angles, apical ridge setose. Prosternal process long, slender, apex angled posteriorly and setose. Metasternum and metepisternum finely rugulose, bearing dense, long, light-brown setae.Abdominal sternites at midlength bearing a row of sparse setae across entire width, suture between sternites 6 and 7 wide and yellow, sternite 7 apically bearing a row of long setae and sparsely on disc. Protibia tridentate, medial and basal denticles closest; meso- and metatibia bicarinate (basal carina ill-defined in smaller specimens), apical cilia long, sharp, widely set; metafemur broad with anterior margin broadly arcuate; metatibia with concave dorsal margin apically and post-medial carina, inner surface broad and flat, metatibial inner spur 1.5 times length of outer spur, inner parallel and curved, outer spur narrow and straight, apices rounded; metatarsomeres combined 1.5 times length of metatibia.
</paragraph>
<caption id="A5F06499FFCEFFA6CBE5D53CFD37FAF4" ID-DOI="http://doi.org/10.5281/zenodo.5749001" ID-Zenodo-Dep="5749001" httpUri="https://zenodo.org/record/5749001/files/figure.png" pageId="3" pageNumber="442" startId="3.[151,264,1304,1328]" targetBox="[196,1432,179,1277]" targetPageId="3">
<paragraph id="F1303411FFCEFFA6CBE5D53CFD37FAF4" blockId="3.[151,1437,1304,1364]" pageId="3" pageNumber="442">
<emphasis id="C3FBE803FFCEFFA6CBE5D53CFEB3FA90" bold="true" box="[151,314,1304,1328]" pageId="3" pageNumber="442">FIGURES 14.</emphasis>
<taxonomicName id="368F4F92FFCEFFA6CA4DD53DFD38FA90" authority="Dechambre, 1999" authorityName="Dechambre" authorityYear="1999" box="[319,689,1304,1328]" class="Insecta" family="Dynastidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="C3FBE803FFCEFFA6CA4DD53DFE7FFA90" box="[319,502,1304,1328]" italics="true" pageId="3" pageNumber="442">Enracius longipes</emphasis>
<bibRefCitation id="951E49E0FFCEFFA6CA89D53CFD38FA90" author="Dechambre, R. - P." box="[507,689,1304,1328]" pageId="3" pageNumber="442" pagination="37 - 38" refId="ref10492" refString="Dechambre, R. - P. (1999) Enracius longipes n. gen., n. sp., nouveau Dynastidae d'Australie (Col. Scarabaeoidea Dynastidae). Bulletin de la Societe Entomologique de France, 104, 37 - 38. https: // doi. org / 10.3406 / bsef. 1999.16541" type="journal article" year="1999">Dechambre, 1999</bibRefCitation>
</taxonomicName>
, paratype male.
<emphasis id="C3FBE803FFCEFFA6C826D53CFCEEFA90" bold="true" box="[852,871,1304,1328]" pageId="3" pageNumber="442">1,</emphasis>
Dorsal view;
<emphasis id="C3FBE803FFCEFFA6C881D53CFB8FFA90" bold="true" box="[1011,1030,1304,1328]" pageId="3" pageNumber="442">2,</emphasis>
pygidium;
<emphasis id="C3FBE803FFCEFFA6CF0BD53CFB05FA90" bold="true" box="[1145,1164,1304,1328]" pageId="3" pageNumber="442">3,</emphasis>
phallobase and parameres, lateral view;
<emphasis id="C3FBE803FFCEFFA6CA6FD518FEA6FAF4" bold="true" box="[285,303,1340,1364]" pageId="3" pageNumber="442">4,</emphasis>
labels (photograph by Fabien Dupuis).
</paragraph>
</caption>
<paragraph id="F1303411FFCEFFA6CBB5D5A6FC11F9A9" blockId="3.[151,1437,1410,2013]" pageId="3" pageNumber="442">
<emphasis id="C3FBE803FFCEFFA6CBB5D5A6FE1BFA3C" bold="true" box="[199,402,1410,1436]" pageId="3" pageNumber="442">
Male (
<figureCitation id="69B42894FFCEFFA6CA66D5A6FE0AFA3C" box="[276,387,1410,1436]" captionStart="FIGURES 14" captionStartId="3.[151,264,1304,1328]" captionTargetBox="[196,1432,179,1277]" captionTargetId="figure-19@3.[156,1432,179,1277]" captionTargetPageId="3" captionText="FIGURES 14. Enracius longipes Dechambre, 1999, paratype male. 1, Dorsal view; 2, pygidium; 3, phallobase and parameres, lateral view; 4, labels (photograph by Fabien Dupuis)." figureDoi="http://doi.org/10.5281/zenodo.5749001" httpUri="https://zenodo.org/record/5749001/files/figure.png" pageId="3" pageNumber="442">Figs. 14</figureCitation>
).
</emphasis>
Description by
<bibRefCitation id="951E49E0FFCEFFA6C93BD5A6FCA9FA3C" author="Dechambre, R. - P." box="[585,800,1410,1436]" pageId="3" pageNumber="442" pagination="37 - 38" refId="ref10492" refString="Dechambre, R. - P. (1999) Enracius longipes n. gen., n. sp., nouveau Dynastidae d'Australie (Col. Scarabaeoidea Dynastidae). Bulletin de la Societe Entomologique de France, 104, 37 - 38. https: // doi. org / 10.3406 / bsef. 1999.16541" type="journal article" year="1999">Dechambre (1999)</bibRefCitation>
. Males differ from females by the following characters: bicolorous brown and black (male dark brown); protarsal claws simple (male internal claw larger and thicker than external claw); anterior margin of pronotum non-tuberculate (male bears a feeble tubercle); metatarsomeres 1.5 times length of tibia (male metatarsomeres 2.0 times length of tibia).
</paragraph>
</subSubSection>
<subSubSection id="B995679AFFCEFFA1CBB5D636FB3AFEBE" lastPageId="4" lastPageNumber="443" pageId="3" pageNumber="442" type="discussion">
<paragraph id="F1303411FFCEFFA1CBB5D636FB3AFEBE" blockId="3.[151,1437,1410,2013]" lastBlockId="4.[151,1436,151,430]" lastPageId="4" lastPageNumber="443" pageId="3" pageNumber="442">
<emphasis id="C3FBE803FFCEFFA6CBB5D636FEB2F98C" bold="true" box="[199,315,1554,1580]" pageId="3" pageNumber="442">Remarks.</emphasis>
Before his death in 1989, Carne passed some dynastines that he thought were undescribed to Dechambre (Fabien Dupuis, personal communication).
<bibRefCitation id="951E49E0FFCEFFA6C861D612FC61F9F0" author="Dechambre, R. - P." box="[787,1000,1590,1616]" pageId="3" pageNumber="442" pagination="37 - 38" refId="ref10492" refString="Dechambre, R. - P. (1999) Enracius longipes n. gen., n. sp., nouveau Dynastidae d'Australie (Col. Scarabaeoidea Dynastidae). Bulletin de la Societe Entomologique de France, 104, 37 - 38. https: // doi. org / 10.3406 / bsef. 1999.16541" type="journal article" year="1999">Dechambre (1999)</bibRefCitation>
later recognised Carne in naming the new genus as
<taxonomicName id="368F4F92FFCEFFA6CA4ED67FFE28F9D4" authorityName="Hutchinson &amp; Allsopp" authorityYear="2021" box="[316,417,1627,1652]" class="Insecta" family="Dynastidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="genus">
<emphasis id="C3FBE803FFCEFFA6CA4ED67FFE28F9D4" box="[316,417,1627,1652]" italics="true" pageId="3" pageNumber="442">Enracius</emphasis>
</taxonomicName>
. Both male specimens that make up the type series of
<taxonomicName id="368F4F92FFCEFFA6CF50D67FFB79F9D4" authorityName="Dechambre" authorityYear="1999" box="[1058,1264,1626,1652]" class="Insecta" family="Dynastidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="C3FBE803FFCEFFA6CF50D67FFB79F9D4" box="[1058,1264,1626,1652]" italics="true" pageId="3" pageNumber="442">Enracius longipes</emphasis>
</taxonomicName>
were listed by Dechambre as housed in the “coll[ection] S. Gaudaire-Thore” (
<bibRefCitation id="951E49E0FFCEFFA6C818D65AFBBBF938" author="Dechambre, R. - P." box="[874,1074,1662,1688]" pageId="3" pageNumber="442" pagination="37 - 38" refId="ref10492" refString="Dechambre, R. - P. (1999) Enracius longipes n. gen., n. sp., nouveau Dynastidae d'Australie (Col. Scarabaeoidea Dynastidae). Bulletin de la Societe Entomologique de France, 104, 37 - 38. https: // doi. org / 10.3406 / bsef. 1999.16541" type="journal article" year="1999">Dechambre 1999</bibRefCitation>
), which was at Étigny (Yonne),
<collectingCountry id="89987481FFCEFFA6CBE5D687FF6AF91D" box="[151,227,1699,1725]" name="France" pageId="3" pageNumber="442">France</collectingCountry>
(
<bibRefCitation id="951E49E0FFCEFFA6CB84D686FE43F91C" author="Dechambre, R. - P." box="[246,458,1698,1724]" pageId="3" pageNumber="442" pagination="245 - 250" refId="ref10550" refString="Dechambre, R. - P. (2003 a) Le genre Onychionyx Arrow, 1914 (Coleoptera, Dynastidae). Coleopteres, 9, 245 - 250." type="journal article" year="2003">Dechambre 2003a</bibRefCitation>
,
<bibRefCitation id="951E49E0FFCEFFA6CAA8D686FDABF91D" author="Dechambre, R. - P." box="[474,546,1698,1725]" pageId="3" pageNumber="442" pagination="377 - 382" refId="ref10581" refString="Dechambre, R. - P. (2003 b) Contribution a la connaissance des Pentodontini australiens (Coleoptera, Dynastidae). Coleopteres, 9, 377 - 382." type="journal article" year="2003">2003b</bibRefCitation>
) and had specimens named from it by
<bibRefCitation id="951E49E0FFCEFFA6C881D686FB2AF91C" author="Endrodi, S." box="[1011,1187,1698,1724]" pageId="3" pageNumber="442" pagination="37 - 42" refId="ref10679" refString="Endrodi, S. (1980) Sechs neue Dynastinen-Arten aus Amerika und Borneo (Coleoptera: Dynastinae). Folia Entomologica Hungarica, 41 (33), 37 - 42." type="journal article" year="1980">Endrődi (1980)</bibRefCitation>
(including
<taxonomicName id="368F4F92FFCEFFA6CE5ED686FE61F940" authority="Endrodi, 1980" authorityName="Endrodi" authorityYear="1980" class="Insecta" family="Dynastidae" genus="Aspidolea" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="gaudairethorei">
<emphasis id="C3FBE803FFCEFFA6CE5ED686FEB4F940" italics="true" pageId="3" pageNumber="442">Aspidolea gaudairethorei</emphasis>
<bibRefCitation id="951E49E0FFCEFFA6CA36D6E2FE61F940" author="Endrodi, S." box="[324,488,1734,1760]" pageId="3" pageNumber="442" pagination="37 - 42" refId="ref10679" refString="Endrodi, S. (1980) Sechs neue Dynastinen-Arten aus Amerika und Borneo (Coleoptera: Dynastinae). Folia Entomologica Hungarica, 41 (33), 37 - 42." type="journal article" year="1980">Endrődi, 1980</bibRefCitation>
</taxonomicName>
). The
<typeStatus id="2E348AB3FFCEFFA6C95CD6E2FD19F940" box="[558,656,1734,1760]" pageId="3" pageNumber="442" type="holotype">holotype</typeStatus>
at least should have been returned to ANIC but it is not there (https:// www.csiro.au/en/Research/Collections/ANIC/Collection-Resources/ANIC-primary_types, accessed
<date id="853112D1FFCEFFA6CE69D6CEFF5AF888" pageId="3" pageNumber="442" value="2021-08-19">19 August 2021</date>
). A third specimen collected at the same time was retained in ANIC and is, presumably, the one illustrated by
<bibRefCitation id="951E49E0FFCEFFA6CBE5D716FEDFF8EC" author="Weir, T. A. &amp; Lawrence, J. F. &amp; Lemann, C. &amp; Gunter, N. L." box="[151,342,1842,1868]" pageId="3" pageNumber="442" pagination="1919" refId="ref11237" refString="Weir, T. A., Lawrence, J. F., Lemann, C. &amp; Gunter, N. L. (2019) Scarabaeidae: Dynastinae Macleay 1919 [sic]. In: Slipinski, A. &amp; Lawrence, J. F. (Eds.), Australian Beetles. Vol. 2. Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing, Clayton South, pp. 516 - 530." type="book chapter" year="2019">
Weir
<emphasis id="C3FBE803FFCEFFA6CBA1D717FE82F8EC" box="[211,267,1842,1868]" italics="true" pageId="3" pageNumber="442">et al.</emphasis>
(2019
</bibRefCitation>
; Plate 54I); it was loaned to and imaged by PMH. The Gaudaire-Thore collection was apparently a euphemism for part of Dechambres collection (Fabien Dupuis, Olivier Montreuil and Antoine Mantilleri, personal communications) at his family home at Étigny.After Dechambres death, most of his type specimens were transferred to the MNHN, as was his wish, but the
<taxonomicName id="368F4F92FFCEFFA6C931D7BBFC85F818" authorityName="Dechambre" authorityYear="1999" box="[579,780,1950,1976]" class="Insecta" family="Dynastidae" genus="Enracius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="C3FBE803FFCEFFA6C931D7BBFC85F818" box="[579,780,1950,1976]" italics="true" pageId="3" pageNumber="442">Enracius longipes</emphasis>
</taxonomicName>
<typeStatus id="2E348AB3FFCEFFA6C860D7BAFCFDF818" box="[786,884,1950,1976]" pageId="3" pageNumber="442" type="holotype">holotype</typeStatus>
was not part of that donation. What was left of the collection was either destroyed by dermestids or sold, and its current location(s) is unknown. However, the
<typeStatus id="2E348AB3FFCEFFA6CE4ED7E7FA15F87D" box="[1340,1436,1987,2013]" pageId="3" pageNumber="442" type="paratype">paratype</typeStatus>
was gifted to and is now housed in FD (Fabien Dupuis, personal communication). Given that we can unambiguously associate the name with a known form, there is no need for a designation of a
<typeStatus id="2E348AB3FFC9FFA1C882D09FFBC0FF75" box="[1008,1097,187,213]" pageId="4" pageNumber="443" type="neotype">neotype</typeStatus>
(
<bibRefCitation id="951E49E0FFC9FFA1CF24D09FFE73FF59" author="International Commission on Zoological Nomenclature" pageId="4" pageNumber="443" refId="ref10923" refString="International Commission on Zoological Nomenclature (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">International Commission on Zoological Nomenclature 1999</bibRefCitation>
, Article 75.3). The image of the
<typeStatus id="2E348AB3FFC9FFA1C818D0FBFC45FF59" box="[874,972,223,249]" pageId="4" pageNumber="443" type="holotype">holotype</typeStatus>
in
<bibRefCitation id="951E49E0FFC9FFA1C883D0FBFB37FF59" author="Dechambre, R. - P." box="[1009,1214,223,249]" pageId="4" pageNumber="443" pagination="37 - 38" refId="ref10492" refString="Dechambre, R. - P. (1999) Enracius longipes n. gen., n. sp., nouveau Dynastidae d'Australie (Col. Scarabaeoidea Dynastidae). Bulletin de la Societe Entomologique de France, 104, 37 - 38. https: // doi. org / 10.3406 / bsef. 1999.16541" type="journal article" year="1999">Dechambre (1999</bibRefCitation>
,
<bibRefCitation id="951E49E0FFC9FFA1CFB8D0FAFA8DFF59" author="Dechambre, R. - P." box="[1226,1284,222,249]" pageId="4" pageNumber="443" refId="ref10613" refString="Dechambre, R. - P. (2005) Dynastidae Australiens et Oceaniens. Les Coleopteres du Monde 30. Hillside Books, Canterbury, 132 pp." type="book" year="2005">2005</bibRefCitation>
) is obviously of a different specimen than the
<typeStatus id="2E348AB3FFC9FFA1C973D127FDE8FEBD" box="[513,609,259,285]" pageId="4" pageNumber="443" type="paratype">paratype</typeStatus>
or the male in ANIC (different arrangement of legs).
</paragraph>
</subSubSection>
<subSubSection id="B995679AFFC9FFA1CBB5D102FCE0FE0D" pageId="4" pageNumber="443" type="distribution">
<paragraph id="F1303411FFC9FFA1CBB5D102FCE0FE0D" blockId="4.[151,1436,151,430]" pageId="4" pageNumber="443">
<emphasis id="C3FBE803FFC9FFA1CBB5D102FE7FFEE0" bold="true" box="[199,502,294,321]" pageId="4" pageNumber="443">Distribution and habitat.</emphasis>
Known only from the southeastern area of
<collectingRegion id="334BFAF3FFC9FFA1C882D103FB33FEE1" box="[1008,1210,295,321]" country="Australia" name="Western Australia" pageId="4" pageNumber="443">Western Australia</collectingRegion>
(
<figureCitation id="69B42894FFC9FFA1CFB9D103FA96FEE1" box="[1227,1311,295,321]" captionStart="FIGURE 14" captionStartId="7.[151,250,1114,1138]" captionTargetBox="[151,1435,309,1090]" captionTargetId="figure-82@7.[151,1435,309,1090]" captionTargetPageId="7" captionText="FIGURE 14. Known distribution of Enracius longipes Dechambre, 1999 and E. richardsi Hutchinson &amp; Allsopp new species in Western Australia." figureDoi="http://doi.org/10.5281/zenodo.5749009" httpUri="https://zenodo.org/record/5749009/files/figure.png" pageId="4" pageNumber="443">Fig. 14</figureCitation>
) (Köppen-Geiger climate of BSk, cold semi-arid;
<bibRefCitation id="951E49E0FFC9FFA1C91ED16FFCBFFEC5" author="Beck, H. E. &amp; Zimmermann, N. E. &amp; McVicar, T. R. &amp; Vergopolan N. &amp; Berg, A. &amp; Wood, E. F." box="[620,822,331,357]" pageId="4" pageNumber="443" pagination="180214" refId="ref10347" refString="Beck, H. E., Zimmermann, N. E., McVicar, T. R., Vergopolan N., Berg, A. &amp; Wood, E. F. (2018) Present and future Koppen- Geiger climate classification maps at 1 - km resolution. Scientific Data, 5, 180214. https: // doi. org / 10.1038 / sdata. 2018.214" type="journal article" year="2018">
Beck
<emphasis id="C3FBE803FFC9FFA1C9C3D16FFD67FEC5" box="[689,750,331,357]" italics="true" pageId="4" pageNumber="443">et al.</emphasis>
2018
</bibRefCitation>
). The
<typeStatus id="2E348AB3FFC9FFA1C8F1D16FFC3AFEC5" box="[899,947,331,357]" pageId="4" pageNumber="443">type</typeStatus>
series would have been captured at light (M.S. Moulds. personal communication), and the locality is open grassland with groups of 16 small
<taxonomicName id="368F4F92FFC9FFA1CE52D14BFEA9FE0D" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="443" phylum="Tracheophyta" rank="genus">
<emphasis id="C3FBE803FFC9FFA1CE52D14BFA15FE29" box="[1312,1436,367,393]" italics="true" pageId="4" pageNumber="443">Eucalyptus</emphasis>
(Myrtaceae)
</taxonomicName>
and some sparse
<taxonomicName id="368F4F92FFC9FFA1CA97D1B7FD3CFE0E" box="[485,693,403,430]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="4" pageNumber="443" phylum="Tracheophyta" rank="genus">
<emphasis id="C3FBE803FFC9FFA1CA97D1B7FDBAFE0D" box="[485,563,403,429]" italics="true" pageId="4" pageNumber="443">Acacia</emphasis>
(Fabaceae)
</taxonomicName>
over limestone.
</paragraph>
</subSubSection>
</treatment>
</document>