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<document id="A75DBA71C4AC065DA042D58FB44F0FDF" ID-CLB-Dataset="3142" ID-DOI="10.11646/zootaxa.5072.5.2" ID-GBIF-Dataset="cdde4106-d3d5-4294-9f6c-1e82afd8d580" ID-ISSN="1175-5326" ID-Zenodo-Dep="5748997" ID-ZooBank="B1D30BB4-8CAB-419E-98DE-02D97726137C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638425477373" checkinUser="plazi" docAuthor="Hutchinson, Paul M. &amp; Allsopp, Peter G." docDate="2021" docId="79268507FFCAFFADCBE5D6A9FD6BFEC5" docLanguage="en" docName="zootaxa.5072.5.2.pdf" docOrigin="Zootaxa 5072 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anomalomorpha Arrow 1908" docType="treatment" docVersion="7" lastPageNumber="446" masterDocId="851FFD7FFFCDFFA5CB72D024FF89FFA0" masterDocTitle="Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina)" masterLastPageNumber="462" masterPageNumber="439" pageNumber="446" updateTime="1699471722501" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="03419C51149CA81278EEF6CB2F5442C3">Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina)</mods:title>
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<mods:namePart id="0D249F8B0DEF012B68B608E89ACC3377">Hutchinson, Paul M.</mods:namePart>
<mods:affiliation id="92932546ACF977183BDF49A9D262CAB7">Department of Primary Industries and Regional Development (Quarantine WA), Locked Bag 69, Welshpool DC, Western Australia 6986, Australia.</mods:affiliation>
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<mods:namePart id="87B422184F919992E2014534FB3B4A01">Allsopp, Peter G.</mods:namePart>
<mods:affiliation id="69289C38EFC23AFA0BE76DBE91CF076F">Queensland Museum, PO Box 3300, South Brisbane, Queensland 4101, Australia.</mods:affiliation>
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<mods:title id="0CFB268D3F6C48B1A1D0FCB9825F45E4">Zootaxa</mods:title>
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<mods:date id="925C20A3819CE276443B0CE03FA9F7CA">2021</mods:date>
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<mods:number id="01F1D6098DB57A47A6E9FE6E4CE67508">2021-12-02</mods:number>
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<mods:identifier id="28177ACFB6679554586B02C9D3A68B61" type="DOI">10.11646/zootaxa.5072.5.2</mods:identifier>
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<treatment id="79268507FFCAFFADCBE5D6A9FD6BFEC5" ID-DOI="http://doi.org/10.5281/zenodo.5748919" ID-GBIF-Taxon="190874934" ID-Zenodo-Dep="5748919" LSID="urn:lsid:plazi:treatment:79268507FFCAFFADCBE5D6A9FD6BFEC5" httpUri="http://treatment.plazi.org/id/79268507FFCAFFADCBE5D6A9FD6BFEC5" lastPageId="8" lastPageNumber="446" pageId="7" pageNumber="446">
<subSubSection id="B995679AFFCAFFA2CBE5D6A9FD8EF907" box="[151,519,1677,1704]" pageId="7" pageNumber="446" type="nomenclature">
<paragraph id="F1303411FFCAFFA2CBE5D6A9FD8EF907" blockId="7.[151,519,1677,1704]" box="[151,519,1677,1704]" pageId="7" pageNumber="446">
<heading id="AA78837DFFCAFFA2CBE5D6A9FD8EF907" bold="true" box="[151,519,1677,1704]" fontSize="11" level="1" pageId="7" pageNumber="446" reason="1">
<taxonomicName id="368F4F92FFCAFFA2CBE5D6A9FD8EF907" authority="Arrow, 1908" authorityName="Arrow" authorityYear="1908" box="[151,519,1677,1704]" class="Insecta" family="Dynastidae" genus="Anomalomorpha" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="446" phylum="Arthropoda" rank="genus">
<emphasis id="C3FBE803FFCAFFA2CBE5D6A9FD8EF907" bold="true" box="[151,519,1677,1704]" pageId="7" pageNumber="446">
<emphasis id="C3FBE803FFCAFFA2CBE5D6A9FEEDF908" bold="true" box="[151,356,1677,1704]" italics="true" pageId="7" pageNumber="446">Anomalomorpha</emphasis>
<bibRefCitation id="951E49E0FFCAFFA2CA1ED6A9FD8EF907" author="Arrow, G. J." box="[364,519,1677,1703]" pageId="7" pageNumber="446" pagination="321 - 358" refId="ref10294" refString="Arrow, G. J. (1908) A contribution to the classification of the coleopterous family Dynastidae. Transactions of the Entomological Society of London, 1908, 321 - 358. https: // doi. org / 10.1111 / j. 1365 - 2311.1908. tb 02151. x" type="journal article" year="1908">Arrow, 1908</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B995679AFFCAFFA2CBE5D6F6FE9BF8A8" pageId="7" pageNumber="446" type="reference_group">
<paragraph id="F1303411FFCAFFA2CBE5D6F6FE9BF8A8" blockId="7.[151,1436,1746,1800]" pageId="7" pageNumber="446">
<treatmentCitationGroup id="D19F133FFFCAFFA2CBE5D6F6FE9BF8A8" pageId="7" pageNumber="446">
<treatmentCitation id="702E1200FFCAFFA2CBE5D6F6FE77F94A" author="Arrow, G. J." box="[151,510,1746,1770]" page="344" pageId="7" pageNumber="446" year="1908">
<taxonomicName id="368F4F92FFCAFFA2CBE5D6F6FE77F94A" authority="Arrow, 1908: 344" authorityName="Arrow" authorityPageNumber="344" authorityYear="1908" box="[151,510,1746,1770]" class="Insecta" family="Dynastidae" genus="Anomalomorpha" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="446" phylum="Arthropoda" rank="genus">
<emphasis id="C3FBE803FFCAFFA2CBE5D6F6FEC8F94A" box="[151,321,1746,1770]" italics="true" pageId="7" pageNumber="446">Anomalomorpha</emphasis>
<bibRefCitation id="951E49E0FFCAFFA2CA35D6F6FE77F94A" author="Arrow, G. J." box="[327,510,1746,1770]" pageId="7" pageNumber="446" pagination="321 - 358" refId="ref10294" refString="Arrow, G. J. (1908) A contribution to the classification of the coleopterous family Dynastidae. Transactions of the Entomological Society of London, 1908, 321 - 358. https: // doi. org / 10.1111 / j. 1365 - 2311.1908. tb 02151. x" type="journal article" year="1908">Arrow, 1908: 344</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="2E348AB3FFCAFFA2C97AD6F6FDB3F94A" box="[520,570,1746,1770]" pageId="7" pageNumber="446">Type</typeStatus>
species:
<taxonomicName id="368F4F92FFCAFFA2C9EBD6F6FBC8F94A" authority="Arrow, 1908" authorityName="Arrow" authorityYear="1908" box="[665,1089,1746,1770]" class="Insecta" family="Dynastidae" genus="Anomalomorpha" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="446" phylum="Arthropoda" rank="species" species="anthracina">
<emphasis id="C3FBE803FFCAFFA2C9EBD6F6FC30F94A" box="[665,953,1746,1770]" italics="true" pageId="7" pageNumber="446">Anomalomorpha anthracina</emphasis>
<bibRefCitation id="951E49E0FFCAFFA2C8CDD6F6FBC8F94A" author="Arrow, G. J." box="[959,1089,1746,1770]" pageId="7" pageNumber="446" pagination="321 - 358" refId="ref10294" refString="Arrow, G. J. (1908) A contribution to the classification of the coleopterous family Dynastidae. Transactions of the Entomological Society of London, 1908, 321 - 358. https: // doi. org / 10.1111 / j. 1365 - 2311.1908. tb 02151. x" type="journal article" year="1908">Arrow, 1908</bibRefCitation>
</taxonomicName>
, subsequent designation by
<bibRefCitation id="951E49E0FFCAFFA2CE12D6F6FE87F8A8" author="Carne, P. B." pageId="7" pageNumber="446" refId="ref10459" refString="Carne, P. B. (1957) Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Commonwealth Scientific and Industrial Research Organisation, Melbourne, 284 pp." type="book" year="1957">Carne (1957)</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B995679AFFCAFFADCBE5D712FD6BFEC5" lastPageId="8" lastPageNumber="447" pageId="7" pageNumber="446" type="diagnosis">
<paragraph id="F1303411FFCAFFADCBE5D712FD6BFEC5" blockId="7.[151,1436,1846,2017]" lastBlockId="8.[151,1436,151,357]" lastPageId="8" lastPageNumber="447" pageId="7" pageNumber="446">
<emphasis id="C3FBE803FFCAFFA2CBE5D712FE3AF8F0" bold="true" box="[151,435,1846,1872]" pageId="7" pageNumber="446">
Diagnosis (
<figureCitation id="69B42894FFCAFFA2CA68D712FE2EF8F0" box="[282,423,1846,1872]" captionStart-0="FIGURES 15–18" captionStart-1="FIGURES 19–22" captionStart-2="FIGURES 23–27" captionStart-3="FIGURES 28–30" captionStart-4="FIGURES 31–34" captionStart-5="FIGURES 35–37" captionStart-6="FIGURES 38–40" captionStart-7="FIGURES 41–43" captionStart-8="FIGURES 44–49" captionStartId-0="8.[151,264,1923,1947]" captionStartId-1="9.[151,264,1764,1788]" captionStartId-2="10.[151,264,1943,1967]" captionStartId-3="12.[151,264,1673,1697]" captionStartId-4="13.[151,264,1982,2006]" captionStartId-5="14.[151,264,1483,1507]" captionStartId-6="15.[151,264,1821,1845]" captionStartId-7="16.[151,264,637,661]" captionStartId-8="16.[151,264,1680,1704]" captionTargetBox-0="[151,1433,396,1900]" captionTargetBox-1="[151,1435,181,1741]" captionTargetBox-2="[272,1345,200,1919]" captionTargetBox-3="[204,1369,207,1650]" captionTargetBox-4="[173,1414,921,1943]" captionTargetBox-5="[186,1394,221,1459]" captionTargetBox-6="[151,1435,741,1798]" captionTargetBox-7="[151,1435,181,613]" captionTargetBox-8="[175,1435,744,1639]" captionTargetId-0="figure-108@8.[151,1435,381,1900]" captionTargetId-1="figure-19@9.[151,1435,181,1741]" captionTargetId-2="figure-26@10.[241,1345,181,1919]" captionTargetId-3="figure-26@12.[162,1424,181,1650]" captionTargetId-4="figure-330@13.[173,1414,921,1958]" captionTargetId-5="figure-26@14.[151,1435,181,1459]" captionTargetId-6="figure-261@15.[151,1435,741,1798]" captionTargetId-7="figure-26@16.[151,1435,181,613]" captionTargetId-8="figure-53@16.[151,1435,722,1656]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="12" captionTargetPageId-4="13" captionTargetPageId-5="14" captionTargetPageId-6="15" captionTargetPageId-7="16" captionTargetPageId-8="16" captionText-0="FIGURES 15–18. Anomalomorpha anthracina Arrow, 1908, male. 15, Dorsal view; 16, ventral view; 17, head dorso-lateral view; 18, pygidium." captionText-1="FIGURES 19–22. Anomalomorpha anthracina Arrow, 1908, female. 19, Dorsal view; 20, ventral view; 21, head dorso-lateral view; 22, pygidium." captionText-2="FIGURES 23–27. Anomalomorpha monteithi Allsopp, 2000, male. 23, Dorsal view; 24, ventral view; 25, lateral view; 26, head dorso-lateral view; 27, pygidium." captionText-3="FIGURES 28–30. Anomalomorpha pilbara Hutchinson &amp; Allsopp new species, holotype male. 28, Dorsal view; 29, ventral view; 30, lateral view." captionText-4="FIGURES 31–34. Anomalomorpha pilbara Hutchinson &amp; Allsopp new species, holotype male. 31, Head dorsal view; 32, head lateral view; 33, pygidium; 34, labels and mounted aedeagus." captionText-5="FIGURES 35–37. Anomalomorpha pilbara Hutchinson &amp; Allsopp new species, allotype female. 35, Dorsal view; 36, ventral view; 37, lateral view." captionText-6="FIGURES 38–40. Anomalomorpha pilbara Hutchinson &amp; Allsopp new species, allotype female. 38, Head dorsal view; 39, head lateral view; 40, pygidium." captionText-7="FIGURES 41–43. Anomalomorpha metacoxae. 41, Anomalomorpha anthracina Arrow, 1908; 42, A. pilbara Hutchinson &amp; Allsopp new species; 43, A. monteithi Allsopp, 2000." captionText-8="FIGURES 44–49. Anomalomorpha male aedeagi. 44–45, Anomalomorpha anthracina Arrow, 1908. 44, parameres dorsal; 45, aedeagus lateral; 46–47, A. pilbara Hutchinson &amp; Allsopp new species. 46, parameres dorsal; 47, aedeagus lateral; 48–49, A. monteithi Allsopp, 2000. 48, parameres dorsal; 49, aedeagus lateral. Arrows show regions of major differences." figureDoi-0="http://doi.org/10.5281/zenodo.5749013" figureDoi-1="http://doi.org/10.5281/zenodo.5749015" figureDoi-2="http://doi.org/10.5281/zenodo.5749019" figureDoi-3="http://doi.org/10.5281/zenodo.5749021" figureDoi-4="http://doi.org/10.5281/zenodo.5749023" figureDoi-5="http://doi.org/10.5281/zenodo.5749025" figureDoi-6="http://doi.org/10.5281/zenodo.5749027" figureDoi-7="http://doi.org/10.5281/zenodo.5749029" figureDoi-8="http://doi.org/10.5281/zenodo.5749031" httpUri-0="https://zenodo.org/record/5749013/files/figure.png" httpUri-1="https://zenodo.org/record/5749015/files/figure.png" httpUri-2="https://zenodo.org/record/5749019/files/figure.png" httpUri-3="https://zenodo.org/record/5749021/files/figure.png" httpUri-4="https://zenodo.org/record/5749023/files/figure.png" httpUri-5="https://zenodo.org/record/5749025/files/figure.png" httpUri-6="https://zenodo.org/record/5749027/files/figure.png" httpUri-7="https://zenodo.org/record/5749029/files/figure.png" httpUri-8="https://zenodo.org/record/5749031/files/figure.png" pageId="7" pageNumber="446">Figs. 15–49</figureCitation>
).
</emphasis>
Body
<quantity id="367799F4FFCAFFA2C973D713FD2DF8F1" box="[513,676,1847,1874]" metricMagnitude="-2" metricUnit="m" metricValue="1.6149999999999998" metricValueMax="2.08" metricValueMin="1.15" pageId="7" pageNumber="446" unit="mm" value="16.15" valueMax="20.8" valueMin="11.5">11.5–20.8 mm</quantity>
long, pronotum and elytra reddish-brown to black, legs yellowishreddish brown (almost white in live specimens);
<taxonomicName id="368F4F92FFCAFFA2C9B2D77FFCB4F8D5" box="[704,829,1883,1909]" form="ovate" pageId="7" pageNumber="446" rank="form">form ovate</taxonomicName>
, low convex, head with recurved frontoclypeal suture often terminating in a tubercle, pronotum unarmed. Maxilla and galea reduced to small conical pieces, unarmed, terminated by pencils of sensory setae; mentum flat, ligula apex rounded, palps slender, elongate and conspicuous; mandibles obscured by clypeus, external margin rounded. Antennae with 10 antennomeres, club with 3 lamellae and shorter than length of shaft. Clypeus subrectangular; clypeofrontal suture posteriorly angulate. Pronotum with anterior margin unarmed, surface evenly convex, lateral margins guttered. Elytra with lateral margins guttered, epipleural plane in anterior half horizontal. Males with all tarsi elongate, claws enlarged, robust and slightly curved; protarsomere 5 longer than protarsomere 1; proclaws of male symmetrical; metafemur of male and female long, slender, protruding beyond elytra; metatibia of female long and slender. Parameres with internal margins untoothed, external margins preapically angulate, apices setose.
</paragraph>
</subSubSection>
</treatment>
</document>
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