169 lines
19 KiB
XML
169 lines
19 KiB
XML
<document id="8BE0DA67F345232A5A2998C6AC9BF7CE" ID-DOI="10.11646/zootaxa.3641.2.1" ID-GBIF-Dataset="eaba4320-60af-4d1d-9f66-73544dc88017" ID-ISSN="1175-5326" ID-Zenodo-Dep="217798" ID-ZooBank="D6ADAE47-BD2E-444D-B1BD-686EE2273A9D" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460595075873" checkinUser="plazi" docAuthor="Altuna, Álvaro" docDate="2013" docId="7911FA2B1068FFB6FF1E72C491DB8CC4" docLanguage="en" docName="zt03641p128.pdf" docOrigin="Zootaxa 3641 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Deltocyathus moseleyi Cairns 1979" docType="treatment" docVersion="6" lastPageNumber="111" masterDocId="852882531062FFBCFF89704B93328B01" masterDocTitle="Scleractinia (Cnidaria: Anthozoa) from ECOMARG 2003, 2008 and 2009 expeditions to bathyal waters off north and northwest Spain (northeast Atlantic)" masterLastPageNumber="128" masterPageNumber="101" pageNumber="111" updateTime="1698326665948" updateUser="plazi">
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<mods:title id="54228785A4219CB732FC0AD49079A7F9">Scleractinia (Cnidaria: Anthozoa) from ECOMARG 2003, 2008 and 2009 expeditions to bathyal waters off north and northwest Spain (northeast Atlantic)</mods:title>
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<mods:name id="FE9001F4A7B9D6461251500B273B35CA" type="personal">
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<mods:roleTerm id="DE9C3ED342BB15ED9AC42BEAD053C8B4">Author</mods:roleTerm>
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<mods:namePart id="E916636AC4340F36CA6D0A1D3BC73531">Altuna, Álvaro</mods:namePart>
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<mods:typeOfResource id="0CCF36C1258169CAE1FE1283E94AE91E">text</mods:typeOfResource>
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<mods:title id="2A480253646719D7A9BFBB5CCFAD54D5">Zootaxa</mods:title>
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<mods:part id="0C0E60C4870684BDC8C30888B36CBC9D">
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<mods:date id="087320EB69881D1E815B3919070B0714">2013</mods:date>
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<mods:detail id="587A0DD8ABE4461B255A83BDCDAD0DC9" type="volume">
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<mods:number id="83CA54ADD8F2B03A2B25C1B60ABB17E5">3641</mods:number>
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<mods:detail id="6FB13C7B4833CA18556EA0F7D1B1F202" type="issue">
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<mods:number id="FD78B14E8CF137BF955E46300780E1DF">2</mods:number>
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<mods:classification id="68A60249964A9E1F0A6F6F8274D3B000">journal article</mods:classification>
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<mods:identifier id="A6A8E09599A3C9AA2DC223F3C4C53F03" type="DOI">10.11646/zootaxa.3641.2.1</mods:identifier>
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<mods:identifier id="12A04490A735D66BC8B6BAD046216785" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="8C71BD0C6B066F76F6A7E564F5AA21AE" type="Zenodo-Dep">217798</mods:identifier>
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<mods:identifier id="ADD3CCC9005FC19AA3F75C392DAEF003" type="ZooBank">D6ADAE47-BD2E-444D-B1BD-686EE2273A9D</mods:identifier>
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<treatment id="7911FA2B1068FFB6FF1E72C491DB8CC4" ID-DOI="http://doi.org/10.5281/zenodo.6146151" ID-GBIF-Taxon="119566459" ID-Zenodo-Dep="6146151" LSID="urn:lsid:plazi:treatment:7911FA2B1068FFB6FF1E72C491DB8CC4" httpUri="http://treatment.plazi.org/id/7911FA2B1068FFB6FF1E72C491DB8CC4" lastPageNumber="111" pageId="10" pageNumber="111">
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<subSubSection id="B9A218B61068FFB6FF1E72C49736886D" pageId="10" pageNumber="111" type="nomenclature">
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<paragraph id="F1074B3D1068FFB6FF1E72C4917B89A8" blockId="10.[151,585,655,714]" box="[151,585,655,681]" pageId="10" pageNumber="111">
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<heading id="AA4FFC511068FFB6FF1E72C4917B89A8" bold="true" box="[151,585,655,681]" fontSize="11" level="1" pageId="10" pageNumber="111" reason="1">
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<taxonomicName id="36B830BE1068FFB6FF1E72C4917B89A8" authority="Cairns, 1979" authorityName="Cairns" authorityYear="1979" box="[151,585,655,681]" class="Anthozoa" family="Caryophylliidae" genus="Deltocyathus" kingdom="Animalia" order="Scleractinia" pageId="10" pageNumber="111" phylum="Cnidaria" rank="species" species="moseleyi">
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<emphasis id="C3CC972F1068FFB6FF1E72C4917B89A8" bold="true" box="[151,585,655,681]" pageId="10" pageNumber="111">
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<emphasis id="C3CC972F1068FFB6FF1E72C4929689A8" bold="true" box="[151,420,655,681]" italics="true" pageId="10" pageNumber="111">Deltocyathus moseleyi</emphasis>
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Cairns, 1979
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="F1074B3D1068FFB6FF1E72F9929689CB" blockId="10.[151,585,655,714]" box="[151,420,690,714]" pageId="10" pageNumber="111">
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(
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<figureCitation id="698357B81068FFB6FF1672F993D989CB" box="[159,235,690,714]" captionStart="FIGURE 3" captionStartId="9.[151,250,1854,1877]" captionTargetBox="[252,1334,1135,1828]" captionTargetId="figure@9.[252,1334,1135,1829]" captionTargetPageId="9" captionText="FIGURE 3. Deltocyathus eccentricus Cairns, 1979 (stn. V 4, Galicia Bank). A) Corallum in calicular view (H = 0.46 cm; GCD = 1.36 cm). B) Same specimen seen from below. The slightly irregular profile of calice is distinctive. Deltocyathus moseleyi Cairns, 1979. C) Corallum in calicular view (H = 0.64 cm; GCD = 1.45 cm, stn. R 2, Galicia Bank). D) Same specimen seen from below. Notice regeneration in D. E) Corallum in calicular view (H = 0.4 cm; GCD = 1.16 cm, stn. V 5, Le Danois Bank). F) Same specimen seen from below. Scale bar: A – B = 0.7 cm; C – D = 0.7 cm; E – F = 0.5 cm." httpUri="https://zenodo.org/record/217801/files/figure.png" pageId="10" pageNumber="111">Figs. 3</figureCitation>
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C–F, tables 1–3)
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</paragraph>
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<paragraph id="F1074B3D1068FFB6FF1E72B19736886D" blockId="10.[151,1436,762,877]" pageId="10" pageNumber="111">
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<taxonomicName id="36B830BE1068FFB6FF1E72B191098811" authority="Cairns, 1979: 100" authorityName="Cairns" authorityPageNumber="100" authorityYear="1979" box="[151,571,762,784]" class="Anthozoa" family="Caryophylliidae" genus="Deltocyathus" kingdom="Animalia" order="Scleractinia" pageId="10" pageNumber="111" phylum="Cnidaria" rank="species" species="moseleyi">
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<emphasis id="C3CC972F1068FFB6FF1E72B1924E8811" box="[151,380,762,784]" italics="true" pageId="10" pageNumber="111">Deltocyathus moseleyi</emphasis>
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Cairns, 1979: 100
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</taxonomicName>
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, pl. 18, figs. 1−3.—Zibrowius 1980: 87, pl. 42, figs. A–K, pl. 43, figs. A–L.—Ramil Blanco & Fernández Pulpeiro 1990: 27.—Álvarez Claudio 1993: 417.—Álvarez Claudio 1994c: 464.—Cairns & Chapman 2001: 35 (tab. 1).—Brito & Ocaña 2004: 383, pl. 82, figs. A–C.—Reveillaud
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<emphasis id="C3CC972F1068FFB6FC777372971E884F" box="[1022,1068,824,846]" italics="true" pageId="10" pageNumber="111">et al</emphasis>
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. 2008: 322 (tab. 1), 325 (fig. 4).— Kitahara & Cairns 2009: 235 (tab. 1).—Altuna 2010: 21.—Louzao
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<emphasis id="C3CC972F1068FFB6FCE4731390AA886C" box="[877,920,855,877]" italics="true" pageId="10" pageNumber="111">et al</emphasis>
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. 2010: S1.
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</paragraph>
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</subSubSection>
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<subSubSection id="B9A218B61068FFB6FF1E73D296B588D4" pageId="10" pageNumber="111" type="materials_examined">
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<paragraph id="F1074B3D1068FFB6FF1E73D296B588D4" blockId="10.[151,1437,921,1989]" pageId="10" pageNumber="111">
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<emphasis id="C3CC972F1068FFB6FF1E73D2924988B3" bold="true" box="[151,379,921,946]" pageId="10" pageNumber="111">Material examined</emphasis>
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. Le Danois Bank, 2003: Stn.
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<date id="85066DFD1068FFB6FD5873D291C488B0" box="[721,758,921,945]" pageId="10" pageNumber="111">
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V
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<quantity id="3640E6D81068FFB6FD6173D291C488B0" box="[744,758,921,945]" metricMagnitude="3" metricUnit="m" metricValue="5.544" pageId="10" pageNumber="111" unit="m" value="5544.0">5</quantity>
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</date>
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, 544 m, three dead specimens. Le Danois Bank, 2008: Stn. D
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<date id="85066DFD1068FFB6FF2273F693FF88D4" box="[171,205,957,981]" pageId="10" pageNumber="111">6V</date>
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,
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<quantity id="3640E6D81068FFB6FF5173F6921388D4" box="[216,289,957,981]" metricMagnitude="2" metricUnit="m" metricValue="5.55" pageId="10" pageNumber="111" unit="m" value="555.0">555 m</quantity>
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, one specimen; Stn. R1R
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<quantity id="3640E6D81068FFB6FDB573F691AD88D4" box="[572,671,957,981]" metricMagnitude="3" metricUnit="m" metricValue="2.751" pageId="10" pageNumber="111" unit="m" value="2751.0">2, 751 m</quantity>
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, one dead specimen. Galicia Bank: Stn. R
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<quantity id="3640E6D81068FFB6FBF173F697E988D4" box="[1144,1243,957,981]" metricMagnitude="3" metricUnit="m" metricValue="2.614" pageId="10" pageNumber="111" unit="m" value="2614.0">2, 614 m</quantity>
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, one specimen.
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</paragraph>
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</subSubSection>
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<subSubSection id="B9A218B61068FFB6FF4E73AA97C68E18" pageId="10" pageNumber="111" type="description">
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<paragraph id="F1074B3D1068FFB6FF4E73AA97C68E18" blockId="10.[151,1437,921,1989]" pageId="10" pageNumber="111">
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<emphasis id="C3CC972F1068FFB6FF4E73AA926B88FB" bold="true" box="[199,345,993,1018]" pageId="10" pageNumber="111">Description.</emphasis>
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Corallum unattached and solitary, discoid, small. Largest specimen examined has GCD=
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<quantity id="3640E6D81068FFB6FACB73AA96A988F8" box="[1346,1435,993,1017]" metricMagnitude="-2" metricUnit="m" metricValue="1.44" pageId="10" pageNumber="111" unit="cm" value="1.44">1.44 cm</quantity>
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and H=
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<quantity id="3640E6D81068FFB6FF7A744E920D8F1C" box="[243,319,1029,1053]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="111" unit="cm" value="0.6">0.6 cm</quantity>
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(stn. R2), but coralla are normally smaller (H=
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<quantity id="3640E6D81068FFB6FCD7744E90CF8F1C" box="[862,1021,1029,1053]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.699999999999999" metricValueMin="4.0" pageId="10" pageNumber="111" unit="cm" value="0.435" valueMax="0.47" valueMin="0.4">0.40−0.47 cm</quantity>
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; GCD= 1.28–
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<quantity id="3640E6D81068FFB6FB13744E97C58F1C" box="[1178,1271,1029,1053]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="10" pageNumber="111" unit="cm" value="1.16">1.16 cm</quantity>
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, stn.
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<date id="85066DFD1068FFB6FAA5744E96638F1C" box="[1324,1361,1029,1053]" pageId="10" pageNumber="111">V5</date>
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). Base almost hemispherical, with a central scar, costate. Costae arranged according to the formula C1−2>C3>C4, reaching the central scar, prominent, well individualized by neat grooves that are deeper near calicular margin, finely granulated. Calices circular.
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<collectingCountry id="89AF0BAD1068FFB6FDCF743A91B68F88" box="[582,644,1137,1161]" name="Spain" pageId="10" pageNumber="111">Septa</collectingCountry>
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arranged hexamerally in six systems and four complete cycles (S1≥S2>S3>S4, 48 septa); S1−S3 reach de columella; S1 and S2 subequal in length, slightly longer than S3, and S3 than S4. S1−S3 paliferous, with P1−P3 separated from the septa by a small notch and forming three crowns; P3 tend to join to P2 near columella.
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<collectingCountry id="89AF0BAD1068FFB6FDA5749691588FF4" box="[556,618,1245,1269]" name="Spain" pageId="10" pageNumber="111">Septa</collectingCountry>
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and pali profusely granulated. Fossa very shallow, containing a small columella formed by a few straight rods. Corallum white (stn. D
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<date id="85066DFD1068FFB6FCEE754A90BA8E18" box="[871,904,1281,1305]" pageId="10" pageNumber="111">6V</date>
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, R2) to light brown (stn. R1R2).
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</paragraph>
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</subSubSection>
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<subSubSection id="B9A218B61068FFB6FF40756E91DB8CC4" pageId="10" pageNumber="111" type="discussion">
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<paragraph id="F1074B3D1068FFB6FF40756E96268E84" blockId="10.[151,1437,921,1989]" pageId="10" pageNumber="111">
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<emphasis id="C3CC972F1068FFB6FF40756E920F8E3C" bold="true" box="[201,317,1317,1341]" pageId="10" pageNumber="111">Remarks.</emphasis>
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The size of the specimen from stn. R2 (GCD=
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<quantity id="3640E6D81068FFB6FCD9756E90998E3C" box="[848,939,1317,1341]" metricMagnitude="-2" metricUnit="m" metricValue="1.44" pageId="10" pageNumber="111" unit="cm" value="1.44">1.44 cm</quantity>
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) is close to the biggest coralla known so far. Specimen from stn. R1R2 has 52 septa and 52 costae, indicating an incomplete S5. Due to the profuse ornamentation, and the closeness of septa and pali, calices appear particularly ‘full’ of radial elements.
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</paragraph>
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<paragraph id="F1074B3D1068FFB6FF4075DA97AA8C10" blockId="10.[151,1437,921,1989]" pageId="10" pageNumber="111">
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<taxonomicName id="36B830BE1068FFB6FF4075DA92F58EA8" box="[201,455,1425,1449]" class="Anthozoa" family="Caryophylliidae" genus="Deltocyathus" kingdom="Animalia" order="Scleractinia" pageId="10" pageNumber="111" phylum="Cnidaria" rank="species" species="moseleyi">
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<emphasis id="C3CC972F1068FFB6FF4075DA92F58EA8" box="[201,455,1425,1449]" italics="true" pageId="10" pageNumber="111">Deltocyathus moseleyi</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="36B830BE1068FFB6FE7775D991938EA8" box="[510,673,1426,1449]" class="Anthozoa" family="Caryophylliidae" genus="Deltocyathus" kingdom="Animalia" order="Scleractinia" pageId="10" pageNumber="111" phylum="Cnidaria" rank="species" species="eccentricus">
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<emphasis id="C3CC972F1068FFB6FE7775D991938EA8" box="[510,673,1426,1449]" italics="true" pageId="10" pageNumber="111">D. eccentricus</emphasis>
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</taxonomicName>
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are very similar and were keyed out recently by Kitahara & Cairns (2009). They are patellate to discoidal, unattached and solitary, parallel-sized coralla. In the ECOMARG surveys they never have been collected at the same stations although they frequently co-occur (Zibrowius 1980).
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<collectingCountry id="89AF0BAD1068FFB6FAF4759196AE8EF3" box="[1405,1436,1498,1522]" name="American Samoa" pageId="10" pageNumber="111">As</collectingCountry>
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significant differences, specimens of
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<taxonomicName id="36B830BE1068FFB6FDBC75B5918B8D14" box="[565,697,1533,1557]" class="Anthozoa" family="Caryophylliidae" genus="Deltocyathus" kingdom="Animalia" order="Scleractinia" pageId="10" pageNumber="111" phylum="Cnidaria" rank="species" species="moseleyi">
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<emphasis id="C3CC972F1068FFB6FDBC75B5918B8D14" box="[565,697,1533,1557]" italics="true" pageId="10" pageNumber="111">D. moseleyi</emphasis>
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</taxonomicName>
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from Le Danois Bank and the Galicia Bank are massive, perfectly circular, with 48 stout septa arranged tightly and S4 well developed. On the contrary, calices of
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<taxonomicName id="36B830BE1068FFB6FB59766996408D38" box="[1232,1394,1570,1593]" class="Anthozoa" family="Caryophylliidae" genus="Deltocyathus" kingdom="Animalia" order="Scleractinia" pageId="10" pageNumber="111" phylum="Cnidaria" rank="species" species="eccentricus">
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<emphasis id="C3CC972F1068FFB6FB59766996408D38" box="[1232,1394,1570,1593]" italics="true" pageId="10" pageNumber="111">D. eccentricus</emphasis>
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</taxonomicName>
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are irregular in oral view, with septa thinner and more loosely arranged, with S4 more rudimentary. Their base is also different. Costae of
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<taxonomicName id="36B830BE1068FFB6FEF2762191328D80" box="[379,512,1641,1665]" class="Anthozoa" family="Caryophylliidae" genus="Deltocyathus" kingdom="Animalia" order="Scleractinia" pageId="10" pageNumber="111" phylum="Cnidaria" rank="species" species="moseleyi">
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<emphasis id="C3CC972F1068FFB6FEF2762191328D80" box="[379,512,1641,1665]" italics="true" pageId="10" pageNumber="111">D. moseleyi</emphasis>
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</taxonomicName>
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are rounded, finely granulated, well individualized, and set apart near margin by deep furrows. Costae of
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<taxonomicName id="36B830BE1068FFB6FE3176C5916C8DA4" box="[440,606,1678,1701]" class="Anthozoa" family="Caryophylliidae" genus="Deltocyathus" kingdom="Animalia" order="Scleractinia" pageId="10" pageNumber="111" phylum="Cnidaria" rank="species" species="eccentricus">
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<emphasis id="C3CC972F1068FFB6FE3176C5916C8DA4" box="[440,606,1678,1701]" italics="true" pageId="10" pageNumber="111">D. eccentricus</emphasis>
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</taxonomicName>
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are ridged, and they appear well individualized only near the calicular margin. The epicentre of the base is subtly pointed, whether in
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<taxonomicName id="36B830BE1068FFB6FCD176F990EE8DC8" box="[856,988,1713,1737]" class="Anthozoa" family="Caryophylliidae" genus="Deltocyathus" kingdom="Animalia" order="Scleractinia" pageId="10" pageNumber="111" phylum="Cnidaria" rank="species" species="moseleyi">
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<emphasis id="C3CC972F1068FFB6FCD176F990EE8DC8" box="[856,988,1713,1737]" italics="true" pageId="10" pageNumber="111">D. moseleyi</emphasis>
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</taxonomicName>
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there is a scar due to a transitory period of attachment to a substrate. Finally, exsertness of septa in
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<taxonomicName id="36B830BE1068FFB6FCD9769D90C78DEC" box="[848,1013,1750,1773]" class="Anthozoa" family="Caryophylliidae" genus="Deltocyathus" kingdom="Animalia" order="Scleractinia" pageId="10" pageNumber="111" phylum="Cnidaria" rank="species" species="eccentricus">
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<emphasis id="C3CC972F1068FFB6FCD9769D90C78DEC" box="[848,1013,1750,1773]" italics="true" pageId="10" pageNumber="111">D. eccentricus</emphasis>
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</taxonomicName>
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is unequal, with S1 and S2 similar, slightly more than S3, and S3 than S4. In
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<taxonomicName id="36B830BE1068FFB6FDE376B191DD8C10" box="[618,751,1785,1809]" class="Anthozoa" family="Caryophylliidae" genus="Deltocyathus" kingdom="Animalia" order="Scleractinia" pageId="10" pageNumber="111" phylum="Cnidaria" rank="species" species="moseleyi">
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<emphasis id="C3CC972F1068FFB6FDE376B191DD8C10" box="[618,751,1785,1809]" italics="true" pageId="10" pageNumber="111">D. moseleyi</emphasis>
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||
</taxonomicName>
|
||
, septa of all cycles are equally exsert.
|
||
</paragraph>
|
||
<paragraph id="F1074B3D1068FFB6FF4E7755962A8C34" blockId="10.[151,1437,921,1989]" box="[199,1304,1821,1846]" pageId="10" pageNumber="111">
|
||
<collectingCountry id="89AF0BAD1068FFB6FF4E775593D48C37" box="[199,230,1822,1846]" name="American Samoa" pageId="10" pageNumber="111">As</collectingCountry>
|
||
occasionally occurs in the species (Zibrowius 1980: 89), one specimen is regenerated (fig. 3D).
|
||
</paragraph>
|
||
<paragraph id="F1074B3D1068FFB6FF4E770A91DB8CC4" blockId="10.[151,1437,921,1989]" pageId="10" pageNumber="111">
|
||
<taxonomicName id="36B830BE1068FFB6FF4E770A92F18C58" box="[199,451,1857,1881]" class="Anthozoa" family="Caryophylliidae" genus="Deltocyathus" kingdom="Animalia" order="Scleractinia" pageId="10" pageNumber="111" phylum="Cnidaria" rank="species" species="moseleyi">
|
||
<emphasis id="C3CC972F1068FFB6FF4E770A92F18C58" box="[199,451,1857,1881]" italics="true" pageId="10" pageNumber="111">Deltocyathus moseleyi</emphasis>
|
||
</taxonomicName>
|
||
is widely distributed in an area between the Celtic Sea, Azores and Madeira (Zibrowius 1980). It is probably common in the bathyal of the southern sector of the Bay of Biscay, and has been collected in the 2003,
|
||
<date id="85066DFD1068FFB6FE8E77C2929D8CA0" box="[263,431,1929,1953]" pageId="10" pageNumber="111" value="2008" valueMax="2009">2008 and 2009</date>
|
||
ECOMARG expeditions. Nevertheless, specimens are never abundant. It was previously recorded by Zibrowius (1980) from Le Danois Bank.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |