142 lines
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142 lines
12 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.714.11503" ID-GBIF-Dataset="82cad059-f2e2-473c-9905-f2bcd7727c2a" ID-PMC="PMC5740425" ID-Pensoft-Pub="1313-2970-714-129" ID-PubMed="29290702" ID-ZBK="82FF5425D41047E5AABC2C9FCE561713" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-714-129" ModsDocOrigin="ZooKeys 714" ModsDocTitle="New species of Metachela Coquillett (Diptera, Empididae) from the Atlantic Forest, Brazil and a key to the Neotropical species" checkinTime="1510066980661" checkinUser="pensoft" docAuthor="Camara, Josenir Teixeira & Rafael, Jose Albertino" docDate="2017" docId="6689701F61B84A85565A202B225070ED" docLanguage="en" docName="ZooKeys 714: 129-140" docOrigin="ZooKeys 714" docSource="http://dx.doi.org/10.3897/zookeys.714.11503" docTitle="Metachela spinulosa Camara & Rafael, 2017, sp. n." docType="treatment" docUuid="740F1C97-E46A-40DE-8F24-97FE22ECBEF5" docUuidSource="ZooBank" docVersion="5" lastPageNumber="136" masterDocId="FFAC1421FFF74A65DC20F416FFB0DA5E" masterDocTitle="New species of Metachela Coquillett (Diptera, Empididae) from the Atlantic Forest, Brazil and a key to the Neotropical species" masterLastPageNumber="140" masterPageNumber="129" pageNumber="133" updateTime="1668165098669" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New species of Metachela Coquillett (Diptera, Empididae) from the Atlantic Forest, Brazil and a key to the Neotropical species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Camara, Josenir Teixeira</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rafael, Jose Albertino</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>714</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>129</mods:start>
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<mods:end>140</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.714.11503</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.714.11503</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-714-129</mods:identifier>
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<mods:identifier type="ZBK">82FF5425D41047E5AABC2C9FCE561713</mods:identifier>
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<mods:identifier type="ZooBank">82FF5425D41047E5AABC2C9FCE561713</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="137252916" LSID="urn:lsid:zoobank.org:act:740F1C97-E46A-40DE-8F24-97FE22ECBEF5" httpUri="http://treatment.plazi.org/id/6689701F61B84A85565A202B225070ED" lastPageId="7" lastPageNumber="136" pageId="4" pageNumber="133">
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<subSubSection pageId="4" pageNumber="133" type="nomenclature">
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<paragraph pageId="4" pageNumber="133">
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<taxonomicName LSID="http://zoobank.org/740F1C97-E46A-40DE-8F24-97FE22ECBEF5" class="Insecta" family="Empididae" genus="Metachela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metachela spinulosa" order="Diptera" pageId="4" pageNumber="133" phylum="Arthropoda" rank="species" species="spinulosa">Metachela spinulosa</taxonomicName>
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<taxonomicNameLabel pageId="4" pageNumber="133">sp. n.</taxonomicNameLabel>
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Figs 12-18, 19-23
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="134" type="type-locality">
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<paragraph pageId="5" pageNumber="134">
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<pageBreakToken pageId="5" pageNumber="134" start="start">Type-locality</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="5" pageNumber="134">
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BRAZIL, Minas Gerais,
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<normalizedToken originalValue="São">Sao</normalizedToken>
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Roque de Minas, Parque Nacional Serra da Canastra, Rio Rolador.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="134" type="type-specimen">
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<paragraph pageId="5" pageNumber="134">Type-specimen.</paragraph>
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<paragraph pageId="5" pageNumber="134">
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Holotype male, pinned, with abdomen in a microvial. Original label: "BRAZIL, MG[Minas Gerais],
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<normalizedToken originalValue="São">Sao</normalizedToken>
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Roque de Minas, Parque Nacional Serra da Canastra, Rio Rolador. Malaise, 15-18.xi.2014. J.L.Nissimian, A.L. Oliveira & A.P.M. Santos." (INPA). Paratypes. Same data as holotype (1 male, 3 females, INPA).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="135" pageId="5" pageNumber="134" type="diagnosis">
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<paragraph pageId="5" pageNumber="134">Diagnosis.</paragraph>
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<paragraph pageId="5" pageNumber="134">Scutum with a mid-longitudinal brown stripe; vein R2+3 fused to R1; male cercus with truncate apex and somewhat scattered spine-like setae; epandrium with an apicoventral acute projection; female tergite 8 with bilobate apex; female tergite 10 divided into two sclerotized plates separated by a membranous area.</paragraph>
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<paragraph pageId="5" pageNumber="134">Male (Fig. 12). Holotype: body length: 3.5 mm; wing length: 2.7 mm.</paragraph>
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<caption pageId="5" pageNumber="134">
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<paragraph pageId="5" pageNumber="134">
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Figures 12-18.
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<taxonomicName class="Insecta" family="Empididae" genus="Metachela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metachela spinulosa" order="Diptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="species" species="spinulosa">Metachela spinulosa</taxonomicName>
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sp. n., holotype male. 12 Habitus, dorsolateral view 13 Right fore femur, anteroventral view 14 Right fore tibia, anteroventral view 15 Abdomen from segments 7-11, lateral view 16 Cercus, dorsal view 17 Subepandrial sclerite and epandrium, dorsal view 18 Hypandrium and epandrium, ventral view.
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</paragraph>
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</caption>
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<paragraph pageId="6" pageNumber="135">
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<pageBreakToken pageId="6" pageNumber="135" start="start">Head</pageBreakToken>
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(Fig. 12): Dark brown to black, setae whitish. Ocellar triangle with two pairs of proclinate bristles, anterior pair stouter. Eyes iridescent black, separated on face. Occiput with scattered fine setae. Mouth parts yellow; proboscis short, right and with yellow setae. Antenna yellow, with scape and pedicel bearing distinct short ventral setulae; postpedicel approximately 2
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide; stylus very short, 0.1
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as postpedicel.
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</paragraph>
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<paragraph pageId="6" pageNumber="135">Thorax (Fig. 12): Elongate, slightly arched dorsally; scutum yellowish except for mid-longitudinal brown stripe, darker posteriorly, and brownish posterolateral spot above wing base; scutum with very small and fine yellow setae except for one notopleural, one postalar, and two pairs of small parallel scutellars; scutellum and mediotergite brown;</paragraph>
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<paragraph pageId="6" pageNumber="135">
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Legs (Fig. 12): Yellow. Fore coxa as long as distance between fore- and mid coxae, 3
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide, with some dorsoapical pale setae. Fore femur (Fig. 13) 1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as fore coxa, 3.5
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide, with anteroventral row offour spines, anteroventral row of 4 denticles, and posteroventral row of 17 denticles, with basal one stouter; anteroventral row of denticles placed on distal half and posteroventral row restricted to distal 0.9, and both rows without apical discontinuity and diverging at apex. Fore tibia (Fig. 14) ~ 0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as fore femur; with decumbent short pale setulae dorsally, denser apically. Mid femur with 2 anteroventral and 15 posteroventral spines, basal pair stouter. Hind legs slender with fine setae, except hind tibia with dorsoapical
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<normalizedToken originalValue="‘comb’">'comb'</normalizedToken>
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of short setae.
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</paragraph>
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<paragraph pageId="6" pageNumber="135">
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Wings (Figs 12 and similar to 20 of female): Membraous, veins yellowish; vein R2+3 short, fused to R1; R4+5 fork angle around 70°; R5 and M1 slightly divergent at extreme apex; cell bm+dm ending beyond apex of R1, ~ 1.4
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as cell br. Halter whitish yellow.
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</paragraph>
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<paragraph pageId="6" pageNumber="135">Abdomen (Fig. 12): Tergites and sternites 1-6 and anterior margin of tergite 7 yellowish, membranous; posterior margin of tergite 7, tergite 8 and sternite 8 brownish; sternite 8 strongly sclerotized, U-shaped with lateral side upward directed posteriorly.</paragraph>
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<paragraph pageId="6" pageNumber="135">Male terminalia: Brown. Cercus wider on basal 1/3, apex truncate in lateral view (Fig. 15), with somewhat scattered spine-like setae (Fig. 16); left and right cerci closely approximated anterodorsally (Fig. 16). Epandrium with a posterodorsal pointed projection apically (Figs 17 and 18) and distinct strong setae on outer face (Fig. 17). Hypandrium membranous medially on basal half, with strong setae (Figs 15 and 18); gonocoxal apodeme projecting anteriorly as a small protuberance. Subepandrial sclerite subrectangular, more sclerotized basally and laterally (Fig. 17). Phallus strongly sclerotized, abruptly acute apically. Ejaculatory apodeme short, trilamellar.</paragraph>
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<paragraph pageId="6" pageNumber="135">Female (Fig. 19, 20). Similar to male. Tergite 7 brown, rectangular, shorter than tergite 8 (Figs 21 and 22); tergite 8 brown, elongate, bilobate posteriorly (Fig. 22); tergite 10 light brown, divided into two sclerotized plates separated by membranous area medially (Fig. 22). Cercus light brown, apex pale (Figs 21, 22 and 23). Sternite 7 light brown, concave on posterior margin (Fig. 23); sternite 8 brown, subrectangular (Fig. 23); sternite 10 light brown, v-shaped (Fig. 23). Body length: 3.9 mm; wing length: 3.1 mm.</paragraph>
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<caption pageId="6" pageNumber="135">
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<paragraph pageId="6" pageNumber="135">
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Figures 19-24. 19-23
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<taxonomicName class="Insecta" family="Empididae" genus="Metachela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metachela spinulosa" order="Diptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="spinulosa">Metachela spinulosa</taxonomicName>
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sp. n., paratype female 19 Habitus, lateral view 20 Wing 21 Segments 6-11, lateral view 22 Tergite 7 until cercus, dorsal view 23 Segments 7-11, ventral view 24
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<taxonomicName class="Insecta" family="Empididae" genus="Metachela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metachela circumdata" order="Diptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="circumdata">Metachela circumdata</taxonomicName>
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, wing modified from
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<bibRefCitation author="Collin, JE" journalOrPublisher="British Museum (Natural History), London" pageId="8" pageNumber="137" title="Diptera of Patagonia and South Chile, Part IV Empididae." year="1933">Collin (1933)</bibRefCitation>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="135" type="geographical records">
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<paragraph pageId="6" pageNumber="135">Geographical records.</paragraph>
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<paragraph pageId="6" pageNumber="135">Brazil (Minas Gerais).</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="136" type="etymology">
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<paragraph pageId="7" pageNumber="136">
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<pageBreakToken pageId="7" pageNumber="136" start="start">Etymology</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="7" pageNumber="136">From the Latin spinosus (spine), referring to the spine-like setae on the male cercus.</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="136" type="remarks">
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<paragraph pageId="7" pageNumber="136">Remarks.</paragraph>
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<paragraph pageId="7" pageNumber="136">
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<taxonomicName class="Insecta" family="Empididae" genus="Metachela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metachela spinulosa" order="Diptera" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="spinulosa">Metachela spinulosa</taxonomicName>
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sp. n. differs from other species especially by the male cercus with scattered spine-like setae (absent in other species) and epandrium with an apicoventral acute projection (absent in other species).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |