220 lines
24 KiB
XML
220 lines
24 KiB
XML
<document ID-CLB-Dataset="118853" ID-DOI="http://dx.doi.org/10.3897/zookeys.1117.86984" ID-GBIF-Dataset="948c3237-6f37-4505-b9fe-025044edcbd0" ID-PMC="PMC9848689" ID-Pensoft-Pub="1313-2970-1117-95" ID-Pensoft-UUID="54DB7D7318935384856B59FD7D22DFF4" ID-PubMed="36761375" ID-ZooBank="C583CC7AB2AD42048FA083C49BB088EA" ModsDocID="1313-2970-1117-95" checkinTime="1660292277777" checkinUser="pensoft" docAuthor="Holzenthal, Ralph W., Blahnik, Roger J. & Rios-Touma, Blanca" docDate="2022" docId="667318E9001558BC90DB06C5AB98ED82" docLanguage="en" docName="ZooKeys 1117: 95-122" docOrigin="ZooKeys 1117" docPubDate="2022-08-11" docSource="http://dx.doi.org/10.3897/zookeys.1117.86984" docTitle="Wormaldia sumaco Holzenthal & Blahnik & Ríos-Touma 2022, sp. nov." docType="treatment" docUuid="E76ACD87-7852-417B-BD24-719118C15619" docUuidSource="ZooBank" docVersion="5" id="54DB7D7318935384856B59FD7D22DFF4" lastPageNumber="95" masterDocId="54DB7D7318935384856B59FD7D22DFF4" masterDocTitle="A new genus and new species of Ecuadorian Philopotamidae (Trichoptera)" masterLastPageNumber="122" masterPageNumber="95" pageNumber="95" updateTime="1732748262665" updateUser="ExternalLinkService">
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<mods:mods id="0AFE634054D817123003FD1E0FBBBEDC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="011A2D56E0A978408862631AA1B1EC35">
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<mods:title id="CC42E8411FBC6F5F6A697745F1D9D427">A new genus and new species of Ecuadorian Philopotamidae (Trichoptera)</mods:title>
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</mods:titleInfo>
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<mods:name id="B6A9281ADF70625B1B77CB9736CFD470" type="personal">
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<mods:roleTerm id="C53B2BDDE42993AD57B1338D1880A139">Author</mods:roleTerm>
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<mods:namePart id="540EDD581FE6D9E0EA14F1932B195E0A">Holzenthal, Ralph W.</mods:namePart>
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<mods:nameIdentifier id="336E9A1ECDFDB1B5696B6BB8DCB81B6E" type="ORCID">https://orcid.org/0000-0003-1853-6340</mods:nameIdentifier>
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<mods:affiliation id="40EC19B7EC6A75A95D737D9BA80C1C73">Department of Entomology, University of Minnesota, 1980 Folwell Avenue, 219 Hodson Hall, St. Paul, Minnesota 55108 USA</mods:affiliation>
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<mods:nameIdentifier id="F86C5FAD55DF63462CFD665E3E5066BD" type="email">holze001@umn.edu</mods:nameIdentifier>
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</mods:name>
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<mods:roleTerm id="C979FFB84A16EA7D742B47AB6060B7B6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6F4EFE5DF3ECDE4D96ABB13A5551938C">Blahnik, Roger J.</mods:namePart>
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<mods:affiliation id="63208298D2B975A939F8F80F0FD772C1">Department of Entomology, University of Minnesota, 1980 Folwell Avenue, 219 Hodson Hall, St. Paul, Minnesota 55108 USA</mods:affiliation>
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<mods:namePart id="8205FD51A571E0A5709EE6311317209D">Rios-Touma, Blanca</mods:namePart>
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<mods:nameIdentifier id="0CB04C4F657276A2294291092CDE3004" type="ORCID">https://orcid.org/0000-0002-3921-0908</mods:nameIdentifier>
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<mods:affiliation id="A9A759531694DD215571807504C7E547">Grupo de Investigacion en Biodiversidad, Medio Ambiente y Salud (BIOMAS), Facultad de Ingenierias y Ciencias Aplicadas, Via Nayon S / N, Campus UDLAPARK, CP 170503, Universidad de Las Americas, Quito, Pichincha, Ecuador</mods:affiliation>
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<mods:typeOfResource id="9A2F84B94B5EF6EA333DBB6D2219A0AC">text</mods:typeOfResource>
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<mods:title id="04D0341A851D76C8541DECBB1B1CD2AF">ZooKeys</mods:title>
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<mods:part id="BB8BE1777C53FC4BED5FF3830EA019F6">
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<mods:date id="F8C8C7DFFD1271DF32F63891E915486F">2022</mods:date>
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<mods:number id="6D040F2BE89383446F613C402CFBB0DD">2022-08-11</mods:number>
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<mods:number id="591826EB2CC387CC488A6B2F318D8D97">1117</mods:number>
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<mods:url id="8846F41D200D3124ED1DCF88544A1001">http://dx.doi.org/10.3897/zookeys.1117.86984</mods:url>
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<mods:classification id="35ECE36B3C832908C9F5B2A61E7B62BA">journal article</mods:classification>
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<mods:identifier id="635A516729ECA62132B6EADE6360D579" type="DOI">http://dx.doi.org/10.3897/zookeys.1117.86984</mods:identifier>
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<mods:identifier id="62EEC83A3559C4D3D31E82D2C0EB91FD" type="Pensoft-Pub">1313-2970-1117-95</mods:identifier>
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<subSection id="A8DDD5EA0813D9B7B62E61CF2D69DCBE" pageId="0" pageNumber="95" type="systematics">
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<treatment id="667318E9001558BC90DB06C5AB98ED82" ID-GBIF-Taxon="198975923" LSID="urn:lsid:zoobank.org:act:E76ACD87-7852-417B-BD24-719118C15619" httpUri="http://treatment.plazi.org/id/667318E9001558BC90DB06C5AB98ED82" lastPageNumber="95" pageId="0" pageNumber="95" scope_family="Philopotamidae" scope_order="Trichoptera">
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<subSubSection id="116FFFE8B1C5F336C0F50D4C2E61963C" pageId="0" pageNumber="95" type="nomenclature">
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<paragraph id="6BC62D3CD9917086DAE063DBA9D7B8DD" pageId="0" pageNumber="95">
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<taxonomicName id="9EE3ED1E4C4550C153799788823366F1" LSID="https://zoobank.org/E76ACD87-7852-417B-BD24-719118C15619" authority="Holzenthal & Blahnik & Ríos-Touma, 2022" authorityName="Holzenthal & Blahnik & Ríos-Touma" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia sumaco" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="sumaco" status="sp. nov.">Wormaldia sumaco</taxonomicName>
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<taxonomicNameLabel id="6E56A8EDC1F5496D77E0CFC64345B5EB" pageId="0" pageNumber="95">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="548FB56DC1C881B976402466CDD41BFE" pageId="0" pageNumber="95" type="description">
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<paragraph id="CB40BA5DFD608D843A19BC977676B112" pageId="0" pageNumber="95">
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<figureCitation id="AA95BF17C3B191883BF86A4E27AB5472" captionStart="Figure 15" captionStartId="F13" captionText="Figure 15. Wormaldia sumaco sp. nov. Male genitalia A segments VII-X, lateral B segments VII-IX, inferior appendage, ventral C segments VIII-X, dorsal (segment VIII offset for clarity) D phallus, lateral." figureDoi="10.3897/zookeys.1117.86984.figure15" httpUri="https://binary.pensoft.net/fig/726442" pageId="0" pageNumber="95">Figs 15</figureCitation>
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<figureCitation id="2D2366B51BBDC5C2E42409F342A52D02" captionStart="Figure 16" captionStartId="F14" captionText="Figure 16. Wormaldia sumaco sp. nov. Male wings A forewing B hind wing." figureDoi="10.3897/zookeys.1117.86984.figure16" httpUri="https://binary.pensoft.net/fig/726443" pageId="0" pageNumber="95">, 16</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="23239965A8A22ECD8098F54492751CE8" pageId="0" pageNumber="95" type="type material">
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<paragraph id="CA5512D97560A2527EF8727D1127A0E4" pageId="0" pageNumber="95">Type material.</paragraph>
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<paragraph id="343AF9874DCE759DBEA9C4A25C0EFA71" pageId="0" pageNumber="95">
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<emphasis id="EC7C7B06CFC048F2D1F1728748DD9604" bold="true" pageId="0" pageNumber="95">
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<emphasis id="F6490E2D705D2D7C1CD06090E4B121B3" italics="true" pageId="0" pageNumber="95">Holotype</emphasis>
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.
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</emphasis>
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Male (pinned). Ecuador: Napo: Wildsumaco Lodge, small stream, Coati Trail @ wooden bridge,
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<geoCoordinate id="606728671E62612D91076BCC66DCA20A" degrees="0.67433" direction="south" orientation="latitude" precision="1" value="-0.67433">0.67433°S</geoCoordinate>
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,
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<geoCoordinate id="0F99547DC38B8F1434C30172FA5CBB00" degrees="77.60260" direction="west" orientation="longitude" precision="1" value="-77.6026">77.60260°W</geoCoordinate>
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, 1420 m a.s.l., 10.iii.2020,
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<normalizedToken id="626B2DC85EC1DE54C5F15974BB2EB05B" originalValue="Ríos">Rios</normalizedToken>
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, Holzenthal, Frandsen, Pauls, Amigo, UMSP000500644 (UMSP).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C3D5C5364D8CD2A43FEA85688AD03" pageId="0" pageNumber="95" type="diagnosis">
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<paragraph id="F2EBAB1C54D7294B3DEE6C5C89A37F87" pageId="0" pageNumber="95">Diagnosis.</paragraph>
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<paragraph id="AF5B6C76A34A935649F02036DBA1E8AA" pageId="0" pageNumber="95">
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Among Neotropical species of
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<taxonomicName id="002C9AF747D7E49E66DC7A6888CF7C42" authorityName="McLachlan" authorityYear="1865" class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="genus">
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<emphasis id="80922424311D24B47E7FF1368D843921" italics="true" pageId="0" pageNumber="95">Wormaldia</emphasis>
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</taxonomicName>
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, this species is unusual in several respects and more closely resembles several North American species (
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<bibRefCitation id="E812E4E515EB231DCACF8D74FE70DE21" author="Munoz-Quesada, FJ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="95" pagination="1 - 75" publicationUrl="https://www.biotaxa.org/Zootaxa/article/view/zootaxa.1838.1.1" refId="B15" refString="Munoz-Quesada, FJ, Holzenthal, RW, 2008. Revision of the Nearctic species of the caddisfly genus Wormaldia McLachlan (Trichoptera: Philopotamidae). Zootaxa 1838: 1 - 75, https://www.biotaxa.org/Zootaxa/article/view/zootaxa.1838.1.1" title="Revision of the Nearctic species of the caddisfly genus Wormaldia McLachlan (Trichoptera: Philopotamidae)." url="https://www.biotaxa.org/Zootaxa/article/view/zootaxa.1838.1.1" volume="1838" year="2008">
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<normalizedToken id="21A40C3A99F86BEB0340A120DB71F148" originalValue="Muñoz-Quesada">Munoz-Quesada</normalizedToken>
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||
and Holzenthal 2008
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</bibRefCitation>
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) than others from the Neotropical region (
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<bibRefCitation id="96486B5469DF15CC9DF83E28A30AE3E1" DOI="https://doi.org/10.11646/zootaxa.3998.1.1" author="Munoz-Quesada, FJ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="95" pagination="1 - 138" refId="B16" refString="Munoz-Quesada, FJ, Holzenthal, RW, 2015. Revision of the Neotropical species of the caddisfly genus Wormaldia McLachlan (Trichoptera: Philopotamidae). Zootaxa 3998 (1): 1 - 138, DOI: https://doi.org/10.11646/zootaxa.3998.1.1" title="Revision of the Neotropical species of the caddisfly genus Wormaldia McLachlan (Trichoptera: Philopotamidae)." url="https://doi.org/10.11646/zootaxa.3998.1.1" volume="3998" year="2015">
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<normalizedToken id="540BC54CEA65869C1A21A0F88464E201" originalValue="Muñoz-Quesada">Munoz-Quesada</normalizedToken>
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||
and Holzenthal 2015
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</bibRefCitation>
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). Particularly unusual is the well-developed ventral projection from sternum VII (Fig.
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<figureCitation id="11DD41F391CC454230428FB4000EBDDD" captionStart="Figure 15" captionStartId="F13" captionText="Figure 15. Wormaldia sumaco sp. nov. Male genitalia A segments VII-X, lateral B segments VII-IX, inferior appendage, ventral C segments VIII-X, dorsal (segment VIII offset for clarity) D phallus, lateral." figureDoi="10.3897/zookeys.1117.86984.figure15" httpUri="https://binary.pensoft.net/fig/726442" pageId="0" pageNumber="95">15A, B</figureCitation>
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), along with the relatively simple tergum X (Fig.
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<figureCitation id="2CA835C5D92495DDECF6A35F50A6EB22" captionStart="Figure 15" captionStartId="F13" captionText="Figure 15. Wormaldia sumaco sp. nov. Male genitalia A segments VII-X, lateral B segments VII-IX, inferior appendage, ventral C segments VIII-X, dorsal (segment VIII offset for clarity) D phallus, lateral." figureDoi="10.3897/zookeys.1117.86984.figure15" httpUri="https://binary.pensoft.net/fig/726442" pageId="0" pageNumber="95">15A, C</figureCitation>
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), absence of dorsal modifications on tergum VIII, and the rather simple, digitate preanal appendages (Fig.
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<figureCitation id="C404965C33273D4C688A91AD83EF52DE" captionStart="Figure 15" captionStartId="F13" captionText="Figure 15. Wormaldia sumaco sp. nov. Male genitalia A segments VII-X, lateral B segments VII-IX, inferior appendage, ventral C segments VIII-X, dorsal (segment VIII offset for clarity) D phallus, lateral." figureDoi="10.3897/zookeys.1117.86984.figure15" httpUri="https://binary.pensoft.net/fig/726442" pageId="0" pageNumber="95">15A, B</figureCitation>
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). The phallus lacks the pair of spines typical of Neotropical species and instead has a tract of granular short spines (Fig.
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<figureCitation id="2D7D05568D70D45C15D8F94669008BBE" captionStart="Figure 15" captionStartId="F13" captionText="Figure 15. Wormaldia sumaco sp. nov. Male genitalia A segments VII-X, lateral B segments VII-IX, inferior appendage, ventral C segments VIII-X, dorsal (segment VIII offset for clarity) D phallus, lateral." figureDoi="10.3897/zookeys.1117.86984.figure15" httpUri="https://binary.pensoft.net/fig/726442" pageId="0" pageNumber="95">15D</figureCitation>
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), more typical of some North American species. Additionally, the phallus is less membranous and tapered apically than most Neotropical
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<taxonomicName id="194F264909E588C3927B7BA714946AA0" authorityName="McLachlan" authorityYear="1865" class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="genus">
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<emphasis id="A4F3EA6CDBAA3CF62F510016FB83BC0F" italics="true" pageId="0" pageNumber="95">Wormaldia</emphasis>
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</taxonomicName>
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, more resembling that of other philopotamid genera (e.g.,
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<taxonomicName id="F56AAB4CCD206C5A2443A7B77C2769A0" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Philopotamidae" genus="Chimarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarra" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="genus">
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<emphasis id="1E201E44613AB12BD54BF6163E0951A8" italics="true" pageId="0" pageNumber="95">Chimarra</emphasis>
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</taxonomicName>
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). Finally, the species lacks R2 in the hind wing and thus fork I (Fig.
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<figureCitation id="BA6352DA70014FA3FA37E0AF4C43015A" captionStart="Figure 16" captionStartId="F14" captionText="Figure 16. Wormaldia sumaco sp. nov. Male wings A forewing B hind wing." figureDoi="10.3897/zookeys.1117.86984.figure16" httpUri="https://binary.pensoft.net/fig/726443" pageId="0" pageNumber="95">16B</figureCitation>
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). In this respect it resembles
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<taxonomicName id="58B6BD92ADA1CFCDA201EB484D489AD8" lsidName="W. gabriella" pageId="0" pageNumber="95" rank="species" species="gabriella">
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<emphasis id="D3F352440CF7E972B2C2D8737EA3B258" italics="true" pageId="0" pageNumber="95">W. gabriella</emphasis>
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</taxonomicName>
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(Banks, 1930),
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<taxonomicName id="C8F3AC9F6FED71C3DA7762F00635437C" lsidName="W. lacerna" pageId="0" pageNumber="95" rank="species" species="lacerna">
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<emphasis id="AD7D86907FF10FF28800343BBC85270F" italics="true" pageId="0" pageNumber="95">W. lacerna</emphasis>
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</taxonomicName>
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Denning, 1958,
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<taxonomicName id="F11B737DA68D5224FBE3423484D639C9" lsidName="W. shawnee" pageId="0" pageNumber="95" rank="species" species="shawnee">
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<emphasis id="BBFF0F839F430B10BF34C9F64F05AB4F" italics="true" pageId="0" pageNumber="95">W. shawnee</emphasis>
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</taxonomicName>
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(Ross, 1938), and
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<taxonomicName id="3A33072AE15802A39F2DFCDD6156FE6B" lsidName="W. strota" pageId="0" pageNumber="95" rank="species" species="strota">
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<emphasis id="645FA3A2BD1F27ECD1046082BC78A794" italics="true" pageId="0" pageNumber="95">W. strota</emphasis>
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</taxonomicName>
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(Ross, 1938) in the North American fauna, which also lack the fork I in the forewing.
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</paragraph>
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<caption id="5982CDE5F3F57817C730A8EADABF097C" doi="10.3897/zookeys.1117.86984.figure15" httpUri="https://binary.pensoft.net/fig/726442" pageId="0" pageNumber="95" start="Figure 15" startId="F13">
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<paragraph id="61239BA6D90EC28033A0BBBB7C93C10C" pageId="0" pageNumber="95">
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<emphasis id="B3C849E38B85C086B6239E4AEDC3A98C" bold="true" pageId="0" pageNumber="95">Figure 15.</emphasis>
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<taxonomicName id="A55BFF86293202B43ADA302893CF0E57" authorityName="Holzenthal & Blahnik & Ríos-Touma" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia sumaco" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="sumaco">
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<emphasis id="83E672B4409A2B175771727A8C78C65F" italics="true" pageId="0" pageNumber="95">Wormaldia sumaco</emphasis>
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</taxonomicName>
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sp. nov. Male genitalia
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<emphasis id="FCBB01700BC0BB8039F2C21B19C7D3D3" bold="true" pageId="0" pageNumber="95">A</emphasis>
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segments VII-X, lateral
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<emphasis id="A097E0806C852E7D692F53A387CA313B" bold="true" pageId="0" pageNumber="95">B</emphasis>
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segments VII-IX, inferior appendage, ventral
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<emphasis id="49490A6E4856D019B59FCE99BDB11939" bold="true" pageId="0" pageNumber="95">C</emphasis>
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segments VIII-X, dorsal (segment VIII offset for clarity)
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<emphasis id="8AA9DBACA9A2AE2EFB10460AE269DBEE" bold="true" pageId="0" pageNumber="95">D</emphasis>
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phallus, lateral.
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</paragraph>
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</caption>
|
||
<caption id="63D98026862210F4B1E04B002CAAD560" doi="10.3897/zookeys.1117.86984.figure16" httpUri="https://binary.pensoft.net/fig/726443" pageId="0" pageNumber="95" start="Figure 16" startId="F14">
|
||
<paragraph id="293B2F29F72E845C4B642330D5F3252B" pageId="0" pageNumber="95">
|
||
<emphasis id="0F4FAA89BD06CE3E7B806667D769DDF6" bold="true" pageId="0" pageNumber="95">Figure 16.</emphasis>
|
||
<taxonomicName id="62C70E8B5E10E25C9C9FF626BD8B61D6" authorityName="Holzenthal & Blahnik & Ríos-Touma" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia sumaco" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="sumaco">
|
||
<emphasis id="6A1B38D82621197EDBA4A718FD42260C" italics="true" pageId="0" pageNumber="95">Wormaldia sumaco</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Male wings
|
||
<emphasis id="EED0C6053B0C098058FC4A29FE41A1B6" bold="true" pageId="0" pageNumber="95">A</emphasis>
|
||
forewing
|
||
<emphasis id="333EB933080BBB4BEFC2611BDE3598F2" bold="true" pageId="0" pageNumber="95">B</emphasis>
|
||
hind wing.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="B256788E1FB9BDACB68A30AE21993CDA" pageId="0" pageNumber="95" type="description">
|
||
<paragraph id="F3EEBF3D46BF67E4509EC064657A70CA" pageId="0" pageNumber="95">Description.</paragraph>
|
||
<paragraph id="DE9878AFBF14EA0B09DB72A2A379EB89" pageId="0" pageNumber="95">
|
||
<emphasis id="18B28E70613E22E8C5097143CCD519C7" bold="true" pageId="0" pageNumber="95">Adult.</emphasis>
|
||
Forewing length male 5.2 mm (
|
||
<emphasis id="214CDB658DC87DC20D6E8497B9098377" italics="true" pageId="0" pageNumber="95">n</emphasis>
|
||
= 1). Head brown, with yellowish setae. Antenna with underlying color yellowish, overlaid with dark brown, somewhat flattened setae, giving antennae an overall dark, but somewhat annulated appearance. Maxillary palps yellowish, with light brown setae. Labial palps yellowish, with light brown setae. Dorsum of thorax brown. Legs medium brown, spurs slightly darker; hind tibiae with rather dense brush of long setae. Forewing with medium brown setae for the most part, except base, apical third, and small spot at base of thyridial cell darker; apical forks I, II, III, IV, and V present. Hind wing translucent, with very few fine, small, brown setae; apical forks II, III, and V present.
|
||
</paragraph>
|
||
<paragraph id="9DDAE369AA47775B65C69A97CD62EAD0" pageId="0" pageNumber="95">
|
||
<emphasis id="45E305A2435172595EE68F1C4D3CB8EE" bold="true" pageId="0" pageNumber="95">Male.</emphasis>
|
||
Sternum VII with flattened, digitate, setose, posteromesal process projecting beyond middle of segment VIII; as viewed ventrally, with process subtriangular, wide basally, subacute apically, set off from segment VII by sclerotized line. Segment VIII moderate in length, both sternum and tergum unmodified. Segment IX lightly sclerotized, with evident sclerotized lines anteriorly and posteriorly, lines connected mid-laterally, converging ventrally; as viewed laterally, segment relatively short, with both anterior and posterior margins moderately, angularly projecting at ca. mid-height. Segment X, in dorsal view, simple in structure, subtriangular, wide basally, narrowed apically, apex rounded, slightly upturned, with numerous sensilla. Preanal appendage elongate, digitate; as viewed dorsally, widely separated, emerging at lateral margins of tergum X, not fused basally. Inferior appendage bi-segmented, segments subequal in length; when viewed laterally, basal segment stout, subrectangular, distinctly wider than apical segment, apical segment nearly uniform in width, slightly expanded and rounded apically; as viewed ventrally, with basal segment bulbous, apical segment much narrower and flatter, with dense patch of short spine-like setae apically, extending basally on ventral margin. Phallus, when viewed laterally, with basodorsal expansion, phallobase relatively short, uniform in width, endotheca with indistinct patch of short, granular spines.
|
||
</paragraph>
|
||
<paragraph id="0A1C9F10DB4374C2389D3956D4AF1142" pageId="0" pageNumber="95">
|
||
<emphasis id="90A2FA26E3CE4169CFA75DEA19E01257" bold="true" pageId="0" pageNumber="95">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E44AF76D5E84015663C0E853B3595FA3" pageId="0" pageNumber="95" type="etymology">
|
||
<paragraph id="4132128D26FC5A9A5132B9884BFD0D56" pageId="0" pageNumber="95">Etymology.</paragraph>
|
||
<paragraph id="3D09C62DF735631031656CFB5F856C6C" pageId="0" pageNumber="95">
|
||
Named for
|
||
<normalizedToken id="A1469900FF91964ECF9985FF522D6862" originalValue="Volcán">Volcan</normalizedToken>
|
||
Sumaco, an isolated stratovolcano in the Ecuadorian Amazon, where this species was discovered.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="25BE5892CD5156F8F64AB06454A70277" pageId="0" pageNumber="95" type="habitat">
|
||
<paragraph id="558BC9E377E15C827CB02E098C3F2DDB" pageId="0" pageNumber="95">Habitat notes.</paragraph>
|
||
<paragraph id="678CF485B43BE696C1D9945CC72E9474" pageId="0" pageNumber="95">
|
||
<taxonomicName id="FA2265F6BA84B58B3C9BB1BE65B6B80E" authorityName="Holzenthal & Blahnik & Ríos-Touma" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Sumacodella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sumacodella elongata" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="elongata">
|
||
<emphasis id="AC50EEF378AD7A8B60D9F92B16BD7FE7" italics="true" pageId="0" pageNumber="95">Sumacodella elongata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="67623F3B7A80F716F91D7252F19C244E" authorityName="Holzenthal & Blahnik & Ríos-Touma" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia natalis" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="natalis">
|
||
<emphasis id="DFC4076B41C89B4893D419A33D83F468" italics="true" pageId="0" pageNumber="95">Wormaldia natalis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="34113DE71E8F6784E7A412C9DA816C1A" authorityName="Holzenthal & Blahnik & Ríos-Touma" authorityYear="2022" class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia sumaco" order="Trichoptera" pageId="0" pageNumber="95" phylum="Arthropoda" rank="species" species="sumaco">
|
||
<emphasis id="74E320455827E695D4686B4AC3E6782C" italics="true" pageId="0" pageNumber="95">Wormaldia sumaco</emphasis>
|
||
</taxonomicName>
|
||
share the same type locality. Small permanent forest streams, similar to the type locality of these species, are common on the mid-elevation slopes of
|
||
<normalizedToken id="2E234FB81C3FCC7E1776EBB1E3F7D054" originalValue="Volcán">Volcan</normalizedToken>
|
||
Sumaco. Leaflitter is abundant and waters are clear with very low conductivity (20-55
|
||
<normalizedToken id="F379FB41381CE823CF7E93A0C33C28C3" originalValue="µS">µS</normalizedToken>
|
||
/cm), highly oxygen saturated (close to 100%), and warm (17-20 °C).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |