treatments-xml/data/5B/50/E9/5B50E916FFBD39054D2DFEA7FDDBFDFC.xml
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<document id="FB97B74DFF3FB89FD967EDD60E47749F" ID-DOI="10.5281/zenodo.155701" ID-GBIF-Dataset="b947e6c9-d6cf-4988-a098-5e01dd69a888" ID-ISSN="1175­5326" ID-Zenodo-Dep="155701" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459438614591" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2002" docId="5B50E916FFBD39054D2DFEA7FDDBFDFC" docLanguage="en" docName="zt00081.pdf" docOrigin="Zootaxa 81" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Geostiba (Sibiota) pluvigena Gusarov, sp. n." docType="treatment" docVersion="14" lastPageNumber="39" masterDocId="A769916EFF9E39234C25FFBCFFE8FFDA" masterDocTitle="A revision of Nearctic species of the genus Geostiba Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="88" masterPageNumber="1" pageNumber="36" updateTime="1698188143631" updateUser="plazi">
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<mods:title id="4C88D992B7FABA4DA50728A47F5FD3F2">A revision of Nearctic species of the genus Geostiba Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="E13AA5D89089F1A7FC1188F590ADFB72">Gusarov, Vladimir I.</mods:namePart>
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<mods:date id="C216C6EC073778D6F5BDF182DF3814A7">2002</mods:date>
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<paragraph id="D3465800FFBD39004D2DFEA7FC16FEEF" blockId="35.[264,1022,283,310]" box="[264,1022,283,310]" pageId="35" pageNumber="36">
<heading id="880EEF6CFFBD39004D2DFEA7FC16FEEF" bold="true" box="[264,1022,283,310]" fontSize="11" level="1" pageId="35" pageNumber="36" reason="1">
<keyStep id="680D40A5FFBD39004D2DFEA7FC16FEEF" box="[264,1022,283,310]" pageId="35" pageNumber="36">
<key id="A7680F31FFBD39004D2DFEA7FC16FEEF" box="[264,1022,283,310]" pageId="35" pageNumber="36">
<keyLead id="6808FB35FFBD39004D2DFEA7FC16FEEF" box="[264,1022,283,310]" pageId="35" pageNumber="36">
<emphasis id="E18D8412FFBD39004D2DFEA7FC16FEEF" bold="true" box="[264,1022,283,310]" pageId="35" pageNumber="36">
5.
<taxonomicName id="14F92383FFBD39004D03FEA0FD07FEEF" ID-CoL="3FSN3" authority="Gusarov" authorityName="Gusarov" box="[294,751,283,310]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="pluvigena" status="sp. nov." subGenus="Sibiota">
<emphasis id="E18D8412FFBD39004D03FEA0FE78FEEC" bold="true" box="[294,400,284,310]" italics="true" pageId="35" pageNumber="36">Geostiba</emphasis>
(
<emphasis id="E18D8412FFBD39004D85FEA0FE1DFEEC" bold="true" box="[416,501,284,310]" italics="true" pageId="35" pageNumber="36">Sibiota</emphasis>
)
<emphasis id="E18D8412FFBD39004E23FEA0FD95FEEC" bold="true" box="[518,637,284,310]" italics="true" pageId="35" pageNumber="36">pluvigena</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="FABE3969FFBD39004ED8FEA7FCD7FEEF" box="[765,831,283,309]" pageId="35" pageNumber="36" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="4BC24485FFBD39004F6BFEA7FC1EFEEF" box="[846,1014,283,309]" captionStart-0="FIGURES 120 ­ 125" captionStart-1="FIGURES 126 ­ 133" captionStart-2="FIGURES 134 ­ 145" captionStartId-0="35.[264,382,1745,1769]" captionStartId-1="36.[264,382,1701,1725]" captionStartId-2="38.[264,383,1559,1583]" captionTargetBox-0="[305,1280,668,1726]" captionTargetBox-1="[308,1279,632,1666]" captionTargetBox-2="[315,1274,655,1520]" captionTargetId-0="figure@35.[302,1286,668,1727]" captionTargetId-1="figure@36.[294,1294,622,1676]" captionTargetId-2="figure@38.[307,1279,649,1533]" captionTargetPageId-0="35" captionTargetPageId-1="36" captionTargetPageId-2="38" captionText-0="FIGURES 120 ­ 125. Abdominal segments 7 ­ 8 of Geostiba pluvigena Gusarov, sp. n. (paratypes from Waterrock Knob (120 ­ 121, 125) and junction of Blue Ridge Parkway and Balsam Mountain Road (122 ­ 124), North Carolina). 120 male tergum 8; 121 male sternum 8; 122 female tergum 8; 123 female sternum 8, 124 apex of female sternum 8; 125 medial portion of male tergum 7, posterior down. Scale bar 0.2 mm (120 ­ 123, 125), 0.1 mm (124)." captionText-1="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 median lobe, ventral view; 127, 131, 133 apex of median lobe, ventral view; 128 median lobe, lateral view; 129 apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." captionText-2="FIGURES 134 ­ 145. Genitalia of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (134, 139 ­ 141) and Waterrock Knob (135 ­ 138, 142 ­ 145), North Carolina). 134 details of internal sac retracted into median lobe, ventral view; 135 copulatory piece of internal sac, dorsal view; 136 medial lamellae of internal sac, ventral view; 137 distal diverticula of everted internal sac, ventral view; 138 everted internal sac, ventral view; 139 spermatheca; 140 ­ 141 details of internal sac retracted into median lobe, lateral view; 142 copulatory piece of internal sac, lateral view; 143 medial lamella of internal sac, lateral view; 144 everted internal sac, lateral view; 145 apex of left paramere, side facing median lobe. Scale bar 0.2 mm (134, 138, 140 ­ 141, 144), 0.1 mm (135 ­ 137, 139, 142 ­ 143, 145)." httpUri-0="https://zenodo.org/record/155719/files/figure.png" httpUri-1="https://zenodo.org/record/155720/files/figure.png" httpUri-2="https://zenodo.org/record/155721/files/figure.png" pageId="35" pageNumber="36">Figs. 120­145</figureCitation>
)
</emphasis>
</keyLead>
</key>
</keyStep>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9BE30B8BFFBD39004D2DFEE3FB82FEAC" box="[264,1130,351,374]" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="D3465800FFBD39004D2DFEE3FB82FEAC" blockId="35.[264,1130,351,374]" box="[264,1130,351,374]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="F3E97F2EFFBD39004D2DFEE3FB82FEAC" box="[264,1130,351,374]" pageId="35" pageNumber="36">
<taxonomicName id="14F92383FFBD39004D2DFEE3FCDCFEAC" ID-CoL="3FSKZ" authority="Lohse &amp; Smetana, 1988: 273" authorityName="Lohse &amp; Smetana" authorityPageNumber="273" authorityYear="1988" box="[264,820,351,374]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="bicarinata">
<emphasis id="E18D8412FFBD39004D2DFEE3FE36FEAC" box="[264,478,351,374]" italics="true" pageId="35" pageNumber="36">Geostiba bicarinata</emphasis>
:
<treatmentCitation id="52587E11FFBD39004DD6FEE3FCDCFEAC" author="Lohse" box="[499,820,351,374]" page="273" pageId="35" pageNumber="36" year="1988">
<bibRefCitation id="B76825F1FFBD39004DD6FEE3FD11FEAC" author="Lohse" box="[499,761,351,374]" pageId="35" pageNumber="88" refString="Lohse, G. A. &amp; Smetana, A. (1988) Four new species of Geostiba Thomson from the Appalachian mountains of North Carolina, with a key to North American species and synonymic notes (Coleoptera: Staphylinidae: Aleocharinae). The Coleopterists Bulletin, 42 (3), 269 - 278." type="journal article" year="1988">Lohse &amp; Smetana, 1988</bibRefCitation>
: 273
</treatmentCitation>
</taxonomicName>
,
<emphasis id="E18D8412FFBD39004F1AFEE3FC7DFEAC" box="[831,917,351,374]" italics="true" pageId="35" pageNumber="36">ex parte</emphasis>
(misidentification).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9BE30B8BFFBD39074D2DFE1AFBFCFDCC" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="D3465800FFBD39004D2DFE1AFE9DFDBB" blockId="35.[264,1323,422,609]" pageId="35" pageNumber="36">
<emphasis id="E18D8412FFBD39004D2DFE1AFE28FE1A" bold="true" box="[264,448,422,448]" pageId="35" pageNumber="36">
<typeStatus id="0C42E6A2FFBD39004D2DFE1AFEABFE1A" box="[264,323,422,448]" pageId="35" pageNumber="36">Type</typeStatus>
material.
</emphasis>
<typeStatus id="0C42E6A2FFBD39004DFCFE1BFDA1FE1B" box="[473,585,423,449]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
,,
<emphasis id="E18D8412FFBD39004EA5FE1AFBB0FE1A" bold="true" box="[640,1112,422,448]" pageId="35" pageNumber="36">
<collectingCountry id="ABEE1890FFBD39004EA5FE1AFC86FE1A" box="[640,878,422,448]" name="United States of America" pageId="35" pageNumber="36">UNITED STATES</collectingCountry>
: North Carolina:
</emphasis>
Haywood Co. / Jackson Co.: junction of Blue Ridge Parkway and Balsam Mountain Road,
<quantity id="1401F5E5FFBD39004891FE73FAEAFE33" box="[1204,1282,463,489]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="35" pageNumber="36" unit="km" value="17.0">17 km</quantity>
W Waynesville,
<geoCoordinate id="B6CD3EC7FFBD39004D83FE4BFDC3FDCB" box="[422,555,503,529]" direction="north" orientation="latitude" pageId="35" pageNumber="36" precision="9" value="35.495335">35°29.72'N</geoCoordinate>
<geoCoordinate id="B6CD3EC7FFBD39004E17FE4BFD57FDCB" box="[562,703,503,529]" direction="west" orientation="longitude" pageId="35" pageNumber="36" precision="9" value="-83.173836">83°10.43'W</geoCoordinate>
,
<quantity id="1401F5E5FFBD39004EEFFE4BFCCDFDCB" box="[714,805,503,529]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="35" pageNumber="36" unit="m" value="1600.0">1600 m</quantity>
, in forest litter,
<taxonomicName id="14F92383FFBD39004FFEFE4BFB9DFDCA" box="[987,1141,503,528]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="rubens">
<emphasis id="E18D8412FFBD39004FFEFE4BFB9DFDCA" box="[987,1141,503,528]" italics="true" pageId="35" pageNumber="36">Picea rubens</emphasis>
</taxonomicName>
,
<taxonomicName id="14F92383FFBD390048A6FE4BFED6FDE2" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="grandifolia">
<emphasis id="E18D8412FFBD390048A6FE4BFED6FDE2" italics="true" pageId="35" pageNumber="36">Fagus grandifolia</emphasis>
</taxonomicName>
,
<taxonomicName id="14F92383FFBD39004D75FDA3FE74FDE2" box="[336,412,543,568]" class="Magnoliopsida" family="Betulaceae" genus="Betula" kingdom="Plantae" order="Fagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="genus">
<emphasis id="E18D8412FFBD39004D75FDA3FE74FDE2" box="[336,412,543,568]" italics="true" pageId="35" pageNumber="36">Betula</emphasis>
</taxonomicName>
,
<taxonomicName id="14F92383FFBD39004D8AFDA3FDB4FDE2" box="[431,604,543,568]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="genus">
<emphasis id="E18D8412FFBD39004D8AFDA3FDB4FDE2" box="[431,604,543,568]" italics="true" pageId="35" pageNumber="36">Rhododendron</emphasis>
</taxonomicName>
,
<taxonomicName id="14F92383FFBD39004E4AFDA3FC77FDE2" box="[623,927,543,568]" class="Liliopsida" family="Asparagaceae" genus="Maianthemum" kingdom="Plantae" order="Asparagales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="species" species="canadense">
<emphasis id="E18D8412FFBD39004E4AFDA3FC77FDE2" box="[623,927,543,568]" italics="true" pageId="35" pageNumber="36">Maianthemum canadense</emphasis>
</taxonomicName>
,
<taxonomicName id="14F92383FFBD39004F94FDA3FB44FDE3" authority="V.I.Gusarov" authorityName="V.I.Gusarov" box="[945,1196,543,569]" class="Magnoliopsida" family="Oxalidaceae" genus="Oxalis" kingdom="Plantae" order="Oxalidales" pageId="35" pageNumber="36" phylum="Tracheophyta" rank="genus">
<emphasis id="E18D8412FFBD39004F94FDA3FC15FDE2" box="[945,1021,543,568]" italics="true" pageId="35" pageNumber="36">Oxalis</emphasis>
(V.I.Gusarov)
</taxonomicName>
,
<date id="A7477EC0FFBD39004899FDA3FAC3FDE3" box="[1212,1323,543,569]" pageId="35" pageNumber="36" value="2001-06-02">2.vi.2001</date>
(
<collectionCode id="B5E8C0C5FFBD39004D35FDFBFE80FDBB" box="[272,360,583,609]" pageId="35" pageNumber="36">KSEM</collectionCode>
).
</paragraph>
<caption id="87860888FFBD39004D2DF96DFBFCF8B2" httpUri="https://zenodo.org/record/155719/files/figure.png" pageId="35" pageNumber="36" targetBox="[305,1280,668,1726]" targetPageId="35">
<paragraph id="D3465800FFBD39004D2DF96DFBFCF8B2" blockId="35.[264,1323,1745,1896]" pageId="35" pageNumber="36">
<emphasis id="E18D8412FFBD39004D2DF96DFE08F933" bold="true" box="[264,480,1745,1769]" pageId="35" pageNumber="36">FIGURES 120­125</emphasis>
. Abdominal segments 7­8 of
<taxonomicName id="14F92383FFBD39004F14F96DFB8FF932" authority="Gusarov" authorityName="Gusarov" box="[817,1127,1745,1768]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="pluvigena" status="sp. nov.">
<emphasis id="E18D8412FFBD39004F14F96DFBEBF932" box="[817,1027,1745,1768]" italics="true" pageId="35" pageNumber="36">Geostiba pluvigena</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="FABE3969FFBD39004850F96DFB5AF933" box="[1141,1202,1745,1769]" pageId="35" pageNumber="36" rank="species">
<emphasis id="E18D8412FFBD39004850F96DFB5AF933" bold="true" box="[1141,1202,1745,1769]" pageId="35" pageNumber="36">sp. n.</emphasis>
</taxonomicNameLabel>
(paratypes from Waterrock Knob (120­121, 125) and junction of Blue Ridge Parkway and Balsam Mountain Road (122­124), North Carolina). 120 male tergum 8; 121 male sternum 8; 122 female tergum 8; 123 female sternum 8, 124 apex of female sternum 8; 125 medial portion of male tergum 7, posterior down. Scale bar 0.2 mm (120­123, 125), 0.1 mm (124).
</paragraph>
</caption>
<paragraph id="D3465800FFBA39074D1DFEA0FBFCFDCC" blockId="36.[264,1326,283,534]" pageId="36" pageNumber="37">
<typeStatus id="0C42E6A2FFBA39074D1DFEA0FE47FEEC" box="[312,431,284,310]" pageId="36" pageNumber="37" type="paratype">Paratypes</typeStatus>
:
<emphasis id="E18D8412FFBA39074DE1FEA7FC7BFEEF" bold="true" box="[452,915,283,309]" pageId="36" pageNumber="37">
<collectingCountry id="ABEE1890FFBA39074DE1FEA7FD58FEEF" box="[452,688,283,309]" name="United States of America" pageId="36" pageNumber="37">UNITED STATES</collectingCountry>
: North Carolina:
</emphasis>
Haywood Co. / Jackson Co.:
<specimenCount id="C5FF9389FFBA3907492BFEA0FE6DFE81" pageId="36" pageNumber="37" type="generic">99 specimens</specimenCount>
, same data as the
<typeStatus id="0C42E6A2FFBA39074E7AFEFDFD2EFE81" box="[607,710,321,347]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
(
<collectionCode id="B5E8C0C5FFBA39074EF2FEFDFCC5FE81" box="[727,813,321,347]" pageId="36" pageNumber="37">KSEM</collectionCode>
,
<collectionCode id="B5E8C0C5FFBA39074F1FFEFDFC7FFE81" LSID="urn:lsid:biocol.org:col:34925" box="[826,919,321,347]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="36" pageNumber="37">AMNH</collectionCode>
,
<collectionCode id="B5E8C0C5FFBA39074F81FEFDFC05FE81" box="[932,1005,321,347]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="36" pageNumber="37">CNCI</collectionCode>
,
<collectionCode id="B5E8C0C5FFBA39074FDEFEFDFBACFE81" box="[1019,1092,321,347]" pageId="36" pageNumber="37">SPSU</collectionCode>
,
<collectionCode id="B5E8C0C5FFBA39074874FEFDFB43FE81" LSID="urn:lsid:biocol.org:col:34795" box="[1105,1195,321,347]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="36" pageNumber="37">FMNH</collectionCode>
);
<specimenCount id="C5FF9389FFBA390748E4FEFDFEA1FE5B" pageId="36" pageNumber="37" type="generic">43 specimens</specimenCount>
, Waterrock Knob, N slope,
<quantity id="1401F5E5FFBA39074EB0FEDBFD37FE5B" box="[661,735,359,385]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="36" pageNumber="37" unit="km" value="13.0">13 km</quantity>
W Waynesville,
<geoCoordinate id="B6CD3EC7FFBA39074F8DFEDBFBF7FE5B" box="[936,1055,359,385]" direction="north" orientation="latitude" pageId="36" pageNumber="37" precision="92" value="35.466667">35°28.0'N</geoCoordinate>
<geoCoordinate id="B6CD3EC7FFBA39074803FEDBFB4CFE5B" box="[1062,1188,359,385]" direction="west" orientation="longitude" pageId="36" pageNumber="37" precision="92" value="-83.136665">83°08.2'W</geoCoordinate>
,
<quantity id="1401F5E5FFBA3907488AFEDBFAE2FE5B" box="[1199,1290,359,385]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="36" pageNumber="37" unit="m" value="1900.0">1900 m</quantity>
, in forest litter,
<taxonomicName id="14F92383FFBA39074DBEFE30FDDFFE7F" box="[411,567,396,421]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="rubens">
<emphasis id="E18D8412FFBA39074DBEFE30FDDFFE7F" box="[411,567,396,421]" italics="true" pageId="36" pageNumber="37">Picea rubens</emphasis>
</taxonomicName>
,
<taxonomicName id="14F92383FFBA39074E62FE30FD37FE7F" box="[583,735,396,421]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="fraseri">
<emphasis id="E18D8412FFBA39074E62FE30FD37FE7F" box="[583,735,396,421]" italics="true" pageId="36" pageNumber="37">Abies fraseri</emphasis>
</taxonomicName>
,
<taxonomicName id="14F92383FFBA39074ED5FE30FCD1FE7F" box="[752,825,396,421]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="genus">
<emphasis id="E18D8412FFBA39074ED5FE30FCD1FE7F" box="[752,825,396,421]" italics="true" pageId="36" pageNumber="37">Rubus</emphasis>
</taxonomicName>
,
<taxonomicName id="14F92383FFBA39074F6CFE30FBAAFE7C" authority="V.I.Gusarov" authorityName="V.I.Gusarov" box="[841,1090,396,422]" class="Magnoliopsida" family="Oxalidaceae" genus="Oxalis" kingdom="Plantae" order="Oxalidales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="genus">
<emphasis id="E18D8412FFBA39074F6CFE30FC7DFE7F" box="[841,917,396,421]" italics="true" pageId="36" pageNumber="37">Oxalis</emphasis>
(V.I.Gusarov)
</taxonomicName>
,
<date id="A7477EC0FFBA39074875FE30FB56FE7C" box="[1104,1214,396,422]" pageId="36" pageNumber="37" value="2001-06-02">2.vi.2001</date>
(
<collectionCode id="B5E8C0C5FFBA390748F5FE30FACEFE7C" box="[1232,1318,396,422]" pageId="36" pageNumber="37">KSEM</collectionCode>
,
<collectionCode id="B5E8C0C5FFBA39074D2DFE0DFEBBFE11" box="[264,339,433,459]" pageId="36" pageNumber="37">SPSU</collectionCode>
); Haywood Co.:, Blue Ridge Parkway, Browning Knob Mt. [
<geoCoordinate id="B6CD3EC7FFBA390748BBFE0DFAC2FE11" box="[1182,1322,433,459]" direction="north" orientation="latitude" pageId="36" pageNumber="37" precision="15" value="35.463055">35°27'47&quot;N</geoCoordinate>
<geoCoordinate id="B6CD3EC7FFBA39074D2DFE6BFE75FE2B" box="[264,413,471,497]" direction="west" orientation="longitude" pageId="36" pageNumber="37" precision="15" value="-83.13194">83°07'55&quot;W</geoCoordinate>
],
<quantity id="1401F5E5FFBA39074D95FE6BFDA4FE2B" box="[432,588,471,497]" metricMagnitude="3" metricUnit="m" metricValue="1.86" metricValueMax="1.89" metricValueMin="1.83" pageId="36" pageNumber="37" unit="m" value="1860.0" valueMax="1890.0" valueMin="1830.0">1830­1890 m</quantity>
(A.Smetana),
<date id="A7477EC0FFBA39074ED3FE6BFC81FE2B" box="[758,873,471,497]" pageId="36" pageNumber="37" value="1986-05-28">28.v.1986</date>
(
<bibRefCitation id="B76825F1FFBA39074F5DFE6BFB5AFE2B" author="Lohse" box="[888,1202,471,497]" pageId="36" pageNumber="88" refString="Lohse, G. A. &amp; Smetana, A. (1988) Four new species of Geostiba Thomson from the Appalachian mountains of North Carolina, with a key to North American species and synonymic notes (Coleoptera: Staphylinidae: Aleocharinae). The Coleopterists Bulletin, 42 (3), 269 - 278." type="journal article" year="1988">Lohse and Smetana (1988)</bibRefCitation>
listed this specimen as a
<typeStatus id="0C42E6A2FFBA39074D97FE40FDFFFDCC" box="[434,535,508,534]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
in the
<typeStatus id="0C42E6A2FFBA39074E42FE40FD71FDCC" box="[615,665,508,534]" pageId="36" pageNumber="37">type</typeStatus>
series of
<taxonomicName id="14F92383FFBA39074F2EFE40FC41FDCF" box="[779,937,508,533]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="bicarinata">
<emphasis id="E18D8412FFBA39074F2EFE40FC41FDCF" box="[779,937,508,533]" italics="true" pageId="36" pageNumber="37">G. bicarinata</emphasis>
</taxonomicName>
)(
<collectionCode id="B5E8C0C5FFBA39074F9FFE40FBE0FDCC" box="[954,1032,508,534]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="36" pageNumber="37">CNCI</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="87860888FFBA39074D2DF919FC40F89D" httpUri="https://zenodo.org/record/155720/files/figure.png" pageId="36" pageNumber="37" targetBox="[308,1279,632,1666]" targetPageId="36">
<paragraph id="D3465800FFBA39074D2DF919FC40F89D" blockId="36.[264,1323,1701,1863]" pageId="36" pageNumber="37">
<emphasis id="E18D8412FFBA39074D2DF919FE36F967" bold="true" box="[264,478,1701,1725]" pageId="36" pageNumber="37">FIGURES 126­133</emphasis>
. Aedeagus of
<taxonomicName id="14F92383FFBA39074E5BF919FC58F966" authority="Gusarov" authorityName="Gusarov" box="[638,944,1701,1724]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pluvigena" status="sp. nov.">
<emphasis id="E18D8412FFBA39074E5BF919FCA6F966" box="[638,846,1701,1724]" italics="true" pageId="36" pageNumber="37">Geostiba pluvigena</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="FABE3969FFBA39074F9FF919FC1DF967" box="[954,1013,1701,1725]" pageId="36" pageNumber="37" rank="species">
<emphasis id="E18D8412FFBA39074F9FF919FC1DF967" bold="true" box="[954,1013,1701,1725]" pageId="36" pageNumber="37">sp. n.</emphasis>
</taxonomicNameLabel>
(paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126­129, 132­133) and Waterrock Knob (130­ 131), North Carolina). 126, 130, 132 median lobe, ventral view; 127, 131, 133 apex of median lobe, ventral view; 128 median lobe, lateral view; 129 apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133).
</paragraph>
</caption>
<subSubSection id="9BE30B8BFFBB39064D1DFEA7FBBEFD34" pageId="37" pageNumber="38" type="diagnosis">
<paragraph id="D3465800FFBB39064D1DFEA7FDF7FE24" blockId="37.[264,1324,283,1870]" pageId="37" pageNumber="38">
<emphasis id="E18D8412FFBB39064D1DFEA7FE50FEEF" bold="true" box="[312,440,283,309]" pageId="37" pageNumber="38">Diagnosis.</emphasis>
<taxonomicName id="14F92383FFBB39064DE9FEA0FD5CFEEF" box="[460,692,284,309]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pluvigena">
<emphasis id="E18D8412FFBB39064DE9FEA0FD5CFEEF" box="[460,692,284,309]" italics="true" pageId="37" pageNumber="38">Geostiba pluvigena</emphasis>
</taxonomicName>
can be distinguished from other Nearctic species of
<taxonomicName id="14F92383FFBB39064D2DFEF8FE9AFE87" box="[264,370,324,349]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="E18D8412FFBB39064D2DFEF8FE9AFE87" box="[264,370,324,349]" italics="true" pageId="37" pageNumber="38">Geostiba</emphasis>
</taxonomicName>
by having small eyes (temple length to eye length ratio 2.7­3.8), pronotal pubescence of
<typeStatus id="0C42E6A2FFBB39064D56FED0FE4DFE5C" box="[371,421,364,390]" pageId="37" pageNumber="38">type</typeStatus>
V, reduced wings, short elytra (pronotum length to elytron length ratio 1.3), the presence of two short parallel carinae in the middle of abdominal tergum
<quantity id="1401F5E5FFBB390648B9FE28FB22FE74" box="[1180,1226,404,430]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="37" pageNumber="38" unit="in" value="7.0">7 in</quantity>
front of posterior margin, the shape of the aedeagus (
<figureCitation id="4BC24485FFBB39064F32FE00FBC6FE0C" box="[791,1070,444,470]" captionStart-0="FIGURES 126 ­ 133" captionStart-1="FIGURES 134 ­ 145" captionStartId-0="36.[264,382,1701,1725]" captionStartId-1="38.[264,383,1559,1583]" captionTargetBox-0="[308,1279,632,1666]" captionTargetBox-1="[315,1274,655,1520]" captionTargetId-0="figure@36.[294,1294,622,1676]" captionTargetId-1="figure@38.[307,1279,649,1533]" captionTargetPageId-0="36" captionTargetPageId-1="38" captionText-0="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 median lobe, ventral view; 127, 131, 133 apex of median lobe, ventral view; 128 median lobe, lateral view; 129 apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." captionText-1="FIGURES 134 ­ 145. Genitalia of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (134, 139 ­ 141) and Waterrock Knob (135 ­ 138, 142 ­ 145), North Carolina). 134 details of internal sac retracted into median lobe, ventral view; 135 copulatory piece of internal sac, dorsal view; 136 medial lamellae of internal sac, ventral view; 137 distal diverticula of everted internal sac, ventral view; 138 everted internal sac, ventral view; 139 spermatheca; 140 ­ 141 details of internal sac retracted into median lobe, lateral view; 142 copulatory piece of internal sac, lateral view; 143 medial lamella of internal sac, lateral view; 144 everted internal sac, lateral view; 145 apex of left paramere, side facing median lobe. Scale bar 0.2 mm (134, 138, 140 ­ 141, 144), 0.1 mm (135 ­ 137, 139, 142 ­ 143, 145)." httpUri-0="https://zenodo.org/record/155720/files/figure.png" httpUri-1="https://zenodo.org/record/155721/files/figure.png" pageId="37" pageNumber="38">Figs. 126­138, 140­145</figureCitation>
) and the shape of the spermatheca (
<figureCitation id="4BC24485FFBB39064D8EFE58FDF9FE24" box="[427,529,484,510]" captionStart="FIGURES 134 ­ 145" captionStartId="38.[264,383,1559,1583]" captionTargetBox="[315,1274,655,1520]" captionTargetId="figure@38.[307,1279,649,1533]" captionTargetPageId="38" captionText="FIGURES 134 ­ 145. Genitalia of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (134, 139 ­ 141) and Waterrock Knob (135 ­ 138, 142 ­ 145), North Carolina). 134 details of internal sac retracted into median lobe, ventral view; 135 copulatory piece of internal sac, dorsal view; 136 medial lamellae of internal sac, ventral view; 137 distal diverticula of everted internal sac, ventral view; 138 everted internal sac, ventral view; 139 spermatheca; 140 ­ 141 details of internal sac retracted into median lobe, lateral view; 142 copulatory piece of internal sac, lateral view; 143 medial lamella of internal sac, lateral view; 144 everted internal sac, lateral view; 145 apex of left paramere, side facing median lobe. Scale bar 0.2 mm (134, 138, 140 ­ 141, 144), 0.1 mm (135 ­ 137, 139, 142 ­ 143, 145)." httpUri="https://zenodo.org/record/155721/files/figure.png" pageId="37" pageNumber="38">Fig. 139</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFBB39064D1DFDB0FBBEFD34" blockId="37.[264,1324,283,1870]" pageId="37" pageNumber="38">
<taxonomicName id="14F92383FFBB39064D1DFDB0FDF5FDFF" box="[312,541,524,549]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pluvigena">
<emphasis id="E18D8412FFBB39064D1DFDB0FDF5FDFF" box="[312,541,524,549]" italics="true" pageId="37" pageNumber="38">Geostiba pluvigena</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="14F92383FFBB39064F2BFDB0FC4FFDFF" box="[782,935,524,549]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="nimbicola">
<emphasis id="E18D8412FFBB39064F2BFDB0FC4FFDFF" box="[782,935,524,549]" italics="true" pageId="37" pageNumber="38">G. nimbicola</emphasis>
</taxonomicName>
,
<taxonomicName id="14F92383FFBB39064F93FDB0FBB4FDFF" box="[950,1116,524,549]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="nebuligena">
<emphasis id="E18D8412FFBB39064F93FDB0FBB4FDFF" box="[950,1116,524,549]" italics="true" pageId="37" pageNumber="38">G. nebuligena</emphasis>
</taxonomicName>
and
<taxonomicName id="14F92383FFBB390648B1FDB0FEB9FD97" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="crepusculigena">
<emphasis id="E18D8412FFBB390648B1FDB0FEB9FD97" italics="true" pageId="37" pageNumber="38">G. crepusculigena</emphasis>
</taxonomicName>
.
<taxonomicName id="14F92383FFBB39064D48FD88FDBCFD97" box="[365,596,564,589]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pluvigena">
<emphasis id="E18D8412FFBB39064D48FD88FDBCFD97" box="[365,596,564,589]" italics="true" pageId="37" pageNumber="38">Geostiba pluvigena</emphasis>
</taxonomicName>
differs from
<taxonomicName id="14F92383FFBB39064ED2FD88FC7AFD97" box="[759,914,564,589]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="nimbicola">
<emphasis id="E18D8412FFBB39064ED2FD88FC7AFD97" box="[759,914,564,589]" italics="true" pageId="37" pageNumber="38">G. nimbicola</emphasis>
</taxonomicName>
in having shorter and broader (in ventral view) apex of median lobe with weak apical denticle (in lateral view) (
<figureCitation id="4BC24485FFBB39064897FDE0FEDEFD44" captionStart="FIGURES 126 ­ 133" captionStartId="36.[264,382,1701,1725]" captionTargetBox="[308,1279,632,1666]" captionTargetId="figure@36.[294,1294,622,1676]" captionTargetPageId="36" captionText="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 median lobe, ventral view; 127, 131, 133 apex of median lobe, ventral view; 128 median lobe, lateral view; 129 apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." httpUri="https://zenodo.org/record/155720/files/figure.png" pageId="37" pageNumber="38">Figs. 126­ 133</figureCitation>
,
<figureCitation id="4BC24485FFBB39064D63FD38FE44FD44" box="[326,428,644,670]" captionStart="FIGURES 100 ­ 107" captionStartId="33.[264,383,1742,1766]" captionTargetBox="[319,1271,659,1705]" captionTargetId="figure@33.[308,1278,649,1715]" captionTargetPageId="33" captionText="FIGURES 100 ­ 107. Aedeagus of Geostiba nimbicola Lohse &amp; Smetana (100 ­ 103 holotype; 104 ­ 107 Mount Sterling Ridge Trail, Tennessee). 100, 104 median lobe, ventral view; 101, 105 apex of median lobe, ventral view; 102, 106 median lobe, lateral view; 103, 107 apex of median lobe, lateral view. Scale bar 0.2 mm (100, 102, 104, 106), 0.1 mm (101, 103, 105, 107)." httpUri="https://zenodo.org/record/155717/files/figure.png" pageId="37" pageNumber="38">100­107</figureCitation>
); from
<taxonomicName id="14F92383FFBB39064E2DFD38FD5AFD47" box="[520,690,644,669]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="nebuligena">
<emphasis id="E18D8412FFBB39064E2DFD38FD5AFD47" box="[520,690,644,669]" italics="true" pageId="37" pageNumber="38">G. nebuligena</emphasis>
</taxonomicName>
in having weaker apical denticle of median lobe of aedeagus (
<figureCitation id="4BC24485FFBB39064DA3FD10FDC6FD1C" box="[390,558,684,710]" captionStart="FIGURES 126 ­ 133" captionStartId="36.[264,382,1701,1725]" captionTargetBox="[308,1279,632,1666]" captionTargetId="figure@36.[294,1294,622,1676]" captionTargetPageId="36" captionText="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 median lobe, ventral view; 127, 131, 133 apex of median lobe, ventral view; 128 median lobe, lateral view; 129 apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." httpUri="https://zenodo.org/record/155720/files/figure.png" pageId="37" pageNumber="38">Figs. 128­129</figureCitation>
,
<figureCitation id="4BC24485FFBB39064E18FD10FD4AFD1C" box="[573,674,684,710]" captionStart="FIGURES 152 ­ 156" captionStartId="41.[264,383,1456,1480]" captionTargetBox="[369,1217,460,1420]" captionTargetId="figure@41.[361,1225,449,1430]" captionTargetPageId="41" captionText="FIGURES 152 ­ 156. Aedeagus of Geostiba nebuligena Gusarov, sp. n. (paratypes from Highways 441 &amp; 71, Tennessee (152, 154 ­ 156) and Appalachian National Scenic Trail W of Clingmans Dome (153), Tennessee / North Carolina). 152 median lobe, ventral view; 153 ­ 154 apex of median lobe, ventral view; 155 median lobe, lateral view; 156 apex of median lobe, lateral view. Scale bar 0.2 mm (152, 155), 0.1 mm (153 ­ 154, 156)." httpUri="https://zenodo.org/record/155723/files/figure.png" pageId="37" pageNumber="38">155­156</figureCitation>
); from
<taxonomicName id="14F92383FFBB39064ED9FD10FC3DFD1F" box="[764,981,684,709]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="crepusculigena">
<emphasis id="E18D8412FFBB39064ED9FD10FC3DFD1F" box="[764,981,684,709]" italics="true" pageId="37" pageNumber="38">G. crepusculigena</emphasis>
</taxonomicName>
in lacking obtuse projection on the ventral side of the apex of median lobe (
<figureCitation id="4BC24485FFBB39064F15FD68FC3FFD34" box="[816,983,724,750]" captionStart="FIGURES 126 ­ 133" captionStartId="36.[264,382,1701,1725]" captionTargetBox="[308,1279,632,1666]" captionTargetId="figure@36.[294,1294,622,1676]" captionTargetPageId="36" captionText="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 median lobe, ventral view; 127, 131, 133 apex of median lobe, ventral view; 128 median lobe, lateral view; 129 apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." httpUri="https://zenodo.org/record/155720/files/figure.png" pageId="37" pageNumber="38">Figs. 128­129</figureCitation>
,
<figureCitation id="4BC24485FFBB39064FC6FD68FBA1FD34" box="[995,1097,724,750]" captionStart="FIGURES 174 ­ 177" captionStartId="45.[264,383,1766,1790]" captionTargetBox="[317,1274,1164,1735]" captionTargetId="figure@45.[305,1283,1154,1742]" captionTargetPageId="45" captionText="FIGURES 174 ­ 177. Aedeagus of Geostiba crepusculigena Gusarov, sp. n. (paratype from Snake Ridge Trail, Tennessee). 174 median lobe, ventral view; 175 apex of median lobe, ventral view; 176 median lobe, lateral view; 177 apex of median lobe, lateral view. Scale bar 0.2 mm (174, 176), 0.1 mm (175, 177)." httpUri="https://zenodo.org/record/155726/files/figure.png" pageId="37" pageNumber="38">176­177</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9BE30B8BFFBB39064D1DFD47FD96F894" pageId="37" pageNumber="38" type="description">
<paragraph id="D3465800FFBB39064D1DFD47FDDEFCBC" blockId="37.[264,1324,283,1870]" pageId="37" pageNumber="38">
<emphasis id="E18D8412FFBB39064D1DFD47FE38FCCF" bold="true" box="[312,464,763,789]" pageId="37" pageNumber="38">Description.</emphasis>
Length
<quantity id="1401F5E5FFBB39064E60FD40FD3CFCCC" box="[581,724,764,790]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="37" pageNumber="38" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.9­2.2 mm</quantity>
. Brown, pronotum, elytra and apex of abdomen often lighter, antennae brownish yellow or light brown, legs and mouthparts brownish yellow. Body parallel­sided.
</paragraph>
<paragraph id="D3465800FFBB39064D1DFCC8FD30FBF4" blockId="37.[264,1324,283,1870]" pageId="37" pageNumber="38">
Head as wide as long, surface on disk with fine isodiametric microsculpture, puncturation very fine, distance between punctures equal to 3­4 times their diameter. Temple length to eye length ratio 2.7­3.8. Antennal article 2 longer than article 3, article 4 transverse (width to length ratio 1.6), articles 5­10 strongly transverse (ratio 1.7­1.9), last article as long as 9 and 10 combined (
<figureCitation id="4BC24485FFBB39064E56FBA8FD22FBF4" box="[627,714,1044,1070]" captionStart="FIGURES 8 ­ 17" captionStartId="6.[264,383,1709,1733]" captionTargetBox="[307,1277,321,1687]" captionTargetId="figure@6.[297,1289,310,1697]" captionTargetPageId="6" captionText="FIGURES 8 ­ 17. Mouthparts of Geostiba circellaris (Gravenhorst) (8, Moscow Region, Russia), G. infirma (Weise) (9, Ivano­Frankovsk Region, Ukraine), G. carteriensis Pace (10 ­ 15, Tennessee), G. appalachigena Gusarov, sp. n. (16, paratype from West Virginia) and G. pluvigena Gusarov, sp. n. (17, paratype from North Carolina). 8 ­ 10, prementum; 11 hypopharynx; 12 left maxilla, ventral view; 13 left galea, dorsal view; 14 left lacinia, dorsal view; 15 mentum; 16 ­ 17 right antenna. Scale bar 0.1 mm (8 ­ 15), 0.2 mm (16 ­ 17). L Length of antennal article." httpUri="https://zenodo.org/record/155703/files/figure.png" pageId="37" pageNumber="38">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFBB39064D1DFB80FDC3FA9C" blockId="37.[264,1324,283,1870]" pageId="37" pageNumber="38">
Pronotum as wide as long, width
<quantity id="1401F5E5FFBB39064EEAFB80FC93FB8C" box="[719,891,1084,1110]" metricMagnitude="-4" metricUnit="m" metricValue="3.95" metricValueMax="4.3" metricValueMin="3.5999999999999996" pageId="37" pageNumber="38" unit="mm" value="0.395" valueMax="0.43" valueMin="0.36">0.36­0.43 mm</quantity>
, wider than head (pronotal width to head width ratio 1.1); microsculpture and puncturation as on head. Pronotal pubescence of
<typeStatus id="0C42E6A2FFBB39064D2DFB30FED2FB7C" box="[264,314,1164,1190]" pageId="37" pageNumber="38">type</typeStatus>
V. Elytra measured from humeral angle shorter than pronotum (pronotal length to elytral length ratio 1.3), wider than long (1.5), with fine isodiametric microsculpture and fine asperate puncturation, distance between punctures equals 2­3 times their diameter. Elytral suture behind scutellum slightly raised in both sexes. Wings reduced to short vestiges, shorter than elytra.
</paragraph>
<paragraph id="D3465800FFBB39064D1DFAE8FB22FA64" blockId="37.[264,1324,283,1870]" pageId="37" pageNumber="38">Abdominal terga with fine microsculpture of transverse meshes, with fine and sparse puncturation, puncturation becoming finer towards abdomen apex, on terga 3­5 distance between punctures equals 2­5 times their diameter. Tergum 7 without white edge.</paragraph>
<paragraph id="D3465800FFBB39064D1DFA70FB8BF9EC" blockId="37.[264,1324,283,1870]" pageId="37" pageNumber="38">
Male tergum 7 with two medial carinae in front of posterior margin. Male tergum 8 with two or four weak and short carinae in front of posterior margin, posterior margin convex (
<figureCitation id="4BC24485FFBB39064D66F9A0FE41F9EC" box="[323,425,1564,1590]" captionStart="FIGURES 120 ­ 125" captionStartId="35.[264,382,1745,1769]" captionTargetBox="[305,1280,668,1726]" captionTargetId="figure@35.[302,1286,668,1727]" captionTargetPageId="35" captionText="FIGURES 120 ­ 125. Abdominal segments 7 ­ 8 of Geostiba pluvigena Gusarov, sp. n. (paratypes from Waterrock Knob (120 ­ 121, 125) and junction of Blue Ridge Parkway and Balsam Mountain Road (122 ­ 124), North Carolina). 120 male tergum 8; 121 male sternum 8; 122 female tergum 8; 123 female sternum 8, 124 apex of female sternum 8; 125 medial portion of male tergum 7, posterior down. Scale bar 0.2 mm (120 ­ 123, 125), 0.1 mm (124)." httpUri="https://zenodo.org/record/155719/files/figure.png" pageId="37" pageNumber="38">Fig. 120</figureCitation>
). Male sternum 8 with convex posterior margin (
<figureCitation id="4BC24485FFBB39064FCAF9A0FBBDF9EC" box="[1007,1109,1564,1590]" captionStart="FIGURES 120 ­ 125" captionStartId="35.[264,382,1745,1769]" captionTargetBox="[305,1280,668,1726]" captionTargetId="figure@35.[302,1286,668,1727]" captionTargetPageId="35" captionText="FIGURES 120 ­ 125. Abdominal segments 7 ­ 8 of Geostiba pluvigena Gusarov, sp. n. (paratypes from Waterrock Knob (120 ­ 121, 125) and junction of Blue Ridge Parkway and Balsam Mountain Road (122 ­ 124), North Carolina). 120 male tergum 8; 121 male sternum 8; 122 female tergum 8; 123 female sternum 8, 124 apex of female sternum 8; 125 medial portion of male tergum 7, posterior down. Scale bar 0.2 mm (120 ­ 123, 125), 0.1 mm (124)." httpUri="https://zenodo.org/record/155719/files/figure.png" pageId="37" pageNumber="38">Fig. 121</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFBB39064D1DF9F8FD67F95C" blockId="37.[264,1324,283,1870]" pageId="37" pageNumber="38">
Female tergum 8 with convex posterior margin (
<figureCitation id="4BC24485FFBB39064F51F9F8FC32F984" box="[884,986,1604,1630]" captionStart="FIGURES 120 ­ 125" captionStartId="35.[264,382,1745,1769]" captionTargetBox="[305,1280,668,1726]" captionTargetId="figure@35.[302,1286,668,1727]" captionTargetPageId="35" captionText="FIGURES 120 ­ 125. Abdominal segments 7 ­ 8 of Geostiba pluvigena Gusarov, sp. n. (paratypes from Waterrock Knob (120 ­ 121, 125) and junction of Blue Ridge Parkway and Balsam Mountain Road (122 ­ 124), North Carolina). 120 male tergum 8; 121 male sternum 8; 122 female tergum 8; 123 female sternum 8, 124 apex of female sternum 8; 125 medial portion of male tergum 7, posterior down. Scale bar 0.2 mm (120 ­ 123, 125), 0.1 mm (124)." httpUri="https://zenodo.org/record/155719/files/figure.png" pageId="37" pageNumber="38">Fig. 122</figureCitation>
), sternum 8 with emarginate posterior margin (
<figureCitation id="4BC24485FFBB39064DFFF9D0FD6AF95C" box="[474,642,1644,1670]" captionStart="FIGURES 120 ­ 125" captionStartId="35.[264,382,1745,1769]" captionTargetBox="[305,1280,668,1726]" captionTargetId="figure@35.[302,1286,668,1727]" captionTargetPageId="35" captionText="FIGURES 120 ­ 125. Abdominal segments 7 ­ 8 of Geostiba pluvigena Gusarov, sp. n. (paratypes from Waterrock Knob (120 ­ 121, 125) and junction of Blue Ridge Parkway and Balsam Mountain Road (122 ­ 124), North Carolina). 120 male tergum 8; 121 male sternum 8; 122 female tergum 8; 123 female sternum 8, 124 apex of female sternum 8; 125 medial portion of male tergum 7, posterior down. Scale bar 0.2 mm (120 ­ 123, 125), 0.1 mm (124)." httpUri="https://zenodo.org/record/155719/files/figure.png" pageId="37" pageNumber="38">Figs. 123­124</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFBB39064D1DF928FD2FF8FC" blockId="37.[264,1324,283,1870]" pageId="37" pageNumber="38">
Aedeagus as in
<figureCitation id="4BC24485FFBB39064DD4F928FCE1F974" box="[497,777,1684,1710]" captionStart-0="FIGURES 126 ­ 133" captionStart-1="FIGURES 134 ­ 145" captionStartId-0="36.[264,382,1701,1725]" captionStartId-1="38.[264,383,1559,1583]" captionTargetBox-0="[308,1279,632,1666]" captionTargetBox-1="[315,1274,655,1520]" captionTargetId-0="figure@36.[294,1294,622,1676]" captionTargetId-1="figure@38.[307,1279,649,1533]" captionTargetPageId-0="36" captionTargetPageId-1="38" captionText-0="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 median lobe, ventral view; 127, 131, 133 apex of median lobe, ventral view; 128 median lobe, lateral view; 129 apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." captionText-1="FIGURES 134 ­ 145. Genitalia of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (134, 139 ­ 141) and Waterrock Knob (135 ­ 138, 142 ­ 145), North Carolina). 134 details of internal sac retracted into median lobe, ventral view; 135 copulatory piece of internal sac, dorsal view; 136 medial lamellae of internal sac, ventral view; 137 distal diverticula of everted internal sac, ventral view; 138 everted internal sac, ventral view; 139 spermatheca; 140 ­ 141 details of internal sac retracted into median lobe, lateral view; 142 copulatory piece of internal sac, lateral view; 143 medial lamella of internal sac, lateral view; 144 everted internal sac, lateral view; 145 apex of left paramere, side facing median lobe. Scale bar 0.2 mm (134, 138, 140 ­ 141, 144), 0.1 mm (135 ­ 137, 139, 142 ­ 143, 145)." httpUri-0="https://zenodo.org/record/155720/files/figure.png" httpUri-1="https://zenodo.org/record/155721/files/figure.png" pageId="37" pageNumber="38">Figs. 126­138, 140­145</figureCitation>
. Apex of median lobe in ventral view narrow, its outline convex or doubly emarginate apically (
<figureCitation id="4BC24485FFBB39064F76F900FB85F90C" box="[851,1133,1724,1750]" captionStart="FIGURES 126 ­ 133" captionStartId="36.[264,382,1701,1725]" captionTargetBox="[308,1279,632,1666]" captionTargetId="figure@36.[294,1294,622,1676]" captionTargetPageId="36" captionText="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 median lobe, ventral view; 127, 131, 133 apex of median lobe, ventral view; 128 median lobe, lateral view; 129 apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." httpUri="https://zenodo.org/record/155720/files/figure.png" pageId="37" pageNumber="38">Figs. 126­127, 130­133</figureCitation>
), in lateral view strait, with weak apical denticle ventrally (
<figureCitation id="4BC24485FFBB39064ED9F958FC4CF924" box="[764,932,1764,1790]" captionStart="FIGURES 126 ­ 133" captionStartId="36.[264,382,1701,1725]" captionTargetBox="[308,1279,632,1666]" captionTargetId="figure@36.[294,1294,622,1676]" captionTargetPageId="36" captionText="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 median lobe, ventral view; 127, 131, 133 apex of median lobe, ventral view; 128 median lobe, lateral view; 129 apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." httpUri="https://zenodo.org/record/155720/files/figure.png" pageId="37" pageNumber="38">Figs. 128­129</figureCitation>
), distal diverticula of internal sac in ventral view broad (
<figureCitation id="4BC24485FFBB39064E37F8B0FD53F8FC" box="[530,699,1804,1830]" captionStart="FIGURES 134 ­ 145" captionStartId="38.[264,383,1559,1583]" captionTargetBox="[315,1274,655,1520]" captionTargetId="figure@38.[307,1279,649,1533]" captionTargetPageId="38" captionText="FIGURES 134 ­ 145. Genitalia of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (134, 139 ­ 141) and Waterrock Knob (135 ­ 138, 142 ­ 145), North Carolina). 134 details of internal sac retracted into median lobe, ventral view; 135 copulatory piece of internal sac, dorsal view; 136 medial lamellae of internal sac, ventral view; 137 distal diverticula of everted internal sac, ventral view; 138 everted internal sac, ventral view; 139 spermatheca; 140 ­ 141 details of internal sac retracted into median lobe, lateral view; 142 copulatory piece of internal sac, lateral view; 143 medial lamella of internal sac, lateral view; 144 everted internal sac, lateral view; 145 apex of left paramere, side facing median lobe. Scale bar 0.2 mm (134, 138, 140 ­ 141, 144), 0.1 mm (135 ­ 137, 139, 142 ­ 143, 145)." httpUri="https://zenodo.org/record/155721/files/figure.png" pageId="37" pageNumber="38">Figs. 137­138</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFBB39064D1DF888FD96F894" blockId="37.[264,1324,283,1870]" box="[312,638,1844,1870]" pageId="37" pageNumber="38">
Spermatheca as in
<figureCitation id="4BC24485FFBB39064E30F888FD92F894" box="[533,634,1844,1870]" captionStart="FIGURES 134 ­ 145" captionStartId="38.[264,383,1559,1583]" captionTargetBox="[315,1274,655,1520]" captionTargetId="figure@38.[307,1279,649,1533]" captionTargetPageId="38" captionText="FIGURES 134 ­ 145. Genitalia of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (134, 139 ­ 141) and Waterrock Knob (135 ­ 138, 142 ­ 145), North Carolina). 134 details of internal sac retracted into median lobe, ventral view; 135 copulatory piece of internal sac, dorsal view; 136 medial lamellae of internal sac, ventral view; 137 distal diverticula of everted internal sac, ventral view; 138 everted internal sac, ventral view; 139 spermatheca; 140 ­ 141 details of internal sac retracted into median lobe, lateral view; 142 copulatory piece of internal sac, lateral view; 143 medial lamella of internal sac, lateral view; 144 everted internal sac, lateral view; 145 apex of left paramere, side facing median lobe. Scale bar 0.2 mm (134, 138, 140 ­ 141, 144), 0.1 mm (135 ­ 137, 139, 142 ­ 143, 145)." httpUri="https://zenodo.org/record/155721/files/figure.png" pageId="37" pageNumber="38">Fig. 139</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9BE30B8BFFB839054D1DFEA7FC98FE74" pageId="38" pageNumber="39" type="distribution">
<paragraph id="D3465800FFB839054D1DFEA7FDA9FE84" blockId="38.[264,1324,283,550]" pageId="38" pageNumber="39">
<emphasis id="E18D8412FFB839054D1DFEA7FE30FEEF" bold="true" box="[312,472,283,309]" pageId="38" pageNumber="39">Distribution.</emphasis>
Known from the Plott Balsams Cataloochee Divide massif (North Carolina) (
<figureCitation id="4BC24485FFB839054DA2FEF8FE10FE84" box="[391,504,324,350]" captionStart="FIGURE 338" captionStartId="79.[264,367,1838,1862]" captionTargetBox="[346,1237,315,1825]" captionTargetId="figure@79.[344,1238,312,1827]" captionTargetPageId="79" captionText="FIGURE 338. Geographical distribution of twelve species of Geostiba in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155750/files/figure.png" pageId="38" pageNumber="39">Figs. 338</figureCitation>
,
<figureCitation id="4BC24485FFB839054E21FEF8FDDDFE84" box="[516,565,324,350]" captionStart="FIGURE 340" captionStartId="82.[264,368,1747,1771]" captionTargetBox="[322,1272,508,1726]" captionTargetId="figure@82.[319,1275,508,1728]" captionTargetPageId="82" captionText="FIGURE 340. Detailed geographical distribution of Geostiba bicarinata Lohse &amp; Smetana (2), G. graveyardensis Pace (3), G. nimbicola Lohse &amp; Smetana (4), G. pluvigena Gusarov, sp. n. (5), G. nebuligena Gusarov, sp. n. (6), G. crepusculigena Gusarov, sp. n. (7), G. balsamensis Pace (8), G. nubigena Lohse &amp; Smetana (9), G. coeligena Gusarov, sp. n. (10) in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155752/files/figure.png" pageId="38" pageNumber="39">340</figureCitation>
).
</paragraph>
<paragraph id="D3465800FFB839054D1DFED7FC98FE74" blockId="38.[264,1324,283,550]" pageId="38" pageNumber="39">
<emphasis id="E18D8412FFB839054D1DFED7FDEDFE5F" bold="true" box="[312,517,363,389]" pageId="38" pageNumber="39">Natural History.</emphasis>
<taxonomicName id="14F92383FFB839054E30FED0FD13FE5F" box="[533,763,364,389]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="nimbicola">
<emphasis id="E18D8412FFB839054E30FED0FD13FE5F" box="[533,763,364,389]" italics="true" pageId="38" pageNumber="39">Geostiba nimbicola</emphasis>
</taxonomicName>
was collected at altitudes above
<quantity id="1401F5E5FFB8390548A5FED0FB31FE5C" box="[1152,1241,364,390]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="38" pageNumber="39" unit="m" value="1600.0">1600 m</quantity>
in leaf litter in mixed forest with red spruce (
<taxonomicName id="14F92383FFB839054EE0FE28FCB7FE77" box="[709,863,404,429]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="38" pageNumber="39" phylum="Tracheophyta" rank="species" species="rubens">
<emphasis id="E18D8412FFB839054EE0FE28FCB7FE77" box="[709,863,404,429]" italics="true" pageId="38" pageNumber="39">Picea rubens</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="9BE30B8BFFB839054D1DFE07FDDBFDFC" pageId="38" pageNumber="39" type="etymology">
<paragraph id="D3465800FFB839054D1DFE07FDDBFDFC" blockId="38.[264,1324,283,550]" pageId="38" pageNumber="39">
<emphasis id="E18D8412FFB839054D1DFE07FE2DFE0F" bold="true" box="[312,453,443,469]" pageId="38" pageNumber="39">Etymology.</emphasis>
The specific name is derived from the Latin adjective
<emphasis id="E18D8412FFB839054868FE00FB4AFE0F" box="[1101,1186,444,469]" italics="true" pageId="38" pageNumber="39">pluvius</emphasis>
(rainy) and the verb
<emphasis id="E18D8412FFB839054D56FE58FE5EFE27" box="[371,438,484,509]" italics="true" pageId="38" pageNumber="39">gigno</emphasis>
(to be born, to arise). It refers to the rains in the Southern Appalachians where the species occurs.
</paragraph>
</subSubSection>
</treatment>
</document>