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<document id="56333E9AD1883DD9D9EE1C65DE67D98A" ID-DOI="10.5281/zenodo.155701" ID-GBIF-Dataset="b947e6c9-d6cf-4988-a098-5e01dd69a888" ID-ISSN="1175­5326" ID-Zenodo-Dep="155701" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459438614591" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2002" docId="5B50E916FF9039324D2DFE42FAC3FE5F" docLanguage="en" docName="zt00081.pdf" docOrigin="Zootaxa 81" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Geostiba" docType="treatment" docVersion="14" lastPageNumber="17" masterDocId="A769916EFF9E39234C25FFBCFFE8FFDA" masterDocTitle="A revision of Nearctic species of the genus Geostiba Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="88" masterPageNumber="1" pageNumber="15" updateTime="1698188143631" updateUser="plazi">
<mods:mods id="A7827C21FD9836411D92C6800ED90ACF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DE7650A8DDE46F05119469F1BAB9538A">
<mods:title id="6AD15A6448E2C6D95447E05E85D5D304">A revision of Nearctic species of the genus Geostiba Thomson, 1858 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="5413704FD12C2E6B8F94F86ACA1C7599">Gusarov, Vladimir I.</mods:namePart>
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<mods:typeOfResource id="C6080464B61479839E322C90745B9BB1">text</mods:typeOfResource>
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<mods:title id="55C3A041FCE8BC57E591730A0FBC3B21">Zootaxa</mods:title>
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<mods:date id="9D4F7CF898DCCAA1A107564DE18D77E4">2002</mods:date>
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<mods:number id="64B7B6435634674C31E6A0542C037865">81</mods:number>
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<mods:classification id="F3E533EEC918BE2F1A1F006C1E9131E1">journal article</mods:classification>
<mods:identifier id="34BABF0740D534E3064F0C929D92EAE4" type="DOI">10.5281/zenodo.155701</mods:identifier>
<mods:identifier id="D9687238C7BA932D1646A662369557CC" type="GBIF-Dataset">b947e6c9-d6cf-4988-a098-5e01dd69a888</mods:identifier>
<mods:identifier id="25F10D07A5AD233BB4A9101AE61A35C1" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="56B90CBE7E6D59C3FBA332F170E1C581" type="Zenodo-Dep">155701</mods:identifier>
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<treatment id="5B50E916FF9039324D2DFE42FAC3FE5F" ID-DOI="http://doi.org/10.5281/zenodo.6277552" ID-GBIF-Taxon="119330226" ID-Zenodo-Dep="6277552" LSID="urn:lsid:plazi:treatment:5B50E916FF9039324D2DFE42FAC3FE5F" httpUri="http://treatment.plazi.org/id/5B50E916FF9039324D2DFE42FAC3FE5F" lastPageId="17" lastPageNumber="17" pageId="14" pageNumber="15">
<subSubSection id="9BE30B8BFF90392D4D2DFE42FC74FDC2" box="[264,924,510,536]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="D3465800FF90392D4D2DFE42FC74FDC2" blockId="14.[264,924,510,536]" box="[264,924,510,536]" pageId="14" pageNumber="15">
<heading id="880EEF6CFF90392D4D2DFE42FC74FDC2" bold="true" box="[264,924,510,536]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="E18D8412FF90392D4D2DFE42FC74FDC2" bold="true" box="[264,924,510,536]" pageId="14" pageNumber="15">
Key for Identification of Nearctic Species of
<taxonomicName id="14F92383FF90392D4F17FE42FC74FDC2" ID-CoL="8VRPP" box="[818,924,510,536]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="E18D8412FF90392D4F17FE42FC74FDC2" bold="true" box="[818,924,510,536]" italics="true" pageId="14" pageNumber="15">Geostiba</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<keyStep id="680D40A5FF90392D4D2DFDF8FAC5FBCC" pageId="14" pageNumber="15">
<subSubSection id="9BE30B8BFF90392D4D2DFDF8FAC3FC94" pageId="14" pageNumber="15" type="key">
<paragraph id="D3465800FF90392D4D2DFDF8FAC3FC94" blockId="14.[264,1325,580,1886]" pageId="14" pageNumber="15">
<keyLead id="6808FB35FF90392D4D2DFDF8FAC3FC94" pageId="14" pageNumber="15">
1 In males pronotum with one medial tubercle at posterior margin; each elytron with a tubercle near scutellum but
<emphasis id="E18D8412FF90392D4EABFDD0FD3EFD5F" box="[654,726,620,645]" italics="true" pageId="14" pageNumber="15">not at</emphasis>
the elytral suture; abdominal tergum 7 with one short medial carina in front of posterior margin. Posterior margin of female sternum 8 distinctly convex, without medial emargination. Aedeagus:
<figureCitation id="4BC24485FF90392D481DFD00FB4FFD0C" box="[1080,1191,700,726]" captionStart="FIGURES 1 ­ 7" captionStartId="5.[264,383,1751,1775]" captionTargetBox="[367,1205,490,1733]" captionTargetId="figure@5.[365,1207,490,1736]" captionTargetPageId="5" captionText="FIGURES 1 ­ 7. Mouthparts of Geostiba circellaris (Gravenhorst) (1 ­ 2, Moscow Region, Russia), G. carteriensis Pace (3 ­ 4, Tennessee), G. appalachigena Gusarov, sp. n. (5 ­ 6, paratype from West Virginia) and G. pluvigena Gusarov, sp. n. (7, paratype from North Carolina). 1, 3 – labrum; 2, 4 – epipharynx; 5 – left mandible, dorsal view; 6 – left mandible, ventral view; 7 – right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155702/files/figure.png" pageId="14" pageNumber="15">
Figs.
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</figureCitation>
in
<bibRefCitation id="B76825F1FF90392D48FFFD00FE62FD24" author="Assing" pageId="14" pageNumber="86" refString="Assing, V. (2001) A revision of the Turkish species of Geostiba Thomson 1858 and Tropimenelytron Pace 1983 (Coleoptera: Staphylinidae, Aleocharinae). Linzer biologische Beitrage, 33 (1), 137 - 185." type="journal article" year="2001">Assing (2001)</bibRefCitation>
. Spermatheca:
<figureCitation id="4BC24485FF90392D4E72FD58FD76FD24" box="[599,670,740,766]" captionStart="FIGURES 1 ­ 7" captionStartId="5.[264,383,1751,1775]" captionTargetBox="[367,1205,490,1733]" captionTargetId="figure@5.[365,1207,490,1736]" captionTargetPageId="5" captionText="FIGURES 1 ­ 7. Mouthparts of Geostiba circellaris (Gravenhorst) (1 ­ 2, Moscow Region, Russia), G. carteriensis Pace (3 ­ 4, Tennessee), G. appalachigena Gusarov, sp. n. (5 ­ 6, paratype from West Virginia) and G. pluvigena Gusarov, sp. n. (7, paratype from North Carolina). 1, 3 – labrum; 2, 4 – epipharynx; 5 – left mandible, dorsal view; 6 – left mandible, ventral view; 7 – right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/155702/files/figure.png" pageId="14" pageNumber="15">
Fig.
<quantity id="1401F5E5FF90392D4EAAFD58FD76FD24" box="[655,670,740,766]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="14" pageNumber="15" unit="in" value="4.0">4</quantity>
</figureCitation>
in
<bibRefCitation id="B76825F1FF90392D4EEFFD58FC9FFD24" author="Assing" box="[714,887,740,766]" pageId="14" pageNumber="86" refString="Assing, V. (2001) A revision of the Turkish species of Geostiba Thomson 1858 and Tropimenelytron Pace 1983 (Coleoptera: Staphylinidae, Aleocharinae). Linzer biologische Beitrage, 33 (1), 137 - 185." type="journal article" year="2001">Assing (2001)</bibRefCitation>
. Body length
<quantity id="1401F5E5FF90392D4808FD58FB54FD24" box="[1069,1212,740,766]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="14" pageNumber="15" unit="mm" value="2.55" valueMax="2.8" valueMin="2.3">2.3­2.8 mm</quantity>
, pronotal width
<quantity id="1401F5E5FF90392D4DA7FCB0FDC4FCFC" box="[386,556,780,806]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.3" metricValueMin="4.3" pageId="14" pageNumber="15" unit="mm" value="0.48" valueMax="0.53" valueMin="0.43">0.43­0.53 mm</quantity>
. Introduced from Europe to Newfoundland (
<figureCitation id="4BC24485FF90392D481BFCB0FB4BFCFC" box="[1086,1187,780,806]" captionStart="FIGURE 336" captionStartId="76.[264,368,1562,1586]" captionTargetBox="[315,1271,753,1543]" captionTargetId="figure@76.[311,1275,751,1546]" captionTargetPageId="76" captionText="FIGURE 336. Geographical distribution of Nearctic species of Geostiba. The ellipse indicates the area where eleven Southern Appalachian species are distributed." httpUri="https://zenodo.org/record/155748/files/figure.png" pageId="14" pageNumber="15">Fig. 336</figureCitation>
) ................ ......................................................................... 15.
<emphasis id="E18D8412FF90392D4FA6FC88FC77FC97" box="[899,927,820,845]" italics="true" pageId="14" pageNumber="15">G.</emphasis>
(
<emphasis id="E18D8412FF90392D4F95FC88FC00FC97" box="[944,1000,820,845]" italics="true" pageId="14" pageNumber="15">s. str</emphasis>
.)
<taxonomicName id="14F92383FF90392D4825FC88FACFFC94" ID-CoL="3FSFC" authority="Gravenhorst" authorityName="Gravenhorst" box="[1024,1319,820,846]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="circellaris">
<emphasis id="E18D8412FF90392D4825FC88FB90FC97" box="[1024,1144,820,845]" italics="true" pageId="14" pageNumber="15">circellaris</emphasis>
(Gravenhorst)
</taxonomicName>
.
</keyLead>
</paragraph>
</subSubSection>
<paragraph id="D3465800FF90392D4D2DFCE0FAC5FBCC" blockId="14.[264,1325,580,1886]" pageId="14" pageNumber="15">
<keyLead id="6808FB35FF90392D4D2DFCE0FAC5FBCC" pageId="14" pageNumber="15">
– In males pronotum without a medial tubercle at posterior margin; elytral suture is raised behind scutellum (flat in
<taxonomicName id="14F92383FF90392D4E99FC38FC8AFC47" box="[700,866,900,925]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="impressula">
<emphasis id="E18D8412FF90392D4E99FC38FC8AFC47" box="[700,866,900,925]" italics="true" pageId="14" pageNumber="15">G. impressula</emphasis>
</taxonomicName>
); abdominal tergum 7 with two short and parallel medial carinae in front of posterior margin or without carinae. Posterior margin of female sternum 8 medially slightly concave or with emargination (
<figureCitation id="4BC24485FF90392D48E4FC68FEB0FBCC" captionStart="FIGURES 29 ­ 33" captionStartId="20.[264,383,1785,1809]" captionTargetBox="[298,1291,699,1756]" captionTargetId="figure@20.[289,1298,689,1763]" captionTargetPageId="20" captionText="FIGURES 29 ­ 33. Abdominal segment 8 of Geostiba appalachigena Gusarov, sp. n. (paratypes from Black Mountain, West Virginia). 29 – male tergum 8; 30 – male sternum 8; 31 – female tergum 8; 32 – female sternum 8, 33 – apex of female sternum 8. Scale bar 0.2 mm (29 ­ 32), 0.1 mm (33)." httpUri="https://zenodo.org/record/155707/files/figure.png" pageId="14" pageNumber="15">Figs. 32­ 33</figureCitation>
,
<figureCitation id="4BC24485FF90392D4D41FC40FE44FBCC" box="[356,428,1020,1046]" captionStart="FIGURES 51 ­ 56" captionStartId="24.[264,383,1769,1793]" captionTargetBox="[321,1256,734,1749]" captionTargetId="figure@24.[318,1259,731,1752]" captionTargetPageId="24" captionText="FIGURES 51 ­ 56. Abdominal segments 7 ­ 8 of Geostiba bicarinata Lohse &amp; Smetana (Richland Balsam Mountain, North Carolina). 51 – male tergum 8; 52 – male sternum 8; 53 – female tergum 8; 54 – female sternum 8, 55 – apex of female sternum 8; 56 – medial portion of male tergum 7, posterior down. Scale bar 0.2 mm (51 ­ 54, 56), 0.1 mm (55)." httpUri="https://zenodo.org/record/155710/files/figure.png" pageId="14" pageNumber="15">54­55</figureCitation>
). Native Nearctic species ........................................................................... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="680D40A5FF90392D4D2DFB98FAC4FBBC" pageId="14" pageNumber="15">
<paragraph id="D3465800FF90392D4D2DFB98FAC5FBE4" blockId="14.[264,1325,580,1886]" box="[264,1325,1060,1086]" pageId="14" pageNumber="15">
<keyLead id="6808FB35FF90392D4D2DFB98FAC5FBE4" box="[264,1325,1060,1086]" pageId="14" pageNumber="15">
2 Species occurring in eastern North
<collectingCountry id="ABEE1890FF90392D4EF1FB98FCD4FBE4" box="[724,828,1060,1086]" name="United States of America" pageId="14" pageNumber="15">America</collectingCountry>
, east of the Rocky Mountains................. 3
</keyLead>
</paragraph>
<paragraph id="D3465800FF90392D4D2DFBF0FAC4FBBC" blockId="14.[264,1325,580,1886]" box="[264,1324,1100,1126]" pageId="14" pageNumber="15">
<keyLead id="6808FB35FF90392D4D2DFBF0FAC4FBBC" box="[264,1324,1100,1126]" pageId="14" pageNumber="15">
– Species occurring in western North
<collectingCountry id="ABEE1890FF90392D4EF9FBF0FCACFBBC" box="[732,836,1100,1126]" name="United States of America" pageId="14" pageNumber="15">America</collectingCountry>
, west of the Rocky Mountains ............ 15
</keyLead>
</paragraph>
</keyStep>
<keyStep id="680D40A5FF90392D4D2DFBC8FAC3F99C" pageId="14" pageNumber="15">
<paragraph id="D3465800FF90392D4D2DFBC8FAC3FA7F" blockId="14.[264,1325,580,1886]" pageId="14" pageNumber="15">
<keyLead id="6808FB35FF90392D4D2DFBC8FAC3FA7F" pageId="14" pageNumber="15">
3 Temples 2.3­2.7 times as long as eyes. Elytra (measured from humeral level) longer than pronotum. Wings fully developed. In males abdominal tergum 7 with two short and parallel medial carinae in front of posterior margin, abdominal tergum 8 with four short and weak carinae in front of posterior margin. Apex of median lobe in lateral view bent ventrally and widened apically (
<figureCitation id="4BC24485FF90392D4F61FAA8FC3CFAF4" box="[836,980,1300,1326]" captionStart="FIGURES 34 ­ 37" captionStartId="21.[264,383,1148,1172]" captionTargetBox="[321,1268,571,1127]" captionTargetId="figure@21.[319,1272,569,1127]" captionTargetPageId="21" captionText="FIGURES 34 ­ 37. Aedeagus of Geostiba appalachigena Gusarov, sp. n. (paratype from Black Mountain, West Virginia). 34 – median lobe, ventral view; 35 – apex of median lobe, ventral view; 36 – median lobe, lateral view; 37 – apex of median lobe, lateral view. Scale bar 0.2 mm (34, 36), 0.1 mm (35, 37)." httpUri="https://zenodo.org/record/155708/files/figure.png" pageId="14" pageNumber="15">Figs. 36­37</figureCitation>
). Body length
<quantity id="1401F5E5FF90392D48B2FAA8FACFFAF4" box="[1175,1319,1300,1326]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.4" metricValueMin="1.9" pageId="14" pageNumber="15" unit="mm" value="2.15" valueMax="2.4" valueMin="1.9">1.9­2.4 mm</quantity>
, pronotal width
<quantity id="1401F5E5FF90392D4DD3FA80FD4CFA8C" box="[502,676,1340,1366]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="4.699999999999999" metricValueMin="4.0" pageId="14" pageNumber="15" unit="mm" value="0.435" valueMax="0.47" valueMin="0.4">0.40­0.47 mm</quantity>
. Widely distributed in eastern North
<collectingCountry id="ABEE1890FF90392D48A5FA80FB0DFA8C" box="[1152,1253,1340,1366]" name="United States of America" pageId="14" pageNumber="15">America</collectingCountry>
from Mount Rogers, Virginia in the south to Maine, Québec and Wisconsin in the north (
<figureCitation id="4BC24485FF90392D4D64FA30FE4DFA7C" box="[321,421,1420,1446]" captionStart="FIGURE 337" captionStartId="77.[264,367,1248,1272]" captionTargetBox="[328,1262,539,1226]" captionTargetId="figure@77.[321,1266,529,1233]" captionTargetPageId="77" captionText="FIGURE 337. Geographical distribution of Geostiba appalachigena Gusarov, sp. n. The ellipse indicates the area where eleven Southern Appalachian species are distributed." httpUri="https://zenodo.org/record/155749/files/figure.png" pageId="14" pageNumber="15">Fig. 337</figureCitation>
)................................................ 1.
<taxonomicName id="14F92383FF90392D4F14FA30FB35FA7C" authority="Gusarov" authorityName="Gusarov" box="[817,1245,1420,1446]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="appalachigena" status="sp. nov." subGenus="Sibiota">
<emphasis id="E18D8412FF90392D4F14FA30FCA6FA7F" box="[817,846,1420,1445]" italics="true" pageId="14" pageNumber="15">G.</emphasis>
(
<emphasis id="E18D8412FF90392D4F7AFA30FC5AFA7F" box="[863,946,1420,1445]" italics="true" pageId="14" pageNumber="15">Sibiota</emphasis>
)
<emphasis id="E18D8412FF90392D4FE6FA30FB99FA7F" box="[963,1137,1420,1445]" italics="true" pageId="14" pageNumber="15">appalachigena</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="FABE3969FF90392D48CCFA37FAC3FA7F" box="[1257,1323,1419,1445]" pageId="14" pageNumber="15" rank="species">
<emphasis id="E18D8412FF90392D48CCFA37FAC3FA7F" bold="true" box="[1257,1323,1419,1445]" pageId="14" pageNumber="15">sp. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="D3465800FF90392D4D2DFA08FAC3F99C" blockId="14.[264,1325,580,1886]" pageId="14" pageNumber="15">
<keyLead id="6808FB35FF90392D4D2DFA08FAC3F99C" pageId="14" pageNumber="15">
– Temples 2.7­6.0 times as long as eyes. Elytra shorter than pronotum. Wings reduced to short vestiges, shorter than elytra. Apex of median lobe of aedeagus has different shape. Distributed in the Southern Appalachians (North Carolina and Tennessee) (
<figureCitation id="4BC24485FF90392D4D64F990FE0DF99C" box="[321,485,1580,1606]" captionStart-0="FIGURE 337" captionStart-1="FIGURE 338" captionStartId-0="77.[264,367,1248,1272]" captionStartId-1="79.[264,367,1838,1862]" captionTargetBox-0="[328,1262,539,1226]" captionTargetBox-1="[346,1237,315,1825]" captionTargetId-0="figure@77.[321,1266,529,1233]" captionTargetId-1="figure@79.[344,1238,312,1827]" captionTargetPageId-0="77" captionTargetPageId-1="79" captionText-0="FIGURE 337. Geographical distribution of Geostiba appalachigena Gusarov, sp. n. The ellipse indicates the area where eleven Southern Appalachian species are distributed." captionText-1="FIGURE 338. Geographical distribution of twelve species of Geostiba in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri-0="https://zenodo.org/record/155749/files/figure.png" httpUri-1="https://zenodo.org/record/155750/files/figure.png" pageId="14" pageNumber="15">Figs. 337­338</figureCitation>
) ............................................................................................................ 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="680D40A5FF90392D4D2DF9E8FDCDF884" pageId="14" pageNumber="15">
<paragraph id="D3465800FF90392D4D2DF9E8FAC4F94C" blockId="14.[264,1325,580,1886]" pageId="14" pageNumber="15">
<keyLead id="6808FB35FF90392D4D2DF9E8FAC4F94C" pageId="14" pageNumber="15">
<specimenCount id="C5FF9389FF90392D4D2DF9E8FE68F9B4" box="[264,384,1620,1646]" pageId="14" pageNumber="15" type="male">4 Males</specimenCount>
(In males posterior margin of abdominal sternum 8 convex and without a fringe of wide setae (
<figureCitation id="4BC24485FF90392D4DC6F9C0FDD1F94C" box="[483,569,1660,1686]" captionStart="FIGURES 29 ­ 33" captionStartId="20.[264,383,1785,1809]" captionTargetBox="[298,1291,699,1756]" captionTargetId="figure@20.[289,1298,689,1763]" captionTargetPageId="20" captionText="FIGURES 29 ­ 33. Abdominal segment 8 of Geostiba appalachigena Gusarov, sp. n. (paratypes from Black Mountain, West Virginia). 29 – male tergum 8; 30 – male sternum 8; 31 – female tergum 8; 32 – female sternum 8, 33 – apex of female sternum 8. Scale bar 0.2 mm (29 ­ 32), 0.1 mm (33)." httpUri="https://zenodo.org/record/155707/files/figure.png" pageId="14" pageNumber="15">Fig. 30</figureCitation>
) and some species with two medial carinae on tergum 7) ....... 5
</keyLead>
</paragraph>
<paragraph id="D3465800FF90392D4D2DF918FDCDF884" blockId="14.[264,1325,580,1886]" pageId="14" pageNumber="15">
<keyLead id="6808FB35FF90392D4D2DF918FDCDF884" pageId="14" pageNumber="15">
– Females (In females posterior margin of abdominal sternum 8 medially emarginate or concave, with a fringe of wide setae (
<figureCitation id="4BC24485FF90392D4EC8F970FC9EF93C" box="[749,886,1740,1766]" captionStart="FIGURES 29 ­ 33" captionStartId="20.[264,383,1785,1809]" captionTargetBox="[298,1291,699,1756]" captionTargetId="figure@20.[289,1298,689,1763]" captionTargetPageId="20" captionText="FIGURES 29 ­ 33. Abdominal segment 8 of Geostiba appalachigena Gusarov, sp. n. (paratypes from Black Mountain, West Virginia). 29 – male tergum 8; 30 – male sternum 8; 31 – female tergum 8; 32 – female sternum 8, 33 – apex of female sternum 8. Scale bar 0.2 mm (29 ­ 32), 0.1 mm (33)." httpUri="https://zenodo.org/record/155707/files/figure.png" pageId="14" pageNumber="15">Figs. 32­33</figureCitation>
,
<figureCitation id="4BC24485FF90392D4FA6F970FC23F93C" box="[899,971,1740,1766]" captionStart="FIGURES 51 ­ 56" captionStartId="24.[264,383,1769,1793]" captionTargetBox="[321,1256,734,1749]" captionTargetId="figure@24.[318,1259,731,1752]" captionTargetPageId="24" captionText="FIGURES 51 ­ 56. Abdominal segments 7 ­ 8 of Geostiba bicarinata Lohse &amp; Smetana (Richland Balsam Mountain, North Carolina). 51 – male tergum 8; 52 – male sternum 8; 53 – female tergum 8; 54 – female sternum 8, 55 – apex of female sternum 8; 56 – medial portion of male tergum 7, posterior down. Scale bar 0.2 mm (51 ­ 54, 56), 0.1 mm (55)." httpUri="https://zenodo.org/record/155710/files/figure.png" pageId="14" pageNumber="15">54­55</figureCitation>
). Females never have carinae on abdominal terga 7­8). Females can be reliably identified only by association with males or by comparing locality data with known geographical distribution of different species of
<taxonomicName id="14F92383FF90392D4D91F8F8FDF6F887" box="[436,542,1860,1885]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="E18D8412FF90392D4D91F8F8FDF6F887" box="[436,542,1860,1885]" italics="true" pageId="14" pageNumber="15">Geostiba</emphasis>
</taxonomicName>
.
</keyLead>
</paragraph>
</keyStep>
<keyStep id="680D40A5FF91392C4D2DFEA0FAC3FE5C" pageId="15" pageNumber="16">
<paragraph id="D3465800FF91392C4D2DFEA0FAC4FE84" blockId="15.[264,1325,284,1870]" pageId="15" pageNumber="16">
<keyLead id="6808FB35FF91392C4D2DFEA0FAC4FE84" pageId="15" pageNumber="16">5 Abdominal tergum 7 with two short and parallel medial carinae in front of posterior margin, tergum 8 with four short and weak carinae in front of posterior margin ....... 6</keyLead>
</paragraph>
<paragraph id="D3465800FF91392C4D2DFED0FAC3FE5C" blockId="15.[264,1325,284,1870]" box="[264,1323,364,390]" pageId="15" pageNumber="16">
<keyLead id="6808FB35FF91392C4D2DFED0FAC3FE5C" box="[264,1323,364,390]" pageId="15" pageNumber="16">– Abdominal terga 7 and 8 without carina or tubercles ................................................ 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="680D40A5FF91392C4D2DFE28FAC3FD44" pageId="15" pageNumber="16">
<paragraph id="D3465800FF91392C4D2DFE28FAC5FE24" blockId="15.[264,1325,284,1870]" pageId="15" pageNumber="16">
<keyLead id="6808FB35FF91392C4D2DFE28FAC5FE24" pageId="15" pageNumber="16">
6 Body length
<quantity id="1401F5E5FF91392C4DF4FE28FDB6FE74" box="[465,606,404,430]" metricMagnitude="-3" metricUnit="m" metricValue="2.0999999999999996" metricValueMax="2.3" metricValueMin="1.9" pageId="15" pageNumber="16" unit="mm" value="2.0999999999999996" valueMax="2.3" valueMin="1.9">1.9­2.3 mm</quantity>
, pronotal width
<quantity id="1401F5E5FF91392C4F3AFE28FC23FE74" box="[799,971,404,430]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" metricValueMax="4.6" metricValueMin="4.0" pageId="15" pageNumber="16" unit="mm" value="0.43" valueMax="0.46" valueMin="0.4">0.40­0.46 mm</quantity>
. Antennal segment 4 slightly transverse (width to length ratio about 1.2;
<figureCitation id="4BC24485FF91392C4F18FE00FC7EFE0C" box="[829,918,444,470]" captionStart="FIGURES 8 ­ 17" captionStartId="6.[264,383,1709,1733]" captionTargetBox="[307,1277,321,1687]" captionTargetId="figure@6.[297,1289,310,1697]" captionTargetPageId="6" captionText="FIGURES 8 ­ 17. Mouthparts of Geostiba circellaris (Gravenhorst) (8, Moscow Region, Russia), G. infirma (Weise) (9, Ivano­Frankovsk Region, Ukraine), G. carteriensis Pace (10 ­ 15, Tennessee), G. appalachigena Gusarov, sp. n. (16, paratype from West Virginia) and G. pluvigena Gusarov, sp. n. (17, paratype from North Carolina). 8 ­ 10, – prementum; 11 – hypopharynx; 12 – left maxilla, ventral view; 13 – left galea, dorsal view; 14 – left lacinia, dorsal view; 15 – mentum; 16 ­ 17 – right antenna. Scale bar 0.1 mm (8 ­ 15), 0.2 mm (16 ­ 17). L – Length of antennal article." httpUri="https://zenodo.org/record/155703/files/figure.png" pageId="15" pageNumber="16">Fig. 16</figureCitation>
). Apex of median lobe in ventral view broad (
<figureCitation id="4BC24485FF91392C4DEEFE58FDBCFE24" box="[459,596,484,510]" captionStart="FIGURES 57 ­ 60" captionStartId="25.[264,383,1274,1298]" captionTargetBox="[312,1277,687,1243]" captionTargetId="figure@25.[300,1287,678,1250]" captionTargetPageId="25" captionText="FIGURES 57 ­ 60. Aedeagus of Geostiba bicarinata Lohse &amp; Smetana (Richland Balsam Mountain, North Carolina). 57 – median lobe, ventral view; 58 – apex of median lobe, ventral view; 59 – median lobe, lateral view; 60 – apex of median lobe, lateral view. Scale bar 0.2 mm (57, 59), 0.1 mm (58, 60)." httpUri="https://zenodo.org/record/155711/files/figure.png" pageId="15" pageNumber="16">Figs. 57­58</figureCitation>
,
<figureCitation id="4BC24485FF91392C4E45FE58FD4EFE24" box="[608,678,484,510]" captionStart="FIGURES 77 ­ 80" captionStartId="29.[264,383,1121,1145]" captionTargetBox="[309,1279,539,1094]" captionTargetId="figure@29.[297,1290,529,1101]" captionTargetPageId="29" captionText="FIGURES 77 ­ 80. Aedeagus of Geostiba graveyardensis Pace (Graveyard Fields, North Carolina). 77 – median lobe, ventral view; 78 – apex of median lobe, ventral view; 79 – median lobe, lateral view; 80 – apex of median lobe, lateral view. Scale bar 0.2 mm (77, 79), 0.1 mm (78, 80)." httpUri="https://zenodo.org/record/155714/files/figure.png" pageId="15" pageNumber="16">77­78</figureCitation>
) .................................................................................. 7
</keyLead>
</paragraph>
<paragraph id="D3465800FF91392C4D2DFDB0FAC3FD44" blockId="15.[264,1325,284,1870]" pageId="15" pageNumber="16">
<keyLead id="6808FB35FF91392C4D2DFDB0FAC3FD44" pageId="15" pageNumber="16">
– Body length
<quantity id="1401F5E5FF91392C4DF5FDB0FDB4FDFC" box="[464,604,524,550]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.3" metricValueMin="1.6" pageId="15" pageNumber="16" unit="mm" value="1.95" valueMax="2.3" valueMin="1.6">1.6­2.3 mm</quantity>
, pronotal width
<quantity id="1401F5E5FF91392C4F39FDB0FC2DFDFC" box="[796,965,524,550]" metricMagnitude="-4" metricUnit="m" metricValue="3.85" metricValueMax="4.3" metricValueMin="3.4" pageId="15" pageNumber="16" unit="mm" value="0.385" valueMax="0.43" valueMin="0.34">0.34­0.43 mm</quantity>
. Antennal segment 4 strongly transverse (width to length ratio about 1.6;
<figureCitation id="4BC24485FF91392C4F19FD88FC7BFD94" box="[828,915,564,590]" captionStart="FIGURES 8 ­ 17" captionStartId="6.[264,383,1709,1733]" captionTargetBox="[307,1277,321,1687]" captionTargetId="figure@6.[297,1289,310,1697]" captionTargetPageId="6" captionText="FIGURES 8 ­ 17. Mouthparts of Geostiba circellaris (Gravenhorst) (8, Moscow Region, Russia), G. infirma (Weise) (9, Ivano­Frankovsk Region, Ukraine), G. carteriensis Pace (10 ­ 15, Tennessee), G. appalachigena Gusarov, sp. n. (16, paratype from West Virginia) and G. pluvigena Gusarov, sp. n. (17, paratype from North Carolina). 8 ­ 10, – prementum; 11 – hypopharynx; 12 – left maxilla, ventral view; 13 – left galea, dorsal view; 14 – left lacinia, dorsal view; 15 – mentum; 16 ­ 17 – right antenna. Scale bar 0.1 mm (8 ­ 15), 0.2 mm (16 ­ 17). L – Length of antennal article." httpUri="https://zenodo.org/record/155703/files/figure.png" pageId="15" pageNumber="16">Fig. 17</figureCitation>
). Apex of median lobe in ventral view narrow (
<figureCitation id="4BC24485FF91392C4DC1FDE0FD67FDAC" box="[484,655,604,630]" captionStart="FIGURES 100 ­ 107" captionStartId="33.[264,383,1742,1766]" captionTargetBox="[319,1271,659,1705]" captionTargetId="figure@33.[308,1278,649,1715]" captionTargetPageId="33" captionText="FIGURES 100 ­ 107. Aedeagus of Geostiba nimbicola Lohse &amp; Smetana (100 ­ 103 – holotype; 104 ­ 107 – Mount Sterling Ridge Trail, Tennessee). 100, 104 – median lobe, ventral view; 101, 105 – apex of median lobe, ventral view; 102, 106 – median lobe, lateral view; 103, 107 – apex of median lobe, lateral view. Scale bar 0.2 mm (100, 102, 104, 106), 0.1 mm (101, 103, 105, 107)." httpUri="https://zenodo.org/record/155717/files/figure.png" pageId="15" pageNumber="16">Figs. 100­101</figureCitation>
,
<figureCitation id="4BC24485FF91392C4E85FDE0FCEDFDAC" box="[672,773,604,630]" captionStart="FIGURES 126 ­ 133" captionStartId="36.[264,382,1701,1725]" captionTargetBox="[308,1279,632,1666]" captionTargetId="figure@36.[294,1294,622,1676]" captionTargetPageId="36" captionText="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 – median lobe, ventral view; 127, 131, 133 – apex of median lobe, ventral view; 128 – median lobe, lateral view; 129 – apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." httpUri="https://zenodo.org/record/155720/files/figure.png" pageId="15" pageNumber="16">126­127</figureCitation>
,
<figureCitation id="4BC24485FF91392C4F30FDE0FC90FDAC" box="[789,888,604,630]" captionStart="FIGURES 152 ­ 156" captionStartId="41.[264,383,1456,1480]" captionTargetBox="[369,1217,460,1420]" captionTargetId="figure@41.[361,1225,449,1430]" captionTargetPageId="41" captionText="FIGURES 152 ­ 156. Aedeagus of Geostiba nebuligena Gusarov, sp. n. (paratypes from Highways 441 &amp; 71, Tennessee (152, 154 ­ 156) and Appalachian National Scenic Trail W of Clingmans Dome (153), Tennessee / North Carolina). 152 – median lobe, ventral view; 153 ­ 154 – apex of median lobe, ventral view; 155 – median lobe, lateral view; 156 – apex of median lobe, lateral view. Scale bar 0.2 mm (152, 155), 0.1 mm (153 ­ 154, 156)." httpUri="https://zenodo.org/record/155723/files/figure.png" pageId="15" pageNumber="16">152­153</figureCitation>
) or broad (in
<taxonomicName id="14F92383FF91392C4808FDE0FB0DFDAF" box="[1069,1253,604,629]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="balsamensis">
<emphasis id="E18D8412FF91392C4808FDE0FB0DFDAF" box="[1069,1253,604,629]" italics="true" pageId="15" pageNumber="16">G. balsamensis</emphasis>
</taxonomicName>
only,
<figureCitation id="4BC24485FF91392C4D1DFD38FE34FD44" box="[312,476,644,670]" captionStart="FIGURES 193 ­ 196" captionStartId="48.[264,383,1727,1751]" captionTargetBox="[311,1277,1075,1699]" captionTargetId="figure@48.[305,1281,1068,1704]" captionTargetPageId="48" captionText="FIGURES 193 ­ 196. Aedeagus of Geostiba balsamensis Pace (Cataloochee Balsam Mountain, North Carolina). 193 – median lobe, ventral view; 194 – apex of median lobe, ventral view; 195 – median lobe, lateral view; 196 – apex of median lobe, lateral view. Scale bar 0.2 mm (193, 195), 0.1 mm (194, 196)." httpUri="https://zenodo.org/record/155729/files/figure.png" pageId="15" pageNumber="16">Figs. 193­194</figureCitation>
) ............................................................................................................. 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="680D40A5FF91392C4D2DFD10FAC2FC04" pageId="15" pageNumber="16">
<paragraph id="D3465800FF91392C4D2DFD10FAC3FCE4" blockId="15.[264,1325,284,1870]" pageId="15" pageNumber="16">
<keyLead id="6808FB35FF91392C4D2DFD10FAC3FCE4" pageId="15" pageNumber="16">
7 Apex of median lobe of aedeagus in lateral view not bent ventrally (
<figureCitation id="4BC24485FF91392C48AAFD10FAF6FD1C" box="[1167,1310,684,710]" captionStart="FIGURES 57 ­ 60" captionStartId="25.[264,383,1274,1298]" captionTargetBox="[312,1277,687,1243]" captionTargetId="figure@25.[300,1287,678,1250]" captionTargetPageId="25" captionText="FIGURES 57 ­ 60. Aedeagus of Geostiba bicarinata Lohse &amp; Smetana (Richland Balsam Mountain, North Carolina). 57 – median lobe, ventral view; 58 – apex of median lobe, ventral view; 59 – median lobe, lateral view; 60 – apex of median lobe, lateral view. Scale bar 0.2 mm (57, 59), 0.1 mm (58, 60)." httpUri="https://zenodo.org/record/155711/files/figure.png" pageId="15" pageNumber="16">Figs. 59­60</figureCitation>
). Body length 2.0­
<quantity id="1401F5E5FF91392C4E24FD68FDB7FD34" box="[513,607,724,750]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="15" pageNumber="16" unit="mm" value="2.3">2.3 mm</quantity>
, pronotal width
<quantity id="1401F5E5FF91392C4F07FD68FC25FD34" box="[802,973,724,750]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" metricValueMax="4.6" metricValueMin="4.0" pageId="15" pageNumber="16" unit="mm" value="0.43" valueMax="0.46" valueMin="0.4">0.40­0.46 mm</quantity>
. Known from the Great Balsam Mountains and Little Pisgah Mountain in the Southern Appalachians(
<figureCitation id="4BC24485FF91392C4859FD40FB02FCCC" box="[1148,1258,764,790]" captionStart="FIGURE 338" captionStartId="79.[264,367,1838,1862]" captionTargetBox="[346,1237,315,1825]" captionTargetId="figure@79.[344,1238,312,1827]" captionTargetPageId="79" captionText="FIGURE 338. Geographical distribution of twelve species of Geostiba in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155750/files/figure.png" pageId="15" pageNumber="16">Figs. 338</figureCitation>
,
<figureCitation id="4BC24485FF91392C48D4FD40FAF7FCCC" box="[1265,1311,764,790]" captionStart="FIGURE 340" captionStartId="82.[264,368,1747,1771]" captionTargetBox="[322,1272,508,1726]" captionTargetId="figure@82.[319,1275,508,1728]" captionTargetPageId="82" captionText="FIGURE 340. Detailed geographical distribution of Geostiba bicarinata Lohse &amp; Smetana (2), G. graveyardensis Pace (3), G. nimbicola Lohse &amp; Smetana (4), G. pluvigena Gusarov, sp. n. (5), G. nebuligena Gusarov, sp. n. (6), G. crepusculigena Gusarov, sp. n. (7), G. balsamensis Pace (8), G. nubigena Lohse &amp; Smetana (9), G. coeligena Gusarov, sp. n. (10) in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155752/files/figure.png" pageId="15" pageNumber="16">340</figureCitation>
) ...................................................................2.
<taxonomicName id="14F92383FF91392C4F63FC98FAC3FCE4" authority="Lohse &amp; Smetana" authorityName="Lohse &amp; Smetana" box="[838,1323,804,830]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bicarinata" subGenus="Sibiota">
<emphasis id="E18D8412FF91392C4F63FC98FC8AFCE7" box="[838,866,804,829]" italics="true" pageId="15" pageNumber="16">G.</emphasis>
(
<emphasis id="E18D8412FF91392C4F56FC98FC2EFCE7" box="[883,966,804,829]" italics="true" pageId="15" pageNumber="16">Sibiota</emphasis>
)
<emphasis id="E18D8412FF91392C4FF2FC98FBB9FCE7" box="[983,1105,804,829]" italics="true" pageId="15" pageNumber="16">bicarinata</emphasis>
Lohse &amp; Smetana
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="D3465800FF91392C4D2DFCF0FAC2FC04" blockId="15.[264,1325,284,1870]" pageId="15" pageNumber="16">
<keyLead id="6808FB35FF91392C4D2DFCF0FAC2FC04" pageId="15" pageNumber="16">
– Median lobe of aedeagus in lateral view bent ventrally (
<figureCitation id="4BC24485FF91392C4FE7FCF0FBA4FCBC" box="[962,1100,844,870]" captionStart="FIGURES 77 ­ 80" captionStartId="29.[264,383,1121,1145]" captionTargetBox="[309,1279,539,1094]" captionTargetId="figure@29.[297,1290,529,1101]" captionTargetPageId="29" captionText="FIGURES 77 ­ 80. Aedeagus of Geostiba graveyardensis Pace (Graveyard Fields, North Carolina). 77 – median lobe, ventral view; 78 – apex of median lobe, ventral view; 79 – median lobe, lateral view; 80 – apex of median lobe, lateral view. Scale bar 0.2 mm (77, 79), 0.1 mm (78, 80)." httpUri="https://zenodo.org/record/155714/files/figure.png" pageId="15" pageNumber="16">Figs. 79­80</figureCitation>
). Body length
<quantity id="1401F5E5FF91392C48D8FCF0FE7DFC54" metricMagnitude="-3" metricUnit="m" metricValue="2.0999999999999996" metricValueMax="2.3" metricValueMin="1.9" pageId="15" pageNumber="16" unit="mm" value="2.0999999999999996" valueMax="2.3" valueMin="1.9">1.9­ 2.3 mm</quantity>
, pronotal width
<quantity id="1401F5E5FF91392C4E76FCC8FD14FC54" box="[595,764,884,910]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.4" metricValueMin="4.1" pageId="15" pageNumber="16" unit="mm" value="0.425" valueMax="0.44" valueMin="0.41">0.41­0.44 mm</quantity>
. Known from Graveyard Ridge and Black Balsam Knob in the Great Balsam Mountains – Pisgah Ridge massif in the Southern Appalachians (
<figureCitation id="4BC24485FF91392C4DC2FC78FDB0FC04" box="[487,600,964,990]" captionStart="FIGURE 338" captionStartId="79.[264,367,1838,1862]" captionTargetBox="[346,1237,315,1825]" captionTargetId="figure@79.[344,1238,312,1827]" captionTargetPageId="79" captionText="FIGURE 338. Geographical distribution of twelve species of Geostiba in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155750/files/figure.png" pageId="15" pageNumber="16">Figs. 338</figureCitation>
,
<figureCitation id="4BC24485FF91392C4E41FC78FD79FC04" box="[612,657,964,990]" captionStart="FIGURE 340" captionStartId="82.[264,368,1747,1771]" captionTargetBox="[322,1272,508,1726]" captionTargetId="figure@82.[319,1275,508,1728]" captionTargetPageId="82" captionText="FIGURE 340. Detailed geographical distribution of Geostiba bicarinata Lohse &amp; Smetana (2), G. graveyardensis Pace (3), G. nimbicola Lohse &amp; Smetana (4), G. pluvigena Gusarov, sp. n. (5), G. nebuligena Gusarov, sp. n. (6), G. crepusculigena Gusarov, sp. n. (7), G. balsamensis Pace (8), G. nubigena Lohse &amp; Smetana (9), G. coeligena Gusarov, sp. n. (10) in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155752/files/figure.png" pageId="15" pageNumber="16">340</figureCitation>
) .............................. 3.
<taxonomicName id="14F92383FF91392C4F8DFC78FAC2FC04" authority="Pace" authorityName="Pace" box="[936,1322,964,990]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="graveyardensis" subGenus="Sibiota">
<emphasis id="E18D8412FF91392C4F8DFC78FC2DFC07" box="[936,965,964,989]" italics="true" pageId="15" pageNumber="16">G.</emphasis>
(
<emphasis id="E18D8412FF91392C4FF3FC78FBC1FC07" box="[982,1065,964,989]" italics="true" pageId="15" pageNumber="16">Sibiota</emphasis>
)
<emphasis id="E18D8412FF91392C481FFC78FB04FC07" box="[1082,1260,964,989]" italics="true" pageId="15" pageNumber="16">graveyardensis</emphasis>
Pace
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="680D40A5FF91392C4D2DFC50FAC5FB14" pageId="15" pageNumber="16">
<paragraph id="D3465800FF91392C4D2DFC50FAC2FBA4" blockId="15.[264,1325,284,1870]" pageId="15" pageNumber="16">
<keyLead id="6808FB35FF91392C4D2DFC50FAC2FBA4" pageId="15" pageNumber="16">
8 Apex of median lobe in ventral view broad (
<figureCitation id="4BC24485FF91392C4F6EFC50FC1DFBDC" box="[843,1013,1004,1030]" captionStart="FIGURES 193 ­ 196" captionStartId="48.[264,383,1727,1751]" captionTargetBox="[311,1277,1075,1699]" captionTargetId="figure@48.[305,1281,1068,1704]" captionTargetPageId="48" captionText="FIGURES 193 ­ 196. Aedeagus of Geostiba balsamensis Pace (Cataloochee Balsam Mountain, North Carolina). 193 – median lobe, ventral view; 194 – apex of median lobe, ventral view; 195 – median lobe, lateral view; 196 – apex of median lobe, lateral view. Scale bar 0.2 mm (193, 195), 0.1 mm (194, 196)." httpUri="https://zenodo.org/record/155729/files/figure.png" pageId="15" pageNumber="16">Figs. 193­194</figureCitation>
). Medial carinae on male tergum 7 weak. Body length
<quantity id="1401F5E5FF91392C4EB7FBA8FCF6FBF4" box="[658,798,1044,1070]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="15" pageNumber="16" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8­2.2 mm</quantity>
, pronotal width
<quantity id="1401F5E5FF91392C4FF9FBA8FB6DFBF4" box="[988,1157,1044,1070]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="3.9" metricValueMin="3.5999999999999996" pageId="15" pageNumber="16" unit="mm" value="0.375" valueMax="0.39" valueMin="0.36">0.36­0.39 mm</quantity>
. Known from the Plott Balsams – Cataloochee Divide massif in the Southern Appalachians (
<figureCitation id="4BC24485FF91392C48D5FB80FE8EFBA4" captionStart="FIGURE 338" captionStartId="79.[264,367,1838,1862]" captionTargetBox="[346,1237,315,1825]" captionTargetId="figure@79.[344,1238,312,1827]" captionTargetPageId="79" captionText="FIGURE 338. Geographical distribution of twelve species of Geostiba in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155750/files/figure.png" pageId="15" pageNumber="16">Figs. 338</figureCitation>
,
<figureCitation id="4BC24485FF91392C4D56FBD8FE48FBA4" box="[371,416,1124,1150]" captionStart="FIGURE 340" captionStartId="82.[264,368,1747,1771]" captionTargetBox="[322,1272,508,1726]" captionTargetId="figure@82.[319,1275,508,1728]" captionTargetPageId="82" captionText="FIGURE 340. Detailed geographical distribution of Geostiba bicarinata Lohse &amp; Smetana (2), G. graveyardensis Pace (3), G. nimbicola Lohse &amp; Smetana (4), G. pluvigena Gusarov, sp. n. (5), G. nebuligena Gusarov, sp. n. (6), G. crepusculigena Gusarov, sp. n. (7), G. balsamensis Pace (8), G. nubigena Lohse &amp; Smetana (9), G. coeligena Gusarov, sp. n. (10) in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155752/files/figure.png" pageId="15" pageNumber="16">340</figureCitation>
) ................................................................... 8.
<taxonomicName id="14F92383FF91392C4FEEFBD8FAC2FBA4" authority="Pace" authorityName="Pace" box="[971,1322,1124,1150]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="balsamensis" subGenus="Sibiota">
<emphasis id="E18D8412FF91392C4FEEFBD8FC0FFBA7" box="[971,999,1124,1149]" italics="true" pageId="15" pageNumber="16">G.</emphasis>
(
<emphasis id="E18D8412FF91392C4FDDFBD8FBA3FBA7" box="[1016,1099,1124,1149]" italics="true" pageId="15" pageNumber="16">Sibiota</emphasis>
)
<emphasis id="E18D8412FF91392C4878FBD8FB04FBA7" box="[1117,1260,1124,1149]" italics="true" pageId="15" pageNumber="16">balsamensis</emphasis>
Pace
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="D3465800FF91392C4D2DFB30FAC5FB14" blockId="15.[264,1325,284,1870]" pageId="15" pageNumber="16">
<keyLead id="6808FB35FF91392C4D2DFB30FAC5FB14" pageId="15" pageNumber="16">
– Apex of median lobe in ventral view narrow (
<figureCitation id="4BC24485FF91392C4FA6FB30FBC7FB7C" box="[899,1071,1164,1190]" captionStart="FIGURES 100 ­ 107" captionStartId="33.[264,383,1742,1766]" captionTargetBox="[319,1271,659,1705]" captionTargetId="figure@33.[308,1278,649,1715]" captionTargetPageId="33" captionText="FIGURES 100 ­ 107. Aedeagus of Geostiba nimbicola Lohse &amp; Smetana (100 ­ 103 – holotype; 104 ­ 107 – Mount Sterling Ridge Trail, Tennessee). 100, 104 – median lobe, ventral view; 101, 105 – apex of median lobe, ventral view; 102, 106 – median lobe, lateral view; 103, 107 – apex of median lobe, lateral view. Scale bar 0.2 mm (100, 102, 104, 106), 0.1 mm (101, 103, 105, 107)." httpUri="https://zenodo.org/record/155717/files/figure.png" pageId="15" pageNumber="16">Figs. 100­101</figureCitation>
,
<figureCitation id="4BC24485FF91392C4867FB30FB4FFB7C" box="[1090,1191,1164,1190]" captionStart="FIGURES 126 ­ 133" captionStartId="36.[264,382,1701,1725]" captionTargetBox="[308,1279,632,1666]" captionTargetId="figure@36.[294,1294,622,1676]" captionTargetPageId="36" captionText="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 – median lobe, ventral view; 127, 131, 133 – apex of median lobe, ventral view; 128 – median lobe, lateral view; 129 – apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." httpUri="https://zenodo.org/record/155720/files/figure.png" pageId="15" pageNumber="16">126­127</figureCitation>
,
<figureCitation id="4BC24485FF91392C489CFB30FAF7FB7C" box="[1209,1311,1164,1190]" captionStart="FIGURES 152 ­ 156" captionStartId="41.[264,383,1456,1480]" captionTargetBox="[369,1217,460,1420]" captionTargetId="figure@41.[361,1225,449,1430]" captionTargetPageId="41" captionText="FIGURES 152 ­ 156. Aedeagus of Geostiba nebuligena Gusarov, sp. n. (paratypes from Highways 441 &amp; 71, Tennessee (152, 154 ­ 156) and Appalachian National Scenic Trail W of Clingmans Dome (153), Tennessee / North Carolina). 152 – median lobe, ventral view; 153 ­ 154 – apex of median lobe, ventral view; 155 – median lobe, lateral view; 156 – apex of median lobe, lateral view. Scale bar 0.2 mm (152, 155), 0.1 mm (153 ­ 154, 156)." httpUri="https://zenodo.org/record/155723/files/figure.png" pageId="15" pageNumber="16">152­153</figureCitation>
). Medial carinae on male tergum 7 strong ..................................................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="680D40A5FF91392C4D2DFB60FAC2FA64" pageId="15" pageNumber="16">
<paragraph id="D3465800FF91392C4D2DFB60FAC3FAB7" blockId="15.[264,1325,284,1870]" pageId="15" pageNumber="16">
<keyLead id="6808FB35FF91392C4D2DFB60FAC3FAB7" pageId="15" pageNumber="16">
9 Apex of median lobe in lateral view with obtuse ventral projection (
<figureCitation id="4BC24485FF91392C4856FB60FAF7FB2C" box="[1139,1311,1244,1270]" captionStart="FIGURES 174 ­ 177" captionStartId="45.[264,383,1766,1790]" captionTargetBox="[317,1274,1164,1735]" captionTargetId="figure@45.[305,1283,1154,1742]" captionTargetPageId="45" captionText="FIGURES 174 ­ 177. Aedeagus of Geostiba crepusculigena Gusarov, sp. n. (paratype from Snake Ridge Trail, Tennessee). 174 – median lobe, ventral view; 175 – apex of median lobe, ventral view; 176 – median lobe, lateral view; 177 – apex of median lobe, lateral view. Scale bar 0.2 mm (174, 176), 0.1 mm (175, 177)." httpUri="https://zenodo.org/record/155726/files/figure.png" pageId="15" pageNumber="16">Figs. 176­177</figureCitation>
). Body length
<quantity id="1401F5E5FF91392C4DFCFAB8FD82FAC4" box="[473,618,1284,1310]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="15" pageNumber="16" unit="mm" value="2.05" valueMax="2.3" valueMin="1.8">1.8­2.3 mm</quantity>
, pronotal width
<quantity id="1401F5E5FF91392C4F12FAB8FC0DFAC4" box="[823,997,1284,1310]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" metricValueMax="4.0" metricValueMin="3.4" pageId="15" pageNumber="16" unit="mm" value="0.37" valueMax="0.4" valueMin="0.34">0.34­0.40 mm</quantity>
. Known from Snake Den Mountain in the Great Smoky Mountains massif (
<figureCitation id="4BC24485FF91392C4FA5FA90FC19FA9C" box="[896,1009,1324,1350]" captionStart="FIGURE 338" captionStartId="79.[264,367,1838,1862]" captionTargetBox="[346,1237,315,1825]" captionTargetId="figure@79.[344,1238,312,1827]" captionTargetPageId="79" captionText="FIGURE 338. Geographical distribution of twelve species of Geostiba in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155750/files/figure.png" pageId="15" pageNumber="16">Figs. 338</figureCitation>
,
<figureCitation id="4BC24485FF91392C4FD8FA90FBC2FA9C" box="[1021,1066,1324,1350]" captionStart="FIGURE 340" captionStartId="82.[264,368,1747,1771]" captionTargetBox="[322,1272,508,1726]" captionTargetId="figure@82.[319,1275,508,1728]" captionTargetPageId="82" captionText="FIGURE 340. Detailed geographical distribution of Geostiba bicarinata Lohse &amp; Smetana (2), G. graveyardensis Pace (3), G. nimbicola Lohse &amp; Smetana (4), G. pluvigena Gusarov, sp. n. (5), G. nebuligena Gusarov, sp. n. (6), G. crepusculigena Gusarov, sp. n. (7), G. balsamensis Pace (8), G. nubigena Lohse &amp; Smetana (9), G. coeligena Gusarov, sp. n. (10) in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155752/files/figure.png" pageId="15" pageNumber="16">340</figureCitation>
) ................................................................................................7.
<taxonomicName id="14F92383FF91392C4F0BFAE8FB35FAB4" authority="Gusarov" authorityName="Gusarov" box="[814,1245,1364,1390]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="crepusculigena" status="sp. nov." subGenus="Sibiota">
<emphasis id="E18D8412FF91392C4F0BFAE8FCA2FAB7" box="[814,842,1364,1389]" italics="true" pageId="15" pageNumber="16">G.</emphasis>
(
<emphasis id="E18D8412FF91392C4F7EFAE8FC46FAB7" box="[859,942,1364,1389]" italics="true" pageId="15" pageNumber="16">Sibiota</emphasis>
)
<emphasis id="E18D8412FF91392C4F9AFAE8FB99FAB7" box="[959,1137,1364,1389]" italics="true" pageId="15" pageNumber="16">crepusculigena</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="FABE3969FF91392C48CCFAEFFAC3FAB7" box="[1257,1323,1363,1389]" pageId="15" pageNumber="16" rank="species">
<emphasis id="E18D8412FF91392C48CCFAEFFAC3FAB7" bold="true" box="[1257,1323,1363,1389]" pageId="15" pageNumber="16">sp. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="D3465800FF91392C4D2DFAC0FAC2FA64" blockId="15.[264,1325,284,1870]" pageId="15" pageNumber="16">
<keyLead id="6808FB35FF91392C4D2DFAC0FAC2FA64" pageId="15" pageNumber="16">
– Apex of median lobe in lateral view without obtuse ventral projection (
<figureCitation id="4BC24485FF91392C48A5FAC0FACFFA4C" box="[1152,1319,1404,1430]" captionStart="FIGURES 100 ­ 107" captionStartId="33.[264,383,1742,1766]" captionTargetBox="[319,1271,659,1705]" captionTargetId="figure@33.[308,1278,649,1715]" captionTargetPageId="33" captionText="FIGURES 100 ­ 107. Aedeagus of Geostiba nimbicola Lohse &amp; Smetana (100 ­ 103 – holotype; 104 ­ 107 – Mount Sterling Ridge Trail, Tennessee). 100, 104 – median lobe, ventral view; 101, 105 – apex of median lobe, ventral view; 102, 106 – median lobe, lateral view; 103, 107 – apex of median lobe, lateral view. Scale bar 0.2 mm (100, 102, 104, 106), 0.1 mm (101, 103, 105, 107)." httpUri="https://zenodo.org/record/155717/files/figure.png" pageId="15" pageNumber="16">Figs. 102­103</figureCitation>
,
<figureCitation id="4BC24485FF91392C4D1DFA18FE74FA64" box="[312,412,1444,1470]" captionStart="FIGURES 126 ­ 133" captionStartId="36.[264,382,1701,1725]" captionTargetBox="[308,1279,632,1666]" captionTargetId="figure@36.[294,1294,622,1676]" captionTargetPageId="36" captionText="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 – median lobe, ventral view; 127, 131, 133 – apex of median lobe, ventral view; 128 – median lobe, lateral view; 129 – apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." httpUri="https://zenodo.org/record/155720/files/figure.png" pageId="15" pageNumber="16">128­129</figureCitation>
,
<figureCitation id="4BC24485FF91392C4D8DFA18FDE3FA64" box="[424,523,1444,1470]" captionStart="FIGURES 152 ­ 156" captionStartId="41.[264,383,1456,1480]" captionTargetBox="[369,1217,460,1420]" captionTargetId="figure@41.[361,1225,449,1430]" captionTargetPageId="41" captionText="FIGURES 152 ­ 156. Aedeagus of Geostiba nebuligena Gusarov, sp. n. (paratypes from Highways 441 &amp; 71, Tennessee (152, 154 ­ 156) and Appalachian National Scenic Trail W of Clingmans Dome (153), Tennessee / North Carolina). 152 – median lobe, ventral view; 153 ­ 154 – apex of median lobe, ventral view; 155 – median lobe, lateral view; 156 – apex of median lobe, lateral view. Scale bar 0.2 mm (152, 155), 0.1 mm (153 ­ 154, 156)." httpUri="https://zenodo.org/record/155723/files/figure.png" pageId="15" pageNumber="16">155­156</figureCitation>
) ..................................................................................................... 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="680D40A5FF91392C4D2DFA70FAC2F924" pageId="15" pageNumber="16">
<paragraph id="D3465800FF91392C4D2DFA70FAC3F95C" blockId="15.[264,1325,284,1870]" pageId="15" pageNumber="16">
<keyLead id="6808FB35FF91392C4D2DFA70FAC3F95C" pageId="15" pageNumber="16">
10 Apex of median lobe longer (
<figureCitation id="4BC24485FF91392C4EB2FA70FCA8FA3C" box="[663,832,1484,1510]" captionStart="FIGURES 100 ­ 107" captionStartId="33.[264,383,1742,1766]" captionTargetBox="[319,1271,659,1705]" captionTargetId="figure@33.[308,1278,649,1715]" captionTargetPageId="33" captionText="FIGURES 100 ­ 107. Aedeagus of Geostiba nimbicola Lohse &amp; Smetana (100 ­ 103 – holotype; 104 ­ 107 – Mount Sterling Ridge Trail, Tennessee). 100, 104 – median lobe, ventral view; 101, 105 – apex of median lobe, ventral view; 102, 106 – median lobe, lateral view; 103, 107 – apex of median lobe, lateral view. Scale bar 0.2 mm (100, 102, 104, 106), 0.1 mm (101, 103, 105, 107)." httpUri="https://zenodo.org/record/155717/files/figure.png" pageId="15" pageNumber="16">Figs. 100­107</figureCitation>
), in ventral view the apex outline convex apically (
<figureCitation id="4BC24485FF91392C4D8FFA48FD88F9D4" box="[426,608,1524,1550]" captionStart="FIGURES 100 ­ 107" captionStartId="33.[264,383,1742,1766]" captionTargetBox="[319,1271,659,1705]" captionTargetId="figure@33.[308,1278,649,1715]" captionTargetPageId="33" captionText="FIGURES 100 ­ 107. Aedeagus of Geostiba nimbicola Lohse &amp; Smetana (100 ­ 103 – holotype; 104 ­ 107 – Mount Sterling Ridge Trail, Tennessee). 100, 104 – median lobe, ventral view; 101, 105 – apex of median lobe, ventral view; 102, 106 – median lobe, lateral view; 103, 107 – apex of median lobe, lateral view. Scale bar 0.2 mm (100, 102, 104, 106), 0.1 mm (101, 103, 105, 107)." httpUri="https://zenodo.org/record/155717/files/figure.png" pageId="15" pageNumber="16">Figs. 101, 105</figureCitation>
), in lateral view without denticle (
<figureCitation id="4BC24485FF91392C482AFA48FB2EF9D4" box="[1039,1222,1524,1550]" captionStart="FIGURES 100 ­ 107" captionStartId="33.[264,383,1742,1766]" captionTargetBox="[319,1271,659,1705]" captionTargetId="figure@33.[308,1278,649,1715]" captionTargetPageId="33" captionText="FIGURES 100 ­ 107. Aedeagus of Geostiba nimbicola Lohse &amp; Smetana (100 ­ 103 – holotype; 104 ­ 107 – Mount Sterling Ridge Trail, Tennessee). 100, 104 – median lobe, ventral view; 101, 105 – apex of median lobe, ventral view; 102, 106 – median lobe, lateral view; 103, 107 – apex of median lobe, lateral view. Scale bar 0.2 mm (100, 102, 104, 106), 0.1 mm (101, 103, 105, 107)." httpUri="https://zenodo.org/record/155717/files/figure.png" pageId="15" pageNumber="16">Figs. 103, 107</figureCitation>
). Body length 1.6­2.0 mm, pronotal width
<quantity id="1401F5E5FF91392C4EF0F9A0FC97F9EC" box="[725,895,1564,1590]" metricMagnitude="-4" metricUnit="m" metricValue="3.85" metricValueMax="4.0" metricValueMin="3.7" pageId="15" pageNumber="16" unit="mm" value="0.385" valueMax="0.4" valueMin="0.37">0.37­0.40 mm</quantity>
. Known from many localities in the Great Smoky Mountains massif (
<figureCitation id="4BC24485FF91392C4E99F9F8FCC5F984" box="[700,813,1604,1630]" captionStart="FIGURE 338" captionStartId="79.[264,367,1838,1862]" captionTargetBox="[346,1237,315,1825]" captionTargetId="figure@79.[344,1238,312,1827]" captionTargetPageId="79" captionText="FIGURE 338. Geographical distribution of twelve species of Geostiba in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155750/files/figure.png" pageId="15" pageNumber="16">Figs. 338</figureCitation>
,
<figureCitation id="4BC24485FF91392C4F1DF9F8FC8DF984" box="[824,869,1604,1630]" captionStart="FIGURE 340" captionStartId="82.[264,368,1747,1771]" captionTargetBox="[322,1272,508,1726]" captionTargetId="figure@82.[319,1275,508,1728]" captionTargetPageId="82" captionText="FIGURE 340. Detailed geographical distribution of Geostiba bicarinata Lohse &amp; Smetana (2), G. graveyardensis Pace (3), G. nimbicola Lohse &amp; Smetana (4), G. pluvigena Gusarov, sp. n. (5), G. nebuligena Gusarov, sp. n. (6), G. crepusculigena Gusarov, sp. n. (7), G. balsamensis Pace (8), G. nubigena Lohse &amp; Smetana (9), G. coeligena Gusarov, sp. n. (10) in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155752/files/figure.png" pageId="15" pageNumber="16">340</figureCitation>
) ..............................................................................................................................4.
<taxonomicName id="14F92383FF91392C4F6FF9D0FAC3F95C" authority="Lohse &amp; Smetana" authorityName="Lohse &amp; Smetana" box="[842,1323,1644,1670]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="nimbicola" subGenus="Sibiota">
<emphasis id="E18D8412FF91392C4F6FF9D0FC8FF95F" box="[842,871,1644,1669]" italics="true" pageId="15" pageNumber="16">G.</emphasis>
(
<emphasis id="E18D8412FF91392C4F5DF9D0FC23F95F" box="[888,971,1644,1669]" italics="true" pageId="15" pageNumber="16">Sibiota</emphasis>
)
<emphasis id="E18D8412FF91392C4FF9F9D0FBB9F95F" box="[988,1105,1644,1669]" italics="true" pageId="15" pageNumber="16">nimbicola</emphasis>
Lohse &amp; Smetana
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="D3465800FF91392C4D2DF928FAC2F924" blockId="15.[264,1325,284,1870]" pageId="15" pageNumber="16">
<keyLead id="6808FB35FF91392C4D2DF928FAC2F924" pageId="15" pageNumber="16">
– Apex of median lobe shorter (
<figureCitation id="4BC24485FF91392C4E85F928FCA0F974" box="[672,840,1684,1710]" captionStart="FIGURES 126 ­ 133" captionStartId="36.[264,382,1701,1725]" captionTargetBox="[308,1279,632,1666]" captionTargetId="figure@36.[294,1294,622,1676]" captionTargetPageId="36" captionText="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 – median lobe, ventral view; 127, 131, 133 – apex of median lobe, ventral view; 128 – median lobe, lateral view; 129 – apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." httpUri="https://zenodo.org/record/155720/files/figure.png" pageId="15" pageNumber="16">Figs. 126­133</figureCitation>
,
<figureCitation id="4BC24485FF91392C4F73F928FC53F974" box="[854,955,1684,1710]" captionStart="FIGURES 152 ­ 156" captionStartId="41.[264,383,1456,1480]" captionTargetBox="[369,1217,460,1420]" captionTargetId="figure@41.[361,1225,449,1430]" captionTargetPageId="41" captionText="FIGURES 152 ­ 156. Aedeagus of Geostiba nebuligena Gusarov, sp. n. (paratypes from Highways 441 &amp; 71, Tennessee (152, 154 ­ 156) and Appalachian National Scenic Trail W of Clingmans Dome (153), Tennessee / North Carolina). 152 – median lobe, ventral view; 153 ­ 154 – apex of median lobe, ventral view; 155 – median lobe, lateral view; 156 – apex of median lobe, lateral view. Scale bar 0.2 mm (152, 155), 0.1 mm (153 ­ 154, 156)." httpUri="https://zenodo.org/record/155723/files/figure.png" pageId="15" pageNumber="16">152­156</figureCitation>
), in ventral view the apex outline often doubly emarginate (
<figureCitation id="4BC24485FF91392C4E8BF900FCCCF90C" box="[686,804,1724,1750]" captionStart="FIGURES 126 ­ 133" captionStartId="36.[264,382,1701,1725]" captionTargetBox="[308,1279,632,1666]" captionTargetId="figure@36.[294,1294,622,1676]" captionTargetPageId="36" captionText="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 – median lobe, ventral view; 127, 131, 133 – apex of median lobe, ventral view; 128 – median lobe, lateral view; 129 – apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." httpUri="https://zenodo.org/record/155720/files/figure.png" pageId="15" pageNumber="16">Figs. 131</figureCitation>
,
<figureCitation id="4BC24485FF91392C4F13F900FC73F90C" box="[822,923,1724,1750]" captionStart="FIGURES 152 ­ 156" captionStartId="41.[264,383,1456,1480]" captionTargetBox="[369,1217,460,1420]" captionTargetId="figure@41.[361,1225,449,1430]" captionTargetPageId="41" captionText="FIGURES 152 ­ 156. Aedeagus of Geostiba nebuligena Gusarov, sp. n. (paratypes from Highways 441 &amp; 71, Tennessee (152, 154 ­ 156) and Appalachian National Scenic Trail W of Clingmans Dome (153), Tennessee / North Carolina). 152 – median lobe, ventral view; 153 ­ 154 – apex of median lobe, ventral view; 155 – median lobe, lateral view; 156 – apex of median lobe, lateral view. Scale bar 0.2 mm (152, 155), 0.1 mm (153 ­ 154, 156)." httpUri="https://zenodo.org/record/155723/files/figure.png" pageId="15" pageNumber="16">153­154</figureCitation>
), in lateral view with a denticle (
<figureCitation id="4BC24485FF91392C4D64F958FE0FF924" box="[321,487,1764,1790]" captionStart="FIGURES 126 ­ 133" captionStartId="36.[264,382,1701,1725]" captionTargetBox="[308,1279,632,1666]" captionTargetId="figure@36.[294,1294,622,1676]" captionTargetPageId="36" captionText="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 – median lobe, ventral view; 127, 131, 133 – apex of median lobe, ventral view; 128 – median lobe, lateral view; 129 – apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." httpUri="https://zenodo.org/record/155720/files/figure.png" pageId="15" pageNumber="16">Figs. 128­129</figureCitation>
,
<figureCitation id="4BC24485FF91392C4DD6F958FDBDF924" box="[499,597,1764,1790]" captionStart="FIGURES 152 ­ 156" captionStartId="41.[264,383,1456,1480]" captionTargetBox="[369,1217,460,1420]" captionTargetId="figure@41.[361,1225,449,1430]" captionTargetPageId="41" captionText="FIGURES 152 ­ 156. Aedeagus of Geostiba nebuligena Gusarov, sp. n. (paratypes from Highways 441 &amp; 71, Tennessee (152, 154 ­ 156) and Appalachian National Scenic Trail W of Clingmans Dome (153), Tennessee / North Carolina). 152 – median lobe, ventral view; 153 ­ 154 – apex of median lobe, ventral view; 155 – median lobe, lateral view; 156 – apex of median lobe, lateral view. Scale bar 0.2 mm (152, 155), 0.1 mm (153 ­ 154, 156)." httpUri="https://zenodo.org/record/155723/files/figure.png" pageId="15" pageNumber="16">155­156</figureCitation>
) ........................................................................................... 11
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<paragraph id="D3465800FF91392C4D2DF8B0FAC3F894" blockId="15.[264,1325,284,1870]" pageId="15" pageNumber="16">
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11 Apex of median lobe in lateral view with a strong ventral denticle (
<figureCitation id="4BC24485FF91392C4873F8B0FB17F8FC" box="[1110,1279,1804,1830]" captionStart="FIGURES 152 ­ 156" captionStartId="41.[264,383,1456,1480]" captionTargetBox="[369,1217,460,1420]" captionTargetId="figure@41.[361,1225,449,1430]" captionTargetPageId="41" captionText="FIGURES 152 ­ 156. Aedeagus of Geostiba nebuligena Gusarov, sp. n. (paratypes from Highways 441 &amp; 71, Tennessee (152, 154 ­ 156) and Appalachian National Scenic Trail W of Clingmans Dome (153), Tennessee / North Carolina). 152 – median lobe, ventral view; 153 ­ 154 – apex of median lobe, ventral view; 155 – median lobe, lateral view; 156 – apex of median lobe, lateral view. Scale bar 0.2 mm (152, 155), 0.1 mm (153 ­ 154, 156)." httpUri="https://zenodo.org/record/155723/files/figure.png" pageId="15" pageNumber="16">Figs. 155­156</figureCitation>
), in ventral view the apex outline strongly doubly emarginate (
<figureCitation id="4BC24485FF91392C4830F888FB2CF894" box="[1045,1220,1844,1870]" captionStart="FIGURES 152 ­ 156" captionStartId="41.[264,383,1456,1480]" captionTargetBox="[369,1217,460,1420]" captionTargetId="figure@41.[361,1225,449,1430]" captionTargetPageId="41" captionText="FIGURES 152 ­ 156. Aedeagus of Geostiba nebuligena Gusarov, sp. n. (paratypes from Highways 441 &amp; 71, Tennessee (152, 154 ­ 156) and Appalachian National Scenic Trail W of Clingmans Dome (153), Tennessee / North Carolina). 152 – median lobe, ventral view; 153 ­ 154 – apex of median lobe, ventral view; 155 – median lobe, lateral view; 156 – apex of median lobe, lateral view. Scale bar 0.2 mm (152, 155), 0.1 mm (153 ­ 154, 156)." httpUri="https://zenodo.org/record/155723/files/figure.png" pageId="15" pageNumber="16">Figs. 152­154</figureCitation>
). Body
</keyLead>
</paragraph>
<paragraph id="D3465800FF8E39334D1DFEA0FAC3FD97" blockId="16.[264,1325,284,1870]" pageId="16" pageNumber="17">
length
<quantity id="1401F5E5FF8E39334DACFEA0FDFCFEEC" box="[393,532,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="16" pageNumber="17" unit="mm" value="1.75" valueMax="1.9" valueMin="1.6">1.6­1.9 mm</quantity>
, pronotal width
<quantity id="1401F5E5FF8E39334EF1FEA0FC95FEEC" box="[724,893,284,310]" metricMagnitude="-4" metricUnit="m" metricValue="3.85" metricValueMax="4.0" metricValueMin="3.7" pageId="16" pageNumber="17" unit="mm" value="0.385" valueMax="0.4" valueMin="0.37">0.37­0.40 mm</quantity>
. Known from many localities in the Great Smoky Mountains massif (
<figureCitation id="4BC24485FF8E39334E99FEF8FCC5FE84" box="[700,813,324,350]" captionStart="FIGURE 338" captionStartId="79.[264,367,1838,1862]" captionTargetBox="[346,1237,315,1825]" captionTargetId="figure@79.[344,1238,312,1827]" captionTargetPageId="79" captionText="FIGURE 338. Geographical distribution of twelve species of Geostiba in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155750/files/figure.png" pageId="16" pageNumber="17">Figs. 338</figureCitation>
,
<figureCitation id="4BC24485FF8E39334F1DFEF8FC8DFE84" box="[824,869,324,350]" captionStart="FIGURE 340" captionStartId="82.[264,368,1747,1771]" captionTargetBox="[322,1272,508,1726]" captionTargetId="figure@82.[319,1275,508,1728]" captionTargetPageId="82" captionText="FIGURE 340. Detailed geographical distribution of Geostiba bicarinata Lohse &amp; Smetana (2), G. graveyardensis Pace (3), G. nimbicola Lohse &amp; Smetana (4), G. pluvigena Gusarov, sp. n. (5), G. nebuligena Gusarov, sp. n. (6), G. crepusculigena Gusarov, sp. n. (7), G. balsamensis Pace (8), G. nubigena Lohse &amp; Smetana (9), G. coeligena Gusarov, sp. n. (10) in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155752/files/figure.png" pageId="16" pageNumber="17">340</figureCitation>
) .......................................................... ...................................................................... 6.
<taxonomicName id="14F92383FF8E39334F7BFED0FB35FE5C" authority="Gusarov" authorityName="Gusarov" box="[862,1245,364,390]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="nebuligena" status="sp. nov." subGenus="Sibiota">
<emphasis id="E18D8412FF8E39334F7BFED0FC92FE5F" box="[862,890,364,389]" italics="true" pageId="16" pageNumber="17">G.</emphasis>
(
<emphasis id="E18D8412FF8E39334FAEFED0FC36FE5F" box="[907,990,364,389]" italics="true" pageId="16" pageNumber="17">Sibiota</emphasis>
)
<emphasis id="E18D8412FF8E39334FCAFED0FB99FE5F" box="[1007,1137,364,389]" italics="true" pageId="16" pageNumber="17">nebuligena</emphasis>
Gusarov
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,
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<emphasis id="E18D8412FF8E393348CCFED7FAC3FE5F" bold="true" box="[1257,1323,363,389]" pageId="16" pageNumber="17">sp. n.</emphasis>
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– Apex of median lobe in lateral view with a weak ventral denticle (
<figureCitation id="4BC24485FF8E39334871FE28FB16FE74" box="[1108,1278,404,430]" captionStart="FIGURES 126 ­ 133" captionStartId="36.[264,382,1701,1725]" captionTargetBox="[308,1279,632,1666]" captionTargetId="figure@36.[294,1294,622,1676]" captionTargetPageId="36" captionText="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 – median lobe, ventral view; 127, 131, 133 – apex of median lobe, ventral view; 128 – median lobe, lateral view; 129 – apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." httpUri="https://zenodo.org/record/155720/files/figure.png" pageId="16" pageNumber="17">Figs. 128­129</figureCitation>
), in ventral view the apex outline weakly doubly emarginate (
<figureCitation id="4BC24485FF8E39334FD2FE00FB77FE0C" box="[1015,1183,444,470]" captionStart="FIGURES 126 ­ 133" captionStartId="36.[264,382,1701,1725]" captionTargetBox="[308,1279,632,1666]" captionTargetId="figure@36.[294,1294,622,1676]" captionTargetPageId="36" captionText="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 – median lobe, ventral view; 127, 131, 133 – apex of median lobe, ventral view; 128 – median lobe, lateral view; 129 – apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." httpUri="https://zenodo.org/record/155720/files/figure.png" pageId="16" pageNumber="17">Figs. 130­131</figureCitation>
) or convex (
<figureCitation id="4BC24485FF8E39334D64FE58FD89FE24" box="[321,609,484,510]" captionStart="FIGURES 126 ­ 133" captionStartId="36.[264,382,1701,1725]" captionTargetBox="[308,1279,632,1666]" captionTargetId="figure@36.[294,1294,622,1676]" captionTargetPageId="36" captionText="FIGURES 126 ­ 133. Aedeagus of Geostiba pluvigena Gusarov, sp. n. (paratypes from the junction of Blue Ridge Parkway and Balsam Mountain Road (126 ­ 129, 132 ­ 133) and Waterrock Knob (130 ­ 131), North Carolina). 126, 130, 132 – median lobe, ventral view; 127, 131, 133 – apex of median lobe, ventral view; 128 – median lobe, lateral view; 129 – apex of median lobe, lateral view. Scale bar 0.2 mm (126, 128, 130, 132), 0.1 mm (127, 129, 131, 133)." httpUri="https://zenodo.org/record/155720/files/figure.png" pageId="16" pageNumber="17">Figs. 126­127, 132­133</figureCitation>
). Body length
<quantity id="1401F5E5FF8E39334F06FE58FC5AFE24" box="[803,946,484,510]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="16" pageNumber="17" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.9­2.2 mm</quantity>
, pronotal width
<quantity id="1401F5E5FF8E39334859FE58FAC1FE24" box="[1148,1321,484,510]" metricMagnitude="-4" metricUnit="m" metricValue="3.95" metricValueMax="4.3" metricValueMin="3.5999999999999996" pageId="16" pageNumber="17" unit="mm" value="0.395" valueMax="0.43" valueMin="0.36">0.36­0.43 mm</quantity>
. Known from the Plott Balsams – Cataloochee Divide massif in the Southern Appalachians (
<figureCitation id="4BC24485FF8E39334DB6FD88FDECFD94" box="[403,516,564,590]" captionStart="FIGURE 338" captionStartId="79.[264,367,1838,1862]" captionTargetBox="[346,1237,315,1825]" captionTargetId="figure@79.[344,1238,312,1827]" captionTargetPageId="79" captionText="FIGURE 338. Geographical distribution of twelve species of Geostiba in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155750/files/figure.png" pageId="16" pageNumber="17">Figs. 338</figureCitation>
,
<figureCitation id="4BC24485FF8E39334E35FD88FDD5FD94" box="[528,573,564,590]" captionStart="FIGURE 340" captionStartId="82.[264,368,1747,1771]" captionTargetBox="[322,1272,508,1726]" captionTargetId="figure@82.[319,1275,508,1728]" captionTargetPageId="82" captionText="FIGURE 340. Detailed geographical distribution of Geostiba bicarinata Lohse &amp; Smetana (2), G. graveyardensis Pace (3), G. nimbicola Lohse &amp; Smetana (4), G. pluvigena Gusarov, sp. n. (5), G. nebuligena Gusarov, sp. n. (6), G. crepusculigena Gusarov, sp. n. (7), G. balsamensis Pace (8), G. nubigena Lohse &amp; Smetana (9), G. coeligena Gusarov, sp. n. (10) in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155752/files/figure.png" pageId="16" pageNumber="17">340</figureCitation>
) .................................. 5.
<taxonomicName id="14F92383FF8E39334F49FD88FB35FD94" authority="Gusarov" authorityName="Gusarov" box="[876,1245,564,590]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pluvigena" status="sp. nov." subGenus="Sibiota">
<emphasis id="E18D8412FF8E39334F49FD88FC61FD97" box="[876,905,564,589]" italics="true" pageId="16" pageNumber="17">G.</emphasis>
(
<emphasis id="E18D8412FF8E39334FBFFD88FC05FD97" box="[922,1005,564,589]" italics="true" pageId="16" pageNumber="17">Sibiota</emphasis>
)
<emphasis id="E18D8412FF8E39334FDBFD88FB99FD97" box="[1022,1137,564,589]" italics="true" pageId="16" pageNumber="17">pluvigena</emphasis>
Gusarov
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,
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<emphasis id="E18D8412FF8E393348CCFD8FFAC3FD97" bold="true" box="[1257,1323,563,589]" pageId="16" pageNumber="17">sp. n.</emphasis>
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</keyStep>
<keyStep id="680D40A5FF8E39334D2DFDE0FAC5FCCC" pageId="16" pageNumber="17">
<paragraph id="D3465800FF8E39334D2DFDE0FAC3FD1C" blockId="16.[264,1325,284,1870]" pageId="16" pageNumber="17">
<keyLead id="6808FB35FF8E39334D2DFDE0FAC3FD1C" pageId="16" pageNumber="17">
12 Body length
<quantity id="1401F5E5FF8E39334DF6FDE0FD88FDAC" box="[467,608,604,630]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="16" pageNumber="17" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8­2.2 mm</quantity>
, pronotal width
<quantity id="1401F5E5FF8E39334F01FDE0FC26FDAC" box="[804,974,604,630]" metricMagnitude="-4" metricUnit="m" metricValue="3.65" metricValueMax="3.9" metricValueMin="3.4" pageId="16" pageNumber="17" unit="mm" value="0.365" valueMax="0.39" valueMin="0.34">0.34­0.39 mm</quantity>
. Lateral sides of the apex of median lobe with shallow emargination (in ventral view) (
<figureCitation id="4BC24485FF8E39334823FD38FACFFD44" box="[1030,1319,644,670]" captionStart="FIGURES 219 ­ 229" captionStartId="54.[264,383,1720,1744]" captionTargetBox="[333,1253,320,1706]" captionTargetId="figure@54.[323,1267,310,1709]" captionTargetPageId="54" captionText="FIGURES 219 ­ 229. Aedeagus of Geostiba nubigena Lohse &amp; Smetana (Richland Balsam Mountain (219 ­ 222), Blue Ridge Parkway (223), Graveyard Fields (224 ­ 227) and Grassy Ridge Mine Overlook (228 ­ 229), North Carolina). 219, 224, 226, 228 – median lobe, ventral view; 220, 223, 225, 227, 229 – apex of median lobe, ventral view; 221 – median lobe, lateral view; 222 – apex of median lobe, lateral view. Scale bar 0.2 mm (219, 221, 224, 226, 228), 0.1 mm (220, 222 ­ 223, 225, 227, 229)." httpUri="https://zenodo.org/record/155733/files/figure.png" pageId="16" pageNumber="17">Figs. 219­220, 223­229</figureCitation>
,
<figureCitation id="4BC24485FF8E39334D1DFD10FE73FD1C" box="[312,411,684,710]" captionStart="FIGURES 247 ­ 252" captionStartId="57.[264,382,1726,1750]" captionTargetBox="[282,1305,745,1687]" captionTargetId="figure@57.[273,1313,729,1700]" captionTargetPageId="57" captionText="FIGURES 247 ­ 252. Abdominal segment 8 and aedeagus of Geostiba coeligena Gusarov, sp. n. (paratype). 247 – male tergum 8; 248 – male sternum 8; 249 – median lobe, ventral view; 250 – apex of median lobe, ventral view; 251 – median lobe, lateral view; 252 – apex of median lobe, lateral view. Scale bar 0.2 mm (247 ­ 249, 251), 0.1 mm (250, 252)." httpUri="https://zenodo.org/record/155735/files/figure.png" pageId="16" pageNumber="17">249­250</figureCitation>
) ..................................................................................................................... 13
</keyLead>
</paragraph>
<paragraph id="D3465800FF8E39334D2DFD68FAC5FCCC" blockId="16.[264,1325,284,1870]" pageId="16" pageNumber="17">
<keyLead id="6808FB35FF8E39334D2DFD68FAC5FCCC" pageId="16" pageNumber="17">
– Body length
<quantity id="1401F5E5FF8E39334DF6FD68FD88FD34" box="[467,608,724,750]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.6" metricValueMin="1.9" pageId="16" pageNumber="17" unit="mm" value="2.25" valueMax="2.6" valueMin="1.9">1.9­2.6 mm</quantity>
, pronotal width
<quantity id="1401F5E5FF8E39334F01FD68FC26FD34" box="[804,974,724,750]" metricMagnitude="-4" metricUnit="m" metricValue="4.15" metricValueMax="4.4" metricValueMin="3.9" pageId="16" pageNumber="17" unit="mm" value="0.415" valueMax="0.44" valueMin="0.39">0.39­0.44 mm</quantity>
. Lateral sides of the apex of median lobe with deep emargination (in ventral view) (
<figureCitation id="4BC24485FF8E39334F9AFD40FB8EFCCC" box="[959,1126,764,790]" captionStart="FIGURES 267 ­ 270" captionStartId="60.[264,382,969,993]" captionTargetBox="[290,1303,319,959]" captionTargetId="figure@60.[278,1309,310,964]" captionTargetPageId="60" captionText="FIGURES 267 ­ 270. Aedeagus of Geostiba alticola Lohse &amp; Smetana (N slope of Mt. Mitchell, North Carolina). 267 – median lobe, ventral view; 268 – apex of median lobe, ventral view; 269 – median lobe, lateral view; 270 – apex of median lobe, lateral view. Scale bar 0.2 mm (267, 269), 0.1 mm (268, 270)." httpUri="https://zenodo.org/record/155738/files/figure.png" pageId="16" pageNumber="17">Figs. 267­268</figureCitation>
,
<figureCitation id="4BC24485FF8E39334857FD40FB3EFCCC" box="[1138,1238,764,790]" captionStart="FIGURES 289 ­ 293" captionStartId="65.[264,383,1756,1780]" captionTargetBox="[342,1242,643,1724]" captionTargetId="figure@65.[330,1257,633,1732]" captionTargetPageId="65" captionText="FIGURES 289 ­ 293. Aedeagus of Geostiba carteriensis Pace (Big Bald (289) and Roan Road Pass (290 ­ 293), Tennessee / North Carolina). 289 ­ 290 – median lobe, ventral view; 291 – apex of median lobe, ventral view; 292 – median lobe, lateral view; 293 – apex of median lobe, lateral view. Scale bar 0.2 mm (289 ­ 290, 292), 0.1 mm (291, 293)." httpUri="https://zenodo.org/record/155741/files/figure.png" pageId="16" pageNumber="17">289­291</figureCitation>
) ..... 14
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</keyStep>
<keyStep id="680D40A5FF8E39334D2DFC98FAC3FB17" pageId="16" pageNumber="17">
<paragraph id="D3465800FF8E39334D2DFC98FAC3FBDC" blockId="16.[264,1325,284,1870]" pageId="16" pageNumber="17">
<keyLead id="6808FB35FF8E39334D2DFC98FAC3FBDC" pageId="16" pageNumber="17">
13 Median lobe of aedeagus in ventral view more narrow (
<figureCitation id="4BC24485FF8E39334FE1FC98FB36FCE4" box="[964,1246,804,830]" captionStart="FIGURES 219 ­ 229" captionStartId="54.[264,383,1720,1744]" captionTargetBox="[333,1253,320,1706]" captionTargetId="figure@54.[323,1267,310,1709]" captionTargetPageId="54" captionText="FIGURES 219 ­ 229. Aedeagus of Geostiba nubigena Lohse &amp; Smetana (Richland Balsam Mountain (219 ­ 222), Blue Ridge Parkway (223), Graveyard Fields (224 ­ 227) and Grassy Ridge Mine Overlook (228 ­ 229), North Carolina). 219, 224, 226, 228 – median lobe, ventral view; 220, 223, 225, 227, 229 – apex of median lobe, ventral view; 221 – median lobe, lateral view; 222 – apex of median lobe, lateral view. Scale bar 0.2 mm (219, 221, 224, 226, 228), 0.1 mm (220, 222 ­ 223, 225, 227, 229)." httpUri="https://zenodo.org/record/155733/files/figure.png" pageId="16" pageNumber="17">Figs. 219­220, 223­229</figureCitation>
). Distal diverticula of internal sac long (
<figureCitation id="4BC24485FF8E39334ECDFCF0FC7BFCBC" box="[744,915,844,870]" captionStart="FIGURES 230 ­ 246" captionStartId="55.[264,382,1503,1527]" captionTargetBox="[316,1271,321,1479]" captionTargetId="figure@55.[305,1282,310,1487]" captionTargetPageId="55" captionText="FIGURES 230 ­ 246. Genitalia of Geostiba nubigena Lohse &amp; Smetana (Richland Balsam Mountain (230 ­ 231, 234 ­ 235, 237 ­ 238, 242, 246) and Blue Ridge Parkway (232 ­ 233, 236, 239 ­ 241, 243 ­ 245), North Carolina). 230 – details of internal sac retracted into median lobe, ventral view; 231 – copulatory piece of internal sac, dorsal view; 232 – medial lamellae of internal sac, ventral view; 233 – distal diverticula of everted internal sac, ventral view; 234 – everted internal sac, ventral view; 235 ­ 236 – spermatheca; 237 ­ 238 – details of internal sac retracted into median lobe, lateral view; 239 – partially everted internal sac, lateral view; 240 – details of internal sac inside partially everted internal sac, lateral view; 241 – everted internal sac, lateral view; 242 – completely everted internal sac, lateral view; 243 – copulatory piece of internal sac, lateral view; 244 – medial lamella of internal sac, lateral view; 245 – right distal diverticulum of everted internal sac, lateral view; 246 – apex of left paramere, side facing median lobe. Scale bar 0.2 mm (230, 234, 237 ­ 242), 0.1 mm (231 ­ 233, 235 ­ 236, 243 ­ 246)." httpUri="https://zenodo.org/record/155734/files/figure.png" pageId="16" pageNumber="17">Figs. 233­234</figureCitation>
), proximal diverticula small (
<figureCitation id="4BC24485FF8E393348DEFCF0FE81FC54" captionStart="FIGURES 230 ­ 246" captionStartId="55.[264,382,1503,1527]" captionTargetBox="[316,1271,321,1479]" captionTargetId="figure@55.[305,1282,310,1487]" captionTargetPageId="55" captionText="FIGURES 230 ­ 246. Genitalia of Geostiba nubigena Lohse &amp; Smetana (Richland Balsam Mountain (230 ­ 231, 234 ­ 235, 237 ­ 238, 242, 246) and Blue Ridge Parkway (232 ­ 233, 236, 239 ­ 241, 243 ­ 245), North Carolina). 230 – details of internal sac retracted into median lobe, ventral view; 231 – copulatory piece of internal sac, dorsal view; 232 – medial lamellae of internal sac, ventral view; 233 – distal diverticula of everted internal sac, ventral view; 234 – everted internal sac, ventral view; 235 ­ 236 – spermatheca; 237 ­ 238 – details of internal sac retracted into median lobe, lateral view; 239 – partially everted internal sac, lateral view; 240 – details of internal sac inside partially everted internal sac, lateral view; 241 – everted internal sac, lateral view; 242 – completely everted internal sac, lateral view; 243 – copulatory piece of internal sac, lateral view; 244 – medial lamella of internal sac, lateral view; 245 – right distal diverticulum of everted internal sac, lateral view; 246 – apex of left paramere, side facing median lobe. Scale bar 0.2 mm (230, 234, 237 ­ 242), 0.1 mm (231 ­ 233, 235 ­ 236, 243 ­ 246)." httpUri="https://zenodo.org/record/155734/files/figure.png" pageId="16" pageNumber="17">Fig. 234</figureCitation>
). Body length
<quantity id="1401F5E5FF8E39334E39FCC8FD4FFC54" box="[540,679,884,910]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="16" pageNumber="17" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8­2.2 mm</quantity>
, pronotal width
<quantity id="1401F5E5FF8E39334F42FCC8FBF9FC54" box="[871,1041,884,910]" metricMagnitude="-4" metricUnit="m" metricValue="3.65" metricValueMax="3.9" metricValueMin="3.4" pageId="16" pageNumber="17" unit="mm" value="0.365" valueMax="0.39" valueMin="0.34">0.34­0.39 mm</quantity>
. Known from the Great Balsam Mountains – Pisgah Ridge massif and from Standing Indian in the Blue Ridge in the Southern Appalachians (
<figureCitation id="4BC24485FF8E39334E84FC78FCF9FC04" box="[673,785,964,990]" captionStart="FIGURE 338" captionStartId="79.[264,367,1838,1862]" captionTargetBox="[346,1237,315,1825]" captionTargetId="figure@79.[344,1238,312,1827]" captionTargetPageId="79" captionText="FIGURE 338. Geographical distribution of twelve species of Geostiba in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155750/files/figure.png" pageId="16" pageNumber="17">Figs. 338</figureCitation>
,
<figureCitation id="4BC24485FF8E39334F3BFC78FCA4FC04" box="[798,844,964,990]" captionStart="FIGURE 340" captionStartId="82.[264,368,1747,1771]" captionTargetBox="[322,1272,508,1726]" captionTargetId="figure@82.[319,1275,508,1728]" captionTargetPageId="82" captionText="FIGURE 340. Detailed geographical distribution of Geostiba bicarinata Lohse &amp; Smetana (2), G. graveyardensis Pace (3), G. nimbicola Lohse &amp; Smetana (4), G. pluvigena Gusarov, sp. n. (5), G. nebuligena Gusarov, sp. n. (6), G. crepusculigena Gusarov, sp. n. (7), G. balsamensis Pace (8), G. nubigena Lohse &amp; Smetana (9), G. coeligena Gusarov, sp. n. (10) in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155752/files/figure.png" pageId="16" pageNumber="17">340</figureCitation>
) .............................................................. .................................................................... 9.
<taxonomicName id="14F92383FF8E39334F76FC50FAC3FBDC" authority="Lohse &amp; Smetana" authorityName="Lohse &amp; Smetana" box="[851,1323,1004,1030]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="nubigena" subGenus="Sibiota">
<emphasis id="E18D8412FF8E39334F76FC50FC87FBDF" box="[851,879,1004,1029]" italics="true" pageId="16" pageNumber="17">G.</emphasis>
(
<emphasis id="E18D8412FF8E39334FA5FC50FC3BFBDF" box="[896,979,1004,1029]" italics="true" pageId="16" pageNumber="17">Sibiota</emphasis>
)
<emphasis id="E18D8412FF8E39334FC1FC50FBB9FBDF" box="[996,1105,1004,1029]" italics="true" pageId="16" pageNumber="17">nubigena</emphasis>
Lohse &amp; Smetana
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="D3465800FF8E39334D2DFBA8FAC3FB17" blockId="16.[264,1325,284,1870]" pageId="16" pageNumber="17">
<keyLead id="6808FB35FF8E39334D2DFBA8FAC3FB17" pageId="16" pageNumber="17">
– Median lobe of aedeagus in ventral view wider (
<figureCitation id="4BC24485FF8E39334F52FBA8FBCAFBF4" box="[887,1058,1044,1070]" captionStart="FIGURES 247 ­ 252" captionStartId="57.[264,382,1726,1750]" captionTargetBox="[282,1305,745,1687]" captionTargetId="figure@57.[273,1313,729,1700]" captionTargetPageId="57" captionText="FIGURES 247 ­ 252. Abdominal segment 8 and aedeagus of Geostiba coeligena Gusarov, sp. n. (paratype). 247 – male tergum 8; 248 – male sternum 8; 249 – median lobe, ventral view; 250 – apex of median lobe, ventral view; 251 – median lobe, lateral view; 252 – apex of median lobe, lateral view. Scale bar 0.2 mm (247 ­ 249, 251), 0.1 mm (250, 252)." httpUri="https://zenodo.org/record/155735/files/figure.png" pageId="16" pageNumber="17">Figs. 249­250</figureCitation>
). Distal diverticula of internal sac short and small (
<figureCitation id="4BC24485FF8E39334EBFFB80FCA7FB8C" box="[666,847,1084,1110]" captionStart="FIGURES 253 ­ 261" captionStartId="58.[264,383,1544,1568]" captionTargetBox="[361,1227,879,1516]" captionTargetId="figure@58.[355,1231,873,1521]" captionTargetPageId="58" captionText="FIGURES 253 ­ 261. Aedeagus of Geostiba coeligena Gusarov, sp. n. (253, 256 ­ 258, 260 ­ 261 – paratype, 254 ­ 255, 259 ­ holotype). 253 – everted internal sac, ventral view; 254 – copulatory piece of internal sac, dorsal view; 255 – medial lamellae of internal sac, ventral view; 256 – distal diverticula of everted internal sac, ventral view; 257 – left proximal diverticulum of everted internal sac, ventral view; 258 – everted internal sac, lateral view; 259 – copulatory piece of internal sac, lateral view; 260 – medial lamella of internal sac, lateral view; 261 – apex of left paramere, side facing median lobe. Scale bar 0.2 mm (253, 258), 0.1 mm (254 ­ 257, 259 ­ 261)." httpUri="https://zenodo.org/record/155736/files/figure.png" pageId="16" pageNumber="17">Figs. 253, 256</figureCitation>
), proximal diverticula very large (
<figureCitation id="4BC24485FF8E393348D5FB80FE32FBA4" captionStart="FIGURES 253 ­ 261" captionStartId="58.[264,383,1544,1568]" captionTargetBox="[361,1227,879,1516]" captionTargetId="figure@58.[355,1231,873,1521]" captionTargetPageId="58" captionText="FIGURES 253 ­ 261. Aedeagus of Geostiba coeligena Gusarov, sp. n. (253, 256 ­ 258, 260 ­ 261 – paratype, 254 ­ 255, 259 ­ holotype). 253 – everted internal sac, ventral view; 254 – copulatory piece of internal sac, dorsal view; 255 – medial lamellae of internal sac, ventral view; 256 – distal diverticula of everted internal sac, ventral view; 257 – left proximal diverticulum of everted internal sac, ventral view; 258 – everted internal sac, lateral view; 259 – copulatory piece of internal sac, lateral view; 260 – medial lamella of internal sac, lateral view; 261 – apex of left paramere, side facing median lobe. Scale bar 0.2 mm (253, 258), 0.1 mm (254 ­ 257, 259 ­ 261)." httpUri="https://zenodo.org/record/155736/files/figure.png" pageId="16" pageNumber="17">Figs. 253, 257­258</figureCitation>
). Body length
<quantity id="1401F5E5FF8E39334EABFBD8FCF1FBA4" box="[654,793,1124,1150]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="16" pageNumber="17" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.8­1.9 mm</quantity>
, pronotal width
<quantity id="1401F5E5FF8E39334FFCFBD8FBAEFBA4" box="[985,1094,1124,1150]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="16" pageNumber="17" unit="mm" value="0.34">0.34 mm</quantity>
. Known only from the Great Smoky Mountains west of Clingmans Dome (
<figureCitation id="4BC24485FF8E39334FE2FB30FBD0FB7C" box="[967,1080,1164,1190]" captionStart="FIGURE 338" captionStartId="79.[264,367,1838,1862]" captionTargetBox="[346,1237,315,1825]" captionTargetId="figure@79.[344,1238,312,1827]" captionTargetPageId="79" captionText="FIGURE 338. Geographical distribution of twelve species of Geostiba in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155750/files/figure.png" pageId="16" pageNumber="17">Figs. 338</figureCitation>
,
<figureCitation id="4BC24485FF8E39334861FB30FB9AFB7C" box="[1092,1138,1164,1190]" captionStart="FIGURE 340" captionStartId="82.[264,368,1747,1771]" captionTargetBox="[322,1272,508,1726]" captionTargetId="figure@82.[319,1275,508,1728]" captionTargetPageId="82" captionText="FIGURE 340. Detailed geographical distribution of Geostiba bicarinata Lohse &amp; Smetana (2), G. graveyardensis Pace (3), G. nimbicola Lohse &amp; Smetana (4), G. pluvigena Gusarov, sp. n. (5), G. nebuligena Gusarov, sp. n. (6), G. crepusculigena Gusarov, sp. n. (7), G. balsamensis Pace (8), G. nubigena Lohse &amp; Smetana (9), G. coeligena Gusarov, sp. n. (10) in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri="https://zenodo.org/record/155752/files/figure.png" pageId="16" pageNumber="17">340</figureCitation>
) ...................... ..................................................................... 10.
<taxonomicName id="14F92383FF8E39334F4BFB08FB35FB14" authority="Gusarov" authorityName="Gusarov" box="[878,1245,1204,1230]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="coeligena" status="sp. nov." subGenus="Sibiota">
<emphasis id="E18D8412FF8E39334F4BFB08FC62FB17" box="[878,906,1204,1229]" italics="true" pageId="16" pageNumber="17">G.</emphasis>
(
<emphasis id="E18D8412FF8E39334FBEFB08FC06FB17" box="[923,1006,1204,1229]" italics="true" pageId="16" pageNumber="17">Sibiota</emphasis>
)
<emphasis id="E18D8412FF8E39334FDAFB08FB99FB17" box="[1023,1137,1204,1229]" italics="true" pageId="16" pageNumber="17">coeligena</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="E18D8412FF8E393348CCFB0FFAC3FB17" bold="true" box="[1257,1323,1203,1229]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="FABE3969FF8E393348CCFB0FFAC3FB17" box="[1257,1323,1203,1229]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="680D40A5FF8E39334D2DFB60FAC2F984" pageId="16" pageNumber="17">
<paragraph id="D3465800FF8E39334D2DFB60FAC3FA4C" blockId="16.[264,1325,284,1870]" pageId="16" pageNumber="17">
<keyLead id="6808FB35FF8E39334D2DFB60FAC3FA4C" pageId="16" pageNumber="17">
14 Median lobe of aedeagus longer (
<figureCitation id="4BC24485FF8E39334EE8FB60FCDFFB2C" box="[717,823,1244,1270]" captionStart="FIGURES 267 ­ 270" captionStartId="60.[264,382,969,993]" captionTargetBox="[290,1303,319,959]" captionTargetId="figure@60.[278,1309,310,964]" captionTargetPageId="60" captionText="FIGURES 267 ­ 270. Aedeagus of Geostiba alticola Lohse &amp; Smetana (N slope of Mt. Mitchell, North Carolina). 267 – median lobe, ventral view; 268 – apex of median lobe, ventral view; 269 – median lobe, lateral view; 270 – apex of median lobe, lateral view. Scale bar 0.2 mm (267, 269), 0.1 mm (268, 270)." httpUri="https://zenodo.org/record/155738/files/figure.png" pageId="16" pageNumber="17">Fig. 269</figureCitation>
), in lateral view more narrow (
<figureCitation id="4BC24485FF8E39334894FB60FE80FAC4" captionStart="FIGURES 267 ­ 270" captionStartId="60.[264,382,969,993]" captionTargetBox="[290,1303,319,959]" captionTargetId="figure@60.[278,1309,310,964]" captionTargetPageId="60" captionText="FIGURES 267 ­ 270. Aedeagus of Geostiba alticola Lohse &amp; Smetana (N slope of Mt. Mitchell, North Carolina). 267 – median lobe, ventral view; 268 – apex of median lobe, ventral view; 269 – median lobe, lateral view; 270 – apex of median lobe, lateral view. Scale bar 0.2 mm (267, 269), 0.1 mm (268, 270)." httpUri="https://zenodo.org/record/155738/files/figure.png" pageId="16" pageNumber="17">Figs. 269­ 270</figureCitation>
), in ventral view the lateral sides of the apex less convex (
<figureCitation id="4BC24485FF8E39334806FAB8FB25FAC4" box="[1059,1229,1284,1310]" captionStart="FIGURES 267 ­ 270" captionStartId="60.[264,382,969,993]" captionTargetBox="[290,1303,319,959]" captionTargetId="figure@60.[278,1309,310,964]" captionTargetPageId="60" captionText="FIGURES 267 ­ 270. Aedeagus of Geostiba alticola Lohse &amp; Smetana (N slope of Mt. Mitchell, North Carolina). 267 – median lobe, ventral view; 268 – apex of median lobe, ventral view; 269 – median lobe, lateral view; 270 – apex of median lobe, lateral view. Scale bar 0.2 mm (267, 269), 0.1 mm (268, 270)." httpUri="https://zenodo.org/record/155738/files/figure.png" pageId="16" pageNumber="17">Figs. 267­268</figureCitation>
). Body length
<quantity id="1401F5E5FF8E39334DADFA90FDFBFA9C" box="[392,531,1324,1350]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.4" metricValueMin="1.9" pageId="16" pageNumber="17" unit="mm" value="2.15" valueMax="2.4" valueMin="1.9">1.9­2.4 mm</quantity>
, pronotal width
<quantity id="1401F5E5FF8E39334EF5FA90FC92FA9C" box="[720,890,1324,1350]" metricMagnitude="-4" metricUnit="m" metricValue="4.15" metricValueMax="4.4" metricValueMin="3.9" pageId="16" pageNumber="17" unit="mm" value="0.415" valueMax="0.44" valueMin="0.39">0.39­0.44 mm</quantity>
. Known only from the Great Craggy Mountains and the Black Mountains in the Southern Appalachians (
<figureCitation id="4BC24485FF8E39334871FAE8FB11FAB4" box="[1108,1273,1364,1390]" captionStart-0="FIGURE 338" captionStart-1="FIGURE 339" captionStartId-0="79.[264,367,1838,1862]" captionStartId-1="80.[264,368,1616,1640]" captionTargetBox-0="[346,1237,315,1825]" captionTargetBox-1="[345,1240,318,1600]" captionTargetId-0="figure@79.[344,1238,312,1827]" captionTargetId-1="figure@80.[344,1241,318,1600]" captionTargetPageId-0="79" captionTargetPageId-1="80" captionText-0="FIGURE 338. Geographical distribution of twelve species of Geostiba in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." captionText-1="FIGURE 339. Detailed geographical distribution of Geostiba alticola Lohse &amp; Smetana (11) and G. carteriensis Pace (12) in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri-0="https://zenodo.org/record/155750/files/figure.png" httpUri-1="https://zenodo.org/record/155751/files/figure.png" pageId="16" pageNumber="17">Figs. 338­339</figureCitation>
) .... ..................................................................... 11.
<taxonomicName id="14F92383FF8E39334F43FAC0FAC3FA4C" authority="Lohse &amp; Smetana" authorityName="Lohse &amp; Smetana" box="[870,1323,1404,1430]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alticola" subGenus="Sibiota">
<emphasis id="E18D8412FF8E39334F43FAC0FC6BFA4F" box="[870,899,1404,1429]" italics="true" pageId="16" pageNumber="17">G.</emphasis>
(
<emphasis id="E18D8412FF8E39334FB1FAC0FC0FFA4F" box="[916,999,1404,1429]" italics="true" pageId="16" pageNumber="17">Sibiota</emphasis>
)
<emphasis id="E18D8412FF8E39334FDDFAC0FBBAFA4F" box="[1016,1106,1404,1429]" italics="true" pageId="16" pageNumber="17">alticola</emphasis>
Lohse &amp; Smetana
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="D3465800FF8E39334D2DFA18FAC2F984" blockId="16.[264,1325,284,1870]" pageId="16" pageNumber="17">
<keyLead id="6808FB35FF8E39334D2DFA18FAC2F984" pageId="16" pageNumber="17">
– Median lobe of aedeagus shorter (
<figureCitation id="4BC24485FF8E39334EE3FA18FCC4FA64" box="[710,812,1444,1470]" captionStart="FIGURES 289 ­ 293" captionStartId="65.[264,383,1756,1780]" captionTargetBox="[342,1242,643,1724]" captionTargetId="figure@65.[330,1257,633,1732]" captionTargetPageId="65" captionText="FIGURES 289 ­ 293. Aedeagus of Geostiba carteriensis Pace (Big Bald (289) and Roan Road Pass (290 ­ 293), Tennessee / North Carolina). 289 ­ 290 – median lobe, ventral view; 291 – apex of median lobe, ventral view; 292 – median lobe, lateral view; 293 – apex of median lobe, lateral view. Scale bar 0.2 mm (289 ­ 290, 292), 0.1 mm (291, 293)." httpUri="https://zenodo.org/record/155741/files/figure.png" pageId="16" pageNumber="17">Fig. 292</figureCitation>
), in lateral view broader (
<figureCitation id="4BC24485FF8E3933487DFA18FAE8FA64" box="[1112,1280,1444,1470]" captionStart="FIGURES 289 ­ 293" captionStartId="65.[264,383,1756,1780]" captionTargetBox="[342,1242,643,1724]" captionTargetId="figure@65.[330,1257,633,1732]" captionTargetPageId="65" captionText="FIGURES 289 ­ 293. Aedeagus of Geostiba carteriensis Pace (Big Bald (289) and Roan Road Pass (290 ­ 293), Tennessee / North Carolina). 289 ­ 290 – median lobe, ventral view; 291 – apex of median lobe, ventral view; 292 – median lobe, lateral view; 293 – apex of median lobe, lateral view. Scale bar 0.2 mm (289 ­ 290, 292), 0.1 mm (291, 293)." httpUri="https://zenodo.org/record/155741/files/figure.png" pageId="16" pageNumber="17">Figs. 292­293</figureCitation>
), in ventral view the lateral sides of the apex more convex (
<figureCitation id="4BC24485FF8E39334FF5FA70FB93FA3C" box="[976,1147,1484,1510]" captionStart="FIGURES 289 ­ 293" captionStartId="65.[264,383,1756,1780]" captionTargetBox="[342,1242,643,1724]" captionTargetId="figure@65.[330,1257,633,1732]" captionTargetPageId="65" captionText="FIGURES 289 ­ 293. Aedeagus of Geostiba carteriensis Pace (Big Bald (289) and Roan Road Pass (290 ­ 293), Tennessee / North Carolina). 289 ­ 290 – median lobe, ventral view; 291 – apex of median lobe, ventral view; 292 – median lobe, lateral view; 293 – apex of median lobe, lateral view. Scale bar 0.2 mm (289 ­ 290, 292), 0.1 mm (291, 293)." httpUri="https://zenodo.org/record/155741/files/figure.png" pageId="16" pageNumber="17">Figs. 289­291</figureCitation>
). Body length
<quantity id="1401F5E5FF8E39334D1DFA48FE2CF9D4" box="[312,452,1524,1550]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.6" metricValueMin="2.2" pageId="16" pageNumber="17" unit="mm" value="2.4" valueMax="2.6" valueMin="2.2">2.2­2.6 mm</quantity>
, pronotal width
<quantity id="1401F5E5FF8E39334EA6FA48FCC5F9D4" box="[643,813,1524,1550]" metricMagnitude="-4" metricUnit="m" metricValue="4.2" metricValueMax="4.4" metricValueMin="4.0" pageId="16" pageNumber="17" unit="mm" value="0.42" valueMax="0.44" valueMin="0.4">0.40­0.44 mm</quantity>
. Known only from Roan Mountain and the Big Bald (Bald Mountains) in the Southern Appalachians (
<figureCitation id="4BC24485FF8E39334FCEF9A0FB78F9EC" box="[1003,1168,1564,1590]" captionStart-0="FIGURE 338" captionStart-1="FIGURE 339" captionStartId-0="79.[264,367,1838,1862]" captionStartId-1="80.[264,368,1616,1640]" captionTargetBox-0="[346,1237,315,1825]" captionTargetBox-1="[345,1240,318,1600]" captionTargetId-0="figure@79.[344,1238,312,1827]" captionTargetId-1="figure@80.[344,1241,318,1600]" captionTargetPageId-0="79" captionTargetPageId-1="80" captionText-0="FIGURE 338. Geographical distribution of twelve species of Geostiba in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." captionText-1="FIGURE 339. Detailed geographical distribution of Geostiba alticola Lohse &amp; Smetana (11) and G. carteriensis Pace (12) in the Southern Appalachians. Isohypses 1200 m and 1500 m are shown. Areas above 1500 m are solid black." httpUri-0="https://zenodo.org/record/155750/files/figure.png" httpUri-1="https://zenodo.org/record/155751/files/figure.png" pageId="16" pageNumber="17">Figs. 338­339</figureCitation>
) .................. ................................................................................... 12.
<taxonomicName id="14F92383FF8E39334FF5F9F8FAC2F984" authority="Pace" authorityName="Pace" box="[976,1322,1604,1630]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="carteriensis" subGenus="Sibiota">
<emphasis id="E18D8412FF8E39334FF5F9F8FC05F987" box="[976,1005,1604,1629]" italics="true" pageId="16" pageNumber="17">G.</emphasis>
(
<emphasis id="E18D8412FF8E39334FDBF9F8FBB9F987" box="[1022,1105,1604,1629]" italics="true" pageId="16" pageNumber="17">Sibiota</emphasis>
)
<emphasis id="E18D8412FF8E39334847F9F8FB04F987" box="[1122,1260,1604,1629]" italics="true" pageId="16" pageNumber="17">carteriensis</emphasis>
Pace
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="D3465800FF8E39334D2DF9D0FAC2F8FC" blockId="16.[264,1325,284,1870]" pageId="16" pageNumber="17">
<keyLead id="6808FB35FF8E39334D2DF9D0FAC2F8FC" pageId="16" pageNumber="17">
<keyStep id="680D40A5FF8E39334D2DF9D0FAC2F8FC" pageId="16" pageNumber="17">
15 Elytra (measured from humeral level) shorter than pronotum. Wings reduced, shorter than elytra. In males pronotum with strong, wide and deep impression along midline. Median lobe of aedeagus with wider apex (in ventral view) (
<figureCitation id="4BC24485FF8E3933480AF900FAF4F90C" box="[1071,1308,1724,1750]" captionStart="FIGURES 310 ­ 315" captionStartId="69.[264,383,1724,1748]" captionTargetBox="[290,1301,520,1688]" captionTargetId="figure@69.[277,1311,510,1700]" captionTargetPageId="69" captionText="FIGURES 310 ­ 315. Aedeagus of Geostiba impressula (Casey) (lectotype of Sipalia fossata Casey (310 ­ 311) and a specimen from Triangle Lake, Oregon (312 ­ 315 )). 310, 312 – median lobe, ventral view; 311, 315 – apex of median lobe, ventral view; 313 – median lobe, lateral view; 314 – apex of median lobe, lateral view. Scale bar 0.4 mm (310, 312 ­ 313), 0.2 mm (311, 314 ­ 315)." httpUri="https://zenodo.org/record/155744/files/figure.png" pageId="16" pageNumber="17">Figs. 310­312, 315</figureCitation>
). Body length
<quantity id="1401F5E5FF8E39334DEEF958FDBCF924" box="[459,596,1764,1790]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="16" pageNumber="17" unit="mm" value="2.45" valueMax="2.7" valueMin="2.2">2.2­2.7 mm</quantity>
, pronotal width
<quantity id="1401F5E5FF8E39334F29F958FC5AF924" box="[780,946,1764,1790]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="5.0" metricValueMin="3.7" pageId="16" pageNumber="17" unit="mm" value="0.435" valueMax="0.5" valueMin="0.37">0.37­0.50 mm</quantity>
. Known from Oregon (
<figureCitation id="4BC24485FF8E393348E5F958FACBF924" box="[1216,1315,1764,1790]" captionStart="FIGURE 336" captionStartId="76.[264,368,1562,1586]" captionTargetBox="[315,1271,753,1543]" captionTargetId="figure@76.[311,1275,751,1546]" captionTargetPageId="76" captionText="FIGURE 336. Geographical distribution of Nearctic species of Geostiba. The ellipse indicates the area where eleven Southern Appalachian species are distributed." httpUri="https://zenodo.org/record/155748/files/figure.png" pageId="16" pageNumber="17">Fig. 336</figureCitation>
) ................................................................................ 13.
<taxonomicName id="14F92383FF8E39334F92F8B0FAC2F8FC" authority="Casey" authorityName="Casey" box="[951,1322,1804,1830]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="impressula" subGenus="Sibiota">
<emphasis id="E18D8412FF8E39334F92F8B0FC3CF8FF" box="[951,980,1804,1829]" italics="true" pageId="16" pageNumber="17">G.</emphasis>
(
<emphasis id="E18D8412FF8E39334FC0F8B0FBD0F8FF" box="[997,1080,1804,1829]" italics="true" pageId="16" pageNumber="17">Sibiota</emphasis>
)
<emphasis id="E18D8412FF8E3933486CF8B0FB20F8FF" box="[1097,1224,1804,1829]" italics="true" pageId="16" pageNumber="17">impressula</emphasis>
(Casey)
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="D3465800FF8E39324D2DF888FAC3FE5F" blockId="16.[264,1325,284,1870]" lastBlockId="17.[312,1325,284,390]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<keyLead id="6808FB35FF8E39334D2DF888FAC3F894" box="[264,1323,1844,1870]" pageId="16" pageNumber="17">– Elytra as long as pronotum. Wings fully developed. In males pronotum without deep</keyLead>
<subSection id="BA7641EBFF8F39324D1DFEA0FAC3FE5F" pageId="17" pageNumber="18" type="multiple">
impression along midline. Median lobe of aedeagus with more narrow apex (in ventral view (
<figureCitation id="4BC24485FF8F39324D93FEF8FD8BFE84" box="[438,611,324,350]" captionStart="FIGURES 321 ­ 326" captionStartId="72.[264,383,1696,1720]" captionTargetBox="[286,1297,660,1656]" captionTargetId="figure@72.[275,1311,649,1670]" captionTargetPageId="72" captionText="FIGURES 321 ­ 326. Abdominal segment 8 and aedeagus of Geostiba silvigena Gusarov, sp. n. (holotype). 321 – male tergum 8; 322 – male sternum 8; 323 – median lobe, ventral view; 324 – apex of median lobe, ventral view; 325 – median lobe, lateral view; 326 – apex of median lobe, lateral view. Scale bar 0.4 mm (321 ­ 323, 325), 0.2 mm (324, 326)." httpUri="https://zenodo.org/record/155746/files/figure.png" pageId="17" pageNumber="18">Figs. 323­324</figureCitation>
). Body length
<quantity id="1401F5E5FF8F39324F06FEF8FC6CFE84" box="[803,900,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="17" pageNumber="18" unit="mm" value="2.4">2.4 mm</quantity>
, pronotal width
<quantity id="1401F5E5FF8F39324868FEF8FB56FE84" box="[1101,1214,324,350]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="17" pageNumber="18" unit="mm" value="0.41">0.41 mm</quantity>
. Known from California (
<figureCitation id="4BC24485FF8F39324DDAFED0FD8BFE5C" box="[511,611,364,390]" captionStart="FIGURE 336" captionStartId="76.[264,368,1562,1586]" captionTargetBox="[315,1271,753,1543]" captionTargetId="figure@76.[311,1275,751,1546]" captionTargetPageId="76" captionText="FIGURE 336. Geographical distribution of Nearctic species of Geostiba. The ellipse indicates the area where eleven Southern Appalachian species are distributed." httpUri="https://zenodo.org/record/155748/files/figure.png" pageId="17" pageNumber="18">Fig. 336</figureCitation>
)............................ 14.
<taxonomicName id="14F92383FF8F39324F53FED0FB35FE5C" authority="Gusarov" authorityName="Gusarov" box="[886,1245,364,390]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="silvigena" status="sp. nov." subGenus="Sibiota">
<emphasis id="E18D8412FF8F39324F53FED0FC7AFE5F" box="[886,914,364,389]" italics="true" pageId="17" pageNumber="18">G.</emphasis>
(
<emphasis id="E18D8412FF8F39324F86FED0FC1EFE5F" box="[931,1014,364,389]" italics="true" pageId="17" pageNumber="18">Sibiota</emphasis>
)
<emphasis id="E18D8412FF8F39324822FED0FB99FE5F" box="[1031,1137,364,389]" italics="true" pageId="17" pageNumber="18">silvigena</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="FABE3969FF8F393248CCFED7FAC3FE5F" box="[1257,1323,363,389]" pageId="17" pageNumber="18" rank="species">
<emphasis id="E18D8412FF8F393248CCFED7FAC3FE5F" bold="true" box="[1257,1323,363,389]" pageId="17" pageNumber="18">sp. n.</emphasis>
</taxonomicNameLabel>
</subSection>
</paragraph>
</treatment>
</document>
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