treatments-xml/data/5B/12/D4/5B12D409FFE2797A157E25D4D0C0FA25.xml

<document id="9440A52508CC1A893C95B4406FC1C7C6" ID-DOI="10.5281/zenodo.170616" ID-GBIF-Dataset="b795d14b-c66d-420b-ad1d-a5c58d09a987" ID-ISSN="1175­5326" ID-Zenodo-Dep="170616" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459714149645" checkinUser="plazi" docAuthor="Lemaitre, Rafael &amp; Watabe, Hajime" docDate="2005" docId="5B12D409FFE2797A157E25D4D0C0FA25" docLanguage="en" docName="zt00819.pdf" docOrigin="Zootaxa 819" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Pagurus ikedai Lemaitre &amp; Watabe, 2005, n. sp." docType="treatment" docVersion="9" lastPageNumber="10" masterDocId="A72BAC71FFE0797314762472D144FFB3" masterDocTitle="Pagurus ikedai (Crustacea: Anomura: Paguridae), a new hermit crab species of the bernhardus group from Japanese waters" masterLastPageNumber="12" masterPageNumber="1" pageNumber="3" updateTime="1698207525134" updateUser="plazi">
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<mods:title id="8CA19BC01FF6E53D7D4A1A0980E35BA0">Pagurus ikedai (Crustacea: Anomura: Paguridae), a new hermit crab species of the bernhardus group from Japanese waters</mods:title>
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<mods:namePart id="4DECF4D82DCD21E2DA12FD026D1466AE">Lemaitre, Rafael</mods:namePart>
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<mods:namePart id="9C89BFB8869434191CD7D43FBBB4EFC1">Watabe, Hajime</mods:namePart>
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<mods:date id="63073467917F020F8612E1BCBD2D30B8">2005</mods:date>
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<treatment id="5B12D409FFE2797A157E25D4D0C0FA25" ID-DOI="http://doi.org/10.5281/zenodo.6265885" ID-GBIF-Taxon="119334172" ID-Zenodo-Dep="6265885" LSID="urn:lsid:plazi:treatment:5B12D409FFE2797A157E25D4D0C0FA25" httpUri="http://treatment.plazi.org/id/5B12D409FFE2797A157E25D4D0C0FA25" lastPageId="9" lastPageNumber="10" pageId="2" pageNumber="3">
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<paragraph id="D304651FFFE27971157E25D4D3CFFE73" blockId="2.[264,676,353,448]" box="[264,651,422,448]" pageId="2" pageNumber="3">
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<emphasis id="E1CFB90DFFE27971157E25D4D3CFFE73" bold="true" box="[264,651,422,448]" pageId="2" pageNumber="3">
<taxonomicName id="14BB1E9CFFE27971157E25D4D0FBFE73" ID-CoL="9c477f13-e7a8-4a95-9674-76479b672bec" box="[264,447,422,448]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ikedai" status="sp. nov.">
<emphasis id="E1CFB90DFFE27971157E25D4D0FBFE73" bold="true" box="[264,447,422,448]" italics="true" pageId="2" pageNumber="3">Pagurus ikedai</emphasis>
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<taxonomicNameLabel id="FAFC0476FFE2797115B125D4D34DFE73" box="[455,521,422,448]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="4B80799AFFE27971166E25D4D3C6FE73" box="[536,642,422,448]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="3.[264,367,1641,1665]" captionStartId-1="4.[264,367,1585,1609]" captionStartId-2="5.[264,368,1775,1799]" captionStartId-3="7.[264,368,1136,1160]" captionStartId-4="7.[264,368,1821,1845]" captionTargetBox-0="[278,1250,596,1621]" captionTargetBox-1="[302,1280,598,1541]" captionTargetBox-2="[334,1249,437,1750]" captionTargetBox-3="[291,1273,341,1094]" captionTargetBox-4="[531,1056,1318,1797]" captionTargetId-0="figure@3.[264,1323,576,1634]" captionTargetId-1="figure@4.[264,1323,578,1561]" captionTargetId-2="figure@5.[333,1253,432,1751]" captionTargetId-3="figure@7.[264,1323,326,1112]" captionTargetId-4="figure@7.[531,1056,1318,1798]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="7" captionTargetPageId-4="7" captionText-0="FIGURE 1. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30: a, paratype male 7.6 mm (USNM 1069118); b – f, i, j, holotype male 8.2 mm (NSMT­Cr 16121); g, h, paratype male 8.5 mm, sta MM­ 30 (USNM 1069118). a, gill lamella; b, shield and cephalic appendages, dorsal view; c, portion of dorsal surface of palm of right cheliped near base of fixed finger and lateral margin, dorsolateral view; d, anterior lobe and posterior portion of thoracic sternite XII (third pereopods), ventral view; e – h, telson in dorsal (e, g) and posterior (f, h) views; i, j, coxae of fifth pereopods and thoracic sternite XXIV in ventral (i) and posterior (j) views. Scale bars: a = 0.25 mm; b, d, i, j = 1.0 mm; c, e – h = 0.5 mm." captionText-1="FIGURE 2. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30, paratype male 7.6 mm (USNM 1069118). Left mouthparts, internal view: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped; g, coxa and ischium of same. Scale bars: a – e, g = 0.5 mm; f = 1.0 mm." captionText-2="FIGURE 3. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30, holotype male 8.2 mm (NSMT­Cr 16121). Right (a – c) and left (d – f) chelipeds: a, d, dorsal view; b, e, mesial view; c, f, ventral view. Scale bars: a – f = 4.0 mm." captionText-3="FIGURE 4. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30, holotype male 8.2 mm (NSMT­Cr 16121): a, second right pereopod, lateral view; b, dactyl of same, mesial view; c, third right pereopod, lateral view; d, dactyl of same, mesial view; e, fourth pereopod, lateral view; f, fifth pereopod, lateral view. Scale bars: a – d = 2.0 mm; e, f = 1.0 mm." captionText-4="FIGURE 5. Pagurus ikedai n. sp., off Jogashima, Japan, live coloration, male 9.8 mm. (Photo: H. Ikeda)." httpUri-0="https://zenodo.org/record/170617/files/figure.png" httpUri-1="https://zenodo.org/record/170618/files/figure.png" httpUri-2="https://zenodo.org/record/170619/files/figure.png" httpUri-3="https://zenodo.org/record/170620/files/figure.png" httpUri-4="https://zenodo.org/record/170621/files/figure.png" pageId="2" pageNumber="3">Figs 1–5</figureCitation>
)
</emphasis>
</heading>
</paragraph>
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material
</emphasis>
. 
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<collectingCountry id="ABAC258FFFE2797115CD2583D343FDB8" box="[443,519,497,523]" name="Japan" pageId="2" pageNumber="3">Japan</collectingCountry>
</emphasis>
. 
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<typeStatus id="0C00DBBDFFE2797116612583D3C6FDB9" box="[535,642,497,522]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
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male 
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, entrance to Tokyo Submarine Canyon, SE of Tsurugi­Zaki, Miura Peninsula, sta MM­30, 
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, 
<geoCoordinate id="B68F03D8FFE2797117A1266BD530FD80" box="[983,1140,537,563]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="9" value="139.74916">139° 44.95'E</geoCoordinate>
, 
<quantity id="1443C8FAFFE2797110F5266BD449FD80" box="[1155,1293,537,563]" metricMagnitude="2" metricUnit="m" metricValue="3.1" metricValueMax="3.3" metricValueMin="2.9" pageId="2" pageNumber="3" unit="m" value="310.0" valueMax="330.0" valueMin="290.0">290–330 m</quantity>
, 
<date id="A70543DFFFE27971116A266BD0E4FDE8" pageId="2" pageNumber="3" value="1994-08-01">1 August 1994</date>
, NSMT­Cr 16121. 
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<typeStatus id="0C00DBBDFFE2797116F62633D3B3FDE9" box="[640,759,577,602]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
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, all from entrance to Tokyo Submarine Canyon: N of Su­no­Saki, Boso Peninsula, sta TT­1, 
<geoCoordinate id="B68F03D8FFE27971175D261BD2F3FD30" box="[811,951,617,643]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="9" value="34.994167">34°59.65’N</geoCoordinate>
, 
<geoCoordinate id="B68F03D8FFE2797117BE261BD51BFD30" box="[968,1119,617,643]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="9" value="139.75833">139°45.50’E</geoCoordinate>
, 
<quantity id="1443C8FAFFE279711006261BD5B9FD30" box="[1136,1277,617,643]" metricMagnitude="2" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="2" pageNumber="3" unit="m" value="315.0" valueMax="350.0" valueMin="280.0">280–350 m</quantity>
, 
<date id="A70543DFFFE27971117B261BD0C0FD18" pageId="2" pageNumber="3" value="1985-05-12">12 May 1985</date>
: 
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6.0– 
<quantity id="1443C8FAFFE27971165526E3D33BFD18" box="[547,639,657,683]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="2" pageNumber="3" unit="mm" value="9.4">9.4 mm</quantity>
, CBM­ZC 8183. SE of Tsurugi­Zaki, Miura Peninsula, sta MS­6, 
<geoCoordinate id="B68F03D8FFE27971152F26CBD0A1FD60" box="[345,485,697,723]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="9" value="35.06333">35°03.80’N</geoCoordinate>
, 
<geoCoordinate id="B68F03D8FFE27971158726CBD3CDFD60" box="[497,649,697,723]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="9" value="139.73967">139°44.38’E</geoCoordinate>
, 
<quantity id="1443C8FAFFE2797116E226CBD25FFD60" box="[660,795,697,723]" metricMagnitude="2" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="2" pageNumber="3" unit="m" value="280.0" valueMax="290.0" valueMin="270.0">270–290 m</quantity>
, 
<date id="A70543DFFFE27971175026CBD2F2FD60" box="[806,950,697,723]" pageId="2" pageNumber="3" value="1992-06-04">4 June 1992</date>
: 
<specimenCount id="C5BDAE96FFE2797117B726CBD55EFD60" box="[961,1050,697,723]" pageId="2" pageNumber="3" type="male">2 males</specimenCount>
8.9, 
<quantity id="1443C8FAFFE27971102226CBD5F4FD60" box="[1108,1200,697,723]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="2" pageNumber="3" unit="mm" value="9.4">9.4 mm</quantity>
, 
<specimenCount id="C5BDAE96FFE2797110CC26CBD46FFD60" box="[1210,1323,697,723]" pageId="2" pageNumber="3" type="female">2 females</specimenCount>
7.5, 
<quantity id="1443C8FAFFE2797115342693D0E2FD48" box="[322,422,737,763]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="2" pageNumber="3" unit="mm" value="8.9">8.9 mm</quantity>
, HSM­Cra 0 134 to 0 137. ESE of Tsurugi­Zaki, sta MU­8, 
<geoCoordinate id="B68F03D8FFE2797110EF2693D462FD48" box="[1177,1318,737,763]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="9" value="35.118332">35°07.10’N</geoCoordinate>
, 
<geoCoordinate id="B68F03D8FFE27971157E277BD0E4FC90" box="[264,416,777,803]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="9" value="139.77884">139°46.73’E</geoCoordinate>
, 
<quantity id="1443C8FAFFE2797115D8277BD373FC90" box="[430,567,777,803]" metricMagnitude="2" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="2" pageNumber="3" unit="m" value="315.0" valueMax="350.0" valueMin="280.0">280–350 m</quantity>
, 
<date id="A70543DFFFE279711633277BD3A0FC90" box="[581,740,777,803]" pageId="2" pageNumber="3" value="1992-07-19">19 July 1992</date>
: 
<specimenCount id="C5BDAE96FFE279711687277BD205FC90" box="[753,833,777,803]" pageId="2" pageNumber="3" type="male">1 male</specimenCount>
<quantity id="1443C8FAFFE27971173C277BD2EDFC90" box="[842,937,777,803]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="2" pageNumber="3" unit="mm" value="8.5">8.5 mm</quantity>
, CBM­ZC 8180. SE of Tsurugi­ Zaki, Miura Peninsula, sta MS­28, 
<geoCoordinate id="B68F03D8FFE2797116CB2743D20EFCF8" box="[701,842,817,843]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="9" value="35.080833">35°04.85’N</geoCoordinate>
, 
<geoCoordinate id="B68F03D8FFE27971172D2743D2B7FCF8" box="[859,1011,817,843]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="9" value="139.74867">139°44.92’E</geoCoordinate>
, 
<quantity id="1443C8FAFFE2797110722743D5D5FCF8" box="[1028,1169,817,843]" metricMagnitude="2" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="2" pageNumber="3" unit="m" value="265.0" valueMax="290.0" valueMin="240.0">240–290 m</quantity>
, 
<date id="A70543DFFFE2797110D42743D003FCC0" pageId="2" pageNumber="3" value="1994-10-27">27 October 1994</date>
: 
<specimenCount id="C5BDAE96FFE279711527272BD0F3FCC0" box="[337,439,857,883]" pageId="2" pageNumber="3" type="female">1 female</specimenCount>
<quantity id="1443C8FAFFE2797115CB272BD35DFCC0" box="[445,537,857,883]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="2" pageNumber="3" unit="mm" value="7.2">7.2 mm</quantity>
, CBM­ZC 8181. SE of Tsurugi­Zaki, Miura Peninsula, sta MM­28, 
<geoCoordinate id="B68F03D8FFE27971157E27F3D0D0FC28" box="[264,404,897,923]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="9" value="35.06333">35°03.80’N</geoCoordinate>
, 
<geoCoordinate id="B68F03D8FFE2797115D727F3D37DFC28" box="[417,569,897,923]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="9" value="139.73967">139°44.38’E</geoCoordinate>
, 
<quantity id="1443C8FAFFE27971163327F3D389FC28" box="[581,717,897,923]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="2" pageNumber="3" unit="m" value="300.0" valueMax="310.0" valueMin="290.0">290–310 m</quantity>
, 
<date id="A70543DFFFE2797116AF27F3D23FFC28" box="[729,891,897,923]" pageId="2" pageNumber="3" value="1994-06-28">28 June 1994</date>
: 
<specimenCount id="C5BDAE96FFE2797117F027F3D2A5FC28" box="[902,993,897,923]" pageId="2" pageNumber="3" type="male">2 males</specimenCount>
8.7, 
<quantity id="1443C8FAFFE27971106A27F3D53EFC28" box="[1052,1146,897,923]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="2" pageNumber="3" unit="mm" value="9.4">9.4 mm</quantity>
, 
<specimenCount id="C5BDAE96FFE2797110F327F3D5B3FC28" box="[1157,1271,897,923]" pageId="2" pageNumber="3" type="female">2 females</specimenCount>
8.5, 9.0 mm, NSMT­Cr 16122; 
<specimenCount id="C5BDAE96FFE27971163027DBD3E4FC70" box="[582,672,937,963]" pageId="2" pageNumber="3" type="male">5 males</specimenCount>
<quantity id="1443C8FAFFE2797116D127DBD273FC70" box="[679,823,937,963]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="8.9" metricValueMin="8.1" pageId="2" pageNumber="3" unit="mm" value="8.5" valueMax="8.9" valueMin="8.1">8.1–8.9 mm</quantity>
, 
<specimenCount id="C5BDAE96FFE27971173527DBD2F0FC70" box="[835,948,937,963]" pageId="2" pageNumber="3" type="female">2 females</specimenCount>
7.2, 
<quantity id="1443C8FAFFE27971179827DBD50EFC70" box="[1006,1098,937,963]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="2" pageNumber="3" unit="mm" value="8.1">8.1 mm</quantity>
, HW. Same station data as 
<typeStatus id="0C00DBBDFFE27971152927A3D08EFC58" box="[351,458,977,1003]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
: 
<specimenCount id="C5BDAE96FFE2797115A227A3D366FC58" box="[468,546,977,1003]" pageId="2" pageNumber="3" type="male">1 male</specimenCount>
<quantity id="1443C8FAFFE27971165F27A3D3C1FC58" box="[553,645,977,1003]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="2" pageNumber="3" unit="mm" value="7.6">7.6 mm</quantity>
, 
<specimenCount id="C5BDAE96FFE2797116E727A3D245FC58" box="[657,769,977,1003]" pageId="2" pageNumber="3" type="female">4 females</specimenCount>
<quantity id="1443C8FAFFE27971177E27A3D2DCFC58" box="[776,920,977,1003]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" metricValueMax="7.6" metricValueMin="7.2" pageId="2" pageNumber="3" unit="mm" value="7.4" valueMax="7.6" valueMin="7.2">7.2–7.6 mm</quantity>
, 
<collectionCode id="B5AAFDDAFFE2797117D527A3D2BCFC58" LSID="urn:lsid:biocol.org:col:34871" box="[931,1016,977,1003]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="3">USNM</collectionCode>
1069118; 
<specimenCount id="C5BDAE96FFE27971100527A3D588FC58" box="[1139,1228,977,1003]" pageId="2" pageNumber="3" type="male">2 males</specimenCount>
9.1, 
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, MNHN­Pg 7263; 
<specimenCount id="C5BDAE96FFE279711661278BD322FBA0" box="[535,614,1017,1043]" pageId="2" pageNumber="3" type="male">1 male</specimenCount>
<quantity id="1443C8FAFFE27971161A278BD38DFBA0" box="[620,713,1017,1043]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="2" pageNumber="3" unit="mm" value="7.4">7.4 mm</quantity>
, 
<specimenCount id="C5BDAE96FFE2797116A2278BD27EFBA0" box="[724,826,1017,1043]" pageId="2" pageNumber="3" type="female">1 female</specimenCount>
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, 
<collectionCode id="B5AAFDDAFFE2797117DE278BD2A7FBA0" LSID="urn:lsid:biocol.org:col:34838" box="[936,995,1017,1043]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="3">SMF</collectionCode>
29993. SE of Tsurugi­Zaki, Miura Peninsula, sta MM­36, 
<geoCoordinate id="B68F03D8FFE2797116182053D3BFFB88" box="[622,763,1057,1083]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="9" value="35.04933">35°02.96’N</geoCoordinate>
, 
<geoCoordinate id="B68F03D8FFE27971177E2053D2E4FB88" box="[776,928,1057,1083]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="9" value="139.74333">139°44.60’E</geoCoordinate>
, 
<quantity id="1443C8FAFFE2797117D82053D573FB88" box="[942,1079,1057,1083]" metricMagnitude="2" metricUnit="m" metricValue="2.9" metricValueMax="3.1" metricValueMin="2.7" pageId="2" pageNumber="3" unit="m" value="290.0" valueMax="310.0" valueMin="270.0">270–310 m</quantity>
, 
<date id="A70543DFFFE2797110352053D44BFB88" box="[1091,1295,1057,1083]" pageId="2" pageNumber="3" value="1995-10-16">16 October 1995</date>
: 
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, CBM­ZC 8182.
</paragraph>
</subSubSection>
<subSubSection id="9BA13694FFE2797B154E2003D2B5FBCD" lastPageId="8" lastPageNumber="9" pageId="2" pageNumber="3" type="description">
<paragraph id="D304651FFFE27971154E2003D3A9FA10" blockId="2.[264,1325,497,1883]" pageId="2" pageNumber="3">
<emphasis id="E1CFB90DFFE27971154E2003D086FB39" box="[312,450,1137,1162]" italics="true" pageId="2" pageNumber="3">Description</emphasis>
. Eleven pairs of biserial gills (
<figureCitation id="4B80799AFFE2797117412003D239FB38" box="[823,893,1137,1163]" captionStart="FIGURE 1" captionStartId="3.[264,367,1641,1665]" captionTargetBox="[278,1250,596,1621]" captionTargetId="figure@3.[264,1323,576,1634]" captionTargetPageId="3" captionText="FIGURE 1. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30: a, paratype male 7.6 mm (USNM 1069118); b – f, i, j, holotype male 8.2 mm (NSMT­Cr 16121); g, h, paratype male 8.5 mm, sta MM­ 30 (USNM 1069118). a, gill lamella; b, shield and cephalic appendages, dorsal view; c, portion of dorsal surface of palm of right cheliped near base of fixed finger and lateral margin, dorsolateral view; d, anterior lobe and posterior portion of thoracic sternite XII (third pereopods), ventral view; e – h, telson in dorsal (e, g) and posterior (f, h) views; i, j, coxae of fifth pereopods and thoracic sternite XXIV in ventral (i) and posterior (j) views. Scale bars: a = 0.25 mm; b, d, i, j = 1.0 mm; c, e – h = 0.5 mm." httpUri="https://zenodo.org/record/170617/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
a): 2 arthrobranchs on each of third maxilliped and first to fourth pereopods, and pleurobranch on each side of somite XIII (above fourth pereopods). Shield (
<figureCitation id="4B80799AFFE2797116EA20B3D39AFB68" box="[668,734,1217,1243]" captionStart="FIGURE 1" captionStartId="3.[264,367,1641,1665]" captionTargetBox="[278,1250,596,1621]" captionTargetId="figure@3.[264,1323,576,1634]" captionTargetPageId="3" captionText="FIGURE 1. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30: a, paratype male 7.6 mm (USNM 1069118); b – f, i, j, holotype male 8.2 mm (NSMT­Cr 16121); g, h, paratype male 8.5 mm, sta MM­ 30 (USNM 1069118). a, gill lamella; b, shield and cephalic appendages, dorsal view; c, portion of dorsal surface of palm of right cheliped near base of fixed finger and lateral margin, dorsolateral view; d, anterior lobe and posterior portion of thoracic sternite XII (third pereopods), ventral view; e – h, telson in dorsal (e, g) and posterior (f, h) views; i, j, coxae of fifth pereopods and thoracic sternite XXIV in ventral (i) and posterior (j) views. Scale bars: a = 0.25 mm; b, d, i, j = 1.0 mm; c, e – h = 0.5 mm." httpUri="https://zenodo.org/record/170617/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
b) about as broad as long; dorsal surface glabrous except for longitudinal row of tufts of short setae on each side; anterior margins between rostrum and lateral projections concave; anterolateral margins sloping; posterior margin truncate. Rostrum triangular, terminating in small spine, reaching to about level of lateral projections; with short setae dorsally. Lateral projections broadly triangular, terminating in small spine often directed anterolaterally.
</paragraph>
<paragraph id="D304651FFFE27971154E21C3D247F9D8" blockId="2.[264,1325,497,1883]" pageId="2" pageNumber="3">Ocular peduncles short, about 0.5 times as long as shield; with few tufts of short setae dorsally. Corneas moderately dilated, width of each about 0.4–0.5 length of ocular peduncle. Ocular acicles narrow, acutely subtriangular, concave dorsally, each with minute to small subterminal spine (often not visible in dorsal view, occasionally absent); separated basally by less than basal width of 1 acicle.</paragraph>
<paragraph id="D304651FFFE27971154E220BD2EDF950" blockId="2.[264,1325,497,1883]" pageId="2" pageNumber="3">Antennular peduncles overreaching corneas by 0.5–0.8 length of ultimate segments. Ultimate segment with scattered short setae dorsally. Penultimate segment with scattered setae. Basal segment with laterodistal angle blunt, setose.</paragraph>
<paragraph id="D304651FFFE27971154E2283D46FF8E8" blockId="2.[264,1325,497,1883]" pageId="2" pageNumber="3">Antennal peduncles overreaching corneas by about 0.5 length of fifth segment; with supernumerary segment. Fifth segment with few setae on lateral and mesial margins. Fourth segment with scattered setae. Third segment with strong spine at ventrodistal angle.</paragraph>
<paragraph id="D304651FFFE37970157E256ED2F4FD95" blockId="3.[264,1323,284,550]" pageId="3" pageNumber="4">Second segment with dorsolateral distal angle prominently produced, reaching nearly to distal margin of fourth segment, and terminating in strong spine (rarely multifid); dorsomesial distal angle with small spine. First segment unarmed or with small spine on lateral surface. Antennal acicles reaching slightly beyond distal margins of corneas, weakly curving outwards (dorsal view), and each terminating in strong spine; mesial margin sparsely setose, lacking spines. Flagellum exceeding outstretched right cheliped; articles naked or with short setae&gt; 0.5 flagellum article in length.</paragraph>
<caption id="87C43597FFE37970157E221BD386F8D3" httpUri="https://zenodo.org/record/170617/files/figure.png" pageId="3" pageNumber="4" targetBox="[278,1250,596,1621]" targetPageId="3">
<paragraph id="D304651FFFE37970157E221BD386F8D3" blockId="3.[264,1323,1641,1888]" pageId="3" pageNumber="4">
<emphasis id="E1CFB90DFFE37970157E221BD0CEF932" bold="true" box="[264,394,1641,1665]" pageId="3" pageNumber="4">FIGURE 1.</emphasis>
<taxonomicName id="14BB1E9CFFE3797015E42218D371F932" box="[402,565,1642,1665]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ikedai" status="sp. nov.">
<emphasis id="E1CFB90DFFE3797015E42218D371F932" box="[402,565,1642,1665]" italics="true" pageId="3" pageNumber="4">Pagurus ikedai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAFC0476FFE37970164A221BD333F933" box="[572,631,1641,1664]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
, Tokyo Submarine Canyon, Japan, sta MM­30: a, paratype male 7.6 mm (USNM 1069118); b–f, i, j, holotype male 8.2 mm (NSMT­Cr 16121); g, h, paratype male 8.5 mm, sta MM­30 (USNM 1069118). a, gill lamella; b, shield and cephalic appendages, dorsal view; c, portion of dorsal surface of palm of right cheliped near base of fixed finger and lateral margin, dorsolateral view; d, anterior lobe and posterior portion of thoracic sternite XII (third pereopods), ventral view; e–h, telson in dorsal (e, g) and posterior (f, h) views; i, j, coxae of fifth pereopods and thoracic sternite XXIV in ventral (i) and posterior (j) views. Scale bars: a = 0.25 mm; b, d, i, j = 1.0 mm; c, e–h = 0.5 mm.
</paragraph>
</caption>
<paragraph id="D304651FFFE47977154E256ED213FD95" blockId="4.[264,1323,284,550]" pageId="4" pageNumber="5">
Mandible (
<figureCitation id="4B80799AFFE4797715CC256ED345FE85" box="[442,513,284,310]" captionStart="FIGURE 2" captionStartId="4.[264,367,1585,1609]" captionTargetBox="[302,1280,598,1541]" captionTargetId="figure@4.[264,1323,578,1561]" captionTargetPageId="4" captionText="FIGURE 2. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30, paratype male 7.6 mm (USNM 1069118). Left mouthparts, internal view: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped; g, coxa and ischium of same. Scale bars: a – e, g = 0.5 mm; f = 1.0 mm." httpUri="https://zenodo.org/record/170618/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
a) with margin of incisor process calcified. Maxillule (
<figureCitation id="4B80799AFFE4797710F8256ED592FE85" box="[1166,1238,284,310]" captionStart="FIGURE 2" captionStartId="4.[264,367,1585,1609]" captionTargetBox="[302,1280,598,1541]" captionTargetId="figure@4.[264,1323,578,1561]" captionTargetPageId="4" captionText="FIGURE 2. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30, paratype male 7.6 mm (USNM 1069118). Left mouthparts, internal view: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped; g, coxa and ischium of same. Scale bars: a – e, g = 0.5 mm; f = 1.0 mm." httpUri="https://zenodo.org/record/170618/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
b) with external lobe of endopod moderately developed, slightly recurved; internal lobe with long terminal bristle. Maxilla (
<figureCitation id="4B80799AFFE479771644251ED333FE35" box="[562,631,364,390]" captionStart="FIGURE 2" captionStartId="4.[264,367,1585,1609]" captionTargetBox="[302,1280,598,1541]" captionTargetId="figure@4.[264,1323,578,1561]" captionTargetPageId="4" captionText="FIGURE 2. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30, paratype male 7.6 mm (USNM 1069118). Left mouthparts, internal view: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped; g, coxa and ischium of same. Scale bars: a – e, g = 0.5 mm; f = 1.0 mm." httpUri="https://zenodo.org/record/170618/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
c) with endopod reaching beyond distal margin of scaphognathite. First maxilliped (
<figureCitation id="4B80799AFFE47977164D25E6D3C1FE1D" box="[571,645,404,430]" captionStart="FIGURE 2" captionStartId="4.[264,367,1585,1609]" captionTargetBox="[302,1280,598,1541]" captionTargetId="figure@4.[264,1323,578,1561]" captionTargetPageId="4" captionText="FIGURE 2. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30, paratype male 7.6 mm (USNM 1069118). Left mouthparts, internal view: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped; g, coxa and ischium of same. Scale bars: a – e, g = 0.5 mm; f = 1.0 mm." httpUri="https://zenodo.org/record/170618/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
d) with endopod reaching distal margin of distal endite. Second maxilliped (
<figureCitation id="4B80799AFFE47977158125CED37BFE65" box="[503,575,444,470]" captionStart="FIGURE 2" captionStartId="4.[264,367,1585,1609]" captionTargetBox="[302,1280,598,1541]" captionTargetId="figure@4.[264,1323,578,1561]" captionTargetPageId="4" captionText="FIGURE 2. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30, paratype male 7.6 mm (USNM 1069118). Left mouthparts, internal view: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped; g, coxa and ischium of same. Scale bars: a – e, g = 0.5 mm; f = 1.0 mm." httpUri="https://zenodo.org/record/170618/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
e) without distinguishing characters. Third maxilliped (
<figureCitation id="4B80799AFFE4797710A425CED45EFE65" box="[1234,1306,444,470]" captionStart="FIGURE 2" captionStartId="4.[264,367,1585,1609]" captionTargetBox="[302,1280,598,1541]" captionTargetId="figure@4.[264,1323,578,1561]" captionTargetPageId="4" captionText="FIGURE 2. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30, paratype male 7.6 mm (USNM 1069118). Left mouthparts, internal view: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped; g, coxa and ischium of same. Scale bars: a – e, g = 0.5 mm; f = 1.0 mm." httpUri="https://zenodo.org/record/170618/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
f, g) with crista dentata on ischium consisting of about 18 corneous­tipped teeth and 1 accessory tooth; carpus unarmed on dorsodistal margin.
</paragraph>
<caption id="87C43597FFE47977157E2243D34BF903" httpUri="https://zenodo.org/record/170618/files/figure.png" pageId="4" pageNumber="5" targetBox="[302,1280,598,1541]" targetPageId="4">
<paragraph id="D304651FFFE47977157E2243D34BF903" blockId="4.[264,1323,1585,1712]" pageId="4" pageNumber="5">
<emphasis id="E1CFB90DFFE47977157E2243D0CDF9FA" bold="true" box="[264,393,1585,1609]" pageId="4" pageNumber="5">FIGURE 2.</emphasis>
<taxonomicName id="14BB1E9CFFE4797715E62240D376F9FA" box="[400,562,1586,1609]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ikedai" status="sp. nov.">
<emphasis id="E1CFB90DFFE4797715E62240D376F9FA" box="[400,562,1586,1609]" italics="true" pageId="4" pageNumber="5">Pagurus ikedai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAFC0476FFE47977164E2243D336F9FB" box="[568,626,1585,1608]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
, Tokyo Submarine Canyon, Japan, sta MM­30, paratype male 7.6 mm (USNM 1069118). Left mouthparts, internal view: a, mandible; b, maxillule; c, maxilla; d, first maxilliped; e, second maxilliped; f, third maxilliped; g, coxa and ischium of same. Scale bars: a–e, g = 0.5 mm; f = 1.0 mm.
</paragraph>
</caption>
<paragraph id="D304651FFFE47975154E2296D253FBE5" blockId="4.[264,1324,1764,1870]" lastBlockId="6.[264,1324,284,1830]" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
Chelipeds massive and stout, unequal, left reaching to about proximal margin of dactyl of right. Right cheliped (
<figureCitation id="4B80799AFFE479771644237ED33FF895" box="[562,635,1804,1830]" captionStart="FIGURE 1" captionStartId="3.[264,367,1641,1665]" captionTargetBox="[278,1250,596,1621]" captionTargetId="figure@3.[264,1323,576,1634]" captionTargetPageId="3" captionText="FIGURE 1. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30: a, paratype male 7.6 mm (USNM 1069118); b – f, i, j, holotype male 8.2 mm (NSMT­Cr 16121); g, h, paratype male 8.5 mm, sta MM­ 30 (USNM 1069118). a, gill lamella; b, shield and cephalic appendages, dorsal view; c, portion of dorsal surface of palm of right cheliped near base of fixed finger and lateral margin, dorsolateral view; d, anterior lobe and posterior portion of thoracic sternite XII (third pereopods), ventral view; e – h, telson in dorsal (e, g) and posterior (f, h) views; i, j, coxae of fifth pereopods and thoracic sternite XXIV in ventral (i) and posterior (j) views. Scale bars: a = 0.25 mm; b, d, i, j = 1.0 mm; c, e – h = 0.5 mm." httpUri="https://zenodo.org/record/170617/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
c, 3a–c) sparsely setose. Chela operculate, dorsoventrally compressed (more so laterally); dorsolateral and dorsomesial margins of palm and fingers sharply defined, finely serrate; fingers each terminating in inwardly directed corneous claw crossed when closed; with tufts of setae inwardly directed near base of larger teeth of cutting edges both dorsally and ventrally. Dactyl about as long as mesial margin of palm; cutting edge with row of usually 3 large calcareous teeth, usually with few smaller ones interspersed, and row of small corneous teeth distally; dorsal surface with numerous short, corneous bristles; ventral surface with numerous small tubercles, and elevated longitudinal median ridge with low corneous­tipped spines. Fixed finger dorsal surface with numerous short, corneous bristles; cutting edge with 3 large calcareous teeth proximally, and 3–5 small calcareous teeth distally; dorsal and ventral surfaces each with longitudinal median ridge with low corneous­tipped spines; ventral surface with numerous small tubercles. Palm dorsal surface with numerous calcareous spines or tubercles diminishing in size towards dorsomesial and dorsolateral margins and terminating in short corneous bristles (
<figureCitation id="4B80799AFFE67975156726F6D012FD2D" box="[273,342,644,670]" captionStart="FIGURE 1" captionStartId="3.[264,367,1641,1665]" captionTargetBox="[278,1250,596,1621]" captionTargetId="figure@3.[264,1323,576,1634]" captionTargetPageId="3" captionText="FIGURE 1. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30: a, paratype male 7.6 mm (USNM 1069118); b – f, i, j, holotype male 8.2 mm (NSMT­Cr 16121); g, h, paratype male 8.5 mm, sta MM­ 30 (USNM 1069118). a, gill lamella; b, shield and cephalic appendages, dorsal view; c, portion of dorsal surface of palm of right cheliped near base of fixed finger and lateral margin, dorsolateral view; d, anterior lobe and posterior portion of thoracic sternite XII (third pereopods), ventral view; e – h, telson in dorsal (e, g) and posterior (f, h) views; i, j, coxae of fifth pereopods and thoracic sternite XXIV in ventral (i) and posterior (j) views. Scale bars: a = 0.25 mm; b, d, i, j = 1.0 mm; c, e – h = 0.5 mm." httpUri="https://zenodo.org/record/170617/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
c), median region with several distinctly larger spines; dorsomesial margin somewhat flared; ventral surface with numerous small tubercles and 2 submedian rows of larger tubercles or spines. Carpus about 0.9 as long as broad, and about 0.8 as long as chela; dorsal surface with numerous spines or tubercles terminating in short corneous bristles, dorsodistal margin spinose; dorsomesial margin well delimited by row of serrate, small or strong, corneous­tipped spines; dorsolateral margin sharply defined, serrate, flaring vetromesially on proximal half; ventral surface with numerous small, low tubercles. Merus usually with 1 or 2 distal spines on dorsal margin, dorsodistal margin spinose; dorsal surface with scattered low tubercles bearing short bristle­like setae; anterior half of ventral surface concave, with numerous small low tubercles, and strongly sloping towards midportion of segment. Ischium with scattered setae; ventromesial margin with minute blunt spines. Coxa with ventromesial row of setae.
</paragraph>
<caption id="87C43597FFE57976157E229DD385F8FF" httpUri="https://zenodo.org/record/170619/files/figure.png" pageId="5" pageNumber="6" targetBox="[334,1249,437,1750]" targetPageId="5">
<paragraph id="D304651FFFE57976157E229DD385F8FF" blockId="5.[264,1323,1775,1868]" pageId="5" pageNumber="6">
<emphasis id="E1CFB90DFFE57976157E229DD0CDF8B4" bold="true" box="[264,393,1775,1799]" pageId="5" pageNumber="6">FIGURE 3.</emphasis>
<taxonomicName id="14BB1E9CFFE5797615E62282D375F8B4" box="[400,561,1776,1799]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="ikedai" status="sp. nov.">
<emphasis id="E1CFB90DFFE5797615E62282D375F8B4" box="[400,561,1776,1799]" italics="true" pageId="5" pageNumber="6">Pagurus ikedai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAFC0476FFE5797616412282D336F8B4" box="[567,626,1776,1799]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
, Tokyo Submarine Canyon, Japan, sta MM­30, holotype male 8.2 mm (NSMT­Cr 16121). Right (a–c) and left (d–f) chelipeds: a, d, dorsal view; b, e, mesial view; c, f, ventral view. Scale bars: a–f = 4.0 mm.
</paragraph>
</caption>
<paragraph id="D304651FFFE67975154E2016D2E7FB45" blockId="6.[264,1324,284,1830]" pageId="6" pageNumber="7">
Left cheliped (
<figureCitation id="4B80799AFFE6797515932016D36EFBCD" box="[485,554,1124,1150]" captionStart="FIGURE 3" captionStartId="5.[264,368,1775,1799]" captionTargetBox="[334,1249,437,1750]" captionTargetId="figure@5.[333,1253,432,1751]" captionTargetPageId="5" captionText="FIGURE 3. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30, holotype male 8.2 mm (NSMT­Cr 16121). Right (a – c) and left (d – f) chelipeds: a, d, dorsal view; b, e, mesial view; c, f, ventral view. Scale bars: a – f = 4.0 mm." httpUri="https://zenodo.org/record/170619/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
d–f) similar to right in shape and armature, differing primarily in size. Dactyl about as long as mesial margin of palm. Cutting edge of dactyl with row of small, closely set corneous spines; cutting edge of fixed finger with row of small, blunt calcareous teeth interspersed with small corneous spines.
</paragraph>
<paragraph id="D304651FFFE67975154E2176D57DF895" blockId="6.[264,1324,284,1830]" pageId="6" pageNumber="7">
Ambulatory legs (
<figureCitation id="4B80799AFFE67975167D2176D314FAAD" box="[523,592,1284,1310]" captionStart="FIGURE 4" captionStartId="7.[264,368,1136,1160]" captionTargetBox="[291,1273,341,1094]" captionTargetId="figure@7.[264,1323,326,1112]" captionTargetPageId="7" captionText="FIGURE 4. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30, holotype male 8.2 mm (NSMT­Cr 16121): a, second right pereopod, lateral view; b, dactyl of same, mesial view; c, third right pereopod, lateral view; d, dactyl of same, mesial view; e, fourth pereopod, lateral view; f, fifth pereopod, lateral view. Scale bars: a – d = 2.0 mm; e, f = 1.0 mm." httpUri="https://zenodo.org/record/170620/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
a–d) similar right from left except for longer meri and stronger spination on dorsal margins of carpi and propodi of right; reaching to, or slightly beyond, distal margins of fingers of right cheliped when extended; meri to dactyls with few setae mostly along dorsal and ventral margins. Dactyl about 1.5 times as long as propodus, terminating in sharp corneous claw; lateral and mesial faces each with longitudinal groove, more marked on mesial surface; ventral margin with 14–22 corneous spines, ventromesial margin with 4–18 corneous spines; dorsal margin with 18–26 corneous spines, dorsomesial margin with 9–20 corneous spines. Propodus with row of corneous­tipped calcareous spines on dorsal margin, and scattered small, low tubercles on dorsolateral surface; ventral margin unarmed except for few setae. Carpus with row of strong, corneous­tipped calcareous spines on dorsal margin, and scattered small, low tubercles on dorsolateral surface. Merus with row of spines on ventral margin. Ischium with small, blunt spines on ventromesial margin. Coxa with ventromesial row of setae. Anterior lobe of thoracic sternite XII (third pereopods; 
<figureCitation id="4B80799AFFE679751671237ED30CF895" box="[519,584,1804,1830]" captionStart="FIGURE 1" captionStartId="3.[264,367,1641,1665]" captionTargetBox="[278,1250,596,1621]" captionTargetId="figure@3.[264,1323,576,1634]" captionTargetPageId="3" captionText="FIGURE 1. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30: a, paratype male 7.6 mm (USNM 1069118); b – f, i, j, holotype male 8.2 mm (NSMT­Cr 16121); g, h, paratype male 8.5 mm, sta MM­ 30 (USNM 1069118). a, gill lamella; b, shield and cephalic appendages, dorsal view; c, portion of dorsal surface of palm of right cheliped near base of fixed finger and lateral margin, dorsolateral view; d, anterior lobe and posterior portion of thoracic sternite XII (third pereopods), ventral view; e – h, telson in dorsal (e, g) and posterior (f, h) views; i, j, coxae of fifth pereopods and thoracic sternite XXIV in ventral (i) and posterior (j) views. Scale bars: a = 0.25 mm; b, d, i, j = 1.0 mm; c, e – h = 0.5 mm." httpUri="https://zenodo.org/record/170617/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
d) subrectangular, with long setae distally.
</paragraph>
<caption id="87C43597FFE77974157E2002D2AEFB43" httpUri="https://zenodo.org/record/170620/files/figure.png" pageId="7" pageNumber="8" targetBox="[291,1273,341,1094]" targetPageId="7">
<paragraph id="D304651FFFE77974157E2002D2AEFB43" blockId="7.[264,1323,1136,1264]" pageId="7" pageNumber="8">
<emphasis id="E1CFB90DFFE77974157E2002D0CDFB3B" bold="true" box="[264,393,1136,1160]" pageId="7" pageNumber="8">FIGURE 4.</emphasis>
<taxonomicName id="14BB1E9CFFE7797415E62003D375FB3B" box="[400,561,1137,1160]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ikedai" status="sp. nov.">
<emphasis id="E1CFB90DFFE7797415E62003D375FB3B" box="[400,561,1137,1160]" italics="true" pageId="7" pageNumber="8">Pagurus ikedai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAFC0476FFE7797416412003D336FB3B" box="[567,626,1137,1160]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
, Tokyo Submarine Canyon, Japan, sta MM­30, holotype male 8.2 mm (NSMT­Cr 16121): a, second right pereopod, lateral view; b, dactyl of same, mesial view; c, third right pereopod, lateral view; d, dactyl of same, mesial view; e, fourth pereopod, lateral view; f, fifth pereopod, lateral view. Scale bars: a–d = 2.0 mm; e, f = 1.0 mm.
</paragraph>
</caption>
<caption id="87C43597FFE77974157E236FD017F8E4" httpUri="https://zenodo.org/record/170621/files/figure.png" pageId="7" pageNumber="8" targetBox="[531,1056,1318,1797]" targetPageId="7">
<paragraph id="D304651FFFE77974157E236FD017F8E4" blockId="7.[264,1323,1821,1879]" pageId="7" pageNumber="8">
<emphasis id="E1CFB90DFFE77974157E236FD0CFF886" bold="true" box="[264,395,1821,1845]" pageId="7" pageNumber="8">FIGURE 5.</emphasis>
<taxonomicName id="14BB1E9CFFE7797415E5236CD372F886" box="[403,566,1822,1845]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="ikedai" status="sp. nov.">
<emphasis id="E1CFB90DFFE7797415E5236CD372F886" box="[403,566,1822,1845]" italics="true" pageId="7" pageNumber="8">Pagurus ikedai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAFC0476FFE77974164B236FD33DF887" box="[573,633,1821,1844]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
, off Jogashima, Japan, live coloration, male 9.8 mm. (Photo: H. Ikeda).
</paragraph>
</caption>
<paragraph id="D304651FFFE8797B154E256ED349FE35" blockId="8.[264,1324,284,1870]" pageId="8" pageNumber="9">
Fourth pereopod (
<figureCitation id="4B80799AFFE8797B167F256ED30BFE85" box="[521,591,284,310]" captionStart="FIGURE 4" captionStartId="7.[264,368,1136,1160]" captionTargetBox="[291,1273,341,1094]" captionTargetId="figure@7.[264,1323,326,1112]" captionTargetPageId="7" captionText="FIGURE 4. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30, holotype male 8.2 mm (NSMT­Cr 16121): a, second right pereopod, lateral view; b, dactyl of same, mesial view; c, third right pereopod, lateral view; d, dactyl of same, mesial view; e, fourth pereopod, lateral view; f, fifth pereopod, lateral view. Scale bars: a – d = 2.0 mm; e, f = 1.0 mm." httpUri="https://zenodo.org/record/170620/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
e) semichelate. Dactyl terminating in corneous claw; with ventrolateral row of small, closely set corneous spines. Propodal rasp with 3 or 4 rows of ovate corneous scales.
</paragraph>
<paragraph id="D304651FFFE8797B154E25E6D298FE65" blockId="8.[264,1324,284,1870]" pageId="8" pageNumber="9">
Fifth pereopod (
<figureCitation id="4B80799AFFE8797B158F25E6D305FE1D" box="[505,577,404,430]" captionStart="FIGURE 4" captionStartId="7.[264,368,1136,1160]" captionTargetBox="[291,1273,341,1094]" captionTargetId="figure@7.[264,1323,326,1112]" captionTargetPageId="7" captionText="FIGURE 4. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30, holotype male 8.2 mm (NSMT­Cr 16121): a, second right pereopod, lateral view; b, dactyl of same, mesial view; c, third right pereopod, lateral view; d, dactyl of same, mesial view; e, fourth pereopod, lateral view; f, fifth pereopod, lateral view. Scale bars: a – d = 2.0 mm; e, f = 1.0 mm." httpUri="https://zenodo.org/record/170620/files/figure.png" pageId="8" pageNumber="9">Fig. 4</figureCitation>
f) chelate. Dactyl with ventrolateral row of small ovate scales. Propodal rasp extending proximally for about 0.3 of segment.
</paragraph>
<paragraph id="D304651FFFE8797B154E2596D5B4FD2D" blockId="8.[264,1324,284,1870]" pageId="8" pageNumber="9">
Uropods and telson asymmetrical. Telson (
<figureCitation id="4B80799AFFE8797B17592596D234FE4D" box="[815,880,484,510]" captionStart="FIGURE 1" captionStartId="3.[264,367,1641,1665]" captionTargetBox="[278,1250,596,1621]" captionTargetId="figure@3.[264,1323,576,1634]" captionTargetPageId="3" captionText="FIGURE 1. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30: a, paratype male 7.6 mm (USNM 1069118); b – f, i, j, holotype male 8.2 mm (NSMT­Cr 16121); g, h, paratype male 8.5 mm, sta MM­ 30 (USNM 1069118). a, gill lamella; b, shield and cephalic appendages, dorsal view; c, portion of dorsal surface of palm of right cheliped near base of fixed finger and lateral margin, dorsolateral view; d, anterior lobe and posterior portion of thoracic sternite XII (third pereopods), ventral view; e – h, telson in dorsal (e, g) and posterior (f, h) views; i, j, coxae of fifth pereopods and thoracic sternite XXIV in ventral (i) and posterior (j) views. Scale bars: a = 0.25 mm; b, d, i, j = 1.0 mm; c, e – h = 0.5 mm." httpUri="https://zenodo.org/record/170617/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
e–h) with transverse suture, and scattered setae on dorsal surface; anterior lobes with long setae laterally; posterior lobes separated by U­shaped cleft; terminal margins oblique, setose, armed with irregular rows of corneous marginal and submarginal spines (sometimes worn out as in 
<typeStatus id="0C00DBBDFFE8797B1033262ED5E8FDC5" box="[1093,1196,604,630]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
right side; 
<figureCitation id="4B80799AFFE8797B157E26F6D00EFD2D" box="[264,330,644,670]" captionStart="FIGURE 1" captionStartId="3.[264,367,1641,1665]" captionTargetBox="[278,1250,596,1621]" captionTargetId="figure@3.[264,1323,576,1634]" captionTargetPageId="3" captionText="FIGURE 1. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30: a, paratype male 7.6 mm (USNM 1069118); b – f, i, j, holotype male 8.2 mm (NSMT­Cr 16121); g, h, paratype male 8.5 mm, sta MM­ 30 (USNM 1069118). a, gill lamella; b, shield and cephalic appendages, dorsal view; c, portion of dorsal surface of palm of right cheliped near base of fixed finger and lateral margin, dorsolateral view; d, anterior lobe and posterior portion of thoracic sternite XII (third pereopods), ventral view; e – h, telson in dorsal (e, g) and posterior (f, h) views; i, j, coxae of fifth pereopods and thoracic sternite XXIV in ventral (i) and posterior (j) views. Scale bars: a = 0.25 mm; b, d, i, j = 1.0 mm; c, e – h = 0.5 mm." httpUri="https://zenodo.org/record/170617/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
e, f), stronger spines near outer angles totally or partially hidden in dorsal view.
</paragraph>
<paragraph id="D304651FFFE8797B154E26DED03EFC3D" blockId="8.[264,1324,284,1870]" pageId="8" pageNumber="9">
Female with paired gonopores; with unpaired pleopods 2–5, pleopod 5 not ovigerous. Male gonopores (
<figureCitation id="4B80799AFFE8797B15AF26A6D35BFD5D" box="[473,543,724,750]" captionStart="FIGURE 1" captionStartId="3.[264,367,1641,1665]" captionTargetBox="[278,1250,596,1621]" captionTargetId="figure@3.[264,1323,576,1634]" captionTargetPageId="3" captionText="FIGURE 1. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30: a, paratype male 7.6 mm (USNM 1069118); b – f, i, j, holotype male 8.2 mm (NSMT­Cr 16121); g, h, paratype male 8.5 mm, sta MM­ 30 (USNM 1069118). a, gill lamella; b, shield and cephalic appendages, dorsal view; c, portion of dorsal surface of palm of right cheliped near base of fixed finger and lateral margin, dorsolateral view; d, anterior lobe and posterior portion of thoracic sternite XII (third pereopods), ventral view; e – h, telson in dorsal (e, g) and posterior (f, h) views; i, j, coxae of fifth pereopods and thoracic sternite XXIV in ventral (i) and posterior (j) views. Scale bars: a = 0.25 mm; b, d, i, j = 1.0 mm; c, e – h = 0.5 mm." httpUri="https://zenodo.org/record/170617/files/figure.png" pageId="8" pageNumber="9">
Fig. 
<date id="A70543DFFFE8797B166426A6D35BFD5D" box="[530,543,724,750]" pageId="8" pageNumber="9">1</date>
</figureCitation>
i, j) with short fringe of setae arising from or near posterior margins of gonopore openings; right coxa with slight papilla or very short (= length of coxa measured on ventral surface) sexual tube protruding from gonopore and directed posteriorly; left coxa at most with slight papilla protruding from gonopore; with unpaired pleopods 3–5.
</paragraph>
<paragraph id="D304651FFFE8797B154E27EED2B5FBCD" blockId="8.[264,1324,284,1870]" pageId="8" pageNumber="9">
<emphasis id="E1CFB90DFFE8797B154E27EED0AFFC06" box="[312,491,924,949]" italics="true" pageId="8" pageNumber="9">Live coloration</emphasis>
(
<figureCitation id="4B80799AFFE8797B158D27EED307FC05" box="[507,579,924,950]" captionStart="FIGURE 5" captionStartId="7.[264,368,1821,1845]" captionTargetBox="[531,1056,1318,1797]" captionTargetId="figure@7.[531,1056,1318,1798]" captionTargetPageId="7" captionText="FIGURE 5. Pagurus ikedai n. sp., off Jogashima, Japan, live coloration, male 9.8 mm. (Photo: H. Ikeda)." httpUri="https://zenodo.org/record/170621/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
). Shield orange with some dark orange spots. Ocular peduncles white basally, then orange gradually fading towards corneas. Antennules and antennas light orange; antennal flagella mostly white. Posterior carapace pink, mottled with reddish. Chelipeds whitish, mottled with light orange. Second and third pereopods whitish except for distal and proximal orange bands on dactyl, proximal orange band on each propodus and carpus, and proximal and subdistal orange bands on merus.
</paragraph>
</subSubSection>
<subSubSection id="9BA13694FFE8797B154E20FED546FAAE" pageId="8" pageNumber="9" type="distribution">
<paragraph id="D304651FFFE8797B154E20FED335FB7D" blockId="8.[264,1324,284,1870]" pageId="8" pageNumber="9">
<emphasis id="E1CFB90DFFE8797B154E20FED083FB16" box="[312,455,1164,1189]" italics="true" pageId="8" pageNumber="9">Distribution</emphasis>
. Presently known only from the Tokyo Submarine Canyon, and off Jogashima, 
<collectingCountry id="ABAC258FFFE8797B15E520C6D09EFB7D" box="[403,474,1204,1230]" name="Japan" pageId="8" pageNumber="9">Japan</collectingCountry>
; 
<quantity id="1443C8FAFFE8797B159320C6D329FB7D" box="[485,621,1204,1230]" metricMagnitude="2" metricUnit="m" metricValue="3.45" metricValueMax="4.5" metricValueMin="2.4" pageId="8" pageNumber="9" unit="m" value="345.0" valueMax="450.0" valueMin="240.0">
240– 
<specimenCount id="C5BDAE96FFE8797B166920C6D329FB7D" box="[543,621,1204,1230]" pageId="8" pageNumber="9" type="generic">450 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="D304651FFFE8797B154E20AED546FAAE" blockId="8.[264,1324,284,1870]" pageId="8" pageNumber="9">
<emphasis id="E1CFB90DFFE8797B154E20AED0D5FB46" box="[312,401,1244,1269]" italics="true" pageId="8" pageNumber="9">Habitat</emphasis>
. Gastropod shells with relatively large aperture: 
<taxonomicName id="14BB1E9CFFE8797B17B920AED462FB45" authority="Watson" authorityName="Watson" box="[975,1318,1244,1270]" class="Gastropoda" family="Calliotropidae" genus="Bathybembix" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="aeola">
<emphasis id="E1CFB90DFFE8797B17B920AED5F5FB46" box="[975,1201,1244,1269]" italics="true" pageId="8" pageNumber="9">Bathybembix aeola</emphasis>
(Watson)
</taxonomicName>
, 
<taxonomicName id="14BB1E9CFFE8797B157E2176D335FAAD" authority="Lischke" authorityName="Lischke" box="[264,625,1284,1310]" class="Gastropoda" family="Buccinidae" genus="Buccinum" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="leucostoma">
<emphasis id="E1CFB90DFFE8797B157E2176D34DFAAE" box="[264,521,1284,1309]" italics="true" pageId="8" pageNumber="9">Buccinum leucostoma</emphasis>
Lischke
</taxonomicName>
, 
<taxonomicName id="14BB1E9CFFE8797B160B2176D2D8FAAD" authority="Okutani" authorityName="Okutani" box="[637,924,1284,1310]" class="Gastropoda" family="Buccinidae" genus="Buccinum" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="sagamianum">
<emphasis id="E1CFB90DFFE8797B160B2176D277FAAE" box="[637,819,1284,1309]" italics="true" pageId="8" pageNumber="9">B. sagamianum</emphasis>
Okutani
</taxonomicName>
, 
<taxonomicName id="14BB1E9CFFE8797B17D12176D546FAAE" box="[935,1026,1284,1309]" class="Gastropoda" family="Calliotropidae" genus="Ginebis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="E1CFB90DFFE8797B17D12176D546FAAE" box="[935,1026,1284,1309]" italics="true" pageId="8" pageNumber="9">Ginebis</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="9BA13694FFE8797B157E215ED3BCFAF5" box="[264,760,1324,1350]" pageId="8" pageNumber="9" type="description">
<paragraph id="D304651FFFE8797B157E215ED3BCFAF5" blockId="8.[264,1324,284,1870]" box="[264,760,1324,1350]" pageId="8" pageNumber="9">
<emphasis id="E1CFB90DFFE8797B157E215ED027FAF6" box="[264,355,1324,1349]" italics="true" pageId="8" pageNumber="9">crumpii</emphasis>
(Pilsbry), and 
<taxonomicName id="14BB1E9CFFE8797B1667215ED3B0FAF5" authority="Dall" authorityName="Dall" box="[529,756,1324,1350]" class="Gastropoda" family="Calliotropidae" genus="Ginebis" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="japonicus">
<emphasis id="E1CFB90DFFE8797B1667215ED3E7FAF6" box="[529,675,1324,1349]" italics="true" pageId="8" pageNumber="9">G. japonicus</emphasis>
(Dall)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="9BA13694FFE8797B154E2126D5CAF985" pageId="8" pageNumber="9" type="etymology">
<paragraph id="D304651FFFE8797B154E2126D353F9BD" blockId="8.[264,1324,284,1870]" pageId="8" pageNumber="9">
<emphasis id="E1CFB90DFFE8797B154E2126D0F7FADE" box="[312,435,1364,1389]" italics="true" pageId="8" pageNumber="9">Etymology</emphasis>
. The name of this new species is dedicated to Hitoshi Ikeda (Hayama Shiosai Museum), who first collected and photographed specimens. The name also is in recognition of his intensive, long­term research on Japanese deep­sea decapods and mollusks, inspired by the earlier work of 
<collectingCountry id="ABAC258FFFE8797B160621BED38AFA55" box="[624,718,1484,1510]" name="Germany" pageId="8" pageNumber="9">German</collectingCountry>
zoologists such as F. Doflein, and H. Balss, as well as the Showa Emperor.
</paragraph>
<paragraph id="D304651FFFE8797B154E226ED5CAF985" blockId="8.[264,1324,284,1870]" box="[312,1166,1564,1590]" pageId="8" pageNumber="9">
<emphasis id="E1CFB90DFFE8797B154E226ED0ACF986" box="[312,488,1564,1589]" italics="true" pageId="8" pageNumber="9">Common name</emphasis>
. “Ikeda­hon­yadokari” or “Ikeda’s 
<taxonomicName id="14BB1E9CFFE8797B17FE226ED2AFF986" box="[904,1003,1564,1589]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="E1CFB90DFFE8797B17FE226ED2AFF986" box="[904,1003,1564,1589]" italics="true" pageId="8" pageNumber="9">Pagurus</emphasis>
</taxonomicName>
hermit crab”.
</paragraph>
</subSubSection>
<subSubSection id="9BA13694FFE8797A154E2236D0C0FA25" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="discussion">
<paragraph id="D304651FFFE8797A154E2236D271FEED" blockId="8.[264,1324,284,1870]" lastBlockId="9.[264,1323,284,1430]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="E1CFB90DFFE8797B154E2236D0DAF9EE" box="[312,414,1604,1629]" italics="true" pageId="8" pageNumber="9">Remarks</emphasis>
. A good number of additional specimens of 
<taxonomicName id="14BB1E9CFFE8797B17CF2236D52AF9EE" box="[953,1134,1604,1629]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ikedai" status="sp. nov.">
<emphasis id="E1CFB90DFFE8797B17CF2236D52AF9EE" box="[953,1134,1604,1629]" italics="true" pageId="8" pageNumber="9">Pagurus ikedai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAFC0476FFE8797B10012236D5F3F9ED" box="[1143,1207,1604,1630]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
collected by H. Ikeda and one of us (HW) from off Jogashima, 
<collectingCountry id="ABAC258FFFE8797B17E5221ED29DF935" box="[915,985,1644,1670]" name="Japan" pageId="8" pageNumber="9">Japan</collectingCountry>
, were sent to Dr. S. Miyake many years prior to commencement of the present study. Unfortunately, Miyake’s material could not located. This includes the male (
<quantity id="1443C8FAFFE8797B177422CED224F965" box="[770,864,1724,1750]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="8" pageNumber="9" unit="mm" value="9.8">9.8 mm</quantity>
) shown in 
<figureCitation id="4B80799AFFE8797B179322CED568F965" box="[997,1068,1724,1750]" captionStart="FIGURE 5" captionStartId="7.[264,368,1821,1845]" captionTargetBox="[531,1056,1318,1797]" captionTargetId="figure@7.[531,1056,1318,1798]" captionTargetPageId="7" captionText="FIGURE 5. Pagurus ikedai n. sp., off Jogashima, Japan, live coloration, male 9.8 mm. (Photo: H. Ikeda)." httpUri="https://zenodo.org/record/170621/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
, from off Jogashima, 
<collectingCountry id="ABAC258FFFE8797B157E2296D00AF94D" box="[264,334,1764,1790]" name="Japan" pageId="8" pageNumber="9">Japan</collectingCountry>
, 
<quantity id="1443C8FAFFE8797B152D2296D0A0F94D" box="[347,484,1764,1790]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="8" pageNumber="9" unit="m" value="300.0" valueMax="320.0" valueMin="280.0">280–320 m</quantity>
, collected in 1980. The remaining specimens that could not be located have the following data: 20+ specimens from South of Jogashima, fishing grounds (Heidashi to Iwadogake by way of Shimashita), 
<quantity id="1443C8FAFFE8797B17312346D295F8FD" box="[839,977,1844,1870]" metricMagnitude="2" metricUnit="m" metricValue="3.1" metricValueMax="3.4" metricValueMin="2.8" pageId="8" pageNumber="9" unit="m" value="310.0" valueMax="340.0" valueMin="280.0">280–340 m</quantity>
, sand to rock, coll. H. Ikeda; and several specimens observed in faunal investigation by HW from top of Misaki Knoll, southwest of Jogashima, 
<quantity id="1443C8FAFFE9797A16592536D3F3FEED" box="[559,695,324,350]" metricMagnitude="2" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" pageId="9" pageNumber="10" unit="m" value="435.0" valueMax="450.0" valueMin="420.0">420–450 m</quantity>
, fine sand.
</paragraph>
<paragraph id="D304651FFFE9797A154E251ED36FFCD5" blockId="9.[264,1323,284,1430]" pageId="9" pageNumber="10">
As previously mentioned, 
<taxonomicName id="14BB1E9CFFE9797A161B251ED25BFE36" box="[621,799,364,389]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ikedai" status="sp. nov.">
<emphasis id="E1CFB90DFFE9797A161B251ED25BFE36" box="[621,799,364,389]" italics="true" pageId="9" pageNumber="10">Pagurus ikedai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAFC0476FFE9797A1751251ED222FE35" box="[807,870,364,390]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
is most closely allied to species of the 
<taxonomicName id="14BB1E9CFFE9797A157E25E6D0CAFE1E" box="[264,398,404,429]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bernhardus">
<emphasis id="E1CFB90DFFE9797A157E25E6D0CAFE1E" box="[264,398,404,429]" italics="true" pageId="9" pageNumber="10">bernhardus</emphasis>
</taxonomicName>
group of 
<taxonomicName id="14BB1E9CFFE9797A167425E6D321FE1E" box="[514,613,404,429]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="E1CFB90DFFE9797A167425E6D321FE1E" box="[514,613,404,429]" italics="true" pageId="9" pageNumber="10">Pagurus</emphasis>
</taxonomicName>
, and except for the lack of a spine in the new species on the dorsodistal margin of the merus of the third maxilliped, fits the definition provided by 
<bibRefCitation id="B72A18EEFFE9797A157E2596D0B6FE4D" author="McLaughlin" box="[264,498,484,510]" pageId="9" pageNumber="11" refString="McLaughlin, P. A. (1974) The hermit crabs (Crustacea Decapoda, Paguroidea) of northwestern North America. Zoologische Verhandelingen, 130, 1 - 396, pl. 1." type="journal article" year="1974">McLaughlin (1974)</bibRefCitation>
. As in males of 
<taxonomicName id="14BB1E9CFFE9797A16C62596D250FE4E" box="[688,788,484,509]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ikedai">
<emphasis id="E1CFB90DFFE9797A16C62596D250FE4E" box="[688,788,484,509]" italics="true" pageId="9" pageNumber="10">P. ikedai</emphasis>
</taxonomicName>
, males of three of the species in the 
<taxonomicName id="14BB1E9CFFE9797A10B42596D074FD96" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bernhardus">
<emphasis id="E1CFB90DFFE9797A10B42596D074FD96" italics="true" pageId="9" pageNumber="10">bernhardus</emphasis>
</taxonomicName>
group, 
<taxonomicName id="14BB1E9CFFE9797A15E4267ED37DFD96" box="[402,569,524,549]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bernhardus">
<emphasis id="E1CFB90DFFE9797A15E4267ED37DFD96" box="[402,569,524,549]" italics="true" pageId="9" pageNumber="10">P. bernhardus</emphasis>
</taxonomicName>
, 
<taxonomicName id="14BB1E9CFFE9797A163D267ED3A4FD96" box="[587,736,524,549]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="E1CFB90DFFE9797A163D267ED3A4FD96" box="[587,736,524,549]" italics="true" pageId="9" pageNumber="10">P. gracilipes</emphasis>
</taxonomicName>
, and 
<taxonomicName id="14BB1E9CFFE9797A1751267ED290FD96" box="[807,980,524,549]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="nipponensis">
<emphasis id="E1CFB90DFFE9797A1751267ED290FD96" box="[807,980,524,549]" italics="true" pageId="9" pageNumber="10">P. nipponensis</emphasis>
</taxonomicName>
, can have very short sexual tubes. However, 
<taxonomicName id="14BB1E9CFFE9797A15AF2646D3D5FDFE" box="[473,657,564,589]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ikedai">
<emphasis id="E1CFB90DFFE9797A15AF2646D3D5FDFE" box="[473,657,564,589]" italics="true" pageId="9" pageNumber="10">Pagurus ikedai</emphasis>
</taxonomicName>
can readily be distinguished from those three by the shape, armature and massiveness of both chelipeds. In 
<taxonomicName id="14BB1E9CFFE9797A17DC262ED557FDC6" box="[938,1043,604,629]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ikedai">
<emphasis id="E1CFB90DFFE9797A17DC262ED557FDC6" box="[938,1043,604,629]" italics="true" pageId="9" pageNumber="10">P. ikedai</emphasis>
</taxonomicName>
, the dorsal surfaces of carpi and chelae are covered with numerous calcareous spines or tubercles that terminate in short corneous bristles (
<figureCitation id="4B80799AFFE9797A163626DED3C7FD75" box="[576,643,684,710]" captionStart="FIGURE 1" captionStartId="3.[264,367,1641,1665]" captionTargetBox="[278,1250,596,1621]" captionTargetId="figure@3.[264,1323,576,1634]" captionTargetPageId="3" captionText="FIGURE 1. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30: a, paratype male 7.6 mm (USNM 1069118); b – f, i, j, holotype male 8.2 mm (NSMT­Cr 16121); g, h, paratype male 8.5 mm, sta MM­ 30 (USNM 1069118). a, gill lamella; b, shield and cephalic appendages, dorsal view; c, portion of dorsal surface of palm of right cheliped near base of fixed finger and lateral margin, dorsolateral view; d, anterior lobe and posterior portion of thoracic sternite XII (third pereopods), ventral view; e – h, telson in dorsal (e, g) and posterior (f, h) views; i, j, coxae of fifth pereopods and thoracic sternite XXIV in ventral (i) and posterior (j) views. Scale bars: a = 0.25 mm; b, d, i, j = 1.0 mm; c, e – h = 0.5 mm." httpUri="https://zenodo.org/record/170617/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
c). The chelae are dorsoventrally flattened, more so laterally, and the dorsolateral and dorsomesial margins of palms and fingers are sharply defined and finely serrate. The left cheliped is very similar to the right in shape and armature, and although distinctly smaller than the right, is much larger relative to that of other congeners in the 
<taxonomicName id="14BB1E9CFFE9797A1527273ED093FCD6" box="[337,471,844,869]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bernhardus">
<emphasis id="E1CFB90DFFE9797A1527273ED093FCD6" box="[337,471,844,869]" italics="true" pageId="9" pageNumber="10">bernhardus</emphasis>
</taxonomicName>
group.
</paragraph>
<paragraph id="D304651FFFE9797A154E2706D0C0FA25" blockId="9.[264,1323,284,1430]" pageId="9" pageNumber="10">
The specimens examined of 
<taxonomicName id="14BB1E9CFFE9797A16E62706D201FC3E" box="[656,837,884,909]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ikedai" status="sp. nov.">
<emphasis id="E1CFB90DFFE9797A16E62706D201FC3E" box="[656,837,884,909]" italics="true" pageId="9" pageNumber="10">Pagurus ikedai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAFC0476FFE9797A17382706D2CBFC3D" box="[846,911,884,910]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
are quite constant in morphology, showing only slight but expected variations overall, such as in the number of spines on the dorsal (18–26), ventral (14–22) and ventromesial margins of the dactyls of the second and third pereopods, and the telson (
<figureCitation id="4B80799AFFE9797A16F5279ED381FBB5" box="[643,709,1004,1030]" captionStart="FIGURE 1" captionStartId="3.[264,367,1641,1665]" captionTargetBox="[278,1250,596,1621]" captionTargetId="figure@3.[264,1323,576,1634]" captionTargetPageId="3" captionText="FIGURE 1. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30: a, paratype male 7.6 mm (USNM 1069118); b – f, i, j, holotype male 8.2 mm (NSMT­Cr 16121); g, h, paratype male 8.5 mm, sta MM­ 30 (USNM 1069118). a, gill lamella; b, shield and cephalic appendages, dorsal view; c, portion of dorsal surface of palm of right cheliped near base of fixed finger and lateral margin, dorsolateral view; d, anterior lobe and posterior portion of thoracic sternite XII (third pereopods), ventral view; e – h, telson in dorsal (e, g) and posterior (f, h) views; i, j, coxae of fifth pereopods and thoracic sternite XXIV in ventral (i) and posterior (j) views. Scale bars: a = 0.25 mm; b, d, i, j = 1.0 mm; c, e – h = 0.5 mm." httpUri="https://zenodo.org/record/170617/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
e–h). No significant variations due to sexual dimorphism were detected. Although the overall size range (shield length 
<quantity id="1443C8FAFFE9797A10582066D593FB9D" box="[1070,1239,1044,1070]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" metricValueMax="9.4" metricValueMin="7.2" pageId="9" pageNumber="10" unit="mm" value="8.3" valueMax="9.4" valueMin="7.2">7.2 to 9.4 mm</quantity>
) of the specimens examined was relatively narrow, no significant allometric variation was observed either. In some males, the coxa of the right fifth pereopods has a slight papilla protruding from the gonopore, whereas in others there is a very short sexual tube (
<figureCitation id="4B80799AFFE9797A10A320FED45EFB15" box="[1237,1306,1164,1190]" captionStart="FIGURE 1" captionStartId="3.[264,367,1641,1665]" captionTargetBox="[278,1250,596,1621]" captionTargetId="figure@3.[264,1323,576,1634]" captionTargetPageId="3" captionText="FIGURE 1. Pagurus ikedai n. sp., Tokyo Submarine Canyon, Japan, sta MM­ 30: a, paratype male 7.6 mm (USNM 1069118); b – f, i, j, holotype male 8.2 mm (NSMT­Cr 16121); g, h, paratype male 8.5 mm, sta MM­ 30 (USNM 1069118). a, gill lamella; b, shield and cephalic appendages, dorsal view; c, portion of dorsal surface of palm of right cheliped near base of fixed finger and lateral margin, dorsolateral view; d, anterior lobe and posterior portion of thoracic sternite XII (third pereopods), ventral view; e – h, telson in dorsal (e, g) and posterior (f, h) views; i, j, coxae of fifth pereopods and thoracic sternite XXIV in ventral (i) and posterior (j) views. Scale bars: a = 0.25 mm; b, d, i, j = 1.0 mm; c, e – h = 0.5 mm." httpUri="https://zenodo.org/record/170617/files/figure.png" pageId="9" pageNumber="10">
Fig. 
<date id="A70543DFFFE9797A117B20FED45EFB15" box="[1293,1306,1164,1190]" pageId="9" pageNumber="10">1</date>
</figureCitation>
i, j); the coxa of the left fifth pereopod invariably has a slight papilla protruding from the gonopore. The males of at least three other species of 
<taxonomicName id="14BB1E9CFFE9797A17D720AED541FB46" box="[929,1029,1244,1269]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="E1CFB90DFFE9797A17D720AED541FB46" box="[929,1029,1244,1269]" italics="true" pageId="9" pageNumber="10">Pagurus</emphasis>
</taxonomicName>
are also known to have short sexual tubes: 
<taxonomicName id="14BB1E9CFFE9797A159D2176D200FAAD" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[491,836,1284,1310]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="constans">
<emphasis id="E1CFB90DFFE9797A159D2176D32AFAAE" box="[491,622,1284,1309]" italics="true" pageId="9" pageNumber="10">P. constans</emphasis>
(
<bibRefCitation id="B72A18EEFFE9797A16082176D27EFAAD" author="Stimpson" box="[638,826,1284,1310]" pageId="9" pageNumber="12" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers Ducibus, observavit et descripsi t. W. Stimpson. Pars VII. Crustacea Anomoura. [Preprint (December 1858) from] Proceedings of the Academy of Natural Sciences of Philadelphia, 10, 225 - 252." type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
)
</taxonomicName>
, 
<emphasis id="E1CFB90DFFE9797A17262176D2F2FAAE" box="[848,950,1284,1309]" italics="true" pageId="9" pageNumber="10">P. h a r t a e</emphasis>
(
<bibRefCitation id="B72A18EEFFE9797A17BE2176D45AFAAD" author="McLaughlin" box="[968,1310,1284,1310]" pageId="9" pageNumber="11" refString="McLaughlin, P. A. &amp; Jensen, G. C. (1996) A new species of hermit crab of the genus Parapagurodes (Decapoda: Anomura: Paguridae) from the eastern Pacific, with a description of its first zoeal stage. Journal of Natural History, 30, 841 - 854." type="journal article" year="1996">McLaughlin &amp; Jensen, 1996</bibRefCitation>
), and 
<emphasis id="E1CFB90DFFE9797A154A215ED0D7FAF6" box="[316,403,1324,1349]" italics="true" pageId="9" pageNumber="10">P. i m a i i</emphasis>
(
<bibRefCitation id="B72A18EEFFE9797A15DE215ED310FAF5" author="Yokoya" box="[424,596,1324,1350]" pageId="9" pageNumber="12" refString="Yokoya, Y. (1939) Macrura and Anomura of decapod Crustacea found in the neighborhood of Onagawa, Miyagi-ken. Scientific Reports of Tohoku University, 14, 261 - 289." type="journal article" year="1939">Yokoya, 1939</bibRefCitation>
); however, when other characters are considered, these three species do not appear to be closely related to 
<taxonomicName id="14BB1E9CFFE9797A17692126D2C0FADE" box="[799,900,1364,1389]" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ikedai">
<emphasis id="E1CFB90DFFE9797A17692126D2C0FADE" box="[799,900,1364,1389]" italics="true" pageId="9" pageNumber="10">P. ikedai</emphasis>
</taxonomicName>
or any of the others in the 
<taxonomicName id="14BB1E9CFFE9797A10B42126D074FA26" class="Malacostraca" family="Paguridae" genus="Pagurus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bernhardus">
<emphasis id="E1CFB90DFFE9797A10B42126D074FA26" italics="true" pageId="9" pageNumber="10">bernhardus</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
</treatment>
</document>